taxonID	type	description	language	source
3C6E87D6924B284CFF14FE2AFCDCFDB8.taxon	materials_examined	Type locality. “ Philippines, nr Baguio, Asin Hot-Spring. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FE2AFCDCFDB8.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Ochi et al. 2016). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FD12FCDCFCE0.taxon	materials_examined	Type locality. “ Philippines, Luzon, nr Manila, Cavite. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FD12FCDCFCE0.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Ochi et al. 2016). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FCFAFC27FBC8.taxon	materials_examined	Type locality. “ Philippines, Luzon, nr Manila, Cavite. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FCFAFC27FBC8.taxon	distribution	Distribution in the Philippines. Luzon (Cavite) (Ochi et al. 2016). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FBA2FC27FB30.taxon	materials_examined	Type locality. “ Philippines, Luzon, nr Manila, Cavite. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FBA2FC27FB30.taxon	distribution	Distribution in the Philippines. Luzon (Cavite) (Ochi et al. 2016). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FA8AFCDCFA18.taxon	materials_examined	Type locality. “ Philippines, C. Luzon, Kinabuhayan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FA8AFCDCFA18.taxon	distribution	Distribution in the Philippines. Luzon (Quezon Province) (Ochi et al. 2016). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FA72FCDCF940.taxon	materials_examined	Type locality. “ Mt. Matotom, Mindanao Is., the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FA72FCDCF940.taxon	distribution	Distribution in the Philippines. Mindanao (South Cotabato) (Ochi et al. 2010; Král et al. 2014). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14F95AFCDCF8A8.taxon	materials_examined	Type locality. “ Mt. Halcon, Mindro [= Mindoro] Is., the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14F95AFCDCF8A8.taxon	distribution	Distribution in the Philippines. Luzon (Ochi et al. 2010; Král et al. 2014). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ Philippines, Negros Is., Manbucal. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Visayas (Ochi et al. 2016). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FEF3FE8DFD5D.taxon	materials_examined	Type locality. “ Yünnan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FEF3FE8DFD5D.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Balthasar 1964; Stebnicka 1986). Overall distribution. The Philippines, China, Nepal, India, Indonesia, Laos, and Thailand (Král et al. 2014).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FD2FFCEBFBC9.taxon	materials_examined	Type locality. “ Java. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FD2FFCEBFBC9.taxon	distribution	Distribution in the Philippines. Luzon and Visayas (Leyte) (Schmidt 1913; Stebnicka 1993; Ochi et al. 2016). Overall distribution. The Philippines, Australia, Indonesia, Malaysia, Thailand, and Vietnam (Stebnicka 1993; Dellacasa & Dellacasa 2006 a; Ochi et al. 2016).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FBA3FCDCFB31.taxon	materials_examined	Type locality. “ Kinabuhayan, Luzon, the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FBA3FCDCFB31.taxon	distribution	Distribution in the Philippines. Luzon (Ochi et al. 2010; Král et al. 2014). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FAC3FCCCF941.taxon	materials_examined	Type locality. “ Tonkin, Hoa Bin et India or., Madras. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FAC3FCCCF941.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Petrovitz 1961 a). Overall distribution. The Philippines, India, Thailand, Sri Lanka, Vietnam, Bhutan, Nepal, and China (including Taiwan) (Stebnicka 1989; Král & Šípek 2013).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14F893FCDCF805.taxon	distribution	Distribution in the Philippines. Luzon (Benguet and Laguna) and Visayas (Leyte) (Petrovitz 1962; Balthasar 1964; Dellacasa 1983). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FF43FD91FE16.taxon	materials_examined	Type locality. “ Die philippinischen Inseln, Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FF43FD91FE16.taxon	distribution	Distribution in the Philippines. Luzon (Harold 1862 b; Boucomont 1914; Stebnicka 1986). Overall distribution. The Philippines, China (Taiwan), Nepal, India, Indonesia, Sri Lanka, Thailand, and Vietnam (Stebnicka 1986).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FE60FAD7FD7E.taxon	materials_examined	Type locality. “ Java. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FE60FAD7FD7E.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993). Overall distribution. The Philippines, Indonesia, Malaysia, and Papua New Guinea (Stebnicka 1993).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FD49FEC5FBD7.taxon	materials_examined	Type locality. “ Coromandel. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FD49FEC5FBD7.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Boucomont 1914; Ochi & Kon 1995). Overall distribution. The Philippines, Bhutan, China (Taiwan), India, Japan, Nepal, Pakistan, Indonesia, Malaysia, Myanmar, Thailand, Vietnam, and Papua New Guinea (Petrovitz 1961; Stebnicka 1992; Ochi & Kon 1995).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FBA6FDE1FAAC.taxon	materials_examined	Type locality. “ Burma. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FBA6FDE1FAAC.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Petrovitz 1961 b). Overall distribution. The Philippines, Cambodia, Laos, Myanmar, Thailand, Vietnam, Japan, Korea, Nepal, and China (including Taiwan) (Petrovitz 1961 b; Stebnicka 1992).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FBA6FDE1FAAC.taxon	discussion	Remarks. This species is widely distributed in the Oriental region, and its “ reported ” occurrence in the Philippines is likely erroneous.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FA74FCDCF94A.taxon	materials_examined	Type locality. “ Philippinen, Luzon, Buranen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FA74FCDCF94A.taxon	distribution	Distribution in the Philippines. Luzon (Petrovitz 1972). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14F95DFCDCF8B3.taxon	materials_examined	Type locality. “ Manila. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14F95DFCDCF8B3.taxon	distribution	Distribution in the Philippines. Luzon (Manila) (Schmidt 1909). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692492841FF14F80AFCDCFF0C.taxon	materials_examined	Type locality. “ Leyte, Visca N Baybay. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692492841FF14F80AFCDCFF0C.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FC4EFA92FAE8.taxon	materials_examined	Type locality. “ Nova Hollandia (Sydney). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FC4EFA92FAE8.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993). Overall distribution. The Philippines, Australia, Japan, China, Indonesia, Malaysia, Myanmar, Thailand, and Vietnam (Stebnicka 1993; Stebnicka & Howden 1997).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FC4EFA92FAE8.taxon	discussion	Remarks. The natural occurrence of this species in the Philippines is doubtful. Although it is commonly found and widely distributed across Australia and mainland Asia, its presence in the Philippines remains highly questionable. Notably, Stebnicka & Howden (1997) in their revision of Ataenius omitted this species from the Philippines, suggesting that earlier records may have resulted from misidentifications.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FAC2FD0CF9AC.taxon	materials_examined	Type locality. “ Mexico. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FAC2FD0CF9AC.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993). Overall distribution. The Philippines, Australian, Oriental, Micronesia, and Nearctic regions (Stebnicka & Howden 1997; Stebnicka & Lago 2005).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14F93EFCF2F84C.taxon	materials_examined	Type locality. “ Celebes, Macassar; Sourabaia; Borneo, Sarawak. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14F93EFCF2F84C.taxon	distribution	Distribution in the Philippines. Luzon (? Manila) and Visayas (Leyte) (Schmidt 1922; Stebnicka 1993). Overall distribution. The Philippines, Australia, Pacific, Oriental, Palearctic, Nearctic, and Neotropical (Stebnicka & Howden 1997; Théry & Bordat 2012).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FED7FCDCFE2C.taxon	materials_examined	Type locality. “ Mindanao, Momungan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FED7FCDCFE2C.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Norte) (Červenka 1994). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FDBFFC4DFCCC.taxon	materials_examined	Type locality. “ Ins. Philippinć. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FDBFFC4DFCCC.taxon	distribution	Distribution. Philippine archipelago (Harold 1860; Balthasar 1964). Overall distribution. The Philippines and Indonesia (Červenka 1994).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FF43FB83FE51.taxon	materials_examined	Type locality. “ Luzon (Imugan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FF43FB83FE51.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Paulian 1944). Overall distribution. The Philippines and China (Taiwan) (Nomura 1973).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FD13FE0DFC59.taxon	materials_examined	Type locality. “ Iles Key. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FD13FE0DFC59.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993). Overall distribution. The Philippines, Indonesia, Christmas Island, Papua New Guinea, and Hawaii (Stebnicka 2012).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FE2BFB42FDB9.taxon	materials_examined	Type locality. “ Sumatra. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FE2BFB42FDB9.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Stebnicka 2012). Overall distribution. The Philippines, Indonesia, Malaysia, and Thailand (Stebnicka 2012).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FC33FB5DFB81.taxon	materials_examined	Type locality. “ Mentawei. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FC33FB5DFB81.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Stebnicka 2012). Overall distribution. The Philippines, Thailand, Indonesia, and Malaysia (Stebnicka 2012).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FB53FCDCFAA1.taxon	materials_examined	Type locality. “ Leyte, Visca N Baybay. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FB53FCDCFAA1.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FA3BFCDCF989.taxon	materials_examined	Type locality. “ Mindanao, N. Mindanao Region, Bukidnon: Kalatungan Mts. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FA3BFCDCF989.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur) and Mindanao (Bukidnon) (Skelley 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14F95BFCDCF8A9.taxon	materials_examined	Type locality. “ Philippines, Palawan Is., Roxas. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14F95BFCDCF8A9.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ochi et al. 2025 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FEF3FCDCFDB9.taxon	materials_examined	Type locality. “ Mindoro, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FEF3FCDCFDB9.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro and Palawan) and Visayas (Leyte) (Rakovič 1987; Stebnicka 1993). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FD13FCDCFCE1.taxon	materials_examined	Type locality. “ Philippines, Panay. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FD13FCDCFCE1.taxon	distribution	Distribution in the Philippines. Visayas (Panay) (Rakovič 1997). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FCFBFCDCFBC9.taxon	materials_examined	Type locality. “ Philipp., Mindanao, Siocom. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FCFBFCDCFBC9.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Rakovič 1997). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FBA3FCDCFB31.taxon	materials_examined	Type locality. “ Philippines, SW Panay. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FBA3FCDCFB31.taxon	distribution	Distribution in the Philippines. Visayas (Panay) (Rakovič 1997). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FA8BFCDCFA3D.taxon	materials_examined	Type locality. “ NE Leyte Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FA8BFCDCFA3D.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993; Rakovič 1987). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14F9C7FCD4F8DD.taxon	materials_examined	Type locality. “ Philippines, Mindanao, Maragusan, Marangig Falls, Candalaga Mts. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14F9C7FCD4F8DD.taxon	distribution	Distribution in the Philippines. Mindanao (Davao de Oro) (Rakovič & Anichtchenko 2021). Overall distribution. Endemic species to the Philippines	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442842FF14F85FFE8DFE51.taxon	materials_examined	Type locality. “ Sumatra, Java, Borneo et aux Celebes. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442842FF14F85FFE8DFE51.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Rakovič et al. 2023). Overall distribution. The Philippines, China (Taiwan), Thailand, Malaysia, Indonesia, French Polynesia, Australia, Fiji, Papua New Guinea, Samoa, Solomon Islands, Seychelles, and Mauritius (Rakovič et al, 2023).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14FE63FCDDFD71.taxon	materials_examined	Type locality. “ Davao, Philippines Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14FE63FCDDFD71.taxon	distribution	Distribution in the Philippines. Mindanao (Davao) (Pittino & Mariani 1986). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14F9C7FCDCF8DD.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is. Mt. Apo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14F9C7FCDCF8DD.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14F8AFFCDCF805.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is. Mt. Apo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14F8AFFCDCF805.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14FC83FDB3FC35.taxon	materials_examined	Type locality. “ India (Bengal). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14FC83FDB3FC35.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Pittino 1984). Overall distribution. The Philippines, China (including Taiwan), India, Thailand, Vietnam, Iran, Nepal, and Pakistan (Pittino 1984).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14FBFFFE6BFA45.taxon	materials_examined	Type locality. “ Ceylon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14FBFFFE6BFA45.taxon	distribution	Distribution in the Philippines. Luzon (Makati and Metro Manila), Visayas (Cebu and Leyte), and Mindanao (Zamboanga del Norte) (Masumoto 1980; Pittino 1984; Stebnicka 1993; Pittino et al. 2013). Overall distribution. The Philippines, Palearctic, Oriental, Madagascan, Australian, and Pacific regions (Pittino et al. 2013).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FF0BFCDCFE98.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., Baracatan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FF0BFCDCFE98.taxon	distribution	Distribution in the Philippines. Mindanao (Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FEF3FBF8FDE4.taxon	materials_examined	Type locality. “ Lanyu Is. Taiwan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FEF3FBF8FDE4.taxon	distribution	Distribution in the Philippines. Luzon (Ifugao) and Mindanao (Bukidnon, Davao, and South Cotabato) (Anichtchenko et al. 2021; Ochi et al. 2021). Overall distribution. The Philippines and China (Taiwan) (Ochi et al. 2021).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FDF7FCDCFCCC.taxon	materials_examined	Type locality. “ Philippines, Basilan Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FDF7FCDCFCCC.taxon	distribution	Distribution in the Philippines. Mindanao (Basilan) (Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FCDFFCDDFBC8.taxon	materials_examined	Type locality. “ Mindanao, Lake Holon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FCDFFCDDFBC8.taxon	distribution	Distribution in the Philippines. Mindanao (South Cotabato and Mt. Apo) (Anichtchenko et al. 2021; Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FBA3FAECFB30.taxon	materials_examined	Type locality. “ Louisiade Archipelago, Sud-Est Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FBA3FAECFB30.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon and Davao) (Anichtchenko et al. 2021). Overall distribution. The Philippines, Papua New Guinea, and Indonesia (Anichtchenko et al. 2021).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FA8BFD99F9AC.taxon	materials_examined	Type locality. “ Borneo occ.: Sambas; Java: Simpar et Kemanglen rés. Tega. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FA8BFD99F9AC.taxon	distribution	Distribution in the Philippines. Luzon (Baguio and Camarines Sur), Visayas (Negros), and Mindanao (Bukidnon, Davao, and Cotabato) (Paulian 1981; Théry & Bordat 2012; Ochi et al. 2021). Overall distribution. The Philippines, Korea, Australia, Indonesia, Japan, China (including Taiwan), Papua New Guinea, New Caledonia, Solomon Islands, Malaysia, New South Wales, and Queensland (Anichtchenko et al. 2021).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14F93FFCDCF894.taxon	materials_examined	Type locality. “ Philippines, Luzon, Ifugao Prov., Dalican. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14F93FFCDCF894.taxon	distribution	Distribution in the Philippines. Luzon (Ifugao) (Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422844FF14F8E7FCDCFF0C.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Quirino, Madera, Sierra Madre Disimungal. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422844FF14F8E7FCDCFF0C.taxon	distribution	Distribution in the Philippines. Luzon (Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FE9FFCDCFE08.taxon	materials_examined	Type locality. “ Philippines, Mindanao, Maragusan, Marangig Falls, Candalaga Mts. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FE9FFCDCFE08.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) and Mindanao (Basilan, Bukidnon, and Davao de Oro) (Anichtchenko et al. 2021; Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FE63FCDCFD70.taxon	materials_examined	Type locality. “ Philippines, Negros Is., Mt. Mandalagan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FE63FCDCFD70.taxon	distribution	Distribution in the Philippines. Visayas (Negros Island) (Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FD4BFCDCFC58.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., Caraga Region, Agusan del sur, Sibagat. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FD4BFCDCFC58.taxon	distribution	Distribution in the Philippines. Mindanao (Agusan del Sur and Bukidnon) (Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FC33FE8AFA8C.taxon	materials_examined	Type locality. “ Malay Peninsula, Penang, Perak. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FC33FE8AFA8C.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993). Overall distribution. The? Philippines, Malaysia, and? China (Taiwan).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FC33FE8AFA8C.taxon	discussion	Remarks. The authors have reviewed the articles of current authors (Mencl et al. 2013; Anichtchenko et al. 2021; Ochi et al. 2021), and it appears that the occurrence of Rhyparus peninsularis in the Philippines is doubtful. Perhaps they are unaware of Stebnicka’s work, or due to the lack of substantial specimens to examine within the archipelago. Notably, Anichtchenko et al. (2021) explicitly listed only two species of Rhyparus (R. helophoroides and R. philippinensis) occurring in the Philippines, suggesting that R. peninsularis was either excluded due to the lack of confirmed specimens or was unaware of Stebnicka’s work.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FA1FFCDCF940.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FA1FFCDCF940.taxon	distribution	Distribution in the Philippines. Luzon (Cagayan Province, Camarines Sur, and Mountain Province), Visayas (Samar), and Mindanao (Davao de Oro, Lanao del Norte, and Lanao del Sur) (Arrow 1905; Anichtchenko et al. 2021; Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14F893FCDCF804.taxon	materials_examined	Type locality. “ Philippines: Mt. Makiling, Laguna Prov. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14F893FCDCF804.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Howden 1995; Skelley 2007). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FDF5FCDCFCEF.taxon	materials_examined	Type locality. “ Mindanao (Dapitan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FDF5FCDCFCEF.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte and Davao) (Moser 1921 b; Sakai 1995; Legrand 2020). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FCFEFCDCFB8C.taxon	materials_examined	Type locality. “ Mt. Dimatabing, Luzon, Tayabas. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FCFEFCDCFB8C.taxon	distribution	Distribution in the Philippines. Luzon (Quezon) (Mikšić 1979; Legrand 2020). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FB1FFCDCFAF5.taxon	materials_examined	Type locality. “ Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FB1FFCDCFAF5.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Moser 1918). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FAC7FC27F9DA.taxon	materials_examined	Type locality. “ Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FAC7FC27F9DA.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Sakai 1995). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14F95DFCDFF823.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14F95DFCDFF823.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur, Ilocos, Laguna, Mt. Banahao, Pampanga, Masbate, and Rizal) and Visayas (Negros Oriental and Panay) (Wallace 1867; Mikšić 1970 b 1970 c; Pavićević 1988). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FF0BFCDDFEBD.taxon	materials_examined	Type locality. “ Philippines, Mindanao Island, Alamada, North Cotabato. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FF0BFCDDFEBD.taxon	distribution	Distribution in the Philippines. Mindanao (Agusan del Sur, Davao de Oro, Lanao del Sur, North and South Cotabato) (Minkina & Kakizoe 2020). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FF0BFCDCFE98.taxon	materials_examined	Type locality. “ Central Visayas, Negros Oriental, Dumaguete. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FF0BFCDCFE98.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Legrand 2018 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FEF3FC27FDC0.taxon	materials_examined	Type locality. “ Mindoro, Mt. Halcon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FEF3FC27FDC0.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Legrand 2018 a). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FDDBFC27FD28.taxon	materials_examined	Type locality. “ Palawan, Brook’s Point. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FDDBFC27FD28.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Antoine 1996; Legrand 2018 a). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FC83FCDCFC34.taxon	materials_examined	Type locality. “ Masbate-Aroroy. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FC83FCDCFC34.taxon	distribution	Distribution in the Philippines. Luzon (Masbate) (Mikšić 1967). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FB87FCDCFAD4.taxon	materials_examined	Type locality. “ Panay (Northwestern part). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FB87FCDCFAD4.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) and Visayas (Negros and Panay) (Moser 1925; Mikšić 1970 b; Legrand 2018 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FAA7FD53F84C.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FAA7FD53F84C.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur and Nueva Vizcaya) and Mindanao (Mohnike 1873; Mikšić 1970 b, 1970 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FAA7FD53F84C.taxon	discussion	Remarks. When Mikšić was studying what is now recognized as the Reineria group, several rounds of revision occurred before it was formally erected to genus rank (Arnaud 2004). In his early work, Reineria Mikšić (1968) was treated as a subgenus of Glycyphana Burmeister (1842), which was synonymized with the genus Astraea Mohnike, 1873. Many species were subsequently reclassified under the genus Reineria. However, for this particular species, the authors were unable to locate the original description supporting its current taxonomic placement. Legrand (2018 a) did not include this species in his catalog of the sub genus Caloglycyphana Mikšić (1968). This species will probably be revised and formally placed in the genus Reineria Mikšić (1968). Furthermore, Mikšić noted that, based on his examination of the available material and the type specimen from the original description, this species exhibits slight morphological differences that may indicate subspeciation.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FF0BFC27FEBD.taxon	materials_examined	Type locality. “ Leyte, Philippines Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FF0BFC27FEBD.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Janson 1881). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FE0FFC27FD15.taxon	materials_examined	Type locality. “ Mindanao parte occidentali. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FE0FFC27FD15.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo and Surigao) (Mohnike 1873; Mikšić 1970 b, 1970 c, 1984). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FCDFFD59FC35.taxon	materials_examined	Type locality. “ Philippines, Oriental Negros, Mt. Canlaon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FCDFFD59FC35.taxon	distribution	Distribution in the Philippines. Visayas (Negros and Panay) (Arnaud, 2019). Overall distribution. Only in the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FB87FC27FAD5.taxon	materials_examined	Type locality. “ Philippines Luzon: Imugan et Banahao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FB87FC27FAD5.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao and Nueva Vizcaya) (Bourgoin 1930; Mikšić 1970 b, 1970 c). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FAA7FC27F9F5.taxon	materials_examined	Type locality. “ Luzon (Mt. Isarog). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FAA7FC27F9F5.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao, Romblon, and Camarines Sur) (Moser 1917 a; Mikšić 1970 b, 1970 c, 1984). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14F9C7FCDCF895.taxon	materials_examined	Type locality. “ Mindanao (Momungan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14F9C7FCDCF895.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Norte and Surigao) (Moser 1922 a; Mikšić 1970 b, 1970 c; Arnaud 2004). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2858FF14F8E7FCDCFE50.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2858FF14F8E7FCDCFE50.taxon	distribution	Distribution in the Philippines. Luzon (Albay, Camarines Sur, Laguna, Masbate, Romblon, and Sibuyan, and Nueva Vizcaya), Visayas (Leyte, Panay, and Samar), and Mindanao (Davao, Lanao del Norte, and Surigao) (Wallace 1867; Mohnike 1873; Mikšić 1970 b, 1970 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14FE63FC27FD70.taxon	materials_examined	Type locality. “ Mindero (= Mindoro). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14FE63FC27FD70.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Mikšić 1968, 1971 b). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14FD4BFDD9FB1C.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14FD4BFDD9FB1C.taxon	distribution	Distribution in the Philippines. Luzon (Camotes Islands, Babuyanes, Palawan, and Polillo), Visayas (Bohol, Leyte, Panay, and Samar), and Mindanao (Basilan, Bukidnon, Camiguin, Zamboanga del Norte, Davao, Lanao del Norte, and Surigao) (Mohnike 1873; Moser 1914; Mikšić 1967, 1971 b). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14FD4BFDD9FB1C.taxon	discussion	Remarks. Wallace (1867) noted that Glycyphana (Glycyphaniola) modesta had been collected from the Philippines (coll. Parry). However, the species is currently known to occur in Indonesia and Malaysia, and the Philippine specimens were likely misidentified as G. modesta instead of G. cuculus, given the subtle morphological differences. Wallace himself remarked, “ this is probably only a variety of E. modesta, ” suggesting taxonomic ambiguity.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14FB6FFC27FA3C.taxon	materials_examined	Type locality. “ Süd Luzon, Panay. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14FB6FFC27FA3C.taxon	distribution	Distribution in the Philippines. Luzon (Masbate and Romblon), Visayas (Negros and Panay), and Mindanao (Surigao del Norte) (Mikšić 1968, 1971 b). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14F98FFC27F938.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14F98FFC27F938.taxon	distribution	Distribution in the Philippines. Luzon (Baguio, Bataan, Ilocos, Laguna, Masbate, Polillo, Mindoro, Mountain Province, Rizal, and Nueva Vizcaya) (Mikšić 1967, 1971 b). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14F893FCDCF860.taxon	materials_examined	Type locality. “ Irawan, Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14F893FCDCF860.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Antoine 2002). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ N. Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Mikšić 1968, 1971 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FEF3FC27FDC1.taxon	materials_examined	Type locality. “ Mindoro, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FEF3FC27FDC1.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Mikšić 1968, 1971 b). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FDDBFC29FCE1.taxon	materials_examined	Type locality. “ Ins. Sibuyan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FDDBFC29FCE1.taxon	distribution	Distribution in the Philippines. Luzon (Sibuyan) (Moser 1922; Mikšić 1971 b; Pavićević 1988). Overall Distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FCFBFC29FBC9.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FCFBFC29FBC9.taxon	distribution	Distribution in the Philippines. Luzon (Benguet, Nueva Vizcaya, Mt. Banahao, and Rizal) (Mikšić 1967). Overall Distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FBA3FC29FA3D.taxon	materials_examined	Type locality. “ Bohol et Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FBA3FC29FA3D.taxon	distribution	Distribution in the Philippines. Luzon (Bataan, Benguet, Nueva Vizcaya, Rizal, and Mt. Banahao), Visayas (Bohol, Negros Island, and Leyte), and Mindanao (Bukidnon, Agusan del Norte, and Lanao del Norte) (Mohnike 1873; Moser 1925; Mikšić 1971 b; Pavićević 1988). Overall Distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14F9C7FCC6F8DD.taxon	materials_examined	Type locality. “ Philippines, île de Leyte. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14F9C7FCC6F8DD.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Legrand 2004). Overall Distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14F8AFFCC6F805.taxon	materials_examined	Type locality. “ Insel Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14F8AFFCC6F805.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Mikšić 1986). Overall Distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285AFF14FF42FCC6FE74.taxon	materials_examined	Type locality. “ Philippines (Palawan Island). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285AFF14FF42FCC6FE74.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Nagai 1984; Mikšić 1986). Overall Distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285AFF14FDBFFCC6FD14.taxon	materials_examined	Type locality. “ Luzon (Imugan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285AFF14FDBFFCC6FD14.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya and Rizal) (Moser 1917 a; Heller 1934). Overall Distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285AFF14FC16FB83FB1C.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285AFF14FC16FB83FB1C.taxon	distribution	Distribution in the Philippines. Visayas (Negros Oriental) and Mindanao (Davao) (White 1856; Wallace 1867; Mohnike 1873; Mikšić 1963 b; Pavićević 1988). Overall distribution. The Philippines and China (Taiwan) (Mikšić 1963 b).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285AFF14FAA6FCDCF9F4.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285AFF14FAA6FCDCF9F4.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) and Visayas (Bohol, Negros Occidental, and Panay) (Mikšić 1963 a, 1963 b, 1987; Jákl 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285DFF14F9C6FCDCFEC5.taxon	materials_examined	Type locality. “ Philippinis. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285DFF14F9C6FCDCFEC5.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur, Laguna, Manila, Marinduque, Ilocos, Nueva Vizcaya, Rizal, Romblon, and Tarlac) and Visayas (Negros Island, Panay) (Fabricius 1775; Gory & Percheron 1833; Wallace 1868; Mohnike 1873; Kraatz 1890; Mikšić 1963 b; Jákl 2022). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14FED7FCDCFDE2.taxon	materials_examined	Type locality. “ Babuynes. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14FED7FCDCFDE2.taxon	distribution	Distribution in the Philippines. Luzon (Babuyan Archipelago) (Mohnike 1873; Mikšić 1963 a; Jákl 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14FDF4FCDCFCCB.taxon	materials_examined	Type locality. “ North Luzon Island, Sierra Madre Mountains. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14FDF4FCDCFCCB.taxon	distribution	Distribution in the Philippines. Luzon (Sierra Madre) (Jákl 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14FC15FCDCFAF5.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14FC15FCDCFAF5.taxon	distribution	Distribution in the Philippines. Luzon (Laguna, Manila, Marinduque, Rizal, Romblon, Ilocos, Camarines Sur, and Sorsogon) and Mindanao (Mt. Apo) (Chevrolat 1841; Hope 1841; Newman 1841; Mikšić 1963 a, 1963 b; Pavićević 1988). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14FAC7FCDCF94A.taxon	materials_examined	Type locality. “ Bohol. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14FAC7FCDCF94A.taxon	distribution	Distribution in the Philippines. Visayas (Bohol, Leyte, and Samar) (Chevrolat 1841; Hope 1841; Newman 1841; Mikšić 1963 a, 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14F95CFCDCF8B3.taxon	materials_examined	Type locality. “ Cebu. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14F95CFCDCF8B3.taxon	distribution	Distribution in the Philippines. Visayas (Cebu) (Mikšić 1963 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285CFF14F805FC27FF0D.taxon	materials_examined	Type locality. “ Philippines, Negros Oriental. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285CFF14F805FC27FF0D.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Arnaud 1989 a). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FE9FFC27FE75.taxon	materials_examined	Type locality. “ Negros Oriental. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FE9FFC27FE75.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Pavićević 1988). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FE44FCDCFD5A.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FE44FCDCFD5A.taxon	distribution	Distribution in the Philippines. Mindanao (Pavićević 1986). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FD2CFCDCFC83.taxon	materials_examined	Type locality. “ Philippinen, Insel Romblon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FD2CFCDCFC83.taxon	distribution	Distribution in the Philippines. Luzon (Sorsogon and Romblon) (Mikšić 1983; Pavićević 1988). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FBC2FCDCFAD1.taxon	materials_examined	Type locality. “ Philippines, Marinduque. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FBC2FCDCFAD1.taxon	distribution	Distribution in the Philippines. Luzon (Marinduque) (Arnaud 1987 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FC15FCDCFBE8.taxon	materials_examined	Type locality. “ Philippines, North Luzon, Calayan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FC15FCDCFBE8.taxon	distribution	Distribution in the Philippines. Luzon (Babuyan Island) (Arnaud 2004). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FAABFCDCF992.taxon	materials_examined	Type locality. “ Dinagat, Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FAABFCDCF992.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) and Mindanao (Agusan del Sur, Dinagat, Lanao del Sur, and Surigao) (Janson 1881; Mikšić 1963 a, 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14F9E4FCDCF8FB.taxon	materials_examined	Type locality. “ Philippines, Mindoro, Mt. Halcon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14F9E4FCDCF8FB.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Arnaud 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14F8CDFCDCF823.taxon	materials_examined	Type locality. “ Philippines, North Samar. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14F8CDFCDCF823.taxon	distribution	Distribution in the Philippines. Visayas (Samar) (Arnaud 1989 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FF0AFCC6FEBC.taxon	materials_examined	Type locality. “ Philippine, Nord Luzon, Nueva Vizcaya. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FF0AFCC6FEBC.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya, Rizal, Manila, and Mountain Province) (Arnaud 1987 a). Overall Distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FE46FCC6FD5C.taxon	materials_examined	Type locality. “ Philippines, Leyte, Mont Balocaue. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FE46FCC6FD5C.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Legrand & Chew Kea Foo 2004). Overall Distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FD66FCDCFC10.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FD66FCDCFC10.taxon	distribution	Distribution in the Philippines. Luzon (Ilocos) (Moser 1909; Mikšić 1963 a, 1987). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FBA2FCDCFAE8.taxon	materials_examined	Type locality. “ Mindanao (Davao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FBA2FCDCFAE8.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) and Mindanao (Agusan del Norte and Davao) (Moser 1918; Mikšić 1963 b; 1987). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FA3AFCDCF9AC.taxon	materials_examined	Type locality. ‘‘ Philippines, Panay Isld., Antique. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FA3AFCDCF9AC.taxon	distribution	Distribution in the Philippines. Visayas (Negros and Panay) (Arnaud 1992; Jákl 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14F93EFCDCF84C.taxon	materials_examined	Type locality. “ Luzon (Bayombong). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14F93EFCDCF84C.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1917 a, 1918; Jákl 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14FF0BFCDCFD5D.taxon	materials_examined	Type locality. “ Philippines Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14FF0BFCDCFD5D.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur, Ifugao, Mountain Province, Laguna, Ilocos, and Nueva Vizcaya) (Wallace 1867; Moser 1906, 1914, 1917 a; Bourgoin 1929; Mikšić 1963 a, 1963 b, 1972 b; Jákl 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14FD2FFCDCFC11.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur and Sorsogon) (Moser 1917 a; Mikšič 1963 a, 1963 b; Jákl 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14FC6BFCDCFB79.taxon	materials_examined	Type locality. “ Panay Island, Antique Province, Mount Madjaas. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14FC6BFCDCFB79.taxon	distribution	Distribution in the Philippines. Visayas (Panay) (Jákl 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14FA8BFCDCFA3D.taxon	materials_examined	Type locality. “ Mindoro, Mt. Halcon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14FA8BFCDCFA3D.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Legrand 2018 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14F98FFCDCF965.taxon	materials_examined	Type locality. “ Philippines, Panay Island, Antique Province, Mount Madjaas. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14F98FFCDCF965.taxon	distribution	Distribution in the Philippines. Visayas (Panay) (Jákl 2022). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692592851FF14F8AFFCDCFF20.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692592851FF14F8AFFCDCFF20.taxon	distribution	Distribution in the Philippines. Luzon (Wallace 1867). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FEBAFCDCFDC0.taxon	materials_examined	Type locality. “ Barbatan Is. - Antique. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FEBAFCDCFDC0.taxon	distribution	Distribution in the Philippines. Visayas (Antique) (Mikšić 1962). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FDDAFCDCFCCC.taxon	materials_examined	Type locality. “ Philippines, Palawan, Puerto Princesa. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FDDAFCDCFCCC.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Jákl 2011, 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FC16FCDCFB78.taxon	materials_examined	Type locality. “ Mindanao (Davao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FC16FCDCFB78.taxon	distribution	Distribution in the Philippines. Visayas (Bohol and Leyte) and Mindanao (Bukidnon, Lanao del Norte, and Davao) (Moser 1917 a; Mikšić 1963 a, 1963 b; Jákl 2017 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FB52FABEFA18.taxon	materials_examined	Type locality. “ Palavan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FB52FABEFA18.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Mikšić 1963 a; Jákl 2017 b). Overall distribution. The Philippines and Indonesia.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FB52FABEFA18.taxon	discussion	Remarks. Jákl (2017 b) noted that the record of this species in Palawan is probably erroneous.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14F9AAFCDCF938.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14F9AAFCDCF938.taxon	distribution	Distribution in the Philippines. Luzon (Cagayan Province) (Mikšić 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14F892FCDCF804.taxon	materials_examined	Type locality. “ Luzon (Imugan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14F892FCDCF804.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1917 a; Mikšić 1963 a, 1963 b, 1970 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FF0AFCDCFE98.taxon	materials_examined	Type locality. “ Philippines, North Luzon, Babuyan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FF0AFCDCFE98.taxon	distribution	Distribution in the Philippines. Luzon (Babuyan Island) (Arnaud 2004). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FEF2FC27FDE4.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FEF2FC27FDE4.taxon	distribution	Distribution in the Philippines. Luzon (Albay) (Moser, 1918; Mikšić 1963 a, 1963 b). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FDF6FC27FC84.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FDF6FC27FC84.taxon	distribution	Distribution in the Philippines. Luzon (Cagayan Province, Catanduanes, Rizal, Manila, and Laguna) and Mindanao (Mohnike 1873; Mikšić 1963 a, 1963 b). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FC4EFCDCFBA4.taxon	materials_examined	Type locality. “ Philippines, Mt. Apo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FC4EFCDCFBA4.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Arnaud 1989 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FB36FCDCF9D0.taxon	materials_examined	Type locality. “ Mindanao (Dapitan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FB36FCDCF9D0.taxon	distribution	Distribution in the Philippines. Luzon (Benguet), Visayas (Negros Oriental), and Mindanao (Agusan del Norte, Bukidnon, Basilan, Davao, and Zamboanga del Norte) (Heller 1895; Kraatz 1895; Mikšić 1963 a, 1963 b, 1987; Pavićević 1988). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14F9E2FCDCF860.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14F9E2FCDCF860.taxon	distribution	Distribution in the Philippines. Luzon (Laguna, Manila, and Nueva Vizcaya) and Visayas (Leyte) (Chevrolat 1841; Mikšić 1963 a, 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ Philippines, Romblon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) (Reichenbach 1995). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FEF3FCDCFC59.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FEF3FCDCFC59.taxon	distribution	Distribution in the Philippines. Luzon (Albay, Camarines Sur, Laguna, Manila, Nueva Vizcaya, Romblon, and Polillo), Visayas (Panay), and Mindanao (Bukidnon and Mt. Apo) (Chevrolat 1841; Newman 1841; Mohnike 1873; Wallace 1868; Haller 1884; Mikšić 1963 a, 1963 b, 1984; Reichenbach 1995). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FC33FCDCFBA5.taxon	materials_examined	Type locality. “ Sibuyan Island, Romblon, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FC33FCDCFBA5.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) (Arnaud 1987 c, 1989 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FB37FCDCFA8D.taxon	materials_examined	Type locality. “ Philippines, Sud Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FB37FCDCFA8D.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) and Visayas (Panay) (Arnaud 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FA1FFCDCF9F5.taxon	materials_examined	Type locality. “ Batanes Island, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FA1FFCDCF9F5.taxon	distribution	Distribution in the Philippines. Luzon (Batanes) (Arnaud 1989 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14F9C7FCDCF805.taxon	materials_examined	Type locality. “ Manila. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14F9C7FCDCF805.taxon	distribution	Distribution in the Philippines. Luzon (Manila), Visayas (Bohol and Leyte), and Mindanao (Camiguin) (Wallace 1867; Dohrn 1872; Kraatz 1880; Mohnike 1873; Schaum 1844; Mikšić 1963 a, 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FF0AFCDCFE99.taxon	materials_examined	Type locality. “ Philippines, Polillo Isld. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FF0AFCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Polillo Island) (Arnaud 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FEF3FCDCFDCE.taxon	materials_examined	Type locality. “ Virac (Insel Catanduanes). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FEF3FCDCFDCE.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur) (Mikšić 1963 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FDD8FCDCFC83.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FDD8FCDCFC83.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao del Norte) (Mohnike 1873; Mikšić 1963 a, 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FC15FCDCFA89.taxon	materials_examined	Type locality. “ Camiguin prope Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FC15FCDCFA89.taxon	distribution	Distribution in the Philippines. Luzon (Romblon), Visayas (Negros Oriental and Samar), and Mindanao (Bukidnon, Misamis Oriental, Camiguin, Davao, Surigao del Norte) (Mohnike 1873; Moser 1914, 1917; Mikšić 1963 a, 1963 b, 1983; Pavićević 1988). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FAE3FCDCF9FE.taxon	materials_examined	Type locality. “ Philippines, Oriental Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FAE3FCDCF9FE.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) and Visayas (Negros Oriental) (Arnaud 2004). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14F9C8FCDCF89F.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14F9C8FCDCF89F.taxon	distribution	Distribution in the Philippines. Luzon (Aurora, Benguet, Babuyan Island, Laguna, Mountain Province, Rizal, and Sorsogon) (Mohnike 1873; Mikšić 1963 a, 1963 b; Pavićević 1988). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552855FF14F8E9FCC6FF68.taxon	materials_examined	Type locality. “ Negros Oriental. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552855FF14F8E9FCC6FF68.taxon	distribution	Distribution in the Philippines. Visayas (Negros Oriental) (Pavićević 1988). Overall Distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FF43FCDCFE51.taxon	materials_examined	Type locality. “ Philippines, Mindanao, Basilan Isld. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FF43FCDCFE51.taxon	distribution	Distribution in the Philippines. Mindanao (Basilan) (Arnaud 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FE28FCDCFD86.taxon	materials_examined	Type locality. “ Mindoro, Mt. Halcon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FE28FCDCFD86.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Arnaud 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FD10FCDCFCEF.taxon	materials_examined	Type locality. “ Batanes Prov., Camiguin Isld. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FD10FCDCFCEF.taxon	distribution	Distribution in the Philippines. Luzon (Babuyan Island) (Arnaud 2000). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FCF9FCDCFA65.taxon	materials_examined	Type locality. “ Des Indes orientalis. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FCF9FCDCFA65.taxon	distribution	Distribution in the Philippines. Luzon (Abra, Albay, Cagayan Province, Camarines Sur, Ilocos, Laguna,? Palawan, Nueva Vizcaya, and Romblon), Visayas (Bohol, Leyte, Cebu, Panay, and Negros Occidental), and Mindanao (Gory & Percheron 1833; Newman 1841; Burmeister 1842; Wallace 1868; Dohrn 1872; Mohnike 1873; Heller 1899 a; Schürhoff 1938; Mikšić 1963 a, 1963 b, 1974 c, 1984, 1986). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FA74FCDCF94A.taxon	materials_examined	Type locality. “ Philippines, Batanes Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FA74FCDCF94A.taxon	distribution	Distribution in the Philippines. Luzon (Batanes) (Arnaud 1989 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522854FF14F95CFCD3FF0C.taxon	materials_examined	Type locality. “ Das Vaterland ist unbekannt. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522854FF14F95CFCD3FF0C.taxon	distribution	Distribution in the Philippines. Luzon (Laguna, Manila, Palawan, Rizal, Isabela, and Romblon), Visayas (Antique, Negros Occidental, Panay, and Samar), and Mindanao (Bukidnon, Zamboanga, Dinagat, and Lanao del Norte) (Wallace 1868; Mikšić 1962, 1963 a, 1963 b, 1983). Overall distribution. This species is widely distributed in the Oriental, Australia, Pacific, and Nearctic regions (Woodruff, 2006; Perissinotto & Clennell, 2021).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FE9EFCDCFD5C.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FE9EFCDCFD5C.taxon	distribution	Distribution in the Philippines. Luzon (Abra, Benguet, Rizal, Manila, Ilocos, Camarines Sur, and Nueva Vizcaya) and Mindanao (Mt. Apo) (Eydoux 1839; Burmeister 1842; 1847; Mohnike 1873; Moser 1909; Mikšić 1963 a, 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FD2EFCDCFC58.taxon	materials_examined	Type locality. “ Luzon-Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FD2EFCDCFC58.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Mikšić 1963 a, 1970 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FC32FCDCFBA4.taxon	materials_examined	Type locality. “ Luzon, Benguet, Berg Santo Tomas. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FC32FCDCFBA4.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) (Schultze 1916 b; Mikšić 1963 a, 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FB36FCDCFA44.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FB36FCDCFA44.taxon	distribution	Distribution in the Philippines. Mindanao (Mohnike 1873). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FA56FCDCF9AC.taxon	materials_examined	Type locality. “ Philippines, Negros Oriental. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FA56FCDCF9AC.taxon	distribution	Distribution in the Philippines. Visayas (Negros Oriental) (Arnaud 1987 a, 1989 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14F93EFCDCF894.taxon	materials_examined	Type locality. “ Philippines, North Province. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14F93EFCDCF894.taxon	distribution	Distribution in the Philippines. Luzon (Ifugao) (Arnaud 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532857FF14F8E6FCDCFF0C.taxon	materials_examined	Type locality. “ Philippines, Cuyo Isld. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532857FF14F8E6FCDCFF0C.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Arnaud 2000). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FE9EFCDCFE2C.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FE9EFCDCFE2C.taxon	distribution	Distribution in the Philippines. Luzon (Mohnike, 1873). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FDBEFCDCFC84.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FDBEFCDCFC84.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon and Surigao) (Mohnike 1873; Mikšić 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FC16FCDCFBEC.taxon	materials_examined	Type locality. “ Philippines, Mindanao, South Cotabato. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FC16FCDCFBEC.taxon	distribution	Distribution in the Philippines. Mindanao (South Cotabato) (Arnaud 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FBFEFCDCFAE8.taxon	materials_examined	Type locality. “ Insel Mindoro. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FBFEFCDCFAE8.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Kraatz 1894 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FAC2FC27F988.taxon	materials_examined	Type locality. “ Negros Occidental, Bago. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FAC2FC27F988.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) and Visayas (Negros Island and Panay) (Moser 1911; Mikšić 1963 a, 1983; Pavićević 1988). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14F9E2FC27F894.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14F9E2FC27F894.taxon	distribution	Distribution in the Philippines. Visayas (? Negros) (Moser 1909; Mikšić 1963 b). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502856FF14F8E6FCDCFF21.taxon	materials_examined	Type locality. “ Philippinen (?). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502856FF14F8E6FCDCFF21.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Haller 1884). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14FEBBFCDCFE16.taxon	materials_examined	Type locality. “ Philippines, Panay, Antique. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14FEBBFCDCFE16.taxon	distribution	Distribution in the Philippines. Visayas (Negros and Panay) (Arnaud 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14FE60FCDCFD37.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14FE60FCDCFD37.taxon	distribution	Distribution in the Philippines. Luzon (Bataan) (Mohnike 1873). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14FC81FCDCFC30.taxon	materials_examined	Type locality. “ Philippines Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14FC81FCDCFC30.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Newman 1841). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14FB8AFC8BFA39.taxon	materials_examined	Type locality. “ Manila. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14FB8AFC8BFA39.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur, Manila, and Mt. Banahao), Visayas (Samar and Leyte), and Mindanao (Agusan, Basilan, Bukidnon, Camiguin, Davao, Dinagat, Lanao del Norte, Surigao, and Zamboanga del Norte) (Dohrn 1872; Mohnike 1873; Moser 1914; Mikšić 1963 a, 1963 b, 1983). Overall distribution. The Philippines and? Vietnam.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14F990FCDCF96E.taxon	materials_examined	Type locality. “ Philippines, Leyte. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14F990FCDCF96E.taxon	distribution	Distribution in the Philippines. Visayas (Leyte and Samar) (Arnaud 2004). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512869FF14F978FCDCFE98.taxon	materials_examined	Type locality. “ Philippines Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512869FF14F978FCDCFE98.taxon	distribution	Distribution in the Philippines. Luzon (Zambales, Bataan, Baguio, Isabela, and Rizal) (Chevrolat 1841; Newman 1841; Burmeister 1842; Mohnike 1873; Mikšić 1963 a, 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FEF2FCDCFDC0.taxon	materials_examined	Type locality. “ Philippines, Batanes Prov., Camiguin Isld. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FEF2FCDCFDC0.taxon	distribution	Distribution in the Philippines. Luzon (Camiguin de Luzon) (Arnaud 2000). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FDDAFCDCFCE0.taxon	materials_examined	Type locality. “ Camiguin de Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FDDAFCDCFCE0.taxon	distribution	Distribution in the Philippines. Mindanao (Camiguin) (Mohnike 1873). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FCFAFCDCFBEC.taxon	materials_examined	Type locality. “ Philippines (?). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FCFAFCDCFBEC.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Haller 1884). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FB36FCDCFA8C.taxon	materials_examined	Type locality. “ Philippines, South Palawan, near Brooks Point. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FB36FCDCFA8C.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Jákl 2022). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FA1EFCC6F9F4.taxon	materials_examined	Type locality. “ Philippines, Oriental Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FA1EFCC6F9F4.taxon	distribution	Distribution in the Philippines. Visayas (Negros Oriental) (Arnaud 2004). Overall Distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14F9C6FCC6F8DC.taxon	materials_examined	Type locality. “ Philippines, Negros Occidental, Mont Canla-on. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14F9C6FCC6F8DC.taxon	distribution	Distribution in the Philippines. Visayas (Negros Island) (Legrand & Chew Kea Foo 2004). Overall Distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2868FF14F8AEFCC6FF20.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2868FF14F8AEFCC6FF20.taxon	distribution	Distribution in the Philippines. Luzon (Ilocos, Rizal, and Mt. Banahao) (Mohnike 1873; Mikšić 1963 b; Jákl 2022). Overall Distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FEF3FE04FCCC.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FEF3FE04FCCC.taxon	distribution	Distribution in the Philippines. Luzon (Mohnike, 1873). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FEF3FE04FCCC.taxon	discussion	Remarks. Jákl (2024) argued that Krajčík (1998) may have “ incorrectly ” placed Protaetiomorpha flavovariegata (Mohnike, 1873) in this genus because it had been historically placed in the genus Prota etia Burmeister, 1842. It was never listed in Protaetiomorpha Mikšíc, 1968, by any other author and is currently regarded as incertae sedis due to the absence of Mohnike’s type. Therefore, its generic status remains dubious.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FC4EFCDCFB1C.taxon	materials_examined	Type locality. “ Mindanao Buk. Lindabon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FC4EFCDCFB1C.taxon	distribution	Distribution in the Philippines. Visayas (Negros Oriental) and Mindanao (Bukidnon) (Mikšić 1979; Pavićević 1988). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FB6EFCDCFA18.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FB6EFCDCFA18.taxon	distribution	Distribution in the Philippines. Luzon (Mohnike 1873). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FA72FCDCF964.taxon	materials_examined	Type locality. “ Negros, Occidental Negros, Cuernos Mountains. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FA72FCDCF964.taxon	distribution	Distribution in the Philippines. Visayas (Negros Occidental) (Moser 1925). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14F976FCDCF860.taxon	materials_examined	Type locality. “ Isabela Archipelagi Sulu. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14F976FCDCF860.taxon	distribution	Distribution in the Philippines. Mindanao (Sulu Archipelago) (Mohnike 1873). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FF0BFCDCFEBD.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FF0BFCDCFEBD.taxon	distribution	Distribution in the Philippines. Luzon (Mikšić 1979). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FE0FFCDCFDE5.taxon	materials_examined	Type locality. “ Philippines, Samar. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FE0FFCDCFDE5.taxon	distribution	Distribution in the Philippines. Visayas (Leyte and Samar) (Arnaud 1996). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FDF7FCDCFCE1.taxon	materials_examined	Type locality. “ Luzon, Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FDF7FCDCFCE1.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Schürhoff 1933 c; Mikšić 1979). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FCFBFCDCFBA5.taxon	materials_examined	Type locality. “ Philippine Islands	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FCFBFCDCFBA5.taxon	distribution	Distribution in the Philippines. Luzon (Laguna and Quezon Province) and Mindanao (Wallace 1867; Mohnike 1873). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FB37FCDCFA8D.taxon	materials_examined	Type locality. “ Ile de Romblon, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FB37FCDCFA8D.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) (Arnaud 1982 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FA1FFCDCF989.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FA1FFCDCF989.taxon	distribution	Distribution in the Philippines. Mindanao (Mohnike 1873). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14F9E3FCC7F895.taxon	materials_examined	Type locality. “ Insel Romblon (Philippinen). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14F9E3FCC7F895.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) (Mikšić 1982 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286AFF14F8E7FCDCFF20.taxon	materials_examined	Type locality. “ Baguio (Luzon). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286AFF14F8E7FCDCFF20.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Schürhoff 1933 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FEF2FCC6FDC0.taxon	materials_examined	Type locality. “ Philippines, Leyte, Mont Balocaue. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FEF2FCC6FDC0.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Legrand & Jákl 2017). Overall Distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FDDAFCDCFCCC.taxon	materials_examined	Type locality. “ Philippines, Oriental Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FDDAFCDCFCCC.taxon	distribution	Distribution in the Philippines. Visayas (Arnaud 1990). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FCDEFE71FB80.taxon	materials_examined	Type locality. “ Philippines, Mindanao, Bukidnon, Cabangsalan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FCDEFE71FB80.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon, Davao Oriental, and South Cotabato) (Legrand & Jákl 2017). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FCDEFE71FB80.taxon	discussion	Remarks. According to Legrand & Jákl (2017), material collected from Palawan may be mislabeled “ provenance douteuse. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FB1AFCDCFA8C.taxon	materials_examined	Type locality. “ Philippines, Or. Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FB1AFCDCFA8C.taxon	distribution	Distribution in the Philippines. Visayas (Negros Island) (Arnaud 1992, 2004). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FA1EFCC6F988.taxon	materials_examined	Type locality. “ Philippines, Mindanao, South Cotabato, Kidapawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FA1EFCC6F988.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon and South Cotabato) (Legrand & Flutsch 2016). Overall Distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14F95AFCDCF84C.taxon	materials_examined	Type locality. “ Philippines, Mindanao, Surigao del Sur. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14F95AFCDCF84C.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao del Sur) (Arnaud 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FF0BFCDCFEBD.taxon	materials_examined	Type locality. “ Luzon, Benguet, Berg Santo Tomas. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FF0BFCDCFEBD.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) (Schultze 1916 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FE0FFCDCFD5D.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FE0FFCDCFD5D.taxon	distribution	Distribution in the Philippines. Luzon (Quezon, Nueva Vizcaya, and Catanduanes) (Mohnike 1873; Mikšić 1981). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FD2FFCDCFC59.taxon	materials_examined	Type locality. “ Philippines, Leyte. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FD2FFCDCFC59.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Arnaud 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FC33FCDCFBA5.taxon	materials_examined	Type locality. “ Philippines, Sibuyan Island, Romblon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FC33FCDCFBA5.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) and Mindanao (? Surigao) (Arnaud 1987 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FB37FCDCFAA1.taxon	materials_examined	Type locality. “ Philippines, Mindanao, Mt. Apo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FB37FCDCFAA1.taxon	distribution	Distribution in the Philippines. Mindanao (Arnaud 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FA3BFCDCF941.taxon	materials_examined	Type locality. “ Imugan, Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FA3BFCDCF941.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya and Camarines Sur) (Schürhoff 1933 a; Mikšić 1981). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14F95BFCDCF84D.taxon	materials_examined	Type locality. “ Imugan, Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14F95BFCDCF84D.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya and Camarines Sur) (Schürhoff 1935). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FF0AFC27FE2A.taxon	materials_examined	Type locality. “ Ins. Philipp. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FF0AFC27FE2A.taxon	distribution	Distribution in the Philippines. Luzon (Laguna, Rizal, Albay, and Polillo) and Visayas (Samar) (Blanchard 1842, 1850; Mohnike 1873; Mikšić 1981) Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FDBCFC27FCCB.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FDBCFC27FCCB.taxon	distribution	Distribution in the Philippines. Luzon (Catanduanes) (Mikšić 1981) Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FCDDFC27FBD4.taxon	materials_examined	Type locality. “ Luzon: Bulusan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FCDDFC27FBD4.taxon	distribution	Distribution in the Philippines. Luzon (Sorsogon) (Mikšić 1981). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FBA6FC27FA89.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FBA6FC27FA89.taxon	distribution	Distribution in the Philippines. Visayas (Samar) (Burmeister 1842; Mohnike 1873). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FAE3FC27F9B6.taxon	materials_examined	Type locality. “ Masbate (Philippinen). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FAE3FC27F9B6.taxon	distribution	Distribution in the Philippines. Luzon (Masbate) (Schürhoff 1933 a). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14F900FC27F8B3.taxon	materials_examined	Type locality. “ Panay. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14F900FC27F8B3.taxon	distribution	Distribution in the Philippines. Visayas (Panay) (Mikšić 1981). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286FFF14F805FCDCFF20.taxon	materials_examined	Type locality. “ Mindanao P. I., Bukidnon, Landabon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286FFF14F805FCDCFF20.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon) (Mikšić 1982 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FEBAFCDCFE2C.taxon	materials_examined	Type locality. “ Luzon (Imugan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FEBAFCDCFE2C.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1922 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FDBEFCDCFCCC.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FDBEFCDCFCCC.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon, Davao, Lanao del Norte, and Zamboanga) (Mohnike 1873; Mikšić 1981). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FCDEFCDCFBC8.taxon	materials_examined	Type locality. “ Philippines, Mindanao, Cotabato, Mt. Apo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FCDEFCDCFBC8.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon, Mt. Apo, and Surigao) (Arnaud 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FBA2FCDCFAD4.taxon	materials_examined	Type locality. “ Luzon: Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FBA2FCDCFAD4.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Bourgoin 1931). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FAA6FCDCF9F4.taxon	materials_examined	Type locality. “ Mindanao, Camp Keithley, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FAA6FCDCF9F4.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Sur, Surigao, and Bukidnon) (Moser 1922 a; Mikšić 1981). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14F9C6FCDCF8F0.taxon	materials_examined	Type locality. “ Mt. Apo, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14F9C6FCDCF8F0.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Mikšić 1981). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286EFF14F8CAFCDCFF0D.taxon	materials_examined	Type locality. “ Luzon (Imugan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286EFF14F8CAFCDCFF0D.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1917 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FB3BFCDCFA89.taxon	materials_examined	Type locality. “ Mindanao, P. I., Galag River / Mt. Apo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FB3BFCDCFA89.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Krikken 1980). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FAE0FCDCF992.taxon	materials_examined	Type locality. “ Luzon Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FAE0FCDCF992.taxon	distribution	Distribution in the Philippines. Luzon (Aurora, Nueva Ecija, and Quirino) (Kobayashi & Sasaki 2016; Sasaki 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14F9E5FCDCF8FB.taxon	materials_examined	Type locality. “ N. Luzon, the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14F9E5FCDCF8FB.taxon	distribution	Distribution in the Philippines. Luzon (Aurora and Quirino) (Kobayashi & Sasaki 2016; Sasaki 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14F8CDFCDCF823.taxon	materials_examined	Type locality. “ Camiguin Is., the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14F8CDFCDCF823.taxon	distribution	Distribution in the Philippines. Luzon (Camiguin) (Kobayashi & Sasaki 2016; Sasaki 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FD2DFCDCFC83.taxon	materials_examined	Type locality. “ Palawan, Mt. Mantaling Jan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FD2DFCDCFC83.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Jákl 2017 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FC1AFCDCFBE8.taxon	materials_examined	Type locality. “ S. Palawan, Brooke’s Point. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FC1AFCDCFBE8.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Jákl 2017 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FE9FFCDCFE2A.taxon	materials_examined	Type locality. “ Luzon: Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FE9FFCDCFE2A.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya and Mt. Banahao) (Bourgoin 1931; Mikšić 1981). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FF0BFD59FE51.taxon	materials_examined	Type locality. “ Mt. Malindong, Panay Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FF0BFD59FE51.taxon	distribution	Distribution in the Philippines. Luzon (Mountain Province) and Visayas (Panay) (Sakai 1997; Kobayashi & Sasaki 2016; Sasaki 2019). Overall distribution. Only in the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FE28FCDCFD86.taxon	materials_examined	Type locality. “ Mt. Barlig, Mountain Province, North Luzon, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FE28FCDCFD86.taxon	distribution	Distribution in the Philippines. Luzon (Mountain Province) (Sasaki 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FD11FCDCFCA7.taxon	materials_examined	Type locality. “ Laguna, Paete. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FD11FCDCFCA7.taxon	distribution	Distribution in the Philippines. Luzon (Laguna, Mindoro, and Rizal) and Visayas (Samar) (Schultze 1916 c; Krikken 1980; Sasaki 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FC36FCDCFBA0.taxon	materials_examined	Type locality. “ Mindanao Is. and Leyte Is., the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FC36FCDCFBA0.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) and Mindanao (Agusan del Sur and Bukidnon) (Kobayashi & Sasaki 2016; Sasaki 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FB3BFCDCFA89.taxon	materials_examined	Type locality. “ N. Luzon, the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FB3BFCDCFA89.taxon	distribution	Distribution in the Philippines. Luzon (Apayao) (Kobayashi & Sasaki 2016; Sasaki 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FAE0FCDCF992.taxon	materials_examined	Type locality. “ Mt. Malindong, Panay Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FAE0FCDCF992.taxon	distribution	Distribution in the Philippines. Visayas (Panay) (Sakai 1997; Kobayashi & Sasaki 2016; Sasaki 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14F9E5FCDCF8B3.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14F9E5FCDCF8B3.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) and Mindanao (Surigao) (Mohnike 1873; Krikken 1980; Kobayashi & Sasaki 2016). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662860FF14F80AFCDCFF0D.taxon	materials_examined	Type locality. “ Philippines, Palawan, Mt. Salakot. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662860FF14F80AFCDCFF0D.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Jákl 2011; Sasaki 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FAC7FCDCF9DA.taxon	materials_examined	Type locality. “ Philippines, N. Luzon, Mt. Province. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FAC7FCDCF9DA.taxon	distribution	Distribution in the Philippines. Luzon (Mountain Province) (Jákl 2011). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FE9FFCDCFE75.taxon	materials_examined	Type locality. “ Catanduanes Is., the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FE9FFCDCFE75.taxon	distribution	Distribution in the Philippines. Luzon (Bicol) (Kobayashi & Sasaki 2016; Sasaki 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FE44FCDCFD7E.taxon	materials_examined	Type locality. “ N. Luzon Is., the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FE44FCDCFD7E.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Ecija, Nueva Vizcaya, Mountain Province, and Ifugao) (Kobayashi & Sasaki 2016; Sasaki 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FD49FC27FCA7.taxon	materials_examined	Type locality. “ Negros Is., the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FD49FC27FCA7.taxon	distribution	Distribution in the Philippines. Visayas (Negros Island) (Kobayashi & Sasaki 2016; Sasaki 2019). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FC36FC27FB8C.taxon	materials_examined	Type locality. “ Leyte Is., the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FC36FC27FB8C.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Kobayashi & Sasaki 2016; Sasaki 2019). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672863FF14F895FC24FE99.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672863FF14F895FC24FE99.taxon	distribution	Distribution in the Philippines. Luzon (Marinduque, Romblon, Rizal, Laguna, and Tarlac), Visayas (Bohol, Negros Oriental, and Samar), and Mindanao (Zamboanga) (Hope 1841; Westwood 1842; Mohnike 1873; Schürhoff 1933 c; Mikšić 1977; Nagai 1985). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FEF0FC27FDCE.taxon	materials_examined	Type locality. “ Palawan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FEF0FC27FDCE.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Nagai 1985). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FD49FAC8FC1F.taxon	materials_examined	Type locality. “ Mindanao Prov. Marinduque. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FD49FAC8FC1F.taxon	distribution	Distribution in the Philippines. Luzon (Marinduque) (Mikšić 1986). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FD49FAC8FC1F.taxon	discussion	Remarks. The earlier distribution record of Heterorhina lumawigi seems confusing and is originally labeled “ Mindanao, Prov. Marinduque. ” However, Marinduque is not located in Mindanao but rather in Luzon.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FC6EFCDCFB3C.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FC6EFCDCFB3C.taxon	distribution	Distribution in the Philippines. Luzon (Laguna, Camarines Sur,? Ifugao,? Mountain Province, and? Sorsogon) and Mindanao (Mohnike 1873; Mikšić 1974 a, 1977). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FAC7FCDCF9FE.taxon	materials_examined	Type locality. “ N. Palawan Bacuit. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FAC7FCDCF9FE.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Schürhoff 1933 c; Mikšić 1977). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14F9C8FCDCF823.taxon	materials_examined	Type locality. “ Philippines Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14F9C8FCDCF823.taxon	distribution	Distribution in the Philippines. Luzon (Laguna and Palawan), Visayas (Bohol and Leyte), and Mindanao (Agusan del Norte, Camiguin, Lanao del Norte, Zamboanga, Cotabato, Davao, Dinagat, and Surigao) (Kirby 1818; Eschscholtz 1822; Schürhoff 1942; Mikšić 1977; Pavićević 1988). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FB37FC27FAA0.taxon	materials_examined	Type locality. “ Bacuit und Binaluan (Nord Palawan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FB37FC27FAA0.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Schürhoff 1942; Mikšić 1977). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FA3BFC7AF964.taxon	materials_examined	Type locality. “ Borneo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FA3BFC7AF964.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) and Mindanao (Zamboanga del Norte, Dinagat, Bukidnon, and Agusan) (Snellen Van Vollenhoven 1864; Mohnike 1873; Mikšić 1977) Overall distribution. The Philippines and Borneo (Mikšić 1977).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14F977FCDCF84C.taxon	materials_examined	Type locality. “ Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14F977FCDCF84C.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Legrand 2022). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FF0BFCDCFE50.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FF0BFCDCFE50.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) and Mindanao (Bukidnon, Davao, Zamboanga, and Surigao) (Mohnike 1873; Mikšić 1977). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FE2BFCDCFD70.taxon	materials_examined	Type locality. “ Insula Busuanga. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FE2BFCDCFD70.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Heller 1895). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FD4BFCDCFC7C.taxon	materials_examined	Type locality. “ Jolo Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FD4BFCDCFC7C.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) and Mindanao (Sulu) (Janson 1888; Mikšić 1977). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FEBBFCDDFDC1.taxon	materials_examined	Type locality. “ Imugan (Nueva Vizcaya). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FEBBFCDDFDC1.taxon	distribution	Distribution in the Philippines. Luzon (Baguio, Nueva Vizcaya, and Laguna) (Moser 1916 c; Mikšić 1972 b, 1977; Ochi & Sakai 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FD13FCDCFCE1.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Asin Hot Spring. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FD13FCDCFCE1.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Ochi & Sakai 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FCFBFCDCFBC9.taxon	materials_examined	Type locality. “ Philippines, Mindoro Isld, Mt. Halcon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FCFBFCDCFBC9.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Arnaud 2000; Ochi & Sakai 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FBA3FCDCFB31.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Bicol, Camarines, Mt. Isalog (= Mt. Isarog). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FBA3FCDCFB31.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur) (Ochi & Kon 2025 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FA8BFCDCF989.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FA8BFCDCF989.taxon	distribution	Distribution in the Philippines. Luzon (Abra, Apayao, Ifugao, Laguna, Mindoro, Mountain Province, and Sibuyan), Visayas (Negros Occidental), and Mindanao (? Mt. Apo) (Waterhouse 1841 a; Gerstaecker 1862; Mikšić 1972 b, 1977). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14F9E3FCDCF8F1.taxon	materials_examined	Type locality. “ Luzon Is. Baguio. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14F9E3FCDCF8F1.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Nagai, 1998). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622864FF14F8CBFCDCFF21.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622864FF14F8CBFCDCFF21.taxon	distribution	Distribution in the Philippines. Luzon (Laguna, Camarines Sur, and Baguio), Visayas (Leyte, Negros, and Samar), and Mindanao (Gerstaecker 1862; Mikšić 1972 b, 1977). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FEBBFCDCFE16.taxon	materials_examined	Type locality. “ Marinduque Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FEBBFCDCFE16.taxon	distribution	Distribution in the Philippines. Luzon (Marinduque) (Arnaud 1989 b; Ochi & Sakai 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FE60FCDCFD7E.taxon	materials_examined	Type locality. “ Iles Philippines, île Romblon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FE60FCDCFD7E.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) (Arnaud 1980). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FD49FCDCFCA7.taxon	materials_examined	Type locality. “ Philippines (Palawan Island). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FD49FCDCFCA7.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Nagai 1984). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FC36FCDCFB44.taxon	materials_examined	Type locality. “ Philippinen-Insel Zebu. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FC36FCDCFB44.taxon	distribution	Distribution in the Philippines. Luzon (Ilocos Sur, Marinduque, Quezon, Cagayan Province, Mountain Province, Laguna, Nueva Vizcaya, Manila, Rizal, Sibuyan, and Romblon), Visayas (Negros Island, Cebu, and Panay), and Mindanao (Mt. Apo) (Kraatz 1893, 1894 a; Mikšić 1972 b, 1977). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FB57FCDCFA41.taxon	materials_examined	Type locality. “ Mindanao Is., Surigao Tandag. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FB57FCDCFA41.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao del Sur and Davao de Oro) (Nagai 1998; Ochi & Sakai 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FA58FC27F94A.taxon	materials_examined	Type locality. “ Philippines, Oriental Negros, Mt. Canlaou. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FA58FC27F94A.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) and Visayas (Negros) (Arnaud 1987 c). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14F95DFC27F8B3.taxon	materials_examined	Type locality. “ Panay, Culasi. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14F95DFC27F8B3.taxon	distribution	Distribution in the Philippines. Visayas (Panay) (Mikšić 1977; Pavićević 1988). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632867FF14F80AFCDCFF20.taxon	materials_examined	Type locality. “ Philippinen-Insel Zebu. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632867FF14F80AFCDCFF20.taxon	distribution	Distribution in the Philippines. Luzon (Sibuyan), Visayas (Cebu), and Mindanao (Kraatz 1893; Mikšić 1972 b, 1977). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FEBBFC27FE08.taxon	materials_examined	Type locality. “ Mindanao Is. Mt. Apo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FEBBFC27FE08.taxon	distribution	Distribution in the Philippines. Mindanao (Davao and Bukidnon) (Nagai 1998; Ochi & Sakai 2021). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FE63FC27FD70.taxon	materials_examined	Type locality. “ Cebu Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FE63FC27FD70.taxon	distribution	Distribution in the Philippines. Visayas (Cebu) and Mindanao (Mt. Apo) (Mikšić 1972 b, 1977). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FCFBFCDCFBC8.taxon	materials_examined	Type locality. “ Philippines, Basilan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FCFBFCDCFBC8.taxon	distribution	Distribution in the Philippines. Mindanao (Basilan) (Jákl 2011). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FBA3FCDCFAE8.taxon	materials_examined	Type locality. “ Mindanao, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FBA3FCDCFAE8.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao, Lanao del Norte, Bukidnon, and Davao) (Schürhoff 1933 b; Mikšić 1977; Krikken 1979). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FAC3FCDCF9AC.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FAC3FCDCF9AC.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) and Mindanao (Lanao del Norte) (Mohnike 1873; Mikšić 1977; Krikken 1979). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14F93FFCDCF860.taxon	materials_examined	Type locality. “ Philippinen-Insel (Zebu). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14F93FFCDCF860.taxon	distribution	Distribution in the Philippines. (Cebu) and Mindanao (Bukidnon) (Kraatz 1895; Schürhoff 1933 b; Mikšić 1977; Krikken 1979). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FD13FCDCFCE1.taxon	materials_examined	Type locality. “ Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FD13FCDCFCE1.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Devecis 2004, 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FCFBFCDCFBC9.taxon	materials_examined	Type locality. “ Brooke’s Point, Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FCFBFCDCFBC9.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Allard 1995; Devecis 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FBA3FC13F9F4.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FBA3FC13F9F4.taxon	distribution	Distribution in the Philippines. Luzon (Bataan, Babuyanes, Cagayan Prov., Marinduque, Mindoro,? Palawan, Pampanga, Rizal, Romblon, and Sibuyan), Visayas (Bohol, Negros, and Leyte), and Mindanao (Dinagat, Surigao, and Mt. Apo) (Eschscholtz 1829; Gory & Percheron 1833; Burmeister 1842; Schultze 1916 a; Mikšić 1977, 1983; Devecis 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FBA3FC13F9F4.taxon	discussion	Remarks. Agestrata splendens from Java may be mislabeled or a lapsus calami. Blanchard (1842) originally noted this species as native to the Philippines, not Java.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14F9C7FC27F8F1.taxon	materials_examined	Type locality. “ Sud Mindanao, Mount Apo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14F9C7FC27F8F1.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Alexis & Delpont 2000; Devecis 2019). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FEBBFCDCFE09.taxon	materials_examined	Type locality. “ Philippines (Palawan Island). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FEBBFCDCFE09.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Nagai 1980). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612879FF14F8CBFC27FF21.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612879FF14F8CBFC27FF21.taxon	distribution	Distribution in the Philippines. Luzon (Manila, Laguna, Marinduque,? Mindoro,? Palawan, and? Sorsogon), Visayas (Cebu and? Leyte), and Mindanao (Surigao) (Mohnike 1873; Mikšić 1977; Devecis 2019) Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FEBBFC27FE16.taxon	materials_examined	Type locality. “ Babuyan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FEBBFC27FE16.taxon	distribution	Distribution in the Philippines. Luzon (Babuyan Island) (Devecis 2019). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FDD8FCDCFCEF.taxon	materials_examined	Type locality. “ Insulas Philippinenses. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FDD8FCDCFCEF.taxon	distribution	Distribution in the Philippines. Luzon (Laguna, Babuyanes, Bataan, and Rizal), Visayas (Samar and Bohol), and Mindanao (Waterhouse 1841 a, Mohnike 1873; Heller 1899 b; Mikšić 1977). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FCF9FC27FBD4.taxon	materials_examined	Type locality. “ Philippines, Leyte Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FCF9FC27FBD4.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Jákl 2008). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FBA6FC27FB3C.taxon	materials_examined	Type locality. “ Luzon, Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FBA6FC27FB3C.taxon	distribution	Distribution in the Philippines. Luzon (Laguna and Nueva Vizcaya) (Mikšić 1977). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FA8EFC27FA65.taxon	materials_examined	Type locality. “ Luzon, Laguna, Paete. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FA8EFC27FA65.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Schultze 1915; Mikšić 1977). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FA77FC27F94A.taxon	materials_examined	Type locality. “ Province de Bukidnon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FA77FC27F94A.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon) (Arnaud & Delpont 2001). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14F95CFCDCF8B3.taxon	materials_examined	Type locality. “ Philippines, île de Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14F95CFCDCF8B3.taxon	distribution	Distribution in the Philippines. Mindanao (Allard 1995). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2878FF14F805FAABFEC4.taxon	materials_examined	Type locality. “ Bengalen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2878FF14F805FAABFEC4.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Burmeister 1842). Overall distribution. This species is found in the Oriental and Palearctic regions (Allard 1995).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FED6FC27FD5C.taxon	materials_examined	Type locality. “ Insulas Philippinenses. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FED6FC27FD5C.taxon	distribution	Distribution in the Philippines. Luzon (Pangasinan, Ilocos Sur, Manila, Rizal, Camarines Sur, Quezon, and Laguna) and Mindanao (Davao, Bukidnon, and Zamboanga) (Eydoux & Souleyet 1841; Waterhouse 1841 a; Heller 1899 b; Mikšić 1977). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FD2EFC27FC84.taxon	materials_examined	Type locality. “ Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FD2EFC27FC84.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Arnaud & Delpont 2001). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FC16FC27FBA4.taxon	materials_examined	Type locality. “ Insula Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FC16FC27FBA4.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) and Mindanao (Bukidnon) (Heller 1899 b; Mikšić 1977; Arnaud & Delpont 2001). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FB36FE2BF988.taxon	materials_examined	Type locality. “ Penang et Sumatra. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FB36FE2BF988.taxon	distribution	Distribution in the Philippines. Luzon (Allard 1995; Sakai & Nagai 1998; Arnaud & Delpont 2001). Overall distribution. The? Philippines and Malaysia.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FB36FE2BF988.taxon	discussion	Remarks. Arnaud & Delpont (2001) argued that Malaysia and the Philippines historically have no physical land connection, and that populations of Macronota pugnator (Heller, 1899) from these regions differ in certain morphological aspects. This raises the question of whether the true Macronota pugnator is present in the Philippines. Furthermore, earlier taxonomic works synonymized Macronota macgregori with M acronota pugnator, and it is most likely that the population previously identified as pugnator in Luzon is macgregori.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14F9E2FC27F894.taxon	materials_examined	Type locality. “ Brooke’s Point, Palawan, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14F9E2FC27F894.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Arnaud 1985). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FEBBFCDCFE09.taxon	materials_examined	Type locality. “ Brooke’s Point, Palawan, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FEBBFCDCFE09.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Arnaud 1985). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FDDBFCDCFCCD.taxon	materials_examined	Type locality. “ Mindanao, Mt. Ape (= Mt. Apo). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FDDBFCDCFCCD.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Mikšić 1973). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FCDFFCDCFC35.taxon	materials_examined	Type locality. “ Mont Kitanglad, Bukidnon Province. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FCDFFCDCFC35.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon) (Devecis 2009). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FBFFFCDCFAD5.taxon	materials_examined	Type locality. “ Panay Island, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FBFFFCDCFAD5.taxon	distribution	Distribution in the Philippines. Visayas (Panay Island) (Devecis 2001). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FAA7FCDCF9F5.taxon	materials_examined	Type locality. “ Philippinen Insel Samar. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FAA7FCDCF9F5.taxon	distribution	Distribution in the Philippines. Visayas (Samar) and Mindanao (Bukidnon, Cotabato, Mt. Apo, and Surigao del Sur) (Mikšić 1974 b, 1976, 1984; Krikken 1978; Pavićević 1988). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14F9C7FCDCF8DD.taxon	materials_examined	Type locality. “ Philippines, North Luzon, Babuyanes, Calayan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14F9C7FCDCF8DD.taxon	distribution	Distribution in the Philippines. Luzon (Babuyanes) (Arnaud 2004). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14F8AFFCDCF805.taxon	materials_examined	Type locality. “ Mount Madias, Panay. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14F8AFFCDCF805.taxon	distribution	Distribution in the Philippines. Visayas (Panay) (Mikšić 1986; Devecis 2009). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ Philippines, Marinduque. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Marinduque) (Arnaud 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FEF0FCDCFDCE.taxon	materials_examined	Type locality. “ Negros, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FEF0FCDCFDCE.taxon	distribution	Distribution in the Philippines. Visayas (Negros Island) (Krikken 1978). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FDD9FCDCFC1F.taxon	materials_examined	Type locality. “ Insulas Philippinenses. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FDD9FCDCFC1F.taxon	distribution	Distribution in the Philippines. Luzon (Bataan, Laguna, Marinduque, Mountain Province, Palawan, and Romblon), Visayas (Cebu and Leyte), and Mindanao (Waterhouse 1841 a; Burmeister 1842; Wallace 1868; Mikšić 1976; Krikken 1978; Devecis 2009). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FC6EFCDCFB44.taxon	materials_examined	Type locality. “ Luzon, Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FC6EFCDCFB44.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao and Nueva Vizcaya) (Mikšić 1983; Devecis 2009). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FB57FCDCFAAD.taxon	materials_examined	Type locality. “ Philippines (Palawan Island). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FB57FCDCFAAD.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Nagai 1984). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FA3CFCDDF992.taxon	materials_examined	Type locality. “ Philippinen, Insel Romblon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FA3CFCDDF992.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) (Mikšić 1983). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14F9E5FCDCF8FB.taxon	materials_examined	Type locality. “ Philippines, Mindoro. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14F9E5FCDCF8FB.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Arnaud 1987 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14F8CDFCDCF823.taxon	materials_examined	Type locality. “ Mount Apo, Sud Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14F8CDFCDCF823.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Devecis 2009). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FEBBFCDCFE08.taxon	materials_examined	Type locality. “ Palawan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FEBBFCDCFE08.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Nagai 1986). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FE63FC27FD70.taxon	materials_examined	Type locality. “ Philippinen (Luzon). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FE63FC27FD70.taxon	distribution	Distribution in the Philippines. Luzon (Sorsogon, Camarines Sur, and Laguna) (Mikšić 1974 d, 1976). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FD4BFC27FBEC.taxon	materials_examined	Type locality. “ Agris Mysoriensibus. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FD4BFC27FBEC.taxon	distribution	Distribution in the Philippines. Visayas (Biliran, Leyte, and Samar) and Mindanao (Mt. Apo, Surigao, Agusan del Norte, and Lanao del Norte) (Hope 1847; Wallace 1867; Mohnike 1873; Mikšić 1974 d, 1976). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FB6FFCDCFA44.taxon	materials_examined	Type locality. “ Philippines, Basilan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FB6FFCDCFA44.taxon	distribution	Distribution in the Philippines. Mindanao (Basilan) (Jákl 2011). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FA57FCDCF940.taxon	materials_examined	Type locality. “ Philippines, Romblon Island, Sibuyan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FA57FCDCF940.taxon	distribution	Distribution in the Philippines. Luzon (Sibuyan) (Arnaud 1989 a; Jákl & Legrand 2018). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14F95BFCDCF860.taxon	materials_examined	Type locality. “ Panaon-Insel (Philippinen). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14F95BFCDCF860.taxon	distribution	Distribution in the Philippines. Visayas (Bohol, Leyte, and Samar) and Mindanao (Dinagat) (Schürhoff 1935; Mikšić 1970 d, 1976; Legrand 2017; Jákl & Legrand 2018). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14FF0BFC27FE51.taxon	materials_examined	Type locality. “ Luzon, Lamao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14FF0BFC27FE51.taxon	distribution	Distribution in the Philippines. Luzon (Bataan, Masbate, and Zambales) and Visayas (Antique, Panay, and Negros Oriental) (Mikšić 1970 d, 1976; Pavićević 1988). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14FE2BFC27FCE1.taxon	materials_examined	Type locality. “ Insulas Philippinarum. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14FE2BFC27FCE1.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur, Laguna, Masbate, Rizal, Marinduque, Mindoro, Mt. Banahao, Polillo, Nueva Vizcaya,? Sorsogon, and Zambales) and Visayas (Bohol, Panay, and Leyte) (Waterhouse 1841 a; Burmeister 1842; Mikšić 1970 d, 1976; Pavićević 1988; Jákl & Legrand 2018). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14FCFBFCDCFBA5.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14FCFBFCDCFBA5.taxon	distribution	Distribution in the Philippines. Mindanao (Agusan del Norte, Bukidnon, Davao, Surigao del Norte, and Zamboanga del Norte) (Mohnike 1873; Mikšić 1970 d, 1976; Jákl & Legrand 2018). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14FB6FFCDCF9D1.taxon	materials_examined	Type locality. “ Luzon (Mt. Isarog). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14FB6FFCDCF9D1.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur, Laguna, and Nueva Vizcaya) (Moser 1917 a; Mikšić 1972 a, 1976). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14F9ABFCDCF8DD.taxon	materials_examined	Type locality. “ Insel Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14F9ABFCDCF8DD.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Mikšić 1986). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287FFF14F8AFFC27FEC5.taxon	materials_examined	Type locality. “ Insulis Philippinensibus. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287FFF14F8AFFC27FEC5.taxon	distribution	Distribution in the Philippines. Luzon (Baguio, Camarines Sur, Masbate, Laguna, Mountain Province, and Nueva Vizcaya), Visayas (Leyte, Negros, and Samar), and Mindanao (Dinagat and Surigao del Norte) (Westwood 1854; Mikšić 1972 a, 1976; Pavićević 1988). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FED4FCDCFE2A.taxon	materials_examined	Type locality. “ Mindanao, Davao del Sur, Kapatagan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FED4FCDCFE2A.taxon	distribution	Distribution in the Philippines. Mindanao (Davao del Sur) (Legrand 2017). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FDBCFCDCFD12.taxon	materials_examined	Type locality. “ Philippines, Leyte, Mt. Balocaue. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FDBCFCDCFD12.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Legrand 2017). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FD65FCDCFBD4.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FD65FCDCFBD4.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur) and Mindanao (Bukidnon and Mt. Apo) (Mohnike 1873; Mikšić 1983; Legrand 2017). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FBA6FCDCFB3C.taxon	materials_examined	Type locality. “ The Philippines, Mindoro Island, Mt. Halcon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FBA6FCDCFB3C.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Jákl 2014). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FA8FFCDCF9DA.taxon	materials_examined	Type locality. “ Palawan, Iwahig, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FA8FFCDCF9DA.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Moser 1910; Mikšić 1972 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14F9ACFC27F902.taxon	materials_examined	Type locality. “ The Philippines, Palawan Island, Magara, Roxas. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14F9ACFC27F902.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Jákl 2020). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287EFF14F895FCDCFF68.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287EFF14F895FCDCFF68.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) and Mindanao (Lanao del Norte and Agusan del Norte) (Wallace 1867; Mikšić 1972 a, 1976). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FEBBFCDCFE16.taxon	materials_examined	Type locality. “ Palawan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FEBBFCDCFE16.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Nagai 1986). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FE60FC27FD12.taxon	materials_examined	Type locality. “ Palawan (Philippines). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FE60FC27FD12.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) and Mindanao (Nagai 1986; Mikšić 1986). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FCDDFCDCFC30.taxon	materials_examined	Type locality. “ Philippines, Oriental Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FCDDFCDCFC30.taxon	distribution	Distribution in the Philippines. Visayas (Negros Oriental) (Arnaud 1990). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FB8AFCDCFB18.taxon	materials_examined	Type locality. “ Philippines, Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FB8AFCDCFB18.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Legrand 2025). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FB73FC27FA65.taxon	materials_examined	Type locality. “ Luzon-Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FB73FC27FA65.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Mikšić 1972 c, 1976). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FA74FC27F926.taxon	materials_examined	Type locality. “ Luzon (Mt. Banahao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FA74FC27F926.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao and Camarines Sur) (Moser 1914; Mikšić 1972 c, 1976; Krikken 1977 b). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692792871FF14F8E9FCDCFE98.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692792871FF14F8E9FCDCFE98.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur, Laguna, Mt. Banahao, Masbate), Visayas (Negros), and Mindanao (Mt. Apo and Zamboanga del Norte) (Mohnike 1873; Thomson 1878; Mikšić 1976, 1984). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FEF3FCDCFDE4.taxon	materials_examined	Type locality. “ N. Palawan (Binaluan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FEF3FCDCFDE4.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Moser 1915 b; Mikšić 1976). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FDF7FCDCFCCC.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FDF7FCDCFCCC.taxon	distribution	Distribution in the Philippines. Luzon (Mohnike 1873). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FCDFFC27FBC8.taxon	materials_examined	Type locality. “ Philippine (Leite). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FCDFFC27FBC8.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Moser 1914; Mikšić 1976). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FBA3FC27FA8C.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FBA3FC27FA8C.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon and Zamboanga Sibugay) (Mohnike 1873; Moser 1914; Mikšić 1976). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FA1FFC27F9F4.taxon	materials_examined	Type locality. “ Quezon, S. Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FA1FFC27F9F4.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Sakai 1996). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14F9C7FC27F8DC.taxon	materials_examined	Type locality. “ Hinabagan, W. Samar. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14F9C7FC27F8DC.taxon	distribution	Distribution in the Philippines. Visayas (Samar) (Sakai 1996). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762870FF14F8AFFC6AFF20.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762870FF14F8AFFC6AFF20.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Mohnike 1873; Schaufuss 1885). Overall distribution. The Philippines and? Malaysia (Sakai 1996).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14FBFFFCDCFAE8.taxon	materials_examined	Type locality. “ Mindanao (Iligan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14FBFFFCDCFAE8.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Norte and Surigao) (Moser 1915 b; Ricchiardi 1996). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14FAC3FCDCF9AC.taxon	materials_examined	Type locality. “ Luzon (Ins. Philippin.). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14FAC3FCDCF9AC.taxon	distribution	Distribution in the Philippines. Luzon (Manila, Laguna, Mindoro, and Palawan) (Kraatz 1883; Ricchiardi 1996). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14F93FFCDCF84C.taxon	materials_examined	Type locality. “ Mindanao, Kabasalan, Zamboanga. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14F93FFCDCF84C.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga Sibugay, Lanao del Norte, and Surigao) (Ricchiardi 1996). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14FE63FCDCFD70.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14FE63FCDCFD70.taxon	distribution	Distribution in the Philippines. Luzon (Ilocos Norte and Quezon) (Antoine 1993). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14FD4BFCDCFC7C.taxon	materials_examined	Type locality. “ Mindanao, Momungan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14FD4BFCDCFC7C.taxon	distribution	Distribution in the Philippines. Luzon (Masbate), Visayas (Leyte) and Mindanao (Lanao del Norte, Zamboanga del Norte, Davao, and Basilan) (Antoine 1993). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FF0BFC89FEBD.taxon	materials_examined	Type locality. “ Luzon (Imugan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FF0BFC89FEBD.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1917 a; Ricchiardi 1996). Overall distribution. The Philippines and Indonesia.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FE0FFCDCFDB9.taxon	materials_examined	Type locality. “ Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FE0FFCDCFDB9.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ricchiardi 1996). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FD13FCDCFC59.taxon	materials_examined	Type locality. “ Philippinen (Panaon). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FD13FCDCFC59.taxon	distribution	Distribution in the Philippines. Visayas (Samar and Leyte) and Mindanao (Basilan) (Moser 1916 b; Ricchiardi 1996). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FC33FCDCFBA5.taxon	materials_examined	Type locality. “ Negros Occ., Mount Canla-on. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FC33FCDCFBA5.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Ricchiardi 1996). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FB37FCDCFAA1.taxon	materials_examined	Type locality. “ Samar Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FB37FCDCFAA1.taxon	distribution	Distribution in the Philippines. Visayas (Samar) (Ricchiardi 1996). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FA73FCDCF941.taxon	materials_examined	Type locality. “ Luzon, Cavite. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FA73FCDCF941.taxon	distribution	Distribution in the Philippines. Luzon (Cavite and Manila) (Miyake 1994). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742872FF14F803FAE6FE51.taxon	materials_examined	Type locality. “ Borneo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742872FF14F803FAE6FE51.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Schultze 1920; Heller 1921; Nakamura 1974; Endrődi 1976). Overall distribution. The Philippines, Malaysia, Myanmar, Indonesia, and Thailand (Endrődi 1976).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FE60FCDCFD7E.taxon	materials_examined	Type locality. “ Nord Luzon, Mountain Province. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FE60FCDCFD7E.taxon	distribution	Distribution in the Philippines. Luzon (Mountain Province) (Silvestre 2002). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FD49FC27FCA7.taxon	materials_examined	Type locality. “ Philippines, Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FD49FC27FCA7.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Silvestre 2006). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FC36FC27FB18.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FC36FC27FB18.taxon	distribution	Distribution in the Philippines. Luzon (Marinduque, Babuyan, Sibuyan, and Mindoro), Visayas (Bohol, Leyte, and Samar), and Mindanao (Dinagat) (Endrődi 1957, 1976; Silvestre 2006). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FB73FC27FA41.taxon	materials_examined	Type locality. “ Philippines, Western Samar, Eastern Visayas. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FB73FC27FA41.taxon	distribution	Distribution in the Philippines. Visayas (Samar) (Silvestre 2006). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FA58FC27F8FB.taxon	materials_examined	Type locality. “ Insulas Philippinenses. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FA58FC27F8FB.taxon	distribution	Distribution in the Philippines. Luzon and Mindanao (Sulu) (Hope 1841; Waterhouse 1841 a, 1841 b; Endrődi 1951, 1976, 1985; Rowland 2003, 2006 a, 2006 b, 2011; Silvestre 2006). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14F8CDFC27F823.taxon	materials_examined	Type locality. “ Philippines, Romblon, Sibuyan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14F8CDFC27F823.taxon	distribution	Distribution in the Philippines. Luzon (Romblon) (Silvestre 2006). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14FD49FCFAFC7B.taxon	materials_examined	Type locality. “ Sumatra: Labouan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14FD49FCFAFC7B.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Endrődi 1974). Overall distribution. The Philippines, India, Indonesia, Malaysia, Myanmar, Sri Lanka, Thailand, Vietnam, and China (including Taiwan) (Drumont et al. 2006).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14FF43FC27FDCE.taxon	materials_examined	Type locality. “ Die Insel Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14FF43FC27FDCE.taxon	distribution	Distribution in the Philippines. Luzon (Babuyanes, Catanduanes, Laguna, Marinduque, Mindoro, Palawan, Romblon, and Pampanga), Visayas (Bohol, Leyte, Negros Island, and Samar), and Mindanao (Basilan, Dinagat, and Misamis Oriental) (Erichson 1834; Arrow 1910; Nakamura 1974; Endrődi 1976). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14FB8AFCDCFB18.taxon	materials_examined	Type locality. “ Harihuron, Mindro Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14FB8AFCDCFB18.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Yamaya 2009). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14FB73FCDCFA41.taxon	materials_examined	Type locality. “ Philippines, Marabut, Western Samar, Eastern Visayas. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14FB73FCDCFA41.taxon	distribution	Distribution in the Philippines. Visayas (Samar) (Dupuis 2002). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14F9C8FE8CF89F.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14F9C8FE8CF89F.taxon	distribution	Distribution in the Philippines. Luzon (Mohnike 1874; Schultze 1916 a; Endrődi 1973; Nakamura 1974). Overall distribution. The Philippines, Indonesia, Malaysia, Sri Lanka, and China (Endrődi 1973; Nakamura 1974).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722874FF14F8E9FBA4FEC5.taxon	materials_examined	Type locality. “ Habitat in Afia. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722874FF14F8E9FBA4FEC5.taxon	distribution	Distribution in the Philippines. This species is found throughout the archipelago. Overall distribution. This species is widespread in the Indo-Pacific region and is also found in the Palearctic, Australian, Neotropical, and Madagascan regions (Hao et al. 2022).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14FC6EFCDCFB44.taxon	materials_examined	Type locality. “ Ditumabo Barangay, Baler Municipality, Aurora Prov., Luzon Is., The Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14FC6EFCDCFB44.taxon	distribution	Distribution in the Philippines. Luzon (Aurora and Quezon) (Geshev 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14F9C8FC55F8B3.taxon	materials_examined	Type locality. “ Celébes centralis. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14F9C8FC55F8B3.taxon	distribution	Distribution in the Philippines. Mindanao (Davao) (Heller 1896; Endrődi, 1959; Kuijten 1994; Geshev 2024). Overall distribution. The Philippines and Indonesia (Geshev 2024).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14FA8FFCDCF9FE.taxon	materials_examined	Type locality. “ Philippine Is. Mt. Makiling, Laguna. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14FA8FFCDCF9FE.taxon	distribution	Distribution in the Philippines. Luzon (Laguna, Marinduque, Sibuyan, Mindoro, Aurora Province, and Sorsogon), Visayas (Panay), and Mindanao (Bukidnon) (Kuijten 1994; Yamaya 2011; Geshev 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14FE0CFB22FD86.taxon	materials_examined	Type locality. “ India (?). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14FE0CFB22FD86.taxon	distribution	Distribution in the Philippines.? Philippine archipelago (Endrődi 1974). Overall distribution. The? Philippines, Indonesia, and Malaysia (Nakamura 1974).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14FD11FC27FCEF.taxon	materials_examined	Type locality. “ Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14FD11FC27FCEF.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Miyake & Yamaya 1999). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732877FF14F80AFCDCFF0D.taxon	materials_examined	Type locality. “ Philippines, Balisan (= Basilan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732877FF14F80AFCDCFF0D.taxon	distribution	Distribution in the Philippines. Mindanao (Basilan) (Voirin 1996). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14FC36FE56FB18.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14FC36FE56FB18.taxon	distribution	Distribution in the Philippines. Luzon (Manila) (Schultze 1916 a; Endrődi 1969 a; Nakamura 1974). Overall distribution. The Philippines, Japan, and China (including Taiwan) (Löbl & Smetana 2006). Genus Dipelicus Hope, 1842	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14FB73FC54FA39.taxon	materials_examined	Type locality. “ Celebes; Tondano; Macassar; Philippinen; Manila; Sumatra. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14FB73FC54FA39.taxon	distribution	Distribution in the Philippines. Luzon (Manila) and Mindanao (Heller 1897 c; Schultze 1916 a; Endrődi 1969, 1971 b). Overall distribution. The Philippines and Indonesia (Heller 1897 c).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14F990FE5DF8FB.taxon	materials_examined	Type locality. “ Sybuyan (= Sibuyan) Island, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14F990FE5DF8FB.taxon	distribution	Distribution in the Philippines. Luzon (Sibuyan), Visayas (Leyte and Samar), and Mindanao (Basilan) (Miyake & Yamaya 1997). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14FE9FFCDCFDCE.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14FE9FFCDCFDCE.taxon	distribution	Distribution in the Philippines. Luzon (Bataan, Camarines Sur, Manila, Quirino, Masbate, Mt. Banahao), Visayas (Leyte), and Mindanao (Fairmaire 1883; Endrődi 1971 a, 1985; Yamaya 2011; Geshev 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14FDD9FCDCFD37.taxon	materials_examined	Type locality. “ Sibuyan Is. and Samar Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14FDD9FCDCFD37.taxon	distribution	Distribution in the Philippines. Luzon (Sibuyan) and Visayas (Samar) (Yamaya 2011). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14F8CCFCDCF823.taxon	materials_examined	Type locality. “ Mt. Canlaon, Negros Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14F8CCFCDCF823.taxon	distribution	Distribution in the Philippines. Visayas (Negros Island) (Yamaya 2009). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FF0AFCDCFE98.taxon	materials_examined	Type locality. “ Mount Apo, Mindanao Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FF0AFCDCFE98.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Yamaya 2006). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FEF2FB3CFDE4.taxon	materials_examined	Type locality. “ Philippinen: Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FEF2FB3CFDE4.taxon	distribution	Distribution in the Philippines. Luzon (Laguna, Benguet, Kalinga) (Arrow 1937 b; Endrődi 1969 a, 1971 b). Overall distribution. The Philippines and China (Taiwan) (Löbl & Smetana 2006).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FDF6FCDCFC84.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FDF6FCDCFC84.taxon	distribution	Distribution in the Philippines. Luzon (Prell 1934; Endrődi 1969 b, 1971 b; Nakamura 1974). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FC4FFCDCFB78.taxon	materials_examined	Type locality. “ Mt. Apo, Mindanao, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FC4FFCDCFB78.taxon	distribution	Distribution in the Philippines. Mindanao (Cotabato) (Miyake & Yamaya 1997; Geshev & Pathomwattananurak 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FB53FCDCFA44.taxon	materials_examined	Type locality. “ Mt. Banahaw, Luzon Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FB53FCDCFA44.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao) (Miyake & Yamaya 2000; Geshev & Pathomwattananurak 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FA57FCDCF940.taxon	materials_examined	Type locality. “ Mt. Halcon, Mindro [Mindoro] Is., Philippines ”.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FA57FCDCF940.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Miyake & Yamaya 2000; Geshev & Pathomwattananurak 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14F95BFCDCF84C.taxon	materials_examined	Type locality. “ Mt. Canlaon, Mambcal, Negros Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14F95BFCDCF84C.taxon	distribution	Distribution in the Philippines. Visayas (Negros Island) (Miyake & Yamaya 1997; Geshev & Pathomwattananurak 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14F98FFCDCF939.taxon	materials_examined	Type locality. “ Luzon: Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14F98FFCDCF939.taxon	distribution	Distribution in the Philippines. Luzon (Ritsema 1897; Heller 1897 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14F893FCDCF861.taxon	materials_examined	Type locality. “ Luzon: Baguio. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14F893FCDCF861.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Moser 1926). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FC83FBAFFBEC.taxon	materials_examined	Type locality. “ Mt. Apo, Mindanao Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FC83FBAFFBEC.taxon	distribution	Distribution in the Philippines. Luzon, Visayas (Leyte), and Mindanao (Mt. Apo) (Muramoto 1995; Jákl & Zídek 2015; Yang et al. 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FC83FBAFFBEC.taxon	discussion	Remarks. Yang et al. (2023) noted that the Eophileurus chinensis identified by Schultze (1916 a) in the Agusan River might be a misidentified specimen of Eophileurus iwasei.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FBFFFCDCFAD5.taxon	materials_examined	Type locality. “ Mt. Halcon, Paitan Barangay, Naujuan Municipality, Oriental Mindoro Prov., Mindoro Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FBFFFCDCFAD5.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro and Palawan) (Yang et al. 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FF0BFCDCFE51.taxon	materials_examined	Type locality. “ Palawan (Süd Palawan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FF0BFCDCFE51.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Prell 1912; Endrődi 1969; Nakamura 1974; Geshev & Pathomwattananurak 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FE2BFCDCFDB9.taxon	materials_examined	Type locality. “ Samar Island, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FE2BFCDCFDB9.taxon	distribution	Distribution in the Philippines. Visayas (Samar) (Miyake & Yamaya 1997). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FF0AFCDCFE99.taxon	materials_examined	Type locality. “ Luzon, Mountain Province, Pauai (Haight’s place). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FF0AFCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Mountain Province) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FEF3FCDCFDCE.taxon	materials_examined	Type locality. “ Luzon (Mt. Banahao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FEF3FCDCFDCE.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao) (Moser 1915 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FDD8FCDCFD37.taxon	materials_examined	Type locality. “ Mt. Balocaue, Leyte Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FDD8FCDCFD37.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Kobayashi 2014). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FC81FCDCFC1F.taxon	materials_examined	Type locality. “ Luzon, Laguna Province, Los Banos. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FC81FCDCFC1F.taxon	distribution	Distribution in the Philippines. Luzon (Los Baños, Laguna) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FC6EFCDCFB44.taxon	materials_examined	Type locality. “ Bazilan Is., Philippines ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FC6EFCDCFB44.taxon	distribution	Distribution in the Philippines. Mindanao (Basilan) (Kobayashi 2014). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FB56FCDCFAAD.taxon	materials_examined	Type locality. “ Luzon, Benguet. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FB56FCDCFAAD.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FA3FFCDCF992.taxon	materials_examined	Type locality. “ Palawan septentrionalis (Binaluan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FA3FFCDCF992.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Moser 1915 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14F9E4FCDCF8FB.taxon	materials_examined	Type locality. “ Luzon, Nueva Vizcaya Province, Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14F9E4FCDCF8FB.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14F8CDFCDCF823.taxon	materials_examined	Type locality. “ La Carlota, Occidental Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14F8CDFCDCF823.taxon	distribution	Distribution in the Philippines. Visayas (Negros Occidental) (Chapin 1931 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FF0AFCDCFE98.taxon	materials_examined	Type locality. “ Mt. Balocaue, Leyte Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FF0AFCDCFE98.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Kobayashi 2013). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FEF2FCDCFDE4.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FEF2FCDCFDE4.taxon	distribution	Distribution in the Philippines. Luzon (Manila) (Blanchard 1851). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FDF6FCDCFCCC.taxon	materials_examined	Type locality. “ Mindanao (Dapitan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FDF6FCDCFCCC.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Moser 1921 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FCDEFCDCFC34.taxon	materials_examined	Type locality. “ Mt. Pulis, N. Luzon Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FCDEFCDCFC34.taxon	distribution	Distribution in the Philippines. Luzon (Mountain Province) (Kobayashi 2013). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FB86FCDCFB1C.taxon	materials_examined	Type locality. “ Mt. Balocaue, Leyte Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FB86FCDCFB1C.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Kobayashi 2014). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FB6EFCDCFA44.taxon	materials_examined	Type locality. “ Mt. Mandaragon, Negros Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FB6EFCDCFA44.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Kobayashi 2014). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FA56FCDCF9AC.taxon	materials_examined	Type locality. “ Mindanao (Dapitan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FA56FCDCF9AC.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Moser 1921 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14F93EFCDCF894.taxon	materials_examined	Type locality. “ Mindanao, Surigao Province, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14F93EFCDCF894.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280AFF14F8E6FC27FF0C.taxon	materials_examined	Type locality. “ Mt. Canla-on, Negros Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280AFF14F8E6FC27FF0C.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Kobayashi 2014). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FE9EFC27FE2C.taxon	materials_examined	Type locality. “ Luzon, Nueva Vizcaya Province, Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FE9EFC27FE2C.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya, Tagaytay, Benguet, Mountain Province, and Quezon) (Moser 1924 b; Kobayashi 2014). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FDBEFC27FD14.taxon	materials_examined	Type locality. “ Mt. Balocaue, Leyte Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FDBEFC27FD14.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Kobayashi 2014). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FD66FC27FC7C.taxon	materials_examined	Type locality. “ Marinduque Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FD66FC27FC7C.taxon	distribution	Distribution in the Philippines. Luzon (Marinduque) (Kobayashi 2014). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FC4EFC27FBA4.taxon	materials_examined	Type locality. “ Mt. Madla-as, Panay Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FC4EFC27FBA4.taxon	distribution	Distribution in the Philippines. Visayas (Panay) (Kobayashi 2014). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FB36FCDCFA8C.taxon	materials_examined	Type locality. “ Mt. Mandalagon, N. Negros Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FB36FCDCFA8C.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Kobayashi 2013). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FA1EFCDCF9F4.taxon	materials_examined	Type locality. “ Mindanao, Surigao Province, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FA1EFCDCF9F4.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14F9C7FCDCF8DC.taxon	materials_examined	Type locality. “ Philippines (Negros Is.). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14F9C7FCDCF8DC.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Kobayashi 2013). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14F8AFFCDCF804.taxon	materials_examined	Type locality. “ Mt. Balocaue, Leyte Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14F8AFFCDCF804.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Kobayashi 2013). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FF0AFCDCFE98.taxon	materials_examined	Type locality. “ Ligao City, Albay, Bicol, Luzon Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FF0AFCDCFE98.taxon	distribution	Distribution in the Philippines. Luzon (Albay) (Kobayashi 2013). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FEF2FCDCFDC0.taxon	materials_examined	Type locality. “ Luzon, Province of Benguet, Irisan River. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FEF2FCDCFDC0.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) (Moser 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FDDAFCDCFD28.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FDDAFCDCFD28.taxon	distribution	Distribution in the Philippines. Luzon (Moser 1915 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FC82FCDCFC10.taxon	materials_examined	Type locality. “ Luzon, Mountain Province, Pauai (Haight’s place). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FC82FCDCFC10.taxon	distribution	Distribution in the Philippines. Luzon (Mountain Province) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FC6AFCDCFB78.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Ritsema 1897). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FB52FCDCFAA0.taxon	materials_examined	Type locality. “ Philippines (Negros Is.). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FB52FCDCFAA0.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Kobayashi 2013). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FA3AFCDCF940.taxon	materials_examined	Type locality. “ Mindanao, Kamp Keithley. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FA3AFCDCF940.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Sur, Agusan del Sur, Misamis Oriental, and Davao del Sur) (Moser 1910; Kobayashi & Medina 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14F95AFCDCF84C.taxon	materials_examined	Type locality. “ Mindanao, Butuan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14F95AFCDCF84C.taxon	distribution	Distribution in the Philippines. This species is endemic to the Philippines and is found in Mindanao (Agusan del Norte) (Moser 1915 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FF0AFCDCFEBD.taxon	materials_examined	Type locality. “ Kabangsalan, Bukidnon, Mindanao Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FF0AFCDCFEBD.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon and Zamboanga del Sur) (Kobayashi & Medina 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FE0FFCDCFDE2.taxon	materials_examined	Type locality. “ Mt. Balocaue, Leyte Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FE0FFCDCFDE2.taxon	distribution	Distribution in the Philippines. Visayas (Leyte and Samar) (Kobayashi 2013). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FDF4FCDCFCCB.taxon	materials_examined	Type locality. “ Luzon (Mt. Makiling). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FDF4FCDCFCCB.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Moser 1915 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FCDDFCDCFC30.taxon	materials_examined	Type locality. “ Philippines (Negros Is.). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FCDDFCDCFC30.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Kobayashi 2013). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FB8AFCDCFB18.taxon	materials_examined	Type locality. “ Mindanao, Kolambugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FB8AFCDCFB18.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Norte) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FB73FCDCFA41.taxon	materials_examined	Type locality. “ Luzon, Province of Benguet, Irisan River. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FB73FCDCFA41.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) (Moser 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FA5BFCDCF9B6.taxon	materials_examined	Type locality. “ Southern Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FA5BFCDCF9B6.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ritsema 1897). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14F900FBD9F8B3.taxon	materials_examined	Type locality. “ Palawan u. N. Borneo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14F900FBD9F8B3.taxon	distribution	Distribution in the Philippines. Luzon (Cavite and Palawan) (Heller 1896; Calcetas et al. 2021). Overall distribution. The Philippines and Malaysia (Mohamedsaid et al. 1997).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280FFF14F805FCDCFF0C.taxon	materials_examined	Type locality. “ Mt. Mandalagon, N. Negros Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280FFF14F805FCDCFF0C.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Kobayashi 2013). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FE9EFCDCFE74.taxon	materials_examined	Type locality. “ Luzon (Mt. Isarog). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FE9EFCDCFE74.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur) (Moser 1917 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FE46FCDCFD5C.taxon	materials_examined	Type locality. “ Luzon (Los Banos). ’	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FE46FCDCFD5C.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Moser 1921 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FD2EFCDCFC84.taxon	materials_examined	Type locality. “ Luzon, Mountain Province, Balbalasang. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FD2EFCDCFC84.taxon	distribution	Distribution in the Philippines. Luzon (Mountain Province) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FC16FCDCFBEC.taxon	materials_examined	Type locality. “ Luzon, Mountain Province, Mount Polis. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FC16FCDCFBEC.taxon	distribution	Distribution in the Philippines. Luzon (Mountain Province) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FBFEFCDCFAE8.taxon	materials_examined	Type locality. “ Bayog, Lanao del sur, Mindanao Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FBFEFCDCFAE8.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) and Mindanao (Lanao del Sur, Davao del Sur, Surigao del Sur, Zamboanga, and Bukidnon) (Kobayashi & Medina 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FAC2FCDCF988.taxon	materials_examined	Type locality. “ Insulae Philippinae, Mindanao, Dapítan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FAC2FCDCF988.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon, Davao, Lanao del Sur, Agusan del Sur, Cotabato, Misamis Oriental, Surigao del Sur, and Zamboanga del Norte) (Heller 1897 b; Kobayashi & Medina 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14F9E2FCDCF8F0.taxon	materials_examined	Type locality. “ Luzon. Province of Benguet, Irisan River. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14F9E2FCDCF8F0.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) (Moser 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280EFF14F8CAFCDCFF21.taxon	materials_examined	Type locality. “ Dominorog, Bukidnon, Mindanao Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280EFF14F8CAFCDCFF21.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon, Lanao del Sur, and Agusan del Sur) (Kobayashi & Medina 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FEBBFCDCFE16.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FEBBFCDCFE16.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Moser 1915 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FE60FCDCFD7E.taxon	materials_examined	Type locality. “ Mindanao, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FE60FCDCFD7E.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FD49FCDCFCA7.taxon	materials_examined	Type locality. “ Mindanao, Bukidnon Province, Tangcolan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FD49FCDCFCA7.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FC31FCDCFB8C.taxon	materials_examined	Type locality. “ Luzon, Cagayan. Camalaniogan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FC31FCDCFB8C.taxon	distribution	Distribution in the Philippines. Luzon (Cagayan Province) (Moser 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FB1EFCDCFA89.taxon	materials_examined	Type locality. “ Insulae Philippinae, Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FB1EFCDCFA89.taxon	distribution	Distribution in the Philippines. Luzon (Heller 1897 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FAE3FCDCF9FE.taxon	materials_examined	Type locality. “ Luzon (Tayabas). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FAE3FCDCF9FE.taxon	distribution	Distribution in the Philippines. Luzon (Quezon Province) (Moser 1916 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14F979FC89F823.taxon	materials_examined	Type locality. “ Philippines Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14F979FC89F823.taxon	distribution	Distribution in the Philippines. Luzon (Catanduanes, Marinduque, and Mindoro), Visayas (Samar), and Mindanao (Dinagat) (Burmeister & Schaum 1841; Waterhouse 1841 a, 1841 ab). Overall distribution. The Philippines and Indonesia.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FEF0FCDCFDCE.taxon	materials_examined	Type locality. “ Mindanao (Dapitan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FEF0FCDCFDCE.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Moser 1921 a, 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FDD9FCDCFD37.taxon	materials_examined	Type locality. “ Mindanao, Surigao Province, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FDD9FCDCFD37.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FC81FCDCFC1F.taxon	materials_examined	Type locality. “ Balocaue, Leyte Island, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FC81FCDCFC1F.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Kobayashi & Fujioka 2014). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FC6EFCDCFB44.taxon	materials_examined	Type locality. “ Luzon, Mountain Province, Baguio. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FC6EFCDCFB44.taxon	distribution	Distribution in the Philippines. Luzon (Baguio and Mountain Province) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FB57FCDCFAAD.taxon	materials_examined	Type locality. “ Luzon, Province of Benguet, Irisan River, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FB57FCDCFAAD.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) (Moser 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FA3CFCDCF992.taxon	materials_examined	Type locality. “ Mt. Halcon, Mindoro Island, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FA3CFCDCF992.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Kobayashi & Fujioka 2014). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14F9E5FCDCF8FB.taxon	materials_examined	Type locality. “ Mt. Canla-on, Negros Island, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14F9E5FCDCF8FB.taxon	distribution	Distribution in the Philippines. Visayas (Negros Island) (Kobayashi & Fujioka 2014). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14F8CDFCDCF823.taxon	materials_examined	Type locality. “ Negros Occidental, Maao, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14F8CDFCDCF823.taxon	distribution	Distribution in the Philippines. Visayas (Negros Occidental) (Moser 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FC33FCDCFB30.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FC33FCDCFB30.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Ecija and Camarines Sur), Visayas, and Mindanao (Misamis Oriental, Lanao del Sur, Bukidnon, Davao del Norte, Davao del Sur, and Cotabato) (Brenske 1896; Moser 1910; Calcetas et al. 2020). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FF0BFCDCFE98.taxon	materials_examined	Type locality. “ Philippinen: Ins. Sibuyan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FF0BFCDCFE98.taxon	distribution	Distribution in the Philippines. Luzon (Sibuyan Island) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FEF3FCDCFDC0.taxon	materials_examined	Type locality. “ Philippines Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FEF3FCDCFDC0.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Sharp 1876). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FDDBFCDCFD28.taxon	materials_examined	Type locality. “ Balocaue, Leyte Island, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FDDBFCDCFD28.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Kobayashi & Fujioka 2014). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FAC3FCDCF9D0.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FAC3FCDCF9D0.taxon	distribution	Distribution in the Philippines. Luzon (Dalla Torre 1912 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14F9ABFB4CF938.taxon	materials_examined	Type locality. “ Sarawak. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14F9ABFB4CF938.taxon	distribution	Distribution in the Philippines. Luzon (Brenske 1894 a; Schultze 1916 a). Overall distribution. The Philippines and Malaysia (Sharp 1876; Calcetas & Adorada 2017).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14F893FCDCF804.taxon	materials_examined	Type locality. “ Manila. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14F893FCDCF804.taxon	distribution	Distribution in the Philippines. Luzon (Manila) (Burmeister 1855; Dalla Torre 1912 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FF0BFCDCFEBD.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FF0BFCDCFEBD.taxon	distribution	Distribution in the Philippines. Luzon (Cagayan Province and Manila) and Visayas (Negros Occidental) (Blanchard 1850; Schultze 1916 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FE44FCDCFD7E.taxon	materials_examined	Type locality. “ Mindanao, Dapitan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FE44FCDCFD7E.taxon	distribution	Distribution in the Philippines. Mindanao (Misamis Occidental and Zamboanga del Norte) (Moser 1924 b; Calcetas & Adorada 2017). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FD49FCDCFC7B.taxon	materials_examined	Type locality. “ Philippines (Mindanao, Zamboanga City). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FD49FCDCFC7B.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga City and Basilan) (Calcetas 2023; Matsumoto 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FC52FCDCFBA0.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FC52FCDCFBA0.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga) (Brenske 1894 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FB3BFCDCFA89.taxon	materials_examined	Type locality. “ Mindanao: Sulu: Basilan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FB3BFCDCFA89.taxon	distribution	Distribution in the Philippines. Mindanao (Sulu Archipelago) (Calcetas & Adorada 2017). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FAE0FCDCF9FE.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FAE0FCDCF9FE.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Frey 1963). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14F9C8FCDCF823.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14F9C8FCDCF823.taxon	distribution	Distribution in the Philippines. Luzon (Antique, Batanes, Cavite, Laguna, Marinduque, Manila, Masbate, Quezon, and Romblon), Visayas (Cebu, Leyte, Negros Island, and Samar), and Mindanao (Bukidnon, Cotabato, Davao del Sur, Davao Oriental, Misamis Oriental, and Surigao del Sur) (Chevrolat 1841; Blanchard 1850; Burmeister 1855; Calcetas & Adorada 2017). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FF0BFB02FEBC.taxon	materials_examined	Type locality. “ Samboangan (ǐle Mindanao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FF0BFB02FEBC.taxon	distribution	Distribution in the Philippines. Mindanao (Sulu and Zamboanga City) (Blanchard 1851; Brenske 1896). Overall distribution. The Philippines and Indonesia (Blanchard 1851; Brenske 1896).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FE0FFBDCFDE4.taxon	materials_examined	Type locality. “ Borneo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FE0FFBDCFDE4.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Schultze 1916 a; Calcetas 2023). Overall distribution. The Philippines and Borneo (Sharp, 1876; Calcetas 2023).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FDF7FCDCFC7C.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FDF7FCDCFC7C.taxon	distribution	Distribution in the Philippines. Luzon (Albay, Antique, Bataan, Batanes, Batangas, Camarines Norte, Cavite, Ilocos Sur, Isabela, Laguna, La Union, Manila, Marinduque, Mountain Province, Oriental Mindoro, Pangasinan, Quezon City, Rizal, Sorsogon, and Zambales), Visayas (Negros Island and Bohol), and Mindanao (Davao) (Burmeister 1855; Brenske 1894 a; Frey 1963; Calcetas & Adorada 2017). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FC4FFCDCFB78.taxon	materials_examined	Type locality. “ Philippines (Mindanao, Misamis Oriental, Cagayan de Oro City). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FC4FFCDCFB78.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon and Misamis Oriental) (Calcetas 2023; Matsumoto 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FB53FCDCFA44.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FB53FCDCFA44.taxon	distribution	Distribution in the Philippines. Luzon (Marinduque), Visayas (Negros Oriental), and Mindanao (Misamis Oriental) (Moser 1924 b; Calcetas & Adorada 2017). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FA57FCDCF964.taxon	materials_examined	Type locality. “ Philippinen, Dugang. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FA57FCDCF964.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) and Mindanao (Mt. Apo and Surigao) (Brenske 1896; Calcetas 2023; Matsumoto 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14F977FCDCF84C.taxon	materials_examined	Type locality. “ Philippines (Mindanao, South Cotabato). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14F977FCDCF84C.taxon	distribution	Distribution in the Philippines. Mindanao (South Cotabato) (Calcetas 2023). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14F8AFFCDCF805.taxon	materials_examined	Type locality. “ The Philippines, Luzon Island, Kalinga Apayao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14F8AFFCDCF805.taxon	distribution	Distribution in the Philippines. Luzon (Apayao) (Calcetas 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FF43FDD9FE2D.taxon	materials_examined	Type locality. “ N O Luzon / Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FF43FDD9FE2D.taxon	distribution	Distribution in the Philippines. Luzon (Prokofiev 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FF43FDD9FE2D.taxon	discussion	Remarks. Zídek (2021) suspected that P. leucothea belongs to a different genus due to its “ less numerous and densely squamosal elytral ribs and short pronotal sulcus, ” which make the species appear taxonomically unfit for its current classification.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FDF7FCDCFC85.taxon	materials_examined	Type locality. “ Philippines (Leyte, Kabeng, Palawan Is.). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FDF7FCDCFC85.taxon	distribution	Distribution in the Philippines. Luzon (Palawan), Visayas (Leyte), and Mindanao (Agusan del Sur, Davao, and Misamis Oriental) (Keith 2006; Matsumoto 2010 b; Prokofiev 2019; Zídek 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FC4FFCDCFBA5.taxon	materials_examined	Type locality. “ La Carlota, Occidental Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FC4FFCDCFBA5.taxon	distribution	Distribution in the Philippines. Visayas (Negros Occidental) (Chapin 1931 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FB37FB91FA45.taxon	materials_examined	Type locality. “ Manila. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FB37FB91FA45.taxon	distribution	Distribution in the Philippines. Luzon (Manila and Benguet) (Brenske 1892 c; Itoh 1993 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FB37FB91FA45.taxon	discussion	Remarks. This species was originally collected from the Philippines, not from South America or South Africa. Itoh (1993) later confirmed that it is indeed from the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FA57FCDCF9AD.taxon	materials_examined	Type locality. “ Philippinen ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FA57FCDCF9AD.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Brenske 1896). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14F977FCDCF860.taxon	materials_examined	Type locality. “ Is. Mindoro, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14F977FCDCF860.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro and Palawan) (Itoh 2003 b; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FF0BFCDCFE98.taxon	materials_examined	Type locality. “ Philippines, Mindanao Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FF0BFCDCFE98.taxon	distribution	Distribution in the Philippines. Mindanao (Calcetas 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FEF3FCDCFDC0.taxon	materials_examined	Type locality. “ The Philippines, Luzon Island, Quezon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FEF3FCDCFDC0.taxon	distribution	Distribution in the Philippines. Luzon (Quezon Province) (Calcetas 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FDDBFCDCFC84.taxon	materials_examined	Type locality. “ Palawan (Binaluan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FDDBFCDCFC84.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro, Palawan, Romblon, and Sibuyan) and Visayas (Leyte, Negros Island, and Samar) (Moser 1915 a; Calcetas 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FC17FCDCFB78.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FC17FCDCFB78.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) and Mindanao (Bukidnon and Surigao) (Brenske 1894 a; Itoh 1992, 1993 b; Calcetas 2019). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FB53FAEAF9F4.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FB53FAEAF9F4.taxon	distribution	Distribution in the Philippines. Luzon (Albay, Batangas, Catanduanes, Cavite, Iloilo, Isabela, Laguna, Masbate, Polillo, and Quezon Province), Visayas (Negros Island and Leyte), and Mindanao (Basilan, Dinagat, and Surigao) (Laporte 1840; Blanchard 1851; Matsumoto 2010 b; Calcetas 2019). Overall distribution. The Philippines and Indonesia (Laporte 1840; Blanchard 1851; Calcetas 2019).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FF0BFCDCFEBD.taxon	materials_examined	Type locality. “ Mt. Apo, Mindanao Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FF0BFCDCFEBD.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo and Mt. Melibengoy) (Matsumoto 2008 b, 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FE0FFCDCFDB9.taxon	materials_examined	Type locality. “ Luzon, Mountain Province, Baguio. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FE0FFCDCFDB9.taxon	distribution	Distribution in the Philippines. Luzon (Baguio and Mountain Province) (Moser 1924 b; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FD13FCDCFC85.taxon	materials_examined	Type locality. “ Luzon, Mount Banahao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FD13FCDCFC85.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao) (Moser 1924 b; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FC17FCDCFB79.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FC17FCDCFB79.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) and Mindanao (Brenske 1894 b; Saylor 1937; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FB53FCDCFA45.taxon	materials_examined	Type locality. “ Mt. Halcon, Mindoro, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FB53FCDCFA45.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Matsumoto 2010, 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FA57FCDCF939.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FA57FCDCF939.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao, Cavite, Manila, and Nueva Ecija) and Mindanao (Brenske 1892 a, 1894 a; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14F893FCDCF805.taxon	materials_examined	Type locality. “ Mt. Canla-on, Negros Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14F893FCDCF805.taxon	distribution	Distribution in the Philippines. Visayas (Negros Island) (Matsumoto 2008 c, 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FF0BFCDCFEBC.taxon	materials_examined	Type locality. “ Southern Cebu, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FF0BFCDCFEBC.taxon	distribution	Distribution in the Philippines. Visayas (Cebu) (Itoh 2003 b; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FE0FFCDCFD5C.taxon	materials_examined	Type locality. “ Mt. Canla-on, Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FE0FFCDCFD5C.taxon	distribution	Distribution in the Philippines. Visayas (Leyte, Negros, and Panay) and Mindanao (Bukidnon) (Matsumoto 2008 c, 2015, 2022). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FD2FFCDCFC7C.taxon	materials_examined	Type locality. “ Palawan, Dumaran, Cuyo, Panay, and Mindoro Isls. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FD2FFCDCFC7C.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro and Palawan) and Visayas (Panay) (Itoh 2003 a; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FC4FFCDCFB1C.taxon	materials_examined	Type locality. “ Manilla. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FC4FFCDCFB1C.taxon	distribution	Distribution in the Philippines. Luzon (Batangas, Manila, Pangasinan, and Quezon Province) and Mindanao (Bukidnon) (Brenske 1892 b, 1894 a; Matsumoto 2015; Calcetas et al. 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FA73FCDCF964.taxon	materials_examined	Type locality. “ Luzon, Nueva Vizcaya Province, Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FA73FCDCF964.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1924 b; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FB6FFCDCFA18.taxon	materials_examined	Type locality. “ Palawan, Bacuit. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FB6FFCDCFA18.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Moser 1911 a; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14F977FCDCF860.taxon	materials_examined	Type locality. “ Luzon: St. Tomas. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14F977FCDCF860.taxon	distribution	Distribution in the Philippines. Luzon (Batangas) (Moser 1926; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FF0BFCDCFEBC.taxon	materials_examined	Type locality. “ Mt. Malindang, Mindanao Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FF0BFCDCFEBC.taxon	distribution	Distribution in the Philippines. Mindanao (Misamis Occidental) (Matsumoto 2010, 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FE0FFCDCFDB8.taxon	materials_examined	Type locality. “ Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FE0FFCDCFDB8.taxon	distribution	Distribution in the Philippines. Mindanao (Misamis Occidental) (Brenske 1894 b; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FD13FCDCFC84.taxon	materials_examined	Type locality. “ Mt. Halcon, Is. Mindoro. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FD13FCDCFC84.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro, Palawan, and Quezon) (Matsumoto 2010, 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FC17FCDCFB80.taxon	materials_examined	Type locality. “ Luzon (Mt. Makiling). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FC17FCDCFB80.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Moser 1915 b; Matsumoto, 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FB1BFCDCFA8C.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FB1BFCDCFA8C.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Brenske 1892 a; Itoh 2003 a; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FA1FFCDCF988.taxon	materials_examined	Type locality. “ Insulae Philippinae. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FA1FFCDCF988.taxon	distribution	Distribution in the Philippines. Luzon (Brenske 1896; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14F9E3FCDCF8A8.taxon	materials_examined	Type locality. “ Cuernos Mountain, Oriental Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14F9E3FCDCF8A8.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro), Visayas (Negros Oriental), and Mindanao (Bukidnon and Cotabato) (Chapin 1931 b; Itoh 2003 b; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2818FF14F803FCDCFE98.taxon	materials_examined	Type locality. “ La Carlota, Occidental Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2818FF14F803FCDCFE98.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro), Visayas (Negros Oriental), and Mindanao (Chapin 1931 b; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FEF3FCDCFDE4.taxon	materials_examined	Type locality. “ Sorsogon, Southern Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FEF3FCDCFDE4.taxon	distribution	Distribution in the Philippines. Luzon (Sorsogon) (Chapin 1931 b; Matsumoto 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FDF7FCDCFC84.taxon	materials_examined	Type locality. “ La Carlota, Occidental Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FDF7FCDCFC84.taxon	distribution	Distribution in the Philippines. Luzon (Palawan and Polillo), Visayas (Leyte, Negros Island, and Panay), and Mindanao (Bukidnon, Misamis Occidental, and Davao) (Chapin 1931 b; Matsumoto 2015, 2022). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FC17FCDCFBA4.taxon	materials_examined	Type locality. “ Mt. Balocawihay, C. Leyte, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FC17FCDCFBA4.taxon	distribution	Distribution in the Philippines. Visayas (Leyte and Samar) and Mindanao (Misamis Occidental, Surigao del Norte, and Surigao del Sur) (Matsumoto 2008 c, 2015, 2022). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FB37FCDCFAA0.taxon	materials_examined	Type locality. “ Mt. Halcon, Is. Mindoro, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FB37FCDCFAA0.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro, Palawan, and Quezon) (Matsumoto 2010, 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FA3BFC92F9AC.taxon	materials_examined	Type locality. “ Borneo (Brunei). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FA3BFC92F9AC.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Moser 1912; Matsumoto 2015). Overall distribution. The Philippines and Borneo.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14F93FFBC2F84C.taxon	materials_examined	Type locality. “ Borneo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14F93FFBC2F84C.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Brenske 1894 a; Schultze 1916 a; Matsumoto 2015). Overall distribution. The Philippines and Borneo (Sharp 1876; Brenske 1894 a).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14F9E3FAB0F84D.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14F9E3FAB0F84D.taxon	distribution	Distribution in the Philippines. Luzon (Bacoor, Benguet, Cavite, Laguna, La Union, Manila, Romblon, and Zambales) and Visayas (Negros) (Blanchard 1850; Pic 1905; Ohaus 1910, 1912 a; Chapin 1931 a). Overall distribution. The Philippines, Malaysia, and Vietnam (Ohaus 1912 a; Chapin 1931 a).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14FAC3FCDCF9D1.taxon	materials_examined	Type locality. “ La Carlota, Occidental Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14FAC3FCDCF9D1.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Chapin 1931 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14FF43FCDCFE75.taxon	materials_examined	Type locality. “ Mt. Apo, Mindanao Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14FF43FCDCFE75.taxon	distribution	Distribution in the Philippines. Visayas (Panay) and Mindanao (Bukidnon, Cotabato, and Davao) (Matsumoto 2005). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14FDBFFCDCFD15.taxon	materials_examined	Type locality. “ Basilan Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14FDBFFCDCFD15.taxon	distribution	Distribution in the Philippines. Mindanao (Basilan and Bukidnon) (Itoh 1996). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14FCDFFCDCFBC9.taxon	materials_examined	Type locality. “ Mindanao, Surigao Province, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14FCDFFCDCFBC9.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FF43FCDCFE51.taxon	materials_examined	Type locality. “ Basilan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FF43FCDCFE51.taxon	distribution	Distribution in the Philippines. Mindanao (Basilan) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FE2BFCDCFDB9.taxon	materials_examined	Type locality. “ Süd Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FE2BFCDCFDB9.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ohaus 1914 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FD13FCDCFC85.taxon	materials_examined	Type locality. “ Luzon; S. Luzon, Albay und Mittel Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FD13FCDCFC85.taxon	distribution	Distribution in the Philippines. Luzon (Albay, Oriental Mindoro, and Romblon) (Ohaus 1912 a; Calcetas et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FC4FFB19FB31.taxon	materials_examined	Type locality. “ Sumatra. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FC4FFB19FB31.taxon	distribution	Distribution in the Philippines. Luzon (Ohaus, 1912 b). Overall distribution. The Philippines, South Africa, India, Sri Lanka, Myanmar, Malaysia, Thailand, Indonesia, Papua New Guinea (Blanchard 1853; Ohaus 1912 b; Calcetas et al. 2021).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FA8BFCB8F989.taxon	materials_examined	Type locality. “ Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FA8BFCB8F989.taxon	distribution	Distribution in the Philippines. Luzon (Manila and Rizal) and Mindanao (Davao del Sur) (Burmeister 1844; Calcetas et al. 2021). Overall distribution. The Philippines,? Malaysia, and? China (Taiwan).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FA8BFCB8F989.taxon	discussion	Remarks. Ohaus (1910) noted that Adoretus ranunculus was also found in Malaysia and Taiwan. These records may represent misidentifications, or the species may indeed be present in those regions. However, it is currently known only from the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14F893FCDCF861.taxon	materials_examined	Type locality. “ Abra Pilar, Luzon, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14F893FCDCF861.taxon	distribution	Distribution in the Philippines. Luzon (Abra Pilar) (Frey 1976). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FF0BFCDCFE51.taxon	materials_examined	Type locality. “ Luzon, Province of Albay, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FF0BFCDCFE51.taxon	distribution	Distribution in the Philippines. Luzon (Albay) (Ohaus 1910, 1915 b; Arrow 1915). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FE2BFCDCFD5D.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FE2BFCDCFD5D.taxon	distribution	Distribution in the Philippines. Luzon (Cagayan Province) (Heller 1893). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FD2FFC50FC7D.taxon	materials_examined	Type locality. “ Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FD2FFC50FC7D.taxon	distribution	Distribution in the Philippines. Luzon (Albay, Batanes, Manila, Mindoro, and Zambales) and Visayas (Negros Occidental) (Burmeister 1844; Ohaus 1915 b, 1918; Blanchard 1851; Schultze 1916 a). Overall distribution. The Philippines and Indonesia (Ohaus, 1910).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FC4FFCDCFBA5.taxon	materials_examined	Type locality. “ Mindanao, Provinz Surigao, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FC4FFCDCFBA5.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FB37FCDCFA19.taxon	materials_examined	Type locality. “ Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FB37FCDCFA19.taxon	distribution	Distribution in the Philippines. Luzon (Manila, Romblon, and Sibuyan) and Visayas (Negros Occidental) (Burmeister 1844; Blanchard 1851; Schultze 1916 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FA73FCDCF939.taxon	materials_examined	Type locality. “ Luzon, Province of Albay, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FA73FCDCF939.taxon	distribution	Distribution in the Philippines. Luzon (Albay, Manila, and Palawan), Visayas (Negros and Samar), and Mindanao (Marawi) (Ohaus 1910, 1918; Schultze 1916 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14F893FCDCF805.taxon	materials_examined	Type locality. “ Luzon, Mt. Banahao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14F893FCDCF805.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Ohaus 1914 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ Mindanao, Agusan, Butuan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Mindanao (Agusan del Norte) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FEF0FCDCFDE2.taxon	materials_examined	Type locality. “ Philippinen: Negros- Insel, Cuernos-Berge. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FEF0FCDCFDE2.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Ohaus 1915 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FDF5FCDCFCCB.taxon	materials_examined	Type locality. “ Island Polilo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FDF5FCDCFCCB.taxon	distribution	Distribution in the Philippines. Luzon (Polillo Island) (Ohaus 1923; Zorn 2006). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FCDDFCDCFBD4.taxon	materials_examined	Type locality. “ Luzon, Province of Camarines, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FCDDFCDCFBD4.taxon	distribution	Distribution in the Philippines. Luzon (Camarines) (Ohaus 1910, 1918). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FBA6FCDCFAD1.taxon	materials_examined	Type locality. “ Mindanao, Zamboanga, Dapitan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FBA6FCDCFAD1.taxon	distribution	Distribution in the Philippines. Visayas (Samar and Bohol) and Mindanao (Surigao del Norte,? Maguindanao, and Zamboanga del Norte) (Ohaus 1923; Zorn 2006). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FAABFCDCF9DA.taxon	materials_examined	Type locality. “ Luzon, Province of Bataan, Lamao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FAABFCDCF9DA.taxon	distribution	Distribution in the Philippines. Luzon (Albay and Bataan) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14F9ACFCDCF926.taxon	materials_examined	Type locality. “ Luzon, Manila. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14F9ACFCDCF926.taxon	distribution	Distribution in the Philippines. Luzon (Abra, Albay, and Manila) and Visayas (Negros Occidental) (Ohaus 1912 a, 1918). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14F8B1FCDCF823.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14F8B1FCDCF823.taxon	distribution	Distribution in the Philippines. Luzon and Mindanao (Burmeister 1844; Schultze 1916 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FF0BFD93FDC0.taxon	materials_examined	Type locality. “ Manilla. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FF0BFD93FDC0.taxon	distribution	Distribution in the Philippines. Luzon (Benguet and Manila) (Burmeister 1844; Blanchard 1850; Harold 1869; Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FF0BFD93FDC0.taxon	discussion	Remarks. Heller (1893) provided detailed descriptions of the morphological variations of the male and female Anomala chalybea.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FDDBFCDCFD28.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FDDBFCDCFD28.taxon	distribution	Distribution in the Philippines. Luzon (Burmeister 1844). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FC83FCDCFC34.taxon	materials_examined	Type locality. “ Mindanao, Agusan River. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FC83FCDCFC34.taxon	distribution	Distribution in the Philippines. Mindanao (Agusan) (Ohaus 1912 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FB87FCDCFB30.taxon	materials_examined	Type locality. “ Nord-Luzon, Mt. Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FB87FCDCFB30.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Ohaus 1916). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FA8BFCDCF9AC.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FA8BFCDCF9AC.taxon	distribution	Distribution in the Philippines. Luzon (Abra, Baguio, Benguet, Palawan, and Manila) (Chevrolat 1841; Burmeister 1844; Blanchard 1851; Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14F93FFCDCF894.taxon	materials_examined	Type locality. “ Mindanao, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14F93FFCDCF894.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281EFF14F8E7FCDCFF0D.taxon	materials_examined	Type locality. “ Northwestern Panay. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281EFF14F8E7FCDCFF0D.taxon	distribution	Distribution in the Philippines. Visayas (Panay) (Ohaus 1924 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FE9FFCDCFE16.taxon	materials_examined	Type locality. “ Mindanao, Dapitan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FE9FFCDCFE16.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Ohaus 1915 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FE60FCDCFD12.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FE60FCDCFD12.taxon	distribution	Distribution in the Philippines. Luzon (Burmeister 1844). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FD65FCDCFC1F.taxon	materials_examined	Type locality. “ Luzon, Manila and Dolores, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FD65FCDCFC1F.taxon	distribution	Distribution in the Philippines. Luzon (Manila and Quezon Province) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FC6EFCDCFB18.taxon	materials_examined	Type locality. “ Luzon: Banahao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FC6EFCDCFB18.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao) (Ohaus 1915 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FB73FCDCFA41.taxon	materials_examined	Type locality. “ Manilla. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FB73FCDCFA41.taxon	distribution	Distribution in the Philippines. Luzon (Manila) and Mindanao (Candèze 1869; Ohaus 1918). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FA58FCDCF9B6.taxon	materials_examined	Type locality. “ Kabasalan, Zamboanga, Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FA58FCDCF9B6.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga Sibugay) (Frey 1976). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14F901FCDCF8B3.taxon	materials_examined	Type locality. “ Luzon, Mt. Banahao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14F901FCDCF8B3.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao) (Ohaus 1914 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692192811FF14F80AFCDCFF0C.taxon	materials_examined	Type locality. “ N. Palawan, Binaluan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692192811FF14F80AFCDCFF0C.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ohaus 1914 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FE9FFCDCFE08.taxon	materials_examined	Type locality. “ Luzon, Province of Cagayan, Cape Engaño, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FE9FFCDCFE08.taxon	distribution	Distribution in the Philippines. Luzon (Cagayan Province) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FE63FCDCFD70.taxon	materials_examined	Type locality. “ Samar. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FE63FCDCFD70.taxon	distribution	Distribution in the Philippines. Visayas (Samar) (Ohaus 1927). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FD4BFCDCFC7C.taxon	materials_examined	Type locality. “ Luzon, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FD4BFCDCFC7C.taxon	distribution	Distribution in the Philippines. Luzon (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FC4FFCDCFBA4.taxon	materials_examined	Type locality. “ Mann, Samar. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FC4FFCDCFBA4.taxon	distribution	Distribution in the Philippines. Visayas (Samar) (Ohaus 1927). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FB37FC5EF988.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FB37FC5EF988.taxon	distribution	Distribution in the Philippines. Luzon (Bulacan, Manila, Mindoro, Palawan, and Tarlac), Visayas (Leyte), and Mindanao (Zamboanga) (Burmeister 1844; Blanchard 1851; Ohaus 1910, 1915 a, 1915 b). Overall distribution. The Philippines and Borneo (Blanchard 1851).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14F9E3FCDCF8F0.taxon	materials_examined	Type locality. “ Negros, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14F9E3FCDCF8F0.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) and Visayas (Negros) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162810FF14F8CBFCDCFF0C.taxon	materials_examined	Type locality. “ Luzon, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162810FF14F8CBFCDCFF0C.taxon	distribution	Distribution in the Philippines. Luzon (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FE9FFCDCFE08.taxon	materials_examined	Type locality. “ Luzon, Mt. Makiling. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FE9FFCDCFE08.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Ohaus 1914 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FE63FCDCFD14.taxon	materials_examined	Type locality. “ Ins. Philippin., S. Luzon: Banahao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FE63FCDCFD14.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao) (Ohaus 1915 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FD67FCDCFBC8.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FD67FCDCFBC8.taxon	distribution	Distribution in the Philippines. Luzon (Cagayan Province and Manila) (Blanchard 1850; Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FBA3FBD8FAA0.taxon	materials_examined	Type locality. “ Bourou. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FBA3FBD8FAA0.taxon	distribution	Distribution in the Philippines. Luzon (Manila) and Visayas (Negros) (Burmeister 1844; Blanchard 1851; Ohaus 1910, 1912 a, 1916 a; Schultze 1916 a). Overall distribution. The Philippines and Indonesia (Ohaus 1916 a; Zorn 2011).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FA3BFCDCF988.taxon	materials_examined	Type locality. “ Mindanao, Zamboanga, Dapitan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FA3BFCDCF988.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14F9E3FCDCF894.taxon	materials_examined	Type locality. “ Luzon, Province of Cagayan, Aparri, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14F9E3FCDCF894.taxon	distribution	Distribution in the Philippines. Luzon (Cagayan Province) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172813FF14F8E7FCDCFF0C.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172813FF14F8E7FCDCFF0C.taxon	distribution	Distribution in the Philippines. Luzon (Ohaus 1912 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FE9FFCDCFE74.taxon	materials_examined	Type locality. “ Mindanao, Surigao, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FE9FFCDCFE74.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FE47FCDCFD5C.taxon	materials_examined	Type locality. “ Philippin. Inseln, Mindanao, Davao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FE47FCDCFD5C.taxon	distribution	Distribution in the Philippines. Mindanao (Davao) (Ohaus 1913). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FD2FFCDCFC58.taxon	materials_examined	Type locality. “ Süd-Luzon, Mt. Isarog. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FD2FFCDCFC58.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur) (Ohaus 1916). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FC33FCDCFB80.taxon	materials_examined	Type locality. “ Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FC33FCDCFB80.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ohaus 1930 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FB1BFCDCFAE8.taxon	materials_examined	Type locality. “ Mindanao, Zamboanga. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FB1BFCDCFAE8.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga) (Ohaus 1930 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FAC3FCDCF9F4.taxon	materials_examined	Type locality. “ Batanes Islands, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FAC3FCDCF9F4.taxon	distribution	Distribution in the Philippines. Luzon (Batanes) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14F9C7FCDCF8DC.taxon	materials_examined	Type locality. “ Luzon, Baguio. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14F9C7FCDCF8DC.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Ohaus 1930 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14F8AFFCDCF804.taxon	materials_examined	Type locality. “ Ins. Philippinenses. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14F8AFFCDCF804.taxon	distribution	Distribution in the Philippines. Luzon (Manila) (Candèze 1869). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FF0BFCDCFE50.taxon	materials_examined	Type locality. “ Mindanao, Camp Keithley P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FF0BFCDCFE50.taxon	distribution	Distribution in the Philippines. Luzon (Palawan), Visayas (Leyte), and Mindanao (Lanao del Sur) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FE2BFCDCFD5C.taxon	materials_examined	Type locality. “ Palawan, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FE2BFCDCFD5C.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FD2FFCDCFC7C.taxon	materials_examined	Type locality. “ Manilla. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FD2FFCDCFC7C.taxon	distribution	Distribution in the Philippines. Luzon (Benguet and Cagayan Province), Visayas (Leyte), and Mindanao (Lanao del Sur) (Candèze 1869; Ohaus 1910, 1918). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FC4FFCDCFBA4.taxon	materials_examined	Type locality. “ Mindanao, Ifugao, Mount Polis. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FC4FFCDCFBA4.taxon	distribution	Distribution in the Philippines. Luzon (Ifugao) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FB37FCDCFAA0.taxon	materials_examined	Type locality. “ Luzon, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FB37FCDCFAA0.taxon	distribution	Distribution in the Philippines. Luzon (Albay) and Visayas (Negros Occidental) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FA3BFCDCF988.taxon	materials_examined	Type locality. “ Luzon, Manila; Polillo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FA3BFCDCF988.taxon	distribution	Distribution in the Philippines. Luzon (Manila and Polillo Island) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14F9E3FCDCF8F0.taxon	materials_examined	Type locality. “ Sibuyan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14F9E3FCDCF8F0.taxon	distribution	Distribution in the Philippines. Luzon (Sibuyan Island) (Ohaus 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152815FF14F8CBFCDCFF20.taxon	materials_examined	Type locality. “ Samar. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152815FF14F8CBFCDCFF20.taxon	distribution	Distribution in the Philippines. Visayas (Samar) and Mindanao (Agusan del Norte and Surigao) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FEBBFCDCFE2C.taxon	materials_examined	Type locality. “ Luzon, Benguet, Baguio. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FEBBFCDCFE2C.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Ohaus 1912 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FDBFFCDCFD14.taxon	materials_examined	Type locality. “ Luzon, Manila, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FDBFFCDCFD14.taxon	distribution	Distribution in the Philippines. Luzon (Manila) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FD67FCDCFC10.taxon	materials_examined	Type locality. “ Luzon, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FD67FCDCFC10.taxon	distribution	Distribution in the Philippines. Luzon (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FC6BFCDCFB78.taxon	materials_examined	Type locality. “ Palawan, Iwahig. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FC6BFCDCFB78.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ohaus 1912 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FB53FCDCFAA0.taxon	materials_examined	Type locality. “ Sibuyan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FB53FCDCFAA0.taxon	distribution	Distribution in the Philippines. Luzon (Sibuyan Island) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FA3BFCDCF938.taxon	materials_examined	Type locality. “ Insel Luçon (Manille). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FA3BFCDCF938.taxon	distribution	Distribution in the Philippines. Luzon (Catanduanes and Manila) (Eschscholtz 1822; Laporte 1840; Ohaus 1912 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122814FF14F893FED0FF0C.taxon	materials_examined	Type locality. “ Manille. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122814FF14F893FED0FF0C.taxon	distribution	Distribution in the Philippines. Luzon (Albay, Bataan, Manila, Palawan, Polillo, Rizal, and Tarlac) and Visayas (Negros Island) (Burmeister 1844; Blanchard 1851; Ohaus 1918). Overall distribution. The Philippines, Borneo, Malaysia, Guam, and Northern Mariana Islands (Dunlap et al. 2016).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FE9FFCDCFE74.taxon	materials_examined	Type locality. “ Mindanao, Surigao, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FE9FFCDCFE74.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FE47FCDCFD5C.taxon	materials_examined	Type locality. “ Ins. Philippin.: Montalban. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FE47FCDCFD5C.taxon	distribution	Distribution in the Philippines. Luzon (Rizal) (Ohaus 1915 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FD2FFCDCFC84.taxon	materials_examined	Type locality. “ Luzon, Zambales; S. Luzon, Albay und Mindoro. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FD2FFCDCFC84.taxon	distribution	Distribution in the Philippines. Luzon (Albay, Mindoro, and Zambales) (Ohaus 1912 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FC17FCDCFB80.taxon	materials_examined	Type locality. “ Luzon, Manila, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FC17FCDCFB80.taxon	distribution	Distribution in the Philippines. Luzon (Manila) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FB1BFCDCFAE8.taxon	materials_examined	Type locality. “ Luzon, Provinz Benguet, Trinidad. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FB1BFCDCFAE8.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) (Ohaus 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FAC3FCDCF9D0.taxon	materials_examined	Type locality. “ Mindanao, Surigao, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FAC3FCDCF9D0.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14F9ABFCDCF8DC.taxon	materials_examined	Type locality. “ Philippinen: Benguet, Baguio. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14F9ABFCDCF8DC.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Ohaus 1915 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132817FF14F8E7FCDCFF0C.taxon	materials_examined	Type locality. “ Nord Palawan: Bacuit. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132817FF14F8E7FCDCFF0C.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ohaus 1914 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FE47FC42FD14.taxon	materials_examined	Type locality. “ Mt. Apo, North Cotabato, Mindanao Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FE47FC42FD14.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FE47FC42FD14.taxon	discussion	Remarks. This species is a junior homonym of Callistethus mindanaoensis Wada, 2002, and in accordance with Article 52 of the ICZN, a replacement name must be proposed.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FE9FFCDCFE74.taxon	materials_examined	Type locality. “ Mt. Apo, Mindanao Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FE9FFCDCFE74.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Wada 2002). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FD67FCDCFC7C.taxon	materials_examined	Type locality. “ Mindanao, Provinz Surigao, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FD67FCDCFC7C.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FC4FFCDCFBA4.taxon	materials_examined	Type locality. “ Bindoyan, Palawan Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FC4FFCDCFBA4.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Wada 1998 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FB37FC53FAA0.taxon	materials_examined	Type locality. “ Manilla. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FB37FC53FAA0.taxon	distribution	Distribution in the Philippines. Luzon (Catanduanes, Laguna, Mindoro, and Palawan) and Visayas (Negros) (Candèze 1869; Ohaus 1915 a). Overall distribution. The Philippines and Malaysia (Ohaus 1915 a).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FA73FE0EF938.taxon	materials_examined	Type locality. “ Java. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FA73FE0EF938.taxon	distribution	Distribution in the Philippines. Luzon (Manila) (Ohaus 1908, 1910; Schultze 1916 a). Overall distribution. The Philippines and Indonesia (Hope 1835; Schultze 1916 a).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FA73FE0EF938.taxon	discussion	Remarks. This species is common in Java, but its presence in the Philippines, as noted by Ohaus (1910), remains doubtful.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14F893FBE4F860.taxon	materials_examined	Type locality. “ Borneo: Kinibalu im Nordosten und Tandjong- Bandjermasin im S-O; Süd Luzon: Albay. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14F893FBE4F860.taxon	distribution	Distribution in the Philippines. Luzon (Albay) (Ohaus, 1908). Overall distribution. The Philippines, Indonesia, and Malaysia (Ohaus 1908).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FE63FCDCFD71.taxon	materials_examined	Type locality. “ Luzon, Nueva Vizcaya, Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FE63FCDCFD71.taxon	distribution	Distribution in the Philippines. Luzon (Baguio and Nueva Vizcaya) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FD4BFCDCFC59.taxon	materials_examined	Type locality. “ Nord-Luzon, Mt. Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FD4BFCDCFC59.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Ohaus 1916). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FC33FCDCFBA5.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FC33FCDCFBA5.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) and Visayas (Negros Occidental) (Newman 1841). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FB37FCDCFA8D.taxon	materials_examined	Type locality. “ Luzon, bei der Lagune su ̈ dlich von Manila gesammelt. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FB37FCDCFA8D.taxon	distribution	Distribution in the Philippines. Luzon (Manila and Laguna) (Ohaus 1905, 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FA1FFCDCF9F5.taxon	materials_examined	Type locality. “ Insulae Philippinae. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FA1FFCDCF9F5.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Kraatz 1892; Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14F9C7FCDCF8DD.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14F9C7FCDCF8DD.taxon	distribution	Distribution in the Philippines. Luzon (Manila and Nueva Vizcaya) (Kraatz 1892; Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14F8AFFCDCF805.taxon	materials_examined	Type locality. “ Süd Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14F8AFFCDCF805.taxon	distribution	Distribution in the Philippines. Luzon (Ohaus 1930). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ Palawan, Bacuit, Mt. Capoas, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ Nord-Luzon, Mt. Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Ohaus 1916). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FEF0FCDCFDCE.taxon	materials_examined	Type locality. “ Mindanao, Surigao, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FEF0FCDCFDCE.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FDD9FCDCFD37.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FDD9FCDCFD37.taxon	distribution	Distribution in the Philippines. Luzon (Ohaus, 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FC81FCDCFC1F.taxon	materials_examined	Type locality. “ Süd-Luzon, Mt. Isarog. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FC81FCDCFC1F.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur) (Ohaus 1916). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FC6EFCDCFB44.taxon	materials_examined	Type locality. “ Nord-Luzon, Mt. Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FC6EFCDCFB44.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Ohaus 1916). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FB57FCDCF9B6.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FB57FCDCF9B6.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Newman 1841; Kraatz 1892; Arrow 1899; Ohaus 1910, 1914 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14F979FCDCF857.taxon	materials_examined	Type locality. “ Mindanao, Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14F979FCDCF857.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1923). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2828FF14F826FCDCFF21.taxon	materials_examined	Type locality. “ Luzon, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2828FF14F826FCDCFF21.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14FEF3FCDCFDC1.taxon	materials_examined	Type locality. “ Ins. Philippin., Negros. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14FEF3FCDCFDC1.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Ohaus 1905). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14FDDBFCDCFCE1.taxon	materials_examined	Type locality. “ Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14FDDBFCDCFCE1.taxon	distribution	Distribution in the Philippines. Luzon (Albay, Cagayan Province, and Benguet) and Mindanao (Burmeister 1844; Ohaus 1910, 1912 a; Schultze 1916 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14FCFBFCDCFAD5.taxon	materials_examined	Type locality. “ Ins. Philippinae. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14FCFBFCDCFAD5.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) and Visayas (Negros, Samar, and Leyte) (Kraatz 1892; Ohaus 1914 b; Schultze 1916 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14FAA7FCDCF9F5.taxon	materials_examined	Type locality. “ Habitat in insulis Philippinis (Luzon et Mindanao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14FAA7FCDCF9F5.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) and Mindanao (Agusan) (Heller 1891; Ohaus 1910, 1912 a, 1918; Schultze 1916). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14F9C7FCDCF8DD.taxon	materials_examined	Type locality. “ Nord Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14F9C7FCDCF8DD.taxon	distribution	Distribution in the Philippines. Luzon (Ohaus 1912 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F282BFF14F8E7FCDCFF0C.taxon	materials_examined	Type locality. “ Luzon, Province of Benguet, Pauai, P. I. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F282BFF14F8E7FCDCFF0C.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) (Ohaus 1910, 1918). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282BFF14FCFAFD33FBA4.taxon	materials_examined	Type locality. “ Pulo-Pinang. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282BFF14FCFAFD33FBA4.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) and Mindanao (Kuijten 1992). Overall distribution. The Philippines, India, Malaysia, Myanmar, Singapore, Thailand, and Indonesia (Hongsuwong & Photaworn 2025).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282BFF14FB36FBD1FA44.taxon	materials_examined	Type locality. “ Mt. Kawi, Java; Sumatra. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282BFF14FB36FBD1FA44.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ohaus 1910; Kuijten 1992). Overall distribution. The Philippines, Malaysia, and Indonesia (Kuijten 1992).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282AFF14FA56FCD5FF0D.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282AFF14FA56FCD5FF0D.taxon	distribution	Distribution in the Philippines. Luzon (Batanes, Benguet, Bataan, Babuyan Islands, Camarines, Mindoro, Romblon, Sibuyan, Marinduque, and Rizal), Visayas (Bohol, Negros, Panay, Leyte, and Samar), and Mindanao (Bukidnon and Marawi) (Westwood 1841; 1842; Blanchard 1851; Ohaus 1910; Wada 1989; Kuijten 1992; Wada 2015). Overall distribution. The Philippines and China (Taiwan) (Wada 2015).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282AFF14FA56FCD5FF0D.taxon	discussion	Remarks. Kuijten (1992) placed Parastasia yukioi Wada, 1989 as a junior synonym of P. canaliculata. However, phylogenetic analyses conducted by Wada (2015), based on morphological characters, supported the recognition of P. yukioi as a separate and valid species. Wada highlighted distinguishing features such as the shape of the male genitalia, the structure of the inner sac, and a generally smaller body size. Therefore, P. yukioi will be treated as a distinct species.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282BFF14FE9FFA8EFE08.taxon	materials_examined	Type locality. “ Ind. Orient. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282BFF14FE9FFA8EFE08.taxon	distribution	Distribution in the Philippines. Luzon (Manila) (Blanchard 1851; Ohaus 1918). Overall distribution. The Philippines, Korea, Vietnam, and China (including Taiwan) (Kim 2011).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282BFF14FE63FCDCFD70.taxon	materials_examined	Type locality. “ Insulae Philippinae, Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282BFF14FE63FCDCFD70.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) (Kraatz 1892; Ohaus 1910). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FE9FFE8DFDC1.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FE9FFE8DFDC1.taxon	distribution	Distribution in the Philippines. Luzon (Manila and Sibuyan) and Mindanao (Marawi) (Westwood 1841; Kuijten 1992; Wada 2015). Overall distribution. The Philippines, Malaysia, Indonesia, and Papua New Guinea (Kuijten 1992; Wada 2015).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FDD8FCB8FC33.taxon	materials_examined	Type locality. “ Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FDD8FCB8FC33.taxon	distribution	Distribution in the Philippines. Luzon (Bicol), Visayas (Panay, Negros, Samar, and Leyte), and Mindanao (Bukidnon, Agusan del Sur, Sarangani, Misamis Oriental, Surigao, and Davao) (Westwood 1841; Kuijten 1992; Wada 2015). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FDD8FCB8FC33.taxon	discussion	Remarks. In the article by Schultze (1916 a), P. discolor scutellaris was noted to occur on the island of Luzon. However, no evidence to date of its existence has been documented. This may be erroneous and mislabeled as ‘ scutellaris ’ instead of ‘ discolor ’.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FB8AFC79FB3C.taxon	materials_examined	Type locality. “ Luzon, Kalinga, Balbalan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FB8AFC79FB3C.taxon	distribution	Distribution in the Philippines. Luzon (Kalinga, Mountain Province, Camiguin, Cagayan Province, and Masbate) (Ohaus 1923; Kuijten 1992; Wada 2015). Overall distribution. The Philippines and Sulawesi (Wada 2015).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FA8FFCDCFA39.taxon	materials_examined	Type locality. “ Mt. Canlaon, Negros Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FA8FFCDCFA39.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Wada 1989, 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14F990FCDCF96E.taxon	materials_examined	Type locality. “ Cuernos Mts., Negros, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14F990FCDCF96E.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Wada 2008). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14F978FCDCF857.taxon	materials_examined	Type locality. “ Mt. Polis, Mt. Province, North Luzon, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14F978FCDCF857.taxon	distribution	Distribution in the Philippines. Luzon (Mountain Province) (Wada 2016). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282DFF14F821FCDCFF21.taxon	materials_examined	Type locality. “ Mt. Dolang dolang, Malaybarlay, Bukidnon, Mindanao Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282DFF14F821FCDCFF21.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon, South Cotabato, and Davao) (Wada 2007, 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FEBBFCDCFE16.taxon	materials_examined	Type locality. “ Mt. Canlaon, Negros Is., Philippine Isls. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FEBBFCDCFE16.taxon	distribution	Distribution in the Philippines. Visayas (Negros Island) (Wada 1997). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FE60FC27FCCB.taxon	materials_examined	Type locality. “ Philippine Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FE60FC27FCCB.taxon	distribution	Distribution in the Philippines. Luzon (Marinduque, Romblon, Bicol, and Mindoro), Visayas (Negros), and Mindanao (Westwood 1841; Fairmaire 1883; Ohaus 1900; Kuijten 1992). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FCDDFCDCFBE8.taxon	materials_examined	Type locality. “ Luzon (Engano). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FCDDFCDCFBE8.taxon	distribution	Distribution in the Philippines. Luzon (Cagayan Province, Laguna, Mindoro, Quirino, and Quezon Province) (Ohaus 1901, 1905, 1930 a; Kuijten 1992; Wada 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FBC2FCDCFAD1.taxon	materials_examined	Type locality. “ Mt. Kitanglad, North Mindanao Is., Philippines Isls. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FBC2FCDCFAD1.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon) (Wada 1997). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FAABFAD3F9DD.taxon	materials_examined	Type locality. “ Sumatra. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FAABFAD3F9DD.taxon	distribution	Distribution in the Philippines. Luzon (Palawan, Mountain Province, Quezon, and Mindoro) and Mindanao (Zamboanga) (Ohaus 1910, 1918; Wada, 2015). Overall distribution. The Philippines, Indonesia, Malaysia, and Thailand (Kuijten 1992; Wada 2015).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14F9ACFCDCF926.taxon	materials_examined	Type locality. “ Bindoyan, Is. Palawan, Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14F9ACFCDCF926.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Wada 1989, 2015). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282CFF14F8E9FCDCFF0C.taxon	materials_examined	Type locality. “ Nord-Luzon, Mt. Imugan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282CFF14F8E9FCDCFF0C.taxon	distribution	Distribution in the Philippines. Luzon (Nueva Vizcaya) (Ohaus 1916). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282CFF14FAA7FCDCFA3C.taxon	distribution	Distribution in the Philippines. Luzon (Mount Banahao) (Boucomont 1925). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282CFF14F98FFCDCF964.taxon	materials_examined	Type locality. “ Dingalan Bay, East Luzon, Philippines Islands. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282CFF14F98FFCDCF964.taxon	distribution	Distribution in the Philippines. Luzon (Aurora) (Pereira 1954). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282CFF14FE9EFDE3FD28.taxon	materials_examined	Type locality. “ Luzon, Mt. Banahao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282CFF14FE9EFDE3FD28.taxon	distribution	Distribution in the Philippines. Luzon (Aurora, Camarines Sur, Laguna, Mindoro, and Mt. Banahao), Visayas (Negros Occidental), and Mindanao (Davao de Oro, Davao del Sur, Kidapawan, and Lanao del Sur) (Ohaus 1930 b; Wada 1998 b; Xu & Zhao 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282CFF14FE9EFDE3FD28.taxon	discussion	Remarks. The longstanding taxonomic confusion regarding the presence of Lutera nigromaculata Ohaus, 1900, in the Philippines was recently resolved by Xu and Zhao (2024). The specimen of L. nigromaculata from Camarines, examined by Ohaus and currently deposited at the MFNB in Berlin, Germany, is likely the first known individual of L. nigrita.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282CFF14FCFBFBE8FBEC.taxon	materials_examined	Type locality. “ Philippinen: Baguio auf Benguet. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282CFF14FCFBFBE8FBEC.taxon	distribution	Distribution in the Philippines. Luzon (Baguio, Mt. Banahao, and Nueva Vizcaya) and Mindanao (? Bukidnon) (Ohaus 1915 b; Kuijten 1988). Overall distribution. The Philippines, Indonesia, and Vietnam (Kuijten 1988).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282FFF14F8E7FCDCFE99.taxon	materials_examined	Type locality. “ Ins. Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282FFF14F8E7FCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Bataan, Benguet, Mindoro, Sibuyan, and Palawan), Visayas (Negros Occidental), and Mindanao (Bukidnon, Davao, Marawi, and Misamis Oriental) (Sharp 1875; Boucomont 1914; Schultze 1916 a; Balthasar 1963 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14FEF0FE2FFCA6.taxon	materials_examined	Type locality. “ India. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14FEF0FE2FFCA6.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Schultze 1916 a). Overall distribution. The Philippines, India, Sri Lanka, Afghanistan, Nepal, China (including Taiwan), Vietnam, Laos, Cambodia, Thailand, Malaysia, Pakistan, and Singapore.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14FEF0FE2FFCA6.taxon	discussion	Remarks. Although records of Catharsius molossus from the Philippines are scarce in the literature, the species is known to occur in Palawan and is entirely absent from other regions of the archipelago, as confirmed by Ochi and Kon (pers. comm.). During a recent entomological survey in Bukidnon, Mindanao, the first author collected Catharsius specimens that differed markedly from other Mindanao populations by having entirely reddish-brown pubescence. These specimens closely resembled C. molossus, and after a detailed examination, assisted by Ochi and Kon, they were confirmed to be C. molossus. A more comprehensive treatment of the geographic distribution of Catharsius species in the Philippines will be presented in a separate paper.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14FBA6FCDCFAF5.taxon	materials_examined	Type locality. “ Iles Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14FBA6FCDCFAF5.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro and Pampanga) and Mindanao (Agusan River) (Gillet 1910; Boucomont 1914; Schultze 1916 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14FA3FFCDCF9B6.taxon	materials_examined	Type locality. “ Philippines (Mindoro Is.). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14FA3FFCDCF9B6.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Ochi, 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14F8B1FCDCF823.taxon	materials_examined	Type locality. “ Mt. Apo, Mindanao Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14F8B1FCDCF823.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi 1990). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14FDD9FCDCFD37.taxon	materials_examined	Type locality. “ Philippines, Luzon, Mountain Province, Sagoda. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14FDD9FCDCFD37.taxon	distribution	Distribution in the Philippines. Luzon (Mountain Province) (Paulian 1981). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14FEF0FCDCFDCE.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Queson City. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14FEF0FCDCFDCE.taxon	distribution	Distribution in the Philippines. Luzon (Quezon) (Ochi et al. 2025 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14FB57FC27FA41.taxon	materials_examined	Type locality. “ Philippines, Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14FB57FC27FA41.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Boucomont 1914; Ochi & Kon 1997). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14FC36FB38FB44.taxon	materials_examined	Type locality. “ Ins. Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14FC36FB38FB44.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Sharp 1875; Boucomont 1914). Overall distribution. Endemic species to the Philippines (Pokorny ́ & Zi ́ dek 2014).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14F9C8FECEF8B3.taxon	materials_examined	Type locality. “ Java. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14F9C8FECEF8B3.taxon	distribution	Distribution in the Philippines. Luzon (Bataan, Laguna, and Rizal) and Visayas (Cebu and Negros Oriental) (Barbero et al. 2011). Overall distribution. The Philippines, China, India, Indonesia, Myanmar, Sri Lanka, and Vietnam (Barbero et al. 2011).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692292821FF14F80AFCDAFEC5.taxon	materials_examined	Type locality. “ Bengalia. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692292821FF14F80AFCDAFEC5.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Barbero et al. 2011). Overall distribution. The Philippines, Pakistan, India, Bhutan, Nepal, Sri Lanka, China, Myanmar, Laos, Thailand, Vietnam, and Indonesia (Barbero et al. 2011).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262821FF14F98FFCDCF939.taxon	materials_examined	Type locality. “ Iles Philippines, Mindanao: Iligan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262821FF14F98FFCDCF939.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Norte) (Boucomont 1921, 1924 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262821FF14FD67FE8CFBED.taxon	materials_examined	Type locality. “ India. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262821FF14FD67FE8CFBED.taxon	distribution	Distribution in the Philippines. Luzon and Mindanao (Lansberge 1875; Balthasar 1963 b; Cheung et al. 2018). Overall distribution. This species is widely distributed in the Oriental and Palearctic regions (Han et al. 2021).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262821FF14FBFFFC46FA8D.taxon	materials_examined	Type locality. “ Iles Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262821FF14FBFFFC46FA8D.taxon	distribution	Distribution in the Philippines. Luzon (Laguna, Manila, and Benguet) (Lansberge 1875; Boucomont 1914; Schultze 1916 a; Balthasar 1963 b). Overall distribution. The Philippines and Indonesia (Janssens 1937).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262821FF14FED7FCCCFDE5.taxon	materials_examined	Type locality. “ Kinb. China. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262821FF14FED7FCCCFDE5.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Barbero et al. 2011). Overall distribution. The Philippines, Bhutan, India, Nepal, China, Myanmar, Laos, Thailand, Vietnam, Singapore, Indonesia, and Malaysia (Barbero et al. 2011).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262820FF14F803FBE1FE74.taxon	materials_examined	Type locality. “ Tranquebar. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262820FF14F803FBE1FE74.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Arrow 1931; Balthasar 1963 b; Nomura 1973; Ochi & Kon 1994). Overall distribution. The Philippines, China, India, Indonesia, Japan, Laos, Malaysia, North Korea, Sri Lanka, and Vietnam (Cheung et al. 2018; Han et al. 2021; Ueda et al. 2022).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692272820FF14FDBFFCDCFD14.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Rizai, Tanay. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692272820FF14FDBFFCDCFD14.taxon	distribution	Distribution in the Philippines. Luzon (Rizal) (Ochi et al. 2025 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692272820FF14FC17FCD8FAE8.taxon	materials_examined	Type locality. “ Borneo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692272820FF14FC17FCD8FAE8.taxon	distribution	Distribution in the Philippines. Luzon (Manila and Mindoro) and Visayas (Negros Occidental) (Blanchard 1853; Lansberge 1875; Boucomont 1914; Balthasar 1944, 1963 b). Overall distribution. The Philippines, Cambodia, China (including Taiwan), Guam, India, Indonesia, Myanmar, and Vietnam (Balthasar 1944; Cheung et al. 2018).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692272820FF14FA3BFCDCF940.taxon	materials_examined	Type locality. “ Mindanao: Davao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692272820FF14FA3BFCDCF940.taxon	distribution	Distribution in the Philippines. Mindanao (Davao, Lanao del Norte, and Misamis Oriental) (Boucomont 1914, 1924 a; Balthasar 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692272820FF14F95BFCDCF860.taxon	materials_examined	Type locality. “ Mindanao: Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692272820FF14F95BFCDCF860.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon, Lanao del Norte, Misamis Oriental, and Surigao) (Boucomont 1925; Balthasar 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FF0BFCDCFE2D.taxon	materials_examined	Type locality. “ Manilla. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FF0BFCDCFE2D.taxon	distribution	Distribution in the Philippines. Luzon (Manila and Laguna) and Mindanao (Agusan del Norte and Misamis Oriental) (Boheman 1858; Harold 1862 a; Boucomont 1914). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FDF7FB2BFC7C.taxon	materials_examined	Type locality. “ Borneo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FDF7FB2BFC7C.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Boucomont 1914, 1921, 1924 b; Balthasar 1963 b; Ochi & Kon 1995). Overall distribution. The Philippines, Indonesia, and Malaysia (Ochi & Kon 1995).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FC4FFCDCFB1D.taxon	materials_examined	Type locality. “ Palawan, Binaluan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FC4FFCDCFB1D.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) and Mindanao (Boucomont 1924 a; Balthasar 1963 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FB6FFB93FA45.taxon	materials_examined	Type locality. “ Rizal, Palawan Is., the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FB6FFB93FA45.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ochi & Kon 2007). Overall distribution. The Philippines and? Thailand (Ochi & Kon 2007).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FA57FD71F939.taxon	materials_examined	Type locality. “ China (Hongkong). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FA57FD71F939.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Balthasar 1963 b). Overall distribution. The Philippines, China (including Taiwan), Cambodia, India, Indonesia, Myanmar, Thailand, and Vietnam (Han et al. 2021).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14F893FCDCF861.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., South Lanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14F893FCDCF861.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Sur) (Ochi & Kon 2025 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ Brooks Point, Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ochi & Kon 2004). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FEF3FCDCFDE5.taxon	materials_examined	Type locality. “ Bicol National Park, Nr. Daet, Central Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FEF3FCDCFDE5.taxon	distribution	Distribution in the Philippines. Luzon (Palawan, Bicol, Baguio, and Negros) (Ochi & Kon 2001, 2007). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FD2FFAD1FC11.taxon	materials_examined	Type locality. “ Borneo: Sandakan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FD2FFAD1FC11.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Norte) (Boucomont 1921, 1924 a; Balthasar 1963 b; Ochi & Kon 1995). Overall distribution. The Philippines, Indonesia, and Malaysia (Ochi & Kon 1995; Ueda et al. 2022).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FC6BFCDCFB79.taxon	materials_examined	Type locality. “ Philippines, Mindanao Isl., Bukidnon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FC6BFCDCFB79.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon and Lanao del Sur) (Ochi et al. 2023 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FA8BFC27FA3D.taxon	materials_examined	Type locality. “ Philippines: Palawan; Los Banos, Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FA8BFC27FA3D.taxon	distribution	Distribution in the Philippines. Luzon (Laguna and Palawan) (Boucomont 1914). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14F98FFCDCF965.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14F98FFCDCF965.taxon	distribution	Distribution in the Philippines. Mindanao (Davao, Agusan, and Cotabato) (Ochi et al. 2023 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14F977FCDCF84D.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Bicol. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14F977FCDCF84D.taxon	distribution	Distribution in the Philippines. Luzon (Bicol) (Ochi et al. 2023 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., Davao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Mindanao (Davao) (Ochi et al. 2023 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FEF3FCDCFDC1.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., Lanao del Sur. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FEF3FCDCFDC1.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Sur) (Ochi et al. 2023 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FDDBFCDCFD29.taxon	materials_examined	Type locality. “ Philippines, Leyte Is., Lake Imeruda. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FDDBFCDCFD29.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Ochi et al. 2023 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FC83FCDCFC11.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Lucena. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FC83FCDCFC11.taxon	distribution	Distribution in the Philippines. Luzon (Lucena) (Ochi et al. 2023 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FC6BFCDCFB79.taxon	materials_examined	Type locality. “ Philippines Batan Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FC6BFCDCFB79.taxon	distribution	Distribution in the Philippines. Luzon (Batan) (Ochi et al. 2021, 2023 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FA8BFCDCFA19.taxon	materials_examined	Type locality. “ Philippines, Mindanao, Mt. Apo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FA8BFCDCFA19.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi et al. 2023 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FA73FCDCF895.taxon	materials_examined	Type locality. “ île de Basilan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FA73FCDCF895.taxon	distribution	Distribution in the Philippines. Mindanao (Davao del Norte, Lanao del Norte, Zamboanga del Norte, Surigao, Bukidnon, and Basilan) (Boucomont 1920, 1921, 1924 a; Ochi et al. 2023 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222824FF14F8E7FCDDFF20.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., North Davao, Masara-Mainit. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222824FF14F8E7FCDDFF20.taxon	distribution	Distribution in the Philippines. Mindanao (Davao del Norte) (Ochi et al. 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FEBBFCDCFE08.taxon	materials_examined	Type locality. “ Philippines, Negros Is., Mt. Kanlaon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FEBBFCDCFE08.taxon	distribution	Distribution in the Philippines. Visayas (Negros and Panay) (Ochi et al. 2023 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FE63FCDCFD70.taxon	materials_examined	Type locality. “ Philippines, Samar Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FE63FCDCFD70.taxon	distribution	Distribution in the Philippines. Visayas (Leyte and Samar) (Ochi et al. 2023 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FD4BFCDCFC58.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Queson, Gen Nakar. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FD4BFCDCFC58.taxon	distribution	Distribution in the Philippines. Luzon (Quezon) (Ochi & Kon 2025 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FC33FCDCFB78.taxon	materials_examined	Type locality. “ Sibuyan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FC33FCDCFB78.taxon	distribution	Distribution in the Philippines. Luzon (Sibuyan) (Boucomont 1924 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FA8BFCDCFA18.taxon	materials_examined	Type locality. “ Philippines, Mindanao Isl., Bukidnon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FA8BFCDCFA18.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon) (Ochi et al. 2023 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FA73FCDCF938.taxon	materials_examined	Type locality. “ Palawan (Binaluan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FA73FCDCF938.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Boucomont 1924 a; Ochi et al. 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14F893FCDCF860.taxon	materials_examined	Type locality. “ Queson National Park, Luzon Is., the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14F893FCDCF860.taxon	distribution	Distribution in the Philippines. Luzon (Quezon) (Ochi & Kon 2006). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FF43FCDCFE51.taxon	materials_examined	Type locality. “ Philippines, Mindanao, Davao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FF43FCDCFE51.taxon	distribution	Distribution in the Philippines. Mindanao (Davao) (Ochi et al. 2025 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FE2BFCDCFD15.taxon	materials_examined	Type locality. “ Iles Philippines, Mindanao: Iligan, Dapitan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FE2BFCDCFD15.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Norte and Zamboanga del Norte) (Boucomont 1921, 1924 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FD67FCDCFC11.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., Zamboanga. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FD67FCDCFC11.taxon	distribution	Distribution in the Philippines. Mindanao (Sarangani and Zamboanga) (Ochi et al. 2023 b, 2025 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FC6BFCDCFB79.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Queson, Atimonan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FC6BFCDCFB79.taxon	distribution	Distribution in the Philippines. Luzon (Quezon) (Ochi et al. 2025 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FA8BFE8DF9D1.taxon	materials_examined	Type locality. “ Hongkong. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FA8BFE8DF9D1.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) and Mindanao (Zamboanga) (Ochi et al. 2024). Overall distribution. The Philippines, Japan, China (including Taiwan), Vietnam, and Laos (Ochi et al. 2024).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14F9E3FB64F8A9.taxon	materials_examined	Type locality. “ Java. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14F9E3FB64F8A9.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) and Mindanao (Agusan del Norte) (Boucomont 1924 a). Overall distribution. The Philippines, Indonesia, Myanmar, and India (Ueda et al. 2022).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FF43FCDCFE75.taxon	materials_examined	Type locality. “ Philippines: Mont Banahao, Los Baños. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FF43FCDCFE75.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Boucomont 1920, 1924 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FE47FCDCFD5D.taxon	materials_examined	Type locality. “ Philippines: Batan Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FE47FCDCFD5D.taxon	distribution	Distribution in the Philippines. Luzon (Batan) (Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FD2FFCDCFC85.taxon	materials_examined	Type locality. “ Philippinen, Bango, Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FD2FFCDCFC85.taxon	distribution	Distribution in the Philippines. Mindanao (Frey 1971). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FC4FFB5AFB1D.taxon	materials_examined	Type locality. “ Borneo: Ile Banguey. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FC4FFB5AFB1D.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Boucomont 1924; Balthasar 1963 b). Overall distribution. The Philippines and Malaysia (Boucomont 1914; Ochi & Kon 1994).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FB6FFCDCFA45.taxon	materials_examined	Type locality. “ Philippines, Palawan, Mt. Salakot. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FB6FFCDCFA45.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ochi et al. 2016). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FA57FCDCF965.taxon	materials_examined	Type locality. “ Sibuyan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FA57FCDCF965.taxon	distribution	Distribution in the Philippines. Luzon (Sibuyan, Quezon, and Baguio) (Boucomont 1924 a; Balthasar 1963 b; Ochi et al. 2016). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14F8AFFCDCF805.taxon	materials_examined	Type locality. “ Philippines: Davao, Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14F8AFFCDCF805.taxon	distribution	Distribution in the Philippines. Mindanao (Davao) (Boucomont 1914). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FF0BFBA4FE74.taxon	materials_examined	Type locality. “ Celebes. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FF0BFBA4FE74.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro), and Mindanao (Davao and Zamboanga del Norte) (Schultze 1916 a; Boucomont 1924). Overall distribution. The Philippines and Indonesia (Boucomont 1924).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FDBFFAB4FC84.taxon	materials_examined	Type locality. “ Insel Luçon (Manilla). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FDBFFAB4FC84.taxon	distribution	Distribution in the Philippines. Luzon (Baguio, Cagayan, Manila, and Rizal), Visayas (Dumaguete), and Mindanao (Basilan, Lanao del Norte, and Surigao) (Eschscholtz 1822; Boucomont 1924 a; Schultze 1916 a). Overall distribution. The Philippines, Indonesia, Malaysia, and Singapore (Balthasar 1963 b).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FC17FC27FBEC.taxon	materials_examined	Type locality. “ Asin hot spring near Baguio, the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FC17FC27FBEC.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Ochi & Kon 2006). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FBFFFC27FAD4.taxon	materials_examined	Type locality. “ Bicol National Park, Luzon, the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FBFFFC27FAD4.taxon	distribution	Distribution in the Philippines. Luzon (Bicol) (Ochi & Kon 2006). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FAA7FCDCFA3C.taxon	materials_examined	Type locality. “ Philippines: Samar. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FAA7FCDCFA3C.taxon	distribution	Distribution in the Philippines. Visayas (Samar) (Krikken 1986). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14F98FFCDCF964.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., South Lanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14F98FFCDCF964.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Sur) (Ochi & Kon 2025 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14F977FCDCF860.taxon	materials_examined	Type locality. “ Ile Mindanao (Philippines): Dapitan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14F977FCDCF860.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Boucomont 1920, 1921, 1924 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FF0BFCDCFEBD.taxon	materials_examined	Type locality. “ Iles Philippines (Luzon): Mont Makiling, Mt. Banahao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FF0BFCDCFEBD.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) and Visayas (Negros Oriental) (Boucomont 1920, 1924 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FE0FFCDCFDB9.taxon	materials_examined	Type locality. “ Ile Mindanao (Philippines): Dapitan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FE0FFCDCFDB9.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Boucomont 1920, 1924 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FD13FCDCFCE1.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., Davao del Norte, Masara-Mainit. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FD13FCDCFCE1.taxon	distribution	Distribution in the Philippines. Mindanao (Davao del Norte) (Ochi et al. 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FCFBFCDCFBC9.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Dingalan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FCFBFCDCFBC9.taxon	distribution	Distribution in the Philippines. Luzon (Ochi et al. 2023 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FBA3FCDCFB31.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., Lanao del Sur. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FBA3FCDCFB31.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Sur) (Ochi et al. 2023 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FA8BFB14F9D1.taxon	materials_examined	Type locality. “ Sumatra. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FA8BFB14F9D1.taxon	distribution	Distribution in the Philippines. Luzon (Laguna, Sibuyan, and Palawan) and Mindanao (Surigao) (Harold 1877; Boucomont 1924 a; Balthasar 1963 b). Overall distribution. The Philippines and Indonesia (Harold 1877; Balthasar 1963 b).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14F9ABFCDCF939.taxon	materials_examined	Type locality. “ Philippines: Bohol: Sierra Bullones: Nan-od. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14F9ABFCDCF939.taxon	distribution	Distribution in the Philippines. Visayas (Bohol) (Krikken & Huijbregts 2011). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14F893FB57F861.taxon	materials_examined	Type locality. “ Java. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14F893FB57F861.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Boucomont 1924 a). Overall distribution. The Philippines and Indonesia (Lansberge 1883; Boucomont 1924 a).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ Banaue, Ifugao, Luzon, the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Ifugao) (Ochi & Kon 2006). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FEF3FCDCFDE5.taxon	materials_examined	Type locality. “ Philippines: Los Banos. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FEF3FCDCFDE5.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Boucomont 1914, 1924 a; Krikken & Huijbregts 2011). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FDF7FCDCFCCD.taxon	materials_examined	Type locality. “ Philippines, Luzon Is. nr. Lucena, Kinabuhayan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FDF7FCDCFCCD.taxon	distribution	Distribution in the Philippines. Luzon (Quezon) (Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FCDFFCDCFC35.taxon	materials_examined	Type locality. “ Philippines, Negros Is., Manbucal. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FCDFFCDCFC35.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Ochi et al. 2024). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FB87FCDCFB1D.taxon	materials_examined	Type locality. “ Mt. Apo, Mindanao Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FB87FCDCFB1D.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi & Kon 2004). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FB6FFCDCFA45.taxon	materials_examined	Type locality. “ Banaue, Ifugao, Luzon, the Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FB6FFCDCFA45.taxon	distribution	Distribution in the Philippines. Luzon (Ifugao) (Ochi & Kon 2006). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FA57FC25F965.taxon	materials_examined	Type locality. “ Hongkong, Bengal. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FA57FC25F965.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Boucomont 1924 a; Balthasar 1963 b). Overall distribution. The Philippines, Cambodia, China (including Taiwan), India, Indonesia, Malaysia, Myanmar, Nepal, Thailand, and Vietnam (Balthasar 1963 b).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14F977FB09F861.taxon	materials_examined	Type locality. “ Palawan, Binaluan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14F977FB09F861.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Balthasar 1959). Overall distribution. Endemic species to the Philippines (Huijbregts & Krikken 2011).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ Palawan Island. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ochi & Kon 2025 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FEF0FCDCFD86.taxon	materials_examined	Type locality. “ Philippines: Mont. Banahao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FEF0FCDCFD86.taxon	distribution	Distribution in the Philippines. This species is endemic to the archipelago and is found in Luzon (Laguna and Sibuyan) and Mindanao (Lanao del Norte) (Boucomont 1920, 1924 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FD11FCDCFCEF.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Quirino. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FD11FCDCFCEF.taxon	distribution	Distribution in the Philippines. Luzon (Quirino) (Ochi et al. 2023 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FCFEFCDCFBD4.taxon	materials_examined	Type locality. “ Philippines: Batan Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FCFEFCDCFBD4.taxon	distribution	Distribution in the Philippines. Luzon (Batan) (Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FBA6FCDCFAD1.taxon	materials_examined	Type locality. “ Palawan, Bacuit. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FBA6FCDCFAD1.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Boucomont 1924 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FAABFCDCF9DA.taxon	materials_examined	Type locality. “ Philippines: Mont Banahao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FAABFCDCF9DA.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao) (Boucomont 1920; 1924 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14F9ACFCDCF8FB.taxon	materials_examined	Type locality. “ Mindanao: Momungan, Pt. Bango. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14F9ACFCDCF8FB.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur) and Mindanao (Lanao del Norte) (Boucomont 1925). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14F8CDFCDCF823.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., South Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14F8CDFCDCF823.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Ochi & Kon 2025 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FF0BFB9BFE74.taxon	materials_examined	Type locality. “ Insel Luçon (Manilla). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FF0BFB9BFE74.taxon	distribution	Distribution in the Philippines. Luzon (Manila and Mindoro) (Eschscholtz 1822; Boucomont 1914; 1924 a). Overall distribution. The Philippines and China (Taiwan) (Krajčík 2006).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FD4BFC27FC58.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., Davao del Norte, Masara-Mainit. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FD4BFC27FC58.taxon	distribution	Distribution in the Philippines. Mindanao (Davao del Norte) (Ochi et al. 2024). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FE47FC27FD70.taxon	materials_examined	Type locality. “ Mindanao: Momungan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FE47FC27FD70.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Norte) (Boucomont 1925). Overall distribution. Endemic subspecies to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FC33FCDCFB80.taxon	materials_examined	Type locality. “ Mindanao, Lanao, Kolobugano. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FC33FCDCFB80.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Norte) (Frey 1972 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FB1BFCDCFAA0.taxon	materials_examined	Type locality. “ Philippines, Los Baños. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FB1BFCDCFAA0.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Boucomont 1914, 1924 a; Krikken & Huijbregts 2011). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FA3BFCDCF988.taxon	materials_examined	Type locality. “ Philippines: Luzon: Los Baños: Laguna. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FA3BFCDCF988.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Krikken 1977 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14F9E3FCDCF84C.taxon	materials_examined	Type locality. “ Iles Philippines: Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14F9E3FCDCF84C.taxon	distribution	Distribution in the Philippines. Mindanao (Bukidnon, Lanao del Norte, and Zamboanga del Norte) (Boucomont 1921, 1924 a; Krikken & Huijbregts 2011). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FEBBFCDCFE16.taxon	materials_examined	Type locality. “ Philippines, Basilan Is., Mindanao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FEBBFCDCFE16.taxon	distribution	Distribution in the Philippines. Mindanao (Basilan and Bukidnon) (Ochi et al. 2024; Ochi & Kon 2025 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FE63FCDCFD37.taxon	materials_examined	Type locality. “ Philippines (Samar). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FE63FCDCFD37.taxon	distribution	Distribution in the Philippines. Visayas (Leyte and Samar) and Mindanao (Surigao) (Krikken 1986; Ochi & Araya 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FC80FCDCFC1F.taxon	materials_examined	Type locality. “ Mt. Apo, Mindanao Is., Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FC80FCDCFC1F.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi & Kon 1999). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FC69FCDCFB44.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., Mt. Apo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FC69FCDCFB44.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi & Araya 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FB56FCDCFAAD.taxon	materials_examined	Type locality. “ Philippines, Leyte. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FB56FCDCFAAD.taxon	distribution	Distribution in the Philippines. Visayas (Leyte) (Ochi & Araya 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FA3FFCDCF992.taxon	materials_examined	Type locality. “ Philippines, Negros Isld., Mambucal. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FA3FFCDCF992.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Ochi & Araya 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14F9E4FCDCF8FB.taxon	materials_examined	Type locality. “ Philippines, Batan Is. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14F9E4FCDCF8FB.taxon	distribution	Distribution in the Philippines. Luzon (Batan) (Ochi et al. 2021). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14F8CCFCDCF823.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Quezon National Park. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14F8CCFCDCF823.taxon	distribution	Distribution in the Philippines. Luzon (Quezon) and Visayas (Negros) (Ochi & Araya 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FF0AFCDCFE99.taxon	materials_examined	Type locality. “ Philippines, Luzon Isld., Quezon National Park. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FF0AFCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Quezon) (Ochi & Araya 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FEF3FE7BFD36.taxon	materials_examined	Type locality. “ Siam. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FEF3FE7BFD36.taxon	distribution	Distribution in the Philippines. Luzon (Laguna and Palawan) and Mindanao (Davao, Lanao del Norte, Surigao, and Zamboanga del Norte) (Boucomont 1914, 1924 a, 1925). Overall distribution. The Philippines, China, India, Indonesia, Malaysia, Myanmar, Thailand, and Vietnam (Ueda et al. 2022).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FC80FCDCFC1F.taxon	materials_examined	Type locality. “ Philippines, Mindanao Isld., Mt. Apo. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FC80FCDCFC1F.taxon	distribution	Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi & Araya 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FC69FCDCFB18.taxon	materials_examined	Type locality. “ Philippines (Quezon). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FC69FCDCFB18.taxon	distribution	Distribution in the Philippines. Luzon (Quezon and Camarines Sur) and Visayas (Negros) (Ochi & Araya 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FB72FCDCFA41.taxon	materials_examined	Type locality. “ Philippines, Luzon Is., Mt. Banahao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FB72FCDCFA41.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao and Camarines Sur) (Ochi & Araya 1992). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FA5BFCDCF96E.taxon	materials_examined	Type locality. “ Philippines, Mindanao Is., Surigao. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FA5BFCDCF96E.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao and Bukidnon) (Ochi & Araya 1992; Ochi & Kon 2025 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14F8B0FB31F823.taxon	materials_examined	Type locality. “ Luzon, Bicol National Park. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14F8B0FB31F823.taxon	distribution	Distribution in the Philippines. Luzon (Palawan and Bicol) (Ochi & Araya 1996). Overall distribution. The Philippines and Malaysia (Krikken & Huijbregts 2009).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69239283EFF14FF0AFE8DFE74.taxon	materials_examined	Type locality. “ Java. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69239283EFF14FF0AFE8DFE74.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Boucomont 1914, 1924 a; Balthasar 1963 b). Overall distribution. The Philippines, Indonesia, China (Taiwan), and Thailand (Krikken & Huijbregts 2009).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69239283EFF14FE46FAB9FD28.taxon	materials_examined	Type locality. “ Java et Sumatra. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69239283EFF14FE46FAB9FD28.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Boucomont 1914, 1924 a; Balthasar 1963 b; Ochi & Kon 1994, 2008). Overall distribution. The Philippines, Indonesia, Malaysia, and Thailand (Ochi & Kon 2008).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69239283EFF14FC82FB3BFB80.taxon	materials_examined	Type locality. “ Borneo, Sarawak. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69239283EFF14FC82FB3BFB80.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Boucomont 1914, 1924 a; Balthasar 1963 b; Ochi et al. 2008; Krikken & Huijbregts 2009). Overall distribution. The Philippines, Indonesia, and Malaysia (Ueda et al. 2022).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69239283EFF14FB52FE8DF9AC.taxon	materials_examined	Type locality. “ Sarawak. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69239283EFF14FB52FE8DF9AC.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Ochi & Araya 1996; Ochi et al. 2008; Huijbregts & Krikken 2009). Overall distribution. The Philippines, India, Bangladesh, Myanmar, Indonesia, Cambodia, Malaysia, Thailand, Nepal, and China (including Taiwan) (Huijbregts & Krikken 2009; Han et al. 2021; Ueda et al. 2022).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692392831FF14F8E7FE20FEC4.taxon	materials_examined	Type locality. “ Singapore. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692392831FF14F8E7FE20FEC4.taxon	distribution	Distribution in the Philippines. Mindanao (Balthasar 1963 a; Ochi & Kon 1994). Overall distribution. The Philippines, Indonesia, Malaysia, Singapore, and Thailand (Balthasar 1963 a; Ochi & Kon 1994).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FDF7FCDCFCCC.taxon	materials_examined	Type locality. “ Philippinen, Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FDF7FCDCFCCC.taxon	distribution	Distribution in the Philippines. Luzon (Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FCDFFCDCFC34.taxon	materials_examined	Type locality. “ Palawan (P. Princesa) ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FCDFFCDCFC34.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Moser 1915 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FB87FCDCFB1C.taxon	materials_examined	Type locality. “ Mindanao (Davao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FB87FCDCFB1C.taxon	distribution	Distribution in the Philippines. Mindanao (Davao) (Moser 1921 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FB6FFCDCFA44.taxon	materials_examined	Type locality. “ Philippinen, Lianga. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FB6FFCDCFA44.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao del Sur) (Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FA57FCDCF9AC.taxon	materials_examined	Type locality. “ Luzon (Montalban). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FA57FCDCF9AC.taxon	distribution	Distribution in the Philippines. Luzon (Rizal) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14F93FFCDCF894.taxon	materials_examined	Type locality. “ Mindanao (Iligan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14F93FFCDCF894.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Norte) (Moser 1917 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362830FF14F8E7FCDCFF0C.taxon	materials_examined	Type locality. “ Luzon (Mt. Isarog). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362830FF14F8E7FCDCFF0C.taxon	distribution	Distribution in the Philippines. Luzon (Camarines Sur) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FE9FFCDCFE74.taxon	materials_examined	Type locality. “ Luzon (Kasiguran). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FE9FFCDCFE74.taxon	distribution	Distribution in the Philippines. Luzon (Aurora) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FE47FCDCFD70.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FE47FCDCFD70.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Brenske 1894 a, 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FD4BFCDCFC7C.taxon	materials_examined	Type locality. “ Philippines. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FD4BFCDCFC7C.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Blanchard 1851; Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FC4FFCDCFB78.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FC4FFCDCFB78.taxon	distribution	Distribution in the Philippines. Luzon (Benguet) (Brenske 1894 a, 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FB53FCDCFAA0.taxon	materials_examined	Type locality. “ Philippinen (Siargao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FB53FCDCFAA0.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao del Norte) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FA3BFCDCF988.taxon	materials_examined	Type locality. “ Mindanao (Siargao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FA3BFCDCF988.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14F9E3FCDCF894.taxon	materials_examined	Type locality. “ South Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14F9E3FCDCF894.taxon	distribution	Distribution in the Philippines. Luzon (Manila and Palawan) and Mindanao (Zamboanga del Norte) (Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372833FF14F8E7FCDCFF0C.taxon	materials_examined	Type locality. “ Mindoro (Subaan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372833FF14F8E7FCDCFF0C.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FED7FCDCFE2C.taxon	materials_examined	Type locality. “ Luzon: Mt. Data. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FED7FCDCFE2C.taxon	distribution	Distribution in the Philippines. Luzon (Mountain Province) (Moser 1926). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FDBFFCDCFD14.taxon	materials_examined	Type locality. “ Baguio. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FDBFFCDCFD14.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Moser 1926). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FCDFFCDCFC34.taxon	materials_examined	Type locality. “ Philippinen, Mindanao, Bitaibitai. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FCDFFCDCFC34.taxon	distribution	Distribution in the Philippines. Mindanao (Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FB86FCDCFB1C.taxon	materials_examined	Type locality. “ Masbate (Arorog). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FB86FCDCFB1C.taxon	distribution	Distribution in the Philippines. Luzon (Masbate) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FB6EFCDCFA44.taxon	materials_examined	Type locality. “ Balabac. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FB6EFCDCFA44.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FA56FCDCF9AC.taxon	materials_examined	Type locality. “ Luzon (Balbalan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FA56FCDCF9AC.taxon	distribution	Distribution in the Philippines. Luzon (Kalinga) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14F93EFCDCF894.taxon	materials_examined	Type locality. “ Palawan (Binaluan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14F93EFCDCF894.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342832FF14F8E6FCDCFF0C.taxon	materials_examined	Type locality. “ Mindoro (Calapan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342832FF14F8E6FCDCFF0C.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FE9EFCDCFE74.taxon	materials_examined	Type locality. “ Mindanao (Dansalan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FE9EFCDCFE74.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FE46FCDCFD5C.taxon	materials_examined	Type locality. “ Palawan (Binaluan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FE46FCDCFD5C.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FD2EFCDCFC58.taxon	materials_examined	Type locality. “ Die Insel Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FD2EFCDCFC58.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Erichson 1834; Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FC32FCDCFB80.taxon	materials_examined	Type locality. “ Philippinen, Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FC32FCDCFB80.taxon	distribution	Distribution in the Philippines. Luzon (Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FB1AFCDCFAE8.taxon	materials_examined	Type locality. “ Mindanao (Kolambugan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FB1AFCDCFAE8.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Norte) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FAC2FCDCF9D0.taxon	materials_examined	Type locality. “ Philippinen, Mindanao, Lianga. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FAC2FCDCF9D0.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao del Sur) (Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14F9AAFCDCF938.taxon	materials_examined	Type locality. “ Philippinen, Mindoro. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14F9AAFCDCF938.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Brenske 1899; Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14F892FCDCF860.taxon	materials_examined	Type locality. “ Philippinen, Negros Isl. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14F892FCDCF860.taxon	distribution	Distribution in the Philippines. Visayas (Negros) (Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FF0BFCDCFEBD.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FF0BFCDCFEBD.taxon	distribution	Distribution in the Philippines. Philippine archipelago (Brenske 1894 a, 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FE0FFCDCFDE5.taxon	materials_examined	Type locality. “ South Palawan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FE0FFCDCFDE5.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FDF7FCDCFCCD.taxon	materials_examined	Type locality. “ N. W. Panay. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FDF7FCDCFCCD.taxon	distribution	Distribution in the Philippines. Visayas (Panay Island) (Moser 1924 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FCDFFCDCFC35.taxon	materials_examined	Type locality. “ Philippinen (Paete). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FCDFFCDCFC35.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Moser 1915 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FB87FCDCFB1D.taxon	materials_examined	Type locality. “ Philippinen, Samar Isl. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FB87FCDCFB1D.taxon	distribution	Distribution in the Philippines. Visayas (Samar) (Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FB6FFE13F9D1.taxon	materials_examined	Type locality. “ Philippinen. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FB6FFE13F9D1.taxon	distribution	Distribution in the Philippines. Luzon (Brenske 1894 a, 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FB6FFE13F9D1.taxon	discussion	Remarks. Brenske observed morphological variations and gave a detailed description of this species (see Brenske 1899).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14F9ABFCDCF939.taxon	materials_examined	Type locality. “ Philippinen: Ins. Sibuyan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14F9ABFCDCF939.taxon	distribution	Distribution in the Philippines. Luzon (Sibuyan) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14F893FCDCF861.taxon	materials_examined	Type locality. “ Mindoro (San Theodoro, Subaan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14F893FCDCF861.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ Masbate (Arorog).	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Luzon (Masbate) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FB87FCDCFB1D.taxon	materials_examined	Type locality. “ Luzon (Mt. Banahao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FB87FCDCFB1D.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FB6FFCDCFA45.taxon	materials_examined	Type locality. “ Balabac. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FB6FFCDCFA45.taxon	distribution	Distribution in the Philippines. Luzon (Palawan) (Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FA57FCDCF9AD.taxon	materials_examined	Type locality. “ Mindanao (Butuan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FA57FCDCF9AD.taxon	distribution	Distribution in the Philippines. Mindanao (Agusan del Norte) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FE63FE0DFC7D.taxon	materials_examined	Type locality. “ Mindanao (Davao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FE63FE0DFC7D.taxon	distribution	Distribution in the Philippines. Mindanao (Davao) (Moser 1917 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FE63FE0DFC7D.taxon	discussion	Remarks. This species is currently placed in the genus Neoserica and subgenus Autoserica. However, this placement is highly doubtful, as most species described under Autoserica [auctorum] by Brenske and Moser are now recognized as Maladera Mulsant, 1871 (Pope, 1960; Ahrens & Fabrizi, 2016). The taxonomic status of all Philippine Sericinae requires a major revision, since most species are known only from their original descriptions. While several taxonomic reappraisals of numerous genera and subgenera have been made in other Oriental and Palearctic regions (e. g., by Ahrens and colleagues), the Philippine fauna remains largely unrevised.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14F93FFCDCF895.taxon	materials_examined	Type locality. “ Philippinen: Bucas Grande (Socorro). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14F93FFCDCF895.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao del Norte) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332837FF14F8E7FCDCFF0C.taxon	materials_examined	Type locality. “ Luzon (Mt. Makiling. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332837FF14F8E7FCDCFF0C.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Moser 1921 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FE9EFCDCFE74.taxon	materials_examined	Type locality. “ Luzon (Bagnio). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FE9EFCDCFE74.taxon	distribution	Distribution in the Philippines. Luzon (Baguio) (Moser 1916 a). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FE46FCDCFD70.taxon	materials_examined	Type locality. “ Philippinen, Mindanao, Dapitan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FE46FCDCFD70.taxon	distribution	Distribution in the Philippines. Luzon (Catanduanes) and Mindanao (Zamboanga del Norte) (Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FD4AFCDCFC58.taxon	materials_examined	Type locality. “ Luzon (Mt. Banahao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FD4AFCDCFC58.taxon	distribution	Distribution in the Philippines. Luzon (Mt. Banahao) (Moser 1915 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FC32FCDCFB80.taxon	materials_examined	Type locality. “ Luzon (Mt. Makiling). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FC32FCDCFB80.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Moser 1915 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FB1AFCDCFAE8.taxon	materials_examined	Type locality. “ Masbate (Arorog). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FB1AFCDCFAE8.taxon	distribution	Distribution in the Philippines. Luzon (Masbate) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FAC2FCDCF9D0.taxon	materials_examined	Type locality. “ Sibuyan. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FAC2FCDCF9D0.taxon	distribution	Distribution in the Philippines. Luzon (Sibuyan) (Moser 1924 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14F9AAFCDCF938.taxon	materials_examined	Type locality. “ Philippinen (Los Banos). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14F9AAFCDCF938.taxon	distribution	Distribution in the Philippines. Luzon (Laguna) (Moser 1913). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14F892FCDCF860.taxon	materials_examined	Type locality. “ Mindanao (Dapitan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14F892FCDCF860.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Moser 1915 c). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FF0BFCDCFE99.taxon	materials_examined	Type locality. “ Mindanao (Surigao). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FF0BFCDCFE99.taxon	distribution	Distribution in the Philippines. Mindanao (Surigao) (Moser 1917 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FEF3FCDCFDC1.taxon	materials_examined	Type locality. “ North Luzon. ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FEF3FCDCFDC1.taxon	distribution	Distribution in the Philippines. Luzon (Brenske 1899). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FD13FCDCFCE1.taxon	materials_examined	Type locality. “ Mindanao (Zamboanga). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FD13FCDCFCE1.taxon	distribution	Distribution in the Philippines. Mindanao (Zamboanga) (Moser 1917 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FC33FCDCFB81.taxon	materials_examined	Type locality. “ Mindoro (Subaan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FC33FCDCFB81.taxon	distribution	Distribution in the Philippines. Luzon (Mindoro) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FB1BFCDCFAE9.taxon	materials_examined	Type locality. “ Mindanao (Surigao, Iligan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FB1BFCDCFAE9.taxon	distribution	Distribution in the Philippines. Mindanao (Lanao del Norte and Surigao) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FAC3FCDCF9D1.taxon	materials_examined	Type locality. “ Mindanao (Butuan). ”	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FAC3FCDCF9D1.taxon	distribution	Distribution in the Philippines. Mindanao (Agusan del Norte) (Moser 1922 b). Overall distribution. Endemic species to the Philippines.	en	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
