taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
45227F36FFB8FFB5B5BA2709FB8EF8FA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171514/files/figure.png	https://doi.org/10.5281/zenodo.19171514	FIGURES 1–4. Type specimens of Chalcoderus and associated labels (1–3). 1. Lectotype of Ateuchus maculatus Fabricius, 1801 (ZMUC). 2. Holotype of Onthophagus chloroderus d’Orbigny, 1905 (MNHN). 3. Neotype ♂ of Chalcoderus signatipennis (de Castelnau, 1840) (MNHN). 4. Distribution of Chalcoderus maculatus (Fabricius, 1801) (varieties with red or green/blue pronotum) and C. signatipennis (de Castelnau, 1840) in West Africa relative to political borders and ecoregions of Olson et al. (2001) (ranges centred mainly on: Western Sudanian Savanna (orange) and Guinean Forest-Savanna Mosaic (light green).	FIGURES 1–4. Type specimens of Chalcoderus and associated labels (1–3). 1. Lectotype of Ateuchus maculatus Fabricius, 1801 (ZMUC). 2. Holotype of Onthophagus chloroderus d’Orbigny, 1905 (MNHN). 3. Neotype ♂ of Chalcoderus signatipennis (de Castelnau, 1840) (MNHN). 4. Distribution of Chalcoderus maculatus (Fabricius, 1801) (varieties with red or green/blue pronotum) and C. signatipennis (de Castelnau, 1840) in West Africa relative to political borders and ecoregions of Olson et al. (2001) (ranges centred mainly on: Western Sudanian Savanna (orange) and Guinean Forest-Savanna Mosaic (light green).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFBCFFB1B5BA2630FE5BFCC7.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171524/files/figure.png	https://doi.org/10.5281/zenodo.19171524	FIGURES 11–15. Type specimens of Chalcoderus and Onthophagus with associated labels. 11. Holotype of Chalcoderus planaticeps (d’Orbigny, 1913) (IRSNB).12. Lectotype of Chalcoderus semicroceus (d’Orbigny, 1915) (MRAC).13. Holotype of Chalcoderus simpliciceps (d’Orbigny, 1905) (ZMHB). 14. Paralectotype of Onthophagus coeruleicollis Arrow, 1907 (NHML). 15. Paralectotype of Onthophagus regalis Arrow, 1907 (MNHN).	FIGURES 11–15. Type specimens of Chalcoderus and Onthophagus with associated labels. 11. Holotype of Chalcoderus planaticeps (d’Orbigny, 1913) (IRSNB).12. Lectotype of Chalcoderus semicroceus (d’Orbigny, 1915) (MRAC).13. Holotype of Chalcoderus simpliciceps (d’Orbigny, 1905) (ZMHB). 14. Paralectotype of Onthophagus coeruleicollis Arrow, 1907 (NHML). 15. Paralectotype of Onthophagus regalis Arrow, 1907 (MNHN).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFBDFFBFB5BA207CFDA6FEFB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171514/files/figure.png	https://doi.org/10.5281/zenodo.19171514	FIGURES 1–4. Type specimens of Chalcoderus and associated labels (1–3). 1. Lectotype of Ateuchus maculatus Fabricius, 1801 (ZMUC). 2. Holotype of Onthophagus chloroderus d’Orbigny, 1905 (MNHN). 3. Neotype ♂ of Chalcoderus signatipennis (de Castelnau, 1840) (MNHN). 4. Distribution of Chalcoderus maculatus (Fabricius, 1801) (varieties with red or green/blue pronotum) and C. signatipennis (de Castelnau, 1840) in West Africa relative to political borders and ecoregions of Olson et al. (2001) (ranges centred mainly on: Western Sudanian Savanna (orange) and Guinean Forest-Savanna Mosaic (light green).	FIGURES 1–4. Type specimens of Chalcoderus and associated labels (1–3). 1. Lectotype of Ateuchus maculatus Fabricius, 1801 (ZMUC). 2. Holotype of Onthophagus chloroderus d’Orbigny, 1905 (MNHN). 3. Neotype ♂ of Chalcoderus signatipennis (de Castelnau, 1840) (MNHN). 4. Distribution of Chalcoderus maculatus (Fabricius, 1801) (varieties with red or green/blue pronotum) and C. signatipennis (de Castelnau, 1840) in West Africa relative to political borders and ecoregions of Olson et al. (2001) (ranges centred mainly on: Western Sudanian Savanna (orange) and Guinean Forest-Savanna Mosaic (light green).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFB3FFBFB5BA25F2FE3CFC63.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171514/files/figure.png	https://doi.org/10.5281/zenodo.19171514	FIGURES 1–4. Type specimens of Chalcoderus and associated labels (1–3). 1. Lectotype of Ateuchus maculatus Fabricius, 1801 (ZMUC). 2. Holotype of Onthophagus chloroderus d’Orbigny, 1905 (MNHN). 3. Neotype ♂ of Chalcoderus signatipennis (de Castelnau, 1840) (MNHN). 4. Distribution of Chalcoderus maculatus (Fabricius, 1801) (varieties with red or green/blue pronotum) and C. signatipennis (de Castelnau, 1840) in West Africa relative to political borders and ecoregions of Olson et al. (2001) (ranges centred mainly on: Western Sudanian Savanna (orange) and Guinean Forest-Savanna Mosaic (light green).	FIGURES 1–4. Type specimens of Chalcoderus and associated labels (1–3). 1. Lectotype of Ateuchus maculatus Fabricius, 1801 (ZMUC). 2. Holotype of Onthophagus chloroderus d’Orbigny, 1905 (MNHN). 3. Neotype ♂ of Chalcoderus signatipennis (de Castelnau, 1840) (MNHN). 4. Distribution of Chalcoderus maculatus (Fabricius, 1801) (varieties with red or green/blue pronotum) and C. signatipennis (de Castelnau, 1840) in West Africa relative to political borders and ecoregions of Olson et al. (2001) (ranges centred mainly on: Western Sudanian Savanna (orange) and Guinean Forest-Savanna Mosaic (light green).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFB0FFBDB5BA224CFCA5FB4A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171514/files/figure.png	https://doi.org/10.5281/zenodo.19171514	FIGURES 1–4. Type specimens of Chalcoderus and associated labels (1–3). 1. Lectotype of Ateuchus maculatus Fabricius, 1801 (ZMUC). 2. Holotype of Onthophagus chloroderus d’Orbigny, 1905 (MNHN). 3. Neotype ♂ of Chalcoderus signatipennis (de Castelnau, 1840) (MNHN). 4. Distribution of Chalcoderus maculatus (Fabricius, 1801) (varieties with red or green/blue pronotum) and C. signatipennis (de Castelnau, 1840) in West Africa relative to political borders and ecoregions of Olson et al. (2001) (ranges centred mainly on: Western Sudanian Savanna (orange) and Guinean Forest-Savanna Mosaic (light green).	FIGURES 1–4. Type specimens of Chalcoderus and associated labels (1–3). 1. Lectotype of Ateuchus maculatus Fabricius, 1801 (ZMUC). 2. Holotype of Onthophagus chloroderus d’Orbigny, 1905 (MNHN). 3. Neotype ♂ of Chalcoderus signatipennis (de Castelnau, 1840) (MNHN). 4. Distribution of Chalcoderus maculatus (Fabricius, 1801) (varieties with red or green/blue pronotum) and C. signatipennis (de Castelnau, 1840) in West Africa relative to political borders and ecoregions of Olson et al. (2001) (ranges centred mainly on: Western Sudanian Savanna (orange) and Guinean Forest-Savanna Mosaic (light green).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFB1FFBDB5BA21D9FCEAF801.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171518/files/figure.png	https://doi.org/10.5281/zenodo.19171518	FIGURES 6–8. Type specimens of Chalcoderus and associated labels. 6. Lectotype ♂ of Chalcoderus longiceps (d’Orbigny, 1904) (MSNG). 7. Holotype of Chalcoderus foliiceps (Quedenfeldt, 1884) (MNHN). 8. Holotype of Chalcoderus graphicus (Wallengren, 1881) (MZLU).	FIGURES 6–8. Type specimens of Chalcoderus and associated labels. 6. Lectotype ♂ of Chalcoderus longiceps (d’Orbigny, 1904) (MSNG). 7. Holotype of Chalcoderus foliiceps (Quedenfeldt, 1884) (MNHN). 8. Holotype of Chalcoderus graphicus (Wallengren, 1881) (MZLU).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFB6FFBAB5BA25C1FDACFC89.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171518/files/figure.png	https://doi.org/10.5281/zenodo.19171518	FIGURES 6–8. Type specimens of Chalcoderus and associated labels. 6. Lectotype ♂ of Chalcoderus longiceps (d’Orbigny, 1904) (MSNG). 7. Holotype of Chalcoderus foliiceps (Quedenfeldt, 1884) (MNHN). 8. Holotype of Chalcoderus graphicus (Wallengren, 1881) (MZLU).	FIGURES 6–8. Type specimens of Chalcoderus and associated labels. 6. Lectotype ♂ of Chalcoderus longiceps (d’Orbigny, 1904) (MSNG). 7. Holotype of Chalcoderus foliiceps (Quedenfeldt, 1884) (MNHN). 8. Holotype of Chalcoderus graphicus (Wallengren, 1881) (MZLU).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFB7FFBBB5BA24E0FE5EFD82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171518/files/figure.png	https://doi.org/10.5281/zenodo.19171518	FIGURES 6–8. Type specimens of Chalcoderus and associated labels. 6. Lectotype ♂ of Chalcoderus longiceps (d’Orbigny, 1904) (MSNG). 7. Holotype of Chalcoderus foliiceps (Quedenfeldt, 1884) (MNHN). 8. Holotype of Chalcoderus graphicus (Wallengren, 1881) (MZLU).	FIGURES 6–8. Type specimens of Chalcoderus and associated labels. 6. Lectotype ♂ of Chalcoderus longiceps (d’Orbigny, 1904) (MSNG). 7. Holotype of Chalcoderus foliiceps (Quedenfeldt, 1884) (MNHN). 8. Holotype of Chalcoderus graphicus (Wallengren, 1881) (MZLU).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFB7FFBBB5BA26FEFC8CFAD3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171522/files/figure.png	https://doi.org/10.5281/zenodo.19171522	FIGURES 9–10. Type specimens of Chalcoderus and associated labels. 9. One of three syntypes of Chalcoderus imbellis (d’Orbigny, 1905) (MSNG). 10. Holotype of Chalcoderus inermiceps (d’Orbigny, 1905a) (NHML).	FIGURES 9–10. Type specimens of Chalcoderus and associated labels. 9. One of three syntypes of Chalcoderus imbellis (d’Orbigny, 1905) (MSNG). 10. Holotype of Chalcoderus inermiceps (d’Orbigny, 1905a) (NHML).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFB4FFB8B5BA24E0FBE9FDAE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171522/files/figure.png	https://doi.org/10.5281/zenodo.19171522	FIGURES 9–10. Type specimens of Chalcoderus and associated labels. 9. One of three syntypes of Chalcoderus imbellis (d’Orbigny, 1905) (MSNG). 10. Holotype of Chalcoderus inermiceps (d’Orbigny, 1905a) (NHML).	FIGURES 9–10. Type specimens of Chalcoderus and associated labels. 9. One of three syntypes of Chalcoderus imbellis (d’Orbigny, 1905) (MSNG). 10. Holotype of Chalcoderus inermiceps (d’Orbigny, 1905a) (NHML).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFB4FFB8B5BA2612FDC0FC59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171524/files/figure.png	https://doi.org/10.5281/zenodo.19171524	FIGURES 11–15. Type specimens of Chalcoderus and Onthophagus with associated labels. 11. Holotype of Chalcoderus planaticeps (d’Orbigny, 1913) (IRSNB).12. Lectotype of Chalcoderus semicroceus (d’Orbigny, 1915) (MRAC).13. Holotype of Chalcoderus simpliciceps (d’Orbigny, 1905) (ZMHB). 14. Paralectotype of Onthophagus coeruleicollis Arrow, 1907 (NHML). 15. Paralectotype of Onthophagus regalis Arrow, 1907 (MNHN).	FIGURES 11–15. Type specimens of Chalcoderus and Onthophagus with associated labels. 11. Holotype of Chalcoderus planaticeps (d’Orbigny, 1913) (IRSNB).12. Lectotype of Chalcoderus semicroceus (d’Orbigny, 1915) (MRAC).13. Holotype of Chalcoderus simpliciceps (d’Orbigny, 1905) (ZMHB). 14. Paralectotype of Onthophagus coeruleicollis Arrow, 1907 (NHML). 15. Paralectotype of Onthophagus regalis Arrow, 1907 (MNHN).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFB4FFB8B5BA2043FC7BFA6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171524/files/figure.png	https://doi.org/10.5281/zenodo.19171524	FIGURES 11–15. Type specimens of Chalcoderus and Onthophagus with associated labels. 11. Holotype of Chalcoderus planaticeps (d’Orbigny, 1913) (IRSNB).12. Lectotype of Chalcoderus semicroceus (d’Orbigny, 1915) (MRAC).13. Holotype of Chalcoderus simpliciceps (d’Orbigny, 1905) (ZMHB). 14. Paralectotype of Onthophagus coeruleicollis Arrow, 1907 (NHML). 15. Paralectotype of Onthophagus regalis Arrow, 1907 (MNHN).	FIGURES 11–15. Type specimens of Chalcoderus and Onthophagus with associated labels. 11. Holotype of Chalcoderus planaticeps (d’Orbigny, 1913) (IRSNB).12. Lectotype of Chalcoderus semicroceus (d’Orbigny, 1915) (MRAC).13. Holotype of Chalcoderus simpliciceps (d’Orbigny, 1905) (ZMHB). 14. Paralectotype of Onthophagus coeruleicollis Arrow, 1907 (NHML). 15. Paralectotype of Onthophagus regalis Arrow, 1907 (MNHN).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFB4FFA6B5BA2250FBF2FD3F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171524/files/figure.png	https://doi.org/10.5281/zenodo.19171524	FIGURES 11–15. Type specimens of Chalcoderus and Onthophagus with associated labels. 11. Holotype of Chalcoderus planaticeps (d’Orbigny, 1913) (IRSNB).12. Lectotype of Chalcoderus semicroceus (d’Orbigny, 1915) (MRAC).13. Holotype of Chalcoderus simpliciceps (d’Orbigny, 1905) (ZMHB). 14. Paralectotype of Onthophagus coeruleicollis Arrow, 1907 (NHML). 15. Paralectotype of Onthophagus regalis Arrow, 1907 (MNHN).	FIGURES 11–15. Type specimens of Chalcoderus and Onthophagus with associated labels. 11. Holotype of Chalcoderus planaticeps (d’Orbigny, 1913) (IRSNB).12. Lectotype of Chalcoderus semicroceus (d’Orbigny, 1915) (MRAC).13. Holotype of Chalcoderus simpliciceps (d’Orbigny, 1905) (ZMHB). 14. Paralectotype of Onthophagus coeruleicollis Arrow, 1907 (NHML). 15. Paralectotype of Onthophagus regalis Arrow, 1907 (MNHN).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFAAFFA6B5BA2769FD3AFB59.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171524/files/figure.png	https://doi.org/10.5281/zenodo.19171524	FIGURES 11–15. Type specimens of Chalcoderus and Onthophagus with associated labels. 11. Holotype of Chalcoderus planaticeps (d’Orbigny, 1913) (IRSNB).12. Lectotype of Chalcoderus semicroceus (d’Orbigny, 1915) (MRAC).13. Holotype of Chalcoderus simpliciceps (d’Orbigny, 1905) (ZMHB). 14. Paralectotype of Onthophagus coeruleicollis Arrow, 1907 (NHML). 15. Paralectotype of Onthophagus regalis Arrow, 1907 (MNHN).	FIGURES 11–15. Type specimens of Chalcoderus and Onthophagus with associated labels. 11. Holotype of Chalcoderus planaticeps (d’Orbigny, 1913) (IRSNB).12. Lectotype of Chalcoderus semicroceus (d’Orbigny, 1915) (MRAC).13. Holotype of Chalcoderus simpliciceps (d’Orbigny, 1905) (ZMHB). 14. Paralectotype of Onthophagus coeruleicollis Arrow, 1907 (NHML). 15. Paralectotype of Onthophagus regalis Arrow, 1907 (MNHN).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
45227F36FFAAFFA6B5BA2142FD9AF934.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19171524/files/figure.png	https://doi.org/10.5281/zenodo.19171524	FIGURES 11–15. Type specimens of Chalcoderus and Onthophagus with associated labels. 11. Holotype of Chalcoderus planaticeps (d’Orbigny, 1913) (IRSNB).12. Lectotype of Chalcoderus semicroceus (d’Orbigny, 1915) (MRAC).13. Holotype of Chalcoderus simpliciceps (d’Orbigny, 1905) (ZMHB). 14. Paralectotype of Onthophagus coeruleicollis Arrow, 1907 (NHML). 15. Paralectotype of Onthophagus regalis Arrow, 1907 (MNHN).	FIGURES 11–15. Type specimens of Chalcoderus and Onthophagus with associated labels. 11. Holotype of Chalcoderus planaticeps (d’Orbigny, 1913) (IRSNB).12. Lectotype of Chalcoderus semicroceus (d’Orbigny, 1915) (MRAC).13. Holotype of Chalcoderus simpliciceps (d’Orbigny, 1905) (ZMHB). 14. Paralectotype of Onthophagus coeruleicollis Arrow, 1907 (NHML). 15. Paralectotype of Onthophagus regalis Arrow, 1907 (MNHN).	2026-01-26	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.		Zenodo	biologists	Moretto, Philippe;Deschodt, Christian M.;Davis, Adrian L. V.			
