taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
42378248165DFFE0BCC15C0440A0F928.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591263/files/figure.png	https://doi.org/10.5281/zenodo.15591263	FIG. 1. — Gecarcinus ruricola (Linnaeus, 1758), dorsal view: A, ♂ 66.4 × 95 mm, Cuba, dry, MNHN-IU-2000-10953 (= MNHN-B10953), with homomorph cheli- peds; B, ♂ 66.2 × 93.3 mm, Cuba, dry, MNHN-IU-2000-10955 (= MNHN-B10955), with conspicuous heterochely and heterodonty. Scale bars: 50 mm. Credits: MNHN-Soubzmaigne.	FIG. 1. — Gecarcinus ruricola (Linnaeus, 1758), dorsal view: A, ♂ 66.4 × 95 mm, Cuba, dry, MNHN-IU-2000-10953 (= MNHN-B10953), with homomorph cheli- peds; B, ♂ 66.2 × 93.3 mm, Cuba, dry, MNHN-IU-2000-10955 (= MNHN-B10955), with conspicuous heterochely and heterodonty. Scale bars: 50 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248165DFFE0BCC15C0440A0F928.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591265/files/figure.png	https://doi.org/10.5281/zenodo.15591265	FIG. 2. — Gecarcinus ruricola (Linnaeus, 1758): neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155): A, carapace; B, frontal view; C, front, proepistome,antennae (recessed) and mesial lobe of infraorbital margin;D, mxp3; E, thoracic sternum. Scale bars: A, B, D, E, 10 mm; C, 5 mm. Credits: MNHN-Soubzmaigne.	FIG. 2. — Gecarcinus ruricola (Linnaeus, 1758): neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155): A, carapace; B, frontal view; C, front, proepistome,antennae (recessed) and mesial lobe of infraorbital margin;D, mxp3; E, thoracic sternum. Scale bars: A, B, D, E, 10 mm; C, 5 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248165DFFE0BCC15C0440A0F928.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591267/files/figure.png	https://doi.org/10.5281/zenodo.15591267	FIG. 3. — Gecarcinus ruricola (Linnaeus, 1758): A-F, neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): A, B, chelipeds, external and internal views; C, pleon; D, penis; E, F, G1, cephalic and caudal views, and G2; G, ♀ 52.8 × 69.9 mm, MNHN-IU-2024-6554: vulvae.Scale bars: A, B, C, 10 mm; D, 5 mm; E, F, G, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 3. — Gecarcinus ruricola (Linnaeus, 1758): A-F, neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): A, B, chelipeds, external and internal views; C, pleon; D, penis; E, F, G1, cephalic and caudal views, and G2; G, ♀ 52.8 × 69.9 mm, MNHN-IU-2024-6554: vulvae.Scale bars: A, B, C, 10 mm; D, 5 mm; E, F, G, 2 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248165DFFE0BCC15C0440A0F928.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591270/files/figure.png	https://doi.org/10.5281/zenodo.15591270	FIG. 4. — Gecarcinus ruricola (Linnaeus,1758).A, B, typical young adult individuals.A, ♀ 41.2 × 54.2,La Désirade; KARUBENTHOS 3, MNHN-IU-2024-7258; B, ♂ 32.7 × 43.4 mm,îles des Saintes,KARUBENTHOS 3, MNHN-IU-2024-4640: general dorsal view.C, D, ♂ 37.8 × 51.4 mm,Curaçao,Caracas Bay,Hato [Hato caves], USNM 56879; C, carapace with a short ridge (not well visible here) on each lateral side; D, asymmetric mxp3. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 4. — Gecarcinus ruricola (Linnaeus,1758).A, B, typical young adult individuals.A, ♀ 41.2 × 54.2,La Désirade; KARUBENTHOS 3, MNHN-IU-2024-7258; B, ♂ 32.7 × 43.4 mm,îles des Saintes,KARUBENTHOS 3, MNHN-IU-2024-4640: general dorsal view.C, D, ♂ 37.8 × 51.4 mm,Curaçao,Caracas Bay,Hato [Hato caves], USNM 56879; C, carapace with a short ridge (not well visible here) on each lateral side; D, asymmetric mxp3. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248165DFFE0BCC15C0440A0F928.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591274/files/figure.png	https://doi.org/10.5281/zenodo.15591274	FIG. 6. — Phenotypic variations of Gecarcinus ruricola (Linnaeus,1758) from various localities:level of mxp3 (resting position) in relation to frontal margin and level of sterno-pleonal cavity’s extremity in relation to thoracic sternal suture 2/3: A, B, Haiti, Navassa Island, NHM 1967.7.1.80-85, CR 66, ♂ 60.0 × 82.7 mm (A), CR 64,♂ 55.6 × 76.6 (B); C, Haiti,♂ 25.4 × 32.0 mm, USNM 1513693;D, ♂ 32.7 × 43.4 mm, Guadeloupe,îles des Saintes;KARUBENTHOS 3, MNHN-IU-2024-4640; E, 1 ♀ 41.2 × 54.2 mm, Guadeloupe, La Désirade, KARUBENTHOS 3, MNHN-IU-2024-7258; F, Dominica, near mouth of Layou River, ♂ 65.6 × 91.0 mm, USNM 126937; G, Colombia, Providence Island, ♀ 63.4 × 83.9 mm, USNM 7343; H, Cuba, Cabanas, ♂ 51.5 × 71.5 mm, USNM 48402, with asymmetric mxp3. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 6. — Phenotypic variations of Gecarcinus ruricola (Linnaeus,1758) from various localities:level of mxp3 (resting position) in relation to frontal margin and level of sterno-pleonal cavity’s extremity in relation to thoracic sternal suture 2/3: A, B, Haiti, Navassa Island, NHM 1967.7.1.80-85, CR 66, ♂ 60.0 × 82.7 mm (A), CR 64,♂ 55.6 × 76.6 (B); C, Haiti,♂ 25.4 × 32.0 mm, USNM 1513693;D, ♂ 32.7 × 43.4 mm, Guadeloupe,îles des Saintes;KARUBENTHOS 3, MNHN-IU-2024-4640; E, 1 ♀ 41.2 × 54.2 mm, Guadeloupe, La Désirade, KARUBENTHOS 3, MNHN-IU-2024-7258; F, Dominica, near mouth of Layou River, ♂ 65.6 × 91.0 mm, USNM 126937; G, Colombia, Providence Island, ♀ 63.4 × 83.9 mm, USNM 7343; H, Cuba, Cabanas, ♂ 51.5 × 71.5 mm, USNM 48402, with asymmetric mxp3. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248165DFFE0BCC15C0440A0F928.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591284/files/figure.png	https://doi.org/10.5281/zenodo.15591284	FIG. 10. — A-C, frontal views,D-F, chelipeds:A, Gecarcinus ruricola (Linnaeus,1758),neotype,♂ 55.0 × 76.0 mm,Cuba,MNHN-IU-2017-8392 (= MNHN-B13155); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126; C, Hartnollius quadratus (Saussure, 1853) n. comb., ♀ 36.0 × 46.0 mm, Mexico, CNCR 24607; D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): see granules on margins of merus and carpus of chelipeds; E, Hartnollius lateralis n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126: see smooth margins of chelipeds; F, Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 34630: see smooth margins of chelipeds. Scale bars: 10 mm. Credits: A, D, MNHN-Soubzmaigne; B, C, E, F, CNCR-Toledano-Carrasco.	FIG. 10. — A-C, frontal views,D-F, chelipeds:A, Gecarcinus ruricola (Linnaeus,1758),neotype,♂ 55.0 × 76.0 mm,Cuba,MNHN-IU-2017-8392 (= MNHN-B13155); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126; C, Hartnollius quadratus (Saussure, 1853) n. comb., ♀ 36.0 × 46.0 mm, Mexico, CNCR 24607; D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): see granules on margins of merus and carpus of chelipeds; E, Hartnollius lateralis n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126: see smooth margins of chelipeds; F, Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 34630: see smooth margins of chelipeds. Scale bars: 10 mm. Credits: A, D, MNHN-Soubzmaigne; B, C, E, F, CNCR-Toledano-Carrasco.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248165DFFE0BCC15C0440A0F928.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591287/files/figure.png	https://doi.org/10.5281/zenodo.15591287	FIG. 11. — Mxp3, internal view, and palp: A, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, MZUSP 18501 (ex USU-279); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Costa Rica, MZUSP 18509 (ex USU-281); C, H. quadratus (Saussure, 1853) n. comb., Mexico, MZUSP 18859 (ex AHF) (see Tavares 1989: fig. 14). Courtesy Marcos Tavares.	FIG. 11. — Mxp3, internal view, and palp: A, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, MZUSP 18501 (ex USU-279); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Costa Rica, MZUSP 18509 (ex USU-281); C, H. quadratus (Saussure, 1853) n. comb., Mexico, MZUSP 18859 (ex AHF) (see Tavares 1989: fig. 14). Courtesy Marcos Tavares.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248165DFFE0BCC15C0440A0F928.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591302/files/figure.png	https://doi.org/10.5281/zenodo.15591302	FIG. 16. — Bayesian inference phylogenetic tree of Gecarcinidae based on the concatenation of mitochondrial COI and 16S rRNA gene fragments.Numbers at nodes are support values for BI (posterior probability) and ML (Ultrafast bootstrap). The clade of crabs with less terrestrial habits (Cardisoma Latreille, 1828, Discoplax A.Milne-Edwards,1867,and Tuerkayana Guinot,N. K. Ng & Rodríguez Moreno, 2018) are highlighted in purple,whereas the clade with markedly terrestrial habits (Gecarcinus Leach, 1814, Hartnollius n. gen. and Johngarthia Türkay, 1970) are highlighted in red. Ocypode rotundata Miers, 1882 was included as outgroup.	FIG. 16. — Bayesian inference phylogenetic tree of Gecarcinidae based on the concatenation of mitochondrial COI and 16S rRNA gene fragments.Numbers at nodes are support values for BI (posterior probability) and ML (Ultrafast bootstrap). The clade of crabs with less terrestrial habits (Cardisoma Latreille, 1828, Discoplax A.Milne-Edwards,1867,and Tuerkayana Guinot,N. K. Ng & Rodríguez Moreno, 2018) are highlighted in purple,whereas the clade with markedly terrestrial habits (Gecarcinus Leach, 1814, Hartnollius n. gen. and Johngarthia Türkay, 1970) are highlighted in red. Ocypode rotundata Miers, 1882 was included as outgroup.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248165DFFE0BCC15C0440A0F928.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591304/files/figure.png	https://doi.org/10.5281/zenodo.15591304	FIG. 17. — Living Gecarcinus ruricola (Linnaeus, 1758): A, ‘dark land crab’ in the field, Martinique, Anse Ceron, forest about 70 m far from the shore, altitude about 4 m; B, ‘red crab’ in the field, Cuba, Bay of Pigs, 22°17’12.28”N, 81°08’18.59”W. Credits: A, F. Lavail (planetedoc.com); B, P. Lindgren (Creative Commons Attribution-Share Alike 4.0).	FIG. 17. — Living Gecarcinus ruricola (Linnaeus, 1758): A, ‘dark land crab’ in the field, Martinique, Anse Ceron, forest about 70 m far from the shore, altitude about 4 m; B, ‘red crab’ in the field, Cuba, Bay of Pigs, 22°17’12.28”N, 81°08’18.59”W. Credits: A, F. Lavail (planetedoc.com); B, P. Lindgren (Creative Commons Attribution-Share Alike 4.0).	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248165DFFE0BCC15C0440A0F928.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591306/files/figure.png	https://doi.org/10.5281/zenodo.15591306	FIG. 18. — Gecarcinus ruricola (Linnaeus,1758) in the field: different morphs; see the red mesial lobe of infraorbital margin:A, B, Saint-Barthelemy,St Jean Beach, back-beach vegetation, 27.VI.2015 and 15.VIII.2024, respectively: ‘green morph’ and ‘red morph’; C, Saint-Barthelemy, Trace de Colombier, coastal xerophilous forest,28.VI.2013; ‘dark morph’; D, Anguilla, Dog Island,flat calcareous islet, xerophilous vegetation,18.X.2024: ‘yellow morph’; E, F, Anguilla,Prickly Pear East, flat calcareous islet, xerophilous vegetation, 26.III.2018: ‘dark morph’; G, H, Dominica, Cabrits National Park, xerophilous forest, 20.I.2022: ‘dark morph’ and ‘mixed colour morph’. Credits: K. Questel (krl-qtl.odoo.com).	FIG. 18. — Gecarcinus ruricola (Linnaeus,1758) in the field: different morphs; see the red mesial lobe of infraorbital margin:A, B, Saint-Barthelemy,St Jean Beach, back-beach vegetation, 27.VI.2015 and 15.VIII.2024, respectively: ‘green morph’ and ‘red morph’; C, Saint-Barthelemy, Trace de Colombier, coastal xerophilous forest,28.VI.2013; ‘dark morph’; D, Anguilla, Dog Island,flat calcareous islet, xerophilous vegetation,18.X.2024: ‘yellow morph’; E, F, Anguilla,Prickly Pear East, flat calcareous islet, xerophilous vegetation, 26.III.2018: ‘dark morph’; G, H, Dominica, Cabrits National Park, xerophilous forest, 20.I.2022: ‘dark morph’ and ‘mixed colour morph’. Credits: K. Questel (krl-qtl.odoo.com).	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248165DFFE0BCC15C0440A0F928.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591312/files/figure.png	https://doi.org/10.5281/zenodo.15591312	FIG. 21. — Gecarcinus ruricola (Linnaeus, 1758), the ‘black land crab’, western Caribbean, San Andrés Archipelago, Providence Island: A, Fresh Water Bay sector, 2020: after a long annual breeding migration from the damp and shaded forest areas, this ovigerous female, with a large pleon and pleopods bearing ripe eggs, lays its eggs in the sea; B, Lighthouse: two females having laid their eggs are setting off again for return migration. Credits: Gabriela Dominguez (MFA/ Filmmaker, bluelizardstudio.co).	FIG. 21. — Gecarcinus ruricola (Linnaeus, 1758), the ‘black land crab’, western Caribbean, San Andrés Archipelago, Providence Island: A, Fresh Water Bay sector, 2020: after a long annual breeding migration from the damp and shaded forest areas, this ovigerous female, with a large pleon and pleopods bearing ripe eggs, lays its eggs in the sea; B, Lighthouse: two females having laid their eggs are setting off again for return migration. Credits: Gabriela Dominguez (MFA/ Filmmaker, bluelizardstudio.co).	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591263/files/figure.png	https://doi.org/10.5281/zenodo.15591263	FIG. 1. — Gecarcinus ruricola (Linnaeus, 1758), dorsal view: A, ♂ 66.4 × 95 mm, Cuba, dry, MNHN-IU-2000-10953 (= MNHN-B10953), with homomorph cheli- peds; B, ♂ 66.2 × 93.3 mm, Cuba, dry, MNHN-IU-2000-10955 (= MNHN-B10955), with conspicuous heterochely and heterodonty. Scale bars: 50 mm. Credits: MNHN-Soubzmaigne.	FIG. 1. — Gecarcinus ruricola (Linnaeus, 1758), dorsal view: A, ♂ 66.4 × 95 mm, Cuba, dry, MNHN-IU-2000-10953 (= MNHN-B10953), with homomorph cheli- peds; B, ♂ 66.2 × 93.3 mm, Cuba, dry, MNHN-IU-2000-10955 (= MNHN-B10955), with conspicuous heterochely and heterodonty. Scale bars: 50 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591265/files/figure.png	https://doi.org/10.5281/zenodo.15591265	FIG. 2. — Gecarcinus ruricola (Linnaeus, 1758): neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155): A, carapace; B, frontal view; C, front, proepistome,antennae (recessed) and mesial lobe of infraorbital margin;D, mxp3; E, thoracic sternum. Scale bars: A, B, D, E, 10 mm; C, 5 mm. Credits: MNHN-Soubzmaigne.	FIG. 2. — Gecarcinus ruricola (Linnaeus, 1758): neotype, ♂ 55.0 × 76.0 mm, Cuba, 1914, MNHN-IU-2017-8392 (= MNHN-B13155): A, carapace; B, frontal view; C, front, proepistome,antennae (recessed) and mesial lobe of infraorbital margin;D, mxp3; E, thoracic sternum. Scale bars: A, B, D, E, 10 mm; C, 5 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591267/files/figure.png	https://doi.org/10.5281/zenodo.15591267	FIG. 3. — Gecarcinus ruricola (Linnaeus, 1758): A-F, neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): A, B, chelipeds, external and internal views; C, pleon; D, penis; E, F, G1, cephalic and caudal views, and G2; G, ♀ 52.8 × 69.9 mm, MNHN-IU-2024-6554: vulvae.Scale bars: A, B, C, 10 mm; D, 5 mm; E, F, G, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 3. — Gecarcinus ruricola (Linnaeus, 1758): A-F, neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): A, B, chelipeds, external and internal views; C, pleon; D, penis; E, F, G1, cephalic and caudal views, and G2; G, ♀ 52.8 × 69.9 mm, MNHN-IU-2024-6554: vulvae.Scale bars: A, B, C, 10 mm; D, 5 mm; E, F, G, 2 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591270/files/figure.png	https://doi.org/10.5281/zenodo.15591270	FIG. 4. — Gecarcinus ruricola (Linnaeus,1758).A, B, typical young adult individuals.A, ♀ 41.2 × 54.2,La Désirade; KARUBENTHOS 3, MNHN-IU-2024-7258; B, ♂ 32.7 × 43.4 mm,îles des Saintes,KARUBENTHOS 3, MNHN-IU-2024-4640: general dorsal view.C, D, ♂ 37.8 × 51.4 mm,Curaçao,Caracas Bay,Hato [Hato caves], USNM 56879; C, carapace with a short ridge (not well visible here) on each lateral side; D, asymmetric mxp3. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 4. — Gecarcinus ruricola (Linnaeus,1758).A, B, typical young adult individuals.A, ♀ 41.2 × 54.2,La Désirade; KARUBENTHOS 3, MNHN-IU-2024-7258; B, ♂ 32.7 × 43.4 mm,îles des Saintes,KARUBENTHOS 3, MNHN-IU-2024-4640: general dorsal view.C, D, ♂ 37.8 × 51.4 mm,Curaçao,Caracas Bay,Hato [Hato caves], USNM 56879; C, carapace with a short ridge (not well visible here) on each lateral side; D, asymmetric mxp3. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591274/files/figure.png	https://doi.org/10.5281/zenodo.15591274	FIG. 6. — Phenotypic variations of Gecarcinus ruricola (Linnaeus,1758) from various localities:level of mxp3 (resting position) in relation to frontal margin and level of sterno-pleonal cavity’s extremity in relation to thoracic sternal suture 2/3: A, B, Haiti, Navassa Island, NHM 1967.7.1.80-85, CR 66, ♂ 60.0 × 82.7 mm (A), CR 64,♂ 55.6 × 76.6 (B); C, Haiti,♂ 25.4 × 32.0 mm, USNM 1513693;D, ♂ 32.7 × 43.4 mm, Guadeloupe,îles des Saintes;KARUBENTHOS 3, MNHN-IU-2024-4640; E, 1 ♀ 41.2 × 54.2 mm, Guadeloupe, La Désirade, KARUBENTHOS 3, MNHN-IU-2024-7258; F, Dominica, near mouth of Layou River, ♂ 65.6 × 91.0 mm, USNM 126937; G, Colombia, Providence Island, ♀ 63.4 × 83.9 mm, USNM 7343; H, Cuba, Cabanas, ♂ 51.5 × 71.5 mm, USNM 48402, with asymmetric mxp3. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 6. — Phenotypic variations of Gecarcinus ruricola (Linnaeus,1758) from various localities:level of mxp3 (resting position) in relation to frontal margin and level of sterno-pleonal cavity’s extremity in relation to thoracic sternal suture 2/3: A, B, Haiti, Navassa Island, NHM 1967.7.1.80-85, CR 66, ♂ 60.0 × 82.7 mm (A), CR 64,♂ 55.6 × 76.6 (B); C, Haiti,♂ 25.4 × 32.0 mm, USNM 1513693;D, ♂ 32.7 × 43.4 mm, Guadeloupe,îles des Saintes;KARUBENTHOS 3, MNHN-IU-2024-4640; E, 1 ♀ 41.2 × 54.2 mm, Guadeloupe, La Désirade, KARUBENTHOS 3, MNHN-IU-2024-7258; F, Dominica, near mouth of Layou River, ♂ 65.6 × 91.0 mm, USNM 126937; G, Colombia, Providence Island, ♀ 63.4 × 83.9 mm, USNM 7343; H, Cuba, Cabanas, ♂ 51.5 × 71.5 mm, USNM 48402, with asymmetric mxp3. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591284/files/figure.png	https://doi.org/10.5281/zenodo.15591284	FIG. 10. — A-C, frontal views,D-F, chelipeds:A, Gecarcinus ruricola (Linnaeus,1758),neotype,♂ 55.0 × 76.0 mm,Cuba,MNHN-IU-2017-8392 (= MNHN-B13155); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126; C, Hartnollius quadratus (Saussure, 1853) n. comb., ♀ 36.0 × 46.0 mm, Mexico, CNCR 24607; D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): see granules on margins of merus and carpus of chelipeds; E, Hartnollius lateralis n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126: see smooth margins of chelipeds; F, Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 34630: see smooth margins of chelipeds. Scale bars: 10 mm. Credits: A, D, MNHN-Soubzmaigne; B, C, E, F, CNCR-Toledano-Carrasco.	FIG. 10. — A-C, frontal views,D-F, chelipeds:A, Gecarcinus ruricola (Linnaeus,1758),neotype,♂ 55.0 × 76.0 mm,Cuba,MNHN-IU-2017-8392 (= MNHN-B13155); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126; C, Hartnollius quadratus (Saussure, 1853) n. comb., ♀ 36.0 × 46.0 mm, Mexico, CNCR 24607; D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): see granules on margins of merus and carpus of chelipeds; E, Hartnollius lateralis n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126: see smooth margins of chelipeds; F, Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 34630: see smooth margins of chelipeds. Scale bars: 10 mm. Credits: A, D, MNHN-Soubzmaigne; B, C, E, F, CNCR-Toledano-Carrasco.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591287/files/figure.png	https://doi.org/10.5281/zenodo.15591287	FIG. 11. — Mxp3, internal view, and palp: A, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, MZUSP 18501 (ex USU-279); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Costa Rica, MZUSP 18509 (ex USU-281); C, H. quadratus (Saussure, 1853) n. comb., Mexico, MZUSP 18859 (ex AHF) (see Tavares 1989: fig. 14). Courtesy Marcos Tavares.	FIG. 11. — Mxp3, internal view, and palp: A, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, MZUSP 18501 (ex USU-279); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Costa Rica, MZUSP 18509 (ex USU-281); C, H. quadratus (Saussure, 1853) n. comb., Mexico, MZUSP 18859 (ex AHF) (see Tavares 1989: fig. 14). Courtesy Marcos Tavares.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591302/files/figure.png	https://doi.org/10.5281/zenodo.15591302	FIG. 16. — Bayesian inference phylogenetic tree of Gecarcinidae based on the concatenation of mitochondrial COI and 16S rRNA gene fragments.Numbers at nodes are support values for BI (posterior probability) and ML (Ultrafast bootstrap). The clade of crabs with less terrestrial habits (Cardisoma Latreille, 1828, Discoplax A.Milne-Edwards,1867,and Tuerkayana Guinot,N. K. Ng & Rodríguez Moreno, 2018) are highlighted in purple,whereas the clade with markedly terrestrial habits (Gecarcinus Leach, 1814, Hartnollius n. gen. and Johngarthia Türkay, 1970) are highlighted in red. Ocypode rotundata Miers, 1882 was included as outgroup.	FIG. 16. — Bayesian inference phylogenetic tree of Gecarcinidae based on the concatenation of mitochondrial COI and 16S rRNA gene fragments.Numbers at nodes are support values for BI (posterior probability) and ML (Ultrafast bootstrap). The clade of crabs with less terrestrial habits (Cardisoma Latreille, 1828, Discoplax A.Milne-Edwards,1867,and Tuerkayana Guinot,N. K. Ng & Rodríguez Moreno, 2018) are highlighted in purple,whereas the clade with markedly terrestrial habits (Gecarcinus Leach, 1814, Hartnollius n. gen. and Johngarthia Türkay, 1970) are highlighted in red. Ocypode rotundata Miers, 1882 was included as outgroup.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591304/files/figure.png	https://doi.org/10.5281/zenodo.15591304	FIG. 17. — Living Gecarcinus ruricola (Linnaeus, 1758): A, ‘dark land crab’ in the field, Martinique, Anse Ceron, forest about 70 m far from the shore, altitude about 4 m; B, ‘red crab’ in the field, Cuba, Bay of Pigs, 22°17’12.28”N, 81°08’18.59”W. Credits: A, F. Lavail (planetedoc.com); B, P. Lindgren (Creative Commons Attribution-Share Alike 4.0).	FIG. 17. — Living Gecarcinus ruricola (Linnaeus, 1758): A, ‘dark land crab’ in the field, Martinique, Anse Ceron, forest about 70 m far from the shore, altitude about 4 m; B, ‘red crab’ in the field, Cuba, Bay of Pigs, 22°17’12.28”N, 81°08’18.59”W. Credits: A, F. Lavail (planetedoc.com); B, P. Lindgren (Creative Commons Attribution-Share Alike 4.0).	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591306/files/figure.png	https://doi.org/10.5281/zenodo.15591306	FIG. 18. — Gecarcinus ruricola (Linnaeus,1758) in the field: different morphs; see the red mesial lobe of infraorbital margin:A, B, Saint-Barthelemy,St Jean Beach, back-beach vegetation, 27.VI.2015 and 15.VIII.2024, respectively: ‘green morph’ and ‘red morph’; C, Saint-Barthelemy, Trace de Colombier, coastal xerophilous forest,28.VI.2013; ‘dark morph’; D, Anguilla, Dog Island,flat calcareous islet, xerophilous vegetation,18.X.2024: ‘yellow morph’; E, F, Anguilla,Prickly Pear East, flat calcareous islet, xerophilous vegetation, 26.III.2018: ‘dark morph’; G, H, Dominica, Cabrits National Park, xerophilous forest, 20.I.2022: ‘dark morph’ and ‘mixed colour morph’. Credits: K. Questel (krl-qtl.odoo.com).	FIG. 18. — Gecarcinus ruricola (Linnaeus,1758) in the field: different morphs; see the red mesial lobe of infraorbital margin:A, B, Saint-Barthelemy,St Jean Beach, back-beach vegetation, 27.VI.2015 and 15.VIII.2024, respectively: ‘green morph’ and ‘red morph’; C, Saint-Barthelemy, Trace de Colombier, coastal xerophilous forest,28.VI.2013; ‘dark morph’; D, Anguilla, Dog Island,flat calcareous islet, xerophilous vegetation,18.X.2024: ‘yellow morph’; E, F, Anguilla,Prickly Pear East, flat calcareous islet, xerophilous vegetation, 26.III.2018: ‘dark morph’; G, H, Dominica, Cabrits National Park, xerophilous forest, 20.I.2022: ‘dark morph’ and ‘mixed colour morph’. Credits: K. Questel (krl-qtl.odoo.com).	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591312/files/figure.png	https://doi.org/10.5281/zenodo.15591312	FIG. 21. — Gecarcinus ruricola (Linnaeus, 1758), the ‘black land crab’, western Caribbean, San Andrés Archipelago, Providence Island: A, Fresh Water Bay sector, 2020: after a long annual breeding migration from the damp and shaded forest areas, this ovigerous female, with a large pleon and pleopods bearing ripe eggs, lays its eggs in the sea; B, Lighthouse: two females having laid their eggs are setting off again for return migration. Credits: Gabriela Dominguez (MFA/ Filmmaker, bluelizardstudio.co).	FIG. 21. — Gecarcinus ruricola (Linnaeus, 1758), the ‘black land crab’, western Caribbean, San Andrés Archipelago, Providence Island: A, Fresh Water Bay sector, 2020: after a long annual breeding migration from the damp and shaded forest areas, this ovigerous female, with a large pleon and pleopods bearing ripe eggs, lays its eggs in the sea; B, Lighthouse: two females having laid their eggs are setting off again for return migration. Credits: Gabriela Dominguez (MFA/ Filmmaker, bluelizardstudio.co).	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591289/files/figure.png	https://doi.org/10.5281/zenodo.15591289	FIG. 12. — Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656): A, frontal view; B, front, orbit,and mesial lobe of infraorbital margin;C, front,proepistome (hidden), antenna (exposed) and mesial lobe of infraorbital margin; D, thoracic sternum and male pleon; E, F, external and internal views of chelipeds. Scale bars: A, D, E, F, 10 mm; B, C, 5 mm. Credits: MNHN-Soubzmaigne.	FIG. 12. — Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656): A, frontal view; B, front, orbit,and mesial lobe of infraorbital margin;C, front,proepistome (hidden), antenna (exposed) and mesial lobe of infraorbital margin; D, thoracic sternum and male pleon; E, F, external and internal views of chelipeds. Scale bars: A, D, E, F, 10 mm; B, C, 5 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591296/files/figure.png	https://doi.org/10.5281/zenodo.15591296	FIG. 14. — Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900): A, frontal view; B, front, proepistome (hidden), antenna (exposed) and mesial lobe of infraorbital margin; C, thoracic sternum and male pleon; D, E, external and internal views of cheli- peds. Scale bars: A, C, D, E, 20 mm; B, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 14. — Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900): A, frontal view; B, front, proepistome (hidden), antenna (exposed) and mesial lobe of infraorbital margin; C, thoracic sternum and male pleon; D, E, external and internal views of cheli- peds. Scale bars: A, C, D, E, 20 mm; B, 2 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591308/files/figure.png	https://doi.org/10.5281/zenodo.15591308	FIG. 19. — Hartnollius lateralis (Fréminville in Guérin,1832) n. comb., living and in the field:A, Guadeloupe,Marie-Galante, KARUBENTHOS 3, Stn IGM25,MNHN- IU-2024-5551; B, C, Guadeloupe, July 2015, on sand; D, E, Martinique, Anse à Prunes, forest about 65 m far from the shore, altitude about 2 m; D, crab bearing a branch; E, crab on a tree. Credits: A, L. Corbari-MNHN-ARBig-LPRig; B, C, J. Poupin; D, E, F. Lavail (planetedoc.com).	FIG. 19. — Hartnollius lateralis (Fréminville in Guérin,1832) n. comb., living and in the field:A, Guadeloupe,Marie-Galante, KARUBENTHOS 3, Stn IGM25,MNHN- IU-2024-5551; B, C, Guadeloupe, July 2015, on sand; D, E, Martinique, Anse à Prunes, forest about 65 m far from the shore, altitude about 2 m; D, crab bearing a branch; E, crab on a tree. Credits: A, L. Corbari-MNHN-ARBig-LPRig; B, C, J. Poupin; D, E, F. Lavail (planetedoc.com).	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591272/files/figure.png	https://doi.org/10.5281/zenodo.15591272	FIG. 5. — Specimens from mainland America identified by Rathbun as Gecarcinus ruricola (Linnaeus, 1758), but being in fact Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb. A, B, ♂ 32.3 × 40.5 mm, Florida, Florida Keys, Loggerhead Key, USNM 71219: A, general dorsal view; B, ventral view; C, D, ♂ 32.4 × 44.2 mm, Nicaragua, Greytown, Rathbun det. Gecarcinus ruricola; labels by Carson and Barnwell indicating G. lateralis, USNM 74612; C, general dorsal view; D, ventral view. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Specimens from mainland America identified by Rathbun as Gecarcinus ruricola (Linnaeus, 1758), but being in fact Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb. A, B, ♂ 32.3 × 40.5 mm, Florida, Florida Keys, Loggerhead Key, USNM 71219: A, general dorsal view; B, ventral view; C, D, ♂ 32.4 × 44.2 mm, Nicaragua, Greytown, Rathbun det. Gecarcinus ruricola; labels by Carson and Barnwell indicating G. lateralis, USNM 74612; C, general dorsal view; D, ventral view. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591276/files/figure.png	https://doi.org/10.5281/zenodo.15591276	FIG. 7. — Crabs from the Antilles in the Catalogue of the old Leipzig Collection identified as Gecarcinus ruricola (Linnaeus,1758),purchased from Felsche [1881], now in the collection of SNSD MTD. Not supposed to be syntypes of Ocypode agricola Reichenbach,1836;it is probably Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb. A-C, ‘specimen 2’, ♂, Antilles, SNSD old catalogue No.Crus 1751; No. 274.6: A, dorsal view; B, frontal view showing mxp3, C, ventral view; D-E, ‘specimen 1’, Antilles, SNSD old catalogue No.Crus 1751; No. 274.6: D, dorsal view; E, frontal view showing mxp3; F-H, ♂, Antilles, SNSD MTK old catalogue Crus 1749; No. 274.4 (nec 274.3, wrong inscription on the specimen): F, dorsal view; G, frontal view showing mxp3; H, ventral view. Credits: André Reimann, SNSD, Dresden.	FIG. 7. — Crabs from the Antilles in the Catalogue of the old Leipzig Collection identified as Gecarcinus ruricola (Linnaeus,1758),purchased from Felsche [1881], now in the collection of SNSD MTD. Not supposed to be syntypes of Ocypode agricola Reichenbach,1836;it is probably Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb. A-C, ‘specimen 2’, ♂, Antilles, SNSD old catalogue No.Crus 1751; No. 274.6: A, dorsal view; B, frontal view showing mxp3, C, ventral view; D-E, ‘specimen 1’, Antilles, SNSD old catalogue No.Crus 1751; No. 274.6: D, dorsal view; E, frontal view showing mxp3; F-H, ♂, Antilles, SNSD MTK old catalogue Crus 1749; No. 274.4 (nec 274.3, wrong inscription on the specimen): F, dorsal view; G, frontal view showing mxp3; H, ventral view. Credits: André Reimann, SNSD, Dresden.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481651FFF2BCDF5D874374FE55.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591278/files/figure.png	https://doi.org/10.5281/zenodo.15591278	FIG. 8. — Syntypes of Gecarcinus depressus Saussure, 1857, Haiti, at MHNG: A-D, ♂ c. 23.1 × 18.7 mm, MHNG-ARTO-16403:A, general dorsal view; B, thoracic sternum:C, anterior ventral view:merus mxp3 not emarginate on anterior margin,mesial lobe infraorbital margin covered by frontal margin,and extremity of sternopleonal cavity at same level as thoracic sternal suture 2/3; D, one cheliped and legs (detached). Probably Gecarcinus ruricola (Linnaeus, 1758); E-H, ♂ c. 17.6 × 14.6 mm (damaged),MHNG-ARTO-16404;E, carapace (damaged),dorsal view;F, ventral view; G, chelae and legs (detached); H, mxp3 (detached),at higher mag- nification: anterior margin with a small notch. Probably Hartnollius lateralis n. comb. Scale bars: A-G, 5 mm, H, 1 mm. Credits: MHNG-Christina Lehmann-Graber.	FIG. 8. — Syntypes of Gecarcinus depressus Saussure, 1857, Haiti, at MHNG: A-D, ♂ c. 23.1 × 18.7 mm, MHNG-ARTO-16403:A, general dorsal view; B, thoracic sternum:C, anterior ventral view:merus mxp3 not emarginate on anterior margin,mesial lobe infraorbital margin covered by frontal margin,and extremity of sternopleonal cavity at same level as thoracic sternal suture 2/3; D, one cheliped and legs (detached). Probably Gecarcinus ruricola (Linnaeus, 1758); E-H, ♂ c. 17.6 × 14.6 mm (damaged),MHNG-ARTO-16404;E, carapace (damaged),dorsal view;F, ventral view; G, chelae and legs (detached); H, mxp3 (detached),at higher mag- nification: anterior margin with a small notch. Probably Hartnollius lateralis n. comb. Scale bars: A-G, 5 mm, H, 1 mm. Credits: MHNG-Christina Lehmann-Graber.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481643FFF0BD1B596346C0FC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591272/files/figure.png	https://doi.org/10.5281/zenodo.15591272	FIG. 5. — Specimens from mainland America identified by Rathbun as Gecarcinus ruricola (Linnaeus, 1758), but being in fact Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb. A, B, ♂ 32.3 × 40.5 mm, Florida, Florida Keys, Loggerhead Key, USNM 71219: A, general dorsal view; B, ventral view; C, D, ♂ 32.4 × 44.2 mm, Nicaragua, Greytown, Rathbun det. Gecarcinus ruricola; labels by Carson and Barnwell indicating G. lateralis, USNM 74612; C, general dorsal view; D, ventral view. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Specimens from mainland America identified by Rathbun as Gecarcinus ruricola (Linnaeus, 1758), but being in fact Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb. A, B, ♂ 32.3 × 40.5 mm, Florida, Florida Keys, Loggerhead Key, USNM 71219: A, general dorsal view; B, ventral view; C, D, ♂ 32.4 × 44.2 mm, Nicaragua, Greytown, Rathbun det. Gecarcinus ruricola; labels by Carson and Barnwell indicating G. lateralis, USNM 74612; C, general dorsal view; D, ventral view. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481643FFF0BD1B596346C0FC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591280/files/figure.png	https://doi.org/10.5281/zenodo.15591280	FIG. 9. — Hartnollius n. gen., general views:A, H. lateralis (Fréminville in Guérin,1832) n. comb.,♂ 42.0 × 56.0 mm,Guadeloupe, MNHN-IU-2017-8390 (= MNHN- B24656); B, H. quadratus (Saussure, 1853) n. comb., ♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900). Scale bars: 20 mm. Credits: MNHN- Soubzmaigne.	FIG. 9. — Hartnollius n. gen., general views:A, H. lateralis (Fréminville in Guérin,1832) n. comb.,♂ 42.0 × 56.0 mm,Guadeloupe, MNHN-IU-2017-8390 (= MNHN- B24656); B, H. quadratus (Saussure, 1853) n. comb., ♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900). Scale bars: 20 mm. Credits: MNHN- Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481643FFF0BD1B596346C0FC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591284/files/figure.png	https://doi.org/10.5281/zenodo.15591284	FIG. 10. — A-C, frontal views,D-F, chelipeds:A, Gecarcinus ruricola (Linnaeus,1758),neotype,♂ 55.0 × 76.0 mm,Cuba,MNHN-IU-2017-8392 (= MNHN-B13155); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126; C, Hartnollius quadratus (Saussure, 1853) n. comb., ♀ 36.0 × 46.0 mm, Mexico, CNCR 24607; D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): see granules on margins of merus and carpus of chelipeds; E, Hartnollius lateralis n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126: see smooth margins of chelipeds; F, Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 34630: see smooth margins of chelipeds. Scale bars: 10 mm. Credits: A, D, MNHN-Soubzmaigne; B, C, E, F, CNCR-Toledano-Carrasco.	FIG. 10. — A-C, frontal views,D-F, chelipeds:A, Gecarcinus ruricola (Linnaeus,1758),neotype,♂ 55.0 × 76.0 mm,Cuba,MNHN-IU-2017-8392 (= MNHN-B13155); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126; C, Hartnollius quadratus (Saussure, 1853) n. comb., ♀ 36.0 × 46.0 mm, Mexico, CNCR 24607; D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): see granules on margins of merus and carpus of chelipeds; E, Hartnollius lateralis n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126: see smooth margins of chelipeds; F, Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 34630: see smooth margins of chelipeds. Scale bars: 10 mm. Credits: A, D, MNHN-Soubzmaigne; B, C, E, F, CNCR-Toledano-Carrasco.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481643FFF0BD1B596346C0FC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591287/files/figure.png	https://doi.org/10.5281/zenodo.15591287	FIG. 11. — Mxp3, internal view, and palp: A, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, MZUSP 18501 (ex USU-279); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Costa Rica, MZUSP 18509 (ex USU-281); C, H. quadratus (Saussure, 1853) n. comb., Mexico, MZUSP 18859 (ex AHF) (see Tavares 1989: fig. 14). Courtesy Marcos Tavares.	FIG. 11. — Mxp3, internal view, and palp: A, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, MZUSP 18501 (ex USU-279); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Costa Rica, MZUSP 18509 (ex USU-281); C, H. quadratus (Saussure, 1853) n. comb., Mexico, MZUSP 18859 (ex AHF) (see Tavares 1989: fig. 14). Courtesy Marcos Tavares.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481643FFF0BD1B596346C0FC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591289/files/figure.png	https://doi.org/10.5281/zenodo.15591289	FIG. 12. — Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656): A, frontal view; B, front, orbit,and mesial lobe of infraorbital margin;C, front,proepistome (hidden), antenna (exposed) and mesial lobe of infraorbital margin; D, thoracic sternum and male pleon; E, F, external and internal views of chelipeds. Scale bars: A, D, E, F, 10 mm; B, C, 5 mm. Credits: MNHN-Soubzmaigne.	FIG. 12. — Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656): A, frontal view; B, front, orbit,and mesial lobe of infraorbital margin;C, front,proepistome (hidden), antenna (exposed) and mesial lobe of infraorbital margin; D, thoracic sternum and male pleon; E, F, external and internal views of chelipeds. Scale bars: A, D, E, F, 10 mm; B, C, 5 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481643FFF0BD1B596346C0FC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591292/files/figure.png	https://doi.org/10.5281/zenodo.15591292	FIG. 13. — Harrtnollius n. gen.: gonopods and vulvae:A, B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656): A, ♂ 40.5 × 55.7 mm: G1, G2, caudal view; B, ♀ 32.6 × 41.8 mm:vulvae;C, H.quadratus (Saussure,1853) n.comb.,♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900):G1, G2, caudal view. Scale bars: A, C, 2 mm; B, 1 mm. Credits: MNHN-Soubzmaigne.	FIG. 13. — Harrtnollius n. gen.: gonopods and vulvae:A, B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656): A, ♂ 40.5 × 55.7 mm: G1, G2, caudal view; B, ♀ 32.6 × 41.8 mm:vulvae;C, H.quadratus (Saussure,1853) n.comb.,♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900):G1, G2, caudal view. Scale bars: A, C, 2 mm; B, 1 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481643FFF0BD1B596346C0FC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591296/files/figure.png	https://doi.org/10.5281/zenodo.15591296	FIG. 14. — Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900): A, frontal view; B, front, proepistome (hidden), antenna (exposed) and mesial lobe of infraorbital margin; C, thoracic sternum and male pleon; D, E, external and internal views of cheli- peds. Scale bars: A, C, D, E, 20 mm; B, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 14. — Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900): A, frontal view; B, front, proepistome (hidden), antenna (exposed) and mesial lobe of infraorbital margin; C, thoracic sternum and male pleon; D, E, external and internal views of cheli- peds. Scale bars: A, C, D, E, 20 mm; B, 2 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481643FFF0BD1B596346C0FC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591300/files/figure.png	https://doi.org/10.5281/zenodo.15591300	FIG. 15. — Hartnollius nobilii (Perger & Wall, 2014) n. comb., paratype, ♀ 36.6 × 44.3 mm, Ecuador, MNHN-IU-2014-11211 (= MNHN-B12314): A, dorsal view; B, ventral view; C, front, orbit, and mesial lobe of infraorbital margin; D, front, proepistome (hidden), antenna (exposed), mesial lobe of infraorbital margin, and mxp3; E, thoracic sternum, female pleon and vulvae; F, left cheliped, internal view; G, chelipeds, external view; H, right P3; I, left P4. Scale bar: 10 mm. Credits: MNHN-Rodríguez Moreno.	FIG. 15. — Hartnollius nobilii (Perger & Wall, 2014) n. comb., paratype, ♀ 36.6 × 44.3 mm, Ecuador, MNHN-IU-2014-11211 (= MNHN-B12314): A, dorsal view; B, ventral view; C, front, orbit, and mesial lobe of infraorbital margin; D, front, proepistome (hidden), antenna (exposed), mesial lobe of infraorbital margin, and mxp3; E, thoracic sternum, female pleon and vulvae; F, left cheliped, internal view; G, chelipeds, external view; H, right P3; I, left P4. Scale bar: 10 mm. Credits: MNHN-Rodríguez Moreno.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481643FFF0BD1B596346C0FC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591302/files/figure.png	https://doi.org/10.5281/zenodo.15591302	FIG. 16. — Bayesian inference phylogenetic tree of Gecarcinidae based on the concatenation of mitochondrial COI and 16S rRNA gene fragments.Numbers at nodes are support values for BI (posterior probability) and ML (Ultrafast bootstrap). The clade of crabs with less terrestrial habits (Cardisoma Latreille, 1828, Discoplax A.Milne-Edwards,1867,and Tuerkayana Guinot,N. K. Ng & Rodríguez Moreno, 2018) are highlighted in purple,whereas the clade with markedly terrestrial habits (Gecarcinus Leach, 1814, Hartnollius n. gen. and Johngarthia Türkay, 1970) are highlighted in red. Ocypode rotundata Miers, 1882 was included as outgroup.	FIG. 16. — Bayesian inference phylogenetic tree of Gecarcinidae based on the concatenation of mitochondrial COI and 16S rRNA gene fragments.Numbers at nodes are support values for BI (posterior probability) and ML (Ultrafast bootstrap). The clade of crabs with less terrestrial habits (Cardisoma Latreille, 1828, Discoplax A.Milne-Edwards,1867,and Tuerkayana Guinot,N. K. Ng & Rodríguez Moreno, 2018) are highlighted in purple,whereas the clade with markedly terrestrial habits (Gecarcinus Leach, 1814, Hartnollius n. gen. and Johngarthia Türkay, 1970) are highlighted in red. Ocypode rotundata Miers, 1882 was included as outgroup.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481643FFF0BD1B596346C0FC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591308/files/figure.png	https://doi.org/10.5281/zenodo.15591308	FIG. 19. — Hartnollius lateralis (Fréminville in Guérin,1832) n. comb., living and in the field:A, Guadeloupe,Marie-Galante, KARUBENTHOS 3, Stn IGM25,MNHN- IU-2024-5551; B, C, Guadeloupe, July 2015, on sand; D, E, Martinique, Anse à Prunes, forest about 65 m far from the shore, altitude about 2 m; D, crab bearing a branch; E, crab on a tree. Credits: A, L. Corbari-MNHN-ARBig-LPRig; B, C, J. Poupin; D, E, F. Lavail (planetedoc.com).	FIG. 19. — Hartnollius lateralis (Fréminville in Guérin,1832) n. comb., living and in the field:A, Guadeloupe,Marie-Galante, KARUBENTHOS 3, Stn IGM25,MNHN- IU-2024-5551; B, C, Guadeloupe, July 2015, on sand; D, E, Martinique, Anse à Prunes, forest about 65 m far from the shore, altitude about 2 m; D, crab bearing a branch; E, crab on a tree. Credits: A, L. Corbari-MNHN-ARBig-LPRig; B, C, J. Poupin; D, E, F. Lavail (planetedoc.com).	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481643FFF0BD1B596346C0FC6A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15608609/files/figure.png	https://doi.org/10.5281/zenodo.15608609	FIG. 20. — Living Hartnollius quadratus (Saussure,1853) n. comb.:A, Mexico,Chiapas,Barra de Boca del Cielo;B, ‘Halloween crab’,Costa Rica,Nosara,on a tree, 2006; C, Panama, on the beach, 2005; D, E, Mexico, Chacahua, La Grúa: two views. Credits: A, J. L. Villalobos Hiriart; B, Bhny, Creative Commons Attribution- Share-Alike License 3.0; C, Dozenist, Creative Commons Attribution-Share-Alike License 3.0; D, E, A. López-Velázquez.	FIG. 20. — Living Hartnollius quadratus (Saussure,1853) n. comb.:A, Mexico,Chiapas,Barra de Boca del Cielo;B, ‘Halloween crab’,Costa Rica,Nosara,on a tree, 2006; C, Panama, on the beach, 2005; D, E, Mexico, Chacahua, La Grúa: two views. Credits: A, J. L. Villalobos Hiriart; B, Bhny, Creative Commons Attribution- Share-Alike License 3.0; C, Dozenist, Creative Commons Attribution-Share-Alike License 3.0; D, E, A. López-Velázquez.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481641FFCBB8265F414390FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591272/files/figure.png	https://doi.org/10.5281/zenodo.15591272	FIG. 5. — Specimens from mainland America identified by Rathbun as Gecarcinus ruricola (Linnaeus, 1758), but being in fact Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb. A, B, ♂ 32.3 × 40.5 mm, Florida, Florida Keys, Loggerhead Key, USNM 71219: A, general dorsal view; B, ventral view; C, D, ♂ 32.4 × 44.2 mm, Nicaragua, Greytown, Rathbun det. Gecarcinus ruricola; labels by Carson and Barnwell indicating G. lateralis, USNM 74612; C, general dorsal view; D, ventral view. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	FIG. 5. — Specimens from mainland America identified by Rathbun as Gecarcinus ruricola (Linnaeus, 1758), but being in fact Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb. A, B, ♂ 32.3 × 40.5 mm, Florida, Florida Keys, Loggerhead Key, USNM 71219: A, general dorsal view; B, ventral view; C, D, ♂ 32.4 × 44.2 mm, Nicaragua, Greytown, Rathbun det. Gecarcinus ruricola; labels by Carson and Barnwell indicating G. lateralis, USNM 74612; C, general dorsal view; D, ventral view. Scale bars: 10 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481641FFCBB8265F414390FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591280/files/figure.png	https://doi.org/10.5281/zenodo.15591280	FIG. 9. — Hartnollius n. gen., general views:A, H. lateralis (Fréminville in Guérin,1832) n. comb.,♂ 42.0 × 56.0 mm,Guadeloupe, MNHN-IU-2017-8390 (= MNHN- B24656); B, H. quadratus (Saussure, 1853) n. comb., ♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900). Scale bars: 20 mm. Credits: MNHN- Soubzmaigne.	FIG. 9. — Hartnollius n. gen., general views:A, H. lateralis (Fréminville in Guérin,1832) n. comb.,♂ 42.0 × 56.0 mm,Guadeloupe, MNHN-IU-2017-8390 (= MNHN- B24656); B, H. quadratus (Saussure, 1853) n. comb., ♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900). Scale bars: 20 mm. Credits: MNHN- Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481641FFCBB8265F414390FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591284/files/figure.png	https://doi.org/10.5281/zenodo.15591284	FIG. 10. — A-C, frontal views,D-F, chelipeds:A, Gecarcinus ruricola (Linnaeus,1758),neotype,♂ 55.0 × 76.0 mm,Cuba,MNHN-IU-2017-8392 (= MNHN-B13155); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126; C, Hartnollius quadratus (Saussure, 1853) n. comb., ♀ 36.0 × 46.0 mm, Mexico, CNCR 24607; D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): see granules on margins of merus and carpus of chelipeds; E, Hartnollius lateralis n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126: see smooth margins of chelipeds; F, Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 34630: see smooth margins of chelipeds. Scale bars: 10 mm. Credits: A, D, MNHN-Soubzmaigne; B, C, E, F, CNCR-Toledano-Carrasco.	FIG. 10. — A-C, frontal views,D-F, chelipeds:A, Gecarcinus ruricola (Linnaeus,1758),neotype,♂ 55.0 × 76.0 mm,Cuba,MNHN-IU-2017-8392 (= MNHN-B13155); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126; C, Hartnollius quadratus (Saussure, 1853) n. comb., ♀ 36.0 × 46.0 mm, Mexico, CNCR 24607; D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): see granules on margins of merus and carpus of chelipeds; E, Hartnollius lateralis n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126: see smooth margins of chelipeds; F, Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 34630: see smooth margins of chelipeds. Scale bars: 10 mm. Credits: A, D, MNHN-Soubzmaigne; B, C, E, F, CNCR-Toledano-Carrasco.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481641FFCBB8265F414390FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591287/files/figure.png	https://doi.org/10.5281/zenodo.15591287	FIG. 11. — Mxp3, internal view, and palp: A, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, MZUSP 18501 (ex USU-279); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Costa Rica, MZUSP 18509 (ex USU-281); C, H. quadratus (Saussure, 1853) n. comb., Mexico, MZUSP 18859 (ex AHF) (see Tavares 1989: fig. 14). Courtesy Marcos Tavares.	FIG. 11. — Mxp3, internal view, and palp: A, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, MZUSP 18501 (ex USU-279); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Costa Rica, MZUSP 18509 (ex USU-281); C, H. quadratus (Saussure, 1853) n. comb., Mexico, MZUSP 18859 (ex AHF) (see Tavares 1989: fig. 14). Courtesy Marcos Tavares.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481641FFCBB8265F414390FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591289/files/figure.png	https://doi.org/10.5281/zenodo.15591289	FIG. 12. — Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656): A, frontal view; B, front, orbit,and mesial lobe of infraorbital margin;C, front,proepistome (hidden), antenna (exposed) and mesial lobe of infraorbital margin; D, thoracic sternum and male pleon; E, F, external and internal views of chelipeds. Scale bars: A, D, E, F, 10 mm; B, C, 5 mm. Credits: MNHN-Soubzmaigne.	FIG. 12. — Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 42.0 × 56.0 mm, Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656): A, frontal view; B, front, orbit,and mesial lobe of infraorbital margin;C, front,proepistome (hidden), antenna (exposed) and mesial lobe of infraorbital margin; D, thoracic sternum and male pleon; E, F, external and internal views of chelipeds. Scale bars: A, D, E, F, 10 mm; B, C, 5 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481641FFCBB8265F414390FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591292/files/figure.png	https://doi.org/10.5281/zenodo.15591292	FIG. 13. — Harrtnollius n. gen.: gonopods and vulvae:A, B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656): A, ♂ 40.5 × 55.7 mm: G1, G2, caudal view; B, ♀ 32.6 × 41.8 mm:vulvae;C, H.quadratus (Saussure,1853) n.comb.,♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900):G1, G2, caudal view. Scale bars: A, C, 2 mm; B, 1 mm. Credits: MNHN-Soubzmaigne.	FIG. 13. — Harrtnollius n. gen.: gonopods and vulvae:A, B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656): A, ♂ 40.5 × 55.7 mm: G1, G2, caudal view; B, ♀ 32.6 × 41.8 mm:vulvae;C, H.quadratus (Saussure,1853) n.comb.,♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900):G1, G2, caudal view. Scale bars: A, C, 2 mm; B, 1 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481641FFCBB8265F414390FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591302/files/figure.png	https://doi.org/10.5281/zenodo.15591302	FIG. 16. — Bayesian inference phylogenetic tree of Gecarcinidae based on the concatenation of mitochondrial COI and 16S rRNA gene fragments.Numbers at nodes are support values for BI (posterior probability) and ML (Ultrafast bootstrap). The clade of crabs with less terrestrial habits (Cardisoma Latreille, 1828, Discoplax A.Milne-Edwards,1867,and Tuerkayana Guinot,N. K. Ng & Rodríguez Moreno, 2018) are highlighted in purple,whereas the clade with markedly terrestrial habits (Gecarcinus Leach, 1814, Hartnollius n. gen. and Johngarthia Türkay, 1970) are highlighted in red. Ocypode rotundata Miers, 1882 was included as outgroup.	FIG. 16. — Bayesian inference phylogenetic tree of Gecarcinidae based on the concatenation of mitochondrial COI and 16S rRNA gene fragments.Numbers at nodes are support values for BI (posterior probability) and ML (Ultrafast bootstrap). The clade of crabs with less terrestrial habits (Cardisoma Latreille, 1828, Discoplax A.Milne-Edwards,1867,and Tuerkayana Guinot,N. K. Ng & Rodríguez Moreno, 2018) are highlighted in purple,whereas the clade with markedly terrestrial habits (Gecarcinus Leach, 1814, Hartnollius n. gen. and Johngarthia Türkay, 1970) are highlighted in red. Ocypode rotundata Miers, 1882 was included as outgroup.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
423782481641FFCBB8265F414390FC8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591308/files/figure.png	https://doi.org/10.5281/zenodo.15591308	FIG. 19. — Hartnollius lateralis (Fréminville in Guérin,1832) n. comb., living and in the field:A, Guadeloupe,Marie-Galante, KARUBENTHOS 3, Stn IGM25,MNHN- IU-2024-5551; B, C, Guadeloupe, July 2015, on sand; D, E, Martinique, Anse à Prunes, forest about 65 m far from the shore, altitude about 2 m; D, crab bearing a branch; E, crab on a tree. Credits: A, L. Corbari-MNHN-ARBig-LPRig; B, C, J. Poupin; D, E, F. Lavail (planetedoc.com).	FIG. 19. — Hartnollius lateralis (Fréminville in Guérin,1832) n. comb., living and in the field:A, Guadeloupe,Marie-Galante, KARUBENTHOS 3, Stn IGM25,MNHN- IU-2024-5551; B, C, Guadeloupe, July 2015, on sand; D, E, Martinique, Anse à Prunes, forest about 65 m far from the shore, altitude about 2 m; D, crab bearing a branch; E, crab on a tree. Credits: A, L. Corbari-MNHN-ARBig-LPRig; B, C, J. Poupin; D, E, F. Lavail (planetedoc.com).	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248167AFFC8BD6858214044F868.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591280/files/figure.png	https://doi.org/10.5281/zenodo.15591280	FIG. 9. — Hartnollius n. gen., general views:A, H. lateralis (Fréminville in Guérin,1832) n. comb.,♂ 42.0 × 56.0 mm,Guadeloupe, MNHN-IU-2017-8390 (= MNHN- B24656); B, H. quadratus (Saussure, 1853) n. comb., ♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900). Scale bars: 20 mm. Credits: MNHN- Soubzmaigne.	FIG. 9. — Hartnollius n. gen., general views:A, H. lateralis (Fréminville in Guérin,1832) n. comb.,♂ 42.0 × 56.0 mm,Guadeloupe, MNHN-IU-2017-8390 (= MNHN- B24656); B, H. quadratus (Saussure, 1853) n. comb., ♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900). Scale bars: 20 mm. Credits: MNHN- Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248167AFFC8BD6858214044F868.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591284/files/figure.png	https://doi.org/10.5281/zenodo.15591284	FIG. 10. — A-C, frontal views,D-F, chelipeds:A, Gecarcinus ruricola (Linnaeus,1758),neotype,♂ 55.0 × 76.0 mm,Cuba,MNHN-IU-2017-8392 (= MNHN-B13155); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126; C, Hartnollius quadratus (Saussure, 1853) n. comb., ♀ 36.0 × 46.0 mm, Mexico, CNCR 24607; D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): see granules on margins of merus and carpus of chelipeds; E, Hartnollius lateralis n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126: see smooth margins of chelipeds; F, Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 34630: see smooth margins of chelipeds. Scale bars: 10 mm. Credits: A, D, MNHN-Soubzmaigne; B, C, E, F, CNCR-Toledano-Carrasco.	FIG. 10. — A-C, frontal views,D-F, chelipeds:A, Gecarcinus ruricola (Linnaeus,1758),neotype,♂ 55.0 × 76.0 mm,Cuba,MNHN-IU-2017-8392 (= MNHN-B13155); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126; C, Hartnollius quadratus (Saussure, 1853) n. comb., ♀ 36.0 × 46.0 mm, Mexico, CNCR 24607; D, Gecarcinus ruricola (Linnaeus, 1758), neotype, ♂ 55.0 × 76.0 mm, Cuba, MNHN-IU-2017-8392 (= MNHN-B13155): see granules on margins of merus and carpus of chelipeds; E, Hartnollius lateralis n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 35126: see smooth margins of chelipeds; F, Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 43.0 × 55.0 mm, Mexico, CNCR 34630: see smooth margins of chelipeds. Scale bars: 10 mm. Credits: A, D, MNHN-Soubzmaigne; B, C, E, F, CNCR-Toledano-Carrasco.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248167AFFC8BD6858214044F868.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591287/files/figure.png	https://doi.org/10.5281/zenodo.15591287	FIG. 11. — Mxp3, internal view, and palp: A, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, MZUSP 18501 (ex USU-279); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Costa Rica, MZUSP 18509 (ex USU-281); C, H. quadratus (Saussure, 1853) n. comb., Mexico, MZUSP 18859 (ex AHF) (see Tavares 1989: fig. 14). Courtesy Marcos Tavares.	FIG. 11. — Mxp3, internal view, and palp: A, Gecarcinus ruricola (Linnaeus, 1758), Jamaica, MZUSP 18501 (ex USU-279); B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Costa Rica, MZUSP 18509 (ex USU-281); C, H. quadratus (Saussure, 1853) n. comb., Mexico, MZUSP 18859 (ex AHF) (see Tavares 1989: fig. 14). Courtesy Marcos Tavares.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248167AFFC8BD6858214044F868.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591292/files/figure.png	https://doi.org/10.5281/zenodo.15591292	FIG. 13. — Harrtnollius n. gen.: gonopods and vulvae:A, B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656): A, ♂ 40.5 × 55.7 mm: G1, G2, caudal view; B, ♀ 32.6 × 41.8 mm:vulvae;C, H.quadratus (Saussure,1853) n.comb.,♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900):G1, G2, caudal view. Scale bars: A, C, 2 mm; B, 1 mm. Credits: MNHN-Soubzmaigne.	FIG. 13. — Harrtnollius n. gen.: gonopods and vulvae:A, B, Hartnollius lateralis (Fréminville in Guérin, 1832) n. comb., Guadeloupe, MNHN-IU-2017-8390 (= MNHN-B24656): A, ♂ 40.5 × 55.7 mm: G1, G2, caudal view; B, ♀ 32.6 × 41.8 mm:vulvae;C, H.quadratus (Saussure,1853) n.comb.,♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900):G1, G2, caudal view. Scale bars: A, C, 2 mm; B, 1 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248167AFFC8BD6858214044F868.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591296/files/figure.png	https://doi.org/10.5281/zenodo.15591296	FIG. 14. — Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900): A, frontal view; B, front, proepistome (hidden), antenna (exposed) and mesial lobe of infraorbital margin; C, thoracic sternum and male pleon; D, E, external and internal views of cheli- peds. Scale bars: A, C, D, E, 20 mm; B, 2 mm. Credits: MNHN-Soubzmaigne.	FIG. 14. — Hartnollius quadratus (Saussure, 1853) n. comb., ♂ 50.0 × 64.0 mm, Mexico, MNHN-IU-2017-8391 (= MNHN-B20900): A, frontal view; B, front, proepistome (hidden), antenna (exposed) and mesial lobe of infraorbital margin; C, thoracic sternum and male pleon; D, E, external and internal views of cheli- peds. Scale bars: A, C, D, E, 20 mm; B, 2 mm. Credits: MNHN-Soubzmaigne.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248167AFFC8BD6858214044F868.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591302/files/figure.png	https://doi.org/10.5281/zenodo.15591302	FIG. 16. — Bayesian inference phylogenetic tree of Gecarcinidae based on the concatenation of mitochondrial COI and 16S rRNA gene fragments.Numbers at nodes are support values for BI (posterior probability) and ML (Ultrafast bootstrap). The clade of crabs with less terrestrial habits (Cardisoma Latreille, 1828, Discoplax A.Milne-Edwards,1867,and Tuerkayana Guinot,N. K. Ng & Rodríguez Moreno, 2018) are highlighted in purple,whereas the clade with markedly terrestrial habits (Gecarcinus Leach, 1814, Hartnollius n. gen. and Johngarthia Türkay, 1970) are highlighted in red. Ocypode rotundata Miers, 1882 was included as outgroup.	FIG. 16. — Bayesian inference phylogenetic tree of Gecarcinidae based on the concatenation of mitochondrial COI and 16S rRNA gene fragments.Numbers at nodes are support values for BI (posterior probability) and ML (Ultrafast bootstrap). The clade of crabs with less terrestrial habits (Cardisoma Latreille, 1828, Discoplax A.Milne-Edwards,1867,and Tuerkayana Guinot,N. K. Ng & Rodríguez Moreno, 2018) are highlighted in purple,whereas the clade with markedly terrestrial habits (Gecarcinus Leach, 1814, Hartnollius n. gen. and Johngarthia Türkay, 1970) are highlighted in red. Ocypode rotundata Miers, 1882 was included as outgroup.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248167AFFC8BD6858214044F868.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15608609/files/figure.png	https://doi.org/10.5281/zenodo.15608609	FIG. 20. — Living Hartnollius quadratus (Saussure,1853) n. comb.:A, Mexico,Chiapas,Barra de Boca del Cielo;B, ‘Halloween crab’,Costa Rica,Nosara,on a tree, 2006; C, Panama, on the beach, 2005; D, E, Mexico, Chacahua, La Grúa: two views. Credits: A, J. L. Villalobos Hiriart; B, Bhny, Creative Commons Attribution- Share-Alike License 3.0; C, Dozenist, Creative Commons Attribution-Share-Alike License 3.0; D, E, A. López-Velázquez.	FIG. 20. — Living Hartnollius quadratus (Saussure,1853) n. comb.:A, Mexico,Chiapas,Barra de Boca del Cielo;B, ‘Halloween crab’,Costa Rica,Nosara,on a tree, 2006; C, Panama, on the beach, 2005; D, E, Mexico, Chacahua, La Grúa: two views. Credits: A, J. L. Villalobos Hiriart; B, Bhny, Creative Commons Attribution- Share-Alike License 3.0; C, Dozenist, Creative Commons Attribution-Share-Alike License 3.0; D, E, A. López-Velázquez.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248167FFFC3BD7B5BA247A2F868.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591300/files/figure.png	https://doi.org/10.5281/zenodo.15591300	FIG. 15. — Hartnollius nobilii (Perger & Wall, 2014) n. comb., paratype, ♀ 36.6 × 44.3 mm, Ecuador, MNHN-IU-2014-11211 (= MNHN-B12314): A, dorsal view; B, ventral view; C, front, orbit, and mesial lobe of infraorbital margin; D, front, proepistome (hidden), antenna (exposed), mesial lobe of infraorbital margin, and mxp3; E, thoracic sternum, female pleon and vulvae; F, left cheliped, internal view; G, chelipeds, external view; H, right P3; I, left P4. Scale bar: 10 mm. Credits: MNHN-Rodríguez Moreno.	FIG. 15. — Hartnollius nobilii (Perger & Wall, 2014) n. comb., paratype, ♀ 36.6 × 44.3 mm, Ecuador, MNHN-IU-2014-11211 (= MNHN-B12314): A, dorsal view; B, ventral view; C, front, orbit, and mesial lobe of infraorbital margin; D, front, proepistome (hidden), antenna (exposed), mesial lobe of infraorbital margin, and mxp3; E, thoracic sternum, female pleon and vulvae; F, left cheliped, internal view; G, chelipeds, external view; H, right P3; I, left P4. Scale bar: 10 mm. Credits: MNHN-Rodríguez Moreno.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
42378248167FFFC3BD7B5BA247A2F868.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15591302/files/figure.png	https://doi.org/10.5281/zenodo.15591302	FIG. 16. — Bayesian inference phylogenetic tree of Gecarcinidae based on the concatenation of mitochondrial COI and 16S rRNA gene fragments.Numbers at nodes are support values for BI (posterior probability) and ML (Ultrafast bootstrap). The clade of crabs with less terrestrial habits (Cardisoma Latreille, 1828, Discoplax A.Milne-Edwards,1867,and Tuerkayana Guinot,N. K. Ng & Rodríguez Moreno, 2018) are highlighted in purple,whereas the clade with markedly terrestrial habits (Gecarcinus Leach, 1814, Hartnollius n. gen. and Johngarthia Türkay, 1970) are highlighted in red. Ocypode rotundata Miers, 1882 was included as outgroup.	FIG. 16. — Bayesian inference phylogenetic tree of Gecarcinidae based on the concatenation of mitochondrial COI and 16S rRNA gene fragments.Numbers at nodes are support values for BI (posterior probability) and ML (Ultrafast bootstrap). The clade of crabs with less terrestrial habits (Cardisoma Latreille, 1828, Discoplax A.Milne-Edwards,1867,and Tuerkayana Guinot,N. K. Ng & Rodríguez Moreno, 2018) are highlighted in purple,whereas the clade with markedly terrestrial habits (Gecarcinus Leach, 1814, Hartnollius n. gen. and Johngarthia Türkay, 1970) are highlighted in red. Ocypode rotundata Miers, 1882 was included as outgroup.	2025-06-03	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba		Zenodo	biologists	Paula, Danièle Guinot;Rodríguez, Paula A.;Atzimba, Moreno I.;Toledano-Carrasco, Atzimba			
