identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
417C8787FFA6FF85FF186811FC96FCB2.text	417C8787FFA6FF85FF186811FC96FCB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Geminimyndus Bahder, Stroinski & Bartlett 2026	<div><p>Genus Geminimyndus Bahder, Stroiński &amp; Bartlett, gen. nov.</p><p>Type species. Geminimyndus spinaparum Bahder, Stroiński &amp; Bartlett, gen. et sp. nov., by present designation.</p><p>Etymology. The name ‘ Geminimyndus ’ is derived from the Latin word ‘gemini’ (twins), a reference to the paired processes arising on the phallobase, combined with Myndus, a genus name and common combining form in the tribe. The name is intended as masculine.</p><p>Diagnosis. Small planthoppers,sexually dimorphic with females having similar pattern as males only significantly darker in color and degree. Large acrometope present, vertex approximately 3X wide as long at midpoint. Posterior margin of head covering pronotum at midpoint, median carina on frons evident, extending from dorsal margin of acrometope to frontoclypeal suture. Phallobase with paired, large processes arising near anterior margin, one on either side of endosoma and a single, spinose process arising on posterior margin.</p><p>Description. Small species, sexually dimorphic with males paler and less patterned, females more fuscous, stronger patterning over body. Forewings weakly to strongly patterned with fuscous patches, patterns stronger in female forewings than males. Head in lateral view rounded, in dorsal view, posterior margin of vertex nearly linear, covering pronotum at midpoint. Frons carinate at midpoint, median ocellus present.</p><p>Tegmen membranous, distinctly elongated (longer than wide) and nearly parallel-sided; slight projection at base of costal vein, clavus long; apical margin widely arcuate with breaking point in the middle of margin (about end of MP 1.1). Basal cell very narrow, distinctly longer than wide. Longitudinal veins distinctly visible dotted with pustules bearing setae. Longitudinal veins ScP+R and MP forming short composite stem from basal cell and forking distinctly before fusion of claval vein Pcu and A1; first fork of ScP+R distinctly before of first fork of MP, slightly basad of CuA stem and about at level of fusion of claval veins Pcu+A1; first fork of MP at the level of clavus apex, first fork of CuA a bit after of first fork of SC+R. Forking sequence of longitudinal veins ScP+R→CuA→MP. Branching pattern RA 1-branched, RP 3-branched, MP 5-branched, CuA 2-branched. Cell C5 the longest; cell C3 shortest, cell C1 bit shorter than C5. Cell length sequence C5→C3→C1. Clavus long reaching 2/3 length of wing, fusion of claval vein after half the clavus. Transverse veinlet icu reaching apex of clavus. Hindwing with “I-type” connection between the MP and CuA (Le Cesne et al. 2022).</p><p>Medioventral process of pygofer present,rounded at apex with two lateral, subtriangular processes; posteriolateral margins of pygofer subtriangular. Phallobase complex, paired, recurved processes arising at anterior margin and a single process arising on posterior margin, angled caudad; endosoma simple, arising in between paired processes of phallobase, processes asymmetrical in length and curvature.</p></div>	https://treatment.plazi.org/id/417C8787FFA6FF85FF186811FC96FCB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bahder, Brian W.;Stroiński, Adam;Labbe, Frederic;Bartlett, Charles R.;Ratsimbazafy, Miangaly Raharimandimby;Randretsiferana, Safidinirina Armande;Razafindrakotomamonjy, Andrianatenaina;Hasinjaka, Rasolondalao Harin’Hala;Pilet, Fabian	Bahder, Brian W., Stroiński, Adam, Labbe, Frederic, Bartlett, Charles R., Ratsimbazafy, Miangaly Raharimandimby, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Hasinjaka, Rasolondalao Harin’Hala, Pilet, Fabian (2026): New planthoppers (Hemiptera: Cixiidae) from Madagascar support the establishment of a new genus, the revised status of Eumyndus perinetensis, and an updated molecular phylogeny of Malagasy Oecleini. Zootaxa 5741 (2): 377-395, DOI: 10.11646/zootaxa.5741.2.9, URL: https://doi.org/10.11646/zootaxa.5741.2.9
417C8787FFA1FF87FF186DE1FB8EF835.text	417C8787FFA1FF87FF186DE1FB8EF835.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Geminimyndus spinaparum Bahder, Stroinski & Bartlett 2026	<div><p>Geminimyndus spinaparum Bahder, Stroiński &amp; Bartlett, gen. et sp. nov.</p><p>(Figs. 2–6)</p><p>Type locality. Madagascar, Alaotra-Mangoro Region, Parc Mitsinjo.</p><p>Etymology. The specific name is derived from the Latin word ‘ spina ’ (thorn) combined with the Latin word ‘ parum ’ (very little, insufficient), a reference to the posterior process of the phallobase being very small.</p><p>Diagnosis. Two strongly recurved processes arising on dorsolateral margin of phallobase, the right process shorter with apex curved dorsally; minute process present on posterior margin of phallobase, significantly shorter than (less than 1/4 length) paired, recurved processes.</p><p>Description. Color. Ground color testaceous, fuscous mottling over entire body, lateral margins of mesonotum with fuscous ring starting along anterior margin, extending postering along lateral carinae, reduced or absent along posterior margin, testaceous from median carina to lateral carinae (Fig. 2). Forewings irregularly patterned, more so in distal third.</p><p>Structure. Body length male: 4.8–5.0 mm with wings, 3.5 mm without wings (n =8); female: 5.2–5.4 mm with wings, 3.8–3.9 mm without wings (n =13) (Table 2).</p><p>Head. In dorsal view, vertex widest at posterior, narrowing slightly, approximately 3X wide as long at midpoint, posterior margin nearly linear, acrometope extending from approximately midpoint of eye to anterior margin (Fig. 3). In lateral view, rounded, slightly elevated above and in front of eyes, midpoint of frons protruding slightly (Fig. 3). In frontal view, frons narrowest at dorsal margin, immediately expanding, widest just below pedicel, constricting slightly at frontoclypeal suture; median carina evident, extending from dorsal margin to frontoclypeal suture; median ocellus present, nearly obsolete (Fig. 3).</p><p>Thorax. In dorsal view, pronotum hidden by posterior margin of vertex at midpoint; lateral carinae present, appearing obsolete at dorsal margin, reaching ventral margin, in lateral view posterior margin strongly sinuate, appearing serpentine (Fig. 3). Mesonotum in dorsal view approximately as wide as long at midpoint, tricarinate, median carina becoming obsolete at posterior margin, lateral carinae poorly developed (Fig. 3).</p><p>Hind tibiae lack lateral spines; spinulation 6-7-6.</p><p>Male terminalia. Pygofer in lateral view reaching dorsal margin, narrowest dorsally, expanding, widest at ventral margin, anterior margin slightly curved, posterior margin irregularly sinuate, forming subtriangular projection at midpoint (Fig. 5). In ventral view, medioventral process present, broad at base, apex rounded, slightly wider at base as long at midpoint, small, subtriangular projections arising on lateral margins approximately at midlength (Fig. 5). Gonostyli in lateral view short, extending just beyond posterior margin of pygofer, clubbed, subtriangular projection arising subapically on dorsal margin. In ventral view, broadest at base, tapering to apex, curved mesad (Fig. 5). Aedeagus simple, endosoma (E) simple (Fig. 6). Phallobase complex, two large, recurved processes present, arising on left dorsolateral margin, first process (PB1) slender, strongly recurved, apex curving dorsad, not reaching aedeagal base, second process (PB2), slightly longer than PB1, slender, apex pointing cephalad, reaching aedeagal base (Fig. 6) and a third process (PB3) arising on posterior margin near ventral margin, small, tooth-like, less than ¼ the length of PB1 and PB2 (Fig. 6). Anal segment, in lateral view, narrowest at base, dorsal margin linear to midpoint angled ventrad approximately 45 degrees, declining margin sinuate, apex broadly rounded, ventral margin linear in posterior ¾, curving at base (Fig. 6).</p><p>Plant associations. Ravenea louvelii Beentje ( Arecaceae).</p><p>Type material. Holotype, male: “ Madagascar, Alaotra-Mangoro Reg. / Parc Mitsinjo / 03.II.2023, Coll.: B.W.Bahder / Host: Ravenea louvelii / Geminimyndus spinaparum /” (FLREC).</p><p>Paratypes: 7 males, 13 females; same locality as holotype (FLREC)</p><p>Distribution. Eastern Madagascar: Alaotra-Mangoro Region, Parc Mitsinjo.</p></div>	https://treatment.plazi.org/id/417C8787FFA1FF87FF186DE1FB8EF835	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bahder, Brian W.;Stroiński, Adam;Labbe, Frederic;Bartlett, Charles R.;Ratsimbazafy, Miangaly Raharimandimby;Randretsiferana, Safidinirina Armande;Razafindrakotomamonjy, Andrianatenaina;Hasinjaka, Rasolondalao Harin’Hala;Pilet, Fabian	Bahder, Brian W., Stroiński, Adam, Labbe, Frederic, Bartlett, Charles R., Ratsimbazafy, Miangaly Raharimandimby, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Hasinjaka, Rasolondalao Harin’Hala, Pilet, Fabian (2026): New planthoppers (Hemiptera: Cixiidae) from Madagascar support the establishment of a new genus, the revised status of Eumyndus perinetensis, and an updated molecular phylogeny of Malagasy Oecleini. Zootaxa 5741 (2): 377-395, DOI: 10.11646/zootaxa.5741.2.9, URL: https://doi.org/10.11646/zootaxa.5741.2.9
417C8787FFADFF8DFF186EFCFBFCF816.text	417C8787FFADFF8DFF186EFCFBFCF816.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Geminimyndus spinamagnus Bahder, Stroinski & Bartlett 2026	<div><p>Geminimyndus spinamagnus Bahder, Stroiński &amp; Bartlett, gen. et sp. nov.</p><p>(Figs. 7–15)</p><p>Type locality. Madagascar, Analamanga Region, Ankazobe district, Ambohitantely Special Reserve.</p><p>Etymology. The specific name is derived from the Latin word ‘ spina ’ (thorn) combined with the Latin word ‘ magnus ’ (large, great), a reference to the large posterior process of the phallobase.</p><p>Diagnosis. Two strongly recurved processes arising on dorsal margin of phallobase, the right process being shorter, both processes with apex curved dorsally; large process present on posterior margin of phallobase, approximately half the length of the paired, recurved processes.</p><p>Description. Color. Ground color testaceous, fuscous mottling over entire body, lateral margins of mesonotum with fuscous ring starting along anterior margin, posteriorly extending along lateral carinae, reduced or absent along posterior margin, testaceous from median carina to lateral carinae in males, fuscous in females (Figs. 7 &amp; 8). Forewings irregularly patterned, more so in distal third.</p><p>Structure. Body length male: 4.8–4.9 mm with wings, 2.8–2.9 mm without wings (n = 23); female: 5.0– 5.1 mm with wings, 4.0 mm without wing (n =27) (Fig. 9) (Table 3).</p><p>Head. In dorsal view, vertex widest at posterior, narrowing slightly, approximately 3X wide as long at midpoint, posterior margin nearly linear, acrometope extending from approximately midpoint of eye to anterior margin (Figs. 10 &amp; 11). In lateral view, rounded, slightly elevated above and in front of eyes, midpoint of frons protruding slightly (Figs. 10 &amp; 11). In frontal view, frons narrowest at dorsal margin, immediately expanding, widest just below second antennal segment, constricting slightly at frontoclypeal suture; median carina evident, extending from dorsal margin to frontoclypeal suture; median ocellus present, nearly obsolete (Figs. 10 &amp; 11).</p><p>Thorax. In dorsal view, pronotum hidden by posterior margin of vertex at midpoint; lateral carinae present, appearing obsolete at dorsal margin, reaching ventral margin, in lateral view posterior margin strongly sinuate, appearing serpentine (Figs. 10 &amp; 11). Mesonotum in dorsal view approximately as wide as long at midpoint, tricarinate, median carina becoming obsolete at posterior margin, lateral carinae poorly developed (Figs. 10 &amp; 11).</p><p>Hind tibiae lack lateral spines; spinulation 6-7-6.</p><p>Male terminalia. Pygofer in lateral view reaching dorsal margin, narrowest dorsally, expanding, widest at ventral margin, anterior margin slightly sinuate, posterior margin irregularly sinuate, forming two subtriangular projections, larger in dorsal half, smaller in ventral half (Figs. 13 &amp; 14). In ventral view, medioventral process present, broad at base, apex rounded, slightly wider at base as long at midpoint, small, subtriangular projections arising on lateral margins approximately at midlength (Figs. 13 &amp; 14). Gonostyli in lateral view short, extending just beyond posterior margin of pygofer, clubbed, subtriangular projection arising subapically on dorsal margin;in ventral view, broadest at base, tapering to apex, curved mesad (Figs. 13 &amp; 14). Aedeagus simple, endosoma (E) simple (Fig. 15). Phallobase complex with two large, recurved processes present, arising on dorsal margin, first process (PB1) slender, strongly recurved, apex curving dorsad and cephalad, not reaching aedeagal base, second process (PB2), slightly longer than PB1, slender, apex curving dorsad and cephalad, reaching aedeagal base (Fig. 15) and a third process (PB3) arising on posterior margin near ventral margin, large, approximately half the length of PB1, curved dorsad at apex (Fig. 12). Anal segment in lateral view narrowest at base, dorsal margin nearly linear to midpoint angled ventrad approximately 45 degrees, declining margin sinuate, apex broadly rounded, ventral margin linear in posterior ¾, curving at base (Figs. 13 &amp; 14).</p><p>Plant associations. Ravenea madagascariensis Becc. ( Arecaceae).</p><p>Type material. Holotype, male: “ Madagascar, Analamanga Reg. / Ambohitantely Reserve / 05.II.2024, Coll.: B.W.Bahder / Host: Ravenea madagascariensis / Geminimyndus spinamagnus” (FLREC) .</p><p>Paratypes: 22 males, 27 females (FLREC), 5 males, 5 females (MIIZ); same locality as holotype .</p><p>Distribution. Central Madagascar: Analamanga, Ambohitantely Special Reserve</p></div>	https://treatment.plazi.org/id/417C8787FFADFF8DFF186EFCFBFCF816	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bahder, Brian W.;Stroiński, Adam;Labbe, Frederic;Bartlett, Charles R.;Ratsimbazafy, Miangaly Raharimandimby;Randretsiferana, Safidinirina Armande;Razafindrakotomamonjy, Andrianatenaina;Hasinjaka, Rasolondalao Harin’Hala;Pilet, Fabian	Bahder, Brian W., Stroiński, Adam, Labbe, Frederic, Bartlett, Charles R., Ratsimbazafy, Miangaly Raharimandimby, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Hasinjaka, Rasolondalao Harin’Hala, Pilet, Fabian (2026): New planthoppers (Hemiptera: Cixiidae) from Madagascar support the establishment of a new genus, the revised status of Eumyndus perinetensis, and an updated molecular phylogeny of Malagasy Oecleini. Zootaxa 5741 (2): 377-395, DOI: 10.11646/zootaxa.5741.2.9, URL: https://doi.org/10.11646/zootaxa.5741.2.9
417C8787FFA8FF8FFF186A7EFB8AFF72.text	417C8787FFA8FF8FFF186A7EFB8AFF72.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Geminimyndus perinetensis (Synave 1965) Bahder & Stroiński & Labbe & Bartlett & Ratsimbazafy & Randretsiferana & Razafindrakotomamonjy & Hasinjaka & Pilet 2026	<div><p>Geminimyndus perinetensis (Synave, 1965) comb. nov.</p><p>(Figs. 16–17)</p><p>Eumyndus perinetensis Synave, 1965</p><p>Type material. “ Sandrangato / Institut Scientifique Madagascar / Para-type / R. I. Sc. N. B. I. G. 23.285 / H. Synave det., 195 (61) / Eumyndus perinetensis . Synave.” (Paratype) (RBINS) (Fig. 16) .</p><p>Additional material. “ Madagascar, Alaotra-Mangoro Reg. / Parc Mitsinjo / 03.II.2023, Coll.: B.W.Bahder / Host: Ravenea madgascariensis / Geminimyndus perinetensis /” (n = 4 males, 3 females) (FLREC) (Fig. 17) .</p><p>Plant associations. Ravenea madagascariensis Becc. ( Arecaceae).</p><p>Distribution. Eastern Madagascar, Atsinanana Region: Parc Mitsinjo, Périnet</p><p>Sequence Data and Analysis. For the novel taxa, sequence data was successfully obtained for all selected markers. Products the size of 315, 529, 633, 1,365, 741 and 795 base pairs (bp) were recovered for the markers H3, COI 5’, COI 3’, 18S, 28S D8 region and 28S D9–D10 region, respectively. All sequences generated for the novel taxa were deposited in GenBank and corresponding accession numbers are presented in Supplementary Table 1.</p><p>The phylogeny constructed from concatenated data of all loci showed very strong statistical support (99.6/100) for the monophyly of Geminimyndus gen. nov. and very strong support (100/100) for it resolving adjacent to Eumyndus (Fig. 18).</p><p>Remarks. Geminimyndus gen. nov. superficially resembles Eumyndus . Historically, Eumyndus was separated from Myndus by the former lacking a median carina on the frons, whereas Myndus possessed this carina.The presence of this carina in G. perinetensis comb. nov. cast doubt on the validity of this character for defining Eumyndus . However, with the proposed status change for G. perinetensis comb. nov., the presence of a median carina is (thus far) a consistent feature for Geminimyndus gen. nov., and with its removal from Eumyndus, the lack of the carina on the frons is now consistent for Eumyndus . Additionally, all three species of Geminimyndus gen. nov. possess a large acrometope, whereas this is lacking in Eumyndus . Furthermore, the presence of paired processes arising from the phallobase in Geminimyndus gen. nov. is a strong character in contrast to the single process observed in Eumyndus to support the establishment of the genus. Thus far, three species have been observed with paired processes and two species of Eumyndus, both possessing a single process, have been observed, supporting the consistency of this character. This distinction is also strongly supported by molecular data for a variety of molecular markers. While specimens of Myndus for molecular study are lacking, the general structure of the genitalia would suggest it is more closely related to Haplaxius than to either Eumyndus or Geminimyndus gen. nov.</p></div>	https://treatment.plazi.org/id/417C8787FFA8FF8FFF186A7EFB8AFF72	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bahder, Brian W.;Stroiński, Adam;Labbe, Frederic;Bartlett, Charles R.;Ratsimbazafy, Miangaly Raharimandimby;Randretsiferana, Safidinirina Armande;Razafindrakotomamonjy, Andrianatenaina;Hasinjaka, Rasolondalao Harin’Hala;Pilet, Fabian	Bahder, Brian W., Stroiński, Adam, Labbe, Frederic, Bartlett, Charles R., Ratsimbazafy, Miangaly Raharimandimby, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Hasinjaka, Rasolondalao Harin’Hala, Pilet, Fabian (2026): New planthoppers (Hemiptera: Cixiidae) from Madagascar support the establishment of a new genus, the revised status of Eumyndus perinetensis, and an updated molecular phylogeny of Malagasy Oecleini. Zootaxa 5741 (2): 377-395, DOI: 10.11646/zootaxa.5741.2.9, URL: https://doi.org/10.11646/zootaxa.5741.2.9
417C8787FFAAFF8EFF186D8AFA32FB60.text	417C8787FFAAFF8EFF186D8AFA32FB60.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Geminimyndus Bahder & Stroiński & Labbe & Bartlett & Ratsimbazafy & Randretsiferana & Razafindrakotomamonjy & Hasinjaka & Pilet 2026	<div><p>Key to the species of Geminimyndus (adult males)</p><p>1. Process on posterior margin of phallobase (PB3) small, less than ¼ length of paired processes (Fig. 6).................................................................................... Geminimyndus spinaparum gen. et sp. nov.</p><p>–. Process on posterior margin of phallobase (PB3) large, half or more the length of the paired processes (Fig. 15).......... 2</p><p>2. Medioventral process of pygofer elongated distad of lateral processes; process on posterior margin extending to the stationary point or beyond of paired processes of the phallobase.................................... G. perinetensis comb. nov.</p><p>–. Medioventral process of pygofer not elongated distad of lateral processes; process on posterior margin not reaching stationary point of paired processes of the phallobase........................................ G. spinamagnus gen. et sp. nov.</p></div>	https://treatment.plazi.org/id/417C8787FFAAFF8EFF186D8AFA32FB60	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bahder, Brian W.;Stroiński, Adam;Labbe, Frederic;Bartlett, Charles R.;Ratsimbazafy, Miangaly Raharimandimby;Randretsiferana, Safidinirina Armande;Razafindrakotomamonjy, Andrianatenaina;Hasinjaka, Rasolondalao Harin’Hala;Pilet, Fabian	Bahder, Brian W., Stroiński, Adam, Labbe, Frederic, Bartlett, Charles R., Ratsimbazafy, Miangaly Raharimandimby, Randretsiferana, Safidinirina Armande, Razafindrakotomamonjy, Andrianatenaina, Hasinjaka, Rasolondalao Harin’Hala, Pilet, Fabian (2026): New planthoppers (Hemiptera: Cixiidae) from Madagascar support the establishment of a new genus, the revised status of Eumyndus perinetensis, and an updated molecular phylogeny of Malagasy Oecleini. Zootaxa 5741 (2): 377-395, DOI: 10.11646/zootaxa.5741.2.9, URL: https://doi.org/10.11646/zootaxa.5741.2.9
