taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
416887E48616E3148EB4FD15FD9E914E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137351/files/figure.png	https://doi.org/10.5281/zenodo.19137351	FIGURE 1. A, D, G, Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali, Indonesia, aquarium trade. B, I, J, Dynomene hispida Guerin-Méneville, 1832, female (11.2 × 8.6 mm) (ZRC 2013.0113), Taiwan; E, H, D. hispida Guerin-Méneville, 1832, male (9.9 × 8.2 mm) (ZRC 2013.0113), Taiwan. C, F, Hirsutodynomene vespertilio McLay & Ng, 2005, male (18.4 × 15.5 mm) (ZRC 2008.1475), Philippines. A–C, dorsal view of carapace; D–F ventral view of cephalothorax; G, H, male pleon; I, outer view of left cheliped; J, dorsal view of right chela and carpus. Abbreviation: bt = basal chelar tubercle.	FIGURE 1. A, D, G, Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali, Indonesia, aquarium trade. B, I, J, Dynomene hispida Guerin-Méneville, 1832, female (11.2 × 8.6 mm) (ZRC 2013.0113), Taiwan; E, H, D. hispida Guerin-Méneville, 1832, male (9.9 × 8.2 mm) (ZRC 2013.0113), Taiwan. C, F, Hirsutodynomene vespertilio McLay & Ng, 2005, male (18.4 × 15.5 mm) (ZRC 2008.1475), Philippines. A–C, dorsal view of carapace; D–F ventral view of cephalothorax; G, H, male pleon; I, outer view of left cheliped; J, dorsal view of right chela and carpus. Abbreviation: bt = basal chelar tubercle.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48616E3148EB4FD15FD9E914E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137361/files/figure.png	https://doi.org/10.5281/zenodo.19137361	FIGURE 4. Colimene pugnatrix (De Man, 1889) comb. nov.. A–E, male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali; F–J, male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali. A, F, dorsal habitus; B, outer view of right cheliped; C, outer view of left cheliped; D, I, dorsal view of carapace; E, J, frontal view of cephalothorax; G, outer view of left cheliped; H, outer view of right cheliped.	FIGURE 4. Colimene pugnatrix (De Man, 1889) comb. nov.. A–E, male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali; F–J, male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali. A, F, dorsal habitus; B, outer view of right cheliped; C, outer view of left cheliped; D, I, dorsal view of carapace; E, J, frontal view of cephalothorax; G, outer view of left cheliped; H, outer view of right cheliped.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48616E3148EB4FD15FD9E914E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137376/files/figure.png	https://doi.org/10.5281/zenodo.19137376	FIGURE 10. A–D, Colimene guamensis (McLay, 2001) comb. nov., holotype male (11.0 × 9.7 mm) (MNHN-IU-2008-11602), Guam; E–H, C. kroppi (McLay, 2001) comb. nov., holotype male (11.9 × 9.7 mm) (MNHN-IU-2008-11604), Guam. A, E, dorsal habitus; B, F, ventral view; C, G, dorsal view of carapace; D, H, frontal view of cephalothorax.	FIGURE 10. A–D, Colimene guamensis (McLay, 2001) comb. nov., holotype male (11.0 × 9.7 mm) (MNHN-IU-2008-11602), Guam; E–H, C. kroppi (McLay, 2001) comb. nov., holotype male (11.9 × 9.7 mm) (MNHN-IU-2008-11604), Guam. A, E, dorsal habitus; B, F, ventral view; C, G, dorsal view of carapace; D, H, frontal view of cephalothorax.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48616E3148EB4FD15FD9E914E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137357/files/figure.png	https://doi.org/10.5281/zenodo.19137357	FIGURE 2. Colimene pugnatrix (De Man, 1889) comb. nov., holotype male (9.8 × 7.2 mm) (SMF 4857), Mauritius. A, dorsal habitus; B, dorsal view of right carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, outer view of right cheliped; F, dorsal view of right cheliped; G, right P4; H, dactylus of right P4; I, right P5; J, plumose setae on leg; K, serrulate seta on leg; L, pleon; M, pleonal somite 5 and telson. A, D, H, J, K, L (after De Man 1889: 444, pl. 10 fig. 13); B, C, E, F, G, I, M (after McLay 2001: fig. 22). Scale bars = 1.0 mm.	FIGURE 2. Colimene pugnatrix (De Man, 1889) comb. nov., holotype male (9.8 × 7.2 mm) (SMF 4857), Mauritius. A, dorsal habitus; B, dorsal view of right carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, outer view of right cheliped; F, dorsal view of right cheliped; G, right P4; H, dactylus of right P4; I, right P5; J, plumose setae on leg; K, serrulate seta on leg; L, pleon; M, pleonal somite 5 and telson. A, D, H, J, K, L (after De Man 1889: 444, pl. 10 fig. 13); B, C, E, F, G, I, M (after McLay 2001: fig. 22). Scale bars = 1.0 mm.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48616E3148EB4FD15FD9E914E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137359/files/figure.png	https://doi.org/10.5281/zenodo.19137359	FIGURE 3. Colimene pugnatrix (De Man, 1889) comb. nov., holotype male (9.8 × 7.2 mm) (SMF 4857), Mauritius. A, dorsal view of carapace; B, ventral view of cephalothorax; C, dorsal view of right cheliped; D, ventral view of right cheliped; E, left P2; F, left P3; G, left P4; H, right P2; I, right P4. Photographs by SMF.	FIGURE 3. Colimene pugnatrix (De Man, 1889) comb. nov., holotype male (9.8 × 7.2 mm) (SMF 4857), Mauritius. A, dorsal view of carapace; B, ventral view of cephalothorax; C, dorsal view of right cheliped; D, ventral view of right cheliped; E, left P2; F, left P3; G, left P4; H, right P2; I, right P4. Photographs by SMF.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48616E3148EB4FD15FD9E914E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137366/files/figure.png	https://doi.org/10.5281/zenodo.19137366	FIGURE 6. Colimene pugnatrix (De Man, 1889) comb. nov., ovigerous female (6.3 × 4.8 mm) (USNM 41047) [holotype of Dynomene pugnatrix brevimana Rathbun, 1911], Providence Island, Madagascar. A, dorsal habitus; B, outer view of right female chela; C, ventral view of cephalothorax and pleon; D, anterior thoracic sternum and sternopleonal cavity.	FIGURE 6. Colimene pugnatrix (De Man, 1889) comb. nov., ovigerous female (6.3 × 4.8 mm) (USNM 41047) [holotype of Dynomene pugnatrix brevimana Rathbun, 1911], Providence Island, Madagascar. A, dorsal habitus; B, outer view of right female chela; C, ventral view of cephalothorax and pleon; D, anterior thoracic sternum and sternopleonal cavity.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48616E3148EB4FD15FD9E914E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137364/files/figure.png	https://doi.org/10.5281/zenodo.19137364	FIGURE 5. Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. A, subhepatic region; epistome, antennae, antennules and orbit; B, left third maxilliped; C, posterior view of carapace showing pleonal somites 1, 2 and P5; D, right chela and merus (dorsal view); E, anterior thoracic sternum and sternopleonal cavity; F, left P4 and P5 coxae, and penis; G, right side of sternum and pleon showing “locked” pleonal condition; H, inner surface of right side of sternum and pleon showing absence of socket on uropod or somite 6. Abbreviations: bt = basal chelar tubercle; cx1–5 = P1–P5 coxa, respectively; pe = penial tube; so6 = pleonal somite 6; st3–st6 = thoracic sternites 3–6, respectively; stu = sternal tubercle for pleonal locking; t = telson; ur = uropod.	FIGURE 5. Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. A, subhepatic region; epistome, antennae, antennules and orbit; B, left third maxilliped; C, posterior view of carapace showing pleonal somites 1, 2 and P5; D, right chela and merus (dorsal view); E, anterior thoracic sternum and sternopleonal cavity; F, left P4 and P5 coxae, and penis; G, right side of sternum and pleon showing “locked” pleonal condition; H, inner surface of right side of sternum and pleon showing absence of socket on uropod or somite 6. Abbreviations: bt = basal chelar tubercle; cx1–5 = P1–P5 coxa, respectively; pe = penial tube; so6 = pleonal somite 6; st3–st6 = thoracic sternites 3–6, respectively; stu = sternal tubercle for pleonal locking; t = telson; ur = uropod.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48616E3148EB4FD15FD9E914E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137368/files/figure.png	https://doi.org/10.5281/zenodo.19137368	FIGURE 7. Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. A, outer and extensor surface of P4 dactylus showing simple and plumose and filiform setae; B, serrate seta from proximal part of flexor margin of ambulatory dactylus; C, serrulate setae with longer setules from surface of pereopods; D, serrulate setae with short setules from parts of carapace and surfaces of pereopods. Abbreviations: s = simple seta; p = plumose seta.	FIGURE 7. Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. A, outer and extensor surface of P4 dactylus showing simple and plumose and filiform setae; B, serrate seta from proximal part of flexor margin of ambulatory dactylus; C, serrulate setae with longer setules from surface of pereopods; D, serrulate setae with short setules from parts of carapace and surfaces of pereopods. Abbreviations: s = simple seta; p = plumose seta.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48616E3148EB4FD15FD9E914E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137374/files/figure.png	https://doi.org/10.5281/zenodo.19137374	FIGURE 9. Colour in life. A–C, Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. D, E, C. pugnatrix (De Man, 1889) comb. nov., male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali. A, D, dorsal habitus; B, E, frontal view; C, ventral habitus.	FIGURE 9. Colour in life. A–C, Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. D, E, C. pugnatrix (De Man, 1889) comb. nov., male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali. A, D, dorsal habitus; B, E, frontal view; C, ventral habitus.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48616E3148EB4FD15FD9E914E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137378/files/figure.png	https://doi.org/10.5281/zenodo.19137378	FIGURE 11. Colimene kroppi (McLay, 2001) comb. nov.. A, paratype female (9.5 × 7.5 mm) (MNHN-IU-2008-11603), Guam; B–F, ovigerous female (10.8 × 9.5 mm) (USNM 291444), Guam. A, frontal view of cephalothorax; B, dorsal habitus; C, outer view of left chela; D, pleon; E, buccal cavity and anterior thoracic sternum; F, sternopleonal cavity.	FIGURE 11. Colimene kroppi (McLay, 2001) comb. nov.. A, paratype female (9.5 × 7.5 mm) (MNHN-IU-2008-11603), Guam; B–F, ovigerous female (10.8 × 9.5 mm) (USNM 291444), Guam. A, frontal view of cephalothorax; B, dorsal habitus; C, outer view of left chela; D, pleon; E, buccal cavity and anterior thoracic sternum; F, sternopleonal cavity.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48616E3148EB4FD15FD9E914E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137380/files/figure.png	https://doi.org/10.5281/zenodo.19137380	FIGURE 12. A–D, Colimene pugnatrix (De Man, 1889) comb. nov., ovigerous female (6.3 × 4.8 mm) (USNM 41047) [holotype of Dynomene pugnatrix brevimana Rathbun, 1911], Providence Island, Madagascar; E, F, C. pugnatrix (De Man, 1889) comb. nov., male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali; G, H, C. pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali; I, J, C. kroppi (McLay, 2001) comb. nov., ovigerous female (10.8 × 9.5 mm) (USNM 291444), Guam; K, L, C. kroppi (McLay, 2001) comb. nov., male (7.1 × 6.5 mm) (USNM 291443), Guam. A, right anterior part of female carapace; B, outer view of right female chela; C, J, right female P5 propodus and dactylus (ventral view); D, female telson (right side damaged), uropods and pleonal somite 6; E, G, right male P5 propodus and dactylus (subventral view); F, left male P5 propodus and dactylus (subventral view); H, K, left male P5 propodus and dactylus (ventral view); I, left female P5 propodus and dactylus (ventral view); L, left male P5 propodus and dactylus (dorsal view). All structures denuded. Scale bars: A–D = 0.5 mm; E–I = 0.2 mm.	FIGURE 12. A–D, Colimene pugnatrix (De Man, 1889) comb. nov., ovigerous female (6.3 × 4.8 mm) (USNM 41047) [holotype of Dynomene pugnatrix brevimana Rathbun, 1911], Providence Island, Madagascar; E, F, C. pugnatrix (De Man, 1889) comb. nov., male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali; G, H, C. pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali; I, J, C. kroppi (McLay, 2001) comb. nov., ovigerous female (10.8 × 9.5 mm) (USNM 291444), Guam; K, L, C. kroppi (McLay, 2001) comb. nov., male (7.1 × 6.5 mm) (USNM 291443), Guam. A, right anterior part of female carapace; B, outer view of right female chela; C, J, right female P5 propodus and dactylus (ventral view); D, female telson (right side damaged), uropods and pleonal somite 6; E, G, right male P5 propodus and dactylus (subventral view); F, left male P5 propodus and dactylus (subventral view); H, K, left male P5 propodus and dactylus (ventral view); I, left female P5 propodus and dactylus (ventral view); L, left male P5 propodus and dactylus (dorsal view). All structures denuded. Scale bars: A–D = 0.5 mm; E–I = 0.2 mm.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48611E3198EB4FF78FC7B9186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137351/files/figure.png	https://doi.org/10.5281/zenodo.19137351	FIGURE 1. A, D, G, Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali, Indonesia, aquarium trade. B, I, J, Dynomene hispida Guerin-Méneville, 1832, female (11.2 × 8.6 mm) (ZRC 2013.0113), Taiwan; E, H, D. hispida Guerin-Méneville, 1832, male (9.9 × 8.2 mm) (ZRC 2013.0113), Taiwan. C, F, Hirsutodynomene vespertilio McLay & Ng, 2005, male (18.4 × 15.5 mm) (ZRC 2008.1475), Philippines. A–C, dorsal view of carapace; D–F ventral view of cephalothorax; G, H, male pleon; I, outer view of left cheliped; J, dorsal view of right chela and carpus. Abbreviation: bt = basal chelar tubercle.	FIGURE 1. A, D, G, Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali, Indonesia, aquarium trade. B, I, J, Dynomene hispida Guerin-Méneville, 1832, female (11.2 × 8.6 mm) (ZRC 2013.0113), Taiwan; E, H, D. hispida Guerin-Méneville, 1832, male (9.9 × 8.2 mm) (ZRC 2013.0113), Taiwan. C, F, Hirsutodynomene vespertilio McLay & Ng, 2005, male (18.4 × 15.5 mm) (ZRC 2008.1475), Philippines. A–C, dorsal view of carapace; D–F ventral view of cephalothorax; G, H, male pleon; I, outer view of left cheliped; J, dorsal view of right chela and carpus. Abbreviation: bt = basal chelar tubercle.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48611E3198EB4FF78FC7B9186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137357/files/figure.png	https://doi.org/10.5281/zenodo.19137357	FIGURE 2. Colimene pugnatrix (De Man, 1889) comb. nov., holotype male (9.8 × 7.2 mm) (SMF 4857), Mauritius. A, dorsal habitus; B, dorsal view of right carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, outer view of right cheliped; F, dorsal view of right cheliped; G, right P4; H, dactylus of right P4; I, right P5; J, plumose setae on leg; K, serrulate seta on leg; L, pleon; M, pleonal somite 5 and telson. A, D, H, J, K, L (after De Man 1889: 444, pl. 10 fig. 13); B, C, E, F, G, I, M (after McLay 2001: fig. 22). Scale bars = 1.0 mm.	FIGURE 2. Colimene pugnatrix (De Man, 1889) comb. nov., holotype male (9.8 × 7.2 mm) (SMF 4857), Mauritius. A, dorsal habitus; B, dorsal view of right carapace; C, frontal view of cephalothorax; D, outer view of right chela; E, outer view of right cheliped; F, dorsal view of right cheliped; G, right P4; H, dactylus of right P4; I, right P5; J, plumose setae on leg; K, serrulate seta on leg; L, pleon; M, pleonal somite 5 and telson. A, D, H, J, K, L (after De Man 1889: 444, pl. 10 fig. 13); B, C, E, F, G, I, M (after McLay 2001: fig. 22). Scale bars = 1.0 mm.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48611E3198EB4FF78FC7B9186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137359/files/figure.png	https://doi.org/10.5281/zenodo.19137359	FIGURE 3. Colimene pugnatrix (De Man, 1889) comb. nov., holotype male (9.8 × 7.2 mm) (SMF 4857), Mauritius. A, dorsal view of carapace; B, ventral view of cephalothorax; C, dorsal view of right cheliped; D, ventral view of right cheliped; E, left P2; F, left P3; G, left P4; H, right P2; I, right P4. Photographs by SMF.	FIGURE 3. Colimene pugnatrix (De Man, 1889) comb. nov., holotype male (9.8 × 7.2 mm) (SMF 4857), Mauritius. A, dorsal view of carapace; B, ventral view of cephalothorax; C, dorsal view of right cheliped; D, ventral view of right cheliped; E, left P2; F, left P3; G, left P4; H, right P2; I, right P4. Photographs by SMF.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48611E3198EB4FF78FC7B9186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137361/files/figure.png	https://doi.org/10.5281/zenodo.19137361	FIGURE 4. Colimene pugnatrix (De Man, 1889) comb. nov.. A–E, male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali; F–J, male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali. A, F, dorsal habitus; B, outer view of right cheliped; C, outer view of left cheliped; D, I, dorsal view of carapace; E, J, frontal view of cephalothorax; G, outer view of left cheliped; H, outer view of right cheliped.	FIGURE 4. Colimene pugnatrix (De Man, 1889) comb. nov.. A–E, male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali; F–J, male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali. A, F, dorsal habitus; B, outer view of right cheliped; C, outer view of left cheliped; D, I, dorsal view of carapace; E, J, frontal view of cephalothorax; G, outer view of left cheliped; H, outer view of right cheliped.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48611E3198EB4FF78FC7B9186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137364/files/figure.png	https://doi.org/10.5281/zenodo.19137364	FIGURE 5. Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. A, subhepatic region; epistome, antennae, antennules and orbit; B, left third maxilliped; C, posterior view of carapace showing pleonal somites 1, 2 and P5; D, right chela and merus (dorsal view); E, anterior thoracic sternum and sternopleonal cavity; F, left P4 and P5 coxae, and penis; G, right side of sternum and pleon showing “locked” pleonal condition; H, inner surface of right side of sternum and pleon showing absence of socket on uropod or somite 6. Abbreviations: bt = basal chelar tubercle; cx1–5 = P1–P5 coxa, respectively; pe = penial tube; so6 = pleonal somite 6; st3–st6 = thoracic sternites 3–6, respectively; stu = sternal tubercle for pleonal locking; t = telson; ur = uropod.	FIGURE 5. Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. A, subhepatic region; epistome, antennae, antennules and orbit; B, left third maxilliped; C, posterior view of carapace showing pleonal somites 1, 2 and P5; D, right chela and merus (dorsal view); E, anterior thoracic sternum and sternopleonal cavity; F, left P4 and P5 coxae, and penis; G, right side of sternum and pleon showing “locked” pleonal condition; H, inner surface of right side of sternum and pleon showing absence of socket on uropod or somite 6. Abbreviations: bt = basal chelar tubercle; cx1–5 = P1–P5 coxa, respectively; pe = penial tube; so6 = pleonal somite 6; st3–st6 = thoracic sternites 3–6, respectively; stu = sternal tubercle for pleonal locking; t = telson; ur = uropod.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48611E3198EB4FF78FC7B9186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137366/files/figure.png	https://doi.org/10.5281/zenodo.19137366	FIGURE 6. Colimene pugnatrix (De Man, 1889) comb. nov., ovigerous female (6.3 × 4.8 mm) (USNM 41047) [holotype of Dynomene pugnatrix brevimana Rathbun, 1911], Providence Island, Madagascar. A, dorsal habitus; B, outer view of right female chela; C, ventral view of cephalothorax and pleon; D, anterior thoracic sternum and sternopleonal cavity.	FIGURE 6. Colimene pugnatrix (De Man, 1889) comb. nov., ovigerous female (6.3 × 4.8 mm) (USNM 41047) [holotype of Dynomene pugnatrix brevimana Rathbun, 1911], Providence Island, Madagascar. A, dorsal habitus; B, outer view of right female chela; C, ventral view of cephalothorax and pleon; D, anterior thoracic sternum and sternopleonal cavity.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48611E3198EB4FF78FC7B9186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137368/files/figure.png	https://doi.org/10.5281/zenodo.19137368	FIGURE 7. Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. A, outer and extensor surface of P4 dactylus showing simple and plumose and filiform setae; B, serrate seta from proximal part of flexor margin of ambulatory dactylus; C, serrulate setae with longer setules from surface of pereopods; D, serrulate setae with short setules from parts of carapace and surfaces of pereopods. Abbreviations: s = simple seta; p = plumose seta.	FIGURE 7. Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. A, outer and extensor surface of P4 dactylus showing simple and plumose and filiform setae; B, serrate seta from proximal part of flexor margin of ambulatory dactylus; C, serrulate setae with longer setules from surface of pereopods; D, serrulate setae with short setules from parts of carapace and surfaces of pereopods. Abbreviations: s = simple seta; p = plumose seta.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48611E3198EB4FF78FC7B9186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137372/files/figure.png	https://doi.org/10.5281/zenodo.19137372	FIGURE 8. Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (mesial view); E, distal part of left G1 (dorsal view); F, left G2 (ventral view); G, distal part of left G2 (ventral view).	FIGURE 8. Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. A, left G1 (ventral view); B, left G1 (dorsal view); C, distal part of left G1 (ventral view); D, distal part of left G1 (mesial view); E, distal part of left G1 (dorsal view); F, left G2 (ventral view); G, distal part of left G2 (ventral view).	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48611E3198EB4FF78FC7B9186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137374/files/figure.png	https://doi.org/10.5281/zenodo.19137374	FIGURE 9. Colour in life. A–C, Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. D, E, C. pugnatrix (De Man, 1889) comb. nov., male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali. A, D, dorsal habitus; B, E, frontal view; C, ventral habitus.	FIGURE 9. Colour in life. A–C, Colimene pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali. D, E, C. pugnatrix (De Man, 1889) comb. nov., male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali. A, D, dorsal habitus; B, E, frontal view; C, ventral habitus.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48611E3198EB4FF78FC7B9186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137380/files/figure.png	https://doi.org/10.5281/zenodo.19137380	FIGURE 12. A–D, Colimene pugnatrix (De Man, 1889) comb. nov., ovigerous female (6.3 × 4.8 mm) (USNM 41047) [holotype of Dynomene pugnatrix brevimana Rathbun, 1911], Providence Island, Madagascar; E, F, C. pugnatrix (De Man, 1889) comb. nov., male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali; G, H, C. pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali; I, J, C. kroppi (McLay, 2001) comb. nov., ovigerous female (10.8 × 9.5 mm) (USNM 291444), Guam; K, L, C. kroppi (McLay, 2001) comb. nov., male (7.1 × 6.5 mm) (USNM 291443), Guam. A, right anterior part of female carapace; B, outer view of right female chela; C, J, right female P5 propodus and dactylus (ventral view); D, female telson (right side damaged), uropods and pleonal somite 6; E, G, right male P5 propodus and dactylus (subventral view); F, left male P5 propodus and dactylus (subventral view); H, K, left male P5 propodus and dactylus (ventral view); I, left female P5 propodus and dactylus (ventral view); L, left male P5 propodus and dactylus (dorsal view). All structures denuded. Scale bars: A–D = 0.5 mm; E–I = 0.2 mm.	FIGURE 12. A–D, Colimene pugnatrix (De Man, 1889) comb. nov., ovigerous female (6.3 × 4.8 mm) (USNM 41047) [holotype of Dynomene pugnatrix brevimana Rathbun, 1911], Providence Island, Madagascar; E, F, C. pugnatrix (De Man, 1889) comb. nov., male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali; G, H, C. pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali; I, J, C. kroppi (McLay, 2001) comb. nov., ovigerous female (10.8 × 9.5 mm) (USNM 291444), Guam; K, L, C. kroppi (McLay, 2001) comb. nov., male (7.1 × 6.5 mm) (USNM 291443), Guam. A, right anterior part of female carapace; B, outer view of right female chela; C, J, right female P5 propodus and dactylus (ventral view); D, female telson (right side damaged), uropods and pleonal somite 6; E, G, right male P5 propodus and dactylus (subventral view); F, left male P5 propodus and dactylus (subventral view); H, K, left male P5 propodus and dactylus (ventral view); I, left female P5 propodus and dactylus (ventral view); L, left male P5 propodus and dactylus (dorsal view). All structures denuded. Scale bars: A–D = 0.5 mm; E–I = 0.2 mm.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48611E3198EB4FF78FC7B9186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137376/files/figure.png	https://doi.org/10.5281/zenodo.19137376	FIGURE 10. A–D, Colimene guamensis (McLay, 2001) comb. nov., holotype male (11.0 × 9.7 mm) (MNHN-IU-2008-11602), Guam; E–H, C. kroppi (McLay, 2001) comb. nov., holotype male (11.9 × 9.7 mm) (MNHN-IU-2008-11604), Guam. A, E, dorsal habitus; B, F, ventral view; C, G, dorsal view of carapace; D, H, frontal view of cephalothorax.	FIGURE 10. A–D, Colimene guamensis (McLay, 2001) comb. nov., holotype male (11.0 × 9.7 mm) (MNHN-IU-2008-11602), Guam; E–H, C. kroppi (McLay, 2001) comb. nov., holotype male (11.9 × 9.7 mm) (MNHN-IU-2008-11604), Guam. A, E, dorsal habitus; B, F, ventral view; C, G, dorsal view of carapace; D, H, frontal view of cephalothorax.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48607E3068EB4FA66FD6597B6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137376/files/figure.png	https://doi.org/10.5281/zenodo.19137376	FIGURE 10. A–D, Colimene guamensis (McLay, 2001) comb. nov., holotype male (11.0 × 9.7 mm) (MNHN-IU-2008-11602), Guam; E–H, C. kroppi (McLay, 2001) comb. nov., holotype male (11.9 × 9.7 mm) (MNHN-IU-2008-11604), Guam. A, E, dorsal habitus; B, F, ventral view; C, G, dorsal view of carapace; D, H, frontal view of cephalothorax.	FIGURE 10. A–D, Colimene guamensis (McLay, 2001) comb. nov., holotype male (11.0 × 9.7 mm) (MNHN-IU-2008-11602), Guam; E–H, C. kroppi (McLay, 2001) comb. nov., holotype male (11.9 × 9.7 mm) (MNHN-IU-2008-11604), Guam. A, E, dorsal habitus; B, F, ventral view; C, G, dorsal view of carapace; D, H, frontal view of cephalothorax.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48605E3038EB4F96FFD6594BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137376/files/figure.png	https://doi.org/10.5281/zenodo.19137376	FIGURE 10. A–D, Colimene guamensis (McLay, 2001) comb. nov., holotype male (11.0 × 9.7 mm) (MNHN-IU-2008-11602), Guam; E–H, C. kroppi (McLay, 2001) comb. nov., holotype male (11.9 × 9.7 mm) (MNHN-IU-2008-11604), Guam. A, E, dorsal habitus; B, F, ventral view; C, G, dorsal view of carapace; D, H, frontal view of cephalothorax.	FIGURE 10. A–D, Colimene guamensis (McLay, 2001) comb. nov., holotype male (11.0 × 9.7 mm) (MNHN-IU-2008-11602), Guam; E–H, C. kroppi (McLay, 2001) comb. nov., holotype male (11.9 × 9.7 mm) (MNHN-IU-2008-11604), Guam. A, E, dorsal habitus; B, F, ventral view; C, G, dorsal view of carapace; D, H, frontal view of cephalothorax.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48605E3038EB4F96FFD6594BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137378/files/figure.png	https://doi.org/10.5281/zenodo.19137378	FIGURE 11. Colimene kroppi (McLay, 2001) comb. nov.. A, paratype female (9.5 × 7.5 mm) (MNHN-IU-2008-11603), Guam; B–F, ovigerous female (10.8 × 9.5 mm) (USNM 291444), Guam. A, frontal view of cephalothorax; B, dorsal habitus; C, outer view of left chela; D, pleon; E, buccal cavity and anterior thoracic sternum; F, sternopleonal cavity.	FIGURE 11. Colimene kroppi (McLay, 2001) comb. nov.. A, paratype female (9.5 × 7.5 mm) (MNHN-IU-2008-11603), Guam; B–F, ovigerous female (10.8 × 9.5 mm) (USNM 291444), Guam. A, frontal view of cephalothorax; B, dorsal habitus; C, outer view of left chela; D, pleon; E, buccal cavity and anterior thoracic sternum; F, sternopleonal cavity.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
416887E48605E3038EB4F96FFD6594BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/19137380/files/figure.png	https://doi.org/10.5281/zenodo.19137380	FIGURE 12. A–D, Colimene pugnatrix (De Man, 1889) comb. nov., ovigerous female (6.3 × 4.8 mm) (USNM 41047) [holotype of Dynomene pugnatrix brevimana Rathbun, 1911], Providence Island, Madagascar; E, F, C. pugnatrix (De Man, 1889) comb. nov., male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali; G, H, C. pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali; I, J, C. kroppi (McLay, 2001) comb. nov., ovigerous female (10.8 × 9.5 mm) (USNM 291444), Guam; K, L, C. kroppi (McLay, 2001) comb. nov., male (7.1 × 6.5 mm) (USNM 291443), Guam. A, right anterior part of female carapace; B, outer view of right female chela; C, J, right female P5 propodus and dactylus (ventral view); D, female telson (right side damaged), uropods and pleonal somite 6; E, G, right male P5 propodus and dactylus (subventral view); F, left male P5 propodus and dactylus (subventral view); H, K, left male P5 propodus and dactylus (ventral view); I, left female P5 propodus and dactylus (ventral view); L, left male P5 propodus and dactylus (dorsal view). All structures denuded. Scale bars: A–D = 0.5 mm; E–I = 0.2 mm.	FIGURE 12. A–D, Colimene pugnatrix (De Man, 1889) comb. nov., ovigerous female (6.3 × 4.8 mm) (USNM 41047) [holotype of Dynomene pugnatrix brevimana Rathbun, 1911], Providence Island, Madagascar; E, F, C. pugnatrix (De Man, 1889) comb. nov., male (10.9 × 9.5 mm) (ZRC 2021.0781), Bali; G, H, C. pugnatrix (De Man, 1889) comb. nov., male (10.0 × 8.3 mm) (ZRC 2021.0722), Bali; I, J, C. kroppi (McLay, 2001) comb. nov., ovigerous female (10.8 × 9.5 mm) (USNM 291444), Guam; K, L, C. kroppi (McLay, 2001) comb. nov., male (7.1 × 6.5 mm) (USNM 291443), Guam. A, right anterior part of female carapace; B, outer view of right female chela; C, J, right female P5 propodus and dactylus (ventral view); D, female telson (right side damaged), uropods and pleonal somite 6; E, G, right male P5 propodus and dactylus (subventral view); F, left male P5 propodus and dactylus (subventral view); H, K, left male P5 propodus and dactylus (ventral view); I, left female P5 propodus and dactylus (ventral view); L, left male P5 propodus and dactylus (dorsal view). All structures denuded. Scale bars: A–D = 0.5 mm; E–I = 0.2 mm.	2025-12-24	Ng, Peter K. L.;Ng, Paul Y. C.		Zenodo	biologists	Ng, Peter K. L.;Ng, Paul Y. C.			
