identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
461DA55ADA0FFFCC236FFBEDC326ECE2.text	461DA55ADA0FFFCC236FFBEDC326ECE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tomthompsonia Wagele & Hain 1991	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Systematics of  Tomthompsonia and  Pleurobranchaeidae</p>
            <p> Pleurobranchidaisrecoveredmonophyleticwithfullsupportand includes three main clades. For the first time in molecular phylogeny,  Tomthompsonia antarctica (Fig. 1A) forms a discrete, early divergent clade with maximum support across matrices and analyses. Thus, we erect  Tomthompsoniidae fam. nov. to encompass a single known genus and species endemic to deep waters in Antarctica (Thiele 1912, Wägele and Hain 1991, Hain et al. 1993). Then, two main clades sister to each other are recovered,  Pleurobranchaeidae and  Pleurobranchidae (bs = 88, pp = 0.95). Within  Pleurobranchaeidae , the type species  Euselenops luniceps (Fig. 1B) is first found as a sister to a clade composed of the type species of both  Pleurobranchaea and  Pleurobranchella . Among the four specimens of  Euselenops available, a conflicting signal is found in the 18S of the specimens sequenced by Wollscheid-Lengeling et al. (2001) and, unfortunately, their relationships are not always well-recovered (see Supporting Information, Figs S1, S2). Excluding these from our concatenated analyses rendered  Pleurobranchaeidae monophyletic without maximum support (bs = 91). COI p -distances for both remaining specimens of  E. luniceps from Singapore and Vietnam seem to represent cases of hidden speciation (86% identity). Considering both type taxa  Pleurobranchaea meckeli (Blainville , 1825) (Fig. 1C) and  Pleurobranchella nicobarica Thiele , 1925 (Fig. 1D) were present in our dataset, and  Pleurobranchella is placed within a clade composed of  P. meckeli ,  P. maculata (Quoy and Gaimard , 1832),  P. californica MacFarland , 1966, and  P. japonica Thiele , 1925 (bs = 100, pp = 1). However,  Pleurobranchella is morphologically distinct from  Pleurobranchaea , including such immediately recognizable features as an ample notum and differences in the reproductive system (Martynov and Schrödl 2009). We, therefore, keep the genus  Pleurobranchella distinct and valid. Finally,  P. cf. novaezealandiae from the Western Pacific clusters with our  P. japonica from Japan with a 99.1% identity, thus suggesting both specimens belong to  P. japonica . </p>
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	https://treatment.plazi.org/id/461DA55ADA0FFFCC236FFBEDC326ECE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Moles, Juan;Brenzinger, Bastian;Berning, Maria I.;Martynov, Alexander;TatianaKorshunova;Schrödl, Michael	Moles, Juan, Brenzinger, Bastian, Berning, Maria I., Martynov, Alexander, TatianaKorshunova, Schrödl, Michael (2024): Systematic rearrangements in an all-genus phylogeny of side-gilled slugs (Heterobranchia: Pleurobranchida). Zoological Journal of the Linnean Society 202 (2): 1-11, DOI: 10.1093/zoolinnean/zlad162, URL: https://doi.org/10.1093/zoolinnean/zlad162
461DA55ADA08FFCD213DFF18C264EA89.text	461DA55ADA08FFCD213DFF18C264EA89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleurobranchidae Gray 1827	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Systematics of  Pleurobranchidae</p>
            <p> Although not supported in the ML and BI phylogenies,  Pleurobranchidae is for the first time recovered as a clade in our analyses. All valid, representative genera within the family are for thefirsttimepresent, namely  Bathyberthella ,  Berthella ,  Berthellina ,  Boreoberthella ,  Pleurehdera , and  Pleurobranchus .  Berthella is consistently found paraphyletic and four clades classifiable as three different genera are recovered. Bearing that in mind, most genera are found monophyletic with maximum support, although how genera are related to each other is not always well supported.  Berthellina is recovered monophyletic with full support, the species  B. edwardsii (Vayssière , 1897) from the Mediterranean and Eastern Atlantic (94.3–99% identity) is the sister-group to the Indo-Pacific species. The latter group is composed of  B. citrina (Rüppell and Leuckart , 1828) from the Red Sea (100% identity),  B. punctata (Quoy and Gaimard , 1832) from Australia, and a clade of  B. delicata (Pease , 1861) species with probabl hidden speciation spanning from the Red Sea, the Maldives (Fig. 1E), and Hawaii (90.7–93.6% identity). All  Bathyberthella antarctica Willan and Bertsch 1987 (Fig. 1F) specimens from across the Weddell Sea and the Scotia Arc conformed to a monophyletic clade (bs = 100, pp = 1) with a 95–100% identity. </p>
            <p> A clade containing  Berthella perforata (Philippi , 1844) specimens from Italy (99.1–100% identity) and the type species of  Berthella ,  B. plumula (Montagu , 1803), with specimens from the Northeast Atlantic and the Mediterranean Sea (98.7–100% identity) was fully recovered only in the ML analysis (bs = 100). This clade also included  B. ocellata (Delle Chiaje , 1830) from Greece (Fig. 1G),  B. medietas Burn , 1962 from Australia,  B. sideralis Lovén , 1846 from Sweden, and  B. platei (Bergh , 1898) from 15 m depth Chile and 250 m depth in the Falkland Islands (99.3– 99.5% identity). A second ‘ Berthella’ clade (bs = 99, pp = 1) is largely composed of the  B. stellata (Risso , 1826) group agreeing to the recently recognized species in Ghanimi et al. (2020b) yet adding some more complexity. The  B. stellata (Fig. 1I) group is found in tropical and temperate waters from the Eastern Pacific, Caribbean, and Mediterranean seas, including  B. andromeda Ghanimi , Schrödl, Goddard et al., 2020,  B. strongi (MacFarland , 1966),  B. nebula Ghanimi et al. , 2020, and  B. vialactea Ghanimi et al. , 2020. This latter group is sister to a Pacific one composed of  B. pellucida (Pease , 1860) from Hawaii,  B. cf. postrema from New Caledonia, our newly sequenced B. cf.  ‘ stellata ’ from Japan, and  Pleurehdera haraldi Ev.Marcus and Er.Marcus , 1970 from Palmyra Atoll in the middle of the Pacific. Since  Pleurehdera haraldi is the type and only species of the genus and nests within this second ‘  Berthella ’ clade, we suggest transferring all the above-mentioned species to the genus  Pleurehdera . Interestingly, only in the BI analysis, a sister relationship between specimens belonging to  Berthella s.s. and the  Pleurehdera clade was recovered yet without full support (pp = 0.95). Sister to both  B. stellata and the Pacific group we found  B. martensi (Pilsbry , 1896), with specimens from Australia and the Pacific side of Panama (99.3% identity) and a likely hidden species from the Maldives (c. 85% identity) (Fig. 1H). This clade deserves the erection of a new genus (see Systematic description). A fourth ‘  Berthella ’ clade (bs = 92, pp = 0.99) is composed of the North Pacific clade that includes specimens of  Berthella californica (Dall , 1900) from California to Panama, two specimens of  Boreoberthella augusta Martynov and Schrödl 2009 (Fig. 1J) from Japan (100% identity), and  Berthella chacei (J.Q.Burch , 1944) from Russia, Canada, and the USA (99–99.7% identity). Given the present phylogenetic scenario and the morphological characters discussed below, we hence establish  Boreoberthella californica comb. nov. (Fig. 1K) and  Boreoberthella chacei comb. nov. Finally, we recovered a monophyletic clade of  Pleurobranchus species (bs = 100, pp = 1) (Fig. 1L), containing the type species  P. peronii Cuvier , 1804 and mainly recovering the species groups already studied by Goodheart et al. (2015). </p>
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	https://treatment.plazi.org/id/461DA55ADA08FFCD213DFF18C264EA89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Moles, Juan;Brenzinger, Bastian;Berning, Maria I.;Martynov, Alexander;TatianaKorshunova;Schrödl, Michael	Moles, Juan, Brenzinger, Bastian, Berning, Maria I., Martynov, Alexander, TatianaKorshunova, Schrödl, Michael (2024): Systematic rearrangements in an all-genus phylogeny of side-gilled slugs (Heterobranchia: Pleurobranchida). Zoological Journal of the Linnean Society 202 (2): 1-11, DOI: 10.1093/zoolinnean/zlad162, URL: https://doi.org/10.1093/zoolinnean/zlad162
461DA55ADA08FFCE23CEFBFBC42DE8BE.text	461DA55ADA08FFCE23CEFBFBC42DE8BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tomoberthella Moles & Brenzinger & Berning & Martynov & TatianaKorshunova & Schrödl 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tomoberthella gen. nov.</p>
            <p>(Fig. 1H)</p>
            <p>urn:lsid:zoobank.org:act: 8D6B1A52-6901-4EF6-853E-0A68B0859EB6.</p>
            <p> Type species:  Tomoberthella martensi (Pilsbry, 1896) comb. nov. </p>
            <p> Pleurobranchus scutatus E. von Martens, 1880 (  non-Pleurobranchus scutatus Forbes, 1844).</p>
            <p> Gymnotoplax martensi Pilsbry , 1896 new name for homonymy; the type species of  Gymnotoplax is considered species dubia according to Willan (1978). </p>
            <p> Berthella martensi (Pilsbry , 1896). </p>
            <p>Diagnosis: Dorsum divided into distinctive, autotomizable areas. Mantle and head delimited by black margins. Jaw platelets’ central cusp broad, with smooth margins.</p>
            <p> Etymology: The genus name is constructed in apposition of tomos (Ancient Greek for slice or section) that denotes cut pieces, alluding to the sectorized mantle, and  Berthella . </p>
            <p>Genre: Feminine.</p>
            <p> Remarks: Originally, ‘  Berthella ’ martensi was included in the genus  Gymnotoplax (Pilsbry, 1896). However, the type species of the genus  Gymnotoplax is  Pleurobranchus americanus A.E.Verrill , 1885, collected off the coast of the NE United States in 457 m. The genus  Gymnotoplax was originally described to include two species:  G. americanus (A.E.Verrill , 1885) and  G. martensi Pilsbry , 1896, but the type species was not designated. Posteriorly, Zilch (1959) designated  G. americanus as the type species of  Gymnotoplax genus, and Marcus (1977) also agreed. An exposed shell was considered characteristic of the genus and thus deserving the erection of  Gymnotoplax . Willan (1978, 1987) revised the original material and consistently showed that  G. americanus consists of a single specimen with an exposed shell because of a damaged dorsum. Thus, Willan (1978) considered, pending clarification of the identity of its type species,  Gymnotoplax as genus dubium. Accordingly, the genus  Gymnotoplax cannot be used as a valid name for ‘  Berthella ’ martensi . Moreover, ‘  B. ’ martensi was described based on a juvenile specimen off Mauritius, and Pilsbry (1896) mentioned that this specimen has a frontal incision. This character may be referring to the partite mantle typical of this  ‘ martensi group’, and thus specimens with a typical partite mantle from the Indo-Pacific were attributed subsequently to this species. Yet the exposed shell of  G. americanus is an artefact that cannot be matched to the partite mantle of ‘  B. ’ martensi . Therefore, according to the morphological diagnosis, we erect the new genus  Tomoberthella gen. nov. for  martensi species described by Pilsbry (1896). Molecular data further reinforces the morphological uniqueness of  Tomoberthella martensi comb. nov. , which appears as a distinct clade sister to the species of  Pleurehdera , and most probably represents a species complex. </p>
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	https://treatment.plazi.org/id/461DA55ADA08FFCE23CEFBFBC42DE8BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Moles, Juan;Brenzinger, Bastian;Berning, Maria I.;Martynov, Alexander;TatianaKorshunova;Schrödl, Michael	Moles, Juan, Brenzinger, Bastian, Berning, Maria I., Martynov, Alexander, TatianaKorshunova, Schrödl, Michael (2024): Systematic rearrangements in an all-genus phylogeny of side-gilled slugs (Heterobranchia: Pleurobranchida). Zoological Journal of the Linnean Society 202 (2): 1-11, DOI: 10.1093/zoolinnean/zlad162, URL: https://doi.org/10.1093/zoolinnean/zlad162
461DA55ADA08FFCD23DCFD35C263E8A3.text	461DA55ADA08FFCD23DCFD35C263E8A3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tomthompsoniidae Moles & Brenzinger & Berning & Martynov & TatianaKorshunova & Schrödl 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tomthompsoniidae fam.nov.</p>
            <p>(Fig. 1A)</p>
            <p>urn:lsid:zoobank.org:act:F71DA1C1-2926-4ED0-9C52-88514B67D4E4.</p>
            <p> Type species:  Tomthompsonia antarctica (Thiele , 1912). </p>
            <p>Diagnosis: Internal helicoid shell. Gland producing sulphuric acid, opening into oral tube and pedal gland absent. Long penial sheath covering penial lobe.</p>
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	https://treatment.plazi.org/id/461DA55ADA08FFCD23DCFD35C263E8A3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Moles, Juan;Brenzinger, Bastian;Berning, Maria I.;Martynov, Alexander;TatianaKorshunova;Schrödl, Michael	Moles, Juan, Brenzinger, Bastian, Berning, Maria I., Martynov, Alexander, TatianaKorshunova, Schrödl, Michael (2024): Systematic rearrangements in an all-genus phylogeny of side-gilled slugs (Heterobranchia: Pleurobranchida). Zoological Journal of the Linnean Society 202 (2): 1-11, DOI: 10.1093/zoolinnean/zlad162, URL: https://doi.org/10.1093/zoolinnean/zlad162
461DA55ADA08FFCD23FDFDB7C5FAE9D5.text	461DA55ADA08FFCD23FDFDB7C5FAE9D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tomthompsonioidea Moles & Brenzinger & Berning & Martynov & TatianaKorshunova & Schrödl 2024	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Tomthompsonioidea superfam.nov.</p>
            <p>urn:lsid:zoobank.org:act:9750A7F4-9CED-4D91-AD7F-BAFE8387115C.</p>
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	https://treatment.plazi.org/id/461DA55ADA08FFCD23FDFDB7C5FAE9D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Moles, Juan;Brenzinger, Bastian;Berning, Maria I.;Martynov, Alexander;TatianaKorshunova;Schrödl, Michael	Moles, Juan, Brenzinger, Bastian, Berning, Maria I., Martynov, Alexander, TatianaKorshunova, Schrödl, Michael (2024): Systematic rearrangements in an all-genus phylogeny of side-gilled slugs (Heterobranchia: Pleurobranchida). Zoological Journal of the Linnean Society 202 (2): 1-11, DOI: 10.1093/zoolinnean/zlad162, URL: https://doi.org/10.1093/zoolinnean/zlad162
