identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
3C6E87D6924B284CFF14FE2AFCDCFDB8.text	3C6E87D6924B284CFF14FE2AFCDCFDB8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gilletianus asinensis Ochi, Kon & Kakizoe 2016	<div><p>Gilletianus asinensis Ochi, Kon &amp; Kakizoe, 2016</p><p>Gilletianus asinensis Ochi, Kon &amp; Kakizoe, 2016: 307 .</p><p>Type locality. “ Philippines, nr Baguio, Asin Hot-Spring.”</p><p>Distribution in the Philippines. Luzon (Baguio) (Ochi et al. 2016).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6924B284CFF14FE2AFCDCFDB8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FD12FCDCFCE0.text	3C6E87D6924B284CFF14FD12FCDCFCE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gilletianus cavitensis Ochi, Kon & Kakizoe 2016	<div><p>Gilletianus cavitensis Ochi, Kon &amp; Kakizoe, 2016</p><p>Gilletianus cavitensis Ochi, Kon &amp; Kakizoe, 2016: 314 .</p><p>Type locality. “ Philippines, Luzon, nr Manila, Cavite.”</p><p>Distribution in the Philippines. Luzon (Baguio) (Ochi et al. 2016).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6924B284CFF14FD12FCDCFCE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FCFAFC27FBC8.text	3C6E87D6924B284CFF14FCFAFC27FBC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gilletianus kinabuhayan subsp. kinabuhayan Ochi, Kon & Kakizoe 2016	<div><p>Gilletianus kinabuhayan kinabuhayan Ochi, Kon &amp; Kakizoe, 2016</p><p>Gilletianus kinabuhayan kinabuhayan Ochi, Kon &amp; Kakizoe, 2016: 310 .</p><p>Type locality. “ Philippines, Luzon, nr Manila, Cavite.”</p><p>Distribution in the Philippines. Luzon (Cavite) (Ochi et al. 2016).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6924B284CFF14FCFAFC27FBC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FBA2FC27FB30.text	3C6E87D6924B284CFF14FBA2FC27FB30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gilletianus kinabuhayan subsp. lucenaensis Ochi, Kon & Kakizoe 2016	<div><p>Gilletianus kinabuhayan lucenaensis Ochi, Kon &amp; Kakizoe, 2016</p><p>Gilletianus kinabuhayan lucenaensis Ochi, Kon &amp; Kakizoe, 2016: 312 .</p><p>Type locality. “ Philippines, Luzon, nr Manila, Cavite.”</p><p>Distribution in the Philippines. Luzon (Cavite) (Ochi et al. 2016).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6924B284CFF14FBA2FC27FB30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FA8AFCDCFA18.text	3C6E87D6924B284CFF14FA8AFCDCFA18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gilletianus luzonensis Ochi, Kon & Kakizoe 2016	<div><p>Gilletianus luzonensis Ochi, Kon &amp; Kakizoe, 2016</p><p>Gilletianus luzonensis Ochi, Kon &amp; Kakizoe, 2016: 308 .</p><p>Type locality. “ Philippines, C. Luzon, Kinabuhayan.”</p><p>Distribution in the Philippines. Luzon (Quezon Province) (Ochi et al. 2016).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6924B284CFF14FA8AFCDCFA18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14FA72FCDCF940.text	3C6E87D6924B284CFF14FA72FCDCF940.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gilletianus matotomensis Ochi, Kon & Kawahara 2010	<div><p>Gilletianus matotomensis Ochi, Kon &amp; Kawahara, 2010</p><p>Gilletianus matotomensis Ochi, Kon &amp; Kawahara, 2010: 7; Král et al. 2014: 123; Ochi et al. 2016: 319.</p><p>Type locality. “Mt. Matotom, Mindanao Is., the Philippines.”</p><p>Distribution in the Philippines. Mindanao (South Cotabato) (Ochi et al. 2010; Král et al. 2014).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6924B284CFF14FA72FCDCF940	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6924B284CFF14F95AFCDCF8A8.text	3C6E87D6924B284CFF14F95AFCDCF8A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gilletianus mindorensis Ochi, Kon & Kawahara 2010	<div><p>Gilletianus mindorensis Ochi, Kon &amp; Kawahara, 2010</p><p>Gilletianus mindorensis Ochi, Kon &amp; Kawahara, 2010: 9; Král et al. 2014: 123; Ochi et al. 2016: 319.</p><p>Type locality. “Mt. Halcon, Mindro [= Mindoro] Is., the Philippines.”</p><p>Distribution in the Philippines. Luzon (Ochi et al. 2010; Král et al. 2014).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6924B284CFF14F95AFCDCF8A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FF0BFCDCFE99.text	3C6E87D69248284FFF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gilletianus negrosensis Ochi, Kon & Kakizoe 2016	<div><p>Gilletianus negrosensis Ochi, Kon &amp; Kakizoe, 2016</p><p>Gilletianus negrosensis Ochi, Kon &amp; Kakizoe, 2016: 312 .</p><p>Type locality. “ Philippines, Negros Is., Manbucal.”</p><p>Distribution in the Philippines. Visayas (Ochi et al. 2016).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69248284FFF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FEF3FE8DFD5D.text	3C6E87D69248284FFF14FEF3FE8DFD5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gilletianus nigrovirgatus (Schmidt 1910)	<div><p>Gilletianus nigrovirgatus (Schmidt, 1910)</p><p>Aphodius nigrovirgatus Schmidt, 1911: 49 .</p><p>Aphodius (Trichaphodius) nigrovirgatus: Balthasar 1964: 169; Stebnicka 1986: 29.</p><p>Gilletianus nigrovirgatus: Král et al. 2014: 123.</p><p>Type locality. “ Yünnan.”</p><p>Distribution in the Philippines. Philippine archipelago (Balthasar 1964; Stebnicka 1986).</p><p>Overall distribution. The Philippines, China, Nepal, India, Indonesia, Laos, and Thailand (Král et al. 2014).</p></div>	https://treatment.plazi.org/id/3C6E87D69248284FFF14FEF3FE8DFD5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FD2FFCEBFBC9.text	3C6E87D69248284FFF14FD2FFCEBFBC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gilletianus reichei (Harold 1859)	<div><p>Gilletianus reichei (Harold, 1859)</p><p>Aphodius reichei Harold, 1859: 210; Boucomont 1914: 342.</p><p>Aphodius (Trichaphodius) reichei: Schmidt 1913: 136; Balthasar 1964: 168; Nomura 1973: 50; Stebnicka 1991: 4; Stebnicka 1992: 6; Stebnicka 1993: 32; Ochi &amp; Kon 1995: 57.</p><p>Gilletianus reichei: Dellacasa &amp; Dellacasa 2006a: 27; Ochi et al. 2016: 319.</p><p>Type locality. “ Java.”</p><p>Distribution in the Philippines. Luzon and Visayas (Leyte) (Schmidt 1913; Stebnicka 1993; Ochi et al. 2016).</p><p>Overall distribution. The Philippines, Australia, Indonesia, Malaysia, Thailand, and Vietnam (Stebnicka 1993; Dellacasa &amp; Dellacasa 2006a; Ochi et al. 2016).</p></div>	https://treatment.plazi.org/id/3C6E87D69248284FFF14FD2FFCEBFBC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FBA3FCDCFB31.text	3C6E87D69248284FFF14FBA3FCDCFB31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gilletianus takeshii Ochi, Kon & Kawahara 2010	<div><p>Gilletianus takeshii Ochi, Kon &amp; Kawahara, 2010</p><p>Gilletianus takeshii Ochi, Kon &amp; Kawahara, 2010: 6; Král et al. 2014: 124; Ochi et al. 2016: 318.</p><p>Type locality. “Kinabuhayan, Luzon, the Philippines.”</p><p>Distribution in the Philippines. Luzon (Ochi et al. 2010; Král et al. 2014).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69248284FFF14FBA3FCDCFB31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14FAC3FCCCF941.text	3C6E87D69248284FFF14FAC3FCCCF941.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Labarrus hoabinhensis Balthasar 1946	<div><p>Labarrus hoabinhensis Balthasar, 1946</p><p>Aphodius (Nialus) hoabinhensis Balthasar, 1946: 67; Stebnicka 1989: 23.</p><p>Aphodius (Nialus) insularis Petrovitz, 1961: 106 . Type locality. “ Philippinen; Ceylon.”; Balthasar 1964: 431.</p><p>Aphodius (Labarrus) hoabinhensis: Král &amp; Šípek 2013: 643.</p><p>Type locality. “Tonkin, Hoa Bin et India or., Madras.”</p><p>Distribution in the Philippines. Philippine archipelago (Petrovitz 1961a).</p><p>Overall distribution. The Philippines, India, Thailand, Sri Lanka, Vietnam, Bhutan, Nepal, and China (including Taiwan) (Stebnicka 1989; Král &amp; Šípek 2013).</p></div>	https://treatment.plazi.org/id/3C6E87D69248284FFF14FAC3FCCCF941	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69248284FFF14F893FCDCF805.text	3C6E87D69248284FFF14F893FCDCF805.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Loboparius semiglobulus (Petrovitz 1962)	<div><p>Loboparius semiglobulus (Petrovitz, 1962)</p><p>Aphodius (Loboparius) semiglobulus Petrovitz, 1962: 104; Balthasar 1964: 61; Dellacasa 1983: 264.</p><p>Type locality. “Luzon, Butac.”</p><p>Distribution in the Philippines. Luzon (Benguet and Laguna) and Visayas (Leyte) (Petrovitz 1962; Balthasar 1964; Dellacasa 1983).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69248284FFF14F893FCDCF805	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FF43FD91FE16.text	3C6E87D69249284EFF14FF43FD91FE16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pharaphodius crenatus (Harold 1862)	<div><p>Pharaphodius crenatus (Harold, 1862)</p><p>Aphodius crenatus Harold, 1862: 142; Boucomont 1914: 341.</p><p>Aphodius (Pharapodius) crenatus: Balthasar 1964: 74; Stebnicka 1986: 14.</p><p>Type locality. “Die philippinischen Inseln, Luzon.”</p><p>Distribution in the Philippines. Luzon (Harold 1862b; Boucomont 1914; Stebnicka 1986).</p><p>Overall distribution. The Philippines, China (Taiwan), Nepal, India, Indonesia, Sri Lanka, Thailand, and Vietnam (Stebnicka 1986).</p></div>	https://treatment.plazi.org/id/3C6E87D69249284EFF14FF43FD91FE16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FE60FAD7FD7E.text	3C6E87D69249284EFF14FE60FAD7FD7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pharaphodius diadema (Wiedemann 1823)	<div><p>Pharaphodius diadema (Wiedemann, 1823)</p><p>Aphodius diadema Wiedemann, 1823: 27; Stebnicka 1993: 32.</p><p>Type locality. “ Java.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993).</p><p>Overall distribution. The Philippines, Indonesia, Malaysia, and Papua New Guinea (Stebnicka 1993).</p></div>	https://treatment.plazi.org/id/3C6E87D69249284EFF14FE60FAD7FD7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FD49FEC5FBD7.text	3C6E87D69249284EFF14FD49FEC5FBD7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pharaphodius marginellus (Fabricius 1781)	<div><p>Pharaphodius marginellus (Fabricius, 1781)</p><p>Scarabaeus marginellus Fabricius, 1781: 21 .</p><p>Aphodius marginellus: Boucomont 1914: 341.</p><p>Aphodius (Pharaphodius) marginellus: Petrovitz 1961: 99; Ochi &amp; Kon 1995: 57.</p><p>Type locality. “Coromandel.”</p><p>Distribution in the Philippines. Philippine archipelago (Boucomont 1914; Ochi &amp; Kon 1995).</p><p>Overall distribution. The Philippines, Bhutan, China (Taiwan), India, Japan, Nepal, Pakistan, Indonesia, Malaysia, Myanmar, Thailand, Vietnam, and Papua New Guinea (Petrovitz 1961; Stebnicka 1992; Ochi &amp; Kon 1995).</p></div>	https://treatment.plazi.org/id/3C6E87D69249284EFF14FD49FEC5FBD7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FBA6FDE1FAAC.text	3C6E87D69249284EFF14FBA6FDE1FAAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pharaphodius priscus (Motschulsky 1858)	<div><p>Pharaphodius priscus (Motschulsky, 1858)</p><p>Aphodius priscus Motschulsky, 1858: 56 .</p><p>Aphodius (Pharaphodius) priscus: Petrovitz 1961b: 100.</p><p>Type locality. “ Burma.”</p><p>Distribution in the Philippines. Philippine archipelago (Petrovitz 1961b).</p><p>Overall distribution. The Philippines, Cambodia, Laos, Myanmar, Thailand, Vietnam, Japan, Korea, Nepal, and China (including Taiwan) (Petrovitz 1961b; Stebnicka 1992).</p><p>Remarks. This species is widely distributed in the Oriental region, and its “reported” occurrence in the Philippines is likely erroneous.</p></div>	https://treatment.plazi.org/id/3C6E87D69249284EFF14FBA6FDE1FAAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14FA74FCDCF94A.text	3C6E87D69249284EFF14FA74FCDCF94A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleuraphodius luzonensis (Petrovitz 1972)	<div><p>Pleuraphodius luzonensis (Petrovitz, 1972)</p><p>Aphodius (Pleuraphodius) luzonensis Petrovitz, 1972: 161 .</p><p>Type locality. “ Philippinen, Luzon, Buranen.”</p><p>Distribution in the Philippines. Luzon (Petrovitz 1972).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69249284EFF14FA74FCDCF94A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69249284EFF14F95DFCDCF8B3.text	3C6E87D69249284EFF14F95DFCDCF8B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleuraphodius tectoformibus (Schmidt 1909)	<div><p>Pleuraphodius tectoformibus (Schmidt, 1909)</p><p>Aphodius tectoformibus Schmidt, 1909:115; Schmidt 1913: 121.</p><p>Type locality. “ Manila.”</p><p>Distribution in the Philippines. Luzon (Manila) (Schmidt 1909).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69249284EFF14F95DFCDCF8B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692492841FF14F80AFCDCFF0C.text	3C6E87D692492841FF14F80AFCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleuraphodius viscaensis (Stebnicka 1993)	<div><p>Pleuraphodius viscaensis (Stebnicka, 1993)</p><p>Aphodius (Pleuraphodius) viscaensis Stebnicka, 1993: 31 .</p><p>Type locality. “ Leyte, Visca N Baybay.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692492841FF14F80AFCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FC4EFA92FAE8.text	3C6E87D692462841FF14FC4EFA92FAE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ataenius australasiae (Boheman 1858)	<div><p>Ataenius australasiae (Boheman, 1858)</p><p>Ataenius australasiae Boheman, 1858: 50; Stebnicka 1993: 31.</p><p>Type locality. “Nova Hollandia (Sydney).”</p><p>Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993).</p><p>Overall distribution. The Philippines, Australia, Japan, China, Indonesia, Malaysia, Myanmar, Thailand, and Vietnam (Stebnicka 1993; Stebnicka &amp; Howden 1997).</p><p>Remarks. The natural occurrence of this species in the Philippines is doubtful. Although it is commonly found and widely distributed across Australia and mainland Asia, its presence in the Philippines remains highly questionable. Notably, Stebnicka &amp; Howden (1997) in their revision of Ataenius omitted this species from the Philippines, suggesting that earlier records may have resulted from misidentifications.</p></div>	https://treatment.plazi.org/id/3C6E87D692462841FF14FC4EFA92FAE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FAC2FD0CF9AC.text	3C6E87D692462841FF14FAC2FD0CF9AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ataenius liogaster Bates 1887	<div><p>Ataenius liogaster Bates, 1887</p><p>Ataenius liogaster Bates, 1887: 94; Stebnicka &amp; Lago 2005: 60.</p><p>Ataenius orbicularis Schmidt, 1914: 697 . Type locality. “ Samoa.”; Stebnicka 1993: 31; Stebnicka &amp; Howden 1997: 743.</p><p>Type locality. “ Mexico.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993).</p><p>Overall distribution. The Philippines, Australian, Oriental, Micronesia, and Nearctic regions (Stebnicka &amp; Howden 1997; Stebnicka &amp; Lago 2005).</p></div>	https://treatment.plazi.org/id/3C6E87D692462841FF14FAC2FD0CF9AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14F93EFCF2F84C.text	3C6E87D692462841FF14F93EFCF2F84C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ataenius peregrinator Harold 1877	<div><p>Ataenius peregrinator Harold, 1877</p><p>Ataenius peregrinator Harold, 1877: 96; Schmidt 1922: 158; Balthasar 1964: 516; Stebnicka 1991: 4; Stebnicka 1993: 30; Stebnicka &amp; Howden 1997: 743; Théry &amp; Bordat 2012: 323.</p><p>Type locality. “ Celebes, Macassar; Sourabaia; Borneo, Sarawak.”</p><p>Distribution in the Philippines. Luzon (? Manila) and Visayas (Leyte) (Schmidt 1922; Stebnicka 1993).</p><p>Overall distribution. The Philippines, Australia, Pacific, Oriental, Palearctic, Nearctic, and Neotropical (Stebnicka &amp; Howden 1997; Théry &amp; Bordat 2012).</p></div>	https://treatment.plazi.org/id/3C6E87D692462841FF14F93EFCF2F84C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FED7FCDCFE2C.text	3C6E87D692462841FF14FED7FCDCFE2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sinaphodius philippinensis Cervenka 1994	<div><p>Sinaphodius philippinensis Červenka, 1994</p><p>Aphodius (Sinaphodius) philippinensis Červenka, 1994: 112 .</p><p>Type locality. “Mindanao, Momungan.”</p><p>Distribution in the Philippines. Mindanao (Lanao del Norte) (Červenka 1994).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692462841FF14FED7FCDCFE2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692462841FF14FDBFFC4DFCCC.text	3C6E87D692462841FF14FDBFFC4DFCCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sinaphodius sinuatus (Harold 1860)	<div><p>Sinaphodius sinuatus (Harold, 1860)</p><p>Aphodius sinuatus Harold, 1860: 614; Boucomont, 1914: 342; Balthasar 1964: 177.</p><p>Aphodius (Trichaphodius) sinuatus: Schmidt 1914: 136.</p><p>Aphodius (Sinaphodius) sinuatus: Červenka 1994: 114.</p><p>Type locality. “Ins. Philippinć.”</p><p>Distribution. Philippine archipelago (Harold 1860; Balthasar 1964).</p><p>Overall distribution. The Philippines and Indonesia (Červenka 1994).</p></div>	https://treatment.plazi.org/id/3C6E87D692462841FF14FDBFFC4DFCCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FF43FB83FE51.text	3C6E87D692472840FF14FF43FB83FE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Saprosites longethorax Paulian 1944	<div><p>Saprosites longethorax Paulian, 1944</p><p>Saprosites longethorax Paulian, 1944: 72; Balthasar 1964: 501; Nomura 1973: 51.</p><p>Type locality. “Luzon (Imugan).”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Paulian 1944).</p><p>Overall distribution. The Philippines and China (Taiwan) (Nomura 1973).</p></div>	https://treatment.plazi.org/id/3C6E87D692472840FF14FF43FB83FE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FD13FE0DFC59.text	3C6E87D692472840FF14FD13FE0DFC59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Saprosites pygmaeus Harold 1877	<div><p>Saprosites pygmaeus Harold, 1877</p><p>Saprosites pygmaeus Harold, 1877: 91; Cartwright &amp; Gordon 1971: 267; Stebnicka 1993: 30; Stebnicka 2012: 18.</p><p>Type locality. “Iles Key.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993).</p><p>Overall distribution. The Philippines, Indonesia, Christmas Island, Papua New Guinea, and Hawaii (Stebnicka 2012).</p></div>	https://treatment.plazi.org/id/3C6E87D692472840FF14FD13FE0DFC59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FE2BFB42FDB9.text	3C6E87D692472840FF14FE2BFB42FDB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Saprosites rufopolitus Gillet 1924	<div><p>Saprosites rufopolitus Gillet, 1924</p><p>Saprosites rufopolitus Gillet, 1924: 103; Stebnicka 2012: 18.</p><p>Type locality. “ Sumatra.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Stebnicka 2012).</p><p>Overall distribution. The Philippines, Indonesia, Malaysia, and Thailand (Stebnicka 2012).</p></div>	https://treatment.plazi.org/id/3C6E87D692472840FF14FE2BFB42FDB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FC33FB5DFB81.text	3C6E87D692472840FF14FC33FB5DFB81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Saprosites verecundus Schmidt 1909	<div><p>Saprosites verecundus Schmidt, 1909</p><p>Saprosites verecundus Schmidt, 1909: 121; Stebnicka 2012: 25.</p><p>Type locality. “Mentawei.”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Stebnicka 2012).</p><p>Overall distribution. The Philippines, Thailand, Indonesia, and Malaysia (Stebnicka 2012).</p></div>	https://treatment.plazi.org/id/3C6E87D692472840FF14FC33FB5DFB81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FB53FCDCFAA1.text	3C6E87D692472840FF14FB53FCDCFAA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Saprovisca leytensis Stebnicka 1993	<div><p>Saprovisca leytensis Stebnicka, 1993</p><p>Saprovisca leytensis Stebnicka, 1993: 29; Skelley 2015: 2.</p><p>Type locality. “ Leyte, Visca N Baybay.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692472840FF14FB53FCDCFAA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14FA3BFCDCF989.text	3C6E87D692472840FF14FA3BFCDCF989.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Saprovisca sarangay Skelley 2015	<div><p>Saprovisca sarangay Skelley, 2015</p><p>Saprovisca sarangay Skelley, 2015: 3 .</p><p>Type locality. “Mindanao, N. Mindanao Region, Bukidnon: Kalatungan Mts. ”</p><p>Distribution in the Philippines. Luzon (Camarines Sur) and Mindanao (Bukidnon) (Skelley 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692472840FF14FA3BFCDCF989	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692472840FF14F95BFCDCF8A9.text	3C6E87D692472840FF14F95BFCDCF8A9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Setylaides palawanensis Ochi, Kon & Cabras 2025	<div><p>Setylaides palawanensis Ochi, Kon &amp; Cabras, 2025</p><p>Setylaides palawanensis Ochi, Kon &amp; Cabras, 2025b: 32 .</p><p>Type locality. “ Philippines, Palawan Is., Roxas.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ochi et al. 2025b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692472840FF14F95BFCDCF8A9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FEF3FCDCFDB9.text	3C6E87D692442843FF14FEF3FCDCFDB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Odochilus (Odochilus) johnsoni Rakovic 1987	<div><p>Odochilus (Odochilus) johnsoni Rakovič, 1987</p><p>Odochilus johnsoni Rakovič, 1987: 38; Stebnicka 1993: 32.</p><p>Odochilus (Odochilus) johnsoni: Rakovič 1997: 163.</p><p>Type locality. “ Mindoro, Philippines.”</p><p>Distribution in the Philippines. Luzon (Mindoro and Palawan) and Visayas (Leyte) (Rakovič 1987; Stebnicka 1993).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692442843FF14FEF3FCDCFDB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FD13FCDCFCE1.text	3C6E87D692442843FF14FD13FCDCFCE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Odochilus (Odochilus) parajohnsoni Rakovic 1997	<div><p>Odochilus (Odochilus) parajohnsoni Rakovič, 1997</p><p>Odochilus (Odochilus) parajohnsoni Rakovič, 1997: 160 .</p><p>Type locality. “ Philippines, Panay.”</p><p>Distribution in the Philippines. Visayas (Panay) (Rakovič 1997).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692442843FF14FD13FCDCFCE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FCFBFCDCFBC9.text	3C6E87D692442843FF14FCFBFCDCFBC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Odochilus (Odochilus) parasetosus Rakovic 1997	<div><p>Odochilus (Odochilus) parasetosus Rakovič, 1997</p><p>Odochilus (Odochilus) parasetosus Rakovič, 1997: 159 .</p><p>Type locality. “Philipp., Mindanao, Siocom.”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Rakovič 1997).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692442843FF14FCFBFCDCFBC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FBA3FCDCFB31.text	3C6E87D692442843FF14FBA3FCDCFB31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Odochilus (Odochilus) parasyntheticus Rakovic 1997	<div><p>Odochilus (Odochilus) parasyntheticus Rakovič, 1997</p><p>Odochilus (Odochilus) parasyntheticus Rakovič, 1997: 158 .</p><p>Type locality. “ Philippines, SW Panay.”</p><p>Distribution in the Philippines. Visayas (Panay) (Rakovič 1997).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692442843FF14FBA3FCDCFB31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14FA8BFCDCFA3D.text	3C6E87D692442843FF14FA8BFCDCFA3D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Odochilus (Odochilus) philippinensis Rakovic 1987	<div><p>Odochilus (Odochilus) philippinensis Rakovič, 1987</p><p>Odochilus syntheticus philippinensis Rakovič, 1987: 34; Stebnicka 1993: 33.</p><p>Odochilus (Odochilus) philippinensis: Rakovič 1997: 162.</p><p>Type locality. “NE Leyte Is.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993; Rakovič 1987).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692442843FF14FA8BFCDCFA3D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442843FF14F9C7FCD4F8DD.text	3C6E87D692442843FF14F9C7FCD4F8DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Odochilus (Parodochilus) shavrini Rakovic & Anichtchenko 2021	<div><p>Odochilus (Parodochilus) shavrini Rakovič &amp; Anichtchenko, 2021</p><p>Odochilus (Parodochilus) shavrini Rakovič &amp; Anichtchenko, 2021: 447 .</p><p>Type locality. “ Philippines, Mindanao, Maragusan, Marangig Falls, Candalaga Mts. ”</p><p>Distribution in the Philippines. Mindanao (Davao de Oro) (Rakovič &amp; Anichtchenko 2021).</p><p>Overall distribution. Endemic species to the Philippines</p></div>	https://treatment.plazi.org/id/3C6E87D692442843FF14F9C7FCD4F8DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692442842FF14F85FFE8DFE51.text	3C6E87D692442842FF14F85FFE8DFE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neotrichiorhyssemus hirsutus (Clouet 1901)	<div><p>Neotrichiorhyssemus hirsutus (Clouët, 1901)</p><p>Trichiorhyssemus hirsutus Clouët, 1901: 35 .</p><p>Neotrichiorhyssemus hirsutus: Rakovič et al. 2023: 134.</p><p>Type locality. “ Sumatra, Java, Borneo et aux Celebes.”</p><p>Distribution in the Philippines. Philippine archipelago (Rakovič et al. 2023).</p><p>Overall distribution. The Philippines, China (Taiwan), Thailand, Malaysia, Indonesia, French Polynesia, Australia, Fiji, Papua New Guinea, Samoa, Solomon Islands, Seychelles, and Mauritius (Rakovič et al, 2023).</p></div>	https://treatment.plazi.org/id/3C6E87D692442842FF14F85FFE8DFE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14FE63FCDDFD71.text	3C6E87D692452842FF14FE63FCDDFD71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleurophorus banksi Pittino & Mariani 1986	<div><p>Pleurophorus banksi Pittino &amp; Mariani, 1986</p><p>Pleurophorus banksi Pittino &amp; Mariani, 1986: 117 .</p><p>Type locality. “ Davao, Philippines Islands.”</p><p>Distribution in the Philippines. Mindanao (Davao) (Pittino &amp; Mariani 1986).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692452842FF14FE63FCDDFD71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14F9C7FCDCF8DD.text	3C6E87D692452842FF14F9C7FCDCF8DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus apoensis Ochi & Kakizoe 2021	<div><p>Rhyparus apoensis Ochi &amp; Kakizoe, 2021</p><p>Rhyparus apoensis Ochi &amp; Kakizoe, 2021: 27 .</p><p>Type locality. “ Philippines, Mindanao Is. Mt. Apo.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692452842FF14F9C7FCDCF8DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14F8AFFCDCF805.text	3C6E87D692452842FF14F8AFFCDCF805.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus apomontanus Ochi & Kakizoe 2021	<div><p>Rhyparus apomontanus Ochi &amp; Kakizoe, 2021</p><p>Rhyparus apomontanus Ochi &amp; Kakizoe, 2021: 19 .</p><p>Type locality. “ Philippines, Mindanao Is. Mt. Apo.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692452842FF14F8AFFCDCF805	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14FC83FDB3FC35.text	3C6E87D692452842FF14FC83FDB3FC35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyssemodes sindicus Pittino 1984	<div><p>Rhyssemodes sindicus Pittino, 1984</p><p>Rhyssemodes sindicus Pittino, 1984: 22 .</p><p>Type locality. “ India ( Bengal).”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Pittino 1984).</p><p>Overall distribution. The Philippines, China (including Taiwan), India, Thailand, Vietnam, Iran, Nepal, and Pakistan (Pittino 1984).</p></div>	https://treatment.plazi.org/id/3C6E87D692452842FF14FC83FDB3FC35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692452842FF14FBFFFE6BFA45.text	3C6E87D692452842FF14FBFFFE6BFA45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyssemus inscitus (Walker 1858)	<div><p>Rhyssemus inscitus (Walker, 1858)</p><p>Psammodius inscitus Walker, 1858: 207 .</p><p>Rhyssemus philippineus Masumoto, 1980: 121 . Type locality. “ Cebu City, Cebu Is. The Philippines.”</p><p>Rhyssemus inscitus: Pittino 1984: 39; Stebnicka 1993: 33; Pittino et al. 2013: 217.</p><p>Type locality. “ Ceylon.”</p><p>Distribution in the Philippines. Luzon (Makati and Metro Manila), Visayas (Cebu and Leyte), and Mindanao (Zamboanga del Norte) (Masumoto 1980; Pittino 1984; Stebnicka 1993; Pittino et al. 2013).</p><p>Overall distribution. The Philippines, Palearctic, Oriental, Madagascan, Australian, and Pacific regions (Pittino et al. 2013).</p></div>	https://treatment.plazi.org/id/3C6E87D692452842FF14FBFFFE6BFA45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FF0BFCDCFE98.text	3C6E87D692422845FF14FF0BFCDCFE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus baracatanensis Ochi & Kakizoe 2021	<div><p>Rhyparus baracatanensis Ochi &amp; Kakizoe, 2021</p><p>Rhyparus baracatanensis Ochi &amp; Kakizoe, 2021: 38 .</p><p>Type locality. “ Philippines, Mindanao Is., Baracatan.”</p><p>Distribution in the Philippines. Mindanao (Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692422845FF14FF0BFCDCFE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FEF3FBF8FDE4.text	3C6E87D692422845FF14FEF3FBF8FDE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus barclayi Masumoto, Ochi & Ho 2019	<div><p>Rhyparus barclayi Masumoto, Ochi &amp; Ho, 2019</p><p>Rhyparus barclayi Masumoto, Ochi &amp; Ho, 2019: 13; Anichtchenko et al. 2021: 100; Ochi et al. 2021: 18.</p><p>Type locality. “Lanyu Is. Taiwan.”</p><p>Distribution in the Philippines. Luzon (Ifugao) and Mindanao (Bukidnon, Davao, and South Cotabato) (Anichtchenko et al. 2021; Ochi et al. 2021).</p><p>Overall distribution. The Philippines and China (Taiwan) (Ochi et al. 2021).</p></div>	https://treatment.plazi.org/id/3C6E87D692422845FF14FEF3FBF8FDE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FDF7FCDCFCCC.text	3C6E87D692422845FF14FDF7FCDCFCCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus basilanensis Ochi & Kakizoe 2021	<div><p>Rhyparus basilanensis Ochi &amp; Kakizoe, 2021</p><p>Rhyparus basilanensis Ochi &amp; Kakizoe, 2021: 29 .</p><p>Type locality. “ Philippines, Basilan Is.”</p><p>Distribution in the Philippines. Mindanao (Basilan) (Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692422845FF14FDF7FCDCFCCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FCDFFCDDFBC8.text	3C6E87D692422845FF14FCDFFCDDFBC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus flavohirtus Anichtchenko, Minkina, Vasiljeva & Medina 2021	<div><p>Rhyparus flavohirtus Anichtchenko, Minkina, Vasiljeva &amp; Medina, 2021</p><p>Rhyparus flavohirtus Anichtchenko, Minkina, Vasiljeva &amp; Medina, 2021: 100; Ochi et al. 2021: 29.</p><p>Type locality. “ Mindanao, Lake Holon. ”</p><p>Distribution in the Philippines. Mindanao (South Cotabato and Mt. Apo) (Anichtchenko et al. 2021; Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692422845FF14FCDFFCDDFBC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FBA3FAECFB30.text	3C6E87D692422845FF14FBA3FAECFB30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus gracilis Arrow 1905	<div><p>Rhyparus gracilis Arrow, 1905</p><p>Rhyparus gracilis Arrow, 1905: 538; Anichtchenko et al. 2021: 102.</p><p>Type locality. “ Louisiade Archipelago, Sud-Est Island.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon and Davao) (Anichtchenko et al. 2021).</p><p>Overall distribution. The Philippines, Papua New Guinea, and Indonesia (Anichtchenko et al. 2021).</p></div>	https://treatment.plazi.org/id/3C6E87D692422845FF14FBA3FAECFB30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14FA8BFD99F9AC.text	3C6E87D692422845FF14FA8BFD99F9AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus helophoroides Fairmaire 1893	<div><p>Rhyparus helophoroides Fairmaire, 1893</p><p>Rhyparus helophoroides Fairmaire, 1893: 145; Mencl et al. 2013: 494; Théry &amp; Bordat 2012: 335; Anichtchenko et al. 2021: 103; Ochi et al. 2021: 18.</p><p>Rhyparus orousseti Paulian, 1981: 111 . Type locality. “ Philippines, Luzon, Mountain Prov., Baguio.”</p><p>Type locality. “Borneo occ.: Sambas; Java: Simpar et Kemanglen rés. Tega.”</p><p>Distribution in the Philippines. Luzon (Baguio and Camarines Sur), Visayas (Negros), and Mindanao (Bukidnon, Davao, and Cotabato) (Paulian 1981; Théry &amp; Bordat 2012; Ochi et al. 2021).</p><p>Overall distribution. The Philippines, Korea, Australia, Indonesia, Japan, China (including Taiwan), Papua New Guinea, New Caledonia, Solomon Islands, Malaysia, New South Wales, and Queensland (Anichtchenko et al. 2021).</p></div>	https://treatment.plazi.org/id/3C6E87D692422845FF14FA8BFD99F9AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422845FF14F93FFCDCF894.text	3C6E87D692422845FF14F93FFCDCF894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus ifugaoensis Ochi & Kakizoe 2021	<div><p>Rhyparus ifugaoensis Ochi &amp; Kakizoe, 2021</p><p>Rhyparus ifugaoensis Ochi &amp; Kakizoe, 2021: 26 .</p><p>Type locality. “ Philippines, Luzon, Ifugao Prov., Dalican.”</p><p>Distribution in the Philippines. Luzon (Ifugao) (Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692422845FF14F93FFCDCF894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692422844FF14F8E7FCDCFF0C.text	3C6E87D692422844FF14F8E7FCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus luzonensis Ochi & Kakizoe 2021	<div><p>Rhyparus luzonensis Ochi &amp; Kakizoe, 2021</p><p>Rhyparus luzonensis Ochi &amp; Kakizoe, 2021: 33 .</p><p>Type locality. “ Philippines, Luzon Is., Quirino, Madera, Sierra Madre Disimungal.”</p><p>Distribution in the Philippines. Luzon (Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692422844FF14F8E7FCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FE9FFCDCFE08.text	3C6E87D692432844FF14FE9FFCDCFE08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus mindanaoensis Anichtchenko, Minkina, Vasiljeva & Medina 2021	<div><p>Rhyparus mindanaoensis Anichtchenko, Minkina, Vasiljeva &amp; Medina, 2021</p><p>Rhyparus mindanaoensis Anichtchenko et al., 2021: 104; Ochi et al. 2021: 18.</p><p>Type locality. “ Philippines, Mindanao, Maragusan, Marangig Falls, Candalaga Mts. ”</p><p>Distribution in the Philippines. Luzon (Palawan) and Mindanao (Basilan, Bukidnon, and Davao de Oro) (Anichtchenko et al. 2021; Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692432844FF14FE9FFCDCFE08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FE63FCDCFD70.text	3C6E87D692432844FF14FE63FCDCFD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus negrosensis Ochi & Kakizoe 2021	<div><p>Rhyparus negrosensis Ochi &amp; Kakizoe, 2021</p><p>Rhyparus negrosensis Ochi &amp; Kakizoe, 2021: 35 .</p><p>Type locality. “ Philippines, Negros Is., Mt. Mandalagan. ”</p><p>Distribution in the Philippines. Visayas (Negros Island) (Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692432844FF14FE63FCDCFD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FD4BFCDCFC58.text	3C6E87D692432844FF14FD4BFCDCFC58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus paraphilippinensis Ochi & Kakizoe 2021	<div><p>Rhyparus paraphilippinensis Ochi &amp; Kakizoe, 2021</p><p>Rhyparus paraphilippinensis Ochi &amp; Kakizoe, 2021: 31; Minkina et al. 2022: 91; Minkina et al. 2023: 5.</p><p>Type locality. “ Philippines, Mindanao Is., Caraga Region, Agusan del sur, Sibagat.”</p><p>Distribution in the Philippines. Mindanao (Agusan del Sur and Bukidnon) (Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692432844FF14FD4BFCDCFC58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FC33FE8AFA8C.text	3C6E87D692432844FF14FC33FE8AFA8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus peninsularis Arrow 1905	<div><p>Rhyparus peninsularis Arrow, 1905</p><p>Rhyparus peninsularis Arrow, 1905: 537; Stebnicka 1993: 33.</p><p>Type locality. “Malay Peninsula, Penang, Perak.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Stebnicka 1993).</p><p>Overall distribution. The? Philippines, Malaysia, and? China (Taiwan).</p><p>Remarks. The authors have reviewed the articles of current authors (Mencl et al. 2013; Anichtchenko et al. 2021; Ochi et al. 2021), and it appears that the occurrence of Rhyparus peninsularis in the Philippines is doubtful. Perhaps they are unaware of Stebnicka’s work, or due to the lack of substantial specimens to examine within the archipelago. Notably, Anichtchenko et al. (2021) explicitly listed only two species of Rhyparus ( R. helophoroides and R. philippinensis) occurring in the Philippines, suggesting that R. peninsularis was either excluded due to the lack of confirmed specimens or was unaware of Stebnicka’s work.</p></div>	https://treatment.plazi.org/id/3C6E87D692432844FF14FC33FE8AFA8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14FA1FFCDCF940.text	3C6E87D692432844FF14FA1FFCDCF940.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyparus philippinensis Arrow 1905	<div><p>Rhyparus philippinensis Arrow, 1905</p><p>Rhyparus philippinensis Arrow, 1905: 538; Boucomont 1914: 345; Balthasar 1964: 612; Mencl et al. 2013: 495; Anichtchenko et al. 2021: 108; Ochi et al. 2021: 37.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Cagayan Province, Camarines Sur, and Mountain Province), Visayas (Samar), and Mindanao (Davao de Oro, Lanao del Norte, and Lanao del Sur) (Arrow 1905; Anichtchenko et al. 2021; Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692432844FF14FA1FFCDCF940	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692432844FF14F893FCDCF804.text	3C6E87D692432844FF14F893FCDCF804.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Termitodiellus luzonensis (Howden 1995)	<div><p>Termitodiellus luzonensis (Howden, 1995)</p><p>Termitodius luzonensis Howden, 1995: 25 .</p><p>Termitodiellus luzonensis: Skelley 2007: 7.</p><p>Type locality. “ Philippines: Mt. Makiling, Laguna Prov. ”</p><p>Distribution in the Philippines. Luzon (Laguna) (Howden 1995; Skelley 2007).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692432844FF14F893FCDCF804	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FDF5FCDCFCEF.text	3C6E87D692402847FF14FDF5FCDCFCEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycosia bakeri Moser 1921	<div><p>Glycosia bakeri Moser, 1921</p><p>Glycosia Bakeri Moser, 1921b: 184; Sakai 1995: 6; Legrand 2020: 51.</p><p>Type locality. “Mindanao (Dapitan).”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte and Davao) (Moser 1921b; Sakai 1995; Legrand 2020).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692402847FF14FDF5FCDCFCEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FCFEFCDCFB8C.text	3C6E87D692402847FF14FCFEFCDCFB8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycosia intermedia (Miksic 1979)	<div><p>Glycosia intermedia (Mikšić, 1979)</p><p>Euglypta intermedia Mikšić, 1979: 30 .</p><p>Astraea (Euglypta) intermedia: Mikšić 1982a: 321.</p><p>Glycosia intermedia: Legrand 2020: 49.</p><p>Type locality. “ Mt. Dimatabing, Luzon, Tayabas.”</p><p>Distribution in the Philippines. Luzon (Quezon) (Mikšić 1979; Legrand 2020).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692402847FF14FCFEFCDCFB8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FB1FFCDCFAF5.text	3C6E87D692402847FF14FB1FFCDCFAF5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycosia palawanica Moser 1918	<div><p>Glycosia palawanica Moser, 1918</p><p>Glycosia palawanica Moser 1918: 177; Sakai 1995: 7.</p><p>Type locality. “ Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Moser 1918).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692402847FF14FB1FFCDCFAF5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FAC7FC27F9DA.text	3C6E87D692402847FF14FAC7FC27F9DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycosia tricolor subsp. latipennis Sakai 1995	<div><p>Glycosia tricolor latipennis Sakai, 1995</p><p>Glycosia tricolor latipennis Sakai, 1995: 10 .</p><p>Type locality. “ Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Sakai 1995).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692402847FF14FAC7FC27F9DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14F95DFCDFF823.text	3C6E87D692402847FF14F95DFCDFF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Caloglycyphana) aethiessida (Wallace 1867)	<div><p>Glycyphana (Caloglycyphana) aethiessida (Wallace, 1867)</p><p>Euryomia aethiessida Wallace, 1867: 95; 1868: 568.</p><p>Glycyphana aethiessida: Gemminger &amp; Harold 1869: 1309; Mohnike 1873: 156; Schoch 1896: 60.</p><p>Glycyphana (Caloglycyphana) aethiessida: Mikšić 1970b: 92; Pavićević 1988: 171; Legrand 2018a: 42.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Camarines Sur, Ilocos, Laguna, Mt. Banahao, Pampanga, Masbate, and Rizal) and Visayas (Negros Oriental and Panay) (Wallace 1867; Mikšić 1970b 1970c; Pavićević 1988).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692402847FF14F95DFCDFF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692402847FF14FF0BFCDDFEBD.text	3C6E87D692402847FF14FF0BFCDDFEBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Termitodiellus mindanaoensis Minkina & Kakizoe 2020	<div><p>Termitodiellus mindanaoensis Minkina &amp; Kakizoe, 2020</p><p>Termitodiellus mindanaoensis Minkina &amp; Kakizoe, 2020: 2 .</p><p>Type locality. “ Philippines, Mindanao Island, Alamada, North Cotabato.”</p><p>Distribution in the Philippines. Mindanao (Agusan del Sur, Davao de Oro, Lanao del Sur, North and South Cotabato) (Minkina &amp; Kakizoe 2020).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692402847FF14FF0BFCDDFEBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FF0BFCDCFE98.text	3C6E87D692412846FF14FF0BFCDCFE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Caloglycyphana) antoinei Legrand 2018	<div><p>Glycyphana (Caloglycyphana) antoinei Legrand, 2018</p><p>Glycyphana (Caloglycyphana) antoinei Legrand, 2018: 38 .</p><p>Type locality. “ Central Visayas, Negros Oriental, Dumaguete.”</p><p>Distribution in the Philippines. Visayas (Negros) (Legrand 2018a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692412846FF14FF0BFCDCFE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FEF3FC27FDC0.text	3C6E87D692412846FF14FEF3FC27FDC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Caloglycyphana) binotata subsp. mindoroana Legrand 2018	<div><p>Glycyphana (Caloglycyphana) binotata mindoroana Legrand, 2018</p><p>Glycyphana (Caloglycyphana) binotata mindoroana Legrand, 2018: 37 .</p><p>Type locality. “ Mindoro, Mt. Halcon. ”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Legrand 2018a).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692412846FF14FEF3FC27FDC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FDDBFC27FD28.text	3C6E87D692412846FF14FDDBFC27FD28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Caloglycyphana) binotata subsp. viossati Antoine 1996	<div><p>Glycyphana (Caloglycyphana) binotata viossati Antoine, 1996</p><p>Glycyphana (Caloglycyphana) binotata viossati Antoine, 1996: 63; Legrand 2018a: 42.</p><p>Type locality. “ Palawan, Brook’s Point.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Antoine 1996; Legrand 2018a).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692412846FF14FDDBFC27FD28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FC83FCDCFC34.text	3C6E87D692412846FF14FC83FCDCFC34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Caloglycyphana) georgijevici Miksic 1967	<div><p>Glycyphana (Caloglycyphana) georgijevici Mikšić, 1967</p><p>Glycyphana pexata georgijevici Mikšić, 1967: 26 .</p><p>Glycyphana (Caloglycyphana) georgijevici: Mikšić 1970b: 68; Mikšić 1970c: 142; Legrand 2018a: 42.</p><p>Type locality. “Masbate-Aroroy.”</p><p>Distribution in the Philippines. Luzon (Masbate) (Mikšić 1967).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692412846FF14FC83FCDCFC34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FB87FCDCFAD4.text	3C6E87D692412846FF14FB87FCDCFAD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Caloglycyphana) immaculata Moser 1925	<div><p>Glycyphana (Caloglycyphana) immaculata Moser, 1925</p><p>Glycyphana aethiessida immaculata Moser, 1925: 69; Mikšić 1970b: 143.</p><p>Glycyphana (Caloglycyphana) immaculata: Legrand 2018a: 36.</p><p>Type locality. “ Panay (Northwestern part).”</p><p>Distribution in the Philippines. Luzon (Romblon) and Visayas (Negros and Panay) (Moser 1925; Mikšić 1970b; Legrand 2018a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692412846FF14FB87FCDCFAD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692412846FF14FAA7FD53F84C.text	3C6E87D692412846FF14FAA7FD53F84C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Caloglycyphana) pulcherrima Mohnike 1873	<div><p>Glycyphana (Caloglycyphana) pulcherrima Mohnike, 1873</p><p>Glycyphana pulcherrima Mohnike, 1873: 157; Schoch 1896: 62.</p><p>Glycyphana (Reineria) pulcherrima Mohnike: Mikšić 1970b: 142; Mikšić 1970c: 22.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Luzon (Camarines Sur and Nueva Vizcaya) and Mindanao (Mohnike 1873; Mikšić 1970b, 1970c).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks.When Mikšić was studying what is now recognized as the Reineria group,several rounds of revision occurred before it was formally erected to genus rank (Arnaud 2004). In his early work, Reineria Mikšić (1968) was treated as a subgenus of Glycyphana Burmeister (1842), which was synonymized with the genus Astraea Mohnike, 1873 . Many species were subsequently reclassified under the genus Reineria . However, for this particular species, the authors were unable to locate the original description supporting its current taxonomic placement. Legrand (2018a) did not include this species in his catalog of the sub genus Caloglycyphana Mikšić (1968) . This species will probably be revised and formally placed in the genus Reineria Mikšić (1968) . Furthermore, Mikšić noted that, based on his examination of the available material and the type specimen from the original description, this species exhibits slight morphological differences that may indicate subspeciation.</p></div>	https://treatment.plazi.org/id/3C6E87D692412846FF14FAA7FD53F84C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FF0BFC27FEBD.text	3C6E87D6925E2859FF14FF0BFC27FEBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Caloglycyphana) rubromarginata subsp. pexata Janson 1881	<div><p>Glycyphana (Caloglycyphana) rubromarginata pexata Janson, 1881</p><p>Glycyphana pexata Janson, 1881: 606 .</p><p>Glycyphana (Caloglycyphana) rubromarginata pexata: Legrand 2018a: 42.</p><p>Type locality. “ Leyte, Philippines Islands.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Janson 1881).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925E2859FF14FF0BFC27FEBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FE0FFC27FD15.text	3C6E87D6925E2859FF14FE0FFC27FD15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Caloglycyphana) rubromarginata subsp. rubromarginata Mohnike 1873	<div><p>Glycyphana (Caloglycyphana) rubromarginata rubromarginata Mohnike, 1873</p><p>Glycyphana rubromarginata Mohnike, 1873: 154 .</p><p>Glycyphana (Caloglycyphana) pexata mindanaoana Mikšić, 1984: 312 . Type locality. “Mindanao, Mt. Apo, Davao.”; Pavićević 1988: 171.</p><p>Glycyphana bipustulata Kraatz, 1899 (nomem dubium).</p><p>Type locality. “Mindanao parte occidentali.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo and Surigao) (Mohnike 1873; Mikšić 1970b, 1970c, 1984).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925E2859FF14FE0FFC27FD15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FCDFFD59FC35.text	3C6E87D6925E2859FF14FCDFFD59FC35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Euglycyphana) boudanti Arnaud 2019	<div><p>Glycyphana (Euglycyphana) boudanti Arnaud, 2019</p><p>Glycyphana (Euglycyphana) boudanti Arnaud 2019: 5 .</p><p>Type locality. “ Philippines, Oriental Negros, Mt. Canlaon. ”</p><p>Distribution in the Philippines. Visayas (Negros and Panay) (Arnaud, 2019).</p><p>Overall distribution. Only in the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925E2859FF14FCDFFD59FC35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FB87FC27FAD5.text	3C6E87D6925E2859FF14FB87FC27FAD5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Euglycyphana) luzonica subsp. banghaasi Bourgoin 1930	<div><p>Glycyphana (Euglycyphana) luzonica banghaasi Bourgoin, 1930</p><p>Glycyphana Bang-Haasi Bourgoin, 1930: 205 .</p><p>Glycyphana (Euglycyphana) luzonica banghaasi: Mikšić 1970b: 39.</p><p>Type locality. “ Philippines Luzon: Imugan et Banahao.”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao and Nueva Vizcaya) (Bourgoin 1930; Mikšić 1970b, 1970c).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925E2859FF14FB87FC27FAD5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14FAA7FC27F9F5.text	3C6E87D6925E2859FF14FAA7FC27F9F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Euglycyphana) luzonica subsp. luzonica Moser 1917	<div><p>Glycyphana (Euglycyphana) luzonica luzonica Moser, 1917</p><p>Glycyphana luzonica Moser, 1917a: 19 .</p><p>Glycyphana (Euglycyphana) luzonica: Mikšić 1970b: 37; Mikšić 1984: 311.</p><p>Type locality. “ Luzon (Mt. Isarog).”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao, Romblon, and Camarines Sur) (Moser 1917a; Mikšić 1970b, 1970c, 1984).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925E2859FF14FAA7FC27F9F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2859FF14F9C7FCDCF895.text	3C6E87D6925E2859FF14F9C7FCDCF895.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Euglycyphana) rubroplagiata Moser 1922	<div><p>Glycyphana (Euglycyphana) rubroplagiata Moser, 1922</p><p>Glycyphana rubroplagiata Moser, 1922a: 113 .</p><p>Glycyphana (Euglycyphana) rubroplagiata: Mikšić 1970b: 142; Mikšić 1970c: 42; Arnaud, 2004: 5.</p><p>Type locality. “Mindanao (Momungan).”</p><p>Distribution in the Philippines. Mindanao (Lanao del Norte and Surigao) (Moser 1922a; Mikšić 1970b, 1970c; Arnaud 2004).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925E2859FF14F9C7FCDCF895	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925E2858FF14F8E7FCDCFE50.text	3C6E87D6925E2858FF14F8E7FCDCFE50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Euglycyphana) vernalis (Wallace 1867)	<div><p>Glycyphana (Euglycyphana) vernalis (Wallace, 1867)</p><p>Euryomia vernalis Wallace, 1867: 96; Wallace 1868: 572.</p><p>Glycyphana vernalis: Gemminger &amp; Harold 1869: 1311; Schoch 1896: 63; Mohnike 1873: 164.</p><p>Glycyphana robusta Mohnike, 1873: 164 . Type locality. “ Samar, Leyte, Mindanao.”; Schoch 1896: 62.</p><p>Glycyphana (Euglycyphana) vernalis: Mikšić 1970b: 142; Mikšić 1970c: 35.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Albay, Camarines Sur, Laguna, Masbate, Romblon, and Sibuyan, and Nueva Vizcaya), Visayas (Leyte, Panay, and Samar), and Mindanao (Davao, Lanao del Norte, and Surigao) (Wallace 1867; Mohnike 1873; Mikšić 1970b, 1970c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925E2858FF14F8E7FCDCFE50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14FE63FC27FD70.text	3C6E87D6925F2858FF14FE63FC27FD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Glycyphaniola) cuculus subsp. basimaculata Miksic 1968	<div><p>Glycyphana (Glycyphaniola) cuculus basimaculata Mikšić, 1968</p><p>Glycyphana (Glycyphaniola) cuculus basimaculata Mikšić, 1968: 306; Mikšić 1971b: 56.</p><p>Type locality. “Mindero (=Mindoro).”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Mikšić 1968, 1971b).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925F2858FF14FE63FC27FD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14FD4BFDD9FB1C.text	3C6E87D6925F2858FF14FD4BFDD9FB1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Glycyphaniola) cuculus subsp. cuculus Burmeister 1842	<div><p>Glycyphana (Glycyphaniola) cuculus cuculus Burmeister, 1842</p><p>Glycyphana cuculus Burmeister, 1842: 352; Mohnike 1873: 161.</p><p>Glycyphana palawana Moser, 1914: 593 . Type locality. “ Palawan.”</p><p>Glycyphana (Glycyphaniola) cuculus cuculus: Mikšić 1971b: 51.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Luzon (Camotes Islands, Babuyanes, Palawan, and Polillo), Visayas (Bohol, Leyte, Panay, and Samar), and Mindanao (Basilan, Bukidnon, Camiguin, Zamboanga del Norte, Davao, Lanao del Norte, and Surigao) (Mohnike 1873; Moser 1914; Mikšić 1967, 1971b).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p><p>Remarks. Wallace (1867) noted that Glycyphana (Glycyphaniola) modesta had been collected from the Philippines (coll. Parry). However, the species is currently known to occur in Indonesia and Malaysia, and the Philippine specimens were likely misidentified as G. modesta instead of G. cuculus, given the subtle morphological differences. Wallace himself remarked, “this is probably only a variety of E. modesta,” suggesting taxonomic ambiguity.</p></div>	https://treatment.plazi.org/id/3C6E87D6925F2858FF14FD4BFDD9FB1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14FB6FFC27FA3C.text	3C6E87D6925F2858FF14FB6FFC27FA3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Glycyphaniola) cuculus subsp. nigropygidialis Miksic 1968	<div><p>Glycyphana (Glycyphaniola) cuculus nigropygidialis Mikšić, 1968</p><p>Glycyphana (Glycyphaniola) cuculus nigropygidialis Mikšić, 1968: 306; Mikšić 1971b: 56; Mikšić 1984: 313.</p><p>Type locality. “Süd Luzon, Panay.”</p><p>Distribution in the Philippines. Luzon (Masbate and Romblon), Visayas (Negros and Panay), and Mindanao (Surigao del Norte) (Mikšić 1968, 1971b).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925F2858FF14FB6FFC27FA3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14F98FFC27F938.text	3C6E87D6925F2858FF14F98FFC27F938.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Glycyphaniola) cuculus subsp. pilifera Miksic 1967	<div><p>Glycyphana (Glycyphaniola) cuculus pilifera Mikšić, 1967</p><p>Glycyphana (Glycyphaniola) cuculus pilifera Mikšić, 1967: 28; Mikšić 1971b: 55.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Baguio, Bataan, Ilocos, Laguna, Masbate, Polillo, Mindoro, Mountain Province, Rizal, and Nueva Vizcaya) (Mikšić 1967, 1971b).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925F2858FF14F98FFC27F938	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925F2858FF14F893FCDCF860.text	3C6E87D6925F2858FF14F893FCDCF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Glycyphaniola) delponti Antoine 2002	<div><p>Glycyphana (Glycyphaniola) delponti Antoine, 2002</p><p>Glycyphana (Glycyphaniola) delponti Antoine, 2002: 12 .</p><p>Type locality. “ Irawan, Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Antoine 2002).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925F2858FF14F893FCDCF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FF0BFCDCFE99.text	3C6E87D6925C285BFF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Glycyphaniola) petrovitzi Miksic 1968	<div><p>Glycyphana (Glycyphaniola) petrovitzi Mikšić, 1968</p><p>Glycyphana (Glycyphaniola) petrovitzi Mikšić, 1968: 306; Mikšić 1971b: 87.</p><p>Type locality. “N. Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Mikšić 1968, 1971b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925C285BFF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FEF3FC27FDC1.text	3C6E87D6925C285BFF14FEF3FC27FDC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Glycyphaniola) quadricolor subsp. mindoroensis Miksic 1968	<div><p>Glycyphana (Glycyphaniola) quadricolor mindoroensis Mikšić, 1968</p><p>Glycyphana (Glycyphaniola) quadricolor mindoroensis Mikšić, 1968: 48; Mikšić 1971b: 71.</p><p>Type locality. “Mindoro, P. I.”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Mikšić 1968, 1971b).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925C285BFF14FEF3FC27FDC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FDDBFC29FCE1.text	3C6E87D6925C285BFF14FDDBFC29FCE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Glycyphaniola) quadricolor subsp. niveopunctata Moser 1922	<div><p>Glycyphana (Glycyphaniola) quadricolor niveopunctata Moser, 1922</p><p>Glycyphana niveopunctata Moser, 1922: 113 .</p><p>Glycyphana (Glycyphaniola) quadricolor niveopunctata: Mikšić, 1971b: 72; Mikšić 1983: 393; Pavićević 1988: 172.</p><p>Type locality. “Ins. Sibuyan.”</p><p>Distribution in the Philippines. Luzon (Sibuyan) (Moser 1922; Mikšić 1971b; Pavićević 1988).</p><p>Overall Distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925C285BFF14FDDBFC29FCE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FCFBFC29FBC9.text	3C6E87D6925C285BFF14FCFBFC29FBC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Glycyphaniola) quadricolor subsp. rimskii Miksic 1967	<div><p>Glycyphana (Glycyphaniola) quadricolor rimskii Mikšić, 1967</p><p>Glycyphana rubroscutellaris rimskii Mikšić, 1967: 27 .</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Benguet, Nueva Vizcaya, Mt. Banahao, and Rizal) (Mikšić 1967).</p><p>Overall Distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925C285BFF14FCFBFC29FBC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14FBA3FC29FA3D.text	3C6E87D6925C285BFF14FBA3FC29FA3D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Glycyphaniola) quadricolor subsp. rubroscutellaris Mohnike 1873	<div><p>Glycyphana (Glycyphaniola) quadricolor rubroscutellaris Mohnike, 1873</p><p>Glycyphana rubroscutellaris Mohnike, 1873: 159 .</p><p>Glycyphana rubroscutellaris immarginicollis Moser, 1925: 69 . Type locality. “Negros, Occidental Negros, Cuernos Mountains.”</p><p>Glycyphana rubroscutellaris nigroscutellaris Moser, 1925: 69 . Type locality. “ Mindanao, Lanao Province, Mumungan.”</p><p>Glycyphana (Glycyphaniola) quadricolor rubroscutellaris: Mikšić 1971b: 73; Pavićević 1988: 172.</p><p>Type locality. “ Bohol et Mindanao.”</p><p>Distribution in the Philippines. Luzon (Bataan, Benguet, Nueva Vizcaya, Rizal, and Mt. Banahao), Visayas (Bohol, Negros Island, and Leyte), and Mindanao (Bukidnon, Agusan del Norte, and Lanao del Norte) (Mohnike 1873; Moser 1925; Mikšić 1971b; Pavićević 1988).</p><p>Overall Distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925C285BFF14FBA3FC29FA3D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14F9C7FCC6F8DD.text	3C6E87D6925C285BFF14F9C7FCC6F8DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Macroglycyphana) mohagani Legrand 2004	<div><p>Glycyphana (Macroglycyphana) mohagani Legrand, 2004</p><p>Glycyphana (Macroglycyphana) mohagani Legrand, 2004: 9 .</p><p>Type locality. “ Philippines, île de Leyte.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Legrand 2004).</p><p>Overall Distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925C285BFF14F9C7FCC6F8DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925C285BFF14F8AFFCC6F805.text	3C6E87D6925C285BFF14F8AFFCC6F805.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycyphana (Macroglycyphana) nigra Miksic 1986	<div><p>Glycyphana (Macroglycyphana) nigra Mikšić, 1986</p><p>Glycyphana (Euglycyphana) nigra Mikšić, 1986: 72 .</p><p>Type locality. “ Insel Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Mikšić 1986).</p><p>Overall Distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925C285BFF14F8AFFCC6F805	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285AFF14FF42FCC6FE74.text	3C6E87D6925D285AFF14FF42FCC6FE74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paraprotaetia juliae (Nagai 1984)	<div><p>Paraprotaetia juliae (Nagai, 1984)</p><p>Protaetia (Poecilophana) juliae Nagai, 1984: 3; Mikšić 1986: 70; Kracjik 1998: 42.</p><p>Paraprotaetia juliae: Sakai &amp; Nagai 1998: 292.</p><p>Type locality. “ Philippines (Palawan Island).”</p><p>Distribution in the Philippines. Luzon (Palawan) (Nagai 1984; Mikšić 1986).</p><p>Overall Distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925D285AFF14FF42FCC6FE74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285AFF14FDBFFCC6FD14.text	3C6E87D6925D285AFF14FDBFFCC6FD14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Podopogonus boettcheri Moser 1917	<div><p>Podopogonus boettcheri Moser, 1917</p><p>Podopogonus boettcheri Moser, 1917a: 18; Heller 1934: 305; Mikšić 1982a: 337.</p><p>Type locality. “Luzon (Imugan).”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya and Rizal) (Moser 1917a; Heller 1934).</p><p>Overall Distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925D285AFF14FDBFFCC6FD14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285AFF14FC16FB83FB1C.text	3C6E87D6925D285AFF14FC16FB83FB1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Caloprotaetia) procera (White 1856)	<div><p>Protaetia (Caloprotaetia) procera (White, 1856)</p><p>Cetonia procera White, 1856: 17; Mohnike 1873: 188.</p><p>Protaetia procera: Schoch 1896: 58.</p><p>Protaetia (Caloprotaetia) procera: Mikšić 1963a: 363; Mikšić 1963b: 416; Mikšić 1987: 437; Pavićević 1988: 172.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Visayas (Negros Oriental) and Mindanao (Davao) (White 1856; Wallace 1867; Mohnike 1873; Mikšić 1963b; Pavićević 1988).</p><p>Overall distribution. The Philippines and China (Taiwan) (Mikšić 1963b).</p></div>	https://treatment.plazi.org/id/3C6E87D6925D285AFF14FC16FB83FB1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285AFF14FAA6FCDCF9F4.text	3C6E87D6925D285AFF14FAA6FCDCF9F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Chalcoprotaetia) franzi Miksic 1963	<div><p>Protaetia (Chalcoprotaetia) franzi Mikšić, 1963</p><p>Protaetia (Chalcoprotaetia) franzi Mikšič, 1963a: 432; Mikšič 1963b: 431; Mikšič 1987: 158; Krajčík 1998: 37; Sakai &amp; Nagai 1998: 275; Jákl 2023: 317.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Romblon) and Visayas (Bohol, Negros Occidental, and Panay) (Mikšić 1963a, 1963b, 1987; Jákl 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925D285AFF14FAA6FCDCF9F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925D285DFF14F9C6FCDCFEC5.text	3C6E87D6925D285DFF14F9C6FCDCFEC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Chalcoprotaetia) philippensis (Fabricius 1775)	<div><p>Protaetia (Chalcoprotaetia) philippensis (Fabricius, 1775)</p><p>Cetonia philippensis Fabricius, 1775: 49; Blanchard 1842: 4; Gory &amp; Percheron 1833: 193; Redtenbacher 1868: 81; Wallace 1868: 581; Gemminger &amp; Harold 1869: 581; Mohnike 1873: 209.</p><p>Protaetia philippensis: Burmeister 1842: 496; Schoch 1895: 115; Schoch 1896: 58; Schenkling 1921: 260.</p><p>Cetonia hieroglyphica Gory &amp; Percheron, 1833: 175 . Type locality. “Des Indes orientalis.” (Ambiguous)</p><p>Cetonia philippensis luzonica Kraatz, 1890: 218 . Type locality. “Luzon.”</p><p>Protaetia (Chalcoprotaetia) philippensis: Mikšič 1963a: 430; Mikšič 1963b: 431; Mikšič 1987: 155; Krajčík 1998: 37; Sakai &amp; Nagai 1998: 275; Jákl 2023: 319.</p><p>Type locality. “Philippinis.”</p><p>Distribution in the Philippines. Luzon (Camarines Sur, Laguna, Manila, Marinduque, Ilocos, Nueva Vizcaya, Rizal, Romblon, and Tarlac) and Visayas (Negros Island, Panay) (Fabricius 1775; Gory &amp; Percheron 1833; Wallace 1868; Mohnike 1873; Kraatz 1890; Mikšić 1963b; Jákl 2022).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925D285DFF14F9C6FCDCFEC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14FED7FCDCFDE2.text	3C6E87D6925A285DFF14FED7FCDCFDE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Chalcoprotaetia) purpurissata (Mohnike 1873)	<div><p>Protaetia (Chalcoprotaetia) purpurissata (Mohnike, 1873)</p><p>Cetonia purpurissata Mohnike, 1873: 211 .</p><p>Protaetia purpurissata: Schenkling 1921: 261</p><p>Protaetia (Chalcoprotaetia) purpurissata: Mikšič 1963a: 429; Jákl 2023: 320.</p><p>Type locality. “Babuynes.”</p><p>Distribution in the Philippines. Luzon (Babuyan Archipelago) (Mohnike 1873; Mikšić 1963a; Jákl 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925A285DFF14FED7FCDCFDE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14FDF4FCDCFCCB.text	3C6E87D6925A285DFF14FDF4FCDCFCCB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Chalcoprotaetia) sierramadrensis Jakl 2023	<div><p>Protaetia (Chalcoprotaetia) sierramadrensis Jákl, 2023</p><p>Protaetia (Chalcoprotaetia) sierramadrensis Jákl, 2023: 320 .</p><p>Type locality. “ North Luzon Island, Sierra Madre Mountains.”</p><p>Distribution in the Philippines. Luzon (Sierra Madre) (Jákl 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925A285DFF14FDF4FCDCFCCB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14FC15FCDCFAF5.text	3C6E87D6925A285DFF14FC15FCDCFAF5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) bifenestrata (Chevrolat 1841)	<div><p>Protaetia (Euprotaetia) bifenestrata (Chevrolat, 1841)</p><p>Cetonia bifenestrata Chevrolat, 1841: 223; Blanchard 1842: 1; Gemminger &amp; Harold 1869: 1322; Wallace 1868: 579; Mohnike 1873: 189.</p><p>Cetonia indra Hope, 1841: 33 . Type locality. “ Manilla.”</p><p>Cetonia gemella Newman, 1841: 170 . Type locality. “ Philippine Islands.”.</p><p>Protaetia bifenestrata: Burmeister 1842: 492; Schoch 1895: 116; Schoch 1896: 56.</p><p>Protaetia (Euprotaetia) bifenestrata: Mikšić 1963a: 415; Mikšić 1963b: 423; Pavićević 1988: 176.</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Laguna, Manila, Marinduque, Rizal, Romblon, Ilocos, Camarines Sur, and Sorsogon) and Mindanao (Mt. Apo) (Chevrolat 1841; Hope 1841; Newman 1841; Mikšić 1963a, 1963b; Pavićević 1988).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925A285DFF14FC15FCDCFAF5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14FAC7FCDCF94A.text	3C6E87D6925A285DFF14FAC7FCDCF94A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) boholica (Mohnike 1873)	<div><p>Protaetia (Euprotaetia) boholica (Mohnike, 1873)</p><p>Cetonia boholica Mohnike, 1873: 231;</p><p>Protaetia boholica: Schoch 1896: 56.</p><p>Protaetia (Euprotaetia) boholica: Mikšić 1963a: 417 (subspecies of Protaetia bifenestrata); Mikšić 1963b: 423.</p><p>Type locality. “ Bohol.”</p><p>Distribution in the Philippines. Visayas (Bohol, Leyte, and Samar) (Chevrolat 1841; Hope 1841; Newman 1841; Mikšić 1963a, 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925A285DFF14FAC7FCDCF94A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285DFF14F95CFCDCF8B3.text	3C6E87D6925A285DFF14F95CFCDCF8B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) endroedii Miksic 1963	<div><p>Protaetia (Euprotaetia) endroedii Mikšić, 1963</p><p>Protaetia bifenestrata endroedii Mikšić, 1963a: 417 .</p><p>Type locality. “ Cebu.”</p><p>Distribution in the Philippines. Visayas (Cebu) (Mikšić 1963a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925A285DFF14F95CFCDCF8B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925A285CFF14F805FC27FF0D.text	3C6E87D6925A285CFF14F805FC27FF0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) gertrudae subsp. gertrudae Arnaud 1989	<div><p>Protaetia (Euprotaetia) gertrudae gertrudae Arnaud, 1989</p><p>Protaetia (Euprotaetia) gertrudae Arnaud, 1989a: 3 .</p><p>Type locality. “ Philippines, Negros Oriental.”</p><p>Distribution in the Philippines. Visayas (Negros) (Arnaud 1989a).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925A285CFF14F805FC27FF0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FE9FFC27FE75.text	3C6E87D6925B285CFF14FE9FFC27FE75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) gertrudae subsp. negricola Pavicevic 1988	<div><p>Protaetia (Euprotaetia) gertrudae negricola Pavićević, 1988</p><p>Protaetia (Euprotaetia) endroedii negricola Pavićević, 1988: 176 .</p><p>Type locality. “ Negros Oriental.”</p><p>Distribution in the Philippines. Visayas (Negros) (Pavićević 1988).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925B285CFF14FE9FFC27FE75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FE44FCDCFD5A.text	3C6E87D6925B285CFF14FE44FCDCFD5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) inexpectata Pavicevic 1986	<div><p>Protaetia (Euprotaetia) inexpectata Pavićević, 1986</p><p>Protaetia (Euprotaetia) inexpectata Pavićević, 1986: 117 .</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Pavićević 1986).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925B285CFF14FE44FCDCFD5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FD2CFCDCFC83.text	3C6E87D6925B285CFF14FD2CFCDCFC83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) kuehbandneri Miksic 1983	<div><p>Protaetia (Euprotaetia) kuehbandneri Mikšić, 1983</p><p>Protaetia (Euprotaetia) kuehbandneri Mikšić, 1983: 387; Kühbandner 1984: 210; Pavićević 1988: 176.</p><p>Type locality. “ Philippinen, Insel Romblon.”</p><p>Distribution in the Philippines. Luzon (Sorsogon and Romblon) (Mikšić 1983; Pavićević 1988).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925B285CFF14FD2CFCDCFC83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FBC2FCDCFAD1.text	3C6E87D6925B285CFF14FBC2FCDCFAD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) lumawigi Arnaud 1987	<div><p>Protaetia (Euprotaetia) lumawigi Arnaud, 1987</p><p>Protaetia (Euprotaetia) lumawigi Arnaud, 1987a: 24; Arnaud 1987c: 4.</p><p>Type locality. “ Philippines, Marinduque.”</p><p>Distribution in the Philippines. Luzon (Marinduque) (Arnaud 1987a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925B285CFF14FBC2FCDCFAD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FC15FCDCFBE8.text	3C6E87D6925B285CFF14FC15FCDCFBE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) mariae Arnaud 2004	<div><p>Protaetia (Euprotaetia) mariae Arnaud, 2004</p><p>Protaetia (Euprotaetia) mariae Arnaud, 2004: 2 .</p><p>Type locality. “ Philippines, North Luzon, Calayan Island.”</p><p>Distribution in the Philippines. Luzon (Babuyan Island) (Arnaud 2004).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925B285CFF14FC15FCDCFBE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14FAABFCDCF992.text	3C6E87D6925B285CFF14FAABFCDCF992.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) nox Janson 1881	<div><p>Protaetia (Euprotaetia) nox Janson, 1881</p><p>Protaetia nox Janson, 1881: 609; Schoch, 1895: 140; Schoch 1896: 58.</p><p>Protaetia (Euprotaetia) nox: Mikšić 1963a: 417 (subspecies of Protaetia bifenestrata); Mikšić 1963b: 424.</p><p>Type locality. “ Dinagat, Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Mindoro) and Mindanao (Agusan del Sur, Dinagat, Lanao del Sur, and Surigao) (Janson 1881; Mikšić 1963a, 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925B285CFF14FAABFCDCF992	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14F9E4FCDCF8FB.text	3C6E87D6925B285CFF14F9E4FCDCF8FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) sebastieni Arnaud 2019	<div><p>Protaetia (Euprotaetia) sebastieni Arnaud, 2019</p><p>Protaetia (Euprotaetia) sebastieni Arnaud, 2019: 3 .</p><p>Type locality. “ Philippines, Mindoro, Mt. Halcon. ”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Arnaud 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925B285CFF14F9E4FCDCF8FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6925B285CFF14F8CDFCDCF823.text	3C6E87D6925B285CFF14F8CDFCDCF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) teisseyrei Arnaud 1989	<div><p>Protaetia (Euprotaetia) teisseyrei Arnaud, 1989</p><p>Protaetia (Euprotaetia) teisseyrei Arnaud, 1989b: 24 .</p><p>Type locality. “ Philippines, North Samar.”</p><p>Distribution in the Philippines. Visayas (Samar) (Arnaud 1989b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6925B285CFF14F8CDFCDCF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FF0AFCC6FEBC.text	3C6E87D69258285FFF14FF0AFCC6FEBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetia) uhligi Arnaud 1987	<div><p>Protaetia (Euprotaetia) uhligi Arnaud, 1987</p><p>Protaetia (Euprotaetia) uhligi Arnaud, 1987a: 24; Arnaud 1987c: 4.</p><p>Type locality. “ Philippine, Nord Luzon, Nueva Vizcaya.”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya, Rizal, Manila, and Mountain Province) (Arnaud 1987a).</p><p>Overall Distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69258285FFF14FF0AFCC6FEBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FE46FCC6FD5C.text	3C6E87D69258285FFF14FE46FCC6FD5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Euprotaetiomima) marlenae Legrand & Chew Kea Foo 2004	<div><p>Protaetia (Euprotaetiomima) marlenae Legrand &amp; Chew Kea Foo, 2004</p><p>Protaetia (Euprotaetiomima) marlenae Legrand &amp; Chew Kea Foo, 2004: 9 .</p><p>Type locality. “ Philippines, Leyte, Mont Balocaue.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Legrand &amp; Chew Kea Foo 2004).</p><p>Overall Distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69258285FFF14FE46FCC6FD5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FD66FCDCFC10.text	3C6E87D69258285FFF14FD66FCDCFC10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Finkia) nigrobrunnea Moser 1909	<div><p>Protaetia (Finkia) nigrobrunnea Moser, 1909</p><p>Protaetia nigrobrunnea Moser, 1909: 319; Moser 1911b: 123; Mikšić 1963a: 445; Mikšić 1987: 490.</p><p>Protaetia (Finkia) nigrobrunnea: Krajčik 1998: 37.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Luzon (Ilocos) (Moser 1909; Mikšić 1963a, 1987).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69258285FFF14FD66FCDCFC10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FBA2FCDCFAE8.text	3C6E87D69258285FFF14FBA2FCDCFAE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Goetzia) luridoguttata Moser 1918	<div><p>Protaetia (Goetzia) luridoguttata Moser, 1918</p><p>Protaetia luridoguttata Moser, 1918: 175; Mikšić 1963a: 443.</p><p>Protaetia (Goetzia) luridoguttata: Mikšić, 1963b: 434; Mikšić 1987: 464.</p><p>Type locality. “ Mindanao (Davao).”</p><p>Distribution in the Philippines. Luzon (Romblon) and Mindanao (Agusan del Norte and Davao) (Moser 1918; Mikšić 1963b; 1987).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69258285FFF14FBA2FCDCFAE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14FA3AFCDCF9AC.text	3C6E87D69258285FFF14FA3AFCDCF9AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Hemiprotaetia) boudanti Arnaud 1992	<div><p>Protaetia (Hemiprotaetia) boudanti Arnaud, 1992</p><p>Hemiprotaetia boudanti Arnaud, 1992: 29; Krajčík 1998: 34; Sakai &amp; Nagai 1998: 291.</p><p>Protaetia (Hemiprotaetia) boudanti: Jákl 2023: 310.</p><p>Type locality. ‘‘ Philippines, Panay Isld., Antique.”</p><p>Distribution in the Philippines. Visayas (Negros and Panay) (Arnaud 1992; Jákl 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69258285FFF14FA3AFCDCF9AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69258285FFF14F93EFCDCF84C.text	3C6E87D69258285FFF14F93EFCDCF84C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Hemiprotaetia) cupriventris Moser 1917	<div><p>Protaetia (Hemiprotaetia) cupriventris Moser, 1917</p><p>Protaetia cupriventris Moser, 1917a: 9; Mikšič 1963a: 431 (= Protaetia dubia); Mikšič 1982a: 163 (= Protaetia dubia); Arnaud 1992: 32; Krajčík 1998: 34 (= Hemiprotaetia dubia).</p><p>Protaetia (Hemiprotaetia) cupriventris: Jákl 2023: 311.</p><p>Type locality. “Luzon (Bayombong).”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1917a, 1918; Jákl 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69258285FFF14F93EFCDCF84C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14FF0BFCDCFD5D.text	3C6E87D69259285EFF14FF0BFCDCFD5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Hemiprotaetia) dubia (Wallace 1867)	<div><p>Protaetia (Hemiprotaetia) dubia (Wallace, 1867)</p><p>Cetonia dubia Wallace, 1867: 97; Wallace 1868: 582; Gemminger &amp; Harold 1869: 1323; Mohnike 1873: 192; Bourgoin 1929: 270.</p><p>Protaetia dubia: Schoch 1896: 57.</p><p>Protaetia gregori Moser, 1906: 278 . Type locality. “Luzon.”; Mikšič 1963a: 431.</p><p>Protaetia flavomaculata Moser, 1914: 590 . Type locality. “Luzon, Mt. Banahao.”; Mikšič 1963a: 431.</p><p>Protaetia monticola Moser, 1917a: 10 . Type locality. “Luzon, Mt. Isarog.”; Mikšič 1963a: 431.</p><p>Protaetia montivaga Moser, 1917a: 8 . Type locality. “Luzon, Mt. Isarog.”; Mikšič 1963a: 431.</p><p>Protaetia (Hemiprotaetia) dubia: Mikšič 1963a: 435; Mikšič 1963b: 431; Jákl 2023: 313.</p><p>Type locality. “ Philippines Islands.”</p><p>Distribution in the Philippines. Luzon (Camarines Sur, Ifugao, Mountain Province, Laguna, Ilocos, and Nueva Vizcaya) (Wallace 1867; Moser 1906, 1914, 1917a; Bourgoin 1929; Mikšić 1963a, 1963b, 1972b; Jákl 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69259285EFF14FF0BFCDCFD5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14FD2FFCDCFC11.text	3C6E87D69259285EFF14FD2FFCDCFC11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Hemiprotaetia) isarogensis Moser 1917	<div><p>Protaetia (Hemiprotaetia) isarogensis Moser, 1917</p><p>Protaetia isarogensis Moser, 1917: 7 .</p><p>Hemiprotaetia isarogensis: Mikšič 1982a: 167; Krajčík 1998: 34; Sakai &amp; Nagai 1998: 291.</p><p>Protaetia (Hemiprotaetia) isarogensis: Mikšič 1963a: 439; Mikšič 1963b: 433; Jákl 2023: 314.</p><p>Type locality. “ Luzon (Mt. Isarog).”</p><p>Distribution in the Philippines. Luzon (Camarines Sur and Sorsogon) (Moser 1917a; Mikšič 1963a, 1963b; Jákl 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69259285EFF14FD2FFCDCFC11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14FC6BFCDCFB79.text	3C6E87D69259285EFF14FC6BFCDCFB79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Hemiprotaetia) madjaasica Jakl 2023	<div><p>Protaetia (Hemiprotaetia) madjaasica Jákl, 2023</p><p>Protaetia (Hemiprotaetia) madjaasica Jákl, 2023: 314 .</p><p>Type locality. “ Panay Island, Antique Province, Mount Madjaas. ”</p><p>Distribution in the Philippines. Visayas (Panay) (Jákl 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69259285EFF14FC6BFCDCFB79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14FA8BFCDCFA3D.text	3C6E87D69259285EFF14FA8BFCDCFA3D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Lawangia) bruvieri Legrand 2018	<div><p>Protaetia (Lawangia) bruvieri Legrand, 2018</p><p>Protaetia (Miksicoprotaetia) bruvieri Legrand, 2018: 4 .</p><p>Protaetia (Lawangia) bruvieri: Jákl 2022: 98.</p><p>Type locality. “ Mindoro, Mt. Halcon. ”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Legrand 2018b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69259285EFF14FA8BFCDCFA3D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69259285EFF14F98FFCDCF965.text	3C6E87D69259285EFF14F98FFCDCF965.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Lawangia) jeanphilippei Jakl 2022	<div><p>Protaetia (Lawangia) jeanphilippei Jákl, 2022</p><p>Protaetia (Lawangia) jeanphilippei Jákl, 2022: 96 .</p><p>Type locality. “ Philippines, Panay Island, Antique Province, Mount Madjaas. ”</p><p>Distribution in the Philippines. Visayas (Panay) (Jákl 2022).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69259285EFF14F98FFCDCF965	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692592851FF14F8AFFCDCFF20.text	3C6E87D692592851FF14F8AFFCDCFF20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Miksicia) arrogans (Wallace 1867)	<div><p>Protaetia (Miksicia) arrogans (Wallace, 1867)</p><p>Cetonia arrogans Wallace, 1867: 97; Wallace 1868: 584; Mohnike 1873: 184; Schoch 1896: 56.</p><p>Protaetia arrogans: Mikšić 1962: 28.</p><p>Protaetia (Urbania) arrogans: Mikšić 1963a: 359.</p><p>Protaetia (Miksicus) arrogans: Özdikmen &amp; Turgut 2009: 227.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Wallace 1867).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692592851FF14F8AFFCDCFF20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FEBAFCDCFDC0.text	3C6E87D692562851FF14FEBAFCDCFDC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Miksicia) balthasari Miksic 1962	<div><p>Protaetia (Miksicia) balthasari Mikšić, 1962</p><p>Protaetia balthasari Mikšić, 1962: 26 .</p><p>Protaetia (Urbania) balthasari: Mikšić 1963a: 359.</p><p>Protaetia (Miksicus) balthasari: Özdikmen &amp; Turgut 2009: 227.</p><p>Type locality. “Barbatan Is.- Antique.”</p><p>Distribution in the Philippines. Visayas (Antique) (Mikšić 1962).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692562851FF14FEBAFCDCFDC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FDDAFCDCFCCC.text	3C6E87D692562851FF14FDDAFCDCFCCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Miksicia) kaorusakaii Jakl 2021	<div><p>Protaetia (Miksicia) kaorusakaii Jákl, 2021</p><p>Miksicus sakaii Ja ́kl, 2011: 208.</p><p>Protaetia (Miksicia) kaorusakaii: Jákl 2021: 309.</p><p>Type locality. “ Philippines, Palawan, Puerto Princesa.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Jákl 2011, 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692562851FF14FDDAFCDCFCCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FC16FCDCFB78.text	3C6E87D692562851FF14FC16FCDCFB78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Netociomima) davaoana Moser 1917	<div><p>Protaetia (Netociomima) davaoana Moser, 1917</p><p>Protaetia davaoana Moser, 1917a: 14 .</p><p>Protaetia (Netociomima) davaoana: Mikšić 1963a: 349; Mikšić 1963b: 415; Mikšić 1987: 362; Jákl 2017b: 20.</p><p>Type locality. “ Mindanao (Davao).”</p><p>Distribution in the Philippines. Visayas (Bohol and Leyte) and Mindanao (Bukidnon, Lanao del Norte, and Davao) (Moser 1917a; Mikšić 1963a, 1963b; Jákl 2017b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692562851FF14FC16FCDCFB78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14FB52FABEFA18.text	3C6E87D692562851FF14FB52FABEFA18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Netociomima) palavanica Miksic 1963	<div><p>Protaetia (Netociomima) palavanica Mikšić, 1963</p><p>Protaetia (Autoprotaetia) bipunctata palavanica Mikšić, 1965c: 163; Mikšić 1987: 375.</p><p>Protaetia (Netociomima) palavanica: Jákl 2017b: 49.</p><p>Type locality. “ Palavan Island .”</p><p>Distribution in the Philippines. Luzon (Palawan) (Mikšić 1963a; Jákl 2017b).</p><p>Overall distribution. The Philippines and Indonesia.</p><p>Remarks. Jákl (2017b) noted that the record of this species in Palawan is probably erroneous.</p></div>	https://treatment.plazi.org/id/3C6E87D692562851FF14FB52FABEFA18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14F9AAFCDCF938.text	3C6E87D692562851FF14F9AAFCDCF938.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Odontoprotaetia) butozani Miksic 1963	<div><p>Protaetia (Odontoprotaetia) butozani Mikšić, 1963</p><p>Protaetia (Odontoprotaetia) butozani Mikšić, 1963b: 430 .</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Cagayan Province) (Mikšić 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692562851FF14F9AAFCDCF938	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692562851FF14F892FCDCF804.text	3C6E87D692562851FF14F892FCDCF804.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Odontoprotaetia) cupriceps Moser 1917	<div><p>Protaetia (Odontoprotaetia) cupriceps Moser, 1917</p><p>Protaetia cupriceps Moser, 1917a: 12 .</p><p>Protaetia (Odontoprotaetia) hiekei Mikšić, 1963b: 427; Mikšić 1970a: 57.</p><p>Type locality. “Luzon (Imugan).”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1917a; Mikšić 1963a, 1963b, 1970a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692562851FF14F892FCDCF804	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FF0AFCDCFE98.text	3C6E87D692572850FF14FF0AFCDCFE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Odontoprotaetia) hallegueni Arnaud 2004	<div><p>Protaetia (Odontoprotaetia) hallegueni Arnaud, 2004</p><p>Protaetia (Odontoprotaetia) hallegueni Arnaud, 2004: 2 .</p><p>Type locality. “ Philippines, North Luzon, Babuyan Island.”</p><p>Distribution in the Philippines. Luzon (Babuyan Island) (Arnaud 2004).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692572850FF14FF0AFCDCFE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FEF2FC27FDE4.text	3C6E87D692572850FF14FEF2FC27FDE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Odontoprotaetia) venerabilis subsp. niveomaculata Moser 1918	<div><p>Protaetia (Odontoprotaetia) venerabilis niveomaculata Moser, 1918</p><p>Protaetia niveomaculata Moser, 1918: 174 .</p><p>Protaetia (Odontoprotaetia) venerabilis grandemaculata Mikšić, 1963a: 424; Mikšić 1963b: 425.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Albay) (Moser, 1918; Mikšić 1963a, 1963b).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692572850FF14FEF2FC27FDE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FDF6FC27FC84.text	3C6E87D692572850FF14FDF6FC27FC84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Odontoprotaetia) venerabilis subsp. venerabilis Mohnike 1873	<div><p>Protaetia (Odontoprotaetia) venerabilis venerabilis Mohnike, 1873</p><p>Cetonia venerabilis Mohnike, 1873: 229 .</p><p>Protaetia (Odontoprotaetia) venerabilis nigrotomentosa Mikšić, 1963a: 424; Mikšić 1963b: 425.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Cagayan Province, Catanduanes, Rizal, Manila, and Laguna) and Mindanao (Mohnike 1873; Mikšić 1963a, 1963b).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692572850FF14FDF6FC27FC84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FC4EFCDCFBA4.text	3C6E87D692572850FF14FC4EFCDCFBA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Poecilophana) audreyae Arnaud 1989	<div><p>Protaetia (Poecilophana) audreyae Arnaud, 1989</p><p>Protaetia (Poecilophana) audreyae Arnaud, 1989b: 23 .</p><p>Type locality. “ Philippines, Mt. Apo.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Arnaud 1989b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692572850FF14FC4EFCDCFBA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14FB36FCDCF9D0.text	3C6E87D692572850FF14FB36FCDCF9D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Poecilophana) ochroplagiata (Heller 1895)	<div><p>Protaetia (Poecilophana) ochroplagiata (Heller, 1895)</p><p>Cetonia ochroplagiata Heller, 1895: 283 .</p><p>Poecilophana ochroplagiata: Schoch 1896: 59.</p><p>Protaetia (Poecilophana) ochroplagiata: Mikšić 1963a: 365; Mikšić 1963b: 417; Mikšić 1987: 184; Pavićević 1988: 173.</p><p>Type locality. “Mindanao (Dapitan).”</p><p>Distribution in the Philippines. Luzon (Benguet), Visayas (Negros Oriental), and Mindanao (Agusan del Norte, Bukidnon, Basilan, Davao, and Zamboanga del Norte) (Heller 1895; Kraatz 1895; Mikšić 1963a, 1963b, 1987; Pavićević 1988).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692572850FF14FB36FCDCF9D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692572850FF14F9E2FCDCF860.text	3C6E87D692572850FF14F9E2FCDCF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) ambigua (Chevrolat 1841)	<div><p>Protaetia (Protaetia) ambigua (Chevrolat, 1841)</p><p>Cetonia ambigua Chevrolat, 1841: 223; Mohnike 1873: 207.</p><p>Protaetia ambigua: Burmeister 1842: 499; Schoch 1896: 56.</p><p>Protaetia ambigua ferrugineotomentosa Mikšić, 1963a: 385 . Type locality. “Bayombong.”</p><p>Protaetia (Protaetia) ambigua: Mikšić 1963b: 419.</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Laguna, Manila, and Nueva Vizcaya) and Visayas (Leyte) (Chevrolat 1841; Mikšić 1963a, 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692572850FF14F9E2FCDCF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FF0BFCDCFE99.text	3C6E87D692542853FF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) anneliae Reichenbach 1995	<div><p>Protaetia (Protaetia) anneliae Reichenbach, 1995</p><p>Protaetia (Protaetia) anneliae Reichenbach, 1995: 45 .</p><p>Type locality. “ Philippines, Romblon.”</p><p>Distribution in the Philippines. Luzon (Romblon) (Reichenbach 1995).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692542853FF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FEF3FCDCFC59.text	3C6E87D692542853FF14FEF3FCDCFC59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) anovittata (Chevrolat 1841)	<div><p>Protaetia (Protaetia) anovittata (Chevrolat, 1841)</p><p>Cetonia anovittata Chevrolat, 1841: 223; Blanchard 1842: 5; Gemminger &amp; Harold 1869: 1321; Mohnike 1873: 220.</p><p>Protaetia anovittata: Burmeister 1842: 497; Schoch 1895: 117; Schoch 1896: 56.</p><p>Cetonia chloris Newman, 1841: 170 . Type locality. “ Philippines Islands.”</p><p>Cetonia anovittata monovittata Haller, 1884: 31 . Type locality. “ Philippines (?).”</p><p>Cetonia anovittata signata Haller, 1884: 32 . Type locality. “ Philippines (?).”</p><p>Cetonia apunctata Haller, 1884: 32 . Type locality. “ Philippines (?).”</p><p>Protaetia (Protaetia) anovittata: Mikšić 1963a: 389; Mikšić 1963b: 419; Mikšić 1984: 303; Reichenbach 1995: 46.</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Albay, Camarines Sur, Laguna, Manila, Nueva Vizcaya, Romblon, and Polillo), Visayas (Panay), and Mindanao (Bukidnon and Mt. Apo) (Chevrolat 1841; Newman 1841; Mohnike 1873; Wallace 1868; Haller 1884; Mikšić 1963a, 1963b, 1984; Reichenbach 1995).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692542853FF14FEF3FCDCFC59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FC33FCDCFBA5.text	3C6E87D692542853FF14FC33FCDCFBA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) antoinei Arnaud 1989	<div><p>Protaetia (Protaetia) antoinei Arnaud, 1989</p><p>Protaetia miksici Arnaud, 1987c: 4 .</p><p>Protaetia (Protaetia) antoinei Arnaud, 1989b: 24 .</p><p>Type locality. “ Sibuyan Island, Romblon, Philippines.”</p><p>Distribution in the Philippines. Luzon (Romblon) (Arnaud 1987c, 1989b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692542853FF14FC33FCDCFBA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FB37FCDCFA8D.text	3C6E87D692542853FF14FB37FCDCFA8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) boudanti Arnaud 1992	<div><p>Protaetia (Protaetia) boudanti Arnaud, 1992</p><p>Protaetia (Protaetia) boudanti Arnaud, 1992: 29 .</p><p>Type locality. “ Philippines, Sud Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) and Visayas (Panay) (Arnaud 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692542853FF14FB37FCDCFA8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14FA1FFCDCF9F5.text	3C6E87D692542853FF14FA1FFCDCF9F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) boulleti Arnaud 1989	<div><p>Protaetia (Protaetia) boulleti Arnaud, 1989</p><p>Protaetia (Protaetia) boulleti Arnaud, 1989a: 1 .</p><p>Type locality. “ Batanes Island, Philippines.”</p><p>Distribution in the Philippines. Luzon (Batanes) (Arnaud 1989a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692542853FF14FA1FFCDCF9F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692542853FF14F9C7FCDCF805.text	3C6E87D692542853FF14F9C7FCDCF805.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) bremei (Schaum 1844)	<div><p>Protaetia (Protaetia) bremei (Schaum, 1844)</p><p>Cetonia (Protaetia) bremei Schaum, 1844: 413; Dohrn 1872: 154.</p><p>Cetonia Bremei: Mohnike 1873: 214.</p><p>Cetonia papalis Mohnike, 1873: 198 . Type locality. “Panaon, Bohol, Leyte, Mindanao.”; Schoch 1896: 58.</p><p>Protaetia bremei inornata Mikšić, 1963a: 380 . Type locality. “ Philippinen.”</p><p>Protaetia (Protaetia) bremei: Mikšić 1963a: 379; Mikšić 1963b: 419.</p><p>Type locality. “ Manila.”</p><p>Distribution in the Philippines. Luzon (Manila), Visayas (Bohol and Leyte), and Mindanao (Camiguin) (Wallace 1867; Dohrn 1872; Kraatz 1880; Mohnike 1873; Schaum 1844; Mikšić 1963a, 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692542853FF14F9C7FCDCF805	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FF0AFCDCFE99.text	3C6E87D692552852FF14FF0AFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) brigitteae Arnaud 2019	<div><p>Protaetia (Protaetia) brigitteae Arnaud, 2019</p><p>Protaetia (Protaetia) brigitteae Arnaud, 2019: 3 .</p><p>Type locality. “ Philippines, Polillo Isld.”</p><p>Distribution in the Philippines. Luzon (Polillo Island) (Arnaud 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692552852FF14FF0AFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FEF3FCDCFDCE.text	3C6E87D692552852FF14FEF3FCDCFDCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) catanduanesiensis Miksic 1963	<div><p>Protaetia (Protaetia) catanduanesiensis Mikšić, 1963</p><p>Protaetia (Protaetia) catanduanesiensis Mikšić, 1963a: 393 .</p><p>Type locality. “ Virac (Insel Catanduanes).”</p><p>Distribution in the Philippines. Luzon (Camarines Sur) (Mikšić 1963a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692552852FF14FEF3FCDCFDCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FDD8FCDCFC83.text	3C6E87D692552852FF14FDD8FCDCFC83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) coeruleosignata (Mohnike 1873)	<div><p>Protaetia (Protaetia) coeruleosignata (Mohnike, 1873)</p><p>Cetonia coeruleosignata Mohnike, 1873: 186 .</p><p>Protaetia coeruleosignata: Schoch 1896: 56.</p><p>Protaetia coeruleosignata flavonotata Mikšić, 1963a: 396 .</p><p>Protaetia (Protaetia) coeruleosignata: Mikšić 1963b: 420.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Surigao del Norte) (Mohnike 1873; Mikšić 1963a, 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692552852FF14FDD8FCDCFC83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FC15FCDCFA89.text	3C6E87D692552852FF14FC15FCDCFA89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) compacta (Mohnike 1873)	<div><p>Protaetia (Protaetia) compacta (Mohnike, 1873)</p><p>Cetonia compacta Mohnike, 1873: 234 .</p><p>Protaetia compacta: Schoch 1896: 56.</p><p>Protaetia talaurensis Moser, 1914: 583 . Type locality. “Ins. Talaur.”</p><p>Protaetia bakeri Moser, 1917: 13 . Type locality. “ Mindanao (Davao).”</p><p>Protaetia pseudobakeri Mikšić, 1963a: 398 .</p><p>Protaetia mindanaona Mikšić, 1963b: 420 .</p><p>Protaetia (Protaetia) compacta: Mikšić 1963b: 420; Mikšić 1983: 386; Pavićević 1988: 173.</p><p>Type locality. “ Camiguin prope Mindanao.”</p><p>Distribution in the Philippines. Luzon (Romblon), Visayas (Negros Oriental and Samar), and Mindanao (Bukidnon, Misamis Oriental, Camiguin, Davao, Surigao del Norte) (Mohnike 1873; Moser 1914, 1917; Mikšić 1963a, 1963b, 1983; Pavićević 1988).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692552852FF14FC15FCDCFA89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14FAE3FCDCF9FE.text	3C6E87D692552852FF14FAE3FCDCF9FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) dubitativa Arnaud 2004	<div><p>Protaetia (Protaetia) dubitativa Arnaud, 2004</p><p>Protaetia (Protaetia) dubitativa Arnaud, 2004: 3 .</p><p>Type locality. “ Philippines, Oriental Negros.”</p><p>Distribution in the Philippines. Luzon (Romblon) and Visayas (Negros Oriental) (Arnaud 2004).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692552852FF14FAE3FCDCF9FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552852FF14F9C8FCDCF89F.text	3C6E87D692552852FF14F9C8FCDCF89F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) ducalis (Mohnike 1873)	<div><p>Protaetia (Protaetia) ducalis (Mohnike, 1873)</p><p>Cetonia ducalis Mohnike, 1873: 196 .</p><p>Protaetia (Protaetia) ducalis: Mikšić 1963a: 387; Mikšić 1963b: 419; Pavićević 1988: 173.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Aurora, Benguet, Babuyan Island, Laguna, Mountain Province, Rizal, and Sorsogon) (Mohnike 1873; Mikšić 1963a, 1963b; Pavićević 1988).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692552852FF14F9C8FCDCF89F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692552855FF14F8E9FCC6FF68.text	3C6E87D692552855FF14F8E9FCC6FF68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) elizabethae Pavicevic 1988	<div><p>Protaetia (Protaetia) elizabethae Pavićević, 1988</p><p>Protaetia (Protaetia) elizabethae Pavićević, 1988: 174 .</p><p>Type locality. “ Negros Oriental.”</p><p>Distribution in the Philippines. Visayas (Negros Oriental) (Pavićević 1988).</p><p>Overall Distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692552855FF14F8E9FCC6FF68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FF43FCDCFE51.text	3C6E87D692522855FF14FF43FCDCFE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) enorae Arnaud 2019	<div><p>Protaetia (Protaetia) enorae Arnaud, 2019</p><p>Protaetia (Protaetia) enorae Arnaud, 2019: 4 .</p><p>Type locality. “ Philippines, Mindanao, Basilan Isld.”</p><p>Distribution in the Philippines. Mindanao (Basilan) (Arnaud 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692522855FF14FF43FCDCFE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FE28FCDCFD86.text	3C6E87D692522855FF14FE28FCDCFD86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) evae Arnaud 2019	<div><p>Protaetia (Protaetia) evae Arnaud, 2019</p><p>Protaetia (Protaetia) evae Arnaud, 2019: 4 .</p><p>Type locality. “ Mindoro, Mt. Halcon. ”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Arnaud 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692522855FF14FE28FCDCFD86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FD10FCDCFCEF.text	3C6E87D692522855FF14FD10FCDCFCEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) evangelinae Arnaud 2000	<div><p>Protaetia (Protaetia) evangelinae Arnaud, 2000</p><p>Protaetia (Protaetia) evangelinae Arnaud, 2000: 5 .</p><p>Type locality. “ Batanes Prov., Camiguin Isld.”</p><p>Distribution in the Philippines. Luzon (Babuyan Island) (Arnaud 2000).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692522855FF14FD10FCDCFCEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FCF9FCDCFA65.text	3C6E87D692522855FF14FCF9FCDCFA65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) ferruginea (Gory & Percheron 1833)	<div><p>Protaetia (Protaetia) ferruginea (Gory &amp; Percheron, 1833)</p><p>Cetonia ferruginea Gory &amp; Percheron, 1833: 196; Blanchard 1842: 5; Wallace 1868: 582; Gemminger &amp; Harold 1869: 1323; Mohnike 1873: 193.</p><p>Protaetia ferruginea: Burmeister 1842: 491; Schoch 1895: 117; Schoch 1896: 57.</p><p>Cetonia sybaritica Newman, 1841: 170 . Type locality. “ Philippines Islands.”</p><p>Protaetia cinnamomea Burmeister, 1842: 491 . Type locality. “Luzon.”</p><p>Protaetia sanguinolenta Heller, 1899a: 4 . Type locality. “ Philippinen.”</p><p>Pachnoda cupriventris Schürhoff, 1938: 191 . Type locality. “ Joko, Kamerun.”</p><p>Protaetia ferruginea ornatissima Mikšić, 1974c: 236 . Type locality. “ Philippines, Torong Corong.”</p><p>Protaetia (Protaetia) ferruginea: Mikšić 1963a: 411; Mikšić 1963b: 423; Mikšić 1983: 387; Mikšić 1984: 303.</p><p>Type locality. “Des Indes orientalis.”</p><p>Distribution in the Philippines. Luzon (Abra, Albay, Cagayan Province, Camarines Sur, Ilocos, Laguna,? Palawan, Nueva Vizcaya, and Romblon), Visayas (Bohol, Leyte, Cebu, Panay, and Negros Occidental), and Mindanao (Gory &amp; Percheron 1833; Newman 1841; Burmeister 1842; Wallace 1868; Dohrn 1872; Mohnike 1873; Heller 1899a; Schürhoff 1938; Mikšić 1963a, 1963b, 1974c, 1984, 1986).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692522855FF14FCF9FCDCFA65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522855FF14FA74FCDCF94A.text	3C6E87D692522855FF14FA74FCDCF94A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) francisi Arnaud 1989	<div><p>Protaetia (Protaetia) francisi Arnaud, 1989</p><p>Protaetia (Protaetia) francisi Arnaud, 1989a: 2 .</p><p>Type locality. “ Philippines, Batanes Island.”</p><p>Distribution in the Philippines. Luzon (Batanes) (Arnaud 1989a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692522855FF14FA74FCDCF94A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692522854FF14F95CFCD3FF0C.text	3C6E87D692522854FF14F95CFCD3FF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) fusca (Herbst 1790)	<div><p>Protaetia (Protaetia) fusca (Herbst, 1790)</p><p>Cetonia fusca Herbst, 1790: 257; Wallace 1868: 584 (= Cetonia madarinea Lichtenstein, 1796).</p><p>Protaetia fusca: Mikšić 1962: 33.</p><p>Protaetia (Heteroprotaetia) fusca: Mikšić 1963a: 356; Mikšić 1963b: 416; Mikšić 1983: 386; Mikšić 1987: 394.</p><p>Type locality. “Das Vaterland ist unbekannt.”</p><p>Distribution in the Philippines. Luzon (Laguna, Manila, Palawan, Rizal, Isabela, and Romblon), Visayas (Antique, Negros Occidental, Panay, and Samar), and Mindanao (Bukidnon, Zamboanga, Dinagat, and Lanao del Norte) (Wallace 1868; Mikšić 1962, 1963a, 1963b, 1983).</p><p>Overall distribution. This species is widely distributed in the Oriental, Australia, Pacific, and Nearctic regions (Woodruff, 2006; Perissinotto &amp; Clennell, 2021).</p></div>	https://treatment.plazi.org/id/3C6E87D692522854FF14F95CFCD3FF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FE9EFCDCFD5C.text	3C6E87D692532854FF14FE9EFCDCFD5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) guerini (Eydoux 1839)	<div><p>Protaetia (Protaetia) guerini (Eydoux, 1839)</p><p>Cetonia guerini Eydoux, 1839: 265; Burmeister 1842: 794.</p><p>Cetonia rogeri Burmeister, 1842: 796 .</p><p>Protaetia viridana Moser, 1909: 317 . Type locality. “ Philippinen.”</p><p>Protaetia guerini: Eydoux &amp; Souleyet 1841: 304; Mohnike 1873: 215; Schoch 1896: 57.</p><p>Protaetia (Protaetia) guerini: Mikšić 1963a: 405; Mikšić 1963b: 421.</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Abra, Benguet, Rizal, Manila, Ilocos, Camarines Sur, and Nueva Vizcaya) and Mindanao (Mt. Apo) (Eydoux 1839; Burmeister 1842; 1847; Mohnike 1873; Moser 1909; Mikšić 1963a, 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692532854FF14FE9EFCDCFD5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FD2EFCDCFC58.text	3C6E87D692532854FF14FD2EFCDCFC58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) hiekeiana Miksic 1970	<div><p>Protaetia (Protaetia) hiekeiana Mikšić, 1970</p><p>Protaetia (Protaetia) cupriceps Mikšić, 1963a: 377 (nec Moser, 1917).</p><p>Protaetia (Protaetia) hiekeiana Mikšić, 1970a: 57 .</p><p>Type locality. “Luzon-Imugan.”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Mikšić 1963a, 1970a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692532854FF14FD2EFCDCFC58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FC32FCDCFBA4.text	3C6E87D692532854FF14FC32FCDCFBA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) igorota Schultze 1916	<div><p>Protaetia (Protaetia) igorota Schultze, 1916</p><p>Protaetia igorota Schultze, 1916b: 294 .</p><p>Protaetia (Protaetia) igorota: Mikšić 1963a: 402; Mikšić 1963b: 421.</p><p>Type locality. “ Luzon, Benguet, Berg Santo Tomas.”</p><p>Distribution in the Philippines. Luzon (Benguet) (Schultze 1916b; Mikšić 1963a, 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692532854FF14FC32FCDCFBA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FB36FCDCFA44.text	3C6E87D692532854FF14FB36FCDCFA44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) incerta (Mohnike 1873)	<div><p>Protaetia (Protaetia) incerta (Mohnike, 1873)</p><p>Cetonia incerta Mohnike, 1873: 239; Mikšić 1963a: 441; Mikšić 1987: 567.</p><p>Protaetia incerta: Schoch 1896: 57.</p><p>Protaetia (Protaetia) incerta: Krajčik 1998: 46.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Mohnike 1873).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692532854FF14FB36FCDCFA44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14FA56FCDCF9AC.text	3C6E87D692532854FF14FA56FCDCF9AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) ismaeli Arnaud 1987	<div><p>Protaetia (Protaetia) ismaeli Arnaud, 1987</p><p>Protaetia (Protaetia) ismaeli Arnaud, 1987a: 27; Arnaud 1989b: 24.</p><p>Type locality. “ Philippines, Negros Oriental.”</p><p>Distribution in the Philippines. Visayas (Negros Oriental) (Arnaud 1987a, 1989b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692532854FF14FA56FCDCF9AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532854FF14F93EFCDCF894.text	3C6E87D692532854FF14F93EFCDCF894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) johani Arnaud 1992	<div><p>Protaetia (Protaetia) johani Arnaud, 1992</p><p>Protaetia (Protaetia) johani Arnaud, 1992: 30 .</p><p>Type locality. “ Philippines, North Province. ”</p><p>Distribution in the Philippines. Luzon (Ifugao) (Arnaud 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692532854FF14F93EFCDCF894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692532857FF14F8E6FCDCFF0C.text	3C6E87D692532857FF14F8E6FCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) labarosae Arnaud 2000	<div><p>Protaetia (Protaetia) labarosae Arnaud, 2000</p><p>Protaetia (Protaetia) labarosae Arnaud, 2000: 5 .</p><p>Type locality. “ Philippines, Cuyo Isld.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Arnaud 2000).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692532857FF14F8E6FCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FE9EFCDCFE2C.text	3C6E87D692502857FF14FE9EFCDCFE2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) leucogramma (Mohnike 1873)	<div><p>Protaetia (Protaetia) leucogramma (Mohnike, 1873)</p><p>Cetonia leucogramma Mohnike, 1873: 201; Mikšić 1963a: 442; Mikšić 1987: 568.</p><p>Protaetia leucogramma: Schoch 1896: 57.</p><p>Protaetia (Protaetia) leucogramma: Krajčik,1998: 46.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Mohnike, 1873).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692502857FF14FE9EFCDCFE2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FDBEFCDCFC84.text	3C6E87D692502857FF14FDBEFCDCFC84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) lineata (Mohnike 1873)	<div><p>Protaetia (Protaetia) lineata (Mohnike, 1873)</p><p>Cetonia lineata Mohnike, 1873: 204; Mikšić 1963a: 443.</p><p>Protaetia lineata: Schoch 1896: 57.</p><p>Protaetia decorata Mikšić, 1962: 6 . Type locality. “Mindanao Buk, Lindabon.”; Arnaud 1992: 32.</p><p>Protaetia (Kuytenia) decorata: Mikšić 1963b: 424.</p><p>Protaetia (Protaetia) lineata: Krajčik 1998: 46.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon and Surigao) (Mohnike 1873; Mikšić 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692502857FF14FDBEFCDCFC84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FC16FCDCFBEC.text	3C6E87D692502857FF14FC16FCDCFBEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) lydieorum Arnaud 2019	<div><p>Protaetia (Protaetia) lydieorum Arnaud, 2019</p><p>Protaetia (Protaetia) lydieorum Arnaud, 2019: 5 .</p><p>Type locality. “ Philippines, Mindanao, South Cotabato.”</p><p>Distribution in the Philippines. Mindanao (South Cotabato) (Arnaud 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692502857FF14FC16FCDCFBEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FBFEFCDCFAE8.text	3C6E87D692502857FF14FBFEFCDCFAE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) mindoroensis Kraatz 1894	<div><p>Protaetia (Protaetia) mindoroensis Kraatz, 1894</p><p>Protaetia mindoroensis Kraatz, 1894b: 298 .</p><p>Protaetia (Protaetia) mindoroensis: Mikšić 1963a: 392.</p><p>Type locality. “ Insel Mindoro.”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Kraatz 1894b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692502857FF14FBFEFCDCFAE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14FAC2FC27F988.text	3C6E87D692502857FF14FAC2FC27F988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) nocturna subsp. banksi Moser 1911	<div><p>Protaetia (Protaetia) nocturna banksi Moser, 1911</p><p>Protaetia banksi Moser, 1911a: 332 .</p><p>Protaetia (Protaetia) nocturna banksi: Mikšić 1963a: 408; Mikšić 1983: 386; Pavićević 1988: 173.</p><p>Type locality. “Negros Occidental, Bago.”</p><p>Distribution in the Philippines. Luzon (Romblon) and Visayas (Negros Island and Panay) (Moser 1911; Mikšić 1963 a, 1983; Pavićević 1988).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692502857FF14FAC2FC27F988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502857FF14F9E2FC27F894.text	3C6E87D692502857FF14F9E2FC27F894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) nocturna subsp. nocturna Moser 1909	<div><p>Protaetia (Protaetia) nocturna nocturna Moser, 1909</p><p>Protaetia nocturna Moser, 1909: 318 .</p><p>Protaetia (Protaetia) nocturna: Mikšić 1963b: 421.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Visayas ( ?Negros) (Moser 1909; Mikšić 1963b).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692502857FF14F9E2FC27F894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692502856FF14F8E6FCDCFF21.text	3C6E87D692502856FF14F8E6FCDCFF21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) pauperata (Haller 1884)	<div><p>Protaetia (Protaetia) pauperata (Haller, 1884)</p><p>Cetonia pauperata Haller, 1884: 33; Mikšić 1963a: 446; Mikšić 1987: 574.</p><p>Protaetia (Protaetia) pauperata: Krajčik 1998: 46.</p><p>Type locality. “ Philippinen (?).”</p><p>Distribution in the Philippines. Philippine archipelago (Haller 1884).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692502856FF14F8E6FCDCFF21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14FEBBFCDCFE16.text	3C6E87D692512856FF14FEBBFCDCFE16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) pavicevici Arnaud 1992	<div><p>Protaetia (Protaetia) pavicevici Arnaud, 1992</p><p>Protaetia (Protaetia) pavicevici Arnaud, 1992: 29 .</p><p>Type locality. “ Philippines, Panay, Antique.”</p><p>Distribution in the Philippines. Visayas (Negros and Panay) (Arnaud 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692512856FF14FEBBFCDCFE16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14FE60FCDCFD37.text	3C6E87D692512856FF14FE60FCDCFD37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) plebeja (Mohnike 1873)	<div><p>Protaetia (Protaetia) plebeja (Mohnike, 1873)</p><p>Cetonia plebeja Mohnike, 1873: 202; Mikšić 1963a: 447; Mikšić 1987: 575.</p><p>Protaetia plebeja: Schoch 1896: 58.</p><p>Protaetia (Protaetia) plebeja: Krajčik 1998: 46.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Bataan) (Mohnike 1873).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692512856FF14FE60FCDCFD37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14FC81FCDCFC30.text	3C6E87D692512856FF14FC81FCDCFC30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) querula (Newman 1841)	<div><p>Protaetia (Protaetia) querula (Newman, 1841)</p><p>Cetonia querula Newman, 1841: 171; Mohnike 1873: 208; Mikšić 1963a: 447; Mikšić 1987: 577.</p><p>Protaetia (Protaetia) tenuicollis: Krajčik 1998: 46.</p><p>Type locality. “ Philippines Islands.”</p><p>Distribution in the Philippines. Philippine archipelago (Newman 1841).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692512856FF14FC81FCDCFC30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14FB8AFC8BFA39.text	3C6E87D692512856FF14FB8AFC8BFA39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) scepsia (Dohrn 1872)	<div><p>Protaetia (Protaetia) scepsia (Dohrn, 1872)</p><p>Cetonia (Protaetia) scepsia Dohrn, 1872: 157 .</p><p>Cetonia moerens Mohnike, 1873: 237 . Type locality. “Mindanao.”; Mikšić 1963a: 444.</p><p>Cetonia satrapa Mohnike, 1873: 222 Type locality. “ Camiguin de Mindanao.”</p><p>Protaetia satrapa: Schoch 1895: 115.</p><p>Protaetia albomaculata Moser, 1914: 589 . Type locality. “ Mindanao (Davao).”</p><p>Protaetia scepsia aethiops Mikšić, 1963a: 373 .</p><p>Protaetia (Protaetia) scepsia: Mikšić 1963b: 417; Mikšić 1983: 386; Bezděk &amp; Hájek 2010b: 634.</p><p>Type locality. “ Manila.”</p><p>Distribution in the Philippines. Luzon (Camarines Sur, Manila, and Mt. Banahao), Visayas (Samar and Leyte), and Mindanao (Agusan, Basilan, Bukidnon, Camiguin, Davao, Dinagat, Lanao del Norte, Surigao, and Zamboanga del Norte) (Dohrn 1872; Mohnike 1873; Moser 1914; Mikšić 1963a, 1963b, 1983).</p><p>Overall distribution. The Philippines and? Vietnam.</p></div>	https://treatment.plazi.org/id/3C6E87D692512856FF14FB8AFC8BFA39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512856FF14F990FCDCF96E.text	3C6E87D692512856FF14F990FCDCF96E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) soulai Arnaud 2004	<div><p>Protaetia (Protaetia) soulai Arnaud, 2004</p><p>Protaetia (Protaetia) soulai Arnaud, 2004: 2 .</p><p>Type locality. “ Philippines, Leyte.”</p><p>Distribution in the Philippines. Visayas (Leyte and Samar) (Arnaud 2004).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692512856FF14F990FCDCF96E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692512869FF14F978FCDCFE98.text	3C6E87D692512869FF14F978FCDCFE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) subviridis (Newman 1841)	<div><p>Protaetia (Protaetia) subviridis (Newman, 1841)</p><p>Cetonia subviridis Newman, 1841: 170 .</p><p>Protaetia subviridis: Schoch 1896: 59.</p><p>Cetonia germana Newman, 1841: 170 . Type locality. “ Philippines Islands.”</p><p>Cetonia manillarum Chevrolat, 1841: 223 Type locality. “ Manille.”; Burmeister 1842: 497 ( Protaetia); Blanchard 1842: 5; Mohnike 1873: 227.</p><p>Protaetia chlorotica Burmeister, 1842: 500 . Type locality. “Luzon.”; Mohnike 1873: 224; Schoch 1896: 56.</p><p>Protaetia (Protaetia) subviridis: Mikšić 1963a: 382; Mikšić 1963b: 419.</p><p>Type locality. “ Philippines Islands.”</p><p>Distribution in the Philippines. Luzon (Zambales, Bataan, Baguio, Isabela, and Rizal) (Chevrolat 1841; Newman 1841; Burmeister 1842; Mohnike 1873; Mikšić 1963a, 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692512869FF14F978FCDCFE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FEF2FCDCFDC0.text	3C6E87D6926E2869FF14FEF2FCDCFDC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) talicici Arnaud 2000	<div><p>Protaetia (Protaetia) talicici Arnaud, 2000</p><p>Protaetia (Protaetia) talicici Arnaud, 2000: 4 .</p><p>Type locality. “ Philippines, Batanes Prov., Camiguin Isld.”</p><p>Distribution in the Philippines. Luzon (Camiguin de Luzon) (Arnaud 2000).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926E2869FF14FEF2FCDCFDC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FDDAFCDCFCE0.text	3C6E87D6926E2869FF14FDDAFCDCFCE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) tenuicollis (Mohnike 1873)	<div><p>Protaetia (Protaetia) tenuicollis (Mohnike, 1873)</p><p>Cetonia tenuicollis Mohnike, 1873: 236; Mikšić 1963a: 448; Mikšić 1987: 580.</p><p>Protaetia tenuicollis: Schoch 1896: 59.</p><p>Protaetia (Protaetia) tenuicollis: Krajčik 1998: 46.</p><p>Type locality. “ Camiguin de Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Camiguin) (Mohnike 1873).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926E2869FF14FDDAFCDCFCE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FCFAFCDCFBEC.text	3C6E87D6926E2869FF14FCFAFCDCFBEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetia) vermifer Haller 1884	<div><p>Protaetia (Protaetia) vermifer Haller, 1884</p><p>Cetonia vermifer Haller, 1884: 32; Mikšić 1963a: 448; Mikšić 1987: 581.</p><p>Protaetia (Protaetia) vermifer: Krajčik 1998: 47.</p><p>Type locality. “ Philippines (?).”</p><p>Distribution in the Philippines. Philippine archipelago (Haller 1884).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926E2869FF14FCFAFCDCFBEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FB36FCDCFA8C.text	3C6E87D6926E2869FF14FB36FCDCFA8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetiola) brookiana Jakl 2022	<div><p>Protaetia (Protaetiola) brookiana Jákl, 2022</p><p>Protaetia (Protaetiola) brookiana Jákl, 2022: 99 .</p><p>Type locality. “ Philippines, South Palawan, near Brooks Point.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Jákl 2022).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926E2869FF14FB36FCDCFA8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14FA1EFCC6F9F4.text	3C6E87D6926E2869FF14FA1EFCC6F9F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetiola) delponti Arnaud 2004	<div><p>Protaetia (Protaetiola) delponti Arnaud, 2004</p><p>Protaetia (Protaetiola) delponti Arnaud, 2004: 3 .</p><p>Type locality. “ Philippines, Oriental Negros.”</p><p>Distribution in the Philippines. Visayas (Negros Oriental) (Arnaud 2004).</p><p>Overall Distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926E2869FF14FA1EFCC6F9F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2869FF14F9C6FCC6F8DC.text	3C6E87D6926E2869FF14F9C6FCC6F8DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetiola) mohagani Legrand & Chew Kea Foo 2004	<div><p>Protaetia (Protaetiola) mohagani Legrand &amp; Chew Kea Foo, 2004</p><p>Protaetia (Protaetiola) mohagani Legrand &amp; Chew Kea Foo, 2004: 4; Jákl 2022: 102.</p><p>Type locality. “ Philippines, Negros Occidental, Mont Canla-on.”</p><p>Distribution in the Philippines. Visayas (Negros Island) (Legrand &amp; Chew Kea Foo 2004).</p><p>Overall Distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926E2869FF14F9C6FCC6F8DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926E2868FF14F8AEFCC6FF20.text	3C6E87D6926E2868FF14F8AEFCC6FF20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetia (Protaetiola) multoguttulata (Mohnike 1873)	<div><p>Protaetia (Protaetiola) multoguttulata (Mohnike, 1873)</p><p>Cetonia multoguttulata Mohnike, 1873: 185 .</p><p>Protaetia (Protaetiola) multoguttulata: Mikšić 1963a: 360; Mikšić 1963b: 416; Mikšić 1987: 474; Jákl 2022: 102.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Ilocos, Rizal, and Mt. Banahao) (Mohnike 1873; Mikšić 1963b; Jákl 2022).</p><p>Overall Distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926E2868FF14F8AEFCC6FF20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FEF3FE04FCCC.text	3C6E87D6926F2868FF14FEF3FE04FCCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Protaetiomorpha flavovariegata (Mohnike 1873)	<div><p>Protaetiomorpha flavovariegata (Mohnike, 1873)</p><p>Cetonia flavovariegata Mohnike, 1873: 205; Mikšíc 1987: 563.</p><p>Protaetia flavovariegata: Schoch 1896: 56.</p><p>Protaetiomorpha flavovariegata: Krajčík 1998: 48.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Mohnike, 1873).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks. Jákl (2024) argued that Krajčík (1998) may have “incorrectly” placed Protaetiomorpha flavovariegata (Mohnike, 1873) in this genus because it had been historically placed in the genus Prota etia Burmeister, 1842. It was never listed in Protaetiomorpha Mikšíc, 1968, by any other author and is currently regarded as incertae sedis due to the absence of Mohnike’s type. Therefore, its generic status remains dubious.</p></div>	https://treatment.plazi.org/id/3C6E87D6926F2868FF14FEF3FE04FCCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FC4EFCDCFB1C.text	3C6E87D6926F2868FF14FC4EFCDCFB1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Euglyptica) aterrima (Miksic 1979)	<div><p>Reineria (Euglyptica) aterrima (Mikšić, 1979)</p><p>Euglypta aterrima Mikšić, 1979: 33; Mikšić 1982b: 103; Pavićević 1988: 171.</p><p>Astraea (Euglypta) aterrima: Mikšić 1982a: 329.</p><p>Type locality. “Mindanao Buk. Lindabon.”</p><p>Distribution in the Philippines. Visayas (Negros Oriental) and Mindanao (Bukidnon) (Mikšić 1979; Pavićević 1988).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926F2868FF14FC4EFCDCFB1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FB6EFCDCFA18.text	3C6E87D6926F2868FF14FB6EFCDCFA18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Euglyptica) attenuata (Mohnike 1873)	<div><p>Reineria (Euglyptica) attenuata (Mohnike, 1873)</p><p>Euglypta attenuata Mohnike, 1873: 177; Schoch 1896: 59; Mikšić 1979: 30.</p><p>Astraea (Euglypta) attenuata: Mikšić 1982a: 331.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Mohnike 1873).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926F2868FF14FB6EFCDCFA18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14FA72FCDCF964.text	3C6E87D6926F2868FF14FA72FCDCF964.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Euglyptica) bakeri (Moser 1925)	<div><p>Reineria (Euglyptica) bakeri (Moser, 1925)</p><p>Euglypta bakeri Moser, 1925: 69; Mikšić 1979: 29.</p><p>Astraea (Euglypta) bakeri: Mikšić 1982a: 329.</p><p>Type locality. “Negros, Occidental Negros, Cuernos Mountains.”</p><p>Distribution in the Philippines. Visayas (Negros Occidental) (Moser 1925).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926F2868FF14FA72FCDCF964	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926F2868FF14F976FCDCF860.text	3C6E87D6926F2868FF14F976FCDCF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Euglyptica) biplagiata (Mohnike 1873)	<div><p>Reineria (Euglyptica) biplagiata (Mohnike, 1873)</p><p>Euglypta biplagiata Mohnike, 1873: 179; Schoch 1896: 59.</p><p>Astraea (Euglypta) biplagiata: Mikšić 1982a: 333.</p><p>Type locality. “ Isabela Archipelagi Sulu.”</p><p>Distribution in the Philippines. Mindanao (Sulu Archipelago) (Mohnike 1873).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926F2868FF14F976FCDCF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FF0BFCDCFEBD.text	3C6E87D6926C286BFF14FF0BFCDCFEBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Euglyptica) coeruleosignata (Miksic 1979)	<div><p>Reineria (Euglyptica) coeruleosignata (Mikšić, 1979)</p><p>Euglypta coeruleosignata Mikšić, 1979: 31 .</p><p>Astraea (Euglypta) coeruleosignata: Mikšić 1982a: 325.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Luzon (Mikšić 1979).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926C286BFF14FF0BFCDCFEBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FE0FFCDCFDE5.text	3C6E87D6926C286BFF14FE0FFCDCFDE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Euglyptica) grimaulti (Arnaud 1996)	<div><p>Reineria (Euglyptica) grimaulti (Arnaud, 1996)</p><p>Euglypta grimaulti Arnaud, 1996: 5 .</p><p>Type locality. “ Philippines, Samar.”</p><p>Distribution in the Philippines. Visayas (Leyte and Samar) (Arnaud 1996).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926C286BFF14FE0FFCDCFDE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FDF7FCDCFCE1.text	3C6E87D6926C286BFF14FDF7FCDCFCE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Euglyptica) kuntzeni (Schurhoff 1933)	<div><p>Reineria (Euglyptica) kuntzeni (Schürhoff, 1933)</p><p>Euglypta kuntzeni Schürhoff, 1933c: 102; Mikšić 1979: 29.</p><p>Astraea (Euglypta) kuntzeni: Mikšić 1982a: 326.</p><p>Type locality. “Luzon, Imugan.”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Schürhoff 1933c; Mikšić 1979).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926C286BFF14FDF7FCDCFCE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FCFBFCDCFBA5.text	3C6E87D6926C286BFF14FCFBFCDCFBA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Euglyptica) megaspilota (Wallace 1867)	<div><p>Reineria (Euglyptica) megaspilota (Wallace, 1867)</p><p>Cetonia megaspilota Wallace, 1867: 97; Wallace 1868: 587.</p><p>Euglypta megaspilota: Mohnike 1873: 176; Schoch 1896: 59; Mikšić 1979: 29.</p><p>Astraea (Euglypta) megaspilota: Mikšić 1982a: 323.</p><p>Type locality. “ Philippine Islands</p><p>Distribution in the Philippines. Luzon (Laguna and Quezon Province) and Mindanao (Wallace 1867; Mohnike 1873).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926C286BFF14FCFBFCDCFBA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FB37FCDCFA8D.text	3C6E87D6926C286BFF14FB37FCDCFA8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Euglyptica) mineti (Arnaud 1982)	<div><p>Reineria (Euglyptica) mineti (Arnaud, 1982)</p><p>Euglypta mineti Arnaud, 1982a: 118 .</p><p>Type locality. “ Ile de Romblon, Philippines.”</p><p>Distribution in the Philippines. Luzon (Romblon) (Arnaud 1982a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926C286BFF14FB37FCDCFA8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14FA1FFCDCF989.text	3C6E87D6926C286BFF14FA1FFCDCF989.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Euglyptica) multoguttata (Mohnike 1873)	<div><p>Reineria (Euglyptica) multoguttata (Mohnike, 1873)</p><p>Euglypta multoguttata Mohnike, 1873: 181; Schoch 1896: 59.</p><p>Astraea (Euglypta) multoguttata: Mikšić 1982a: 334.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Mohnike 1873).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926C286BFF14FA1FFCDCF989	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286BFF14F9E3FCC7F895.text	3C6E87D6926C286BFF14F9E3FCC7F895.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Euglyptica) romblonica (Miksic 1982)	<div><p>Reineria (Euglyptica) romblonica (Mikšić, 1982)</p><p>Euglypta romblonica Mikšić, 1982b: 101 .</p><p>Astraea (Euglypta) romblonica: Mikšić 1982a: 327; Mikšić 1983: 393.</p><p>Type locality. “ Insel Romblon (Philippinen).”</p><p>Distribution in the Philippines. Luzon (Romblon) (Mikšić 1982b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926C286BFF14F9E3FCC7F895	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926C286AFF14F8E7FCDCFF20.text	3C6E87D6926C286AFF14F8E7FCDCFF20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Euglyptica) schaueri (Schurhoff 1933)	<div><p>Reineria (Euglyptica) schaueri (Schürhoff, 1933)</p><p>Euglypta schaueri Schürhoff, 1933a: 32; Mikšić 1979: 30.</p><p>Astraea (Euglypta) schaueri: Mikšić 1982a: 332.</p><p>Type locality. “ Baguio ( Luzon).”</p><p>Distribution in the Philippines. Luzon (Baguio) (Schürhoff 1933a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926C286AFF14F8E7FCDCFF20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FEF2FCC6FDC0.text	3C6E87D6926D286AFF14FEF2FCC6FDC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Reineria) expleta Legrand & Jakl 2017	<div><p>Reineria (Reineria) expleta Legrand &amp; Jákl, 2017</p><p>Reineria (Reineria) expleta Legrand &amp; Jákl, 2017: 19 .</p><p>Type locality. “ Philippines, Leyte, Mont Balocaue.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Legrand &amp; Jákl 2017).</p><p>Overall Distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926D286AFF14FEF2FCC6FDC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FDDAFCDCFCCC.text	3C6E87D6926D286AFF14FDDAFCDCFCCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Reineria) lumawigi (Arnaud 1990)	<div><p>Reineria (Reineria) lumawigi (Arnaud, 1990)</p><p>Euglypta (Astraea) lumawigi Arnaud, 1990: 8 .</p><p>Reineria (Reineria) lumawigi: Arnaud 2004: 6.</p><p>Type locality. “ Philippines, Oriental Negros.”</p><p>Distribution in the Philippines. Visayas (Arnaud 1990).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926D286AFF14FDDAFCDCFCCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FCDEFE71FB80.text	3C6E87D6926D286AFF14FCDEFE71FB80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Reineria) mucksteini Legrand & Jakl 2017	<div><p>Reineria (Reineria) mucksteini Legrand &amp; Jákl, 2017</p><p>Reineria (Reineria) mucksteini Legrand &amp; Jákl, 2017: 20 .</p><p>Type locality. “ Philippines, Mindanao, Bukidnon, Cabangsalan.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon, Davao Oriental, and South Cotabato) (Legrand &amp; Jákl 2017).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks. According to Legrand &amp; Jákl (2017), material collected from Palawan may be mislabeled “ provenance douteuse.”</p></div>	https://treatment.plazi.org/id/3C6E87D6926D286AFF14FCDEFE71FB80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FB1AFCDCFA8C.text	3C6E87D6926D286AFF14FB1AFCDCFA8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Reineria) romeoi (Arnaud 2004)	<div><p>Reineria (Reineria) romeoi (Arnaud, 2004)</p><p>Astraea (Euglypta) lumawigi Arnaud, 1992: 31 .</p><p>Reineria (Reineria) romeoi Arnaud, 2004: 6 .</p><p>Type locality. “ Philippines, Or. Negros .”</p><p>Distribution in the Philippines. Visayas (Negros Island) (Arnaud 1992, 2004).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926D286AFF14FB1AFCDCFA8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14FA1EFCC6F988.text	3C6E87D6926D286AFF14FA1EFCC6F988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria (Reineria) thieuleuxi Legrand & Flutsch 2016	<div><p>Reineria (Reineria) thieuleuxi Legrand &amp; Flutsch, 2016</p><p>Reineria (Euglyptica) thieuleuxi Legrand &amp; Flutsch, 2016: 80 .</p><p>Reineria (Reineria) thieuleuxi: Legrand &amp; Flutsch 2017: 27.</p><p>Type locality. “ Philippines, Mindanao, South Cotabato, Kidapawan.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon and South Cotabato) (Legrand &amp; Flutsch 2016).</p><p>Overall Distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926D286AFF14FA1EFCC6F988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926D286AFF14F95AFCDCF84C.text	3C6E87D6926D286AFF14F95AFCDCF84C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria alaini (Arnaud 1992)	<div><p>Reineria (sensu lato) alaini (Arnaud, 1992)</p><p>Astraea (Astraea) alaini Arnaud, 1992: 31 .</p><p>Euglypta (Reineria) alaini: Krajčik 1998: 29.</p><p>Type locality. “ Philippines, Mindanao, Surigao del Sur.”</p><p>Distribution in the Philippines. Mindanao (Surigao del Sur) (Arnaud 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926D286AFF14F95AFCDCF84C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FF0BFCDCFEBD.text	3C6E87D6926A286DFF14FF0BFCDCFEBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria benquetia (Schultze 1916)	<div><p>Reineria (sensu lato) benquetia (Schultze, 1916)</p><p>Astraea benquetia Schultze, 1916b: 293; Mikšić 1981: 231; Mikšić 1982a: 317.</p><p>Euglypta (Reineria) benguetia: Krajčik 1998: 29 (incorrectly spelled).</p><p>Type locality. “ Luzon, Benguet, Berg Santo Tomas.”</p><p>Distribution in the Philippines. Luzon (Benguet) (Schultze 1916b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926A286DFF14FF0BFCDCFEBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FE0FFCDCFD5D.text	3C6E87D6926A286DFF14FE0FFCDCFD5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria biguttulata (Mohnike 1873)	<div><p>Reineria (sensu lato) biguttulata (Mohnike, 1873)</p><p>Astraea biguttulata Mohnike, 1873: 171; Schoch 1896: 59; Mikšić 1981: 211; Mikšić 1982a: 292.</p><p>Euglypta (Reineria) biguttulata: Krajčik 1998: 29.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Quezon, Nueva Vizcaya, and Catanduanes) (Mohnike 1873; Mikšić 1981).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926A286DFF14FE0FFCDCFD5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FD2FFCDCFC59.text	3C6E87D6926A286DFF14FD2FFCDCFC59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria boudanti (Arnaud 1992)	<div><p>Reineria (sensu lato) boudanti (Arnaud, 1992)</p><p>Astraea (Astraea) boudanti Arnaud, 1992: 30 .</p><p>Euglypta (Reineria) boudanti: Krajčik 1998: 29.</p><p>Type locality. “ Philippines, Leyte.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Arnaud 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926A286DFF14FD2FFCDCFC59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FC33FCDCFBA5.text	3C6E87D6926A286DFF14FC33FCDCFBA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria cartereti (Arnaud 1987)	<div><p>Reineria (sensu lato) cartereti (Arnaud, 1987)</p><p>Astraea (Astraea) cartereti Arnaud, 1987c: 4 .</p><p>Euglypta (Reineria) cartereti: Krajčik 1998: 29.</p><p>Type locality. “ Philippines, Sibuyan Island, Romblon.”</p><p>Distribution in the Philippines. Luzon (Romblon) and Mindanao (?Surigao) (Arnaud 1987c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926A286DFF14FC33FCDCFBA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FB37FCDCFAA1.text	3C6E87D6926A286DFF14FB37FCDCFAA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria dumasi (Arnaud 1992)	<div><p>Reineria (sensu lato) dumasi (Arnaud, 1992)</p><p>Astraea (Astraea) dumasi Arnaud, 1992: 30 .</p><p>Euglypta (Reineria) dumasi: Krajčik 1998: 29.</p><p>Type locality. “ Philippines, Mindanao, Mt. Apo.”</p><p>Distribution in the Philippines. Mindanao (Arnaud 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926A286DFF14FB37FCDCFAA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14FA3BFCDCF941.text	3C6E87D6926A286DFF14FA3BFCDCF941.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria eckeli (Schurhoff 1933)	<div><p>Reineria (sensu lato) eckeli (Schürhoff, 1933)</p><p>Astraea eckeli Schürhoff, 1933a: 31; Mikšić 1981: 215; Mikšić 1982a: 298.</p><p>Euglypta (Reineria) eckeli: Krajčik 1998: 29.</p><p>Type locality. “Imugan, Luzon.”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya and Camarines Sur) (Schürhoff 1933a; Mikšić 1981).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926A286DFF14FA3BFCDCF941	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926A286DFF14F95BFCDCF84D.text	3C6E87D6926A286DFF14F95BFCDCF84D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria fasciata (Schurhoff 1935)	<div><p>Reineria (sensu lato) fasciata (Schürhoff, 1935)</p><p>Astraea fasciata Schürhoff, 1935: 26; Mikšić 1981: 219; Mikšić 1982a: 302.</p><p>Euglypta (Reineria) fasciata: Krajčik 1998: 29.</p><p>Type locality. “Imugan, Luzon.”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya and Camarines Sur) (Schürhoff 1935).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926A286DFF14F95BFCDCF84D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FF0AFC27FE2A.text	3C6E87D6926B286CFF14FF0AFC27FE2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria flavopunctata subsp. flavopunctata Blanchard 1842	<div><p>Reineria (sensu lato) flavopunctata flavopunctata Blanchard, 1842</p><p>Macronota flavopunctata Blanchard, 1842: 19; Blanchard 1850: 41; Wallace 1868: 555; Schoch 1896: 35; Mikšić 1976: 353.</p><p>Astraea tigrina Mohnike, 1873: 172 . Type locality. “Luzon et Samar.”; Schoch 1896: 59.</p><p>Astraea flavopunctata: Mikšić 1981: 224; Mikšić 1982a: 306.</p><p>Euglypta (Reineria) flavopunctata: Krajčik 1998: 29.</p><p>Type locality. “Ins. Philipp.”</p><p>Distribution in the Philippines. Luzon (Laguna, Rizal, Albay, and Polillo) and Visayas (Samar) (Blanchard 1842, 1850; Mohnike 1873; Mikšić 1981)</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926B286CFF14FF0AFC27FE2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FDBCFC27FCCB.text	3C6E87D6926B286CFF14FDBCFC27FCCB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria flavopunctata subsp. maculiventris (Heller 1899) Blanchard 1842	<div><p>Reineria (sensu lato) flavopunctata maculiventris (Heller, 1899)</p><p>Astraea tigrina maculiventris Heller, 1899a: 5 .</p><p>Astraea flavopunctata maculiventris: Mikšić 1981: 234; Mikšić 1982a: 307.</p><p>Euglypta (Reineria) flavopunctata maculiventris: Krajčik 1998: 29.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Luzon (Catanduanes) (Mikšić 1981)</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926B286CFF14FDBCFC27FCCB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FCDDFC27FBD4.text	3C6E87D6926B286CFF14FCDDFC27FBD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria francolina subsp. bulusanica (Miksic 1981)	<div><p>Reineria (sensu lato) francolina bulusanica (Mikšić, 1981)</p><p>Astraea francolina bulusanica Mikšić, 1981: 227; Mikšić 1982a: 311.</p><p>Euglypta (Reineria) francolina bulusanica: Krajčik 1998: 29.</p><p>Type locality. “Luzon: Bulusan.”</p><p>Distribution in the Philippines. Luzon (Sorsogon) (Mikšić 1981).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926B286CFF14FCDDFC27FBD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FBA6FC27FA89.text	3C6E87D6926B286CFF14FBA6FC27FA89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria francolina subsp. francolina (Burmeister 1842)	<div><p>Reineria (sensu lato) francolina francolina (Burmeister, 1842)</p><p>Cetonia francolina Burmeister, 1842: 794 .</p><p>Astraea francolina: Mohnike 1873: 168; Schoch 1896: 59 (francolinica); Mikšić 1981: 226; Mikšić 1982a: 309.</p><p>Euglypta (Reineria) francolina: Krajčik 1998: 29.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Visayas (Samar) (Burmeister 1842; Mohnike 1873).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926B286CFF14FBA6FC27FA89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14FAE3FC27F9B6.text	3C6E87D6926B286CFF14FAE3FC27F9B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria francolina subsp. knothi (Schurhoff 1933)	<div><p>Reineria (sensu lato) francolina knothi (Schürhoff, 1933)</p><p>Astraea knothi Schürhoff, 1933a: 32 .</p><p>Astraea francolina knothi: Mikšić 1981: 227; Mikšić 1982a: 313.</p><p>Euglypta (Reineria) francolina knothi: Krajčik 1998: 29.</p><p>Type locality. “ Masbate (Philippinen).”</p><p>Distribution in the Philippines. Luzon (Masbate) (Schürhoff 1933a).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926B286CFF14FAE3FC27F9B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286CFF14F900FC27F8B3.text	3C6E87D6926B286CFF14F900FC27F8B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria francolina subsp. panayana (Miksic 1981)	<div><p>Reineria (sensu lato) francolina panayana (Mikšić, 1981)</p><p>Astraea francolina panayana Mikšić, 1981: 228; Mikšić 1982a: 314.</p><p>Euglypta (Reineria) francolina panayana: Krajčik 1998: 29.</p><p>Type locality. “Panay.”</p><p>Distribution in the Philippines. Visayas (Panay) (Mikšić 1981).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926B286CFF14F900FC27F8B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6926B286FFF14F805FCDCFF20.text	3C6E87D6926B286FFF14F805FCDCFF20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria immaculata (Miksic 1982)	<div><p>Reineria (sensu lato) immaculata (Mikšić, 1982)</p><p>Astraea immaculata Mikšić, 1982b: 104; Mikšić 1982a: 297.</p><p>Euglypta (Reineria) immaculata: Krajčik 1998: 29.</p><p>Type locality. “Mindanao P. I., Bukidnon, Landabon.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon) (Mikšić 1982b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6926B286FFF14F805FCDCFF20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FEBAFCDCFE2C.text	3C6E87D69268286FFF14FEBAFCDCFE2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria luzonica (Moser 1922)	<div><p>Reineria (sensu lato) luzonica (Moser, 1922)</p><p>Astraea luzonica Moser, 1922a: 113; Mikšić 1981: 210; Mikšić 1982a: 291.</p><p>Euglypta (Reineria) luzonica: Krajčik 1998: 29.</p><p>Type locality. “Luzon (Imugan).”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1922a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69268286FFF14FEBAFCDCFE2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FDBEFCDCFCCC.text	3C6E87D69268286FFF14FDBEFCDCFCCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria margaritacea (Mohnike 1873)	<div><p>Reineria (sensu lato) margaritacea (Mohnike, 1873)</p><p>Astraea margaritacea Mohnike, 1873: 170; Schoch 1896: 59; Mikšić 1981: 229; Mikšić 1982a: 315.</p><p>Euglypta (Reineria) margaritacea: Krajčik 1998: 29.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon, Davao, Lanao del Norte, and Zamboanga) (Mohnike 1873; Mikšić 1981).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69268286FFF14FDBEFCDCFCCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FCDEFCDCFBC8.text	3C6E87D69268286FFF14FCDEFCDCFBC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria miksici (Arnaud 1992)	<div><p>Reineria (sensu lato) miksici (Arnaud, 1992)</p><p>Astraea (Astraea) miksici Arnaud, 1992: 30 .</p><p>Euglypta (Reineria) miksici: Krajčik 1998: 29.</p><p>Type locality. “ Philippines, Mindanao, Cotabato, Mt. Apo.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon, Mt. Apo, and Surigao) (Arnaud 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69268286FFF14FCDEFCDCFBC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FBA2FCDCFAD4.text	3C6E87D69268286FFF14FBA2FCDCFAD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria moseri (Bourgoin 1931)	<div><p>Reineria (sensu lato) moseri (Bourgoin, 1931)</p><p>Astraea moseri Bourgoin, 1931: 176; Mikšić 1981: 220; Mikšić 1982a: 304.</p><p>Euglypta (Reineria) moseri: Krajčik 1998: 29.</p><p>Type locality. “Luzon: Imugan.”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Bourgoin 1931).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69268286FFF14FBA2FCDCFAD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14FAA6FCDCF9F4.text	3C6E87D69268286FFF14FAA6FCDCF9F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria multimaculata (Moser 1910)	<div><p>Reineria (sensu lato) multimaculata (Moser, 1910)</p><p>Astraea multimaculata Moser, 1910: 183; Mikšić 1981: 212; Mikšić 1982a: 294.</p><p>Euglypta (Reineria) multimaculata: Krajčik 1998: 29.</p><p>Type locality. “Mindanao, Camp Keithley, P. I.”</p><p>Distribution in the Philippines. Mindanao (Lanao del Sur, Surigao, and Bukidnon) (Moser 1922a; Mikšić 1981).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69268286FFF14FAA6FCDCF9F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286FFF14F9C6FCDCF8F0.text	3C6E87D69268286FFF14F9C6FCDCF8F0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria pehrsoni (Miksic 1981)	<div><p>Reineria (sensu lato) pehrsoni (Mikšić, 1981)</p><p>Astraea pehrsoni Mikšić, 1981: 215; Mikšić 1982a: 297.</p><p>Euglypta (Reineria) pehrsoni: Krajčik 1998: 29.</p><p>Type locality. “Mt. Apo, Philippines.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Mikšić 1981).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69268286FFF14F9C6FCDCF8F0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69268286EFF14F8CAFCDCFF0D.text	3C6E87D69268286EFF14F8CAFCDCFF0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria punctiventris (Moser 1917)	<div><p>Reineria (sensu lato) punctiventris (Moser, 1917)</p><p>Astraea punctiventris Moser, 1917a: 16; Mikšić 1981: 209; Mikšić 1982a: 289.</p><p>Euglypta (Reineria) punctiventris: Krajčik 1998: 29.</p><p>Type locality. “Luzon (Imugan).”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1917a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69268286EFF14F8CAFCDCFF0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FB3BFCDCFA89.text	3C6E87D69269286EFF14FB3BFCDCFA89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes apo Krikken 1980	<div><p>Callynomes apo Krikken, 1980</p><p>Callynomes apo Krikken, 1980: 15; Kobayashi &amp; Sasaki 2016: 62; Sasaki 2019: 70.</p><p>Type locality. “Mindanao, P. I., Galag River / Mt. Apo.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Krikken 1980).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69269286EFF14FB3BFCDCFA89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FAE0FCDCF992.text	3C6E87D69269286EFF14FAE0FCDCF992.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes borealis Kobayashi & Sasaki 2016	<div><p>Callynomes borealis Kobayashi &amp; Sasaki, 2016</p><p>Callynomes borealis Kobayashi &amp; Sasaki, 2016: 50; Sasaki 2019: 70.</p><p>Type locality. “Luzon Is., Philippines.”</p><p>Distribution in the Philippines. Luzon (Aurora, Nueva Ecija, and Quirino) (Kobayashi &amp; Sasaki 2016; Sasaki 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69269286EFF14FAE0FCDCF992	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14F9E5FCDCF8FB.text	3C6E87D69269286EFF14F9E5FCDCF8FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes brevispinus Kobayashi & Sasaki 2016	<div><p>Callynomes brevispinus Kobayashi &amp; Sasaki, 2016</p><p>Callynomes brevispinus Kobayashi &amp; Sasaki, 2016: 51; Sasaki 2019: 70.</p><p>Type locality. “ N. Luzon, the Philippines.”</p><p>Distribution in the Philippines. Luzon (Aurora and Quirino) (Kobayashi &amp; Sasaki 2016; Sasaki 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69269286EFF14F9E5FCDCF8FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14F8CDFCDCF823.text	3C6E87D69269286EFF14F8CDFCDCF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes camiguinana Kobayashi & Sasaki 2016	<div><p>Callynomes camiguinana Kobayashi &amp; Sasaki, 2016</p><p>Callynomes camiguinana Kobayashi &amp; Sasaki, 2016: 53; Sasaki 2019: 69.</p><p>Type locality. “ Camiguin Is., the Philippines.”</p><p>Distribution in the Philippines. Luzon (Camiguin) (Kobayashi &amp; Sasaki 2016; Sasaki 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69269286EFF14F8CDFCDCF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FD2DFCDCFC83.text	3C6E87D69269286EFF14FD2DFCDCFC83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clinterocera bancoi Jakl 2017	<div><p>Clinterocera bancoi Jákl, 2017</p><p>Clinterocera bancoi Jákl, 2017: 6 .</p><p>Type locality. “ Palawan, Mt. Mantaling Jan. ”</p><p>Distribution in the Philippines. Luzon (Palawan) (Jákl 2017a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69269286EFF14FD2DFCDCFC83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FC1AFCDCFBE8.text	3C6E87D69269286EFF14FC1AFCDCFBE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clinterocera palawanica Jakl 2017	<div><p>Clinterocera palawanica Jákl, 2017</p><p>Clinterocera palawanica Jákl, 2017: 12 .</p><p>Type locality. “S. Palawan, Brooke’s Point.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Jákl 2017a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69269286EFF14FC1AFCDCFBE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69269286EFF14FE9FFCDCFE2A.text	3C6E87D69269286EFF14FE9FFCDCFE2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Reineria simillima (Bourgoin 1931)	<div><p>Reineria (sensu lato) simillima (Bourgoin, 1931)</p><p>Astraea simillima Bourgoin, 1931: 177; Mikšić 1981: 216; Mikšić 1982a: 299.</p><p>Astraea simillima rubra Bourgoin, 1931: 177 .</p><p>Euglypta (Reineria) simillima: Krajčik 1998: 29.</p><p>Type locality. “Luzon: Imugan.”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya and Mt. Banahao) (Bourgoin 1931; Mikšić 1981).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69269286EFF14FE9FFCDCFE2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FF0BFD59FE51.text	3C6E87D692662861FF14FF0BFD59FE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes fujiokai Sakai 1997	<div><p>Callynomes fujiokai Sakai, 1997</p><p>Callynomes fujiokai Sakai, 1997: 12; Sakai &amp; Nagai 1998: 156; Kobayashi &amp; Sasaki 2016: 62; Sasaki 2019: 70.</p><p>Type locality. “ Mt. Malindong, Panay Is., Philippines.”</p><p>Distribution in the Philippines. Luzon (Mountain Province) and Visayas (Panay) (Sakai 1997; Kobayashi &amp; Sasaki 2016; Sasaki 2019).</p><p>Overall distribution. Only in the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692662861FF14FF0BFD59FE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FE28FCDCFD86.text	3C6E87D692662861FF14FE28FCDCFD86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes kobayashii Sasaki 2019	<div><p>Callynomes kobayashii Sasaki, 2019</p><p>Callynomes kobayashii Sasaki, 2019: 67 .</p><p>Type locality. “ Mt.Barlig, Mountain Province, North Luzon, Philippines.”</p><p>Distribution in the Philippines. Luzon (Mountain Province) (Sasaki 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692662861FF14FE28FCDCFD86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FD11FCDCFCA7.text	3C6E87D692662861FF14FD11FCDCFCA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes luzonica (Schultze 1916)	<div><p>Callynomes luzonica (Schultze, 1916)</p><p>Coenochilus luzonica Schultze, 1916c: 349 .</p><p>Callynomes luzonica: Krikken 1980: 18; Kobayashi &amp; Sasaki 2016: 63; Sasaki 2019: 70.</p><p>Type locality. “ Laguna, Paete.”</p><p>Distribution in the Philippines. Luzon (Laguna, Mindoro, and Rizal) and Visayas (Samar) (Schultze 1916c; Krikken 1980; Sasaki 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692662861FF14FD11FCDCFCA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FC36FCDCFBA0.text	3C6E87D692662861FF14FC36FCDCFBA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes meridionalis Kobayashi & Sasaki 2016	<div><p>Callynomes meridionalis Kobayashi &amp; Sasaki, 2016</p><p>Callynomes meridionalis Kobayashi &amp; Sasaki, 2016: 56; Sasaki 2019: 70.</p><p>Type locality. “Mindanao Is. and Leyte Is., the Philippines.”</p><p>Distribution in the Philippines. Visayas (Leyte) and Mindanao (Agusan del Sur and Bukidnon) (Kobayashi &amp; Sasaki 2016; Sasaki 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692662861FF14FC36FCDCFBA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FB3BFCDCFA89.text	3C6E87D692662861FF14FB3BFCDCFA89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes nishiguroi Kobayashi & Sasaki 2016	<div><p>Callynomes nishiguroi Kobayashi &amp; Sasaki, 2016</p><p>Callynomes nishiguroi Kobayashi &amp; Sasaki, 2016: 57; Sasaki 2019: 69.</p><p>Type locality. “ N. Luzon, the Philippines.”</p><p>Distribution in the Philippines. Luzon (Apayao) (Kobayashi &amp; Sasaki 2016; Sasaki 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692662861FF14FB3BFCDCFA89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14FAE0FCDCF992.text	3C6E87D692662861FF14FAE0FCDCF992.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes niveoguttata Sakai 1997	<div><p>Callynomes niveoguttata Sakai, 1997</p><p>Callynomes niveoguttata Sakai, 1997: 12; Sakai &amp; Nagai 1998: 156; Kobayashi &amp; Sasaki 2016: 63; Sasaki 2019: 68.</p><p>Type locality. “ Mt. Malindong, Panay Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Panay) (Sakai 1997; Kobayashi &amp; Sasaki 2016; Sasaki 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692662861FF14FAE0FCDCF992	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662861FF14F9E5FCDCF8B3.text	3C6E87D692662861FF14F9E5FCDCF8B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes niveosparsa Mohnike 1873	<div><p>Callynomes niveosparsa Mohnike, 1873</p><p>Callynomes niveosparsa Mohnike, 1873: 241; Schoch 1896: 80 (niveisparsa); Heller 1897d: 182 (niveosparsus); Krikken 1980: 18; Kobayashi &amp; Sasaki 2016: 63; Sasaki 2019: 70.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Visayas (Leyte) and Mindanao (Surigao) (Mohnike 1873; Krikken 1980; Kobayashi &amp; Sasaki 2016).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692662861FF14F9E5FCDCF8B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692662860FF14F80AFCDCFF0D.text	3C6E87D692662860FF14F80AFCDCFF0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes palawanicus Jakl 2011	<div><p>Callynomes palawanicus Jákl, 2011</p><p>Callynomes palawanicus Jákl, 2011: 212; Kobayashi &amp; Sasaki 2016: 64; Sasaki 2019: 70.</p><p>Type locality. “ Philippines, Palawan, Mt. Salakot. ”</p><p>Distribution in the Philippines. Luzon (Palawan) (Jákl 2011; Sasaki 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692662860FF14F80AFCDCFF0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FAC7FCDCF9DA.text	3C6E87D692672860FF14FAC7FCDCF9DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anthracophora (Anthracophora) philippinica Jakl 2011	<div><p>Anthracophora (Anthracophora) philippinica Jákl, 2011</p><p>Anthracophora (Anthracophora) philippinica Jákl, 2011: 217 .</p><p>Type locality. “ Philippines, N. Luzon, Mt. Province. ”</p><p>Distribution in the Philippines. Luzon (Mountain Province) (Jákl 2011).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692672860FF14FAC7FCDCF9DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FE9FFCDCFE75.text	3C6E87D692672860FF14FE9FFCDCFE75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes parva Kobayashi & Sasaki 2016	<div><p>Callynomes parva Kobayashi &amp; Sasaki, 2016</p><p>Callynomes parva Kobayashi &amp; Sasaki, 2016: 57; Sasaki 2019: 69.</p><p>Type locality. “ Catanduanes Is., the Philippines.”</p><p>Distribution in the Philippines. Luzon (Bicol) (Kobayashi &amp; Sasaki 2016; Sasaki 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692672860FF14FE9FFCDCFE75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FE44FCDCFD7E.text	3C6E87D692672860FF14FE44FCDCFD7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes sakaii Kobayashi & Sasaki 2016	<div><p>Callynomes sakaii Kobayashi &amp; Sasaki, 2016</p><p>Callynomes sakaii Kobayashi &amp; Sasaki, 2016: 59; Sasaki 2019: 68.</p><p>Type locality. “ N. Luzon Is., the Philippines.”</p><p>Distribution in the Philippines. Luzon (Nueva Ecija, Nueva Vizcaya, Mountain Province, and Ifugao) (Kobayashi &amp; Sasaki 2016; Sasaki 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692672860FF14FE44FCDCFD7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FD49FC27FCA7.text	3C6E87D692672860FF14FD49FC27FCA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes variabilis subsp. negrosensis Kobayashi & Sasaki 2016	<div><p>Callynomes variabilis negrosensis Kobayashi &amp; Sasaki, 2016</p><p>Callynomes variabilis negrosensis Kobayashi &amp; Sasaki, 2016: 62; Sasaki 2019: 70.</p><p>Type locality. “Negros Is., the Philippines.”</p><p>Distribution in the Philippines. Visayas (Negros Island) (Kobayashi &amp; Sasaki 2016; Sasaki 2019).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692672860FF14FD49FC27FCA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672860FF14FC36FC27FB8C.text	3C6E87D692672860FF14FC36FC27FB8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callynomes variabilis subsp. variabilis Kobayashi & Sasaki 2016	<div><p>Callynomes variabilis variabilis Kobayashi &amp; Sasaki, 2016</p><p>Callynomes variabilis variabilis Kobayashi &amp; Sasaki, 2016: 61; Sasaki 2019: 70.</p><p>Type locality. “ Leyte Is., the Philippines.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Kobayashi &amp; Sasaki 2016; Sasaki 2019).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692672860FF14FC36FC27FB8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692672863FF14F895FC24FE99.text	3C6E87D692672863FF14F895FC24FE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicheros (Dicheros) bicornis subsp. ornatus Hope 1841	<div><p>Dicheros (Dicheros) bicornis ornatus Hope, 1841</p><p>Dicheros ornatus Hope, 1841: 33; Burmeister 1842: 219; Mohnike 1873: 119; Schoch 1896: 17; Mikšić 1977: 207; Pavićević 1988: 171.</p><p>Heterorhina ornata Hope, 1846: 140; Wallace 1867: 525.</p><p>Heterorhina biguttata Westwood, 1842: 141 .</p><p>Diceros samarensis Schürhoff, 1933c: 99 . Type locality. “ Samara Insel (= Samar).”</p><p>Dicheros (Dicheros) bicornis ornatus: Nagai 1985: 174.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Marinduque, Romblon, Rizal, Laguna, and Tarlac), Visayas (Bohol, Negros Oriental, and Samar), and Mindanao (Zamboanga) (Hope 1841; Westwood 1842; Mohnike 1873; Schürhoff 1933c; Mikšić 1977; Nagai 1985).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692672863FF14F895FC24FE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FEF0FC27FDCE.text	3C6E87D692642863FF14FEF0FC27FDCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicheros (Dicheros) bicornis subsp. rowelli Nagai 1985	<div><p>Dicheros (Dicheros) bicornis rowelli Nagai, 1985</p><p>Dicheros bicornis rowelli Nagai, 1985: 174 .</p><p>Type locality. “ Palawan Island.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Nagai 1985).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692642863FF14FEF0FC27FDCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FD49FAC8FC1F.text	3C6E87D692642863FF14FD49FAC8FC1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterorhina (Eoheterorrhina) lumawigi Miksic 1986	<div><p>Heterorhina (Eoheterorrhina) lumawigi Mikšić, 1986</p><p>Heterorhina (Eoheterorrhina) lumawigi Mikšić, 1986: 69 .</p><p>Type locality. “ Mindanao Prov. Marinduque.”</p><p>Distribution in the Philippines. Luzon (Marinduque) (Mikšić 1986).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks. The earlier distribution record of Heterorhina lumawigi seems confusing and is originally labeled “Mindanao, Prov. Marinduque.” However, Marinduque is not located in Mindanao but rather in Luzon.</p></div>	https://treatment.plazi.org/id/3C6E87D692642863FF14FD49FAC8FC1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FC6EFCDCFB3C.text	3C6E87D692642863FF14FC6EFCDCFB3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterorhina (Eoheterorrhina) simillima (Mohnike 1873)	<div><p>Heterorhina (Eoheterorrhina) simillima (Mohnike, 1873)</p><p>Coryphocera simillima Mohnike, 1873: 122; Schoch 1896: 18.</p><p>Heterorhina (Eoheterorrhina) simillima: Mikšić 1977: 300.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Luzon (Laguna, Camarines Sur,? Ifugao,? Mountain Province, and? Sorsogon) and Mindanao (Mohnike 1873; Mikšić 1974a, 1977).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692642863FF14FC6EFCDCFB3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14FAC7FCDCF9FE.text	3C6E87D692642863FF14FAC7FCDCF9FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterorhina (Heterorhina) kuntzeni Schurhoff 1933	<div><p>Heterorhina (Heterorhina) kuntzeni Schürhoff, 1933</p><p>Heterorhina kuntzeni Schürhoff, 1933c: 98 .</p><p>Heterorhina (Heterorhina) kuntzeni: Mikšić 1977: 324.</p><p>Type locality. “N. Palawan Bacuit.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Schürhoff 1933c; Mikšić 1977).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692642863FF14FAC7FCDCF9FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692642863FF14F9C8FCDCF823.text	3C6E87D692642863FF14F9C8FCDCF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterorhina (Heterorhina) macleayi (Kirby 1818)	<div><p>Heterorhina (Heterorhina) macleayi (Kirby, 1818)</p><p>Heterorhina macleaii Kirby, 1818: 408 ( Cetonia); Wallace 1867: 527.</p><p>Cetonia pretiosa Eschscholtz, 1822: 23 . Type locality. “ Insel Luzon (Manila).” (Ambiguous).</p><p>Heterorhina macleayi suturalis Schürhoff, 1942: 283 . Type locality. “Mindanao (Davao, Butuan).”</p><p>Coryphocera Mac Leayi: Burmeister 1842: 226 ( Mac leaji); Mohnike 1873: 121.</p><p>Heterorhina macleayi: Pavićević 1988: 171.</p><p>Heterorhina (Heterorhina) macleayi: Mikšić 1977: 329.</p><p>Type locality. “ Philippines Islands.”</p><p>Distribution in the Philippines. Luzon (Laguna and Palawan), Visayas (Bohol and Leyte), and Mindanao (Agusan del Norte, Camiguin, Lanao del Norte, Zamboanga, Cotabato, Davao, Dinagat, and Surigao) (Kirby 1818; Eschscholtz 1822; Schürhoff 1942; Mikšić 1977; Pavićević 1988).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692642863FF14F9C8FCDCF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FB37FC27FAA0.text	3C6E87D692652862FF14FB37FC27FAA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clinteria atra (Wiedemann 1823) Schurhoff 1942	<div><p>Clinteria (sensu lato) atra boettcheri Schürhoff, 1942</p><p>Clinteria imperialis boettcheri Schürhoff, 1942: 288 .</p><p>Clinteria atra boettcheri: Mikšić 1977: 91; Legrand 2022: 9.</p><p>Type locality. “ Bacuit und Binaluan ( Nord Palawan).”</p><p>Distribution in the Philippines. Luzon (Palawan) (Schürhoff 1942; Mikšić 1977).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692652862FF14FB37FC27FAA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FA3BFC7AF964.text	3C6E87D692652862FF14FA3BFC7AF964.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clinteria atra (Wiedemann 1823) Snellen Van Vollenhoven 1864	<div><p>Clinteria (sensu lato) atra egens Snellen Van Vollenhoven, 1864</p><p>Clinteria egens Snellen Van Vollenhoven, 1864: 152 .</p><p>Clinteria formosa Mohnike, 1873: 125 . Type locality. “Mindanao.”; Schoch 1896: 29.</p><p>Clinteria atra egens: Mikšić 1977: 90.</p><p>Type locality. “Borneo.”</p><p>Distribution in the Philippines. Luzon (Palawan) and Mindanao (Zamboanga del Norte, Dinagat, Bukidnon, and Agusan) (Snellen Van Vollenhoven 1864; Mohnike 1873; Mikšić 1977)</p><p>Overall distribution. The Philippines and Borneo (Mikšić 1977).</p></div>	https://treatment.plazi.org/id/3C6E87D692652862FF14FA3BFC7AF964	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14F977FCDCF84C.text	3C6E87D692652862FF14F977FCDCF84C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clinteria jakli Legrand 2022	<div><p>Clinteria (sensu lato) jakli Legrand, 2022</p><p>Clinteria jakli Legrand, 2022: 8 .</p><p>Type locality. “ Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Legrand 2022).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692652862FF14F977FCDCF84C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FF0BFCDCFE50.text	3C6E87D692652862FF14FF0BFCDCFE50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterorhina (Heterorhina) paupera (Mohnike 1873)	<div><p>Heterorhina (Heterorhina) paupera (Mohnike, 1873)</p><p>Coryphocera paupera Mohnike, 1873: 124; Schoch 1895: 17 ( pauperata); Schoch 1896: 18.</p><p>Heterorhina (Heterorhina) paupera: Mikšić 1977: 319.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Luzon (Laguna) and Mindanao (Bukidnon, Davao, Zamboanga, and Surigao) (Mohnike 1873; Mikšić 1977).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692652862FF14FF0BFCDCFE50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FE2BFCDCFD70.text	3C6E87D692652862FF14FE2BFCDCFD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterorhina (Heterorhina) schadenbergi Heller 1895	<div><p>Heterorhina (Heterorhina) schadenbergi Heller, 1895</p><p>Heterorhina schadenbergi Heller, 1895: 281 .</p><p>Coryphocera schadenbergi: Schoch 1896: 18.</p><p>Heterorhina (Heterorhina) schadenbergi: Mikšić 1977: 326.</p><p>Type locality. “Insula Busuanga.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Heller 1895).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692652862FF14FE2BFCDCFD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692652862FF14FD4BFCDCFC7C.text	3C6E87D692652862FF14FD4BFCDCFC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterorhina (Heterorhina) versicolor (Janson 1888)	<div><p>Heterorhina (Heterorhina) versicolor (Janson, 1888)</p><p>Coryphocera versicolor Janson, 1888: 207; Schoch 1896: 18.</p><p>Heterorhina (Heterorhina) versicolor: Mikšić 1977: 328.</p><p>Type locality. “ Jolo Island .”</p><p>Distribution in the Philippines. Luzon (Palawan) and Mindanao (Sulu) (Janson 1888; Mikšić 1977).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692652862FF14FD4BFCDCFC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FEBBFCDDFDC1.text	3C6E87D692622865FF14FEBBFCDDFDC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemiphaedimus inermis (Moser 1916)	<div><p>Hemiphaedimus inermis (Moser, 1916)</p><p>Phaedimus inermis Moser, 1916c: 385 .</p><p>Hemiphaedimus inermis: Mikšić 1972b: 73; Mikšić 1977: 132; Ochi &amp; Sakai 2021: 28.</p><p>Type locality. “ Imugan (Nueva Vizcaya).”</p><p>Distribution in the Philippines. Luzon (Baguio, Nueva Vizcaya, and Laguna) (Moser 1916c; Mikšić 1972b, 1977; Ochi &amp; Sakai 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692622865FF14FEBBFCDDFDC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FD13FCDCFCE1.text	3C6E87D692622865FF14FD13FCDCFCE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus asinensis Ochi & Sakai 2021	<div><p>Phaedimus asinensis Ochi &amp; Sakai, 2021</p><p>Phaedimus asinensis Ochi &amp; Sakai, 2021: 29 .</p><p>Type locality. “ Philippines, Luzon Is., Asin Hot Spring.”</p><p>Distribution in the Philippines. Luzon (Baguio) (Ochi &amp; Sakai 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692622865FF14FD13FCDCFCE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FCFBFCDCFBC9.text	3C6E87D692622865FF14FCFBFCDCFBC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus boudanti Arnaud 2000	<div><p>Phaedimus boudanti Arnaud 2000</p><p>Phaedimus boudanti Arnaud, 2000: 4; Ochi &amp; Sakai 2021: 34.</p><p>Type locality. “ Philippines, Mindoro Isld, Mt. Halcon. ”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Arnaud 2000; Ochi &amp; Sakai 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692622865FF14FCFBFCDCFBC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FBA3FCDCFB31.text	3C6E87D692622865FF14FBA3FCDCFB31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus cabrasae Ochi & Kon 2025	<div><p>Phaedimus cabrasae Ochi &amp; Kon, 2025</p><p>Phaedimus cabrasae Ochi &amp; Kon, 2025a: 59 .</p><p>Type locality. “ Philippines, Luzon Is., Bicol, Camarines, Mt. Isalog (= Mt. Isarog).”</p><p>Distribution in the Philippines. Luzon (Camarines Sur) (Ochi &amp; Kon 2025a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692622865FF14FBA3FCDCFB31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14FA8BFCDCF989.text	3C6E87D692622865FF14FA8BFCDCF989.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus cumingi (Waterhouse 1841)	<div><p>Phaedimus cumingi (Waterhouse, 1841)</p><p>Mycteristes cumingi Waterhouse, 1841a: 36; Hope 1841: 26; Wallace 1868: 523.</p><p>Phaedimus cumingi: Burmeister 1842: 176; Gerstaecker 1862: 364; Gemminger &amp; Harold 1869: 1275; Mohnike 1873: 114; Mikšić 1972b: 73; Mikšić 1977: 137; Ochi &amp; Sakai 2021: 34.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Abra, Apayao, Ifugao, Laguna, Mindoro, Mountain Province, and Sibuyan), Visayas (Negros Occidental), and Mindanao (?Mt. Apo) (Waterhouse 1841a; Gerstaecker 1862; Mikšić 1972b, 1977).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692622865FF14FA8BFCDCF989	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622865FF14F9E3FCDCF8F1.text	3C6E87D692622865FF14F9E3FCDCF8F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus ishidai Nagai 1998	<div><p>Phaedimus ishidai Nagai, 1998</p><p>Phaedimus ishidai Nagai, 1998: 2; Ochi &amp; Sakai 2021: 35.</p><p>Type locality. “Luzon Is. Baguio.”</p><p>Distribution in the Philippines. Luzon (Baguio) (Nagai, 1998).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692622865FF14F9E3FCDCF8F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692622864FF14F8CBFCDCFF21.text	3C6E87D692622864FF14F8CBFCDCFF21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus jagori Gerstaecker 1862	<div><p>Phaedimus jagori Gerstaecker, 1862</p><p>Phaedimus jagori Gerstaecker, 1862: 364; Gemminger &amp; Harold 1869: 1275; Mikšić 1972b: 73; Mikšić 1977: 141; Ochi &amp; Sakai 2021: 36.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Laguna, Camarines Sur, and Baguio), Visayas (Leyte, Negros, and Samar), and Mindanao (Gerstaecker 1862; Mikšić 1972b, 1977).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692622864FF14F8CBFCDCFF21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FEBBFCDCFE16.text	3C6E87D692632864FF14FEBBFCDCFE16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus lumawigi Arnaud 1989	<div><p>Phaedimus lumawigi Arnaud, 1989</p><p>Phaedimus lumawigi Arnaud, 1989b: 25; Ochi &amp; Sakai 2021: 36.</p><p>Type locality. “ Marinduque Is.”</p><p>Distribution in the Philippines. Luzon (Marinduque) (Arnaud 1989b; Ochi &amp; Sakai 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692632864FF14FEBBFCDCFE16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FE60FCDCFD7E.text	3C6E87D692632864FF14FE60FCDCFD7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus lydiae Arnaud 1980	<div><p>Phaedimus lydiae Arnaud, 1980</p><p>Phaedimus lydiae Arnaud, 1980: 123; Ochi &amp; Sakai 2021: 37.</p><p>Type locality. “ Iles Philippines, île Romblon.”</p><p>Distribution in the Philippines. Luzon (Romblon) (Arnaud 1980).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692632864FF14FE60FCDCFD7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FD49FCDCFCA7.text	3C6E87D692632864FF14FD49FCDCFCA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus meridionalis Nagai 1984	<div><p>Phaedimus meridionalis Nagai, 1984</p><p>Phaedimus meridionalis Nagai, 1984: 1; Ochi &amp; Sakai 2021: 37.</p><p>Type locality. “ Philippines (Palawan Island).”</p><p>Distribution in the Philippines. Luzon (Palawan) (Nagai 1984).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692632864FF14FD49FCDCFCA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FC36FCDCFB44.text	3C6E87D692632864FF14FC36FCDCFB44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus mohnikei Kraatz 1893	<div><p>Phaedimus mohnikei Kraatz, 1893</p><p>Phaedimus mohnikei Kraatz, 1893: 76; Mikšić 1972b: 73; Mikšić 1977: 139; Ochi &amp; Sakai 2021: 38.</p><p>Type locality. “Philippinen-Insel Zebu.”</p><p>Distribution in the Philippines. Luzon (Ilocos Sur, Marinduque, Quezon, Cagayan Province, Mountain Province, Laguna, Nueva Vizcaya, Manila, Rizal, Sibuyan, and Romblon), Visayas (Negros Island, Cebu, and Panay), and Mindanao (Mt. Apo) (Kraatz 1893, 1894a; Mikšić 1972b, 1977).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692632864FF14FC36FCDCFB44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FB57FCDCFA41.text	3C6E87D692632864FF14FB57FCDCFA41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus orientalis Nagai 1998	<div><p>Phaedimus orientalis Nagai, 1998</p><p>Phaedimus orientalis Nagai, 1998: 9; Ochi &amp; Sakai 2021: 39.</p><p>Type locality. “Mindanao Is., Surigao Tandag.”</p><p>Distribution in the Philippines. Mindanao (Surigao del Sur and Davao de Oro) (Nagai 1998; Ochi &amp; Sakai 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692632864FF14FB57FCDCFA41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14FA58FC27F94A.text	3C6E87D692632864FF14FA58FC27F94A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus rafaelii subsp. howdeni Arnaud 1987	<div><p>Phaedimus rafaelii howdeni Arnaud, 1987</p><p>Phaedimus howdeni Arnaud, 1987c: 4; Arnaud 1989b: 24.</p><p>Phaedimus rafaelii howdeni: Ochi &amp; Sakai 2021: 40.</p><p>Type locality. “ Philippines, Oriental Negros, Mt. Canlaou. ”</p><p>Distribution in the Philippines. Luzon (Romblon) and Visayas (Negros) (Arnaud 1987c).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692632864FF14FA58FC27F94A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632864FF14F95DFC27F8B3.text	3C6E87D692632864FF14F95DFC27F8B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus rafaelii subsp. rafaelii Miksic 1977	<div><p>Phaedimus rafaelii rafaelii Mikšić, 1977</p><p>Phaedimus rafaelii Mikšić, 1977: 142; Pavićević 1988: 171; Arnaud 1989b: 25; Ochi &amp; Sakai 2021: 39.</p><p>Type locality. “Panay, Culasi.”</p><p>Distribution in the Philippines. Visayas (Panay) (Mikšić 1977; Pavićević 1988).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692632864FF14F95DFC27F8B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692632867FF14F80AFCDCFF20.text	3C6E87D692632867FF14F80AFCDCFF20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus wittei Kraatz 1893	<div><p>Phaedimus wittei Kraatz, 1893</p><p>Phaedimus wittei Kraatz, 1893: 76; Mikšić 1972b: 74; Mikšić 1977: 140; Ochi &amp; Sakai 2021: 41.</p><p>Type locality. “Philippinen-Insel Zebu.”</p><p>Distribution in the Philippines. Luzon (Sibuyan), Visayas (Cebu), and Mindanao (Kraatz 1893; Mikšić 1972b, 1977).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692632867FF14F80AFCDCFF20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FEBBFC27FE08.text	3C6E87D692602867FF14FEBBFC27FE08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus zebuanus subsp. kachikun Nagai 1998	<div><p>Phaedimus zebuanus kachikun Nagai, 1998</p><p>Phaedimus zebuanus kachikun Nagai, 1998: 9; Ochi &amp; Sakai 2021: 42</p><p>Type locality. “Mindanao Is. Mt. Apo.”</p><p>Distribution in the Philippines. Mindanao (Davao and Bukidnon) (Nagai 1998; Ochi &amp; Sakai 2021).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692602867FF14FEBBFC27FE08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FE63FC27FD70.text	3C6E87D692602867FF14FE63FC27FD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedimus zebuanus subsp. zebuanus Miksic 1972	<div><p>Phaedimus zebuanus zebuanus Mikšić, 1972</p><p>Phaedimus zebuanus Mikšić, 1972b: 74; Mikšić 1977: 143; Ochi &amp; Sakai 2021: 41.</p><p>Type locality. “ Cebu Is.”</p><p>Distribution in the Philippines. Visayas (Cebu) and Mindanao (Mt. Apo) (Mikšić 1972b, 1977).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692602867FF14FE63FC27FD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FCFBFCDCFBC8.text	3C6E87D692602867FF14FCFBFCDCFBC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philistina (Rhinacosmus) fujiokai (Jakl 2011)	<div><p>Philistina (Rhinacosmus) fujiokai (Jákl, 2011)</p><p>Mycteristes (Rhinacosmus) fujiokai Jákl, 2011: 215 .</p><p>Type locality. “ Philippines, Basilan Island.”</p><p>Distribution in the Philippines. Mindanao (Basilan) (Jákl 2011).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692602867FF14FCFBFCDCFBC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FBA3FCDCFAE8.text	3C6E87D692602867FF14FBA3FCDCFAE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philistina (Rhinacosmus) knirschi (Schurhoff 1933)	<div><p>Philistina (Rhinacosmus) knirschi (Schürhoff, 1933)</p><p>Rhinacosmus knirschi Schürhoff, 1933b: 90; Krikken 1979: 286 (= Rhinacosmus pilosus).</p><p>Mycteristes (Rhinacosmus) knirschi: Mikšić 1977: 109.</p><p>Type locality. “Mindanao, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao, Lanao del Norte, Bukidnon, and Davao) (Schürhoff 1933b; Mikšić 1977; Krikken 1979).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692602867FF14FBA3FCDCFAE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14FAC3FCDCF9AC.text	3C6E87D692602867FF14FAC3FCDCF9AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philistina (Rhinacosmus) pilosus (Mohnike 1873)	<div><p>Philistina (Rhinacosmus) pilosus (Mohnike, 1873)</p><p>Macronota pilosa Mohnike, 1873: 148; Schoch 1896: 35.</p><p>Mycteristes (Rhinacosmus) pilosa: Mikšić 1977: 112.</p><p>Rhinacosmus pilosus: Krikken 1979: 287.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) and Mindanao (Lanao del Norte) (Mohnike 1873; Mikšić 1977; Krikken 1979).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692602867FF14FAC3FCDCF9AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692602867FF14F93FFCDCF860.text	3C6E87D692602867FF14F93FFCDCF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philistina (Rhinacosmus) zebuanus Kraatz 1895	<div><p>Philistina (Rhinacosmus) zebuanus Kraatz, 1895</p><p>Rhinacosmus zebuanus Kraatz, 1895: 109; Krikken 1979: 287.</p><p>Rhinacosmus kuntzeni Schürhoff, 1933b: 90 . Type locality. “Mindanao, Tangcolan.”</p><p>Mycteristes (Rhinacosmus) zebuanus: Mikšić 1977: 110.</p><p>Type locality. “Philippinen-Insel (Zebu).”</p><p>Distribution in the Philippines. (Cebu) and Mindanao (Bukidnon) (Kraatz 1895; Schürhoff 1933b; Mikšić 1977; Krikken 1979).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692602867FF14F93FFCDCF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FD13FCDCFCE1.text	3C6E87D692612866FF14FD13FCDCFCE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agestrata alexisi Devecis 2004	<div><p>Agestrata alexisi Devecis, 2004</p><p>Agestrata alexisi Devecis, 2004: 8; Devecis 2019: 52.</p><p>Type locality. “Negros.”</p><p>Distribution in the Philippines. Visayas (Negros) (Devecis 2004, 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692612866FF14FD13FCDCFCE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FCFBFCDCFBC9.text	3C6E87D692612866FF14FCFBFCDCFBC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agestrata antoinei Allard 1995	<div><p>Agestrata antoinei Allard, 1995</p><p>Agestrata antoinei Allard, 1995: 10; Devecis 2019: 52.</p><p>Type locality. “ Brooke’s Point, Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Allard 1995; Devecis 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692612866FF14FCFBFCDCFBC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FBA3FC13F9F4.text	3C6E87D692612866FF14FBA3FC13F9F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agestrata luconica (Eschscholtz 1829)	<div><p>Agestrata luconica (Eschscholtz, 1829)</p><p>Cetonia (Agestrata) luconica Eschscholtz, 1829: 11 .</p><p>Agestrata splendens Gory &amp; Percheron, 1833: 506 . Type locality. “De Java.”; Blanchard 1842: 17.</p><p>Agestrata luzonica: Wallace 1867: 534; Mohnike 1873: 127; Schoch 1896: 30; Mikšić 1977: 38; Mikšić 1983: 386.</p><p>Agestrata luconica: Burmeister 1842: 309; Allard 1995: 11; Devecis 2019: 50.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Bataan, Babuyanes, Cagayan Prov., Marinduque, Mindoro,? Palawan, Pampanga, Rizal, Romblon, and Sibuyan), Visayas (Bohol, Negros, and Leyte), and Mindanao (Dinagat, Surigao, and Mt. Apo) (Eschscholtz 1829; Gory &amp; Percheron 1833; Burmeister 1842; Schultze 1916a; Mikšić 1977, 1983; Devecis 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks. Agestrata splendens from Java may be mislabeled or a lapsus calami. Blanchard (1842) originally noted this species as native to the Philippines, not Java.</p></div>	https://treatment.plazi.org/id/3C6E87D692612866FF14FBA3FC13F9F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14F9C7FC27F8F1.text	3C6E87D692612866FF14F9C7FC27F8F1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agestrata semperi subsp. boudanti Alexis & Delpont 2000	<div><p>Agestrata semperi boudanti Alexis &amp; Delpont, 2000</p><p>Agestrata boudanti Alexis &amp; Delpont, 2000: 53 .</p><p>Agestrata semperi boudanti: Devecis 2019: 48.</p><p>Type locality. “ Sud Mindanao, Mount Apo. ”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Alexis &amp; Delpont 2000; Devecis 2019).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692612866FF14F9C7FC27F8F1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612866FF14FEBBFCDCFE09.text	3C6E87D692612866FF14FEBBFCDCFE09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Theodosia rodorigesi Nagai 1980	<div><p>Theodosia (sensu lato) rodorigesi Nagai, 1980</p><p>Theodosia rodorigesi Nagai, 1980: 49; Krajčik 1998: 87.</p><p>Type locality. “ Philippines (Palawan Island).”</p><p>Distribution in the Philippines. Luzon (Palawan) (Nagai 1980).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692612866FF14FEBBFCDCFE09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692612879FF14F8CBFC27FF21.text	3C6E87D692612879FF14F8CBFC27FF21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agestrata semperi subsp. semperi Mohnike 1873	<div><p>Agestrata semperi semperi Mohnike, 1873</p><p>Agestrata semperi Mohnike, 1873: 127; Schoch 1896: 30; Mikšić 1977: 38; Allard 1995: 11.</p><p>Agestrata semperi semperi: Devecis 2019: 47.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Luzon (Manila, Laguna, Marinduque,?Mindoro,? Palawan, and? Sorsogon), Visayas (Cebu and? Leyte), and Mindanao (Surigao) (Mohnike 1873; Mikšić 1977; Devecis 2019)</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692612879FF14F8CBFC27FF21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FEBBFC27FE16.text	3C6E87D6927E2879FF14FEBBFC27FE16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agestrata semperi subsp. soucioui Devecis 2019	<div><p>Agestrata semperi soucioui Devecis, 2019</p><p>Agestrata semperi soucioui Devecis, 2019: 49 .</p><p>Type locality. “ Babuyan Island .”</p><p>Distribution in the Philippines. Luzon (Babuyan Island) (Devecis 2019).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927E2879FF14FEBBFC27FE16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FDD8FCDCFCEF.text	3C6E87D6927E2879FF14FDD8FCDCFCEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronota cupripes (Waterhouse 1841)	<div><p>Macronota cupripes (Waterhouse, 1841)</p><p>Lomaptera cupripes Waterhouse, 1841a: 38; Burmeister 1842: 316; Hope 1841: 27; Mohnike 1873: 129.</p><p>Thaumastopeus cupripes: Heller 1899b: 356; Mikšić 1977: 14; Allard 1995: 14.</p><p>Type locality. “Insulas Philippinenses.”</p><p>Distribution in the Philippines. Luzon (Laguna, Babuyanes, Bataan, and Rizal), Visayas (Samar and Bohol), and Mindanao (Waterhouse 1841a, Mohnike 1873; Heller 1899b; Mikšić 1977).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927E2879FF14FDD8FCDCFCEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FCF9FC27FBD4.text	3C6E87D6927E2879FF14FCF9FC27FBD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronota inexpectatus subsp. glabratus (Jakl 2008)	<div><p>Macronota inexpectatus glabratus (Jákl, 2008)</p><p>Thaumastopeus inexpectatus glabratus Jákl, 2008: 115 .</p><p>Type locality. “ Philippines, Leyte Is.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Jákl 2008).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927E2879FF14FCF9FC27FBD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FBA6FC27FB3C.text	3C6E87D6927E2879FF14FBA6FC27FB3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronota inexpectatus subsp. inexpectatus (Miksic 1977)	<div><p>Macronota inexpectatus inexpectatus (Mikšić, 1977)</p><p>Thaumastopeus inexpectatus Mikšić, 1977: 16; Allard 1995: 17.</p><p>Type locality. “Luzon, Imugan.”</p><p>Distribution in the Philippines. Luzon (Laguna and Nueva Vizcaya) (Mikšić 1977).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927E2879FF14FBA6FC27FB3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FA8EFC27FA65.text	3C6E87D6927E2879FF14FA8EFC27FA65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronota macgregori subsp. macgregori (Schultze 1915)	<div><p>Macronota macgregori macgregori (Schultze, 1915)</p><p>Thaumastopeus mcgregori Schultze, 1915: 271; Mikšić 1977: 13.</p><p>Type locality. “ Luzon, Laguna, Paete.”</p><p>Distribution in the Philippines. Luzon (Laguna) (Schultze 1915; Mikšić 1977).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927E2879FF14FA8EFC27FA65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14FA77FC27F94A.text	3C6E87D6927E2879FF14FA77FC27F94A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronota macgregori subsp. mindanaoensis (Arnaud & Delpont 2001)	<div><p>Macronota macgregori mindanaoensis (Arnaud &amp; Delpont, 2001)</p><p>Thaumastopeus macgregori mindanaoensis Arnaud &amp; Delpont, 2001: 74 .</p><p>Type locality. “ Province de Bukidnon.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon) (Arnaud &amp; Delpont 2001).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927E2879FF14FA77FC27F94A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2879FF14F95CFCDCF8B3.text	3C6E87D6927E2879FF14F95CFCDCF8B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronota miksici (Allard 1995)	<div><p>Macronota miksici (Allard, 1995)</p><p>Thaumastopeus miksici Allard, 1995: 17 .</p><p>Type locality. “ Philippines, île de Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Allard 1995).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927E2879FF14F95CFCDCF8B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927E2878FF14F805FAABFEC4.text	3C6E87D6927E2878FF14F805FAABFEC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronota nigritus subsp. nigritus (Frolich 1792)	<div><p>Macronota nigritus nigritus (Frölich, 1792)</p><p>Cetonia nigrita Frölich, 1792: 110 .</p><p>Lomaptera ebena Burmeister, 1842: 315 . Type locality. “ Philippinen.”</p><p>Thaumastopeus nigritus: Allard 1995: 16.</p><p>Type locality. “ Bengalen.”</p><p>Distribution in the Philippines. Philippine archipelago (Burmeister 1842).</p><p>Overall distribution. This species is found in the Oriental and Palearctic regions (Allard 1995).</p></div>	https://treatment.plazi.org/id/3C6E87D6927E2878FF14F805FAABFEC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FED6FC27FD5C.text	3C6E87D6927F2878FF14FED6FC27FD5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronota nigritus subsp. nigroaeneus (Waterhouse 1841)	<div><p>Macronota nigritus nigroaeneus (Waterhouse, 1841)</p><p>Lomaptera nigroaenea Waterhouse, 1841a: 38; Burmeister 1842: 316; Hope 1841: 27.</p><p>Lomaptera schaumi Eydoux &amp; Souleyet, 1841: 302 . Type locality. “ Manille.”</p><p>Thaumastopeus nigritus nigroaeneus: Heller 1899b: 355; Mikšić 1977: 15.</p><p>Type locality. “Insulas Philippinenses.”</p><p>Distribution in the Philippines. Luzon (Pangasinan, Ilocos Sur, Manila, Rizal, Camarines Sur, Quezon, and Laguna) and Mindanao (Davao, Bukidnon, and Zamboanga) (Eydoux &amp; Souleyet 1841; Waterhouse 1841a; Heller 1899b; Mikšić 1977).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927F2878FF14FED6FC27FD5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FD2EFC27FC84.text	3C6E87D6927F2878FF14FD2EFC27FC84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronota palawanicus subsp. hangayi (Arnaud & Delpont 2001)	<div><p>Macronota palawanicus hangayi (Arnaud &amp; Delpont, 2001)</p><p>Thaumastopeus palawanicus hangayi Arnaud &amp; Delpont, 2001: 78 .</p><p>Type locality. “ Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Arnaud &amp; Delpont 2001).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927F2878FF14FD2EFC27FC84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FC16FC27FBA4.text	3C6E87D6927F2878FF14FC16FC27FBA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronota palawanicus subsp. palawanicus (Heller 1899)	<div><p>Macronota palawanicus palawanicus (Heller, 1899)</p><p>Thaumastopeus palawanicus Heller, 1899b: 357; Mikšić 1977: 20 (= Thaumastopeus higginsi); Allard 1995: 14; Arnaud &amp; Delpont 2001: 77.</p><p>Type locality. “ Insula Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) and Mindanao (Bukidnon) (Heller 1899b; Mikšić 1977; Arnaud &amp; Delpont 2001).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927F2878FF14FC16FC27FBA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14FB36FE2BF988.text	3C6E87D6927F2878FF14FB36FE2BF988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronota pugnator subsp. pugnator (Heller 1899)	<div><p>Macronota pugnator pugnator (Heller, 1899)</p><p>Thaumastopeus pugnator Heller, 1899: 362; Allard 1995: 13; Sakai &amp; Nagai 1998: 195; Arnaud &amp; Delpont 2001: 73.</p><p>Type locality. “ Penang et Sumatra.”</p><p>Distribution in the Philippines. Luzon (Allard 1995; Sakai &amp; Nagai 1998; Arnaud &amp; Delpont 2001).</p><p>Overall distribution. The? Philippines and Malaysia.</p><p>Remarks. Arnaud &amp; Delpont (2001) argued that Malaysia and the Philippines historically have no physical land connection, and that populations of Macronota pugnator (Heller, 1899) from these regions differ in certain morphological aspects. This raises the question of whether the true Macronota pugnator is present in the Philippines. Furthermore, earlier taxonomic works synonymized Macronota macgregori with M acronota pugnator, and it is most likely that the population previously identified as pugnator in Luzon is macgregori .</p></div>	https://treatment.plazi.org/id/3C6E87D6927F2878FF14FB36FE2BF988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927F2878FF14F9E2FC27F894.text	3C6E87D6927F2878FF14F9E2FC27F894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macronota striatus subsp. rubyi (Arnaud 1985)	<div><p>Macronota striatus rubyi (Arnaud, 1985)</p><p>Thaumastopeus rubyi Arnaud, 1985: 18 .</p><p>Thaumastopeus striatus rubyi: Sakai &amp; Nagai 1998: 195.</p><p>Type locality. “ Brooke’s Point, Palawan, Philippines.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Arnaud 1985).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927F2878FF14F9E2FC27F894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FEBBFCDCFE09.text	3C6E87D6927C287BFF14FEBBFCDCFE09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clerota rodriguezi Arnaud 1985	<div><p>Clerota rodriguezi Arnaud, 1985</p><p>Clerota rodriguezi Arnaud, 1985: 18 .</p><p>Type locality. “ Brooke’s Point, Palawan, Philippines.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Arnaud 1985).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927C287BFF14FEBBFCDCFE09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FDDBFCDCFCCD.text	3C6E87D6927C287BFF14FDDBFCDCFCCD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paraplectrone crassa (Miksic 1973)	<div><p>Paraplectrone crassa (Mikšić, 1973)</p><p>Plectrone crassa Mikšić, 1973: 78; Mikšić 1974: 162; Mikšić 1976: 389; Krikken 1978: 31.</p><p>Paraplectrone crassa: Mikšić 1985: 141; Devecis 2009: 84.</p><p>Type locality. “ Mindanao, Mt. Ape (=Mt. Apo).”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Mikšić 1973).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927C287BFF14FDDBFCDCFCCD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FCDFFCDCFC35.text	3C6E87D6927C287BFF14FCDFFCDCFC35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paraplectrone kitangladensis Devecis 2009	<div><p>Paraplectrone kitangladensis Devecis, 2009</p><p>Paraplectrone kitangladensis Devecis, 2009: 85 .</p><p>Type locality. “ Mont Kitanglad, Bukidnon Province.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon) (Devecis 2009).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927C287BFF14FCDFFCDCFC35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FBFFFCDCFAD5.text	3C6E87D6927C287BFF14FBFFFCDCFAD5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectrone delponti Devecis 2001	<div><p>Plectrone delponti Devecis, 2001</p><p>Plectrone delponti Devecis, 2001: 82; Devecis 2009: 78.</p><p>Type locality. “ Panay Island, Philippines.”</p><p>Distribution in the Philippines. Visayas (Panay Island) (Devecis 2001).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927C287BFF14FBFFFCDCFAD5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14FAA7FCDCF9F5.text	3C6E87D6927C287BFF14FAA7FCDCF9F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectrone endroedii Miksic 1974	<div><p>Plectrone endroedii Mikšić, 1974</p><p>Plectrone endroedii Mikšić, 1974b: 162; Mikšić 1976: 388; Krikken 1978: 32; Mikšić 1984: 300; Pavićević 1988: 170; Devecis 2009: 68.</p><p>Type locality. “ Philippinen Insel Samar.”</p><p>Distribution in the Philippines. Visayas (Samar) and Mindanao (Bukidnon, Cotabato, Mt. Apo, and Surigao del Sur) (Mikšić 1974b, 1976, 1984; Krikken 1978; Pavićević 1988).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927C287BFF14FAA7FCDCF9F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14F9C7FCDCF8DD.text	3C6E87D6927C287BFF14F9C7FCDCF8DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectrone ismaeli Arnaud 2004	<div><p>Plectrone ismaeli Arnaud, 2004</p><p>Plectrone ismaeli Arnaud, 2004: 3; Devecis 2009: 70.</p><p>Type locality. “ Philippines, North Luzon, Babuyanes, Calayan Island.”</p><p>Distribution in the Philippines. Luzon (Babuyanes) (Arnaud 2004).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927C287BFF14F9C7FCDCF8DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927C287BFF14F8AFFCDCF805.text	3C6E87D6927C287BFF14F8AFFCDCF805.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectrone lumawigi Miksic 1986	<div><p>Plectrone lumawigi Mikšić, 1986</p><p>Plectrone lumawigi Mikšić, 1986: 357; Devecis 2009: 71.</p><p>Type locality. “ Mount Madias, Panay.”</p><p>Distribution in the Philippines. Visayas (Panay) (Mikšić 1986; Devecis 2009).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927C287BFF14F8AFFCDCF805	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FF0BFCDCFE99.text	3C6E87D6927D287AFF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectrone malvari Arnaud 1992	<div><p>Plectrone malvari Arnaud, 1992</p><p>Plectrone malvari Arnaud, 1992: 31; Devecis 2009: 65.</p><p>Type locality. “ Philippines, Marinduque.”</p><p>Distribution in the Philippines. Luzon (Marinduque) (Arnaud 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927D287AFF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FEF0FCDCFDCE.text	3C6E87D6927D287AFF14FEF0FCDCFDCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectrone negricola Krikken 1978	<div><p>Plectrone negricola Krikken, 1978</p><p>Plectrone negricola Krikken, 1978: 30; Devecis 2001: 83; Devecis 2009: 75.</p><p>Type locality. “Negros, Philippines.”</p><p>Distribution in the Philippines. Visayas (Negros Island) (Krikken 1978).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927D287AFF14FEF0FCDCFDCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FDD9FCDCFC1F.text	3C6E87D6927D287AFF14FDD9FCDCFC1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectrone nigrocoerulea (Waterhouse 1841)	<div><p>Plectrone nigrocoerulea (Waterhouse, 1841)</p><p>Macronota nigrocoerulea Waterhouse, 1841: 40; Hope 1841: 27; Wallace 1867: 546.</p><p>Chalcothea Barrotiana Burmeister, 1842: 319 . Type locality. “Luzon.”</p><p>Plectrone nigrocoerulea rugipes Devecis, 2009: 65 . Type locality. “Mont Banahaw, Sud Luzon.”</p><p>Plectrone nigrocoerulea: Wallace 1868: 546; Mohnike 1873: 133; Schoch 1896: 34; Mikšić 1976: 386; Krikken 1978: 32; Devecis 2009: 64.</p><p>Type locality. “Insulas Philippinenses.”</p><p>Distribution in the Philippines. Luzon (Bataan, Laguna, Marinduque, Mountain Province, Palawan, and Romblon), Visayas (Cebu and Leyte), and Mindanao (Waterhouse 1841a; Burmeister 1842; Wallace 1868; Mikšić 1976; Krikken 1978; Devecis 2009).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927D287AFF14FDD9FCDCFC1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FC6EFCDCFB44.text	3C6E87D6927D287AFF14FC6EFCDCFB44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectrone nonveilleri Miksic 1983	<div><p>Plectrone nonveilleri Mikšić, 1983</p><p>Plectrone nonveilleri Mikšić, 1983: 383; Devecis 2009: 69.</p><p>Type locality. “Luzon, Imugan.”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao and Nueva Vizcaya) (Mikšić 1983; Devecis 2009).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927D287AFF14FC6EFCDCFB44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FB57FCDCFAAD.text	3C6E87D6927D287AFF14FB57FCDCFAAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectrone rodriguezi Nagai 1984	<div><p>Plectrone rodriguezi Nagai, 1984</p><p>Plectrone rodriguezi Nagai, 1984: 6; Devecis 2009: 81.</p><p>Type locality. “ Philippines (Palawan Island).”</p><p>Distribution in the Philippines. Luzon (Palawan) (Nagai 1984).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927D287AFF14FB57FCDCFAAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14FA3CFCDDF992.text	3C6E87D6927D287AFF14FA3CFCDDF992.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectrone romblonica Miksic 1983	<div><p>Plectrone romblonica Mikšić, 1983</p><p>Plectrone romblonica Mikšić, 1983: 384; Devecis 2009: 71.</p><p>Type locality. “ Philippinen, Insel Romblon.”</p><p>Distribution in the Philippines. Luzon (Romblon) (Mikšić 1983).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927D287AFF14FA3CFCDDF992	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14F9E5FCDCF8FB.text	3C6E87D6927D287AFF14F9E5FCDCF8FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectrone romeoi Arnaud 1987	<div><p>Plectrone romeoi Arnaud, 1987</p><p>Plectrone romeoi Arnaud, 1987a: 27; Arnaud 1992: 31; Devecis 2009: 71.</p><p>Type locality. “ Philippines, Mindoro.”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Arnaud 1987a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927D287AFF14F9E5FCDCF8FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927D287AFF14F8CDFCDCF823.text	3C6E87D6927D287AFF14F8CDFCDCF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Plectrone viridipurpurea Devecis 2009	<div><p>Plectrone viridipurpurea Devecis, 2009</p><p>Plectrone viridipurpurea Devecis, 2009: 79 .</p><p>Type locality. “ Mount Apo, Sud Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Devecis 2009).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927D287AFF14F8CDFCDCF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FEBBFCDCFE08.text	3C6E87D6927A287DFF14FEBBFCDCFE08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euselates dissimilis Nagai 1986	<div><p>Euselates (sensu lato) dissimilis Nagai, 1986</p><p>Euselates dissimilis Nagai, 1986: 184 .</p><p>Type locality. “ Palawan Island.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Nagai 1986).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927A287DFF14FEBBFCDCFE08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FE63FC27FD70.text	3C6E87D6927A287DFF14FE63FC27FD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euselates stictica (Hope 1847) Miksic 1974	<div><p>Euselates (sensu lato) stictica luzonica Mikšić, 1974</p><p>Euselates stictica luzonica Mikšić, 1974d: 120; Mikšić 1976: 343.</p><p>Type locality. “ Philippinen ( Luzon).”</p><p>Distribution in the Philippines. Luzon (Sorsogon, Camarines Sur, and Laguna) (Mikšić 1974d, 1976).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927A287DFF14FE63FC27FD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FD4BFC27FBEC.text	3C6E87D6927A287DFF14FD4BFC27FBEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euselates stictica (Hope 1847) stictica (Hope 1847	<div><p>Euselates (sensu lato) stictica stictica (Hope, 1847)</p><p>Macronota stictica Hope, 1847: 34 .</p><p>Macronota guttulata Wallace, 1867: 94 . Type locality. “ Philippine Islands.”; Wallace 1868: 551; Mohnike 1873: 150.</p><p>Macronota lugubris Mohnike, 1873: 152 . Type locality. “ Panaon, Leyte, Luzon.”</p><p>Euselates stictica: Mikšić 1974d: 110; Mikšić 1976: 341.</p><p>Type locality. “Agris Mysoriensibus.”</p><p>Distribution in the Philippines. Visayas (Biliran, Leyte, and Samar) and Mindanao (Mt. Apo, Surigao, Agusan del Norte, and Lanao del Norte) (Hope 1847; Wallace 1867; Mohnike 1873; Mikšić 1974d, 1976).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927A287DFF14FD4BFC27FBEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FB6FFCDCFA44.text	3C6E87D6927A287DFF14FB6FFCDCFA44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Ixorida) basilanensis Jakl 2011	<div><p>Ixorida (Ixorida) basilanensis Jákl, 2011</p><p>Ixorida (Ixorida) basilanensis Jákl, 2011: 218; Legrand 2017: 53; Jákl &amp; Legrand 2018: 87.</p><p>Type locality. “ Philippines, Basilan Island.”</p><p>Distribution in the Philippines. Mindanao (Basilan) (Jákl 2011).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927A287DFF14FB6FFCDCFA44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14FA57FCDCF940.text	3C6E87D6927A287DFF14FA57FCDCF940.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Ixorida) florenti Arnaud 1989	<div><p>Ixorida (Ixorida) florenti Arnaud, 1989</p><p>Ixorida florenti Arnaud, 1989a: 62; Legrand 2017: 53.</p><p>Ixorida (Ixorida) florenti: Jákl &amp; Legrand 2018: 87.</p><p>Type locality. “ Philippines, Romblon Island, Sibuyan.”</p><p>Distribution in the Philippines. Luzon (Sibuyan) (Arnaud 1989a; Jákl &amp; Legrand 2018).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927A287DFF14FA57FCDCF940	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927A287DFF14F95BFCDCF860.text	3C6E87D6927A287DFF14F95BFCDCF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Ixorida) guentheri Schurhoff 1935	<div><p>Ixorida (Ixorida) guentheri Schürhoff, 1935</p><p>Ixorida guentheri Schürhoff, 1935: 26; Mikšić 1970d: 124; Mikšić 1976: 193.</p><p>Ixorida (Ixorida) guentheri: Legrand 2017: 53; Jákl &amp; Legrand 2018: 87.</p><p>Type locality. “ Panaon-Insel (Philippinen).”</p><p>Distribution in the Philippines. Visayas (Bohol, Leyte, and Samar) and Mindanao (Dinagat) (Schürhoff 1935; Mikšić 1970d, 1976; Legrand 2017; Jákl &amp; Legrand 2018).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927A287DFF14F95BFCDCF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14FF0BFC27FE51.text	3C6E87D6927B287CFF14FF0BFC27FE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Ixorida) philippinensis subsp. nigripennis Miksic 1970	<div><p>Ixorida (Ixorida) philippinensis nigripennis Mikšić, 1970</p><p>Ixorida philippinensis nigripennis Mikšić, 1970d: 124; Mikšić 1976: 192; Pavićević 1988: 170.</p><p>Ixorida (Ixorida) philippinensis nigripennis: Jákl &amp; Legrand 2018: 87.</p><p>Type locality. “Luzon, Lamao.”</p><p>Distribution in the Philippines. Luzon (Bataan, Masbate, and Zambales) and Visayas (Antique, Panay, and Negros Oriental) (Mikšić 1970d, 1976; Pavićević 1988).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927B287CFF14FF0BFC27FE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14FE2BFC27FCE1.text	3C6E87D6927B287CFF14FE2BFC27FCE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Ixorida) philippinensis subsp. philippinensis (Waterhouse 1841)	<div><p>Ixorida (Ixorida) philippinensis philippinensis (Waterhouse, 1841)</p><p>Macronota philippinensis Waterhouse, 1841: 39; Hope 1841: 27; Mohnike 1873: 136.</p><p>Macronota auroguttata Burmeister, 1842: 323 . Type locality. “ Philippinen.”</p><p>Ixorida philippensis: Schoch 1897: 466.</p><p>Ixorida philippinensis philippinensis: Mikšić 1970d: 123; Mikšić 1976: 190.</p><p>Ixorida (Ixorida) philippinensis philippinensis: Legrand 2017: 53; Jákl &amp; Legrand 2018: 87.</p><p>Type locality. “Insulas Philippinarum.”</p><p>Distribution in the Philippines. Luzon (Camarines Sur, Laguna, Masbate, Rizal, Marinduque, Mindoro, Mt. Banahao, Polillo, Nueva Vizcaya,? Sorsogon, and Zambales) and Visayas (Bohol, Panay, and Leyte) (Waterhouse 1841a; Burmeister 1842; Mikšić 1970d, 1976; Pavićević 1988; Jákl &amp; Legrand 2018).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927B287CFF14FE2BFC27FCE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14FCFBFCDCFBA5.text	3C6E87D6927B287CFF14FCFBFCDCFBA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Ixorida) propinqua (Mohnike 1873)	<div><p>Ixorida (Ixorida) propinqua (Mohnike, 1873)</p><p>Macronota propinqua Mohnike, 1873: 139 .</p><p>Ixorida propinqua: Mikšić 1970d: 124; Mikšić 1976: 193.</p><p>Ixorida (Ixorida) propinqua: Legrand 2017: 53; Jákl &amp; Legrand 2018: 87.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Agusan del Norte, Bukidnon, Davao, Surigao del Norte, and Zamboanga del Norte) (Mohnike 1873; Mikšić 1970d, 1976; Jákl &amp; Legrand 2018).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927B287CFF14FCFBFCDCFBA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14FB6FFCDCF9D1.text	3C6E87D6927B287CFF14FB6FFCDCF9D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Mecinonota) albomaculata (Moser 1917)	<div><p>Ixorida (Mecinonota) albomaculata (Moser, 1917)</p><p>Macronota albomaculata Moser, 1917a: 6 .</p><p>Mecinonota albomaculata: Mikšić 1972a: 118; Mikšić 1976: 245.</p><p>Ixorida (Mecinonota) albomaculata: Legrand 2017: 55.</p><p>Type locality. “ Luzon (Mt. Isarog).”</p><p>Distribution in the Philippines. Luzon (Camarines Sur, Laguna, and Nueva Vizcaya) (Moser 1917a; Mikšić 1972a, 1976).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927B287CFF14FB6FFCDCF9D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287CFF14F9ABFCDCF8DD.text	3C6E87D6927B287CFF14F9ABFCDCF8DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Mecinonota) antoinei (Miksic 1986)	<div><p>Ixorida (Mecinonota) antoinei (Mikšić, 1986)</p><p>Mecinonota antoinei Mikšić, 1986: 67 .</p><p>Ixorida (Mecinonota) antoinei: Legrand 2017: 55.</p><p>Type locality. “ Insel Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Mikšić 1986).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927B287CFF14F9ABFCDCF8DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6927B287FFF14F8AFFC27FEC5.text	3C6E87D6927B287FFF14F8AFFC27FEC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Mecinonota) fraterna subsp. fraterna (Westwood 1854)	<div><p>Ixorida (Mecinonota) fraterna fraterna (Westwood, 1854)</p><p>Macronota fraterna Westwood, 1854: 71 .</p><p>Mecinonota fraterna: Mikšić 1972a: 100; Mikšić 1976: 231; Pavićević 1988: 170.</p><p>Ixorida (Mecinonota) fraterna: Legrand 2017: 55.</p><p>Type locality. “Insulis Philippinensibus.”</p><p>Distribution in the Philippines. Luzon (Baguio, Camarines Sur, Masbate, Laguna, Mountain Province, and Nueva Vizcaya), Visayas (Leyte, Negros, and Samar), and Mindanao (Dinagat and Surigao del Norte) (Westwood 1854; Mikšić 1972a, 1976; Pavićević 1988).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6927B287FFF14F8AFFC27FEC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FED4FCDCFE2A.text	3C6E87D69278287FFF14FED4FCDCFE2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Mecinonota) incongrua Legrand 2017	<div><p>Ixorida (Mecinonota) incongrua Legrand, 2017</p><p>Ixorida (Mecinonota) incongrua Legrand, 2017: 56 .</p><p>Type locality. “ Mindanao, Davao del Sur, Kapatagan.”</p><p>Distribution in the Philippines. Mindanao (Davao del Sur) (Legrand 2017).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69278287FFF14FED4FCDCFE2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FDBCFCDCFD12.text	3C6E87D69278287FFF14FDBCFCDCFD12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Mecinonota) leytensis Legrand 2017	<div><p>Ixorida (Mecinonota) leytensis Legrand, 2017</p><p>Ixorida (Mecinonota) leytensis Legrand, 2017: 55 .</p><p>Type locality. “ Philippines, Leyte, Mt. Balocaue. ”</p><p>Distribution in the Philippines. Visayas (Leyte) (Legrand 2017).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69278287FFF14FDBCFCDCFD12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FD65FCDCFBD4.text	3C6E87D69278287FFF14FD65FCDCFBD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Mecinonota) mindanaoensis (Mohnike 1873)	<div><p>Ixorida (Mecinonota) mindanaoensis (Mohnike, 1873)</p><p>Macronota mindanaoensis Mohnike, 1873: 140; Schoch 1896: 35; Mikšić 1976: 356.</p><p>Mecinonota albomaculata mindanaoana Mikšić, 1983: 383 . Type locality. “Silipon-Bukidnon.”</p><p>Ixorida (Mecinonota) mindanaoensis: Legrand 2017: 55.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Luzon (Camarines Sur) and Mindanao (Bukidnon and Mt. Apo) (Mohnike 1873; Mikšić 1983; Legrand 2017).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69278287FFF14FD65FCDCFBD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FBA6FCDCFB3C.text	3C6E87D69278287FFF14FBA6FCDCFB3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Mecinonota) mindoroana Jakl 2014	<div><p>Ixorida (Mecinonota) mindoroana Jákl, 2014</p><p>Ixorida (Mecinonota) mindoroana Jákl, 2014: 444; Legrand 2017: 55.</p><p>Type locality. “ The Philippines, Mindoro Island, Mt. Halcon. ”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Jákl 2014).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69278287FFF14FBA6FCDCFB3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14FA8FFCDCF9DA.text	3C6E87D69278287FFF14FA8FFCDCF9DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Mecinonota) palawanica (Moser 1910)	<div><p>Ixorida (Mecinonota) palawanica (Moser, 1910)</p><p>Macronota luctuosa palawanica Moser, 1910: 183 .</p><p>Mecinonota luctuosa palavanica: Mikšić 1972a: 115; Mikšić 1976: 241.</p><p>Ixorida (Mecinonota) palawanica: Legrand 2017: 55.</p><p>Type locality. “ Palawan, Iwahig, P. I.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Moser 1910; Mikšić 1972a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69278287FFF14FA8FFCDCF9DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287FFF14F9ACFC27F902.text	3C6E87D69278287FFF14F9ACFC27F902.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Mecinonota) regia subsp. palawana Jakl 2020	<div><p>Ixorida (Mecinonota) regia palawana Jákl, 2020</p><p>Ixorida (Mecinonota) regia palawana Jákl, 2020: 399 .</p><p>Type locality. “ The Philippines, Palawan Island, Magara, Roxas.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Jákl 2020).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69278287FFF14F9ACFC27F902	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69278287EFF14F895FCDCFF68.text	3C6E87D69278287EFF14F895FCDCFF68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixorida (Mecinonota) vidua (Wallace 1867)	<div><p>Ixorida (Mecinonota) vidua (Wallace, 1867)</p><p>Macronota vidua Wallace, 1867: 94; Wallace 1868: 550; Mohnike 1873: 137; Schoch 1896: 36.</p><p>Mecinonota vidua: Mikšić 1972a: 116; Mikšić 1976: 243.</p><p>Ixorida (Mecinonota) vidua: Legrand 2017: 55.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Romblon) and Mindanao (Lanao del Norte and Agusan del Norte) (Wallace 1867; Mikšić 1972a, 1976).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69278287EFF14F895FCDCFF68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FEBBFCDCFE16.text	3C6E87D69279287EFF14FEBBFCDCFE16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Meroloba spadix Nagai 1986	<div><p>Meroloba spadix Nagai, 1986</p><p>Meroloba spadix Nagai, 1986: 186 .</p><p>Type locality. “ Palawan Island.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Nagai 1986).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69279287EFF14FEBBFCDCFE16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FE60FC27FD12.text	3C6E87D69279287EFF14FE60FC27FD12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Meroloba suturalis subsp. quadrilineata Nagai 1984	<div><p>Meroloba suturalis quadrilineata Nagai, 1984</p><p>Meroloba suturalis: Mikšić 1971c: 394; Mikšić 1976: 215.</p><p>Meloroba quadrilineata Nagai, 1984: 4; Mikšić 1986: 66.</p><p>Type locality. “ Palawan (Philippines).”</p><p>Distribution in the Philippines. Luzon (Palawan) and Mindanao (Nagai 1986; Mikšić 1986).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69279287EFF14FE60FC27FD12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FCDDFCDCFC30.text	3C6E87D69279287EFF14FCDDFCDCFC30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleuronota lumawigi Arnaud 1990	<div><p>Pleuronota lumawigi Arnaud, 1990</p><p>Pleuronota lumawigi Arnaud, 1990: 8 .</p><p>Type locality. “ Philippines, Oriental Negros.”</p><p>Distribution in the Philippines. Visayas (Negros Oriental) (Arnaud 1990).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69279287EFF14FCDDFCDCFC30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FB8AFCDCFB18.text	3C6E87D69279287EFF14FB8AFCDCFB18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleuronota mimetica Legrand 2025	<div><p>Pleuronota mimetica Legrand, 2025</p><p>Pleuronota mimetica Legrand, 2025: 1 .</p><p>Type locality. “ Philippines, Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Legrand 2025).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69279287EFF14FB8AFCDCFB18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FB73FC27FA65.text	3C6E87D69279287EFF14FB73FC27FA65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleuronota setosa subsp. imuganica (Miksic 1972)	<div><p>Pleuronota setosa imuganica (Mikšić, 1972)</p><p>Macronotiola setosa imuganica Mikšić, 1972c: 131; Mikšić 1976: 161.</p><p>Pleuronota setosa imuganica: Krikken 1977b: 202.</p><p>Type locality. “Luzon-Imugan.”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Mikšić 1972c, 1976).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69279287EFF14FB73FC27FA65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69279287EFF14FA74FC27F926.text	3C6E87D69279287EFF14FA74FC27F926.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pleuronota setosa subsp. setosa (Moser 1914)	<div><p>Pleuronota setosa setosa (Moser, 1914)</p><p>Macronota setosa Moser, 1914: 574 .</p><p>Macronotiola setosa: Mikšić 1972c: 130; Mikšić 1976: 159.</p><p>Pleuronota setosa setosa: Krikken 1977b: 202.</p><p>Type locality. “ Luzon (Mt. Banahao).”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao and Camarines Sur) (Moser 1914; Mikšić 1972c, 1976; Krikken 1977b).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69279287EFF14FA74FC27F926	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692792871FF14F8E9FCDCFE98.text	3C6E87D692792871FF14F8E9FCDCFE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taeniodera abdominalis (Mohnike 1873)	<div><p>Taeniodera abdominalis (Mohnike, 1873)</p><p>Macronota abdominalis Mohnike, 1873: 142; Schoch 1896: 34.</p><p>Macronota domina Thomson, 1878: 16 . Type locality. “Ins. Philipp.”; Schoch 1896: 35; Sakai 1996: 27 (= Taeniodera tricolor).</p><p>Taeniodera abdominalis: Mikšić 1976: 78; Mikšić 1984: 300; Sakai,1996: 25.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Luzon (Camarines Sur, Laguna, Mt. Banahao, Masbate), Visayas (Negros), and Mindanao (Mt. Apo and Zamboanga del Norte) (Mohnike 1873; Thomson 1878; Mikšić 1976, 1984).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692792871FF14F8E9FCDCFE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FEF3FCDCFDE4.text	3C6E87D692762871FF14FEF3FCDCFDE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taeniodera boettcheri (Moser 1915)	<div><p>Taeniodera boettcheri (Moser, 1915)</p><p>Macronota boettcheri Moser, 1915b: 590 .</p><p>Taeniodera boettcheri: Mikšić 1976: 62.</p><p>Type locality. “N. Palawan ( Binaluan).”</p><p>Distribution in the Philippines. Luzon (Palawan) (Moser 1915b; Mikšić 1976).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692762871FF14FEF3FCDCFDE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FDF7FCDCFCCC.text	3C6E87D692762871FF14FDF7FCDCFCCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taeniodera gratiosa (Mohnike 1873)	<div><p>Taeniodera gratiosa (Mohnike, 1873)</p><p>Macronota gratiosa Mohnike, 1873: 149; Mikšić 1976: 354.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Mohnike 1873).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692762871FF14FDF7FCDCFCCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FCDFFC27FBC8.text	3C6E87D692762871FF14FCDFFC27FBC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taeniodera jucunda subsp. flavosignata (Moser 1914)	<div><p>Taeniodera jucunda flavosignata (Moser, 1914)</p><p>Macronota flavosignata Moser, 1914: 573 .</p><p>Taeniodera jucunda flavosignata: Mikšić 1976: 66.</p><p>Type locality. “ Philippine (Leite).”</p><p>Distribution in the Philippines. Visayas (Leyte) (Moser 1914; Mikšić 1976).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692762871FF14FCDFFC27FBC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FBA3FC27FA8C.text	3C6E87D692762871FF14FBA3FC27FA8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taeniodera jucunda subsp. jucunda (Mohnike 1873)	<div><p>Taeniodera jucunda jucunda (Mohnike, 1873)</p><p>Macronota jucunda Mohnike, 1873: 145; Schoch 1896: 35.</p><p>Macronota sponsa Mohnike, 1873: 146 . Type locality. “Mindanao.”; Schoch 1896: 35.</p><p>Taeniodera jucunda: Mikšić 1976: 63.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon and Zamboanga Sibugay) (Mohnike 1873; Moser 1914; Mikšić 1976).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692762871FF14FBA3FC27FA8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14FA1FFC27F9F4.text	3C6E87D692762871FF14FA1FFC27F9F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taeniodera tricolor subsp. palawanica Sakai 1996	<div><p>Taeniodera tricolor palawanica Sakai, 1996</p><p>Taeniodera tricolor palawanica Sakai, 1996: 28 .</p><p>Type locality. “ Quezon, S. Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Sakai 1996).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692762871FF14FA1FFC27F9F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762871FF14F9C7FC27F8DC.text	3C6E87D692762871FF14F9C7FC27F8DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taeniodera tricolor subsp. samarensis Sakai 1996	<div><p>Taeniodera tricolor samarensis Sakai, 1996</p><p>Taeniodera tricolor samarensis Sakai, 1996: 27 .</p><p>Type locality. “Hinabagan, W. Samar.”</p><p>Distribution in the Philippines. Visayas (Samar) (Sakai 1996).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692762871FF14F9C7FC27F8DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692762870FF14F8AFFC6AFF20.text	3C6E87D692762870FF14F8AFFC6AFF20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Taeniodera tricolor subsp. tricolor (Mohnike 1873)	<div><p>Taeniodera tricolor tricolor (Mohnike, 1873)</p><p>Macronota tricolor Mohnike, 1873: 143; Schoch 1896: 36.</p><p>Macronota tricolor bispilota Schaufuss, 1885: 195 .</p><p>Macronota tricolor lacernae Schaufuss, 1885: 195 .</p><p>Taeniodera tricolor: Sakai 1996: 27.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Mohnike 1873; Schaufuss 1885).</p><p>Overall distribution. The Philippines and? Malaysia (Sakai 1996).</p></div>	https://treatment.plazi.org/id/3C6E87D692762870FF14F8AFFC6AFF20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14FBFFFCDCFAE8.text	3C6E87D692772870FF14FBFFFCDCFAE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasyvalgus (Dasyvalgus) biplagiatus Moser 1915	<div><p>Dasyvalgus (Dasyvalgus) biplagiatus Moser, 1915</p><p>Dasyvalgus biplagiatus Moser, 1915b: 602; Ricchiardi 1996: 368.</p><p>Dasyvalgus (Dasyvalgus) biplagiatus: Krajčik 1999: 60.</p><p>Type locality. “ Mindanao (Iligan).”</p><p>Distribution in the Philippines. Mindanao (Lanao del Norte and Surigao) (Moser 1915b; Ricchiardi 1996).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692772870FF14FBFFFCDCFAE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14FAC3FCDCF9AC.text	3C6E87D692772870FF14FAC3FCDCF9AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasyvalgus (Dasyvalgus) luzonicus (Kraatz 1883)	<div><p>Dasyvalgus (Dasyvalgus) luzonicus (Kraatz, 1883)</p><p>Valgus sellatus luzonicus Kraatz, 1883: 375 .</p><p>Dasyvalgus luzonicus: Ricchiardi 1996: 369.</p><p>Dasyvalgus (Dasyvalgus) luzonicus: Krajčik 1999: 60.</p><p>Type locality. “Luzon (Ins. Philippin.).”</p><p>Distribution in the Philippines. Luzon (Manila, Laguna, Mindoro, and Palawan) (Kraatz 1883; Ricchiardi 1996).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692772870FF14FAC3FCDCF9AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14F93FFCDCF84C.text	3C6E87D692772870FF14F93FFCDCF84C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasyvalgus (Dasyvalgus) mindanaonensis Ricchiardi 1996	<div><p>Dasyvalgus (Dasyvalgus) mindanaonensis Ricchiardi, 1996</p><p>Dasyvalgus mindanaonensis Ricchiardi, 1996: 369 .</p><p>Dasyvalgus (Dasyvalgus) mindanaonensis: Krajčik 1999: 60.</p><p>Type locality. “ Mindanao, Kabasalan, Zamboanga.”</p><p>Distribution in the Philippines. Mindanao (Zamboanga Sibugay, Lanao del Norte, and Surigao) (Ricchiardi 1996).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692772870FF14F93FFCDCF84C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14FE63FCDCFD70.text	3C6E87D692772870FF14FE63FCDCFD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microvalgus (Paediovalgus) luzonensis Antoine 1993	<div><p>Microvalgus (Paediovalgus) luzonensis Antoine, 1993</p><p>Microvalgus (Paediovalgus) luzonensis Antoine, 1993: 336; Krajčik 1999: 58.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Ilocos Norte and Quezon) (Antoine 1993).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692772870FF14FE63FCDCFD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692772870FF14FD4BFCDCFC7C.text	3C6E87D692772870FF14FD4BFCDCFC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microvalgus (Paediovalgus) mindanaoensis Antoine 1993	<div><p>Microvalgus (Paediovalgus) mindanaoensis Antoine, 1993</p><p>Microvalgus (Paediovalgus) mindanaoensis Antoine, 1993: 335; Krajčik 1999: 58.</p><p>Type locality. “Mindanao, Momungan.”</p><p>Distribution in the Philippines. Luzon (Masbate), Visayas (Leyte) and Mindanao (Lanao del Norte, Zamboanga del Norte, Davao, and Basilan) (Antoine 1993).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692772870FF14FD4BFCDCFC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FF0BFC89FEBD.text	3C6E87D692742873FF14FF0BFC89FEBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasyvalgus (Dasyvalgus) obsoletus Moser 1917	<div><p>Dasyvalgus (Dasyvalgus) obsoletus Moser, 1917</p><p>Dasyvalgus obsoletus Moser, 1917a: 21; Ricchiardi 1996: 372.</p><p>Dasyvalgus (Dasyvalgus) obsoletus: Krajčik 1999: 61.</p><p>Type locality. “Luzon (Imugan).”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1917a; Ricchiardi 1996).</p><p>Overall distribution. The Philippines and Indonesia.</p></div>	https://treatment.plazi.org/id/3C6E87D692742873FF14FF0BFC89FEBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FE0FFCDCFDB9.text	3C6E87D692742873FF14FE0FFCDCFDB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasyvalgus (Dasyvalgus) palawanus Ricchiardi 1996	<div><p>Dasyvalgus (Dasyvalgus) palawanus Ricchiardi, 1996</p><p>Dasyvalgus palawanus Ricchiardi, 1996: 372 .</p><p>Dasyvalgus (Dasyvalgus) palawanus: Krajčik 1999: 61.</p><p>Type locality. “ Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ricchiardi 1996).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692742873FF14FE0FFCDCFDB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FD13FCDCFC59.text	3C6E87D692742873FF14FD13FCDCFC59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasyvalgus (Dasyvalgus) panaonus Moser 1916	<div><p>Dasyvalgus (Dasyvalgus) panaonus Moser, 1916</p><p>Dasyvalgus panaonus Moser, 1916b: 155; Ricchiardi 1996: 373.</p><p>Dasyvalgus (Dasyvalgus) panaonus: Krajčik 1999: 61.</p><p>Type locality. “ Philippinen ( Panaon).”</p><p>Distribution in the Philippines. Visayas (Samar and Leyte) and Mindanao (Basilan) (Moser 1916b; Ricchiardi 1996).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692742873FF14FD13FCDCFC59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FC33FCDCFBA5.text	3C6E87D692742873FF14FC33FCDCFBA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasyvalgus (Dasyvalgus) philippinus Ricchiardi 1996	<div><p>Dasyvalgus (Dasyvalgus) philippinus Ricchiardi, 1996</p><p>Dasyvalgus philippinus Ricchiardi, 1996: 374 .</p><p>Dasyvalgus (Dasyvalgus) philippinus: Krajčik 1999: 61.</p><p>Type locality. “Negros Occ., Mount Canla-on.”</p><p>Distribution in the Philippines. Visayas (Negros) (Ricchiardi 1996).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692742873FF14FC33FCDCFBA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FB37FCDCFAA1.text	3C6E87D692742873FF14FB37FCDCFAA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dasyvalgus (Dasyvalgus) samariensis Ricchiardi 1996	<div><p>Dasyvalgus (Dasyvalgus) samariensis Ricchiardi, 1996</p><p>Dasyvalgus samariensis Ricchiardi, 1996: 375 .</p><p>Dasyvalgus (Dasyvalgus) samariensis: Krajčik 1999: 61.</p><p>Type locality. “ Samar Is.”</p><p>Distribution in the Philippines. Visayas (Samar) (Ricchiardi 1996).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692742873FF14FB37FCDCFAA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742873FF14FA73FCDCF941.text	3C6E87D692742873FF14FA73FCDCF941.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterovalgus itohi Miyake 1994	<div><p>Heterovalgus itohi Miyake, 1994</p><p>Heterovalgus itohi Miyake, 1994: 152; Krajčik 1999: 62.</p><p>Type locality. “ Luzon, Cavite.”</p><p>Distribution in the Philippines. Luzon (Cavite and Manila) (Miyake 1994).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692742873FF14FA73FCDCF941	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692742872FF14F803FAE6FE51.text	3C6E87D692742872FF14F803FAE6FE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allomyrina pfeifferi subsp. pfeifferi (Redtenbacher 1868)	<div><p>Allomyrina pfeifferi pfeifferi (Redtenbacher, 1868)</p><p>Myrina pfeifferi Redtenbacher, 1868: 79 .</p><p>Xylotrupes mindanaoensis Schultze, 1920: 191 . Type locality. “Mindanao, Surigao.”; Heller 1921: 195.</p><p>Allomyrina pfeifferi: Endrődi 1976: 236; Nakamura 1974: 6.</p><p>Type locality. “Borneo.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Schultze 1920; Heller 1921; Nakamura 1974; Endrődi 1976).</p><p>Overall distribution. The Philippines, Malaysia, Myanmar, Indonesia, and Thailand (Endrődi 1976).</p></div>	https://treatment.plazi.org/id/3C6E87D692742872FF14F803FAE6FE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FE60FCDCFD7E.text	3C6E87D692752872FF14FE60FCDCFD7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xylotrupes lumawigi Silvestre 2002	<div><p>Xylotrupes lumawigi Silvestre, 2002</p><p>Xylotrupes lumawigi Silvestre, 2002: 247; Silvestre 2006: 372.</p><p>Type locality. “ Nord Luzon, Mountain Province.”</p><p>Distribution in the Philippines. Luzon (Mountain Province) (Silvestre 2002).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692752872FF14FE60FCDCFD7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FD49FC27FCA7.text	3C6E87D692752872FF14FD49FC27FCA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xylotrupes philippinensis subsp. boudanti Silvestre 2006	<div><p>Xylotrupes philippinensis boudanti Silvestre, 2006</p><p>Xylotrupes philippinensis boudanti Silvestre, 2006: 371 .</p><p>Type locality. “ Philippines, Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Silvestre 2006).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692752872FF14FD49FC27FCA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FC36FC27FB18.text	3C6E87D692752872FF14FC36FC27FB18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xylotrupes philippinensis subsp. philippinensis Endrodi 1957	<div><p>Xylotrupes philippinensis philippinensis Endrődi, 1957</p><p>Xylotrupes gideon philippinensis Endrődi, 1957: 65; Endrődi 1976: 226; Silvestre 2002: 247.</p><p>Xylotrupes philippinensis: Silvestre 2006: 367.</p><p>Xylotrupes philippinensis philippinensis: Rowland 2011: 3.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Luzon (Marinduque, Babuyan, Sibuyan, and Mindoro), Visayas (Bohol, Leyte, and Samar), and Mindanao (Dinagat) (Endrődi 1957, 1976; Silvestre 2006).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692752872FF14FC36FC27FB18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FB73FC27FA41.text	3C6E87D692752872FF14FB73FC27FA41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xylotrupes pubescens subsp. beaudeti Silvestre 2006	<div><p>Xylotrupes pubescens beaudeti Silvestre, 2006</p><p>Xylotrupes pubescens beaudeti Silvestre, 2006: 366 .</p><p>Type locality. “ Philippines, Western Samar, Eastern Visayas. ”</p><p>Distribution in the Philippines. Visayas (Samar) (Silvestre 2006).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692752872FF14FB73FC27FA41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14FA58FC27F8FB.text	3C6E87D692752872FF14FA58FC27F8FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xylotrupes pubescens subsp. pubescens Waterhouse 1841	<div><p>Xylotrupes pubescens pubescens Waterhouse 1841</p><p>Xylotrupes pubescens Waterhouse, 1841a: 41; Waterhouse 1841b: 539; Hope 1841: 22; Silvestre 2006: 363.</p><p>Xylotrupes gideon pubescens: Endrődi 1951: 245; Endrődi 1976: 229; Endrődi 1985: 623.</p><p>Xylotrupes pubescens pubescens: Rowland 2003: 230; Rowland 2006a: 12; Rowland 2006b: 13; Rowland 2011: 3.</p><p>Type locality. “Insulas Philippinenses.”</p><p>Distribution in the Philippines. Luzon and Mindanao (Sulu) (Hope 1841; Waterhouse 1841a, 1841b; Endrődi 1951, 1976, 1985; Rowland 2003, 2006a, 2006b, 2011; Silvestre 2006).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692752872FF14FA58FC27F8FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692752872FF14F8CDFC27F823.text	3C6E87D692752872FF14F8CDFC27F823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Xylotrupes pubescens subsp. sibuyanensis Silvestre 2006	<div><p>Xylotrupes pubescens sibuyanensis Silvestre, 2006</p><p>Xylotrupes pubescens sibuyanensis Silvestre, 2006: 365 .</p><p>Type locality. “ Philippines, Romblon, Sibuyan.”</p><p>Distribution in the Philippines. Luzon (Romblon) (Silvestre 2006).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692752872FF14F8CDFC27F823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14FD49FCFAFC7B.text	3C6E87D692722875FF14FD49FCFAFC7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Blabephorus pinguis Fairmaire 1898	<div><p>Blabephorus pinguis Fairmaire, 1898</p><p>Blabephorus pinguis Fairmaire, 1898: 382; Endrődi 1974: 337; Drumont et al. 2006: 444.</p><p>Type locality. “ Sumatra: Labouan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Endrődi 1974).</p><p>Overall distribution. The Philippines, India, Indonesia, Malaysia, Myanmar, Sri Lanka, Thailand, Vietnam, and China (including Taiwan) (Drumont et al. 2006).</p></div>	https://treatment.plazi.org/id/3C6E87D692722875FF14FD49FCFAFC7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14FF43FC27FDCE.text	3C6E87D692722875FF14FF43FC27FDCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chalcosoma atlas subsp. hesperus (Erichson 1834)	<div><p>Chalcosoma atlas hesperus (Erichson, 1834)</p><p>Dynastes hesperus Erichson, 1834: 238 .</p><p>Chalcosoma atlas: Arrow 1910: 267; Nakamura 1974: 6.</p><p>Chalcosoma atlas hesperus: Endrődi 1976: 248.</p><p>Type locality. “ Die Insel Luzon.”</p><p>Distribution in the Philippines. Luzon (Babuyanes, Catanduanes, Laguna, Marinduque, Mindoro, Palawan, Romblon, and Pampanga), Visayas (Bohol, Leyte, Negros Island, and Samar), and Mindanao (Basilan, Dinagat, and Misamis Oriental) (Erichson 1834; Arrow 1910; Nakamura 1974; Endrődi 1976).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692722875FF14FF43FC27FDCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14FB8AFCDCFB18.text	3C6E87D692722875FF14FB8AFCDCFB18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clyster hirokiharukae Yamaya 2009	<div><p>Clyster hirokiharukae Yamaya, 2009</p><p>Clyster hirokiharukae Yamaya, 2009: 26 .</p><p>Type locality. “ Harihuron, Mindro Island.”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Yamaya 2009).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692722875FF14FB8AFCDCFB18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14FB73FCDCFA41.text	3C6E87D692722875FF14FB73FCDCFA41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clyster rosaemariae Dupuis 2002	<div><p>Clyster rosaemariae Dupuis, 2002</p><p>Clyster rosaemariae Dupuis, 2002: 169; Yamaya 2009: 24.</p><p>Type locality. “ Philippines, Marabut, Western Samar, Eastern Visayas. ”</p><p>Distribution in the Philippines. Visayas (Samar) (Dupuis 2002).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692722875FF14FB73FCDCFA41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722875FF14F9C8FE8CF89F.text	3C6E87D692722875FF14F9C8FE8CF89F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oryctes (Rykanes) gnu Mohnike 1874	<div><p>Oryctes (Rykanes) gnu Mohnike, 1874</p><p>Oryctes gnu Mohnike, 1874: 261; Schultze 1916a: 170; Nakamura 1974: 5.</p><p>Oryctes (Rykanes) gnu: Endrődi 1973: 47.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Mohnike 1874; Schultze 1916a; Endrődi 1973; Nakamura 1974).</p><p>Overall distribution. The Philippines, Indonesia, Malaysia, Sri Lanka, and China (Endrődi 1973; Nakamura 1974).</p></div>	https://treatment.plazi.org/id/3C6E87D692722875FF14F9C8FE8CF89F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692722874FF14F8E9FBA4FEC5.text	3C6E87D692722874FF14F8E9FBA4FEC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oryctes (Rykanes) rhinoceros (Linnaeus 1758)	<div><p>Oryctes (Rykanes) rhinoceros (Linnaeus, 1758)</p><p>Scarabaeus rhinoceros Linnaeus, 1758: 346 .</p><p>Oryctes rhinoceros: Arrow 1910: 278; Schultze 1916a: 170; Nakamura 1974: 5.</p><p>Oryctes (Rykanes) rhinoceros: Endrődi 1973: 74.</p><p>Type locality. “ Habitat in Afia. ”</p><p>Distribution in the Philippines. This species is found throughout the archipelago.</p><p>Overall distribution. This species is widespread in the Indo-Pacific region and is also found in the Palearctic, Australian, Neotropical, and Madagascan regions (Hao et al. 2022).</p></div>	https://treatment.plazi.org/id/3C6E87D692722874FF14F8E9FBA4FEC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14FC6EFCDCFB44.text	3C6E87D692732874FF14FC6EFCDCFB44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceratohyphus luzonicus Geshev 2024	<div><p>Ceratohyphus luzonicus Geshev, 2024</p><p>Ceratohyphus luzonicus Geshev, 2024: 2 .</p><p>Type locality. “Ditumabo Barangay, Baler Municipality, Aurora Prov., Luzon Is., The Philippines.”</p><p>Distribution in the Philippines. Luzon (Aurora and Quezon) (Geshev 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692732874FF14FC6EFCDCFB44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14F9C8FC55F8B3.text	3C6E87D692732874FF14F9C8FC55F8B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neohyphus celebesus Heller 1896	<div><p>Neohyphus celebesus Heller, 1896</p><p>Neohyphus celebesus Heller, 1896: 13; Kuijten 1975: 105; Kuijten 1994: 59; Geshev 2024: 5.</p><p>Neohyphus celebesus philippinus Endrődi, 1959: 122 . Type locality. “ Davao.”; Endrődi 1971a: 214 (synonymized with Neohyphus semivelutinus).</p><p>Type locality. “ Celébes centralis.”</p><p>Distribution in the Philippines. Mindanao (Davao) (Heller 1896; Endrődi, 1959; Kuijten 1994; Geshev 2024).</p><p>Overall distribution. The Philippines and Indonesia (Geshev 2024).</p></div>	https://treatment.plazi.org/id/3C6E87D692732874FF14F9C8FC55F8B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14FA8FFCDCF9FE.text	3C6E87D692732874FF14FA8FFCDCF9FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neohyphus philippinus (Arrow 1931)	<div><p>Neohyphus philippinus (Arrow, 1931)</p><p>Peltonotus philippinus Arrow, 1931: 611; Endrődi 1971a: 214 (synonymized with Neohyphus semivelutinus).</p><p>Neohyphus philippinus: Kuijten 1994: 59; Yamaya 2011: 5.</p><p>Type locality. “ Philippine Is. Mt. Makiling, Laguna.”</p><p>Distribution in the Philippines. Luzon (Laguna, Marinduque, Sibuyan, Mindoro, Aurora Province, and Sorsogon), Visayas (Panay), and Mindanao (Bukidnon) (Kuijten 1994; Yamaya 2011; Geshev 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692732874FF14FA8FFCDCF9FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14FE0CFB22FD86.text	3C6E87D692732874FF14FE0CFB22FD86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trichogomphus bronchus (Herbst 1785)	<div><p>Trichogomphus bronchus (Herbst, 1785)</p><p>Scarabaeus bronchus Herbst, 1785: 305 .</p><p>Trichogomphus bronchus: Endrődi 1974: 346; Nakamura 1974: 4.</p><p>Type locality. “ India (?).”</p><p>Distribution in the Philippines.?Philippine archipelago (Endrődi 1974).</p><p>Overall distribution. The? Philippines, Indonesia, and Malaysia (Nakamura 1974).</p></div>	https://treatment.plazi.org/id/3C6E87D692732874FF14FE0CFB22FD86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732874FF14FD11FC27FCEF.text	3C6E87D692732874FF14FD11FC27FCEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trichogomphus lunicollis subsp. fujiokai Miyake & Yamaya 1999	<div><p>Trichogomphus lunicollis fujiokai Miyake &amp; Yamaya, 1999</p><p>Trichogomphus lunicollis fujiokai Miyake &amp; Yamaya, 1999: 108 .</p><p>Type locality. “ Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Miyake &amp; Yamaya 1999).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692732874FF14FD11FC27FCEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692732877FF14F80AFCDCFF0D.text	3C6E87D692732877FF14F80AFCDCFF0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neohyphus quadridentatus Voirin 1996	<div><p>Neohyphus quadridentatus Voirin, 1996</p><p>Neohyphus quadridentatus Voirin, 1996: 215; Yamaya 2011: 5.</p><p>Type locality. “ Philippines, Balisan (= Basilan).”</p><p>Distribution in the Philippines. Mindanao (Basilan) (Voirin 1996).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692732877FF14F80AFCDCFF0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14FC36FE56FB18.text	3C6E87D692702877FF14FC36FE56FB18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alissonotum pauperum (Burmeister 1847)	<div><p>Alissonotum pauperum (Burmeister, 1847)</p><p>Heteronychus pauper Burmeister, 1847: 94 .</p><p>Alissonotum pauper: Schultze 1916a: 169; Endrődi 1969a: 424; Nakamura 1974: 1; Endrődi 1985: 320.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Luzon (Manila) (Schultze 1916a; Endrődi 1969a; Nakamura 1974).</p><p>Overall distribution. The Philippines, Japan, and China (including Taiwan) (Löbl &amp; Smetana 2006).</p><p>Genus Dipelicus Hope, 1842</p></div>	https://treatment.plazi.org/id/3C6E87D692702877FF14FC36FE56FB18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14FB73FC54FA39.text	3C6E87D692702877FF14FB73FC54FA39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dipelicus deiphobus subsp. robustus (Heller 1897)	<div><p>Dipelicus deiphobus robustus (Heller, 1897)</p><p>Camelonotus robustus Heller, 1897c:172 .</p><p>Dipelicus deiphobus robustus: Endrődi 1969: 458; Endrődi 1971b: 298; Endrődi 1985: 340.</p><p>Type locality. “ Celebes; Tondano; Macassar; Philippinen; Manila; Sumatra.”</p><p>Distribution in the Philippines. Luzon (Manila) and Mindanao (Heller 1897c; Schultze 1916a; Endrődi 1969, 1971b).</p><p>Overall distribution. The Philippines and Indonesia (Heller 1897c).</p></div>	https://treatment.plazi.org/id/3C6E87D692702877FF14FB73FC54FA39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14F990FE5DF8FB.text	3C6E87D692702877FF14F990FE5DF8FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dipelicus monotuberculatus Miyake & Yamaya 1997	<div><p>Dipelicus monotuberculatus Miyake &amp; Yamaya, 1997</p><p>Dipelicus monotuberculatus Miyake &amp; Yamaya, 1997: 16 .</p><p>Type locality. “ Sybuyan (= Sibuyan) Island, Philippines.”</p><p>Distribution in the Philippines. Luzon (Sibuyan), Visayas (Leyte and Samar), and Mindanao (Basilan) (Miyake &amp; Yamaya 1997).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Genus Papuana Arrow, 1911</p></div>	https://treatment.plazi.org/id/3C6E87D692702877FF14F990FE5DF8FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14FE9FFCDCFDCE.text	3C6E87D692702877FF14FE9FFCDCFDCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neohyphus semivelutinus (Fairmaire 1883)	<div><p>Neohyphus semivelutinus (Fairmaire, 1883)</p><p>Melanhyphus semivelutinus Fairmaire, 1883: 12 .</p><p>Neohyphus semivelutinus: Endrődi 1971a: 214; Nakamura 1974: 1; Endrődi 1985: 202; Kuijten 1994: 59; Yamaya 2011: 5.</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Bataan, Camarines Sur, Manila, Quirino, Masbate, Mt. Banahao), Visayas (Leyte), and Mindanao (Fairmaire 1883; Endrődi 1971a, 1985; Yamaya 2011; Geshev 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692702877FF14FE9FFCDCFDCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14FDD9FCDCFD37.text	3C6E87D692702877FF14FDD9FCDCFD37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neohyphus suzukii Yamaya 2011	<div><p>Neohyphus suzukii Yamaya, 2011</p><p>Neohyphus suzukii Yamaya, 2011: 9 .</p><p>Type locality. “Sibuyan Is. and Samar Is.”</p><p>Distribution in the Philippines. Luzon (Sibuyan) and Visayas (Samar) (Yamaya 2011).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692702877FF14FDD9FCDCFD37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692702877FF14F8CCFCDCF823.text	3C6E87D692702877FF14F8CCFCDCF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Papuana matsumotoi Yamaya 2009	<div><p>Papuana matsumotoi Yamaya, 2009</p><p>Papuana matsumotoi Yamaya, 2009: 25 .</p><p>Type locality. “ Mt. Canlaon, Negros Island.”</p><p>Distribution in the Philippines. Visayas (Negros Island) (Yamaya 2009).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692702877FF14F8CCFCDCF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FF0AFCDCFE98.text	3C6E87D692712876FF14FF0AFCDCFE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Papuana miyakei Yamaya 2006	<div><p>Papuana miyakei Yamaya, 2006</p><p>Papuana miyakei Yamaya, 2006: 49 .</p><p>Type locality. “Mount Apo, Mindanao Island.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Yamaya 2006).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692712876FF14FF0AFCDCFE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FEF2FB3CFDE4.text	3C6E87D692712876FF14FEF2FB3CFDE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Papuana philippinica Arrow 1937	<div><p>Papuana philippinica Arrow, 1937</p><p>Papuana philippinica Arrow, 1937: 47; Endrődi 1969a: 445; Endrődi 1971b: 255; Nakamura 1974: 2; Endrődi 1985: 322; Löbl &amp; Smetana 2006: 281.</p><p>Type locality. “ Philippinen: Luzon.”</p><p>Distribution in the Philippines. Luzon (Laguna, Benguet, Kalinga) (Arrow 1937b; Endrődi 1969a, 1971b).</p><p>Overall distribution. The Philippines and China (Taiwan) (Löbl &amp; Smetana 2006).</p></div>	https://treatment.plazi.org/id/3C6E87D692712876FF14FEF2FB3CFDE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FDF6FCDCFC84.text	3C6E87D692712876FF14FDF6FCDCFC84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Papuana uniformis (Prell 1934)	<div><p>Papuana uniformis (Prell, 1934)</p><p>Heteroclyster uniformis Prell, 1934: 163; Nakamura 1974: 4.</p><p>Papuana cognata Endrődi, 1969b: 5 . Type locality. “ Philippinen; North Luzon.”; Endrődi 1971b: 255.</p><p>Papuana uniformis: Endrődi 1985: 323.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Prell 1934; Endrődi 1969b, 1971b; Nakamura 1974).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692712876FF14FDF6FCDCFC84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FC4FFCDCFB78.text	3C6E87D692712876FF14FC4FFCDCFB78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx apoensis Miyake & Yamaya 1997	<div><p>Pseudohomonyx apoensis Miyake &amp; Yamaya, 1997</p><p>Pseudohomonyx apoensis Miyake &amp; Yamaya, 1997: 15; Geshev &amp; Pathomwattananurak 2023: 118.</p><p>Type locality. “ Mt. Apo, Mindanao, Philippines.”</p><p>Distribution in the Philippines. Mindanao (Cotabato) (Miyake &amp; Yamaya 1997; Geshev &amp; Pathomwattananurak 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692712876FF14FC4FFCDCFB78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FB53FCDCFA44.text	3C6E87D692712876FF14FB53FCDCFA44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx luzonicus Miyake & Yamaya 2000	<div><p>Pseudohomonyx luzonicus Miyake &amp; Yamaya, 2000</p><p>Pseudohomonyx luzonicus Miyake &amp; Yamaya, 2000: 7; Geshev &amp; Pathomwattananurak 2023: 122.</p><p>Type locality. “ Mt. Banahaw, Luzon Is., Philippines.”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao) (Miyake &amp; Yamaya 2000; Geshev &amp; Pathomwattananurak 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692712876FF14FB53FCDCFA44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14FA57FCDCF940.text	3C6E87D692712876FF14FA57FCDCF940.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx mindroensis Miyake & Yamaya 2000	<div><p>Pseudohomonyx mindroensis Miyake &amp; Yamaya, 2000</p><p>Pseudohomonyx mindroensis Miyake &amp; Yamaya, 2000: 9; Geshev &amp; Pathomwattananurak 2023: 122.</p><p>Type locality. “Mt. Halcon, Mindro [Mindoro] Is., Philippines ”.</p><p>Distribution in the Philippines. Luzon (Mindoro) (Miyake &amp; Yamaya 2000; Geshev &amp; Pathomwattananurak 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692712876FF14FA57FCDCF940	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692712876FF14F95BFCDCF84C.text	3C6E87D692712876FF14F95BFCDCF84C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx negros Miyake & Yamaya 1997	<div><p>Pseudohomonyx negros Miyake &amp; Yamaya, 1997</p><p>Pseudohomonyx negros Miyake &amp; Yamaya, 1997: 14; Geshev &amp; Pathomwattananurak 2023: 122.</p><p>Type locality. “ Mt. Canlaon, Mambcal, Negros Island.”</p><p>Distribution in the Philippines. Visayas (Negros Island) (Miyake &amp; Yamaya 1997; Geshev &amp; Pathomwattananurak 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692712876FF14F95BFCDCF84C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14F98FFCDCF939.text	3C6E87D6920E2809FF14F98FFCDCF939.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia adoretoides Ritsema 1897	<div><p>Apogonia adoretoides Ritsema, 1897</p><p>Apogonia adoretoides Ritsema, 1897a:123; Heller 1897a: 4; Ritsema 1904: 111; Dalla Torre 1912c:136; Bezděk 2004: 6.</p><p>Type locality. “ Luzon: Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Ritsema 1897; Heller 1897a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920E2809FF14F98FFCDCF939	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14F893FCDCF861.text	3C6E87D6920E2809FF14F893FCDCF861.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia angusta Moser 1926	<div><p>Apogonia angusta Moser, 1926</p><p>Apogonia angusta Moser, 1926: 203; Bezděk 2004: 8.</p><p>Type locality. “ Luzon: Baguio.”</p><p>Distribution in the Philippines. Luzon (Baguio) (Moser 1926).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920E2809FF14F893FCDCF861	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FC83FBAFFBEC.text	3C6E87D6920E2809FF14FC83FBAFFBEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eophileurus iwasei Muramoto 1995	<div><p>Eophileurus iwasei Muramoto, 1995</p><p>Eophileurus iwasei Muramoto, 1995: 225; Jákl &amp; Zídek 2015: 19.</p><p>Type locality. “ Mt. Apo, Mindanao Is. ”</p><p>Distribution in the Philippines. Luzon, Visayas (Leyte), and Mindanao (Mt. Apo) (Muramoto 1995; Jákl &amp; Zídek 2015; Yang et al. 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks. Yang et al. (2023) noted that the Eophileurus chinensis identified by Schultze (1916a) in the Agusan River might be a misidentified specimen of Eophileurus iwasei .</p></div>	https://treatment.plazi.org/id/3C6E87D6920E2809FF14FC83FBAFFBEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FBFFFCDCFAD5.text	3C6E87D6920E2809FF14FBFFFCDCFAD5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eophileurus varipunctatus Yang, Pathomwattananurak & Zhao 2022	<div><p>Eophileurus varipunctatus Yang, Pathomwattananurak &amp; Zhao, 2022</p><p>Eophileurus varipunctatus Yang, Pathomwattananurak &amp; Zhao, 2022: 291 .</p><p>Type locality. “ Mt. Halcon, Paitan Barangay, Naujuan Municipality, Oriental Mindoro Prov., Mindoro Is.”</p><p>Distribution in the Philippines. Luzon (Mindoro and Palawan) (Yang et al. 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920E2809FF14FBFFFCDCFAD5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FF0BFCDCFE51.text	3C6E87D6920E2809FF14FF0BFCDCFE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx palavanus Prell 1912	<div><p>Pseudohomonyx palavanus Prell, 1912</p><p>Pseudohomonyx palavanus Prell, 1912: 186; Endrődi 1969a: 440; Nakamura 1974: 2; Geshev &amp; Pathomwattananurak 2023: 123.</p><p>Type locality. “ Palawan ( Süd Palawan).”</p><p>Distribution in the Philippines. Luzon (Palawan) (Prell 1912; Endrődi 1969; Nakamura 1974; Geshev &amp; Pathomwattananurak 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920E2809FF14FF0BFCDCFE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920E2809FF14FE2BFCDCFDB9.text	3C6E87D6920E2809FF14FE2BFCDCFDB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudohomonyx samarensis Miyake & Yamaya 1997	<div><p>Pseudohomonyx samarensis Miyake &amp; Yamaya, 1997</p><p>Pseudohomonyx samarensis Miyake &amp; Yamaya, 1997: 13; Geshev &amp; Pathomwattananurak 2023: 123.</p><p>Type locality. “ Samar Island, Philippines.”</p><p>Distribution in the Philippines. Visayas (Samar) (Miyake &amp; Yamaya 1997).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920E2809FF14FE2BFCDCFDB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FF0AFCDCFE99.text	3C6E87D6920F2808FF14FF0AFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia angustipes Moser 1924	<div><p>Apogonia angustipes Moser, 1924</p><p>Apogonia angustipes Moser, 1924b: 203; Bezděk 2004: 8.</p><p>Type locality. “ Luzon, Mountain Province, Pauai ( Haight’s place).”</p><p>Distribution in the Philippines. Luzon (Mountain Province) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920F2808FF14FF0AFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FEF3FCDCFDCE.text	3C6E87D6920F2808FF14FEF3FCDCFDCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia bakeri Moser 1915	<div><p>Apogonia bakeri Moser, 1915</p><p>Apogonia Bakeri Moser, 1915a: 127; Bezděk 2004: 9.</p><p>Type locality. “ Luzon (Mt. Banahao).”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao) (Moser 1915a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920F2808FF14FEF3FCDCFDCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FDD8FCDCFD37.text	3C6E87D6920F2808FF14FDD8FCDCFD37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia balocauensis Kobayashi 2014	<div><p>Apogonia balocauensis Kobayashi, 2014</p><p>Apogonia balocauensis Kobayashi, 2014: 56 .</p><p>Type locality. “ Mt. Balocaue, Leyte Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Kobayashi 2014).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920F2808FF14FDD8FCDCFD37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FC81FCDCFC1F.text	3C6E87D6920F2808FF14FC81FCDCFC1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia banosana Moser 1924	<div><p>Apogonia banosana Moser, 1924</p><p>Apogonia banosa Moser, 1924b: 350; Bezděk 2004: 10.</p><p>Type locality. “ Luzon, Laguna Province, Los Banos.”</p><p>Distribution in the Philippines. Luzon (Los Baños, Laguna) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920F2808FF14FC81FCDCFC1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FC6EFCDCFB44.text	3C6E87D6920F2808FF14FC6EFCDCFB44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia bazilanensis Kobayashi 2014	<div><p>Apogonia bazilanensis Kobayashi, 2014</p><p>Apogonia bazilanensis Kobayashi, 2014: 50 .</p><p>Type locality. “Bazilan Is., Philippines ”</p><p>Distribution in the Philippines. Mindanao (Basilan) (Kobayashi 2014).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920F2808FF14FC6EFCDCFB44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FB56FCDCFAAD.text	3C6E87D6920F2808FF14FB56FCDCFAAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia benguetana Moser 1924	<div><p>Apogonia benguetana Moser, 1924</p><p>Apogonia benguetana Moser, 1924b: 351; Bezděk 2004: 10.</p><p>Type locality. “ Luzon, Benguet.”</p><p>Distribution in the Philippines. Luzon (Benguet) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920F2808FF14FB56FCDCFAAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14FA3FFCDCF992.text	3C6E87D6920F2808FF14FA3FFCDCF992.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia boettcheri Moser 1915	<div><p>Apogonia boettcheri Moser, 1915</p><p>Apogonia boettcheri Moser, 1915a: 123; Bezděk 2004: 11.</p><p>Type locality. “ Palawan septentrionalis ( Binaluan).”</p><p>Distribution in the Philippines. Luzon (Palawan) (Moser 1915a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920F2808FF14FA3FFCDCF992	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14F9E4FCDCF8FB.text	3C6E87D6920F2808FF14F9E4FCDCF8FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia callosifrons Moser 1924	<div><p>Apogonia callosifrons Moser, 1924</p><p>Apogonia callosifrons Moser, 1924b: 350; Bezděk 2004: 13.</p><p>Type locality. “ Luzon, Nueva Vizcaya Province, Imugan.”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920F2808FF14F9E4FCDCF8FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920F2808FF14F8CDFCDCF823.text	3C6E87D6920F2808FF14F8CDFCDCF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia carlotae Chapin 1931	<div><p>Apogonia carlotae Chapin, 1931</p><p>Apogonia carlotae Chapin, 1931b: 310; Bezděk 2004: 14.</p><p>Type locality. “La Carlota, Occidental Negros.”</p><p>Distribution in the Philippines. Visayas (Negros Occidental) (Chapin 1931b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920F2808FF14F8CDFCDCF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FF0AFCDCFE98.text	3C6E87D6920C280BFF14FF0AFCDCFE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia crassitarsis Kobayashi 2013	<div><p>Apogonia crassitarsis Kobayashi, 2013</p><p>Apogonia crassitarsis Kobayashi, 2013: 69 .</p><p>Type locality. “ Mt. Balocaue, Leyte Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Kobayashi 2013).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920C280BFF14FF0AFCDCFE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FEF2FCDCFDE4.text	3C6E87D6920C280BFF14FEF2FCDCFDE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia cuprescens Blanchard 1850	<div><p>Apogonia cuprescens Blanchard, 1850</p><p>Apogonia cuprescens Blanchard, 1850: 228; Gemminger &amp; Harold 1869: 1156; Ritsema 1891: 94; Dalla Torre 1912c: 137; Bezděk 2004: 18.</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Manila) (Blanchard 1851).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920C280BFF14FEF2FCDCFDE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FDF6FCDCFCCC.text	3C6E87D6920C280BFF14FDF6FCDCFCCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia dapitana Moser 1921	<div><p>Apogonia dapitana Moser, 1921</p><p>Apogonia dapitana Moser, 1921a: 57; Bezděk 2004: 19.</p><p>Type locality. “Mindanao (Dapitan).”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Moser 1921a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920C280BFF14FDF6FCDCFCCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FCDEFCDCFC34.text	3C6E87D6920C280BFF14FCDEFCDCFC34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia depressifrons Kobayashi 2013	<div><p>Apogonia depressifrons Kobayashi, 2013</p><p>Apogonia depressifrons Kobayashi, 2013: 62 .</p><p>Type locality. “ Mt. Pulis, N. Luzon Is., Philippines.”</p><p>Distribution in the Philippines. Luzon (Mountain Province) (Kobayashi 2013).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920C280BFF14FCDEFCDCFC34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FB86FCDCFB1C.text	3C6E87D6920C280BFF14FB86FCDCFB1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia exasperata Kobayashi 2014	<div><p>Apogonia exasperata Kobayashi, 2014</p><p>Apogonia exasperata Kobayashi, 2014: 52 .</p><p>Type locality. “ Mt. Balocaue, Leyte Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Kobayashi 2014).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920C280BFF14FB86FCDCFB1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FB6EFCDCFA44.text	3C6E87D6920C280BFF14FB6EFCDCFA44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia fallaciosa Kobayashi 2014	<div><p>Apogonia fallaciosa Kobayashi, 2014</p><p>Apogonia fallaciosa Kobayashi, 2014: 55 .</p><p>Type locality. “ Mt. Mandaragon, Negros Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Negros) (Kobayashi 2014).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920C280BFF14FB6EFCDCFA44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14FA56FCDCF9AC.text	3C6E87D6920C280BFF14FA56FCDCF9AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia fuscula Moser 1921	<div><p>Apogonia fuscula Moser, 1921</p><p>Apogonia fuscula Moser, 1921a: 57; Bezděk 2004: 23.</p><p>Type locality. “Mindanao (Dapitan).”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Moser 1921a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920C280BFF14FA56FCDCF9AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280BFF14F93EFCDCF894.text	3C6E87D6920C280BFF14F93EFCDCF894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia griseosquamosa Moser 1924	<div><p>Apogonia griseosquamosa Moser, 1924</p><p>Apogonia griseosquamosa Moser, 1924b: 354; Bezděk 2004: 24.</p><p>Type locality. “ Mindanao, Surigao Province, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920C280BFF14F93EFCDCF894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920C280AFF14F8E6FC27FF0C.text	3C6E87D6920C280AFF14F8E6FC27FF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia imugana subsp. canlaonensis Kobayashi 2014	<div><p>Apogonia imugana canlaonensis Kobayashi, 2014</p><p>Apogonia imugana canlaonensis Kobayashi, 2014: 59</p><p>Type locality. “ Mt. Canla-on, Negros Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Negros) (Kobayashi 2014).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920C280AFF14F8E6FC27FF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FE9EFC27FE2C.text	3C6E87D6920D280AFF14FE9EFC27FE2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia imugana subsp. imugana Moser 1924	<div><p>Apogonia imugana imugana Moser, 1924</p><p>Apogonia imugana Moser, 1924b: 349; Bezděk 2004: 26.</p><p>Apogonia imugana imugana: Kobayashi, 2014: 59</p><p>Type locality. “ Luzon, Nueva Vizcaya Province, Imugan.”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya, Tagaytay, Benguet, Mountain Province, and Quezon) (Moser 1924b; Kobayashi 2014).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920D280AFF14FE9EFC27FE2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FDBEFC27FD14.text	3C6E87D6920D280AFF14FDBEFC27FD14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia imugana subsp. leytecola Kobayashi 2014	<div><p>Apogonia imugana leytecola Kobayashi, 2014</p><p>Apogonia imugana leytecola Kobayashi, 2014: 59</p><p>Type locality. “ Mt. Balocaue, Leyte Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Kobayashi 2014).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920D280AFF14FDBEFC27FD14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FD66FC27FC7C.text	3C6E87D6920D280AFF14FD66FC27FC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia imugana subsp. marinduqueana Kobayashi 2014	<div><p>Apogonia imugana marinduqueana Kobayashi, 2014</p><p>Apogonia imugana marinduqueana Kobayashi, 2014: 59 .</p><p>Type locality. “ Marinduque Is., Philippines.”</p><p>Distribution in the Philippines. Luzon (Marinduque) (Kobayashi 2014).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920D280AFF14FD66FC27FC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FC4EFC27FBA4.text	3C6E87D6920D280AFF14FC4EFC27FBA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia imugana subsp. panayensis Kobayashi 2014	<div><p>Apogonia imugana panayensis Kobayashi, 2014</p><p>Apogonia imugana panayensis Kobayashi, 2014: 59 .</p><p>Type locality. “ Mt. Madla-as, Panay Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Panay) (Kobayashi 2014).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920D280AFF14FC4EFC27FBA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FB36FCDCFA8C.text	3C6E87D6920D280AFF14FB36FCDCFA8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia laevicarinata Kobayashi 2013	<div><p>Apogonia laevicarinata Kobayashi, 2013</p><p>Apogonia laevicarinata Kobayashi, 2013: 65 .</p><p>Type locality. “ Mt. Mandalagon, N. Negros Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Kobayashi 2013).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920D280AFF14FB36FCDCFA8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14FA1EFCDCF9F4.text	3C6E87D6920D280AFF14FA1EFCDCF9F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia latitarsis Moser 1924	<div><p>Apogonia latitarsis Moser, 1924</p><p>Apogonia latitarsis Moser, 1924b: 351; Bezděk 2004: 29.</p><p>Type locality. “ Mindanao, Surigao Province, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920D280AFF14FA1EFCDCF9F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14F9C7FCDCF8DC.text	3C6E87D6920D280AFF14F9C7FCDCF8DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia latiuscula Kobayashi 2013	<div><p>Apogonia latiuscula Kobayashi, 2013</p><p>Apogonia latiuscula Kobayashi, 2013: 60 .</p><p>Type locality. “ Philippines (Negros Is.).”</p><p>Distribution in the Philippines. Visayas (Negros) (Kobayashi 2013).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920D280AFF14F9C7FCDCF8DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920D280AFF14F8AFFCDCF804.text	3C6E87D6920D280AFF14F8AFFCDCF804.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia leytensis Kobayashi 2013	<div><p>Apogonia leytensis Kobayashi, 2013</p><p>Apogonia leytensis Kobayashi, 2013: 70 .</p><p>Type locality. “ Mt. Balocaue, Leyte Is.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Kobayashi 2013).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920D280AFF14F8AFFCDCF804	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FF0AFCDCFE98.text	3C6E87D6920A280DFF14FF0AFCDCFE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia ligaoensis Kobayashi 2013	<div><p>Apogonia ligaoensis Kobayashi, 2013</p><p>Apogonia ligaoensis Kobayashi, 2013: 64 .</p><p>Type locality. “Ligao City, Albay, Bicol, Luzon Is.”</p><p>Distribution in the Philippines. Luzon (Albay) (Kobayashi 2013).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920A280DFF14FF0AFCDCFE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FEF2FCDCFDC0.text	3C6E87D6920A280DFF14FEF2FCDCFDC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia lutea Moser 1910	<div><p>Apogonia lutea Moser, 1910</p><p>Apogonia lutea Moser, 1910: 188; Dalla Torre 1912c: 139; Ritsema 1912: 128; Bezděk 2004: 30.</p><p>Type locality. “ Luzon, Province of Benguet, Irisan River .”</p><p>Distribution in the Philippines. Luzon (Benguet) (Moser 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920A280DFF14FEF2FCDCFDC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FDDAFCDCFD28.text	3C6E87D6920A280DFF14FDDAFCDCFD28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia luzonica Moser 1915	<div><p>Apogonia luzonica Moser, 1915</p><p>Apogonia luzonica Moser, 1915b: 581; Bezděk 2004: 31.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Moser 1915b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920A280DFF14FDDAFCDCFD28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FC82FCDCFC10.text	3C6E87D6920A280DFF14FC82FCDCFC10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia maculipennis Moser 1924	<div><p>Apogonia maculipennis Moser, 1924</p><p>Apogonia maculipennis Moser, 1924b: 352; Bezděk 2004: 31.</p><p>Type locality. “ Luzon, Mountain Province, Pauai ( Haight’s place).”</p><p>Distribution in the Philippines. Luzon (Mountain Province) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920A280DFF14FC82FCDCFC10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FC6AFCDCFB78.text	3C6E87D6920A280DFF14FC6AFCDCFB78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia magnifica Ritsema 1897	<div><p>Apogonia magnifica Ritsema, 1897</p><p>Apogonia magnifica Ritsema, 1897: 122; Ritsema 1904: 113; Dalla Torre 1912c: 139; Bezděk 2004: 31.</p><p>Type locality. “Philippine Island.”</p><p>Distribution in the Philippines. Philippine archipelago (Ritsema 1897).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920A280DFF14FC6AFCDCFB78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FB52FCDCFAA0.text	3C6E87D6920A280DFF14FB52FCDCFAA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia mandaragonensis Kobayashi 2013	<div><p>Apogonia mandaragonensis Kobayashi, 2013</p><p>Apogonia mandaragonensis Kobayashi, 2013: 59 .</p><p>Type locality. “ Philippines (Negros Is.).”</p><p>Distribution in the Philippines. Visayas (Negros) (Kobayashi 2013).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920A280DFF14FB52FCDCFAA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14FA3AFCDCF940.text	3C6E87D6920A280DFF14FA3AFCDCF940.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia metallescens Moser 1910	<div><p>Apogonia metallescens Moser, 1910</p><p>Apogonia metallescens Moser, 1910: 186; Dalla Torre 1912c: 140; Ritsema 1912: 128; Bezděk 2004: 32; Kobayashi &amp; Medina 2024: 32.</p><p>Type locality. “Mindanao, Kamp Keithley.”</p><p>Distribution in the Philippines. Mindanao (Lanao del Sur, Agusan del Sur, Misamis Oriental, and Davao del Sur) (Moser 1910; Kobayashi &amp; Medina 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920A280DFF14FA3AFCDCF940	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920A280DFF14F95AFCDCF84C.text	3C6E87D6920A280DFF14F95AFCDCF84C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia mindanaoana Moser 1915	<div><p>Apogonia mindanaoana Moser, 1915</p><p>Apogonia mindanaoana Moser, 1915b: 581; Bezděk 2004: 32.</p><p>Type locality. “ Mindanao, Butuan.”</p><p>Distribution in the Philippines. This species is endemic to the Philippines and is found in Mindanao (Agusan del Norte) (Moser 1915b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920A280DFF14F95AFCDCF84C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FF0AFCDCFEBD.text	3C6E87D6920B280CFF14FF0AFCDCFEBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia mindanaoensis Kobayashi & Medina 2024	<div><p>Apogonia mindanaoensis Kobayashi &amp; Medina, 2024</p><p>Apogonia mindanaoensis Kobayashi &amp; Medina, 2024: 29 .</p><p>Type locality. “Kabangsalan, Bukidnon, Mindanao Is.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon and Zamboanga del Sur) (Kobayashi &amp; Medina 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920B280CFF14FF0AFCDCFEBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FE0FFCDCFDE2.text	3C6E87D6920B280CFF14FE0FFCDCFDE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia mohagani Kobayashi 2013	<div><p>Apogonia mohagani Kobayashi, 2013</p><p>Apogonia mohagani Kobayashi, 2013: 71 .</p><p>Type locality. “ Mt. Balocaue, Leyte Is.”</p><p>Distribution in the Philippines. Visayas (Leyte and Samar) (Kobayashi 2013).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920B280CFF14FE0FFCDCFDE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FDF4FCDCFCCB.text	3C6E87D6920B280CFF14FDF4FCDCFCCB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia monticola Moser 1915	<div><p>Apogonia monticola Moser, 1915</p><p>Apogonia monticola Moser, 1915b: 582; Bezděk 2004: 34.</p><p>Type locality. “ Luzon (Mt. Makiling).”</p><p>Distribution in the Philippines. Luzon (Laguna) (Moser 1915b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920B280CFF14FDF4FCDCFCCB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FCDDFCDCFC30.text	3C6E87D6920B280CFF14FCDDFCDCFC30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia negrosensis Kobayashi 2013	<div><p>Apogonia negrosensis Kobayashi, 2013</p><p>Apogonia negrosensis Kobayashi, 2013: 57 .</p><p>Type locality. “ Philippines (Negros Is.).”</p><p>Distribution in the Philippines. Visayas (Negros) (Kobayashi 2013).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920B280CFF14FCDDFCDCFC30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FB8AFCDCFB18.text	3C6E87D6920B280CFF14FB8AFCDCFB18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia nigripennis Moser 1924	<div><p>Apogonia nigripennis Moser, 1924</p><p>Apogonia nigripennis Moser, 1924b: 351; Bezděk 2004: 35.</p><p>Type locality. “Mindanao, Kolambugan.”</p><p>Distribution in the Philippines. Mindanao (Lanao del Norte) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920B280CFF14FB8AFCDCFB18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FB73FCDCFA41.text	3C6E87D6920B280CFF14FB73FCDCFA41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia nigrobrunnea Moser 1910	<div><p>Apogonia nigrobrunnea Moser, 1910</p><p>Apogonia nigrobrunnea Moser, 1910: 187; Dalla Torre 1912c: 140; Ritsema 1912: 128; Bezděk 2004: 35.</p><p>Type locality. “Luzon, Province of Benguet, Irisan River.”</p><p>Distribution in the Philippines. Luzon (Benguet) (Moser 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920B280CFF14FB73FCDCFA41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14FA5BFCDCF9B6.text	3C6E87D6920B280CFF14FA5BFCDCF9B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia oberthuerii Ritsema 1897	<div><p>Apogonia oberthuerii Ritsema, 1897</p><p>Apogonia oberthuerii Ritsema, 1897: 121; Bezděk 2004: 36.</p><p>Type locality. “ Southern Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ritsema 1897).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920B280CFF14FA5BFCDCF9B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280CFF14F900FBD9F8B3.text	3C6E87D6920B280CFF14F900FBD9F8B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia palawana Heller 1897	<div><p>Apogonia palawana Heller, 1897</p><p>Apogonia palawana Heller, 1897: 8; Ritsema 1904: 113; Dalla Torre 1912c: 141; Mohamedsaid et al. 1997: 215; Bezděk 2004: 36; Calcetas et al. 2021: 1201.</p><p>Type locality. “ Palawan u. N. Borneo. ”</p><p>Distribution in the Philippines. Luzon (Cavite and Palawan) (Heller 1896; Calcetas et al. 2021).</p><p>Overall distribution. The Philippines and Malaysia (Mohamedsaid et al. 1997).</p></div>	https://treatment.plazi.org/id/3C6E87D6920B280CFF14F900FBD9F8B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6920B280FFF14F805FCDCFF0C.text	3C6E87D6920B280FFF14F805FCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia paralaevicarinata Kobayashi 2013	<div><p>Apogonia paralaevicarinata Kobayashi, 2013</p><p>Apogonia paralaevicarinata Kobayashi, 2013: 67 .</p><p>Type locality. “ Mt. Mandalagon, N. Negros Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Negros) (Kobayashi 2013).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6920B280FFF14F805FCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FE9EFCDCFE74.text	3C6E87D69208280FFF14FE9EFCDCFE74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia pectoralis Moser 1917	<div><p>Apogonia pectoralis Moser, 1917</p><p>Apogonia pectoralis Moser, 1917: 73; Bezděk 2004: 38.</p><p>Type locality. “ Luzon (Mt. Isarog).”</p><p>Distribution in the Philippines. Luzon (Camarines Sur) (Moser 1917b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69208280FFF14FE9EFCDCFE74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FE46FCDCFD5C.text	3C6E87D69208280FFF14FE46FCDCFD5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia philippinica Moser 1921	<div><p>Apogonia philippinica Moser, 1921</p><p>Apogonia philippinica Moser, 1921a: 58; Bezděk 2004: 38.</p><p>Type locality. “Luzon (Los Banos).’</p><p>Distribution in the Philippines. Luzon (Laguna) (Moser 1921a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69208280FFF14FE46FCDCFD5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FD2EFCDCFC84.text	3C6E87D69208280FFF14FD2EFCDCFC84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia philochlaenioides Moser 1924	<div><p>Apogonia philochlaenioides Moser, 1924</p><p>Apogonia philochlaenioides Moser, 1924b: 353; Bezděk 2004: 39.</p><p>Type locality. “ Luzon, Mountain Province, Balbalasang.”</p><p>Distribution in the Philippines. Luzon (Mountain Province) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69208280FFF14FD2EFCDCFC84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FC16FCDCFBEC.text	3C6E87D69208280FFF14FC16FCDCFBEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia polisana Moser 1924	<div><p>Apogonia polisana Moser, 1924</p><p>Apogonia polisana Moser, 1924b: 352; Bezděk 2004: 39.</p><p>Type locality. “ Luzon, Mountain Province, Mount Polis. ”</p><p>Distribution in the Philippines. Luzon (Mountain Province) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69208280FFF14FC16FCDCFBEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FBFEFCDCFAE8.text	3C6E87D69208280FFF14FBFEFCDCFAE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia pseudorizali Kobayashi & Medina 2024	<div><p>Apogonia pseudorizali Kobayashi &amp; Medina, 2024</p><p>Apogonia pseudorizali Kobayashi &amp; Medina, 2024: 27 .</p><p>Type locality. “Bayog, Lanao del sur, Mindanao Is., Philippines.”</p><p>Distribution in the Philippines. Luzon (Palawan) and Mindanao (Lanao del Sur, Davao del Sur, Surigao del Sur, Zamboanga, and Bukidnon) (Kobayashi &amp; Medina 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69208280FFF14FBFEFCDCFAE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14FAC2FCDCF988.text	3C6E87D69208280FFF14FAC2FCDCF988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia rizali Heller 1897	<div><p>Apogonia rizali Heller, 1897</p><p>Apogonia rizali Heller, 1897b: 191; Dalla Torre 1912c: 141; Ritsema 1904 b: 114; Bezděk 2004: 42; Kobayashi &amp; Medina 2024: 22.</p><p>Type locality. “Insulae Philippinae, Mindanao, Dapítan.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon, Davao, Lanao del Sur, Agusan del Sur, Cotabato, Misamis Oriental, Surigao del Sur, and Zamboanga del Norte) (Heller 1897b; Kobayashi &amp; Medina 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69208280FFF14FAC2FCDCF988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280FFF14F9E2FCDCF8F0.text	3C6E87D69208280FFF14F9E2FCDCF8F0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia rugipennis Moser 1910	<div><p>Apogonia rugipennis Moser, 1910</p><p>Apogonia rugipennis Moser, 1910: 189; Dalla Torre 1912c: 141; Ritsema 1912: 128; Bezděk 2004: 43.</p><p>Type locality. “Luzon. Province of Benguet, Irisan River.”</p><p>Distribution in the Philippines. Luzon (Benguet) (Moser 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69208280FFF14F9E2FCDCF8F0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69208280EFF14F8CAFCDCFF21.text	3C6E87D69208280EFF14F8CAFCDCFF21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia rugositas Kobayashi & Medina 2024	<div><p>Apogonia rugositas Kobayashi &amp; Medina, 2024</p><p>Apogonia rugositas Kobayashi &amp; Medina, 2024: 30 .</p><p>Type locality. “Dominorog, Bukidnon, Mindanao Is., Philippines.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon, Lanao del Sur, and Agusan del Sur) (Kobayashi &amp; Medina 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69208280EFF14F8CAFCDCFF21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FEBBFCDCFE16.text	3C6E87D69209280EFF14FEBBFCDCFE16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia squamifera Moser 1915	<div><p>Apogonia squamifera Moser, 1915</p><p>Apogonia squamifera Moser, 1915a: 129; Bezděk 2004: 47.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Philippine archipelago (Moser 1915a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69209280EFF14FEBBFCDCFE16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FE60FCDCFD7E.text	3C6E87D69209280EFF14FE60FCDCFD7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia surigaoana Moser 1924	<div><p>Apogonia surigaoana Moser, 1924</p><p>Apogonia surigaoana Moser, 1924b: 352; Bezděk 2004: 49.</p><p>Type locality. “Mindanao, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69209280EFF14FE60FCDCFD7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FD49FCDCFCA7.text	3C6E87D69209280EFF14FD49FCDCFCA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia tangcolana Moser 1924	<div><p>Apogonia tangcolana Moser, 1924</p><p>Apogonia tangcolana Moser, 1924b: 350; Bezděk, 2004: 50.</p><p>Type locality. “ Mindanao, Bukidnon Province, Tangcolan.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69209280EFF14FD49FCDCFCA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FC31FCDCFB8C.text	3C6E87D69209280EFF14FC31FCDCFB8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia viridana Moser 1910	<div><p>Apogonia viridana Moser, 1910</p><p>Apogonia viridana Moser, 1910: 187; Dalla Torre 1912c: 143; Ritsema 1912: 128; Bezděk 2004: 53.</p><p>Type locality. “Luzon, Cagayan. Camalaniogan.”</p><p>Distribution in the Philippines. Luzon (Cagayan Province) (Moser 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69209280EFF14FC31FCDCFB8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FB1EFCDCFA89.text	3C6E87D69209280EFF14FB1EFCDCFA89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia viridifulva Heller 1897	<div><p>Apogonia viridifulva Heller, 1897</p><p>Apogonia viridi-fulva Heller, 1897b: 189; Ritsema 1904: 115.</p><p>Apogonia viridifulva: Dalla Torre 1912c: 143; Bezděk 2004: 53; Bezděk &amp; Hájek 2010a: 287.</p><p>Type locality. “Insulae Philippinae, Luzon.”</p><p>Distribution in the Philippines. Luzon (Heller 1897b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69209280EFF14FB1EFCDCFA89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14FAE3FCDCF9FE.text	3C6E87D69209280EFF14FAE3FCDCF9FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apogonia viridimicans Moser 1916	<div><p>Apogonia viridimicans Moser, 1916</p><p>Apogonia viridimicans Moser, 1916a: 181; Bezděk 2004: 53.</p><p>Type locality. “Luzon (Tayabas).”</p><p>Distribution in the Philippines. Luzon (Quezon Province) (Moser 1916a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69209280EFF14FAE3FCDCF9FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69209280EFF14F979FC89F823.text	3C6E87D69209280EFF14F979FC89F823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euchirus dupontianus Burmeister 1841	<div><p>Euchirus dupontianus Burmeister, 1841</p><p>Euchirus dupontianus Burmeister, 1841: 227; Burmeister 1842: 701.</p><p>Euchirus quadrilineatus Waterhouse, 1841a: 41 . Type locality. “ Philippines Islands.”; Waterhouse 1841b: 539; Hope 1841: 22.</p><p>Type locality. “ Philippines Islands.”</p><p>Distribution in the Philippines. Luzon (Catanduanes, Marinduque, and Mindoro), Visayas (Samar), and Mindanao (Dinagat) (Burmeister &amp; Schaum 1841; Waterhouse 1841 a, 1841ab).</p><p>Overall distribution. The Philippines and Indonesia.</p></div>	https://treatment.plazi.org/id/3C6E87D69209280EFF14F979FC89F823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FEF0FCDCFDCE.text	3C6E87D692062801FF14FEF0FCDCFDCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hoplia bakeri Moser 1921	<div><p>Hoplia (sensu lato) bakeri Moser, 1921</p><p>Hoplia bakeri Moser, 1921a: 69 .</p><p>Type locality. “Mindanao (Dapitan).”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Moser 1921 a, 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692062801FF14FEF0FCDCFDCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FDD9FCDCFD37.text	3C6E87D692062801FF14FDD9FCDCFD37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hoplia griseovestita Moser 1924	<div><p>Hoplia (sensu lato) griseovestita Moser, 1924</p><p>Hoplia griseovestita Moser, 1924b: 356 .</p><p>Type locality. “ Mindanao, Surigao Province, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692062801FF14FDD9FCDCFD37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FC81FCDCFC1F.text	3C6E87D692062801FF14FC81FCDCFC1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hoplia leytensis Kobayashi & Fujioka 2014	<div><p>Hoplia (sensu lato) leytensis Kobayashi &amp; Fujioka, 2014</p><p>Hoplia leytensis Kobayashi &amp; Fujioka, 2014: 26 .</p><p>Type locality. “ Balocaue, Leyte Island, Philippines.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Kobayashi &amp; Fujioka 2014).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692062801FF14FC81FCDCFC1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FC6EFCDCFB44.text	3C6E87D692062801FF14FC6EFCDCFB44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hoplia luzonica Moser 1924	<div><p>Hoplia (sensu lato) luzonica Moser, 1924</p><p>Hoplia luzonica Moser, 1924b: 356 .</p><p>Type locality. “Luzon, Mountain Province, Baguio.”</p><p>Distribution in the Philippines. Luzon (Baguio and Mountain Province) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692062801FF14FC6EFCDCFB44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FB57FCDCFAAD.text	3C6E87D692062801FF14FB57FCDCFAAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hoplia maculifera Moser 1910	<div><p>Hoplia (sensu lato) maculifera Moser, 1910</p><p>Hoplia maculifera Moser, 1910: 185 .</p><p>Type locality. “ Luzon, Province of Benguet, Irisan River, P. I.”</p><p>Distribution in the Philippines. Luzon (Benguet) (Moser 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692062801FF14FB57FCDCFAAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14FA3CFCDCF992.text	3C6E87D692062801FF14FA3CFCDCF992.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hoplia mindoroensis Kobayashi & Fujioka 2014	<div><p>Hoplia (sensu lato) mindoroensis Kobayashi &amp; Fujioka, 2014</p><p>Hoplia mindoroensis Kobayashi &amp; Fujioka, 2014: 25 .</p><p>Type locality. “ Mt. Halcon, Mindoro Island, Philippines.”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Kobayashi &amp; Fujioka 2014).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692062801FF14FA3CFCDCF992	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14F9E5FCDCF8FB.text	3C6E87D692062801FF14F9E5FCDCF8FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hoplia negrosensis Kobayashi & Fujioka 2014	<div><p>Hoplia (sensu lato) negrosensis Kobayashi &amp; Fujioka, 2014</p><p>Hoplia negrosensis Kobayashi &amp; Fujioka, 2014: 29 .</p><p>Type locality. “ Mt. Canla-on, Negros Island, Philippines.”</p><p>Distribution in the Philippines. Visayas (Negros Island) (Kobayashi &amp; Fujioka 2014).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692062801FF14F9E5FCDCF8FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692062801FF14F8CDFCDCF823.text	3C6E87D692062801FF14F8CDFCDCF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hoplia philippinensis Moser 1910	<div><p>Hoplia (sensu lato) philippinensis Moser, 1910</p><p>Hoplia philippinensis Moser, 1910: 184 .</p><p>Type locality. “Negros Occidental, Maao, P. I. ”</p><p>Distribution in the Philippines. Visayas (Negros Occidental) (Moser 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692062801FF14F8CDFCDCF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FC33FCDCFB30.text	3C6E87D692072800FF14FC33FCDCFB30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Exopholis philippinica (Brenske 1896)	<div><p>Exopholis philippinica (Brenske, 1896)</p><p>Exopholis philippinica Brenske, 1896: 179; Dalla Torre 1912c: 169; Calcetas et al. 2020: 529.</p><p>Lepidiota corpulenta Moser, 1910: 185 . Type locality. “Mindanao, Camp Keithley, P.I.”; Dalla Torre 1912c: 170.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Luzon (Nueva Ecija and Camarines Sur), Visayas, and Mindanao (Misamis Oriental, Lanao del Sur, Bukidnon, Davao del Norte, Davao del Sur, and Cotabato) (Brenske 1896; Moser 1910; Calcetas et al. 2020).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692072800FF14FC33FCDCFB30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FF0BFCDCFE98.text	3C6E87D692072800FF14FF0BFCDCFE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hoplia sibuyana Moser 1924	<div><p>Hoplia (sensu lato) sibuyana Moser, 1924</p><p>Hoplia sibuyana Moser, 1924b: 164 .</p><p>Type locality. “ Philippinen: Ins. Sibuyan.”</p><p>Distribution in the Philippines. Luzon (Sibuyan Island) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692072800FF14FF0BFCDCFE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FEF3FCDCFDC0.text	3C6E87D692072800FF14FEF3FCDCFDC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hoplia simplex Sharp 1876	<div><p>Hoplia (sensu lato) simplex Sharp, 1876</p><p>Hoplia simplex Sharp, 1876: 66 .</p><p>Type locality. “ Philippines Islands.”</p><p>Distribution in the Philippines. Philippine archipelago (Sharp 1876).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692072800FF14FEF3FCDCFDC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FDDBFCDCFD28.text	3C6E87D692072800FF14FDDBFCDCFD28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hoplia variabilis Kobayashi & Fujioka 2014	<div><p>Hoplia (sensu lato) variabilis Kobayashi &amp; Fujioka, 2014</p><p>Hoplia variabilis Kobayashi &amp; Fujioka, 2014: 28 .</p><p>Type locality. “ Balocaue, Leyte Island, Philippines.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Kobayashi &amp; Fujioka 2014).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692072800FF14FDDBFCDCFD28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14FAC3FCDCF9D0.text	3C6E87D692072800FF14FAC3FCDCF9D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidiota blanchardi Dalla Torre 1912	<div><p>Lepidiota blanchardi Dalla Torre, 1912</p><p>Lepidiota blanchardi Dalla Torre, 1912c: 170 .</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Dalla Torre 1912c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692072800FF14FAC3FCDCF9D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14F9ABFB4CF938.text	3C6E87D692072800FF14F9ABFB4CF938.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidiota munda Sharp 1876	<div><p>Lepidiota munda Sharp, 1876</p><p>Lepidiota munda Sharp, 1876: 71; Dalla Torre 1912c: 171; Schultze 1916a: 176.</p><p>Type locality. “ Sarawak.”</p><p>Distribution in the Philippines. Luzon (Brenske 1894a; Schultze 1916a).</p><p>Overall distribution. The Philippines and Malaysia (Sharp 1876; Calcetas &amp; Adorada 2017).</p></div>	https://treatment.plazi.org/id/3C6E87D692072800FF14F9ABFB4CF938	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692072800FF14F893FCDCF804.text	3C6E87D692072800FF14F893FCDCF804.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidiota philippinica (Burmeister 1855)	<div><p>Lepidiota philippinica (Burmeister, 1855)</p><p>Leucopholis philippinica Burmeister, 1855: 537 .</p><p>Lepidiota philippinica: Dalla Torre 1912c: 172.</p><p>Type locality. “ Manila.”</p><p>Distribution in the Philippines. Luzon (Manila) (Burmeister 1855; Dalla Torre 1912c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692072800FF14F893FCDCF804	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FF0BFCDCFEBD.text	3C6E87D692042803FF14FF0BFCDCFEBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lepidiota punctum Blanchard 1851	<div><p>Lepidiota punctum Blanchard, 1851</p><p>Lepidiota punctum Blanchard, 1850: 157; Dalla Torre 1912c: 172; Schultze 1916a: 172.</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Cagayan Province and Manila) and Visayas (Negros Occidental) (Blanchard 1850; Schultze 1916a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692042803FF14FF0BFCDCFEBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FE44FCDCFD7E.text	3C6E87D692042803FF14FE44FCDCFD7E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis bakeri Moser 1924	<div><p>Leucopholis bakeri Moser, 1924</p><p>Leucopholis bakeri Moser, 1924b: 354; Calcetas &amp; Adorada 2017: 90; Calcetas 2023: 200.</p><p>Type locality. “Mindanao, Dapitan.”</p><p>Distribution in the Philippines. Mindanao (Misamis Occidental and Zamboanga del Norte) (Moser 1924b; Calcetas &amp; Adorada 2017).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692042803FF14FE44FCDCFD7E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FD49FCDCFC7B.text	3C6E87D692042803FF14FD49FCDCFC7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis bezdeki Calcetas 2023	<div><p>Leucopholis bezdeki Calcetas, 2023</p><p>Leucopholis bezdeki Calcetas, 2023: 190; Matsumoto 2024: 34.</p><p>Type locality. “ Philippines (Mindanao, Zamboanga City).”</p><p>Distribution in the Philippines. Mindanao (Zamboanga City and Basilan) (Calcetas 2023; Matsumoto 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692042803FF14FD49FCDCFC7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FC52FCDCFBA0.text	3C6E87D692042803FF14FC52FCDCFBA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis fontainei Brenske 1894	<div><p>Leucopholis fontainei Brenske, 1894</p><p>Leucopholis fontainei Brenske, 1894a: 31; Dalla Torre 1912: 177; Frey 1963: 551.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Mindanao (Zamboanga) (Brenske 1894a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692042803FF14FC52FCDCFBA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FB3BFCDCFA89.text	3C6E87D692042803FF14FB3BFCDCFA89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis guevarai Calcetas & Adorada 2017	<div><p>Leucopholis guevarai Calcetas &amp; Adorada, 2017</p><p>Leucopholis guevarai Calcetas &amp; Adorada, 2017: 96; Calcetas 2023: 200.</p><p>Type locality. “ Mindanao: Sulu: Basilan Island.”</p><p>Distribution in the Philippines. Mindanao (Sulu Archipelago) (Calcetas &amp; Adorada 2017).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692042803FF14FB3BFCDCFA89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14FAE0FCDCF9FE.text	3C6E87D692042803FF14FAE0FCDCF9FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis hirtiventris Frey 1963	<div><p>Leucopholis hirtiventris Frey, 1963</p><p>Leucopholis hirtiventris Frey, 1963: 550 .</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Philippine archipelago (Frey 1963).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692042803FF14FAE0FCDCF9FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692042803FF14F9C8FCDCF823.text	3C6E87D692042803FF14F9C8FCDCF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis irrorata Chevrolat 1841	<div><p>Leucopholis irrorata Chevrolat, 1841</p><p>Leucopholis irrorata Chevrolat, 1841: 222; Dalla Torre 1912c: 177; Frey 1963: 551; Calcetas &amp; Adorada 2017: 93; Calcetas, 2023: 200.</p><p>Leucopholis simillima Blanchard, 1851: 158 . Type locality. “ Manille.”</p><p>Leucopholis pollinosa Burmeister, 1855: 304 . Type locality. “Luzon.”</p><p>Leucopholis pollinosa furfurosa Burmeister, 1855: 304 .</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Antique, Batanes, Cavite, Laguna, Marinduque, Manila, Masbate, Quezon, and Romblon), Visayas (Cebu, Leyte, Negros Island, and Samar), and Mindanao (Bukidnon, Cotabato, Davao del Sur, Davao Oriental, Misamis Oriental, and Surigao del Sur) (Chevrolat 1841; Blanchard 1850; Burmeister 1855; Calcetas &amp; Adorada 2017).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692042803FF14F9C8FCDCF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FF0BFB02FEBC.text	3C6E87D692052802FF14FF0BFB02FEBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis jacquinoti Blanchard 1851	<div><p>Leucopholis jacquinoti Blanchard, 1851</p><p>Leucopholis jacquinoti Blanchard, 1851: 158; Brenske 1896: 187; Dalla Torre 1912c: 177; Frey 1963: 551.</p><p>Type locality. “Samboangan (ǐle Mindanao).”</p><p>Distribution in the Philippines. Mindanao (Sulu and Zamboanga City) (Blanchard 1851; Brenske 1896).</p><p>Overall distribution. The Philippines and Indonesia (Blanchard 1851; Brenske 1896).</p></div>	https://treatment.plazi.org/id/3C6E87D692052802FF14FF0BFB02FEBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FE0FFBDCFDE4.text	3C6E87D692052802FF14FE0FFBDCFDE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis pollens Sharp 1876	<div><p>Leucopholis pollens Sharp, 1876</p><p>Leucopholis pollens Sharp, 1876: 80; Schultze 1916a: 179.</p><p>Type locality. “Borneo.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Schultze 1916a; Calcetas 2023).</p><p>Overall distribution. The Philippines and Borneo (Sharp, 1876; Calcetas 2023).</p></div>	https://treatment.plazi.org/id/3C6E87D692052802FF14FE0FFBDCFDE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FDF7FCDCFC7C.text	3C6E87D692052802FF14FDF7FCDCFC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis pulverulenta Burmeister 1855	<div><p>Leucopholis pulverulenta Burmeister, 1855</p><p>Leucopholis pulverulenta Burmeister, 1855: 305; Frey 1963: 551; Calcetas &amp; Adorada 2017: 98; Calcetas 2023: 200.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Albay, Antique, Bataan, Batanes, Batangas, Camarines Norte, Cavite, Ilocos Sur, Isabela, Laguna, La Union, Manila, Marinduque, Mountain Province, Oriental Mindoro, Pangasinan, Quezon City, Rizal, Sorsogon, and Zambales), Visayas (Negros Island and Bohol), and Mindanao (Davao) (Burmeister 1855; Brenske 1894a; Frey 1963; Calcetas &amp; Adorada 2017).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692052802FF14FDF7FCDCFC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FC4FFCDCFB78.text	3C6E87D692052802FF14FC4FFCDCFB78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis ratcliffei Calcetas 2023	<div><p>Leucopholis ratcliffei Calcetas, 2023</p><p>Leucopholis ratcliffei Calcetas, 2023: 197; Matsumoto 2024: 36.</p><p>Type locality. “ Philippines ( Mindanao, Misamis Oriental, Cagayan de Oro City).”</p><p>Distribution in the Philippines. Mindanao (Bukidnon and Misamis Oriental) (Calcetas 2023; Matsumoto 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692052802FF14FC4FFCDCFB78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FB53FCDCFA44.text	3C6E87D692052802FF14FB53FCDCFA44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis reflexa Moser 1924	<div><p>Leucopholis reflexa Moser, 1924</p><p>Leucopholis reflexa Moser, 1924b: 354; Calcetas &amp; Adorada 2017: 90; Calcetas 2023: 200.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Luzon (Marinduque), Visayas (Negros Oriental), and Mindanao (Misamis Oriental) (Moser 1924b; Calcetas &amp; Adorada 2017).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692052802FF14FB53FCDCFA44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14FA57FCDCF964.text	3C6E87D692052802FF14FA57FCDCF964.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis semperi Brenske 1896	<div><p>Leucopholis semperi Brenske, 1896</p><p>Leucopholis semperi Brenske, 1896: 194; Dalla Torre 1912c: 178; Schultze 1916a: 179; Frey 1963: 551; Calcetas 2023: 192; Matsumoto 2024: 36.</p><p>Type locality. “ Philippinen, Dugang.”</p><p>Distribution in the Philippines. Visayas (Leyte) and Mindanao (Mt. Apo and Surigao) (Brenske 1896; Calcetas 2023; Matsumoto 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692052802FF14FA57FCDCF964	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692052802FF14F977FCDCF84C.text	3C6E87D692052802FF14F977FCDCF84C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucopholis stainesi Calcetas 2023	<div><p>Leucopholis stainesi Calcetas, 2023</p><p>Leucopholis stainesi Calcetas, 2023: 186 .</p><p>Type locality. “ Philippines ( Mindanao, South Cotabato).”</p><p>Distribution in the Philippines. Mindanao (South Cotabato) (Calcetas 2023).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692052802FF14F977FCDCF84C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14F8AFFCDCF805.text	3C6E87D692022805FF14F8AFFCDCF805.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carlschoenherria adoradae Calcetas 2019	<div><p>Carlschoenherria adoradae Calcetas, 2019</p><p>Carlschoenherria adoradae Calcetas, 2019: 331 .</p><p>Type locality. “ The Philippines, Luzon Island, Kalinga Apayao.”</p><p>Distribution in the Philippines. Luzon (Apayao) (Calcetas 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692022805FF14F8AFFCDCF805	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FF43FDD9FE2D.text	3C6E87D692022805FF14FF43FDD9FE2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philacelota leucothea Prokofiev 2019	<div><p>Philacelota leucothea Prokofiev, 2019</p><p>Philacelota leucothea Prokofiev, 2019: 6; Zídek 2021: 400.</p><p>Type locality. “N O Luzon/ Philippines.”</p><p>Distribution in the Philippines. Luzon (Prokofiev 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks. Zídek (2021) suspected that P. leucothea belongs to a different genus due to its “less numerous and densely squamosal elytral ribs and short pronotal sulcus,” which make the species appear taxonomically unfit for its current classification.</p></div>	https://treatment.plazi.org/id/3C6E87D692022805FF14FF43FDD9FE2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FDF7FCDCFC85.text	3C6E87D692022805FF14FDF7FCDCFC85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philgertia lii (Keith 2006)	<div><p>Philgertia lii (Keith, 2006)</p><p>Engertia lii Keith, 2006: 91; Prokofiev 2019: 12.</p><p>Philgertia lii: Zídek 2021: 404.</p><p>Type locality. “ Philippines (Leyte, Kabeng, Palawan Is.).”</p><p>Distribution in the Philippines. Luzon (Palawan), Visayas (Leyte), and Mindanao (Agusan del Sur, Davao, and Misamis Oriental) (Keith 2006; Matsumoto 2010b; Prokofiev 2019; Zídek 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692022805FF14FDF7FCDCFC85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FC4FFCDCFBA5.text	3C6E87D692022805FF14FC4FFCDCFBA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stephanopholis lopezi Chapin 1931	<div><p>Stephanopholis lopezi Chapin, 1931</p><p>Stephanopholis lopezi Chapin, 1931b: 309 .</p><p>Type locality. “La Carlota, Occidental Negros.”</p><p>Distribution in the Philippines. Visayas (Negros Occidental) (Chapin 1931b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692022805FF14FC4FFCDCFBA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FB37FB91FA45.text	3C6E87D692022805FF14FB37FB91FA45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stephanopholis melolonthoides (Brenske 1892)	<div><p>Stephanopholis melolonthoides (Brenske, 1892)</p><p>Rhabdopholis melolonthoides Brenske, 1892c: 157 .</p><p>Type locality. “ Manila.”</p><p>Distribution in the Philippines. Luzon (Manila and Benguet) (Brenske 1892c; Itoh 1993a).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks. This species was originally collected from the Philippines, not from South America or South Africa. Itoh (1993) later confirmed that it is indeed from the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692022805FF14FB37FB91FA45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692022805FF14FA57FCDCF9AD.text	3C6E87D692022805FF14FA57FCDCF9AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stephanopholis philippinensis Brenske 1896	<div><p>Stephanopholis philippinensis Brenske, 1896</p><p>Stephanopholis philippinensis Brenske, 1896: 180; Dalla Torre 1912c: 166.</p><p>Type locality. “ Philippinen ”</p><p>Distribution in the Philippines. Philippine archipelago (Brenske 1896).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692022805FF14FA57FCDCF9AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14F977FCDCF860.text	3C6E87D692032804FF14F977FCDCF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia angulicalcarata (Itoh 2003)	<div><p>Amphitrichia angulicalcarata (Itoh, 2003)</p><p>Holotrichia angulicalcarata Itoh, 2003b: 371 .</p><p>Amphitrichia angulicalcarata: Matsumoto 2015: 16.</p><p>Type locality. “ Is. Mindoro, Philippines.”</p><p>Distribution in the Philippines. Luzon (Mindoro and Palawan) (Itoh 2003b; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692032804FF14F977FCDCF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FF0BFCDCFE98.text	3C6E87D692032804FF14FF0BFCDCFE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carlschoenherria gapudi Calcetas 2019	<div><p>Carlschoenherria gapudi Calcetas, 2019</p><p>Carlschoenherria gapudi Calcetas, 2019: 335 .</p><p>Type locality. “ Philippines, Mindanao Island .”</p><p>Distribution in the Philippines. Mindanao (Calcetas 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692032804FF14FF0BFCDCFE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FEF3FCDCFDC0.text	3C6E87D692032804FF14FEF3FCDCFDC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carlschoenherria hadsallae Calcetas 2019	<div><p>Carlschoenherria hadsallae Calcetas, 2019</p><p>Carlschoenherria hadsallae Calcetas, 2019: 333 .</p><p>Type locality. “ The Philippines, Luzon Island, Quezon.”</p><p>Distribution in the Philippines. Luzon (Quezon Province) (Calcetas 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692032804FF14FEF3FCDCFDC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FDDBFCDCFC84.text	3C6E87D692032804FF14FDDBFCDCFC84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carlschoenherria palawana (Moser 1915)	<div><p>Carlschoenherria palawana (Moser, 1915)</p><p>Schöenherria palawana Moser, 1915: 149; Schultze 1916a: 180.</p><p>Schoenherria palawana: Itoh 1992: 10; Itoh 1993b: 10.</p><p>Carlschoenherria palawana: Calcetas 2019: 339.</p><p>Type locality. “ Palawan ( Binaluan).”</p><p>Distribution in the Philippines. Luzon (Mindoro, Palawan, Romblon, and Sibuyan) and Visayas (Leyte, Negros Island, and Samar) (Moser 1915a; Calcetas 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692032804FF14FDDBFCDCFC84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FC17FCDCFB78.text	3C6E87D692032804FF14FC17FCDCFB78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carlschoenherria philippinica (Brenske 1894)	<div><p>Carlschoenherria philippinica (Brenske, 1894)</p><p>Schönherria hispida var. philippinica Brenske, 1894: 27; Schultze 1916a: 180.</p><p>Schoenherria philippinica: Itoh 1992: 9 (valid species); Itoh 1993b: 6.</p><p>Carlschoenherria philippinica: Calcetas 2019: 340.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Visayas (Leyte) and Mindanao (Bukidnon and Surigao) (Brenske 1894a; Itoh 1992, 1993b; Calcetas 2019).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692032804FF14FC17FCDCFB78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692032804FF14FB53FAEAF9F4.text	3C6E87D692032804FF14FB53FAEAF9F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carlschoenherria sulcipennis (Laporte 1840)	<div><p>Carlschoenherria sulcipennis (Laporte, 1840)</p><p>Melolontha sulcipennis Laporte, 1840: 131; Blanchard 1851: 160.</p><p>Schönherria sulcipennis: Burmeister 1855: 418; Schultze 1916a: 180.</p><p>Schoenherria sulcipennis: Itoh 1992: 9; Itoh 1993b: 4.</p><p>Carlschoenherria sulcipennis: Calcetas 2019: 337.</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Albay, Batangas, Catanduanes, Cavite, Iloilo, Isabela, Laguna, Masbate, Polillo, and Quezon Province), Visayas (Negros Island and Leyte), and Mindanao (Basilan, Dinagat, and Surigao) (Laporte 1840; Blanchard 1851; Matsumoto 2010b; Calcetas 2019).</p><p>Overall distribution. The Philippines and Indonesia (Laporte 1840; Blanchard 1851; Calcetas 2019).</p></div>	https://treatment.plazi.org/id/3C6E87D692032804FF14FB53FAEAF9F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FF0BFCDCFEBD.text	3C6E87D692002807FF14FF0BFCDCFEBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia apoensis (Matsumoto 2008)	<div><p>Amphitrichia apoensis (Matsumoto, 2008)</p><p>Holotrichia apoensis Matsumoto, 2008b: 16 .</p><p>Amphitrichia apoensis: Matsumoto 2015: 16.</p><p>Type locality. “ Mt. Apo, Mindanao Island.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo and Mt. Melibengoy) (Matsumoto 2008b, 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692002807FF14FF0BFCDCFEBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FE0FFCDCFDB9.text	3C6E87D692002807FF14FE0FFCDCFDB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia bakeri (Moser 1924)	<div><p>Amphitrichia bakeri (Moser, 1924)</p><p>Holotrichia bakeri Moser, 1924b: 355 .</p><p>Amphitrichia bakeri: Matsumoto 2015: 16.</p><p>Type locality. “Luzon, Mountain Province, Baguio.”</p><p>Distribution in the Philippines. Luzon (Baguio and Mountain Province) (Moser 1924b; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692002807FF14FE0FFCDCFDB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FD13FCDCFC85.text	3C6E87D692002807FF14FD13FCDCFC85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia banahaoensis (Moser 1924)	<div><p>Amphitrichia banahaoensis (Moser, 1924)</p><p>Holotrichia banahaoensis Moser, 1924b: 355 .</p><p>Amphitrichia banahaoensis: Matsumoto 2015: 16.</p><p>Type locality. “ Luzon, Mount Banahao. ”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao) (Moser 1924b; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692002807FF14FD13FCDCFC85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FC17FCDCFB79.text	3C6E87D692002807FF14FC17FCDCFB79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia barda (Brenske 1894)	<div><p>Amphitrichia barda (Brenske, 1894)</p><p>Holotrichia barda Brenske, 1894b: 358; Dalla Torre 1912c: 210.</p><p>Phyllophaga bardana Saylor, 1937: 321 .</p><p>Amphitrichia barda: Matsumoto 2015: 16.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Visayas (Leyte) and Mindanao (Brenske 1894b; Saylor 1937; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692002807FF14FC17FCDCFB79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FB53FCDCFA45.text	3C6E87D692002807FF14FB53FCDCFA45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia bicallosicephala (Matsumoto 2010)	<div><p>Amphitrichia bicallosicephala (Matsumoto, 2010)</p><p>Holotrichia bicallosicephala Matsumoto, 2010: 43 .</p><p>Amphitrichia bicallosicephala: Matsumoto 2015: 16.</p><p>Type locality. “ Mt. Halcon, Mindoro, Philippines.”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Matsumoto 2010, 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692002807FF14FB53FCDCFA45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14FA57FCDCF939.text	3C6E87D692002807FF14FA57FCDCF939.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia bipunctata (Brenske 1892)	<div><p>Amphitrichia bipunctata (Brenske, 1892)</p><p>Holotrichia bipunctata Brenske, 1892b: 187; Dalla Torre 1912c: 210; Matsumoto 2008b: 16.</p><p>Holotrichia bipunctata minor Brenske, 1894: 23 . Type locality. “ Philippinen.”</p><p>Amphitrichia bipunctata: Matsumoto 2015: 16.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao, Cavite, Manila, and Nueva Ecija) and Mindanao (Brenske 1892a, 1894a; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692002807FF14FA57FCDCF939	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692002807FF14F893FCDCF805.text	3C6E87D692002807FF14F893FCDCF805.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia canlaonensis (Matsumoto 2008)	<div><p>Amphitrichia canlaonensis (Matsumoto, 2008)</p><p>Holotrichia canlaonensis Matsumoto, 2008c: 101 .</p><p>Amphitrichia canlaonensis: Matsumoto 2015: 16.</p><p>Type locality. “ Mt. Canla-on, Negros Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Negros Island) (Matsumoto 2008c, 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692002807FF14F893FCDCF805	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FF0BFCDCFEBC.text	3C6E87D692012806FF14FF0BFCDCFEBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia cebuana (Itoh 2003)	<div><p>Amphitrichia cebuana (Itoh, 2003)</p><p>Holotrichia cebuana Itoh, 2003b: 374 .</p><p>Amphitrichia cebuana: Matsumoto 2015: 16.</p><p>Type locality. “ Southern Cebu, Philippines.”</p><p>Distribution in the Philippines. Visayas (Cebu) (Itoh 2003b; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692012806FF14FF0BFCDCFEBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FE0FFCDCFD5C.text	3C6E87D692012806FF14FE0FFCDCFD5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia dannymohagani (Matsumoto 2008)	<div><p>Amphitrichia dannymohagani (Matsumoto, 2008)</p><p>Holotrichia dannymohagani Matsumoto, 2008c: 106 .</p><p>Amphitrichia dannymohagani: Matsumoto 2015: 16; Matsumoto 2022: 31.</p><p>Type locality. “ Mt. Canla-on, Negros.”</p><p>Distribution in the Philippines. Visayas (Leyte, Negros, and Panay) and Mindanao (Bukidnon) (Matsumoto 2008c, 2015, 2022).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692012806FF14FE0FFCDCFD5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FD2FFCDCFC7C.text	3C6E87D692012806FF14FD2FFCDCFC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia excavaticollis (Itoh 2003)	<div><p>Amphitrichia excavaticollis (Itoh, 2003)</p><p>Holotrichia excavaticollis Itoh, 2003a: 8 .</p><p>Amphitrichia excavaticollis: Matsumoto 2015: 16.</p><p>Type locality. “ Palawan, Dumaran, Cuyo, Panay, and Mindoro Isls.”</p><p>Distribution in the Philippines. Luzon (Mindoro and Palawan) and Visayas (Panay) (Itoh 2003a; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692012806FF14FD2FFCDCFC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FC4FFCDCFB1C.text	3C6E87D692012806FF14FC4FFCDCFB1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia flachi (Brenske 1892)	<div><p>Amphitrichia flachi (Brenske, 1892)</p><p>Holotrichia flachi Brenske, 1892b: 165; Brenske 1894b: 358.</p><p>Amphitrichia flachi: Matsumoto 2015: 16.</p><p>Type locality. “ Manilla.”</p><p>Distribution in the Philippines. Luzon (Batangas, Manila, Pangasinan, and Quezon Province) and Mindanao (Bukidnon) (Brenske 1892b, 1894a; Matsumoto 2015; Calcetas et al. 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692012806FF14FC4FFCDCFB1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FA73FCDCF964.text	3C6E87D692012806FF14FA73FCDCF964.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia imugana (Moser 1924)	<div><p>Amphitrichia imugana (Moser, 1924)</p><p>Holotrichia imugana Moser, 1924b: 354 .</p><p>Amphitrichia imugana: Matsumoto 2015: 16.</p><p>Type locality. “ Luzon, Nueva Vizcaya Province, Imugan.”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Moser 1924b; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692012806FF14FA73FCDCF964	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14FB6FFCDCFA18.text	3C6E87D692012806FF14FB6FFCDCFA18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia latecostata (Moser 1911)	<div><p>Amphitrichia latecostata (Moser, 1911)</p><p>Holotrichia latecostata Moser, 1911a: 331 .</p><p>Amphitrichia latecostata: Matsumoto 2015: 16.</p><p>Type locality. “ Palawan, Bacuit.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Moser 1911a; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692012806FF14FB6FFCDCFA18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692012806FF14F977FCDCF860.text	3C6E87D692012806FF14F977FCDCF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia luzonica (Moser 1926)	<div><p>Amphitrichia luzonica (Moser, 1926)</p><p>Holotrichia luzonica Moser, 1926: 203 .</p><p>Amphitrichia luzonica: Matsumoto 2015: 16.</p><p>Type locality. “ Luzon: St. Tomas. ”</p><p>Distribution in the Philippines. Luzon (Batangas) (Moser 1926; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692012806FF14F977FCDCF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FF0BFCDCFEBC.text	3C6E87D6921E2819FF14FF0BFCDCFEBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia malindangensis (Matsumoto 2010)	<div><p>Amphitrichia malindangensis (Matsumoto, 2010)</p><p>Holotrichia malindangensis Matsumoto, 2010: 48 .</p><p>Amphitrichia malindangensis: Matsumoto 2015: 16.</p><p>Type locality. “ Mt. Malindang, Mindanao Is., Philippines.”</p><p>Distribution in the Philippines. Mindanao (Misamis Occidental) (Matsumoto 2010, 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921E2819FF14FF0BFCDCFEBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FE0FFCDCFDB8.text	3C6E87D6921E2819FF14FE0FFCDCFDB8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia mindanaona (Brenske 1894)	<div><p>Amphitrichia mindanaona (Brenske, 1894)</p><p>Holotrichia mindanaona Brenske, 1894b: 358; Dalla Torre 1912c: 204.</p><p>Amphitrichia mindanaona: Matsumoto 2015: 165.</p><p>Type locality. “Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Misamis Occidental) (Brenske 1894b; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921E2819FF14FE0FFCDCFDB8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FD13FCDCFC84.text	3C6E87D6921E2819FF14FD13FCDCFC84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia mindoroensis (Matsumoto 2010)	<div><p>Amphitrichia mindoroensis (Matsumoto, 2010)</p><p>Holotrichia mindoroensis Matsumoto, 2010: 46 .</p><p>Amphitrichia mindoroensis: Matsumoto 2015: 16.</p><p>Type locality. “ Mt. Halcon, Is. Mindoro. ”</p><p>Distribution in the Philippines. Luzon (Mindoro, Palawan, and Quezon) (Matsumoto 2010, 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921E2819FF14FD13FCDCFC84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FC17FCDCFB80.text	3C6E87D6921E2819FF14FC17FCDCFB80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia monticola (Moser 1915)	<div><p>Amphitrichia monticola (Moser, 1915)</p><p>Holotrichia monticola Moser, 1915b: 583 .</p><p>Amphitrichia monticola: Matsumoto 2015 b: 16.</p><p>Type locality. “ Luzon (Mt. Makiling).”</p><p>Distribution in the Philippines. Luzon (Laguna) (Moser 1915b; Matsumoto, 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921E2819FF14FC17FCDCFB80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FB1BFCDCFA8C.text	3C6E87D6921E2819FF14FB1BFCDCFA8C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia philippinica (Brenske 1892)	<div><p>Amphitrichia philippinica (Brenske, 1892)</p><p>Holotrichia philippinica Brenske, 1892b: 188; Dalla Torre 1912c: 205; Itoh 2003a: 13.</p><p>Amphitrichia philippinica: Matsumoto 2015: 16.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Brenske 1892a; Itoh 2003a; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921E2819FF14FB1BFCDCFA8C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14FA1FFCDCF988.text	3C6E87D6921E2819FF14FA1FFCDCF988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia quadrangulata (Brenske 1896)	<div><p>Amphitrichia quadrangulata (Brenske, 1896)</p><p>Holotrichia quadrangulata Brenske, 1896: 198; Dalla Torre 1912c: 206.</p><p>Amphitrichia quadrangulata: Matsumoto 2015: 16.</p><p>Type locality. “Insulae Philippinae.”</p><p>Distribution in the Philippines. Luzon (Brenske 1896; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921E2819FF14FA1FFCDCF988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2819FF14F9E3FCDCF8A8.text	3C6E87D6921E2819FF14F9E3FCDCF8A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia sexspecula (Chapin 1931)	<div><p>Amphitrichia sexspecula (Chapin, 1931)</p><p>Holotrichia sexspecula Chapin, 1931b: 311; Itoh 2003b: 377.</p><p>Amphitrichia sexspecula: Matsumoto 2015: 16.</p><p>Type locality. “ Cuernos Mountain, Oriental Negros.”</p><p>Distribution in the Philippines. Luzon (Mindoro), Visayas (Negros Oriental), and Mindanao (Bukidnon and Cotabato) (Chapin 1931b; Itoh 2003b; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921E2819FF14F9E3FCDCF8A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921E2818FF14F803FCDCFE98.text	3C6E87D6921E2818FF14F803FCDCFE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia negrosiana (Chapin 1931)	<div><p>Amphitrichia negrosiana (Chapin, 1931)</p><p>Holotrichia negrosiana Chapin, 1931b: 312 .</p><p>Amphitrichia negrosiana: Matsumoto 2015: 16</p><p>Type locality. “La Carlota, Occidental Negros.”</p><p>Distribution in the Philippines. Luzon (Mindoro), Visayas (Negros Oriental), and Mindanao (Chapin 1931b; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921E2818FF14F803FCDCFE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FEF3FCDCFDE4.text	3C6E87D6921F2818FF14FEF3FCDCFDE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia sorsogona (Chapin 1931)	<div><p>Amphitrichia sorsogona (Chapin, 1931)</p><p>Holotrichia sorsogona Chapin, 1931b: 312 .</p><p>Amphitrichia sorsogona: Matsumoto 2015: 16</p><p>Type locality. “ Sorsogon, Southern Luzon. ”</p><p>Distribution in the Philippines. Luzon (Sorsogon) (Chapin 1931b; Matsumoto 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921F2818FF14FEF3FCDCFDE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FDF7FCDCFC84.text	3C6E87D6921F2818FF14FDF7FCDCFC84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia stylifer (Chapin 1931)	<div><p>Amphitrichia stylifer (Chapin, 1931)</p><p>Holotrichia stylifer Chapin, 1931b: 312 .</p><p>Amphitrichia stylifer: Matsumoto 2015: 16; Matsumoto 2022: 29.</p><p>Type locality. “La Carlota, Occidental Negros.”</p><p>Distribution in the Philippines. Luzon (Palawan and Polillo), Visayas (Leyte, Negros Island, and Panay), and Mindanao (Bukidnon, Misamis Occidental, and Davao) (Chapin 1931b; Matsumoto 2015, 2022).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921F2818FF14FDF7FCDCFC84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FC17FCDCFBA4.text	3C6E87D6921F2818FF14FC17FCDCFBA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia succedanea (Matsumoto 2008)	<div><p>Amphitrichia succedanea (Matsumoto, 2008)</p><p>Holotrichia succedanea Matsumoto, 2008c: 104 .</p><p>Amphitrichia succedanea: Matsumoto 2015: 16; Matsumoto 2022: 30.</p><p>Type locality. “ Mt. Balocawihay, C. Leyte, Philippines.”</p><p>Distribution in the Philippines. Visayas (Leyte and Samar) and Mindanao (Misamis Occidental, Surigao del Norte, and Surigao del Sur) (Matsumoto 2008c, 2015, 2022).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921F2818FF14FC17FCDCFBA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FB37FCDCFAA0.text	3C6E87D6921F2818FF14FB37FCDCFAA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia tenuitibialis (Matsumoto 2010)	<div><p>Amphitrichia tenuitibialis (Matsumoto, 2010)</p><p>Holotrichia tenuitibialis Matsumoto, 2010: 50 .</p><p>Amphitrichia tenuitibialis: Matsumoto 2015: 16</p><p>Type locality. “ Mt. Halcon, Is. Mindoro, Philippines.”</p><p>Distribution in the Philippines. Luzon (Mindoro, Palawan, and Quezon) (Matsumoto 2010, 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921F2818FF14FB37FCDCFAA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14FA3BFC92F9AC.text	3C6E87D6921F2818FF14FA3BFC92F9AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia unguicularis (Moser 1912)	<div><p>Amphitrichia unguicularis (Moser, 1912)</p><p>Holotrichia unguicularis Moser, 1912: 449</p><p>Amphitrichia unguicularis: Matsumoto 2015: 16</p><p>Type locality. “ Borneo (Brunei).”</p><p>Distribution in the Philippines. Luzon (Palawan) (Moser 1912; Matsumoto 2015).</p><p>Overall distribution. The Philippines and Borneo.</p></div>	https://treatment.plazi.org/id/3C6E87D6921F2818FF14FA3BFC92F9AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921F2818FF14F93FFBC2F84C.text	3C6E87D6921F2818FF14F93FFBC2F84C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphitrichia vidua (Sharp 1876)	<div><p>Amphitrichia vidua (Sharp, 1876)</p><p>Lachnosterna vidua Sharp, 1876: 85 .</p><p>Holotrichia vidua: Brenske 1892b: 166; Brenske 1894b: 358; Dalla Torre 1912c: 205.</p><p>Amphitrichia vidua: Matsumoto 2015: 16.</p><p>Type locality. “Borneo.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Brenske 1894a; Schultze 1916a; Matsumoto 2015).</p><p>Overall distribution. The Philippines and Borneo (Sharp 1876; Brenske 1894a).</p></div>	https://treatment.plazi.org/id/3C6E87D6921F2818FF14F93FFBC2F84C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14F9E3FAB0F84D.text	3C6E87D6921C281BFF14F9E3FAB0F84D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Adoretus (Adoretus) luridus Blanchard 1851	<div><p>Adoretus (Adoretus) luridus Blanchard, 1851</p><p>Adoretus luridus Blanchard, 1851: 234; Ohaus 1910: 262; Ohaus 1912a: 267.</p><p>Adoretus philippinicus Pic, 1905: 131 . Type locality. “Luzon.”; Ohaus 1910: 262; Chapin 1931a: 306.</p><p>Adoretus (Adoretus) luridus: Ohaus 1914a: 471; Ohaus 1918: 188.</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Bacoor, Benguet, Cavite, Laguna, La Union, Manila, Romblon, and Zambales) and Visayas (Negros) (Blanchard 1850; Pic 1905; Ohaus 1910, 1912a; Chapin 1931a).</p><p>Overall distribution. The Philippines, Malaysia, and Vietnam (Ohaus 1912a; Chapin 1931a).</p></div>	https://treatment.plazi.org/id/3C6E87D6921C281BFF14F9E3FAB0F84D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14FAC3FCDCF9D1.text	3C6E87D6921C281BFF14FAC3FCDCF9D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Adoretus lopezi Chapin 1931	<div><p>Adoretus (sensu lato) lopezi Chapin, 1931</p><p>Adoretus lopezi Chapin, 1931a: 306 .</p><p>Type locality. “La Carlota, Occidental Negros.”</p><p>Distribution in the Philippines. Visayas (Negros) (Chapin 1931a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921C281BFF14FAC3FCDCF9D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14FF43FCDCFE75.text	3C6E87D6921C281BFF14FF43FCDCFE75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Latipalpus dannymohagani Matsumoto 2005	<div><p>Latipalpus dannymohagani Matsumoto, 2005</p><p>Latipalpus dannymohagani Matsumoto, 2005: 219 .</p><p>Type locality. “ Mt. Apo, Mindanao Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Panay) and Mindanao (Bukidnon, Cotabato, and Davao) (Matsumoto 2005).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921C281BFF14FF43FCDCFE75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14FDBFFCDCFD15.text	3C6E87D6921C281BFF14FDBFFCDCFD15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Octoplasia basilanensis Itoh 1996	<div><p>Octoplasia basilanensis Itoh, 1996</p><p>Octoplasia basilanensis Itoh, 1996: 776 .</p><p>Type locality. “ Basilan Is., Philippines.”</p><p>Distribution in the Philippines. Mindanao (Basilan and Bukidnon) (Itoh 1996).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921C281BFF14FDBFFCDCFD15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921C281BFF14FCDFFCDCFBC9.text	3C6E87D6921C281BFF14FCDFFCDCFBC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sophrops mindanaoensis (Moser 1924)	<div><p>Sophrops mindanaoensis (Moser, 1924)</p><p>Microtrichia mindanaoensis Moser, 1924b: 356 .</p><p>Sophrops mindanaoensis: Frey 1972b: 355; Bunalski 2022: 126.</p><p>Type locality. “ Mindanao, Surigao Province, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921C281BFF14FCDFFCDCFBC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FF43FCDCFE51.text	3C6E87D6921D281AFF14FF43FCDCFE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Adoretus (Chaetadoretus) bakeri Ohaus 1923	<div><p>Adoretus (Chaetadoretus) bakeri Ohaus, 1923</p><p>Chaetadoretus bakeri Ohaus, 1923: 590 .</p><p>Type locality. “ Basilan.”</p><p>Distribution in the Philippines. Mindanao (Basilan) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921D281AFF14FF43FCDCFE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FE2BFCDCFDB9.text	3C6E87D6921D281AFF14FE2BFCDCFDB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Adoretus (Chaetadoretus) palawanus Ohaus 1914	<div><p>Adoretus (Chaetadoretus) palawanus Ohaus, 1914</p><p>Chaetadoretus palawanus Ohaus, 1914a: 494 .</p><p>Type locality. “ Süd Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ohaus 1914a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921D281AFF14FE2BFCDCFDB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FD13FCDCFC85.text	3C6E87D6921D281AFF14FD13FCDCFC85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Adoretus (Chaetadoretus) semperi Ohaus 1912	<div><p>Adoretus (Chaetadoretus) semperi Ohaus, 1912</p><p>Adoretus semperi Ohaus, 1912a: 268 .</p><p>Type locality. “Luzon; S. Luzon, Albay und Mittel Luzon.”</p><p>Distribution in the Philippines. Luzon (Albay, Oriental Mindoro, and Romblon) (Ohaus 1912a; Calcetas et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921D281AFF14FD13FCDCFC85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FC4FFB19FB31.text	3C6E87D6921D281AFF14FC4FFB19FB31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Adoretus (Lepadoretus) compressus (Weber 1801)	<div><p>Adoretus (Lepadoretus) compressus (Weber, 1801)</p><p>Melolontha compressa Weber, 1801: 72 .</p><p>Adoretus umbrosus Burmeister, 1844: 475 . Type locality. “ Manille.”; Blanchard 1851: 233; Ohaus 1910: 262; Ohaus 1916a: 269.</p><p>Type locality. “ Sumatra.”</p><p>Distribution in the Philippines. Luzon (Ohaus, 1912b).</p><p>Overall distribution. The Philippines, South Africa, India, Sri Lanka, Myanmar, Malaysia, Thailand, Indonesia, Papua New Guinea (Blanchard 1853; Ohaus 1912b; Calcetas et al. 2021).</p></div>	https://treatment.plazi.org/id/3C6E87D6921D281AFF14FC4FFB19FB31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14FA8BFCB8F989.text	3C6E87D6921D281AFF14FA8BFCB8F989.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Adoretus (Lepadoretus) ranunculus Burmeister 1844	<div><p>Adoretus (Lepadoretus) ranunculus Burmeister, 1844</p><p>Adoretus ranunculus Burmeister, 1844: 474; Ohaus 1910: 262.</p><p>Type locality. “ Philippines.”</p><p>Distribution in the Philippines. Luzon (Manila and Rizal) and Mindanao (Davao del Sur) (Burmeister 1844; Calcetas et al. 2021).</p><p>Overall distribution. The Philippines,? Malaysia, and? China (Taiwan).</p><p>Remarks. Ohaus (1910) noted that Adoretus ranunculus was also found in Malaysia and Taiwan. These records may represent misidentifications, or the species may indeed be present in those regions. However, it is currently known only from the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921D281AFF14FA8BFCB8F989	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921D281AFF14F893FCDCF861.text	3C6E87D6921D281AFF14F893FCDCF861.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala abraensis Frey 1976	<div><p>Anomala abraensis Frey, 1976</p><p>Anomala (s. str.) abraensis Frey, 1976: 297 .</p><p>Type locality. “ Abra Pilar, Luzon, Philippines.”</p><p>Distribution in the Philippines. Luzon (Abra Pilar) (Frey 1976).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921D281AFF14F893FCDCF861	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FF0BFCDCFE51.text	3C6E87D6921A281DFF14FF0BFCDCFE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala albaya Arrow 1915	<div><p>Anomala albaya Arrow, 1915</p><p>Anomala albaya Arrow, 1915: 231;</p><p>Anomala (s.str.) albaya: Ohaus 1918: 79.</p><p>Anomala whiteheadiana Ohaus, 1915a: 108 .</p><p>Type locality. “ Luzon, Province of Albay, P. I.”</p><p>Distribution in the Philippines. Luzon (Albay) (Ohaus 1910, 1915b; Arrow 1915).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921A281DFF14FF0BFCDCFE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FE2BFCDCFD5D.text	3C6E87D6921A281DFF14FE2BFCDCFD5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala andradei Heller 1893	<div><p>Anomala andradei Heller, 1893</p><p>Anomala andradei Heller, 1893: 323; Ohaus 1910: 248.</p><p>Anomala (s. str.) andradei: Ohaus 1918: 79.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Cagayan Province) (Heller 1893).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921A281DFF14FE2BFCDCFD5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FD2FFC50FC7D.text	3C6E87D6921A281DFF14FD2FFC50FC7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala anoguttata subsp. anoguttata Burmeister 1844	<div><p>Anomala anoguttata anoguttata Burmeister, 1844</p><p>Anomala anoguttata Burmeister, 1844: 280; Blanchard 1851: 194.</p><p>Anomala (Euchlora) anoguttata: Ohaus 1915b: 341; Schultze 1916a: 173; Ohaus 1918: 102.</p><p>Type locality. “ Philippines.”</p><p>Distribution in the Philippines. Luzon (Albay, Batanes, Manila, Mindoro, and Zambales) and Visayas (Negros Occidental) (Burmeister 1844; Ohaus 1915b, 1918; Blanchard 1851; Schultze 1916a).</p><p>Overall distribution. The Philippines and Indonesia (Ohaus, 1910).</p></div>	https://treatment.plazi.org/id/3C6E87D6921A281DFF14FD2FFC50FC7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FC4FFCDCFBA5.text	3C6E87D6921A281DFF14FC4FFCDCFBA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala apogonioides Ohaus 1924	<div><p>Anomala apogonioides Ohaus, 1924</p><p>Anomala apogonioides Ohaus, 1924b: 53 .</p><p>Type locality. “Mindanao, Provinz Surigao, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921A281DFF14FC4FFCDCFBA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FB37FCDCFA19.text	3C6E87D6921A281DFF14FB37FCDCFA19.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala atrocyanea Burmeister 1844	<div><p>Anomala atrocyanea Burmeister, 1844</p><p>Anomala atrocyanea Burmeister, 1844: 277; Schultze 1916a: 173.</p><p>Anomala (Euchlora) nitidissima Blanchard, 1851: 194 . Type locality. “ Philippines.”; Ohaus 1912a: 260; Dalla Torre 1912b: 103; Ohaus 1918: 102.</p><p>Type locality. “ Philippines.”</p><p>Distribution in the Philippines. Luzon (Manila, Romblon, and Sibuyan) and Visayas (Negros Occidental) (Burmeister 1844; Blanchard 1851; Schultze 1916a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921A281DFF14FB37FCDCFA19	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14FA73FCDCF939.text	3C6E87D6921A281DFF14FA73FCDCF939.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala baeri Ohaus 1910	<div><p>Anomala baeri Ohaus, 1910</p><p>Anomala (Euchlora) baeri Ohaus, 1910: 252; Schultze 1916a: 173.</p><p>Anomala (s. str.) baeri: Ohaus 1918: 80.</p><p>Type locality. “ Luzon, Province of Albay, P. I.”</p><p>Distribution in the Philippines. Luzon (Albay, Manila, and Palawan), Visayas (Negros and Samar), and Mindanao (Marawi) (Ohaus 1910, 1918; Schultze 1916a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921A281DFF14FA73FCDCF939	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921A281DFF14F893FCDCF805.text	3C6E87D6921A281DFF14F893FCDCF805.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala bakeri Ohaus 1914	<div><p>Anomala bakeri Ohaus, 1914</p><p>Anomala Bakeri Ohaus, 1914b: 197 .</p><p>Anomala (s. str.) Bakeri: Ohaus 1918: 80.</p><p>Type locality. “ Luzon, Mt. Banahao. ”</p><p>Distribution in the Philippines. Luzon (Laguna) (Ohaus 1914b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921A281DFF14F893FCDCF805	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FF0BFCDCFE99.text	3C6E87D6921B281CFF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala butuana Ohaus 1923	<div><p>Anomala butuana Ohaus, 1923</p><p>Anomala butuana Ohaus, 1923: 583 .</p><p>Type locality. “ Mindanao, Agusan, Butuan.”</p><p>Distribution in the Philippines. Mindanao (Agusan del Norte) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921B281CFF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FEF0FCDCFDE2.text	3C6E87D6921B281CFF14FEF0FCDCFDE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala caduca Ohaus 1915	<div><p>Anomala caduca Ohaus, 1915</p><p>Anomala caduca Ohaus, 1915a: 107 .</p><p>Anomala (s. str.) caduca: Ohaus 1918: 80.</p><p>Type locality. “ Philippinen: Negros- Insel, Cuernos-Berge.”</p><p>Distribution in the Philippines. Visayas (Negros) (Ohaus 1915a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921B281CFF14FEF0FCDCFDE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FDF5FCDCFCCB.text	3C6E87D6921B281CFF14FDF5FCDCFCCB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala calpurnia Ohaus 1923	<div><p>Anomala calpurnia Ohaus, 1923</p><p>Anomala calpurnia Ohaus, 1923: 587; Zorn 2006: 50.</p><p>Type locality. “Island Polilo.”</p><p>Distribution in the Philippines. Luzon (Polillo Island) (Ohaus 1923; Zorn 2006).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921B281CFF14FDF5FCDCFCCB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FCDDFCDCFBD4.text	3C6E87D6921B281CFF14FCDDFCDCFBD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala camarinensis Ohaus 1910	<div><p>Anomala camarinensis Ohaus, 1910</p><p>Anomala camarinensis Ohaus, 1910: 246 .</p><p>Anomala (s. str.) camarinensis: Ohaus 1918: 80.</p><p>Type locality. “ Luzon, Province of Camarines, P. I.”</p><p>Distribution in the Philippines. Luzon (Camarines) (Ohaus 1910, 1918).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921B281CFF14FCDDFCDCFBD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FBA6FCDCFAD1.text	3C6E87D6921B281CFF14FBA6FCDCFAD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala cassiana Ohaus 1923	<div><p>Anomala cassiana Ohaus, 1923</p><p>Anomala cassiana Ohaus, 1923: 587; Zorn 2006: 49.</p><p>Type locality. “ Mindanao, Zamboanga, Dapitan.”</p><p>Distribution in the Philippines. Visayas (Samar and Bohol) and Mindanao (Surigao del Norte,? Maguindanao, and Zamboanga del Norte) (Ohaus 1923; Zorn 2006).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921B281CFF14FBA6FCDCFAD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14FAABFCDCF9DA.text	3C6E87D6921B281CFF14FAABFCDCF9DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala catenatopunctata Ohaus 1910	<div><p>Anomala catenatopunctata Ohaus, 1910</p><p>Anomala catenatopunctata Ohaus, 1910: 244 .</p><p>Anomala (s. str.) catenatopunctata: Ohaus 1918: 80.</p><p>Type locality. “ Luzon, Province of Bataan, Lamao.”</p><p>Distribution in the Philippines. Luzon (Albay and Bataan) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921B281CFF14FAABFCDCF9DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14F9ACFCDCF926.text	3C6E87D6921B281CFF14F9ACFCDCF926.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala ceramopyga Ohaus 1912	<div><p>Anomala ceramopyga Ohaus, 1912</p><p>Anomala (Euchlora) ceramopyga Ohaus, 1912a: 261; Ohaus 1918: 102.</p><p>Type locality. “ Luzon, Manila.”</p><p>Distribution in the Philippines. Luzon (Abra, Albay, and Manila) and Visayas (Negros Occidental) (Ohaus 1912a, 1918).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921B281CFF14F9ACFCDCF926	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6921B281CFF14F8B1FCDCF823.text	3C6E87D6921B281CFF14F8B1FCDCF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala chalcoptera Burmeister 1844	<div><p>Anomala chalcoptera Burmeister, 1844</p><p>Anomala chalcoptera Burmeister, 1844: 281; Schultze 1916a: 173.</p><p>Anomala (Euchlora) chalcoptera: Ohaus 1918: 102.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon and Mindanao (Burmeister 1844; Schultze 1916a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6921B281CFF14F8B1FCDCF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FF0BFD93FDC0.text	3C6E87D69218281FFF14FF0BFD93FDC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala chalybaea Burmeister 1844	<div><p>Anomala chalybaea Burmeister, 1844</p><p>Anomala chalybaea Burmeister, 1844: 282; Heller 1893: 21; Ohaus 1910: 250.</p><p>Anomala (s. str.) chalybaea: Ohaus 1918: 80.</p><p>Aprosterna polita Blanchard, 1851: 196 . Type locality. “ Manille.”</p><p>Anomala relucens Harold, 1869: 123; Ohaus, 1910: 250.</p><p>Type locality. “ Manilla.”</p><p>Distribution in the Philippines. Luzon (Benguet and Manila) (Burmeister 1844; Blanchard 1850; Harold 1869; Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks. Heller (1893) provided detailed descriptions of the morphological variations of the male and female Anomala chalybea .</p></div>	https://treatment.plazi.org/id/3C6E87D69218281FFF14FF0BFD93FDC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FDDBFCDCFD28.text	3C6E87D69218281FFF14FDDBFCDCFD28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala chloropyga Burmeister 1844	<div><p>Anomala chloropyga Burmeister, 1844</p><p>Anomala (Euchlora) chloropyga Burmeister, 1844: 281; Ohaus 1910: 252; Ohaus 1918: 102.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Burmeister 1844).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69218281FFF14FDDBFCDCFD28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FC83FCDCFC34.text	3C6E87D69218281FFF14FC83FCDCFC34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala cladera Ohaus 1912	<div><p>Anomala cladera Ohaus, 1912</p><p>Anomala cladera Ohaus, 1912a: 257; Schultze 1916a: 173.</p><p>Anomala (s. str.) cladera: Ohaus 1918: 82 (eladera).</p><p>Type locality. “ Mindanao, Agusan River.”</p><p>Distribution in the Philippines. Mindanao (Agusan) (Ohaus 1912a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69218281FFF14FC83FCDCFC34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FB87FCDCFB30.text	3C6E87D69218281FFF14FB87FCDCFB30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala cobala Ohaus 1916	<div><p>Anomala cobala Ohaus, 1916</p><p>Anomala cobala Ohaus, 1916: 358 .</p><p>Anomala (s. str.) cobala: Ohaus 1918: 81.</p><p>Type locality. “ Nord-Luzon, Mt. Imugan. ”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Ohaus 1916).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69218281FFF14FB87FCDCFB30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14FA8BFCDCF9AC.text	3C6E87D69218281FFF14FA8BFCDCF9AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala corruscans (Chevrolat 1841)	<div><p>Anomala corruscans (Chevrolat, 1841)</p><p>Euchlora corruscans Chevrolat, 1841: 222 .</p><p>Anomala corruscans cuprea Ohaus, 1910: 250 . Type locality. “ Palawan.”</p><p>Anomala corruscans: Burmeister 1844: 282; Blanchard 1851: 196; Redtenbacher 1868: 71; Heller 1893: 323.</p><p>Anomala (s. str.) corruscans: Ohaus 1918: 81.</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Abra, Baguio, Benguet, Palawan, and Manila) (Chevrolat 1841; Burmeister 1844; Blanchard 1851; Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69218281FFF14FA8BFCDCF9AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281FFF14F93FFCDCF894.text	3C6E87D69218281FFF14F93FFCDCF894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala cucusa Ohaus 1923	<div><p>Anomala cucusa Ohaus, 1923</p><p>Anomala cucusa Ohaus, 1923: 586 .</p><p>Type locality. “Mindanao, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69218281FFF14F93FFCDCF894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69218281EFF14F8E7FCDCFF0D.text	3C6E87D69218281EFF14F8E7FCDCFF0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala cupriventris Ohaus 1924	<div><p>Anomala cupriventris Ohaus, 1924</p><p>Euchlora cupriventris Ohaus, 1924: 103 .</p><p>Type locality. “ Northwestern Panay.”</p><p>Distribution in the Philippines. Visayas (Panay) (Ohaus 1924a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69218281EFF14F8E7FCDCFF0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FE9FFCDCFE16.text	3C6E87D69219281EFF14FE9FFCDCFE16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala dapitana Ohaus 1915	<div><p>Anomala dapitana Ohaus, 1915</p><p>Anomala dapitana Ohaus, 1915b: 339 .</p><p>Anomala (s. str.) dapitana: Ohaus 1918: 81.</p><p>Type locality. “Mindanao, Dapitan.”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Ohaus 1915b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69219281EFF14FE9FFCDCFE16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FE60FCDCFD12.text	3C6E87D69219281EFF14FE60FCDCFD12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala dasypyga Burmeister 1844	<div><p>Anomala dasypyga Burmeister, 1844</p><p>Anomala dasypyga Burmeister, 1844: 280 .</p><p>Anomala (Euchlora) dasypyga: Ohaus 1918: 103.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Burmeister 1844).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69219281EFF14FE60FCDCFD12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FD65FCDCFC1F.text	3C6E87D69219281EFF14FD65FCDCFC1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala despumata Ohaus 1910	<div><p>Anomala despumata Ohaus, 1910</p><p>Anomala despumata Ohaus, 1910: 243 .</p><p>Anomala (s. str.) despumata: Ohaus 1918: 81.</p><p>Type locality. “ Luzon, Manila and Dolores, P. I.”</p><p>Distribution in the Philippines. Luzon (Manila and Quezon Province) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69219281EFF14FD65FCDCFC1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FC6EFCDCFB18.text	3C6E87D69219281EFF14FC6EFCDCFB18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala ecbolima Ohaus 1915	<div><p>Anomala ecbolima Ohaus, 1915</p><p>Anomala ecbolima Ohaus, 1915a: 106 .</p><p>Anomala (s. str.) ecbolima: Ohaus 1918: 82.</p><p>Type locality. “Luzon: Banahao.”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao) (Ohaus 1915a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69219281EFF14FC6EFCDCFB18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FB73FCDCFA41.text	3C6E87D69219281EFF14FB73FCDCFA41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala encausta Candeze 1869	<div><p>Anomala encausta Candèze, 1869</p><p>Anomala (Euchlora) encausta Candèze, 1869: 42; Ohaus 1918: 103.</p><p>Type locality. “ Manilla.”</p><p>Distribution in the Philippines. Luzon (Manila) and Mindanao (Candèze 1869; Ohaus 1918).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69219281EFF14FB73FCDCFA41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14FA58FCDCF9B6.text	3C6E87D69219281EFF14FA58FCDCF9B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala eucardia Frey 1976	<div><p>Anomala eucardia Frey, 1976</p><p>Anomala eucardia Frey, 1976: 297 .</p><p>Type locality. “ Kabasalan, Zamboanga, Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Zamboanga Sibugay) (Frey 1976).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69219281EFF14FA58FCDCF9B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69219281EFF14F901FCDCF8B3.text	3C6E87D69219281EFF14F901FCDCF8B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala exanthematica Ohaus 1914	<div><p>Anomala exanthematica Ohaus, 1914</p><p>Anomala exanthematica Ohaus, 1914b: 187 .</p><p>Anomala (s. str.) exanthematica: Ohaus 1918: 82.</p><p>Type locality. “ Luzon, Mt. Banahao. ”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao) (Ohaus 1914b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69219281EFF14F901FCDCF8B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692192811FF14F80AFCDCFF0C.text	3C6E87D692192811FF14F80AFCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala expedita Ohaus 1914	<div><p>Anomala expedita Ohaus, 1914</p><p>Anomala (Euchlora) expedita Ohaus, 1914b: 191; Ohaus 1918: 103.</p><p>Type locality. “N. Palawan, Binaluan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ohaus 1914b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692192811FF14F80AFCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FE9FFCDCFE08.text	3C6E87D692162811FF14FE9FFCDCFE08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala flavoscutellata Ohaus 1910	<div><p>Anomala flavoscutellata Ohaus, 1910</p><p>Anomala (Heteroplia) flavoscutellata Ohaus, 1910: 237 .</p><p>Anomala (s. str.) flavoscutellata: Ohaus 1918: 82.</p><p>Type locality. “Luzon, Province of Cagayan, Cape Engaño, P. I.”</p><p>Distribution in the Philippines. Luzon (Cagayan Province) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692162811FF14FE9FFCDCFE08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FE63FCDCFD70.text	3C6E87D692162811FF14FE63FCDCFD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala fuscaoaerea Ohaus 1927	<div><p>Anomala fuscaoaerea Ohaus, 1927</p><p>Anomala fuscaoaerea Ohaus, 1927: 61 .</p><p>Type locality. “ Samar.”</p><p>Distribution in the Philippines. Visayas (Samar) (Ohaus 1927).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692162811FF14FE63FCDCFD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FD4BFCDCFC7C.text	3C6E87D692162811FF14FD4BFCDCFC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala heteroglypha Ohaus 1910	<div><p>Anomala heteroglypha Ohaus, 1910</p><p>Anomala (Aprosterna) heteroglypha Ohaus, 1910: 248 .</p><p>Anomala (s. str.) heteroglypha: Ohaus 1918: 82.</p><p>Type locality. “Luzon, P. I.”</p><p>Distribution in the Philippines. Luzon (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692162811FF14FD4BFCDCFC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FC4FFCDCFBA4.text	3C6E87D692162811FF14FC4FFCDCFBA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala hortensia (Ohaus 1927)	<div><p>Anomala hortensia (Ohaus, 1927)</p><p>Euchlora hortensia Ohaus, 1927: 61 .</p><p>Type locality. “ Mann, Samar.”</p><p>Distribution in the Philippines. Visayas (Samar) (Ohaus 1927).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692162811FF14FC4FFCDCFBA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14FB37FC5EF988.text	3C6E87D692162811FF14FB37FC5EF988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala humeralis (Burmeister 1844)	<div><p>Anomala humeralis (Burmeister, 1844)</p><p>Spilota humeralis Burmeister, 1844: 262; Blanchard 1851: 191.</p><p>Spilota eydouxi Blanchard, 1851: 192 . Type locality. “ Manille.”</p><p>Spilota pallidipennis Blanchard, 1851: 192 . Type locality. “Bonjer-Massin (Borneo); Samboangan (Mindanao).”; Ohaus, 1915a: 108 ( Anomala).</p><p>Anomala humeralis infuscata Ohaus, 1915a: 107 . Type locality. “Luzon: Los Baños.”</p><p>Anomala humeralis: Ohaus 1910: 242.</p><p>Anomala (s. str.) humeralis: Ohaus 1918: 83.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Bulacan, Manila, Mindoro, Palawan, and Tarlac), Visayas (Leyte), and Mindanao (Zamboanga) (Burmeister 1844; Blanchard 1851; Ohaus 1910, 1915a, 1915b).</p><p>Overall distribution. The Philippines and Borneo (Blanchard 1851).</p></div>	https://treatment.plazi.org/id/3C6E87D692162811FF14FB37FC5EF988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162811FF14F9E3FCDCF8F0.text	3C6E87D692162811FF14F9E3FCDCF8F0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala inconsueta Ohaus 1910	<div><p>Anomala inconsueta Ohaus, 1910</p><p>Anomala (Euchlora) inconsueta Ohaus, 1910: 250; Ohaus 1918: 103.</p><p>Type locality. “Negros, P. I.”</p><p>Distribution in the Philippines. Luzon (Benguet) and Visayas (Negros) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692162811FF14F9E3FCDCF8F0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692162810FF14F8CBFCDCFF0C.text	3C6E87D692162810FF14F8CBFCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala infans Ohaus 1910	<div><p>Anomala infans Ohaus, 1910</p><p>Anomala (Rhinoplia) infans Ohaus, 1910: 236 .</p><p>Anomala (Aprosterna) infans: Ohaus 1918: 63.</p><p>Type locality. “Luzon, P. I.”</p><p>Distribution in the Philippines. Luzon (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692162810FF14F8CBFCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FE9FFCDCFE08.text	3C6E87D692172810FF14FE9FFCDCFE08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala inopinata Ohaus 1914	<div><p>Anomala inopinata Ohaus, 1914</p><p>Anomala inopinata Ohaus, 1914b: 188 .</p><p>Anomala (Euchlora) inopinata: Ohaus 1918: 103.</p><p>Type locality. “ Luzon, Mt. Makiling. ”</p><p>Distribution in the Philippines. Luzon (Laguna) (Ohaus 1914b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692172810FF14FE9FFCDCFE08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FE63FCDCFD14.text	3C6E87D692172810FF14FE63FCDCFD14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala laetabilis Ohaus 1915	<div><p>Anomala laetabilis Ohaus, 1915</p><p>Anomala laetabilis Ohaus, 1915a: 105 .</p><p>Anomala (s. str.) laetabilis: Ohaus 1918: 83.</p><p>Type locality. “Ins. Philippin., S. Luzon: Banahao.”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao) (Ohaus 1915a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692172810FF14FE63FCDCFD14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FD67FCDCFBC8.text	3C6E87D692172810FF14FD67FCDCFBC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala leotaudi Blanchard 1851	<div><p>Anomala leotaudi Blanchard, 1851</p><p>Anomala leotaudi Blanchard, 1851: 191 .</p><p>Anomala leotaudii fuscoviridis Ohaus, 1910: 242 . Type locality. “Luzon, Province of Cagayan, Cape Engaño.”</p><p>Anomala (s. str.) leotaudi: Ohaus 1918: 83.</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Cagayan Province and Manila) (Blanchard 1850; Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692172810FF14FD67FCDCFBC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FBA3FBD8FAA0.text	3C6E87D692172810FF14FBA3FBD8FAA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala lucidula (Guerin-Meneville 1830)	<div><p>Anomala lucidula (Guérin-Méneville, 1830)</p><p>Euchlora lucidula Guérin-Méneville, 1830: 88 .</p><p>Anomala exarata Burmeister, 1844: 260 . Type locality. “ Philippines.”; Blanchard 1851: 192; Ohaus 1912a: 256; Ohaus 1918: 82.</p><p>Anomala lucidula: Ohaus 1916a: 108; Zorn 2011: 354.</p><p>Type locality. “Bourou.”</p><p>Distribution in the Philippines. Luzon (Manila) and Visayas (Negros) (Burmeister 1844; Blanchard 1851; Ohaus 1910, 1912a, 1916a; Schultze 1916a).</p><p>Overall distribution. The Philippines and Indonesia (Ohaus 1916a; Zorn 2011).</p></div>	https://treatment.plazi.org/id/3C6E87D692172810FF14FBA3FBD8FAA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14FA3BFCDCF988.text	3C6E87D692172810FF14FA3BFCDCF988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala macrophalla Ohaus 1923	<div><p>Anomala macrophalla Ohaus, 1923</p><p>Anomala macrophalla Ohaus, 1923: 584 .</p><p>Type locality. “ Mindanao, Zamboanga, Dapitan.”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692172810FF14FA3BFCDCF988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172810FF14F9E3FCDCF894.text	3C6E87D692172810FF14F9E3FCDCF894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala macrophthalma Ohaus 1910	<div><p>Anomala macrophthalma Ohaus, 1910</p><p>Anomala (Heteroplia) macrophthalma Ohaus, 1910: 238 .</p><p>Anomala (s. str.) macrophthalma: Ohaus 1918: 84.</p><p>Type locality. “ Luzon, Province of Cagayan, Aparri, P. I.”</p><p>Distribution in the Philippines. Luzon (Cagayan Province) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692172810FF14F9E3FCDCF894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692172813FF14F8E7FCDCFF0C.text	3C6E87D692172813FF14F8E7FCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala maculifemorata Ohaus 1912	<div><p>Anomala maculifemorata Ohaus, 1912</p><p>Anomala (Euchlora) maculifemorata Ohaus, 1912a: 258; Ohaus 1918: 103.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Ohaus 1912a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692172813FF14F8E7FCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FE9FFCDCFE74.text	3C6E87D692142813FF14FE9FFCDCFE74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala melitta Ohaus 1923	<div><p>Anomala melitta Ohaus, 1923</p><p>Anomala melitta Ohaus, 1923: 582 .</p><p>Type locality. “Mindanao, Surigao, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692142813FF14FE9FFCDCFE74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FE47FCDCFD5C.text	3C6E87D692142813FF14FE47FCDCFD5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala micholitzi Ohaus 1913	<div><p>Anomala micholitzi Ohaus, 1913</p><p>Anomala (Euchlora) micholitzi Ohaus, 1913: 69 .</p><p>Type locality. “Philippin. Inseln, Mindanao, Davao.”</p><p>Distribution in the Philippines. Mindanao (Davao) (Ohaus 1913).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692142813FF14FE47FCDCFD5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FD2FFCDCFC58.text	3C6E87D692142813FF14FD2FFCDCFC58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala misandria Ohaus 1916	<div><p>Anomala misandria Ohaus, 1916</p><p>Anomala misandria Ohaus, 1916: 258 .</p><p>Anomala (s. str.) misandria: Ohaus 1918: 84.</p><p>Type locality. “ Süd-Luzon, Mt. Isarog. ”</p><p>Distribution in the Philippines. Luzon (Camarines Sur) (Ohaus 1916).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692142813FF14FD2FFCDCFC58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FC33FCDCFB80.text	3C6E87D692142813FF14FC33FCDCFB80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala muricata Ohaus 1930	<div><p>Anomala muricata Ohaus, 1930</p><p>Anomala muricata Ohaus, 1930a: 556 .</p><p>Type locality. “ Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ohaus 1930a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692142813FF14FC33FCDCFB80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FB1BFCDCFAE8.text	3C6E87D692142813FF14FB1BFCDCFAE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala nerissa (Ohaus 1930)	<div><p>Anomala nerissa (Ohaus, 1930)</p><p>Euchlora nerissa Ohaus, 1930a: 558 .</p><p>Type locality. “ Mindanao, Zamboanga.”</p><p>Distribution in the Philippines. Mindanao (Zamboanga) (Ohaus 1930a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692142813FF14FB1BFCDCFAE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14FAC3FCDCF9F4.text	3C6E87D692142813FF14FAC3FCDCF9F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala noctivaga Ohaus 1910	<div><p>Anomala noctivaga Ohaus, 1910</p><p>Anomala noctivaga Ohaus, 1910: 246 .</p><p>Anomala (s. str.) noctivaga: Ohaus 1918: 84.</p><p>Type locality. “ Batanes Islands, P. I.”</p><p>Distribution in the Philippines. Luzon (Batanes) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692142813FF14FAC3FCDCF9F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14F9C7FCDCF8DC.text	3C6E87D692142813FF14F9C7FCDCF8DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala nocturna Ohaus 1930	<div><p>Anomala nocturna Ohaus, 1930</p><p>Anomala nocturna Ohaus, 1930a: 557 .</p><p>Type locality. “ Luzon, Baguio.”</p><p>Distribution in the Philippines. Luzon (Baguio) (Ohaus 1930a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692142813FF14F9C7FCDCF8DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692142813FF14F8AFFCDCF804.text	3C6E87D692142813FF14F8AFFCDCF804.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala obesa Candeze 1869	<div><p>Anomala obesa Candèze, 1869</p><p>Anomala (Euchlora) obesa Candèze, 1869: 41; Ohaus 1918: 104.</p><p>Type locality. “Ins. Philippinenses.”</p><p>Distribution in the Philippines. Luzon (Manila) (Candèze 1869).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692142813FF14F8AFFCDCF804	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FF0BFCDCFE50.text	3C6E87D692152812FF14FF0BFCDCFE50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala ovatula Ohaus 1910	<div><p>Anomala ovatula Ohaus, 1910</p><p>Anomala ovatula Ohaus, 1910: 240 .</p><p>Anomala (s. str.) ovatula: Ohaus 1918: 84.</p><p>Type locality. “Mindanao, Camp Keithley P. I.”</p><p>Distribution in the Philippines. Luzon (Palawan), Visayas (Leyte), and Mindanao (Lanao del Sur) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692152812FF14FF0BFCDCFE50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FE2BFCDCFD5C.text	3C6E87D692152812FF14FE2BFCDCFD5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala palawana Ohaus 1910	<div><p>Anomala palawana Ohaus, 1910</p><p>Anomala palawana Ohaus, 1910: 241 .</p><p>Anomala (s. str.) palawana: Ohaus 1918: 84.</p><p>Type locality. “ Palawan, P. I.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692152812FF14FE2BFCDCFD5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FD2FFCDCFC7C.text	3C6E87D692152812FF14FD2FFCDCFC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala planata Candeze 1869	<div><p>Anomala planata Candèze, 1869</p><p>Anomala planata Candèze, 1869: 42; Ohaus 1910: 248.</p><p>Anomala (s. str.) planata: Ohaus 1918: 84.</p><p>Type locality. “ Manilla.”</p><p>Distribution in the Philippines. Luzon (Benguet and Cagayan Province), Visayas (Leyte), and Mindanao (Lanao del Sur) (Candèze 1869; Ohaus 1910, 1918).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692152812FF14FD2FFCDCFC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FC4FFCDCFBA4.text	3C6E87D692152812FF14FC4FFCDCFBA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala polina Ohaus 1923	<div><p>Anomala polina Ohaus, 1923</p><p>Anomala polina Ohaus, 1923: 585 .</p><p>Type locality. “ Mindanao, Ifugao, Mount Polis. ”</p><p>Distribution in the Philippines. Luzon (Ifugao) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692152812FF14FC4FFCDCFBA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FB37FCDCFAA0.text	3C6E87D692152812FF14FB37FCDCFAA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala praematura Ohaus 1910	<div><p>Anomala praematura Ohaus, 1910</p><p>Anomala (Euchlora) praematura Ohaus, 1910: 251 .</p><p>Anomala (s. str.) praematura: Ohaus 1918: 84.</p><p>Type locality. “Luzon, P. I.”</p><p>Distribution in the Philippines. Luzon (Albay) and Visayas (Negros Occidental) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692152812FF14FB37FCDCFAA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14FA3BFCDCF988.text	3C6E87D692152812FF14FA3BFCDCF988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala proctolasia Ohaus 1910	<div><p>Anomala proctolasia Ohaus, 1910</p><p>Anomala proctolasia Ohaus, 1910: 239; Ohaus 1918: 84.</p><p>Type locality. “ Luzon, Manila; Polillo.”</p><p>Distribution in the Philippines. Luzon (Manila and Polillo Island) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692152812FF14FA3BFCDCF988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152812FF14F9E3FCDCF8F0.text	3C6E87D692152812FF14F9E3FCDCF8F0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala quadricalcarata Ohaus 1924	<div><p>Anomala quadricalcarata Ohaus, 1924</p><p>Anomala quadricalcarata Ohaus, 1924b: 54 .</p><p>Type locality. “Sibuyan.”</p><p>Distribution in the Philippines. Luzon (Sibuyan Island) (Ohaus 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692152812FF14F9E3FCDCF8F0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692152815FF14F8CBFCDCFF20.text	3C6E87D692152815FF14F8CBFCDCFF20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala samarensis Ohaus 1923	<div><p>Anomala samarensis Ohaus, 1923</p><p>Anomala samarensis Ohaus, 1923: 585 .</p><p>Type locality. “ Samar.”</p><p>Distribution in the Philippines. Visayas (Samar) and Mindanao (Agusan del Norte and Surigao) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692152815FF14F8CBFCDCFF20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FEBBFCDCFE2C.text	3C6E87D692122815FF14FEBBFCDCFE2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala sanchezi Ohaus 1912	<div><p>Anomala sanchezi Ohaus, 1912</p><p>Anomala (Heteroplia) sanchezi Ohaus, 1912a: 255 .</p><p>Anomala (Aprosterna) sanchezi: Ohaus 1918: 64.</p><p>Type locality. “Luzon, Benguet, Baguio.”</p><p>Distribution in the Philippines. Luzon (Baguio) (Ohaus 1912a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692122815FF14FEBBFCDCFE2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FDBFFCDCFD14.text	3C6E87D692122815FF14FDBFFCDCFD14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala schultzeana Ohaus 1910	<div><p>Anomala schultzeana Ohaus, 1910</p><p>Anomala schultzeana Ohaus, 1910: 247; Ohaus 1918: 86.</p><p>Type locality. “ Luzon, Manila, P. I.”</p><p>Distribution in the Philippines. Luzon (Manila) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692122815FF14FDBFFCDCFD14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FD67FCDCFC10.text	3C6E87D692122815FF14FD67FCDCFC10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala semperiana Ohaus 1910	<div><p>Anomala semperiana Ohaus, 1910</p><p>Anomala semperiana Ohaus, 1910: 241 .</p><p>Anomala (s. str.) semperiana: Ohaus 1918: 86.</p><p>Type locality. “Luzon, P. I.”</p><p>Distribution in the Philippines. Luzon (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692122815FF14FD67FCDCFC10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FC6BFCDCFB78.text	3C6E87D692122815FF14FC6BFCDCFB78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala seticrus Ohaus 1912	<div><p>Anomala seticrus Ohaus, 1912</p><p>Anomala (Euchlora) seticrus Ohaus, 1912a: 259; Ohaus 1918: 104.</p><p>Type locality. “ Palawan, Iwahig.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ohaus 1912a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692122815FF14FC6BFCDCFB78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FB53FCDCFAA0.text	3C6E87D692122815FF14FB53FCDCFAA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala sibuyana Ohaus 1923	<div><p>Anomala sibuyana Ohaus, 1923</p><p>Anomala sibuyana Ohaus, 1923: 582 .</p><p>Type locality. “Sibuyan.”</p><p>Distribution in the Philippines. Luzon (Sibuyan Island) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692122815FF14FB53FCDCFAA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122815FF14FA3BFCDCF938.text	3C6E87D692122815FF14FA3BFCDCF938.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala smaragdina (Eschscholtz 1822)	<div><p>Anomala smaragdina (Eschscholtz, 1822)</p><p>Anomala smaragdina Eschscholtz, 1822: 18; Burmeister 1844: 276; Blanchard 1851: 194; Ohaus 1912a: 263.</p><p>Euchlora smaragdula Laporte, 1840: 153 .</p><p>Anomala (Euchlora) smaragdina: Ohaus 1918: 104.</p><p>Type locality. “ Insel Luçon (Manille).”</p><p>Distribution in the Philippines. Luzon (Catanduanes and Manila) (Eschscholtz 1822; Laporte 1840; Ohaus 1912a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692122815FF14FA3BFCDCF938	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692122814FF14F893FED0FF0C.text	3C6E87D692122814FF14F893FED0FF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala sulcatula Burmeister 1844	<div><p>Anomala sulcatula Burmeister, 1844</p><p>Anomala sulcatula Burmeister, 1844: 261; Blanchard 1851: 192; Cartwright &amp; Gordon 1971: 284.</p><p>Anomala (s. str.) sulcatula: Ohaus 1918: 86.</p><p>Type locality. “ Manille.”</p><p>Distribution in the Philippines. Luzon (Albay, Bataan, Manila, Palawan, Polillo, Rizal, and Tarlac) and Visayas (Negros Island) (Burmeister 1844; Blanchard 1851; Ohaus 1918).</p><p>Overall distribution. The Philippines, Borneo, Malaysia, Guam, and Northern Mariana Islands (Dunlap et al. 2016).</p></div>	https://treatment.plazi.org/id/3C6E87D692122814FF14F893FED0FF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FE9FFCDCFE74.text	3C6E87D692132814FF14FE9FFCDCFE74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala surigana Ohaus 1923	<div><p>Anomala surigana Ohaus, 1923</p><p>Anomala surigana Ohaus, 1923: 583 .</p><p>Type locality. “Mindanao, Surigao, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692132814FF14FE9FFCDCFE74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FE47FCDCFD5C.text	3C6E87D692132814FF14FE47FCDCFD5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala trichopyga Ohaus 1915	<div><p>Anomala trichopyga Ohaus, 1915</p><p>Anomala (Euchlora) trichopyga Ohaus, 1915b: 340; Ohaus 1918: 104.</p><p>Type locality. “Ins. Philippin.: Montalban.”</p><p>Distribution in the Philippines. Luzon (Rizal) (Ohaus 1915b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692132814FF14FE47FCDCFD5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FD2FFCDCFC84.text	3C6E87D692132814FF14FD2FFCDCFC84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala trigonopyga Ohaus 1912	<div><p>Anomala trigonopyga Ohaus, 1912</p><p>Anomala (Euchlora) trigonopyga Ohaus, 1912a: 263; Ohaus 1918: 104;</p><p>Type locality. “Luzon, Zambales; S. Luzon, Albay und Mindoro.”</p><p>Distribution in the Philippines. Luzon (Albay, Mindoro, and Zambales) (Ohaus 1912a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692132814FF14FD2FFCDCFC84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FC17FCDCFB80.text	3C6E87D692132814FF14FC17FCDCFB80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala vietipennis Ohaus 1910	<div><p>Anomala vietipennis Ohaus, 1910</p><p>Anomala vietipennis Ohaus, 1910: 245 .</p><p>Anomala (s. str.) vietipennis: Ohaus 1918: 87.</p><p>Type locality. “ Luzon, Manila, P. I.”</p><p>Distribution in the Philippines. Luzon (Manila) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692132814FF14FC17FCDCFB80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FB1BFCDCFAE8.text	3C6E87D692132814FF14FB1BFCDCFAE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala vigilax Ohaus 1924	<div><p>Anomala vigilax Ohaus, 1924</p><p>Anomala vigilax Ohaus, 1924b: 53</p><p>Type locality. “ Luzon, Provinz Benguet, Trinidad.”</p><p>Distribution in the Philippines. Luzon (Benguet) (Ohaus 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692132814FF14FB1BFCDCFAE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14FAC3FCDCF9D0.text	3C6E87D692132814FF14FAC3FCDCF9D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala volanda Ohaus 1923	<div><p>Anomala volanda Ohaus, 1923</p><p>Anomala volanda Ohaus, 1923: 584 .</p><p>Type locality. “Mindanao, Surigao, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692132814FF14FAC3FCDCF9D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132814FF14F9ABFCDCF8DC.text	3C6E87D692132814FF14F9ABFCDCF8DC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anomala xestopyga Ohaus 1915	<div><p>Anomala xestopyga Ohaus, 1915</p><p>Anomala xestopyga Ohaus, 1915b: 340 .</p><p>Anomala (s. str.) xestopyga: Ohaus 1918: 87.</p><p>Type locality. “ Philippinen: Benguet, Baguio.”</p><p>Distribution in the Philippines. Luzon (Baguio) (Ohaus 1915b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692132814FF14F9ABFCDCF8DC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692132817FF14F8E7FCDCFF0C.text	3C6E87D692132817FF14F8E7FCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callistethus boettcheri (Ohaus 1914)	<div><p>Callistethus boettcheri (Ohaus, 1914)</p><p>Anomala (Spilota) boettcheri Ohaus, 1914b: 189; Ohaus 1918: 106.</p><p>Type locality. “ Nord Palawan: Bacuit.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ohaus 1914b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692132817FF14F8E7FCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FE47FC42FD14.text	3C6E87D692102817FF14FE47FC42FD14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callistethus mindanaoensis Kobayashi & Fujioka 2021	<div><p>Callistethus mindanaoensis Kobayashi &amp; Fujioka, 2021 [Junior homonymy]</p><p>Callistethus mindanaoensis Kobayashi &amp; Fujioka, 2021: 117 .</p><p>Type locality. “Mt. Apo, North Cotabato, Mindanao Is., Philippines.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks. This species is a junior homonym of Callistethus mindanaoensis Wada, 2002, and in accordance with Article 52 of the ICZN, a replacement name must be proposed.</p></div>	https://treatment.plazi.org/id/3C6E87D692102817FF14FE47FC42FD14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FE9FFCDCFE74.text	3C6E87D692102817FF14FE9FFCDCFE74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callistethus mindanaoensis Wada 2002	<div><p>Callistethus mindanaoensis Wada, 2002</p><p>Callistethus mindanaoensis Wada, 2002: 178 .</p><p>Type locality. “ Mt. Apo, Mindanao Is. ”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Wada 2002).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692102817FF14FE9FFCDCFE74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FD67FCDCFC7C.text	3C6E87D692102817FF14FD67FCDCFC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callistethus moseri (Ohaus 1924)	<div><p>Callistethus moseri (Ohaus, 1924)</p><p>Anomala (Spilota) moseri Ohaus, 1924b: 54 .</p><p>Type locality. “Mindanao, Provinz Surigao, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692102817FF14FD67FCDCFC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FC4FFCDCFBA4.text	3C6E87D692102817FF14FC4FFCDCFBA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callistethus palawanensis Wada 1998	<div><p>Callistethus palawanensis Wada, 1998</p><p>Callistethus palawanensis Wada, 1998: 146 .</p><p>Type locality. “ Bindoyan, Palawan Is., Philippines.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Wada 1998a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692102817FF14FC4FFCDCFBA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FB37FC53FAA0.text	3C6E87D692102817FF14FB37FC53FAA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Callistethus picturatus (Candeze 1869)	<div><p>Callistethus picturatus (Candèze, 1869)</p><p>Anomala (Spilota) picturata Candèze, 1869: 42; Ohaus 1915a: 112; Ohaus 1918: 107.</p><p>Type locality. “ Manilla.”</p><p>Distribution in the Philippines. Luzon (Catanduanes, Laguna, Mindoro, and Palawan) and Visayas (Negros) (Candèze 1869; Ohaus 1915a).</p><p>Overall distribution. The Philippines and Malaysia (Ohaus 1915a).</p></div>	https://treatment.plazi.org/id/3C6E87D692102817FF14FB37FC53FAA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14FA73FE0EF938.text	3C6E87D692102817FF14FA73FE0EF938.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mimela blumei Hope 1835	<div><p>Mimela blumei Hope, 1835</p><p>Mimela blumei Hope, 1835: 112; Ohaus 1908: 642, Ohaus 1910: 260; Schultze 1916a: 174.</p><p>Type locality. “ Java.”</p><p>Distribution in the Philippines. Luzon (Manila) (Ohaus 1908, 1910; Schultze 1916a).</p><p>Overall distribution. The Philippines and Indonesia (Hope 1835; Schultze 1916a).</p><p>Remarks. This species is common in Java, but its presence in the Philippines, as noted by Ohaus (1910), remains doubtful.</p></div>	https://treatment.plazi.org/id/3C6E87D692102817FF14FA73FE0EF938	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692102817FF14F893FBE4F860.text	3C6E87D692102817FF14F893FBE4F860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mimela maculicollis Ohaus 1908	<div><p>Mimela maculicollis Ohaus, 1908</p><p>Mimela maculicollis Ohaus, 1908: 636 .</p><p>Type locality. “Borneo: Kinibalu im Nordosten und Tandjong- Bandjermasin im S-O; Süd Luzon: Albay.”</p><p>Distribution in the Philippines. Luzon (Albay) (Ohaus, 1908).</p><p>Overall distribution. The Philippines, Indonesia, and Malaysia (Ohaus 1908).</p></div>	https://treatment.plazi.org/id/3C6E87D692102817FF14F893FBE4F860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FE63FCDCFD71.text	3C6E87D692112816FF14FE63FCDCFD71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) alticola (Ohaus 1923)	<div><p>Malaia (Falsopopillia) alticola (Ohaus, 1923)</p><p>Popillia alticola Ohaus, 1923: 589 .</p><p>Type locality. “ Luzon, Nueva Vizcaya, Imugan.”</p><p>Distribution in the Philippines. Luzon (Baguio and Nueva Vizcaya) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692112816FF14FE63FCDCFD71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FD4BFCDCFC59.text	3C6E87D692112816FF14FD4BFCDCFC59.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) boettcheri (Ohaus 1916)	<div><p>Malaia (Falsopopillia) boettcheri (Ohaus, 1916)</p><p>Popillia boettcheri Ohaus, 1916: 356; Ohaus 1918: 13</p><p>Type locality. “ Nord-Luzon, Mt. Imugan. ”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Ohaus 1916).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692112816FF14FD4BFCDCFC59	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FC33FCDCFBA5.text	3C6E87D692112816FF14FC33FCDCFBA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) cetrata (Newman 1841)	<div><p>Malaia (Falsopopillia) cetrata (Newman, 1841)</p><p>Popillia cetrata Newman, 1841: 223; Burmeister 1844: 546; Kraatz 1892: 284; Ohaus 1910: 255; Ohaus 1918: 136.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Benguet) and Visayas (Negros Occidental) (Newman 1841).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692112816FF14FC33FCDCFBA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FB37FCDCFA8D.text	3C6E87D692112816FF14FB37FCDCFA8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) conopyga (Ohaus 1905)	<div><p>Malaia (Falsopopillia) conopyga (Ohaus, 1905)</p><p>Popillia conopyga Ohaus, 1905: 92; Ohaus 1910: 260.</p><p>Type locality. “Luzon, bei der Lagune südlich von Manila gesammelt.”</p><p>Distribution in the Philippines. Luzon (Manila and Laguna) (Ohaus 1905, 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692112816FF14FB37FCDCFA8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FA1FFCDCF9F5.text	3C6E87D692112816FF14FA1FFCDCF9F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) depressa (Kraatz 1892)	<div><p>Malaia (Falsopopillia) depressa (Kraatz, 1892)</p><p>Popillia depressa Kraatz, 1892: 287; Ohaus 1910: 261; Ohaus 1918: 137.</p><p>Type locality. “Insulae Philippinae.”</p><p>Distribution in the Philippines. Luzon (Baguio) (Kraatz 1892; Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692112816FF14FA1FFCDCF9F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14F9C7FCDCF8DD.text	3C6E87D692112816FF14F9C7FCDCF8DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) depressiuscula (Kraatz 1892)	<div><p>Malaia (Falsopopillia) depressiuscula (Kraatz, 1892)</p><p>Popillia depressiuscula Kraatz, 1892: 286; Ohaus 1910: 261; Ohaus 1918: 137.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Manila and Nueva Vizcaya) (Kraatz 1892; Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692112816FF14F9C7FCDCF8DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14F8AFFCDCF805.text	3C6E87D692112816FF14F8AFFCDCF805.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) furcula (Ohaus 1930)	<div><p>Malaia (Falsopopillia) furcula (Ohaus, 1930)</p><p>Popillia furcula Ohaus, 1930a: 555 .</p><p>Type locality. “Süd Luzon.”</p><p>Distribution in the Philippines. Luzon (Ohaus 1930).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692112816FF14F8AFFCDCF805	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692112816FF14FF0BFCDCFE99.text	3C6E87D692112816FF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mimela palawana Ohaus 1910	<div><p>Mimela palawana Ohaus, 1910</p><p>Mimela palawana Ohaus, 1910: 252; Ohaus 1918: 131; Ohaus 1943: 74 (palauana).</p><p>Type locality. “ Palawan, Bacuit, Mt. Capoas, P. I.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692112816FF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FF0BFCDCFE99.text	3C6E87D6922E2829FF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) macronyx (Ohaus 1916)	<div><p>Malaia (Falsopopillia) macronyx (Ohaus, 1916)</p><p>Popillia macronyx Ohaus, 1916: 355; Ohaus 1918: 138 (macrony).</p><p>Type locality. “ Nord-Luzon, Mt. Imugan. ”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Ohaus 1916).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922E2829FF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FEF0FCDCFDCE.text	3C6E87D6922E2829FF14FEF0FCDCFDCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) nona (Ohaus 1923)	<div><p>Malaia (Falsopopillia) nona (Ohaus, 1923)</p><p>Popillia nona Ohaus, 1923: 589 .</p><p>Type locality. “Mindanao, Surigao, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922E2829FF14FEF0FCDCFDCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FDD9FCDCFD37.text	3C6E87D6922E2829FF14FDD9FCDCFD37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) oculata (Ohaus 1910)	<div><p>Malaia (Falsopopillia) oculata (Ohaus, 1910)</p><p>Popillia oculata Ohaus, 1910: 255; Ohaus 1918: 139.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Ohaus,1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922E2829FF14FDD9FCDCFD37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FC81FCDCFC1F.text	3C6E87D6922E2829FF14FC81FCDCFC1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) pulla (Ohaus 1916)	<div><p>Malaia (Falsopopillia) pulla (Ohaus, 1916)</p><p>Popillia pulla Ohaus, 1916: 357; Ohaus 1918: 140.</p><p>Type locality. “ Süd-Luzon, Mt. Isarog. ”</p><p>Distribution in the Philippines. Luzon (Camarines Sur) (Ohaus 1916).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922E2829FF14FC81FCDCFC1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FC6EFCDCFB44.text	3C6E87D6922E2829FF14FC6EFCDCFB44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) rugithorax (Ohaus 1916)	<div><p>Malaia (Falsopopillia) rugithorax (Ohaus, 1916)</p><p>Popillia rugithorax Ohaus, 1916: 354; Ohaus 1918: 140.</p><p>Type locality. “ Nord-Luzon, Mt. Imugan. ”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Ohaus 1916).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922E2829FF14FC6EFCDCFB44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14FB57FCDCF9B6.text	3C6E87D6922E2829FF14FB57FCDCF9B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Falsopopillia) scalpta (Newman 1841)	<div><p>Malaia (Falsopopillia) scalpta (Newman, 1841)</p><p>Popilia scalpta Newman, 1841: 222; Burmeister 1844: 546; Kraatz 1892: 286; Ohaus 1910: 254; Ohaus 1918: 140.</p><p>Popillia aemula Newman, 1841: 222 . Type locality. “ Philippine Islands.”; Burmeister 1844: 546; Kraatz 1892: 285.</p><p>Popillia picticollis Kraatz, 1892: 284 . Type locality. “Luzon.”</p><p>Popillia scalpta caeca Ohaus, 1914b: 192 . Type locality. “Luzon, Mt. Banahao.”</p><p>Popillia scalpta microps Ohaus, 1914b: 192 . Type locality. “Luzon, Mt. Makiling.”</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Laguna) (Newman 1841; Kraatz 1892; Arrow 1899; Ohaus 1910, 1914b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922E2829FF14FB57FCDCF9B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2829FF14F979FCDCF857.text	3C6E87D6922E2829FF14F979FCDCF857.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Malaia) stigma Ohaus 1923	<div><p>Malaia (Malaia) stigma Ohaus, 1923</p><p>Malaia stigma Ohaus, 1923: 588 .</p><p>Type locality. “Mindanao, Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Ohaus 1923).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922E2829FF14F979FCDCF857	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922E2828FF14F826FCDCFF21.text	3C6E87D6922E2828FF14F826FCDCFF21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Malaia) thoracica Ohaus 1910	<div><p>Malaia (Malaia) thoracica Ohaus, 1910</p><p>Malaia thoracica Ohaus, 1910: 253; Ohaus 1918: 151.</p><p>Type locality. “Luzon, P. I.”</p><p>Distribution in the Philippines. Luzon (Baguio) (Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922E2828FF14F826FCDCFF21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14FEF3FCDCFDC1.text	3C6E87D6922F2828FF14FEF3FCDCFDC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Pseudomalaia) flavopilosa (Ohaus 1905)	<div><p>Malaia (Pseudomalaia) flavopilosa (Ohaus, 1905)</p><p>Pseudomalaia flavopilosa Ohaus, 1905: 91; Ohaus 1918: 151.</p><p>Type locality. “Ins. Philippin., Negros.”</p><p>Distribution in the Philippines. Visayas (Negros) (Ohaus 1905).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922F2828FF14FEF3FCDCFDC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14FDDBFCDCFCE1.text	3C6E87D6922F2828FF14FDDBFCDCFCE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Pseudomalaia) pilifera (Burmeister 1844)	<div><p>Malaia (Pseudomalaia) pilifera (Burmeister, 1844)</p><p>Popillia pilifera Burmeister, 1844: 309; Ohaus 1918: 151.</p><p>Pseudomalaia pilifera: Ohaus 1910: 260; Ohaus 1912a: 264; Schultze 1916a: 174.</p><p>Type locality. “Luzon.”</p><p>Distribution in the Philippines. Luzon (Albay, Cagayan Province, and Benguet) and Mindanao (Burmeister 1844; Ohaus 1910, 1912a; Schultze 1916a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922F2828FF14FDDBFCDCFCE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14FCFBFCDCFAD5.text	3C6E87D6922F2828FF14FCFBFCDCFAD5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Pseudomalaia) semperi Kraatz 1892	<div><p>Malaia (Pseudomalaia) semperi Kraatz, 1892</p><p>Malaia semperi Kraatz, 1892: 178 .</p><p>Pseudomalaia semperi: Ohaus 1914b: 193; Ohaus 1918: 152.</p><p>Malaia semperi marginipennis Kraatz, 1892: 179 . Type locality. “Ins. Philippinae.”</p><p>Pseudomalaia semperi marginipennis: Schultze 1916a: 174.</p><p>Malaia semperi pallidipennis Kraatz, 1892: 179 . Type locality. “Ins. Philippinae.”</p><p>Pseudomalaia semperi pallidipennis: Schultze 1916a: 174.</p><p>Pseudomalaia semperi nigripennis Ohaus, 1914b: 192 . Type locality. “ Luzon, Mt. Banahao. ”</p><p>Type locality. “Ins. Philippinae.”</p><p>Distribution in the Philippines. Luzon (Laguna) and Visayas (Negros, Samar, and Leyte) (Kraatz 1892; Ohaus 1914b; Schultze 1916a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922F2828FF14FCFBFCDCFAD5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14FAA7FCDCF9F5.text	3C6E87D6922F2828FF14FAA7FCDCF9F5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Pseudomalaia) tagala (Heller 1891)	<div><p>Malaia (Pseudomalaia) tagala (Heller, 1891)</p><p>Popilia tagala Heller, 1891: 305 .</p><p>Pseudomalaia tagala: Ohaus, 1910: 260; Ohaus 1912a: 265; Schultze 1916a: 174; Ohaus 1918: 152.</p><p>Type locality. “Habitat in insulis Philippinis (Luzon et Mindanao).”</p><p>Distribution in the Philippines. Luzon (Benguet) and Mindanao (Agusan) (Heller 1891; Ohaus 1910, 1912a, 1918; Schultze 1916).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922F2828FF14FAA7FCDCF9F5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F2828FF14F9C7FCDCF8DD.text	3C6E87D6922F2828FF14F9C7FCDCF8DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Malaia (Pseudomalaia) whiteheadi Ohaus 1912	<div><p>Malaia (Pseudomalaia) whiteheadi Ohaus, 1912</p><p>Pseudomalaia whiteheadi Ohaus, 1912a: 267; Schultze 1916a: 174; Ohaus 1918: 152.</p><p>Type locality. “Nord Luzon.”</p><p>Distribution in the Philippines. Luzon (Ohaus 1912a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922F2828FF14F9C7FCDCF8DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922F282BFF14F8E7FCDCFF0C.text	3C6E87D6922F282BFF14F8E7FCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Popillia mcgregori Ohaus 1910	<div><p>Popillia mcgregori Ohaus, 1910</p><p>Popillia mcgregori Ohaus, 1910: 254; Ohaus 1918: 139.</p><p>Type locality. “ Luzon, Province of Benguet, Pauai, P. I.”</p><p>Distribution in the Philippines. Luzon (Benguet) (Ohaus 1910, 1918).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922F282BFF14F8E7FCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282BFF14FCFAFD33FBA4.text	3C6E87D6922C282BFF14FCFAFD33FBA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia bimaculata subsp. bimaculata (Guerin-Meneville 1843)	<div><p>Parastasia bimaculata bimaculata (Guérin-Méneville, 1843)</p><p>Barymorpha bimaculata Guérin-Méneville, 1843: 41 .</p><p>Parastasia bimaculata bimaculata: Kuijten 1992: 36.</p><p>Parastasia bimaculata: Hongsuwong &amp; Photaworn 2025: 92.</p><p>Type locality. “Pulo-Pinang.”</p><p>Distribution in the Philippines. Luzon (Palawan) and Mindanao (Kuijten 1992).</p><p>Overall distribution. The Philippines, India, Malaysia, Myanmar, Singapore, Thailand, and Indonesia (Hongsuwong &amp; Photaworn 2025).</p></div>	https://treatment.plazi.org/id/3C6E87D6922C282BFF14FCFAFD33FBA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282BFF14FB36FBD1FA44.text	3C6E87D6922C282BFF14FB36FBD1FA44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia burmeisteri Ohaus 1898	<div><p>Parastasia burmeisteri Ohaus, 1898</p><p>Parastasia burmeisteri Ohaus, 1898: 10; Ohaus 1900: 253; Ohaus 1918: 36; Ohaus 1934: 104; Kuijten 1992: 45.</p><p>Parastasia nonfriedi Ohaus, 1898: 10 . Type locality. “Mt. Kawi, Java.”; Ohaus 1910: 262.</p><p>Type locality. “ Mt. Kawi, Java; Sumatra.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ohaus 1910; Kuijten 1992).</p><p>Overall distribution. The Philippines, Malaysia, and Indonesia (Kuijten 1992).</p></div>	https://treatment.plazi.org/id/3C6E87D6922C282BFF14FB36FBD1FA44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282AFF14FA56FCD5FF0D.text	3C6E87D6922C282AFF14FA56FCD5FF0D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia canaliculata Westwood 1841	<div><p>Parastasia canaliculata Westwood, 1841</p><p>Parastasia canaliculata Westwood, 1841: 304; Hope 1841: 34; Hope 1846: 93; Blanchard 1851: 217; Ohaus, 1898: 18; Ohaus 1910: 261; Ohaus 1918: 35.</p><p>Parastasia bipunctata Westwood, 1841; 304. Type locality. “ Philippine Islands.”; Hope 1841: 34; Hope 1846: 94; Ohaus, 1898: 18.</p><p>Parastasia rubrotessellata Blanchard, 1851: 217 . Type locality. “ Philippines.”</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Batanes, Benguet, Bataan, Babuyan Islands, Camarines, Mindoro, Romblon, Sibuyan, Marinduque, and Rizal), Visayas (Bohol, Negros, Panay, Leyte, and Samar), and Mindanao (Bukidnon and Marawi) (Westwood 1841; 1842; Blanchard 1851; Ohaus 1910; Wada 1989; Kuijten 1992; Wada 2015).</p><p>Overall distribution. The Philippines and China (Taiwan) (Wada 2015).</p><p>Remarks. Kuijten (1992) placed Parastasia yukioi Wada, 1989 as a junior synonym of P. canaliculata . However, phylogenetic analyses conducted by Wada (2015), based on morphological characters, supported the recognition of P. yukioi as a separate and valid species. Wada highlighted distinguishing features such as the shape of the male genitalia, the structure of the inner sac, and a generally smaller body size. Therefore, P. yukioi will be treated as a distinct species.</p></div>	https://treatment.plazi.org/id/3C6E87D6922C282AFF14FA56FCD5FF0D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282BFF14FE9FFA8EFE08.text	3C6E87D6922C282BFF14FE9FFA8EFE08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Popillia mutans Newman 1838	<div><p>Popillia mutans Newman, 1838</p><p>Popillia mutans Newman, 1838: 337; Kraatz 1892: 240; Ohaus 1918: 139; Kim 2011: 71.</p><p>Popillia relucens Blanchard, 1851: 199 . Type locality. “ Manille.”</p><p>Type locality. “Ind. Orient.”</p><p>Distribution in the Philippines. Luzon (Manila) (Blanchard 1851; Ohaus 1918).</p><p>Overall distribution. The Philippines, Korea, Vietnam, and China (including Taiwan) (Kim 2011).</p></div>	https://treatment.plazi.org/id/3C6E87D6922C282BFF14FE9FFA8EFE08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922C282BFF14FE63FCDCFD70.text	3C6E87D6922C282BFF14FE63FCDCFD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Popillia variabilis Kraatz 1892	<div><p>Popillia variabilis Kraatz, 1892</p><p>Popillia variabilis Kraatz, 1892: 283; Ohaus 1910: 261; Ohaus 1918: 141.</p><p>Type locality. “Insulae Philippinae, Luzon.”</p><p>Distribution in the Philippines. Luzon (Benguet) (Kraatz 1892; Ohaus 1910).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922C282BFF14FE63FCDCFD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FE9FFE8DFDC1.text	3C6E87D6922D282AFF14FE9FFE8DFDC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia confluens Westwood 1841	<div><p>Parastasia confluens Westwood, 1841</p><p>Parastasia confluens Westwood, 1841: 304; Hope 1841: 35; Hope 1846: 95; Ohaus 1898: 12; Ohaus 1910: 261; Ohaus 1918: 32; Kuijten 1992: 54; Wada 2015: 49.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Manila and Sibuyan) and Mindanao (Marawi) (Westwood 1841; Kuijten 1992; Wada 2015).</p><p>Overall distribution. The Philippines, Malaysia, Indonesia, and Papua New Guinea (Kuijten 1992; Wada 2015).</p></div>	https://treatment.plazi.org/id/3C6E87D6922D282AFF14FE9FFE8DFDC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FDD8FCB8FC33.text	3C6E87D6922D282AFF14FDD8FCB8FC33.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia discolor subsp. discolor Westwood 1841	<div><p>Parastasia discolor discolor Westwood, 1841</p><p>Parastasia discolor Westwood, 1841: 304; Hope 1841: 34; Hope 1846: 94; Ohaus 1900: 244; Schultze 1916a: 175; Ohaus 1918: 34.</p><p>Parastasia discolor discolor: Wada 2015: 132.</p><p>Type locality. “ Philippines.”</p><p>Distribution in the Philippines. Luzon (Bicol), Visayas (Panay, Negros, Samar, and Leyte), and Mindanao (Bukidnon, Agusan del Sur, Sarangani, Misamis Oriental, Surigao, and Davao) (Westwood 1841; Kuijten 1992; Wada 2015).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p><p>Remarks. In the article by Schultze (1916a), P. discolor scutellaris was noted to occur on the island of Luzon. However, no evidence to date of its existence has been documented. This may be erroneous and mislabeled as ‘ scutellaris ’ instead of ‘ discolor ’.</p></div>	https://treatment.plazi.org/id/3C6E87D6922D282AFF14FDD8FCB8FC33	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FB8AFC79FB3C.text	3C6E87D6922D282AFF14FB8AFC79FB3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia incurva Ohaus 1923	<div><p>Parastasia incurva Ohaus, 1923</p><p>Parastasia incurva Ohaus, 1923: 581; Kuijten 1992: 89; Wada 2015: 235.</p><p>Type locality. “ Luzon, Kalinga, Balbalan.”</p><p>Distribution in the Philippines. Luzon (Kalinga, Mountain Province, Camiguin, Cagayan Province, and Masbate) (Ohaus 1923; Kuijten 1992; Wada 2015).</p><p>Overall distribution. The Philippines and Sulawesi (Wada 2015).</p></div>	https://treatment.plazi.org/id/3C6E87D6922D282AFF14FB8AFC79FB3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14FA8FFCDCFA39.text	3C6E87D6922D282AFF14FA8FFCDCFA39.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia isidai Wada 1989	<div><p>Parastasia isidai Wada, 1989</p><p>Parastasia isidai Wada, 1989: 2; Kuijten 1992: 85 (synonymized with Parastasia helleri); Wada 2003: 3 (valid species); Wada 2015: 145.</p><p>Type locality. “ Mt. Canlaon, Negros Is., Philippines.”</p><p>Distribution in the Philippines. Visayas (Negros) (Wada 1989, 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922D282AFF14FA8FFCDCFA39	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14F990FCDCF96E.text	3C6E87D6922D282AFF14F990FCDCF96E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia jamesonae Wada 2008	<div><p>Parastasia jamesonae Wada, 2008</p><p>Parastasia jamesonae Wada, 2008: 97; Wada 2015: 198.</p><p>Type locality. “ Cuernos Mts., Negros, Philippines.”</p><p>Distribution in the Philippines. Visayas (Negros) (Wada 2008).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922D282AFF14F990FCDCF96E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282AFF14F978FCDCF857.text	3C6E87D6922D282AFF14F978FCDCF857.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia luzonensis Wada 2016	<div><p>Parastasia luzonensis Wada, 2016</p><p>Parastasia luzonensis Wada, 2016: 123 .</p><p>Type locality. “ Mt. Polis, Mt. Province, North Luzon, Philippines.”</p><p>Distribution in the Philippines. Luzon (Mountain Province) (Wada 2016).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922D282AFF14F978FCDCF857	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922D282DFF14F821FCDCFF21.text	3C6E87D6922D282DFF14F821FCDCFF21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia mitsumata Wada 2007	<div><p>Parastasia mitsumata Wada, 2007</p><p>Parastasia mitsumata Wada, 2007: 498; Wada 2015: 203.</p><p>Type locality. “ Mt. Dolang dolang, Malaybarlay, Bukidnon, Mindanao Is.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon, South Cotabato, and Davao) (Wada 2007, 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922D282DFF14F821FCDCFF21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FEBBFCDCFE16.text	3C6E87D6922A282DFF14FEBBFCDCFE16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia negrosensis Wada 1997	<div><p>Parastasia negrosensis Wada, 1997</p><p>Parastasia negrosensis Wada, 1997: 422; Wada 2015: 96.</p><p>Type locality. “ Mt. Canlaon, Negros Is., Philippine Isls.”</p><p>Distribution in the Philippines. Visayas (Negros Island) (Wada 1997).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922A282DFF14FEBBFCDCFE16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FE60FC27FCCB.text	3C6E87D6922A282DFF14FE60FC27FCCB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia nigriceps subsp. nigriceps Westwood 1841	<div><p>Parastasia nigriceps nigriceps Westwood, 1841</p><p>Parastasia nigriceps Westwood, 1841: 304; Hope 1841: 35; Hope 1846: 94; Ohaus 1900: 233; Ohaus 1910: 262; Dalla Torre 1912a: 33; Kuijten 1992: 118.</p><p>Parastasia nigriceps nigriceps: Wada 2015: 98.</p><p>Type locality. “ Philippine Islands.”</p><p>Distribution in the Philippines. Luzon (Marinduque, Romblon, Bicol, and Mindoro), Visayas (Negros), and Mindanao (Westwood 1841; Fairmaire 1883; Ohaus 1900; Kuijten 1992).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922A282DFF14FE60FC27FCCB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FCDDFCDCFBE8.text	3C6E87D6922A282DFF14FCDDFCDCFBE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia nigroscutellata Ohaus 1901	<div><p>Parastasia nigroscutellata Ohaus, 1901</p><p>Parastasia nigroscutellata Ohaus, 1901: 125; Ohaus 1905: 96; Schultze 1916a: 176; Ohaus 1918: 33; Ohaus 1930a: 555; Wada 2015: 104.</p><p>Type locality. “Luzon (Engano).”</p><p>Distribution in the Philippines. Luzon (Cagayan Province, Laguna, Mindoro, Quirino, and Quezon Province) (Ohaus 1901, 1905, 1930a; Kuijten 1992; Wada 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922A282DFF14FCDDFCDCFBE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FBC2FCDCFAD1.text	3C6E87D6922A282DFF14FBC2FCDCFAD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia takeshii Wada 1997	<div><p>Parastasia takeshii Wada, 1997</p><p>Parastasia takeshii Wada, 1997: 420; Wada 2015: 116.</p><p>Type locality. “ Mt. Kitanglad, North Mindanao Is., Philippines Isls.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon) (Wada 1997).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922A282DFF14FBC2FCDCFAD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14FAABFAD3F9DD.text	3C6E87D6922A282DFF14FAABFAD3F9DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia westwoodi Westwood 1841	<div><p>Parastasia westwoodi Westwood, 1841</p><p>Parastasia westwoodi Westwood, 1841: 304; Ohaus 1910: 262; Ohaus 1918: 36; Wada 2015: 252.</p><p>Type locality. “ Sumatra.”</p><p>Distribution in the Philippines. Luzon (Palawan, Mountain Province, Quezon, and Mindoro) and Mindanao (Zamboanga) (Ohaus 1910, 1918; Wada,2015).</p><p>Overall distribution. The Philippines, Indonesia, Malaysia, and Thailand (Kuijten 1992; Wada 2015).</p></div>	https://treatment.plazi.org/id/3C6E87D6922A282DFF14FAABFAD3F9DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282DFF14F9ACFCDCF926.text	3C6E87D6922A282DFF14F9ACFCDCF926.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parastasia yukioi Wada 1989	<div><p>Parastasia yukioi Wada, 1989</p><p>Parastasia yukioi Wada, 1989: 3; Kuijten 1992: 48 (= Parastasia canaliculata); Wada 2015: 217 (valid species).</p><p>Type locality. “Bindoyan, Is. Palawan, Philippines.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Wada 1989, 2015).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922A282DFF14F9ACFCDCF926	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922A282CFF14F8E9FCDCFF0C.text	3C6E87D6922A282CFF14F8E9FCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lutera boettcheri Ohaus 1916	<div><p>Lutera boettcheri Ohaus, 1916</p><p>Lutera boettcheri Ohaus, 1916: 353; Ohaus 1918: 39; Ohaus 1934: 112; Machatschke 1972: 45; Kuijten 1988: 86.</p><p>Type locality. “ Nord-Luzon, Mt. Imugan. ”</p><p>Distribution in the Philippines. Luzon (Nueva Vizcaya) (Ohaus 1916).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922A282CFF14F8E9FCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282CFF14FAA7FCDCFA3C.text	3C6E87D6922B282CFF14FAA7FCDCFA3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haroldius globosus Boucomont 1925	<div><p>Haroldius globosus Boucomont, 1925</p><p>Haroldius globosus Boucomont, 1925: 151; Pereira 1954: 7; Balthasar 1963a: 250; Krajčík 2006: 68.</p><p>Type locality. “Luzon: Mont Banahao.”</p><p>Distribution in the Philippines. Luzon (Mount Banahao) (Boucomont 1925).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922B282CFF14FAA7FCDCFA3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282CFF14F98FFCDCF964.text	3C6E87D6922B282CFF14F98FFCDCF964.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haroldius philippinensis Pereira 1954	<div><p>Haroldius philippinensis Pereira, 1954</p><p>Haroldius philippinensis Pereira, 1954: 1; Balthasar 1963a: 250; Krajčík 2006: 69.</p><p>Type locality. “ Dingalan Bay, East Luzon, Philippines Islands.”</p><p>Distribution in the Philippines. Luzon (Aurora) (Pereira 1954).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922B282CFF14F98FFCDCF964	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282CFF14FE9EFDE3FD28.text	3C6E87D6922B282CFF14FE9EFDE3FD28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lutera nigrita Ohaus 1930	<div><p>Lutera nigrita Ohaus, 1930</p><p>Lutera nigromaculata (nec Ohaus, 1900); Ohaus 1905: 97; Ohaus 1918: 39.</p><p>Lutera nigromaculata nigrita Ohaus, 1930b: 140 .</p><p>Parastasia nigrita: Machatschke 1972: 45.</p><p>Lutera nigrita: Wada 1998b: 360; Xu &amp; Zhao 2024: 538.</p><p>Type locality. “ Luzon, Mt. Banahao. ”</p><p>Distribution in the Philippines. Luzon (Aurora, Camarines Sur, Laguna, Mindoro, and Mt. Banahao), Visayas (Negros Occidental), and Mindanao (Davao de Oro, Davao del Sur, Kidapawan, and Lanao del Sur) (Ohaus 1930b; Wada 1998b; Xu &amp; Zhao 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks. The longstanding taxonomic confusion regarding the presence of Lutera nigromaculata Ohaus, 1900, in the Philippines was recently resolved by Xu and Zhao(2024).The specimen of L.nigromaculata from Camarines, examined by Ohaus and currently deposited at the MFNB in Berlin, Germany, is likely the first known individual of L. nigrita .</p></div>	https://treatment.plazi.org/id/3C6E87D6922B282CFF14FE9EFDE3FD28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282CFF14FCFBFBE8FBEC.text	3C6E87D6922B282CFF14FCFBFBE8FBEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rutelarcha bakeri Ohaus 1915	<div><p>Rutelarcha bakeri Ohaus, 1915</p><p>Rutelarcha bakeri Ohaus, 1915b: 343; Ohaus 1934: 113; Machatschke 1972: 46; Kuijten 1988: 82.</p><p>Type locality. “ Philippinen: Baguio auf Benguet.”</p><p>Distribution in the Philippines. Luzon (Baguio, Mt. Banahao, and Nueva Vizcaya) and Mindanao (? Bukidnon) (Ohaus 1915b; Kuijten 1988).</p><p>Overall distribution. The Philippines, Indonesia, and Vietnam (Kuijten 1988).</p></div>	https://treatment.plazi.org/id/3C6E87D6922B282CFF14FCFBFBE8FBEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6922B282FFF14F8E7FCDCFE99.text	3C6E87D6922B282FFF14F8E7FCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catharsius aethiops Sharp 1875	<div><p>Catharsius aethiops Sharp, 1875</p><p>Catharsius aethiops Sharp, 1875: 41; Schultze 1916a: 160; Balthasar 1963a: 309; Krajčík 2006: 33.</p><p>Catharsius molossus var. aethiops Boucomont 1914: 330 .</p><p>Type locality. “Ins. Philippines.”</p><p>Distribution in the Philippines. Luzon (Bataan, Benguet, Mindoro, Sibuyan, and Palawan), Visayas (Negros Occidental), and Mindanao (Bukidnon, Davao, Marawi, and Misamis Oriental) (Sharp 1875; Boucomont 1914; Schultze 1916a; Balthasar 1963a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6922B282FFF14F8E7FCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14FEF0FE2FFCA6.text	3C6E87D69228282FFF14FEF0FE2FFCA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Catharsius molossus (Linnaeus 1758)	<div><p>Catharsius molossus (Linnaeus, 1758)</p><p>Scarabaeus molossus Linnaeus, 1758: 347 .</p><p>Catharsius molossus: Schultze 1916a: 160.</p><p>Type locality. “ India.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Schultze 1916a).</p><p>Overall distribution. The Philippines, India, Sri Lanka, Afghanistan, Nepal, China (including Taiwan), Vietnam, Laos, Cambodia, Thailand, Malaysia, Pakistan, and Singapore.</p><p>Remarks.Although records of Catharsius molossus from the Philippines are scarce in the literature, the species is known to occur in Palawan and is entirely absent from other regions of the archipelago, as confirmed by Ochi and Kon (pers. comm.). During a recent entomological survey in Bukidnon, Mindanao, the first author collected Catharsius specimens that differed markedly from other Mindanao populations by having entirely reddish-brown pubescence. These specimens closely resembled C. molossus, and after a detailed examination, assisted by Ochi and Kon, they were confirmed to be C. molossus . A more comprehensive treatment of the geographic distribution of Catharsius species in the Philippines will be presented in a separate paper.</p></div>	https://treatment.plazi.org/id/3C6E87D69228282FFF14FEF0FE2FFCA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14FBA6FCDCFAF5.text	3C6E87D69228282FFF14FBA6FCDCFAF5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copris tetraodon Gillet 1910	<div><p>Copris (sensu lato) tetraodon Gillet, 1910</p><p>Copris tetraodon Gillet, 1910: 11; Boucomont 1914: 334; Schultze 1916a: 160; Balthasar 1963a: 344; Krajčík 2006: 45; Zidek 2020: 11.</p><p>Type locality. “ Iles Philippines.”</p><p>Distribution in the Philippines. Luzon (Mindoro and Pampanga) and Mindanao (Agusan River) (Gillet 1910; Boucomont 1914; Schultze 1916a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69228282FFF14FBA6FCDCFAF5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14FA3FFCDCF9B6.text	3C6E87D69228282FFF14FA3FFCDCF9B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microcopris mindorensis Ochi 1992	<div><p>Microcopris mindorensis Ochi, 1992</p><p>Copris (Microcopris) mindorensis Ochi, 1992: 13; Krajčík 2006: 43; Kaneko &amp; Wada 2020: 39.</p><p>Microcopris mindorensis: Zidek 2020: 13.</p><p>Type locality. “ Philippines (Mindoro Is.).”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Ochi, 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69228282FFF14FA3FFCDCF9B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69228282FFF14F8B1FCDCF823.text	3C6E87D69228282FFF14F8B1FCDCF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ochicanthon philippinense (Ochi 1990)	<div><p>Ochicanthon philippinense (Ochi, 1990)</p><p>Phacosoma philippinense Ochi, 1990: 217 .</p><p>Ochicanthon philippinense: Krajčík 2006: 77; Krikken &amp; Huijbregts 2007: 426 ( philippinensis).</p><p>Type locality. “ Mt. Apo, Mindanao Is., Philippines.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi 1990).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69228282FFF14F8B1FCDCF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14FDD9FCDCFD37.text	3C6E87D69229282EFF14FDD9FCDCFD37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Panelus orousseti Paulian 1981	<div><p>Panelus (sensu lato) orousseti Paulian, 1981</p><p>Panelus orousseti Paulian, 1981: 110; Krajčík 2006: 144.</p><p>Type locality. “ Philippines, Luzon, Mountain Province, Sagoda.”</p><p>Distribution in the Philippines. Luzon (Mountain Province) (Paulian 1981).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69229282EFF14FDD9FCDCFD37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14FEF0FCDCFDCE.text	3C6E87D69229282EFF14FEF0FCDCFDCE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Panelus philippinensis Ochi, Kon & Cabras 2025	<div><p>Panelus (sensu lato) philippinensis Ochi, Kon &amp; Cabras, 2025</p><p>Panelus (incertae sedis) philippinensis Ochi, Kon &amp; Cabras, 2025a: 285 .</p><p>Type locality. “ Philippines, Luzon Is., Queson City.”</p><p>Distribution in the Philippines. Luzon (Quezon) (Ochi et al. 2025a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69229282EFF14FEF0FCDCFDCE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14FB57FC27FA41.text	3C6E87D69229282EFF14FB57FC27FA41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paragymnopleurus sparsus subsp. palawanicus Ochi & Kon 1997	<div><p>Paragymnopleurus sparsus palawanicus Ochi &amp; Kon, 1997</p><p>Gymnopleurus (Paragymnopleurus) sparsus: Balthasar 1963a: 221; Ochi &amp; Kon 1994: 282;</p><p>Paragymnopleurus sparsus palawanicus Ochi &amp; Kon, 1997: 236; Krajčík 2006: 67.</p><p>Type locality. “ Philippines, Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Boucomont 1914; Ochi &amp; Kon 1997).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69229282EFF14FB57FC27FA41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14FC36FB38FB44.text	3C6E87D69229282EFF14FC36FB38FB44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paragymnopleurus stipes (Sharp 1875)	<div><p>Paragymnopleurus stipes (Sharp, 1875)</p><p>Gymnopleurus stipes Sharp, 1875: 35; Krajčík 2006: 67.</p><p>Gymnopleurus (Paragymnopleurus) stipes: Balthasar 1963a: 217.</p><p>Paragymnopleurus stipes stipes: Pokorný &amp; Zídek 2014.</p><p>Type locality. “Ins. Philippines.”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Sharp 1875; Boucomont 1914).</p><p>Overall distribution. Endemic species to the Philippines (Pokorný &amp; Zídek 2014).</p></div>	https://treatment.plazi.org/id/3C6E87D69229282EFF14FC36FB38FB44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69229282EFF14F9C8FECEF8B3.text	3C6E87D69229282EFF14F9C8FECEF8B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tibiodrepanus hircus (Wiedemann 1823)	<div><p>Tibiodrepanus hircus (Wiedemann, 1823)</p><p>Copris hircus Wiedemann, 1823: 19 .</p><p>Tibiodrepanus hircus: Krikken 2009: 21; Barbero et al. 2011: 33.</p><p>Type locality. “Java.”</p><p>Distribution in the Philippines. Luzon (Bataan, Laguna, and Rizal) and Visayas (Cebu and Negros Oriental) (Barbero et al. 2011).</p><p>Overall distribution. The Philippines, China, India, Indonesia, Myanmar, Sri Lanka, and Vietnam (Barbero et al. 2011).</p></div>	https://treatment.plazi.org/id/3C6E87D69229282EFF14F9C8FECEF8B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692292821FF14F80AFCDAFEC5.text	3C6E87D692292821FF14F80AFCDAFEC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tibiodrepanus setosus (Wiedemann 1823)	<div><p>Tibiodrepanus setosus (Wiedemann, 1823)</p><p>Copris setosa Wiedemann, 1823: 19 .</p><p>Tibiodrepanus setosus: Barbero et al. 2011: 36.</p><p>Type locality. “Bengalia.”</p><p>Distribution in the Philippines. Luzon (Laguna) (Barbero et al. 2011).</p><p>Overall distribution. The Philippines, Pakistan, India, Bhutan, Nepal, Sri Lanka, China, Myanmar, Laos, Thailand, Vietnam, and Indonesia (Barbero et al. 2011).</p></div>	https://treatment.plazi.org/id/3C6E87D692292821FF14F80AFCDAFEC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262821FF14F98FFCDCF939.text	3C6E87D692262821FF14F98FFCDCF939.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anoctus rasilis (Boucomont 1921)	<div><p>Anoctus rasilis (Boucomont, 1921)</p><p>Onthophagus rasilis Boucomont, 1921: 91; Boucomont 1924a: 676.</p><p>Anoctus rasilis: Balthasar 1963b: 150; Krajčík 2006: 11.</p><p>Type locality. “ Iles Philippines, Mindanao: Iligan.”</p><p>Distribution in the Philippines. Mindanao (Lanao del Norte) (Boucomont 1921, 1924a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692262821FF14F98FFCDCF939	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262821FF14FD67FE8CFBED.text	3C6E87D692262821FF14FD67FE8CFBED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onitis falcatus (Wulfen 1786)	<div><p>Onitis falcatus (Wulfen, 1786)</p><p>Scarabaeus falcatus Wulfen, 1786: 14 .</p><p>Onitis falcatus: Lansberge 1875: 126; Janssens 1937: 44; Balthasar 1963b: 33; Cheung et al. 2018: 245; Han et al. 2021: 138.</p><p>Type locality. “ India.”</p><p>Distribution in the Philippines. Luzon and Mindanao (Lansberge 1875; Balthasar 1963b; Cheung et al. 2018).</p><p>Overall distribution. This species is widely distributed in the Oriental and Palearctic regions (Han et al. 2021).</p></div>	https://treatment.plazi.org/id/3C6E87D692262821FF14FD67FE8CFBED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262821FF14FBFFFC46FA8D.text	3C6E87D692262821FF14FBFFFC46FA8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onitis phartopus Lansberge 1875	<div><p>Onitis phartopus Lansberge, 1875</p><p>Onitis phartopus Lansberge, 1875: 128; Boucomont 1914: 336; Janssens 1937: 45; Balthasar 1963b: 33; Krajčík 2006: 82.</p><p>Type locality. “ Iles Philippines.”</p><p>Distribution in the Philippines. Luzon (Laguna, Manila, and Benguet) (Lansberge 1875; Boucomont 1914; Schultze 1916a; Balthasar 1963b).</p><p>Overall distribution. The Philippines and Indonesia (Janssens 1937).</p></div>	https://treatment.plazi.org/id/3C6E87D692262821FF14FBFFFC46FA8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262821FF14FED7FCCCFDE5.text	3C6E87D692262821FF14FED7FCCCFDE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tibiodrepanus sinicus (Harold 1868)	<div><p>Tibiodrepanus sinicus (Harold, 1868)</p><p>Drepanocerus sinicus Harold, 1868: 104 .</p><p>Tibiodrepanus sinicus: Barbero et al. 2011: 40; Han et al. 2021: 134.</p><p>Type locality. “Kinb. China.”</p><p>Distribution in the Philippines. Luzon (Laguna) (Barbero et al. 2011).</p><p>Overall distribution. The Philippines, Bhutan, India, Nepal, China, Myanmar, Laos, Thailand, Vietnam, Singapore, Indonesia, and Malaysia (Barbero et al. 2011).</p></div>	https://treatment.plazi.org/id/3C6E87D692262821FF14FED7FCCCFDE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692262820FF14F803FBE1FE74.text	3C6E87D692262820FF14F803FBE1FE74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caccobius (Caccophilus) unicornis (Fabricius 1798)	<div><p>Caccobius (Caccophilus) unicornis (Fabricius, 1798)</p><p>Copris unicornis Fabricius, 1798: 33 .</p><p>Caccobius (Caccophilus) unicornis: Balthasar 1963b: 142; Nomura 1973: 47; Ochi &amp; Kon 1994: 293; Chen et al. 2002: 55; Chandra &amp; Gupta 2013: 343; Cheung et al. 2018: 247; Han et al. 2021: 134; Ueda et al. 2022: 180.</p><p>Type locality. “ Tranquebar.”</p><p>Distribution in the Philippines. Philippine archipelago (Arrow 1931; Balthasar 1963b; Nomura 1973; Ochi &amp; Kon 1994).</p><p>Overall distribution. The Philippines, China, India, Indonesia, Japan, Laos, Malaysia, North Korea, Sri Lanka, and Vietnam (Cheung et al. 2018; Han et al. 2021; Ueda et al. 2022).</p></div>	https://treatment.plazi.org/id/3C6E87D692262820FF14F803FBE1FE74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692272820FF14FDBFFCDCFD14.text	3C6E87D692272820FF14FDBFFCDCFD14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyobius philippinensis Ochi, Kon & Cabras 2025	<div><p>Cyobius philippinensis Ochi, Kon &amp; Cabras, 2025</p><p>Cyobius philippinensis Och, Kon &amp; Cabras, 2025a: 286 .</p><p>Type locality. “ Philippines, Luzon Is., Rizai, Tanay.”</p><p>Distribution in the Philippines. Luzon (Rizal) (Ochi et al. 2025a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692272820FF14FDBFFCDCFD14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692272820FF14FC17FCD8FAE8.text	3C6E87D692272820FF14FC17FCD8FAE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Colobonthophagus) armatus Blanchard 1853	<div><p>Onthophagus (Colobonthophagus) armatus Blanchard, 1853</p><p>Onthophagus armatus Blanchard, 1853: 98; Boucomont 1924a: 671.</p><p>Onthophagus luzonicus Lansberge, 1883: 41 . Type locality. “Java, Sumatra.”; Krajčík 2006: 90.</p><p>Onthophagus pseudoarmatus Balthasar, 1944: 93 . Type locality. “ China, Fukien.”</p><p>Onthophagus (Onthophagus) armatus: Balthasar 1963b: 277; Nomura 1973: 49.</p><p>Onthophagus (Colobonthophagus) armatus: Löbl &amp; Smetana 2006: 163; Cheung et al. 2018: 249; Han et al. 2021: 135; Ueda et al. 2022: 185.</p><p>Type locality. “Borneo.”</p><p>Distribution in the Philippines. Luzon (Manila and Mindoro) and Visayas (Negros Occidental) (Blanchard 1853; Lansberge 1875; Boucomont 1914; Balthasar 1944, 1963b).</p><p>Overall distribution. The Philippines, Cambodia, China (including Taiwan), Guam, India, Indonesia, Myanmar, and Vietnam (Balthasar 1944; Cheung et al. 2018).</p></div>	https://treatment.plazi.org/id/3C6E87D692272820FF14FC17FCD8FAE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692272820FF14FA3BFCDCF940.text	3C6E87D692272820FF14FA3BFCDCF940.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Endrodius) baeri Boucomont 1914	<div><p>Onthophagus (Endrodius) baeri Boucomont, 1914</p><p>Onthophagus baeri Boucomont, 1914: 307; Boucomont 1924a: 669; Krajčík 2006: 91.</p><p>Onthophagus (Endrodius) baeri: Balthasar 1963b: 284.</p><p>Type locality. “ Mindanao: Davao.”</p><p>Distribution in the Philippines. Mindanao (Davao, Lanao del Norte, and Misamis Oriental) (Boucomont 1914, 1924a; Balthasar 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692272820FF14FA3BFCDCF940	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692272820FF14F95BFCDCF860.text	3C6E87D692272820FF14F95BFCDCF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Endrodius) gaesatus Boucomont 1925	<div><p>Onthophagus (Endrodius) gaesatus Boucomont, 1925</p><p>Onthophagus gaesatus Boucomont, 1925: 152; Krajčík 2006: 106.</p><p>Onthophagus (Endrodius) gaesatus: Balthasar 1963b: 363.</p><p>Type locality. “Mindanao: Surigao.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon, Lanao del Norte, Misamis Oriental, and Surigao) (Boucomont 1925; Balthasar 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692272820FF14F95BFCDCF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FF0BFCDCFE2D.text	3C6E87D692242823FF14FF0BFCDCFE2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Endrodius) praedatus Harold 1862	<div><p>Onthophagus (Endrodius) praedatus Harold, 1862</p><p>Onthophagus praedatus Harold, 1862: 403; Boucomont 1914: 307; Boucomont 1924a: 670; Krajčík 2006: 126.</p><p>Onthophagus verticalis Boheman, 1858: 44 .</p><p>Onthophagus ternatensis Boucomont, 1914: 307 . Type locality. “ Ternate.”</p><p>Onthophagus (Endrodius) praedatus: Balthasar 1963b: 482.</p><p>Type locality. “ Manilla.”</p><p>Distribution in the Philippines. Luzon (Manila and Laguna) and Mindanao (Agusan del Norte and Misamis Oriental) (Boheman 1858; Harold 1862a; Boucomont 1914).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692242823FF14FF0BFCDCFE2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FDF7FB2BFC7C.text	3C6E87D692242823FF14FDF7FB2BFC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Gibbonthophagus) cervicapra Boucomont 1914	<div><p>Onthophagus (Gibbonthophagus) cervicapra Boucomont, 1914</p><p>Onthophagus cervicapra Boucomont, 1914: 294; Boucomont 1924b: 114.</p><p>Onthophagus verecundus Boucomont, 1921: 89 . Type locality. “ Bornéo: Martapura: île Banguey; Philippines: Palawan; Java: Boeloe Lawang.”</p><p>Onthophagus (Onthophagus) cervicapra: Balthasar 1963b: 306; Ochi &amp; Kon 1995: 55.</p><p>Type locality. “Borneo.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Boucomont 1914, 1921, 1924b; Balthasar 1963b; Ochi &amp; Kon 1995).</p><p>Overall distribution. The Philippines, Indonesia, and Malaysia (Ochi &amp; Kon 1995).</p></div>	https://treatment.plazi.org/id/3C6E87D692242823FF14FDF7FB2BFC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FC4FFCDCFB1D.text	3C6E87D692242823FF14FC4FFCDCFB1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Gibbonthophagus) eschscholtzi Boucomont 1924	<div><p>Onthophagus (Gibbonthophagus) eschscholtzi Boucomont, 1924</p><p>Onthophagus eschscholtzi Boucomont, 1924a: 675 .</p><p>Onthophagus (Onthophagus) eschscholtzi: Balthasar 1963b: 343; Krajčík 2006: 102.</p><p>Onthophagus (Gibbonthophagus) eschscholtzi: Ochi &amp; Kon 2025c: 48.</p><p>Type locality. “ Palawan, Binaluan.”</p><p>Distribution in the Philippines. Luzon (Palawan) and Mindanao (Boucomont 1924a; Balthasar 1963b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692242823FF14FC4FFCDCFB1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FB6FFB93FA45.text	3C6E87D692242823FF14FB6FFB93FA45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Gibbonthophagus) kiyoshii Ochi & Kon 2007	<div><p>Onthophagus (Gibbonthophagus) kiyoshii Ochi &amp; Kon, 2007</p><p>Onthophagus (Gibbonthophagus) kiyoshii Ochi &amp; Kon, 2007: 89 .</p><p>Type locality. “ Rizal, Palawan Is., the Philippines.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ochi &amp; Kon 2007).</p><p>Overall distribution. The Philippines and? Thailand (Ochi &amp; Kon 2007).</p></div>	https://treatment.plazi.org/id/3C6E87D692242823FF14FB6FFB93FA45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14FA57FD71F939.text	3C6E87D692242823FF14FA57FD71F939.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Gibbonthophagus) luridipennis Boheman 1858	<div><p>Onthophagus (Gibbonthophagus) luridipennis Boheman, 1858</p><p>Onthophagus luridipennis Boheman, 1858: 45 .</p><p>Onthophagus (Onthophagus) luridipennis: Balthasar 1963b: 424.</p><p>Onthophagus (Gibbonthophagus) luridipennis: Han et al. 2021: 135.</p><p>Type locality. “ China (Hongkong).”</p><p>Distribution in the Philippines. Philippine archipelago (Balthasar 1963b).</p><p>Overall distribution. The Philippines, China (including Taiwan), Cambodia, India, Indonesia, Myanmar, Thailand, and Vietnam (Han et al. 2021).</p></div>	https://treatment.plazi.org/id/3C6E87D692242823FF14FA57FD71F939	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692242823FF14F893FCDCF861.text	3C6E87D692242823FF14F893FCDCF861.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Gibbonthophagus) megasubcornutus Ochi & Kon 2025	<div><p>Onthophagus (Gibbonthophagus) megasubcornutus Ochi &amp; Kon, 2025</p><p>Onthophagus (Gibbonthophagus) megasubcornutus Ochi &amp; Kon, 2025c: 44 .</p><p>Type locality. “ Philippines, Mindanao Is., South Lanao. ”</p><p>Distribution in the Philippines. Mindanao (Lanao del Sur) (Ochi &amp; Kon 2025c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692242823FF14F893FCDCF861	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FF0BFCDCFE99.text	3C6E87D692252822FF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Gibbonthophagus) palawanicus Ochi & Kon 2004	<div><p>Onthophagus (Gibbonthophagus) palawanicus Ochi &amp; Kon, 2004</p><p>Onthophagus (Gibbonthophagus) palawanicus Ochi &amp; Kon, 2004: 34; Krajčík 2006: 123.</p><p>Type locality. “ Brooks Point, Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ochi &amp; Kon 2004).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692252822FF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FEF3FCDCFDE5.text	3C6E87D692252822FF14FEF3FCDCFDE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Gibbonthophagus) sasajii Ochi & Kon 2001	<div><p>Onthophagus (Gibbonthophagus) sasajii Ochi &amp; Kon, 2001</p><p>Onthophagus (Gibbonthophagus) sasajii Ochi &amp; Kon, 2001: 372; Krajčík 2006: 131; Ochi &amp; Kon 2007: 91.</p><p>Type locality. “ Bicol National Park, Nr. Daet, Central Luzon. ”</p><p>Distribution in the Philippines. Luzon (Palawan, Bicol, Baguio, and Negros) (Ochi &amp; Kon 2001, 2007).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692252822FF14FEF3FCDCFDE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FD2FFAD1FC11.text	3C6E87D692252822FF14FD2FFAD1FC11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Hikidaeus) pastillatus Boucomont 1919	<div><p>Onthophagus (Hikidaeus) pastillatus Boucomont, 1919</p><p>Onthophagus pastillatus Boucomont, 1919: 318; Boucomont 1921: 92; Boucomont 1924a: 674.</p><p>Onthophagus (Onthophagus) pastillatus: Balthasar 1963b: 469; Ochi &amp; Kon 1995: 52.</p><p>Onthophagus (Hikidaeus) pastillatus: Ueda et al. 2022: 185.</p><p>Type locality. “Borneo: Sandakan.”</p><p>Distribution in the Philippines. Mindanao (Lanao del Norte) (Boucomont 1921, 1924a; Balthasar 1963b; Ochi &amp; Kon 1995).</p><p>Overall distribution. The Philippines, Indonesia, and Malaysia (Ochi &amp; Kon 1995; Ueda et al. 2022).</p></div>	https://treatment.plazi.org/id/3C6E87D692252822FF14FD2FFAD1FC11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FC6BFCDCFB79.text	3C6E87D692252822FF14FC6BFCDCFB79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Hikidaeus) semiazusae Ochi, Kon & Cabras 2023	<div><p>Onthophagus (Hikidaeus) semiazusae Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Hikidaeus) semiazusae Ochi, Kon &amp; Cabras, 2023b: 52 .</p><p>Type locality. “ Philippines, Mindanao Isl., Bukidnon.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon and Lanao del Sur) (Ochi et al. 2023b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692252822FF14FC6BFCDCFB79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14FA8BFC27FA3D.text	3C6E87D692252822FF14FA8BFC27FA3D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Indachorius) aereopictus subsp. aereopictus Boucomont 1914	<div><p>Onthophagus (Indachorius) aereopictus aereopictus Boucomont, 1914</p><p>Onthophagus aereopictus Boucomont, 1914: 326; Boucomont 1924a: 678.</p><p>Onthophagus (Indachorius) aereopictus: Balthasar 1963b: 265; Krajčík 2006: 87; Ochi et al. 2023a: 47.</p><p>Type locality. “ Philippines: Palawan; Los Banos, Luzon.”</p><p>Distribution in the Philippines. Luzon (Laguna and Palawan) (Boucomont 1914).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692252822FF14FA8BFC27FA3D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14F98FFCDCF965.text	3C6E87D692252822FF14F98FFCDCF965.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Indachorius) agusanensis Ochi, Kon & Cabras 2023	<div><p>Onthophagus (Indachorius) agusanensis Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Indachorius) agusanensis Ochi, Kon &amp; Cabras, 2023a: 38 .</p><p>Type locality. “ Philippines, Mindanao Is.”</p><p>Distribution in the Philippines. Mindanao (Davao, Agusan, and Cotabato) (Ochi et al. 2023a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692252822FF14F98FFCDCF965	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692252822FF14F977FCDCF84D.text	3C6E87D692252822FF14F977FCDCF84D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Indachorius) basimaculatus Ochi, Kon & Cabras 2023	<div><p>Onthophagus (Indachorius) basimaculatus Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Indachorius) basimaculatus Ochi, Kon &amp; Cabras, 2023a: 42 .</p><p>Type locality. “ Philippines, Luzon Is., Bicol.”</p><p>Distribution in the Philippines. Luzon (Bicol) (Ochi et al. 2023a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692252822FF14F977FCDCF84D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FF0BFCDCFE99.text	3C6E87D692222825FF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Indachorius) davaoensis Ochi, Kon & Cabras 2023	<div><p>Onthophagus (Indachorius) davaoensis Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Indachorius) davaoensis Ochi, Kon &amp; Cabras, 2023a: 39 .</p><p>Type locality. “ Philippines, Mindanao Is., Davao.”</p><p>Distribution in the Philippines. Mindanao (Davao) (Ochi et al. 2023a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692222825FF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FEF3FCDCFDC1.text	3C6E87D692222825FF14FEF3FCDCFDC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Indachorius) lanaodelsurensis Ochi, Kon & Cabras 2023	<div><p>Onthophagus (Indachorius) lanaodelsurensis Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Indachorius) leyteinsulanus Ochi, Kon &amp; Cabras, 2023a: 46 .</p><p>Type locality. “ Philippines, Mindanao Is., Lanao del Sur.”</p><p>Distribution in the Philippines. Mindanao (Lanao del Sur) (Ochi et al. 2023a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692222825FF14FEF3FCDCFDC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FDDBFCDCFD29.text	3C6E87D692222825FF14FDDBFCDCFD29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Indachorius) leyteinsulanus Ochi, Kon & Cabras 2023	<div><p>Onthophagus (Indachorius) leyteinsulanus Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Indachorius) leyteinsulanus Ochi, Kon &amp; Cabras, 2023a: 45 .</p><p>Type locality. “ Philippines, Leyte Is., Lake Imeruda. ”</p><p>Distribution in the Philippines. Visayas (Leyte) (Ochi et al. 2023a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692222825FF14FDDBFCDCFD29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FC83FCDCFC11.text	3C6E87D692222825FF14FC83FCDCFC11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Indachorius) lucenaensis Ochi, Kon & Cabras 2023	<div><p>Onthophagus (Indachorius) lucenaensis Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Indachorius) lucenaensis Ochi, Kon &amp; Cabras, 2023a: 40 .</p><p>Type locality. “ Philippines, Luzon Is., Lucena.”</p><p>Distribution in the Philippines. Luzon (Lucena) (Ochi et al. 2023a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692222825FF14FC83FCDCFC11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FC6BFCDCFB79.text	3C6E87D692222825FF14FC6BFCDCFB79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Indachorius) semiaereopictus Ochi, Kawahara & Cabras 2021	<div><p>Onthophagus (Indachorius) semiaereopictus Ochi, Kawahara &amp; Cabras, 2021</p><p>Onthophagus (Indachorius) semiaereopictus Ochi, Kawahara &amp; Cabras, 2021: 60; Ochi et al. 2023a: 48.</p><p>Type locality. “ Philippines Batan Is.”</p><p>Distribution in the Philippines. Luzon (Batan) (Ochi et al. 2021, 2023a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692222825FF14FC6BFCDCFB79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FA8BFCDCFA19.text	3C6E87D692222825FF14FA8BFCDCFA19.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Matashia) apomontanus Ochi, Kon & Cabras 2023	<div><p>Onthophagus (Matashia) apomontanus Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Matashia) apomontanus Ochi, Kon &amp; Cabras, 2023c: 654 .</p><p>Type locality. “ Philippines, Mindanao, Mt. Apo.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi et al. 2023c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692222825FF14FA8BFCDCFA19	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222825FF14FA73FCDCF895.text	3C6E87D692222825FF14FA73FCDCF895.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Matashia) galeatus Boucomont 1919	<div><p>Onthophagus (Matashia) galeatus Boucomont, 1919</p><p>Onthophagus galeatus Boucomont, 1919: 315; Boucomont 1921: 92; Boucomont 1924a: 671; Krajčík 2006: 106.</p><p>Onthophagus sus Boucomont, 1919: 317 . Type locality. “ Ile Mindano (Philippines): Dapitan.”; Krajčík 2006: 106.</p><p>Onthophagus (Onthophagus) galeatus: Balthasar 1963b: 364.</p><p>Type locality. “île de Basilan.”</p><p>Distribution in the Philippines. Mindanao (Davao del Norte, Lanao del Norte, Zamboanga del Norte, Surigao, Bukidnon, and Basilan) (Boucomont 1920, 1921, 1924a; Ochi et al. 2023c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692222825FF14FA73FCDCF895	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692222824FF14F8E7FCDDFF20.text	3C6E87D692222824FF14F8E7FCDDFF20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Matashia) semigaleatus Ochi, Kon & Cabras 2024	<div><p>Onthophagus (Matashia) semigaleatus Ochi, Kon &amp; Cabras, 2024</p><p>Onthophagus (Matashia) galeatus: Ochi, Kon &amp; Cabras 2023a: 655.</p><p>Onthophagus (Matashia) semigaleatus Ochi, Kon &amp; Cabras, 2024: 47 .</p><p>Type locality. “ Philippines, Mindanao Is., North Davao, Masara-Mainit.”</p><p>Distribution in the Philippines. Mindanao (Davao del Norte) (Ochi et al. 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692222824FF14F8E7FCDDFF20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FEBBFCDCFE08.text	3C6E87D692232824FF14FEBBFCDCFE08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Matashia) negrosensis Ochi, Kon & Cabras 2023	<div><p>Onthophagus (Matashia) negrosensis Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Matashia) negrosensis Ochi, Kon &amp; Cabras, 2023c: 647 .</p><p>Type locality. “ Philippines, Negros Is., Mt. Kanlaon. ”</p><p>Distribution in the Philippines. Visayas (Negros and Panay) (Ochi et al. 2023c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692232824FF14FEBBFCDCFE08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FE63FCDCFD70.text	3C6E87D692232824FF14FE63FCDCFD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Matashia) samarensis Ochi, Kon & Cabras 2023	<div><p>Onthophagus (Matashia) samarensis Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Matashia) samarensis Ochi, Kon &amp; Cabras, 2023c: 650 .</p><p>Type locality. “ Philippines, Samar Is.”</p><p>Distribution in the Philippines. Visayas (Leyte and Samar) (Ochi et al. 2023c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692232824FF14FE63FCDCFD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FD4BFCDCFC58.text	3C6E87D692232824FF14FD4BFCDCFC58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Matashia) semisibuyanus Ochi & Kon 2025	<div><p>Onthophagus (Matashia) semisibuyanus Ochi &amp; Kon, 2025</p><p>Onthophagus (Matashia) semisibuyanus Ochi &amp; Kon, 2025c: 54 .</p><p>Type locality. “ Philippines, Luzon Is., Queson, Gen Nakar.”</p><p>Distribution in the Philippines. Luzon (Quezon) (Ochi &amp; Kon 2025c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692232824FF14FD4BFCDCFC58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FC33FCDCFB78.text	3C6E87D692232824FF14FC33FCDCFB78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Matashia) sibuyanus Boucomont 1924	<div><p>Onthophagus (Matashia) sibuyanus Boucomont, 1924</p><p>Onthophagus sibuyanus Boucomont, 1924a: 676 .</p><p>Onthophagus (Onthophagus) sibuyanus: Balthasar 1963b: 525; Krajčík 2006: 132.</p><p>Onthophagus (Matashia) sibuyanus: Ochi &amp; Kon 2025c: 54.</p><p>Type locality. “Sibuyan.”</p><p>Distribution in the Philippines. Luzon (Sibuyan) (Boucomont 1924a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692232824FF14FC33FCDCFB78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FA8BFCDCFA18.text	3C6E87D692232824FF14FA8BFCDCFA18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Micronthophagus) bukidnonensis Ochi, Kon & Cabras 2023	<div><p>Onthophagus (Micronthophagus) bukidnonensis Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Micronthophagus) bukidnonensis Ochi, Kon &amp; Cabras, 2023b: 54 .</p><p>Type locality. “ Philippines, Mindanao Isl., Bukidnon.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon) (Ochi et al. 2023b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692232824FF14FA8BFCDCFA18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14FA73FCDCF938.text	3C6E87D692232824FF14FA73FCDCF938.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Micronthophagus) cuneus Boucomont 1924	<div><p>Onthophagus (Micronthophagus) cuneus Boucomont, 1924</p><p>Onthophagus cuneus Boucomont, 1924a: 673; Krajčík 2006: 99.</p><p>Onthophagus (Onthophagus) cuneus: Balthasar 1963b: 323.</p><p>Onthophagus (Micronthophagus) cuneus: Ochi et al. 2024: 56.</p><p>Type locality. “ Palawan ( Binaluan).”</p><p>Distribution in the Philippines. Luzon (Palawan) (Boucomont 1924a; Ochi et al. 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692232824FF14FA73FCDCF938	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692232824FF14F893FCDCF860.text	3C6E87D692232824FF14F893FCDCF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Micronthophagus) taoi Ochi & Kon 2006	<div><p>Onthophagus (Micronthophagus) taoi Ochi &amp; Kon, 2006</p><p>Onthophagus (Micronthophagus) taoi Ochi &amp; Kon, 2006: 47 .</p><p>Type locality. “ Queson National Park, Luzon Is., the Philippines.”</p><p>Distribution in the Philippines. Luzon (Quezon) (Ochi &amp; Kon 2006).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692232824FF14F893FCDCF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FF43FCDCFE51.text	3C6E87D692202827FF14FF43FCDCFE51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Onthophagiellus) calinanensis Ochi, Kon & Cabras 2025	<div><p>Onthophagus (Onthophagiellus) calinanensis Ochi, Kon &amp; Cabras, 2025</p><p>Onthophagus (Onthophagiellus) calinanensis Ochi, Kon &amp; Cabras, 2025b: 40 .</p><p>Type locality. “ Philippines, Mindanao, Davao.”</p><p>Distribution in the Philippines. Mindanao (Davao) (Ochi et al. 2025b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692202827FF14FF43FCDCFE51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FE2BFCDCFD15.text	3C6E87D692202827FF14FE2BFCDCFD15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Onthophagiellus) inermivertex Boucomont 1921	<div><p>Onthophagus (Onthophagiellus) inermivertex Boucomont, 1921</p><p>Onthophagus inermivertex Boucomont, 1921: 90; Boucomont 1924a: 671; Krajčík 2006: 111.</p><p>Onthophagus (Indachorius) inermivertex: Balthasar 1963b: 392.</p><p>Onthophagus (Onthophagiellus) inermivertex: Ochi et al. 2025b: 42.</p><p>Type locality. “ Iles Philippines, Mindanao: Iligan, Dapitan.”</p><p>Distribution in the Philippines. Mindanao (Lanao del Norte and Zamboanga del Norte) (Boucomont 1921, 1924a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692202827FF14FE2BFCDCFD15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FD67FCDCFC11.text	3C6E87D692202827FF14FD67FCDCFC11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Onthophagiellus) pseudocrassicollis Ochi, Kon & Cabras 2023	<div><p>Onthophagus (Onthophagiellus) pseudocrassicollis Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Onthophagiellus) pseudocrassicollis Ochi, Kon &amp; Cabras, 2023b: 57; Ochi et al. 2025b: 41.</p><p>Type locality. “ Philippines, Mindanao Is., Zamboanga.”</p><p>Distribution in the Philippines. Mindanao (Sarangani and Zamboanga) (Ochi et al. 2023b, 2025b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692202827FF14FD67FCDCFC11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FC6BFCDCFB79.text	3C6E87D692202827FF14FC6BFCDCFB79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Onthophagiellus) spinimetatibialis Ochi, Kon & Cabras 2025	<div><p>Onthophagus (Onthophagiellus) spinimetatibialis Ochi, Kon &amp; Cabras, 2025</p><p>Onthophagus (Onthophagiellus) spinimetatibialis Ochi, Kon &amp; Cabras, 2025b: 36 .</p><p>Type locality. “ Philippines, Luzon Is., Queson, Atimonan.”</p><p>Distribution in the Philippines. Luzon (Quezon) (Ochi et al. 2025b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692202827FF14FC6BFCDCFB79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14FA8BFE8DF9D1.text	3C6E87D692202827FF14FA8BFE8DF9D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Paraphanaeomorphus) argyropygus Gillet 1927	<div><p>Onthophagus (Paraphanaeomorphus) argyropygus Gillet, 1927</p><p>Onthophagus argyropygus Gillet, 1927: 256 .</p><p>Onthophagus (Paraphanaeomorphus) argyropygus: Ochi et al. 2024: 58.</p><p>Type locality. “ Hongkong.”</p><p>Distribution in the Philippines. Luzon (Mindoro) and Mindanao (Zamboanga) (Ochi et al. 2024).</p><p>Overall distribution. The Philippines, Japan, China (including Taiwan), Vietnam, and Laos (Ochi et al. 2024).</p></div>	https://treatment.plazi.org/id/3C6E87D692202827FF14FA8BFE8DF9D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692202827FF14F9E3FB64F8A9.text	3C6E87D692202827FF14F9E3FB64F8A9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Parentius) lilliputanus Lansberge 1883	<div><p>Onthophagus (Parentius) lilliputanus Lansberge, 1883</p><p>Onthophagus lilliputanus Lansberge, 1883: 69; Boucomont 1924a: 675.</p><p>Onthophagus (Onthophagus) lilliputanus: Balthasar 1963b: 420.</p><p>Onthophagus (Furconthophagus) lilliputanus: Ueda et al. 2022: 184.</p><p>Type locality. “ Java.”</p><p>Distribution in the Philippines. Luzon (Laguna) and Mindanao (Agusan del Norte) (Boucomont 1924a).</p><p>Overall distribution. The Philippines, Indonesia, Myanmar, and India (Ueda et al. 2022).</p></div>	https://treatment.plazi.org/id/3C6E87D692202827FF14F9E3FB64F8A9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FF43FCDCFE75.text	3C6E87D692212826FF14FF43FCDCFE75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Phanaeomorphus) bakeri Boucomont 1919	<div><p>Onthophagus (Phanaeomorphus) bakeri Boucomont, 1919</p><p>Onthophagus bakeri Boucomont, 1919: 312; Boucomont 1924a: 675; Krajčík 2006: 92.</p><p>Onthophagus (Phanaeomorphus) bakeri: Balthasar, 1963b: 285.</p><p>Type locality. “ Philippines: Mont Banahao, Los Baños.”</p><p>Distribution in the Philippines. Luzon (Laguna) (Boucomont 1920, 1924a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692212826FF14FF43FCDCFE75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FE47FCDCFD5D.text	3C6E87D692212826FF14FE47FCDCFD5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Phanaeomorphus) batanensis Ochi, Kawahara & Cabras 2021	<div><p>Onthophagus (Phanaeomorphus) batanensis Ochi, Kawahara &amp; Cabras, 2021</p><p>Onthophagus (Phanaeomorphus) batanensis Ochi, Kawahara &amp; Cabras, 2021: 59 .</p><p>Type locality. “ Philippines: Batan Is.”</p><p>Distribution in the Philippines. Luzon (Batan) (Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692212826FF14FE47FCDCFD5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FD2FFCDCFC85.text	3C6E87D692212826FF14FD2FFCDCFC85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Phanaeomorphus) mindanaoensis Frey 1971	<div><p>Onthophagus (Phanaeomorphus) mindanaoensis Frey, 1971</p><p>Onthophagus (Phanaeomorphus) mindanaoensis Frey, 1971: 95 .</p><p>Type locality. “ Philippinen, Bango, Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Frey 1971).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692212826FF14FD2FFCDCFC85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FC4FFB5AFB1D.text	3C6E87D692212826FF14FC4FFB5AFB1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Pseudophanaeomorphus) bangueyensis Boucomont 1914	<div><p>Onthophagus (Pseudophanaeomorphus) bangueyensis Boucomont, 1914</p><p>Onthophagus bangueyensis Boucomont, 1914: 323 .</p><p>Onthophagus (Onthophagus) bangueyensis: Balthasar 1963b: 286.</p><p>Onthophagus (Phanaeomorphus) bangueyensis: Ochi &amp; Kon 1994: 298.</p><p>Type locality. “Borneo: Ile Banguey.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Boucomont 1924; Balthasar 1963b).</p><p>Overall distribution. The Philippines and Malaysia (Boucomont 1914; Ochi &amp; Kon 1994).</p></div>	https://treatment.plazi.org/id/3C6E87D692212826FF14FC4FFB5AFB1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FB6FFCDCFA45.text	3C6E87D692212826FF14FB6FFCDCFA45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Pseudophanaeomorphus) palawantagal Ochi, Kon & Barclay 2016	<div><p>Onthophagus (Pseudophanaeomorphus) palawantagal Ochi, Kon &amp; Barclay, 2016</p><p>Onthophagus (Pseudophanaeomorphus) palawantagal Ochi, Kon &amp; Barclay, 2016: 431 .</p><p>Type locality. “ Philippines, Palawan, Mt. Salakot. ”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ochi et al. 2016).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692212826FF14FB6FFCDCFA45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14FA57FCDCF965.text	3C6E87D692212826FF14FA57FCDCF965.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Pseudophanaeomorphus) tagal Boucomont 1924	<div><p>Onthophagus (Pseudophanaeomorphus) tagal Boucomont, 1924</p><p>Onthophagus tagal Boucomont, 1924a: 672; Krajčík 2006: 135.</p><p>Onthophagus (Phanaeomorphus) tagal: Balthasar 1963b: 547; Ochi et al. 2016: 429.</p><p>Type locality. “ Sibuyan Island .”</p><p>Distribution in the Philippines. Luzon (Sibuyan, Quezon, and Baguio) (Boucomont 1924a; Balthasar 1963b; Ochi et al. 2016).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692212826FF14FA57FCDCF965	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692212826FF14F8AFFCDCF805.text	3C6E87D692212826FF14F8AFFCDCF805.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Sunenaga) mindanaensis Boucomont 1914	<div><p>Onthophagus (Sunenaga) mindanaensis Boucomont, 1914</p><p>Onthophagus wallacei mindanaensis Boucomont, 1914 .</p><p>Type locality. “ Philippines: Davao, Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Davao) (Boucomont 1914).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692212826FF14F8AFFCDCF805	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FF0BFBA4FE74.text	3C6E87D6923E2839FF14FF0BFBA4FE74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus (Sunenaga) wallacei Harold 1871	<div><p>Onthophagus (Sunenaga) wallacei Harold, 1871</p><p>Onthophagus wallacei Harold, 1871: 114; Boucomont 1924: 681.</p><p>Onthophagus simulans Sharp, 1875: 60 . Type locality. “Menado.”; Boucomont 1913: 418; Schultze 1916a: 160.</p><p>Type locality. “ Celebes.”</p><p>Distribution in the Philippines. Luzon (Mindoro), and Mindanao (Davao and Zamboanga del Norte) (Schultze 1916a; Boucomont 1924).</p><p>Overall distribution. The Philippines and Indonesia (Boucomont 1924).</p></div>	https://treatment.plazi.org/id/3C6E87D6923E2839FF14FF0BFBA4FE74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FDBFFAB4FC84.text	3C6E87D6923E2839FF14FDBFFAB4FC84.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus babirussa (Eschscholtz 1822)	<div><p>Onthophagus (sensu lato) babirussa (Eschscholtz, 1822)</p><p>Copris babirussa Eschscholtz, 1822: 31 .</p><p>Onthophagus babirussa: Boucomont 1924a: 67; Krajčík 2006: 91.</p><p>Onthophagus (Onthophagus) babirussa: Balthasar 1963b: 283.</p><p>Type locality. “ Insel Luçon (Manilla).”</p><p>Distribution in the Philippines. Luzon (Baguio, Cagayan, Manila, and Rizal), Visayas (Dumaguete), and Mindanao (Basilan, Lanao del Norte, and Surigao) (Eschscholtz 1822; Boucomont 1924a; Schultze 1916a).</p><p>Overall distribution. The Philippines, Indonesia, Malaysia, and Singapore (Balthasar 1963b).</p></div>	https://treatment.plazi.org/id/3C6E87D6923E2839FF14FDBFFAB4FC84	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FC17FC27FBEC.text	3C6E87D6923E2839FF14FC17FC27FBEC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus bicolensis subsp. asin Ochi & Kon 2006	<div><p>Onthophagus (sensu lato) bicolensis asin Ochi &amp; Kon, 2006</p><p>Onthophagus (Onthophagus) bicolensis asin Ochi &amp; Kon, 2006: 45 .</p><p>Type locality. “ Asin hot spring near Baguio, the Philippines.”</p><p>Distribution in the Philippines. Luzon (Baguio) (Ochi &amp; Kon 2006).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923E2839FF14FC17FC27FBEC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FBFFFC27FAD4.text	3C6E87D6923E2839FF14FBFFFC27FAD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus bicolensis subsp. bicolensis Ochi & Kon 2006	<div><p>Onthophagus (sensu lato) bicolensis bicolensis Ochi &amp; Kon, 2006</p><p>Onthophagus (Onthophagus) bicolensis bicolensis Ochi &amp; Kon, 2006: 46 .</p><p>Type locality. “ Bicol National Park, Luzon, the Philippines.”</p><p>Distribution in the Philippines. Luzon (Bicol) (Ochi &amp; Kon 2006).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923E2839FF14FBFFFC27FAD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14FAA7FCDCFA3C.text	3C6E87D6923E2839FF14FAA7FCDCFA3C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus bistiniocelloides Krikken 1986	<div><p>Onthophagus (sensu lato) bistiniocelloides Krikken, 1986</p><p>Onthophagus bistiniocelloides Krikken, 1986: 281; Krajčík 2006: 93.</p><p>Type locality. “ Philippines: Samar.”</p><p>Distribution in the Philippines. Visayas (Samar) (Krikken 1986).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923E2839FF14FAA7FCDCFA3C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14F98FFCDCF964.text	3C6E87D6923E2839FF14F98FFCDCF964.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus cabrasae Ochi & Kon 2025	<div><p>Onthophagus (sensu lato) cabrasae Ochi &amp; Kon, 2025</p><p>Onthophagus (incertae sedis) cabrasae Ochi &amp; Kon, 2025c: 51 .</p><p>Type locality. “ Philippines, Mindanao Is., South Lanao. ”</p><p>Distribution in the Philippines. Mindanao (Lanao del Sur) (Ochi &amp; Kon 2025c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923E2839FF14F98FFCDCF964	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923E2839FF14F977FCDCF860.text	3C6E87D6923E2839FF14F977FCDCF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus calcaratus Boucomont 1919	<div><p>Onthophagus (sensu lato) calcaratus Boucomont, 1919</p><p>Onthophagus calcaratus Boucomont, 1919: 316; Boucomont 1921: 92; Boucomont 1924a: 674.</p><p>Onthophagus (Onthophagus) calcaratus: Balthasar 1963b: 301; Krajčík 2006: 95.</p><p>Type locality. “ Ile Mindanao (Philippines): Dapitan.”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Boucomont 1920, 1921, 1924a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923E2839FF14F977FCDCF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FF0BFCDCFEBD.text	3C6E87D6923F2838FF14FF0BFCDCFEBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus citreum Boucomont 1919	<div><p>Onthophagus (sensu lato) citreum Boucomont, 1919</p><p>Onthophagus citreum Boucomont, 1919: 312; Boucomont 1924a: 673.</p><p>Onthophagus (Onthophagus) citreum: Balthasar 1963b: 311; Krajčík 2006: 96.</p><p>Type locality. “ Iles Philippines ( Luzon): Mont Makiling, Mt. Banahao. ”</p><p>Distribution in the Philippines. Luzon (Laguna) and Visayas (Negros Oriental) (Boucomont 1920, 1924a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923F2838FF14FF0BFCDCFEBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FE0FFCDCFDB9.text	3C6E87D6923F2838FF14FE0FFCDCFDB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus dapitanensis Boucomont 1919	<div><p>Onthophagus (sensu lato) dapitanensis Boucomont, 1919</p><p>Onthophagus dapitanensis Boucomont, 1919: 318; Boucomont 1924a: 670.</p><p>Onthophagus (Onthophagus) dapitanensis: Balthasar 1963b: 327; Krajčík 2006: 99.</p><p>Type locality. “ Ile Mindanao (Philippines): Dapitan.”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Boucomont 1920, 1924a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923F2838FF14FE0FFCDCFDB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FD13FCDCFCE1.text	3C6E87D6923F2838FF14FD13FCDCFCE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus davaoimbutus Ochi, Kon & Cabras 2024	<div><p>Onthophagus (sensu lato) davaoimbutus Ochi, Kon &amp; Cabras, 2024</p><p>Onthophagus (Onthophagus) davaoimbutus Ochi, Kon &amp; Cabras, 2024: 49 .</p><p>Type locality. “ Philippines, Mindanao Is., Davao del Norte, Masara-Mainit.”</p><p>Distribution in the Philippines. Mindanao (Davao del Norte) (Ochi et al. 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923F2838FF14FD13FCDCFCE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FCFBFCDCFBC9.text	3C6E87D6923F2838FF14FCFBFCDCFBC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus dingalanensis Ochi, Kon & Cabras 2023	<div><p>Onthophagus (sensu lato) dingalanensis Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Onthophagus) dingalanensis Ochi, Kon &amp; Cabras, 2023b: 61 .</p><p>Type locality. “ Philippines, Luzon Is., Dingalan.”</p><p>Distribution in the Philippines. Luzon (Ochi et al. 2023b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923F2838FF14FCFBFCDCFBC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FBA3FCDCFB31.text	3C6E87D6923F2838FF14FBA3FCDCFB31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus falsicyobius Ochi, Kon & Cabras 2023	<div><p>Onthophagus (sensu lato) falsicyobius Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Onthophagus) falsicyobius Ochi, Kon &amp; Cabras, 2023b: 60 .</p><p>Type locality. “ Philippines, Mindanao Is., Lanao del Sur.”</p><p>Distribution in the Philippines. Mindanao (Lanao del Sur) (Ochi et al. 2023b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923F2838FF14FBA3FCDCFB31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14FA8BFB14F9D1.text	3C6E87D6923F2838FF14FA8BFB14F9D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus gestroi Harold 1877	<div><p>Onthophagus (sensu lato) gestroi Harold, 1877</p><p>Onthophagus Gestroi Harold, 1877: 59; Boucomont 1924a: 673.</p><p>Onthophagus (Onthophagus) gestroi: Balthasar 1963b: 367.</p><p>Type locality. “ Sumatra.”</p><p>Distribution in the Philippines. Luzon (Laguna, Sibuyan, and Palawan) and Mindanao (Surigao) (Harold 1877; Boucomont 1924a; Balthasar 1963b).</p><p>Overall distribution. The Philippines and Indonesia (Harold 1877; Balthasar 1963b).</p></div>	https://treatment.plazi.org/id/3C6E87D6923F2838FF14FA8BFB14F9D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14F9ABFCDCF939.text	3C6E87D6923F2838FF14F9ABFCDCF939.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus hielkemai Krikken & Huijbregts 2011	<div><p>Onthophagus (sensu lato) hielkemai Krikken &amp; Huijbregts, 2011</p><p>Onthophagus hielkemai Krikken &amp; Huijbregts, 2011: 209 .</p><p>Type locality. “ Philippines: Bohol: Sierra Bullones: Nan-od.”</p><p>Distribution in the Philippines. Visayas (Bohol) (Krikken &amp; Huijbregts 2011).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923F2838FF14F9ABFCDCF939	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923F2838FF14F893FB57F861.text	3C6E87D6923F2838FF14F893FB57F861.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus hirsutulus Lansberge 1883	<div><p>Onthophagus (sensu lato) hirsutulus Lansberge, 1883</p><p>Onthophagus hirsutulus Lansberge, 1883: 66; Boucomont 1924a: 672.</p><p>Type locality. “ Java.”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Boucomont 1924a).</p><p>Overall distribution. The Philippines and Indonesia (Lansberge 1883; Boucomont 1924a).</p></div>	https://treatment.plazi.org/id/3C6E87D6923F2838FF14F893FB57F861	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FF0BFCDCFE99.text	3C6E87D6923C283BFF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus ifugaoensis Ochi & Kon 2006	<div><p>Onthophagus (sensu lato) ifugaoensis Ochi &amp; Kon, 2006</p><p>Onthophagus (Onthophagus) ifugaoensis Ochi &amp; Kon, 2006: 41 .</p><p>Type locality. “ Banaue, Ifugao, Luzon, the Philippines.”</p><p>Distribution in the Philippines. Luzon (Ifugao) (Ochi &amp; Kon 2006).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923C283BFF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FEF3FCDCFDE5.text	3C6E87D6923C283BFF14FEF3FCDCFDE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus lunulifer Boucomont 1914	<div><p>Onthophagus (sensu lato) lunulifer Boucomont, 1914</p><p>Onthophagus lunulifer Boucomont, 1914: 309; Boucomont 1924a: 677; Krikken &amp; Huijbregts 2011: 206.</p><p>Onthophagus (Onthophagus) lunulifer: Balthasar 1963b: 423; Krajčík 2006: 116.</p><p>Type locality. “ Philippines: Los Banos.”</p><p>Distribution in the Philippines. Luzon (Laguna) (Boucomont 1914, 1924a; Krikken &amp; Huijbregts 2011).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923C283BFF14FEF3FCDCFDE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FDF7FCDCFCCD.text	3C6E87D6923C283BFF14FDF7FCDCFCCD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus luzonwallacei Ochi, Kawahara & Cabras 2021	<div><p>Onthophagus (sensu lato) luzonwallacei Ochi, Kawahara &amp; Cabras, 2021</p><p>Onthophagus (incertae sedis) luzonwallacei Ochi, Kawahara &amp; Cabras, 2021: 64 .</p><p>Type locality. “ Philippines, Luzon Is. nr. Lucena, Kinabuhayan.”</p><p>Distribution in the Philippines. Luzon (Quezon) (Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923C283BFF14FDF7FCDCFCCD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FCDFFCDCFC35.text	3C6E87D6923C283BFF14FCDFFCDCFC35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus manbucalensis Ochi, Kon & Cabras 2024	<div><p>Onthophagus (sensu lato) manbucalensis Ochi, Kon &amp; Cabras, 2024</p><p>Onthophagus (incertae sedis) manbucalensis Ochi, Kon &amp; Cabras, 2024: 55 .</p><p>Type locality. “ Philippines, Negros Is., Manbucal. ”</p><p>Distribution in the Philippines. Visayas (Negros) (Ochi et al. 2024).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923C283BFF14FCDFFCDCFC35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FB87FCDCFB1D.text	3C6E87D6923C283BFF14FB87FCDCFB1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus matsudai Ochi & Kon 2004	<div><p>Onthophagus (sensu lato) matsudai Ochi &amp; Kon, 2004</p><p>Onthophagus (Onthophagus) matsudai Ochi &amp; Kon, 2004: 31; Krajčík 2006: 117.</p><p>Type locality. “ Mt. Apo, Mindanao Island.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi &amp; Kon 2004).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923C283BFF14FB87FCDCFB1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FB6FFCDCFA45.text	3C6E87D6923C283BFF14FB6FFCDCFA45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus ohkuboi Ochi & Kon 2006	<div><p>Onthophagus (sensu lato) ohkuboi Ochi &amp; Kon, 2006</p><p>Onthophagus (Onthophagus) ohkuboi Ochi &amp; Kon, 2006: 42 .</p><p>Type locality. “ Banaue, Ifugao, Luzon, the Philippines.”</p><p>Distribution in the Philippines. Luzon (Ifugao) (Ochi &amp; Kon 2006).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923C283BFF14FB6FFCDCFA45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14FA57FC25F965.text	3C6E87D6923C283BFF14FA57FC25F965.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus orientalis Harold 1868	<div><p>Onthophagus (sensu lato) orientalis Harold, 1868</p><p>Onthophagus orientalis Harold, 1868: 83; Boucomont 1924a: 673.</p><p>Onthophagus (Onthophagus) orientalis: Balthasar 1963b: 463.</p><p>Type locality. “ Hongkong, Bengal.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Boucomont 1924a; Balthasar 1963b).</p><p>Overall distribution. The Philippines, Cambodia, China (including Taiwan), India, Indonesia, Malaysia, Myanmar, Nepal, Thailand, and Vietnam (Balthasar 1963b).</p></div>	https://treatment.plazi.org/id/3C6E87D6923C283BFF14FA57FC25F965	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923C283BFF14F977FB09F861.text	3C6E87D6923C283BFF14F977FB09F861.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus palavanus Balthasar 1959	<div><p>Onthophagus (sensu lato) palavanus Balthasar, 1959</p><p>Onthophagus javanensis palavanus Balthasar, 1959: 193 .</p><p>Onthophagus palavanus: Huijbregts &amp; Krikken 2011: 49.</p><p>Type locality. “ Palawan, Binaluan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Balthasar 1959).</p><p>Overall distribution. Endemic species to the Philippines (Huijbregts &amp; Krikken 2011).</p></div>	https://treatment.plazi.org/id/3C6E87D6923C283BFF14F977FB09F861	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FF0BFCDCFE99.text	3C6E87D6923D283AFF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus palawanfalsicyobius Ochi & Kon 2025	<div><p>Onthophagus (sensu lato) palawanfalsicyobius Ochi &amp; Kon, 2025</p><p>Onthophagus (incertae sedis) palawanfalsicyobius Ochi &amp; Kon, 2025c: 48 .</p><p>Type locality. “ Palawan Island.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ochi &amp; Kon 2025c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923D283AFF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FEF0FCDCFD86.text	3C6E87D6923D283AFF14FEF0FCDCFD86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus philippinensis Boucomont 1919	<div><p>Onthophagus (sensu lato) philippinensis Boucomont, 1919</p><p>Onthophagus philippinensis Boucomont, 1919: 313; Boucomont 1924: 676.</p><p>Onthophagus (Onthophagus) philippinensis: Balthasar 1963b: 478; Krajčík 2006: 124.</p><p>Type locality. “ Philippines: Mont. Banahao.”</p><p>Distribution in the Philippines. This species is endemic to the archipelago and is found in Luzon (Laguna and Sibuyan) and Mindanao (Lanao del Norte) (Boucomont 1920, 1924a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923D283AFF14FEF0FCDCFD86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FD11FCDCFCEF.text	3C6E87D6923D283AFF14FD11FCDCFCEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus quirinoensis Ochi, Kon & Cabras 2023	<div><p>Onthophagus (sensu lato) quirinoensis Ochi, Kon &amp; Cabras, 2023</p><p>Onthophagus (Onthophagus) quirinoensis Ochi, Kon &amp; Cabras, 2023b: 59 .</p><p>Type locality. “ Philippines, Luzon Is., Quirino.”</p><p>Distribution in the Philippines. Luzon (Quirino) (Ochi et al. 2023b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923D283AFF14FD11FCDCFCEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FCFEFCDCFBD4.text	3C6E87D6923D283AFF14FCFEFCDCFBD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus rizali Ochi, Kawahara & Cabras 2021	<div><p>Onthophagus (sensu lato) rizali Ochi, Kawahara &amp; Cabras, 2021</p><p>Onthophagus (incertae sedis) rizali Ochi, Kawahara &amp; Cabras, 2021: 62 .</p><p>Type locality. “ Philippines: Batan Is.”</p><p>Distribution in the Philippines. Luzon (Batan) (Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923D283AFF14FCFEFCDCFBD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FBA6FCDCFAD1.text	3C6E87D6923D283AFF14FBA6FCDCFAD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus sceptrifer Boucomont 1924	<div><p>Onthophagus (sensu lato) sceptrifer Boucomont, 1924</p><p>Onthophagus sceptrifer Boucomont, 1924a: 677; Krikken &amp; Huijbregts 2011: 206.</p><p>Onthophagus (Onthophagus) sceptrifer: Balthasar 1963b: 513; Krajčík 2006: 131.</p><p>Type locality. “ Palawan, Bacuit.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Boucomont 1924a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923D283AFF14FBA6FCDCFAD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14FAABFCDCF9DA.text	3C6E87D6923D283AFF14FAABFCDCF9DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus sexdentatus Boucomont 1919	<div><p>Onthophagus (sensu lato) sexdentatus Boucomont, 1919</p><p>Onthophagus sexdentatus Boucomont, 1919: 321; Boucomont 1924a: 674.</p><p>Onthophagus (Onthophagus) sexdentatus: Balthasar 1963b: 524; Krajčík 2006: 132.</p><p>Type locality. “ Philippines: Mont Banahao.”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao) (Boucomont 1920; 1924a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923D283AFF14FAABFCDCF9DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14F9ACFCDCF8FB.text	3C6E87D6923D283AFF14F9ACFCDCF8FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus surdus Boucomont 1925	<div><p>Onthophagus (sensu lato) surdus Boucomont, 1925</p><p>Onthophagus surdus Boucomont, 1925: 153 .</p><p>Onthophagus (Onthophagus) surdus: Balthasar 1963b: 542; Krajčík 2006: 135.</p><p>Type locality. “Mindanao: Momungan, Pt. Bango. ”</p><p>Distribution in the Philippines. Luzon (Camarines Sur) and Mindanao (Lanao del Norte) (Boucomont 1925).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923D283AFF14F9ACFCDCF8FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923D283AFF14F8CDFCDCF823.text	3C6E87D6923D283AFF14F8CDFCDCF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus surigaoensis Ochi & Kon 2025	<div><p>Onthophagus (sensu lato) surigaoensis Ochi &amp; Kon, 2025</p><p>Onthophagus (incertae sedis) surigaoensis Ochi &amp; Kon, 2025c: 53 .</p><p>Type locality. “ Philippines, Mindanao Is., South Surigao. ”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Ochi &amp; Kon 2025c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923D283AFF14F8CDFCDCF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FF0BFB9BFE74.text	3C6E87D6923A283DFF14FF0BFB9BFE74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus terminatus (Eschscholtz 1822)	<div><p>Onthophagus (sensu lato) terminatus (Eschscholtz, 1822)</p><p>Copris terminata Eschscholtz, 1822: 33 .</p><p>Onthophagus terminatus: Boucomont 1914: 318; Boucomont 1924a: 676.</p><p>Onthophagus (Onthophagus) terminatus: Balthasar 1963b: 551; Nomura 1973: 49; Krajčík 2006: 136.</p><p>Type locality. “ Insel Luçon (Manilla).”</p><p>Distribution in the Philippines. Luzon (Manila and Mindoro) (Eschscholtz 1822; Boucomont 1914; 1924a).</p><p>Overall distribution. The Philippines and China (Taiwan) (Krajčík 2006).</p></div>	https://treatment.plazi.org/id/3C6E87D6923A283DFF14FF0BFB9BFE74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FD4BFC27FC58.text	3C6E87D6923A283DFF14FD4BFC27FC58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus tiniocelloides subsp. mamasensis Ochi, Kon & Cabras 2024	<div><p>Onthophagus (sensu lato) tiniocelloides mamasensis Ochi, Kon &amp; Cabras, 2024</p><p>Onthophagus (incertae sedis) tiniocelloides mamasensis Ochi, Kon &amp; Cabras, 2024: 54.</p><p>Type locality. “ Philippines, Mindanao Is., Davao del Norte, Masara-Mainit.”</p><p>Distribution in the Philippines. Mindanao (Davao del Norte) (Ochi et al. 2024).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923A283DFF14FD4BFC27FC58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FE47FC27FD70.text	3C6E87D6923A283DFF14FE47FC27FD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus tiniocelloides subsp. tiniocelloides Boucomont 1925	<div><p>Onthophagus (sensu lato) tiniocelloides tiniocelloides Boucomont, 1925</p><p>Onthophagus tiniocelloides Boucomont, 1925: 152; Krikken, 1986: 282.</p><p>Onthophagus (Onthophagus) tiniocelloides: Balthasar 1963b: 554; Krajčík 2006: 137.</p><p>Type locality. “Mindanao: Momungan.”</p><p>Distribution in the Philippines. Mindanao (Lanao del Norte) (Boucomont 1925).</p><p>Overall distribution. Endemic subspecies to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923A283DFF14FE47FC27FD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FC33FCDCFB80.text	3C6E87D6923A283DFF14FC33FCDCFB80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus traginus Frey 1972	<div><p>Onthophagus (sensu lato) traginus Frey, 1972</p><p>Onthophagus traginus Frey, 1972a: 268; Krajčík 2006: 137.</p><p>Type locality. “Mindanao, Lanao, Kolobugano.”</p><p>Distribution in the Philippines. Mindanao (Lanao del Norte) (Frey 1972a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923A283DFF14FC33FCDCFB80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FB1BFCDCFAA0.text	3C6E87D6923A283DFF14FB1BFCDCFAA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus unidentatus Boucomont 1927	<div><p>Onthophagus (sensu lato) unidentatus Boucomont, 1927</p><p>Onthophagus unidens Boucomont, 1914: 308; Boucomont 1924a: 677.</p><p>Onthophagus (Onthophagus) unidentatus: Balthasar 1963b: 570; Krajčík 2006: 139;</p><p>Onthophagus unidentatus: Krikken &amp; Huijbregts 2011: 212 (redescription).</p><p>Type locality. “ Philippines, Los Baños.”</p><p>Distribution in the Philippines. Luzon (Laguna) (Boucomont 1914, 1924a; Krikken &amp; Huijbregts 2011).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923A283DFF14FB1BFCDCFAA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14FA3BFCDCF988.text	3C6E87D6923A283DFF14FA3BFCDCF988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus wiebesi Krikken 1977	<div><p>Onthophagus (sensu lato) wiebesi Krikken, 1977</p><p>Onthophagus wiebesi Krikken, 1977a: 177; Krajčík 2006: 142.</p><p>Type locality. “ Philippines: Luzon: Los Baños: Laguna.”</p><p>Distribution in the Philippines. Luzon (Laguna) (Krikken 1977a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923A283DFF14FA3BFCDCF988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923A283DFF14F9E3FCDCF84C.text	3C6E87D6923A283DFF14F9E3FCDCF84C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onthophagus zebu Boucomont 1921	<div><p>Onthophagus (sensu lato) zebu Boucomont, 1921</p><p>Onthophagus zebu Boucomont, 1921: 89; Boucomont 1924a: 677.</p><p>Onthophagus (Onthophagus) zebu: Balthasar, 1963b: 594; Krajčík 2006: 142;</p><p>Onthophagus zebu: Krikken &amp; Huijbregts 2011: 212.</p><p>Type locality. “ Iles Philippines: Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Bukidnon, Lanao del Norte, and Zamboanga del Norte) (Boucomont 1921, 1924a; Krikken &amp; Huijbregts 2011).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923A283DFF14F9E3FCDCF84C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FEBBFCDCFE16.text	3C6E87D6923B283CFF14FEBBFCDCFE16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) basilanensis Ochi, Kon & Cabras 2024	<div><p>Parascatonomus (Granulidorsum) basilanensis Ochi, Kon &amp; Cabras, 2024</p><p>Parascatonomus (Granulidorsum) basilanensis Ochi, Kon &amp; Cabras, 2024: 58; Ochi &amp; Kon 2025c: 56.</p><p>Type locality. “ Philippines, Basilan Is., Mindanao.”</p><p>Distribution in the Philippines. Mindanao (Basilan and Bukidnon) (Ochi et al. 2024; Ochi &amp; Kon 2025c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923B283CFF14FEBBFCDCFE16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FE63FCDCFD37.text	3C6E87D6923B283CFF14FE63FCDCFD37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) dejongi (Krikken 1986)	<div><p>Parascatonomus (Granulidorsum) dejongi (Krikken, 1986)</p><p>Onthophagus dejongi Krikken, 1986: 279; Krajčík 2006: 100.</p><p>Onthophagus (Parascatonomus) dejongi: Ochi &amp; Araya 1992: 99.</p><p>Type locality. “ Philippines (Samar).”</p><p>Distribution in the Philippines. Visayas (Leyte and Samar) and Mindanao (Surigao) (Krikken 1986; Ochi &amp; Araya 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923B283CFF14FE63FCDCFD37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FC80FCDCFC1F.text	3C6E87D6923B283CFF14FC80FCDCFC1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) hayashimasaoi (Ochi & Kon 1999)	<div><p>Parascatonomus (Granulidorsum) hayashimasaoi (Ochi &amp; Kon, 1999)</p><p>Onthophagus (Parascatonomus) hayashimasaoi Ochi &amp; Kon, 1999: 137; Krajčík 2006: 108.</p><p>Type locality. “ Mt. Apo, Mindanao Is., Philippines.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi &amp; Kon 1999).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923B283CFF14FC80FCDCFC1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FC69FCDCFB44.text	3C6E87D6923B283CFF14FC69FCDCFB44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) koichii (Ochi & Araya 1992)	<div><p>Parascatonomus (Granulidorsum) koichii (Ochi &amp; Araya, 1992)</p><p>Onthophagus (Parascatonomus) koichii Ochi &amp; Araya, 1992: 94; Krajčík 2006: 112.</p><p>Type locality. “ Philippines, Mindanao Is., Mt. Apo.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi &amp; Araya 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923B283CFF14FC69FCDCFB44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FB56FCDCFAAD.text	3C6E87D6923B283CFF14FB56FCDCFAAD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) leytensis (Ochi & Araya 1992)	<div><p>Parascatonomus (Granulidorsum) leytensis (Ochi &amp; Araya, 1992)</p><p>Onthophagus (Parascatonomus) leytensis Ochi &amp; Araya, 1992: 103; Krajčík 2006: 115.</p><p>Type locality. “ Philippines, Leyte.”</p><p>Distribution in the Philippines. Visayas (Leyte) (Ochi &amp; Araya 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923B283CFF14FB56FCDCFAAD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14FA3FFCDCF992.text	3C6E87D6923B283CFF14FA3FFCDCF992.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) negrosensis (Ochi & Araya 1992)	<div><p>Parascatonomus (Granulidorsum) negrosensis (Ochi &amp; Araya, 1992)</p><p>Onthophagus (Parascatonomus) negrosensis Ochi &amp; Araya, 1992: 100; Krajčík 2006: 120.</p><p>Type locality. “ Philippines, Negros Isld., Mambucal. ”</p><p>Distribution in the Philippines. Visayas (Negros) (Ochi &amp; Araya 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923B283CFF14FA3FFCDCF992	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14F9E4FCDCF8FB.text	3C6E87D6923B283CFF14F9E4FCDCF8FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) pararudis Ochi, Kawahara & Cabras 2021	<div><p>Parascatonomus (Granulidorsum) pararudis Ochi, Kawahara &amp; Cabras, 2021</p><p>Parascatonomus (Granulidorsum) pararudis Ochi, Kawahara &amp; Cabras, 2021: 65 .</p><p>Type locality. “ Philippines, Batan Is.”</p><p>Distribution in the Philippines. Luzon (Batan) (Ochi et al. 2021).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923B283CFF14F9E4FCDCF8FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D6923B283CFF14F8CCFCDCF823.text	3C6E87D6923B283CFF14F8CCFCDCF823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) pseudorudis (Ochi & Araya 1992)	<div><p>Parascatonomus (Granulidorsum) pseudorudis (Ochi &amp; Araya, 1992)</p><p>Onthophagus (Parascatonomus) pseudorudis Ochi &amp; Araya, 1992: 94; Krajčík 2006: 126.</p><p>Type locality. “ Philippines, Luzon Is., Quezon National Park.”</p><p>Distribution in the Philippines. Luzon (Quezon) and Visayas (Negros) (Ochi &amp; Araya 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D6923B283CFF14F8CCFCDCF823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FF0AFCDCFE99.text	3C6E87D69238283FFF14FF0AFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) quezonensis (Ochi & Araya 1992)	<div><p>Parascatonomus (Granulidorsum) quezonensis (Ochi &amp; Araya, 1992)</p><p>Onthophagus (Parascatonomus) quezonensis Ochi &amp; Araya, 1992: 97; Krajčík 2006: 128.</p><p>Type locality. “ Philippines, Luzon Isld., Quezon National Park.”</p><p>Distribution in the Philippines. Luzon (Quezon) (Ochi &amp; Araya 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69238283FFF14FF0AFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FEF3FE7BFD36.text	3C6E87D69238283FFF14FEF3FE7BFD36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) rudis (Sharp 1875)	<div><p>Parascatonomus (Granulidorsum) rudis (Sharp, 1875)</p><p>Onthophagus rudis Sharp, 1875: 58; Boucomont 1913: 418; Boucomont 1924a: 669; Boucomont 1925: 153.</p><p>Onthophagus (Onthophagus) rudis: Balthasar 1963b: 505.</p><p>Onthophagus (Parascatonomus) rudis: Ochi &amp; Kon 1994: 294; Ochi et al. 2008: 201.</p><p>Parascatonomus (Granulidorsum) rudis: Ueda et al. 2022: 181.</p><p>Type locality. “ Siam.”</p><p>Distribution in the Philippines. Luzon (Laguna and Palawan) and Mindanao (Davao, Lanao del Norte, Surigao, and Zamboanga del Norte) (Boucomont 1914, 1924 a, 1925).</p><p>Overall distribution. The Philippines, China, India, Indonesia, Malaysia, Myanmar, Thailand, and Vietnam (Ueda et al. 2022).</p></div>	https://treatment.plazi.org/id/3C6E87D69238283FFF14FEF3FE7BFD36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FC80FCDCFC1F.text	3C6E87D69238283FFF14FC80FCDCFC1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) shizumui (Ochi & Araya 1992)	<div><p>Parascatonomus (Granulidorsum) shizumui (Ochi &amp; Araya, 1992)</p><p>Onthophagus (Parascatonomus) shizumui Ochi &amp; Araya, 1992: 104; Krajčík 2006: 132.</p><p>Type locality. “ Philippines, Mindanao Isld., Mt. Apo.”</p><p>Distribution in the Philippines. Mindanao (Mt. Apo) (Ochi &amp; Araya 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69238283FFF14FC80FCDCFC1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FC69FCDCFB18.text	3C6E87D69238283FFF14FC69FCDCFB18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) takedai (Ochi & Araya 1992)	<div><p>Parascatonomus (Granulidorsum) takedai (Ochi &amp; Araya, 1992)</p><p>Onthophagus (Parascatonomus) takedai Ochi &amp; Araya, 1992: 99; Krajčík 2006: 135.</p><p>Type locality. “ Philippines (Quezon).”</p><p>Distribution in the Philippines. Luzon (Quezon and Camarines Sur) and Visayas (Negros) (Ochi &amp; Araya 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69238283FFF14FC69FCDCFB18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FB72FCDCFA41.text	3C6E87D69238283FFF14FB72FCDCFA41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) takeshii (Ochi & Araya 1992)	<div><p>Parascatonomus (Granulidorsum) takeshii (Ochi &amp; Araya, 1992)</p><p>Onthophagus (Parascatonomus) takeshii Ochi &amp; Araya, 1992: 102; Krajčík 2006: 135.</p><p>Type locality. “ Philippines, Luzon Is., Mt. Banahao. ”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao and Camarines Sur) (Ochi &amp; Araya 1992).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69238283FFF14FB72FCDCFA41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14FA5BFCDCF96E.text	3C6E87D69238283FFF14FA5BFCDCF96E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Granulidorsum) yvescambeforti (Ochi & Araya 1992)	<div><p>Parascatonomus (Granulidorsum) yvescambeforti (Ochi &amp; Araya, 1992)</p><p>Onthophagus (Parascatonomus) yvescambeforti Ochi &amp; Araya, 1992: 96; Krajčík 2006: 142.</p><p>Parascatonomus (Granulidorsum) yvescambeforti: Ochi &amp; Kon 2025c: 56.</p><p>Type locality. “ Philippines, Mindanao Is., Surigao.”</p><p>Distribution in the Philippines. Mindanao (Surigao and Bukidnon) (Ochi &amp; Araya 1992; Ochi &amp; Kon 2025c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D69238283FFF14FA5BFCDCF96E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69238283FFF14F8B0FB31F823.text	3C6E87D69238283FFF14F8B0FB31F823.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Necramator) katoi (Ochi & Araya 1996)	<div><p>Parascatonomus (Necramator) katoi (Ochi &amp; Araya, 1996)</p><p>Onthophagus (Parascatonomus) katoi Ochi &amp; Araya, 1996: 13; Krajčík 2006: 111; Krikken &amp; Huijbregts 2009: 54.</p><p>Type locality. “ Luzon, Bicol National Park.”</p><p>Distribution in the Philippines. Luzon (Palawan and Bicol) (Ochi &amp; Araya 1996).</p><p>Overall distribution. The Philippines and Malaysia (Krikken &amp; Huijbregts 2009).</p></div>	https://treatment.plazi.org/id/3C6E87D69238283FFF14F8B0FB31F823	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69239283EFF14FF0AFE8DFE74.text	3C6E87D69239283EFF14FF0AFE8DFE74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Necramator) pilularius (Lansberge 1883)	<div><p>Parascatonomus (Necramator) pilularius (Lansberge, 1883)</p><p>Onthophagus pilularius Lansberge, 1883: 70; Boucomont 1914: 277; Boucomont 1924a: 670.</p><p>Onthophagus (Onthophagus) pilularius: Balthasar 1963b: 480.</p><p>Onthophagus (Parascatonomus) pilularius: Krikken &amp; Huijbregts 2009: 48.</p><p>Type locality. “ Java.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Boucomont 1914, 1924a; Balthasar 1963b).</p><p>Overall distribution. The Philippines, Indonesia, China (Taiwan), and Thailand (Krikken &amp; Huijbregts 2009).</p></div>	https://treatment.plazi.org/id/3C6E87D69239283EFF14FF0AFE8DFE74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69239283EFF14FE46FAB9FD28.text	3C6E87D69239283EFF14FE46FAB9FD28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Necramator) semiaureus (Lansberge 1883)	<div><p>Parascatonomus (Necramator) semiaureus (Lansberge, 1883)</p><p>Onthophagus semiaureus Lansberge, 1883: 75; Boucomont 1924a: 670.</p><p>Onthophagus (Onthophagus) semiaureus: Balthasar 1963b: 517.</p><p>Onthophagus (Parascatonomus) semiaureus: Ochi &amp; Kon 1994: 294; Ochi et al. 2008: 208.</p><p>Type locality. “ Java et Sumatra.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Boucomont 1914, 1924a; Balthasar 1963b; Ochi &amp; Kon 1994, 2008).</p><p>Overall distribution. The Philippines, Indonesia, Malaysia, and Thailand (Ochi &amp; Kon 2008).</p></div>	https://treatment.plazi.org/id/3C6E87D69239283EFF14FE46FAB9FD28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69239283EFF14FC82FB3BFB80.text	3C6E87D69239283EFF14FC82FB3BFB80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Necramator) semicupreus (Harold 1877)	<div><p>Parascatonomus (Necramator) semicupreus (Harold, 1877)</p><p>Onthophagus semicupreus Harold, 1877: 81; Boucomont 1924a: 670.</p><p>Onthophagus (Onthophagus) semicupreus: Balthasar 1963b: 519.</p><p>Onthophagus (Parascatonomus) semicupreus: Ochi et al. 2008: 220; Krikken &amp; Huijbregts 2009: 48.</p><p>Parascatonomus (Necramator) semicupreus: Ueda et al. 2022: 181.</p><p>Type locality. “ Borneo, Sarawak.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Boucomont 1914, 1924a; Balthasar 1963b; Ochi et al. 2008; Krikken &amp; Huijbregts 2009).</p><p>Overall distribution. The Philippines, Indonesia, and Malaysia (Ueda et al. 2022).</p></div>	https://treatment.plazi.org/id/3C6E87D69239283EFF14FC82FB3BFB80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D69239283EFF14FB52FE8DF9AC.text	3C6E87D69239283EFF14FB52FE8DF9AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parascatonomus (Parascatonomus) discedens (Sharp 1875)	<div><p>Parascatonomus (Parascatonomus) discedens (Sharp, 1875)</p><p>Onthophagus discedens Sharp, 1875: 49 .</p><p>Onthophagus (Parascatonomus) discedens: Ochi &amp; Araya 1996: 12; Ochi et al. 2008: 205; Huijbregts &amp; Krikken 2009: 8.</p><p>Parascatonomus discedens: Han et al. 2021: 136.</p><p>Parascatonomus (Parascatonomus) discedens: Ueda et al. 2022: 181.</p><p>Type locality. “ Sarawak.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Ochi &amp; Araya 1996; Ochi et al. 2008; Huijbregts &amp; Krikken 2009).</p><p>Overall distribution. The Philippines, India, Bangladesh, Myanmar, Indonesia, Cambodia, Malaysia, Thailand, Nepal, and China (including Taiwan) (Huijbregts &amp; Krikken 2009; Han et al. 2021; Ueda et al. 2022).</p></div>	https://treatment.plazi.org/id/3C6E87D69239283EFF14FB52FE8DF9AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692392831FF14F8E7FE20FEC4.text	3C6E87D692392831FF14F8E7FE20FEC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sisyphus (Sisyphus) thoracicus subsp. thoracicus Sharp 1875	<div><p>Sisyphus (Sisyphus) thoracicus thoracicus Sharp, 1875</p><p>Sisyphus thoracicus Sharp, 1875: 39; Balthasar 1963a: 241.</p><p>Sisyphus (Sisyphus) thoracicus: Ochi &amp; Kon 1994: 283.</p><p>Type locality. “ Singapore.”</p><p>Distribution in the Philippines. Mindanao (Balthasar 1963a; Ochi &amp; Kon 1994).</p><p>Overall distribution. The Philippines, Indonesia, Malaysia, Singapore, and Thailand (Balthasar 1963a; Ochi &amp; Kon 1994).</p></div>	https://treatment.plazi.org/id/3C6E87D692392831FF14F8E7FE20FEC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FDF7FCDCFCCC.text	3C6E87D692362831FF14FDF7FCDCFCCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera analis (Brenske 1899)	<div><p>Maladera (sensu lato) analis (Brenske, 1899)</p><p>Autoserica analis Brenske, 1899: 207; Dalla Torre 1912a: 20.</p><p>Type locality. “ Philippinen, Luzon.”</p><p>Distribution in the Philippines. Luzon (Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692362831FF14FDF7FCDCFCCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FCDFFCDCFC34.text	3C6E87D692362831FF14FCDFFCDCFC34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera bakeri (Moser 1915)	<div><p>Maladera (sensu lato) bakeri (Moser, 1915)</p><p>Autoserica bakeri Moser, 1915c: 159 .</p><p>Type locality. “ Palawan (P. Princesa)”</p><p>Distribution in the Philippines. Luzon (Palawan) (Moser 1915c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692362831FF14FCDFFCDCFC34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FB87FCDCFB1C.text	3C6E87D692362831FF14FB87FCDCFB1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera davaoana (Moser 1921)	<div><p>Maladera (sensu lato) davaoana (Moser, 1921)</p><p>Autoserica davaoana Moser, 1921: 48 .</p><p>Type locality. “ Mindanao (Davao).”</p><p>Distribution in the Philippines. Mindanao (Davao) (Moser 1921a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692362831FF14FB87FCDCFB1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FB6FFCDCFA44.text	3C6E87D692362831FF14FB6FFCDCFA44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera eremita (Brenske 1899)	<div><p>Maladera (sensu lato) eremita (Brenske, 1899)</p><p>Autoserica eremita Brenske, 1899: 209; Dalla Torre 1912a: 22.</p><p>Type locality. “ Philippinen, Lianga.”</p><p>Distribution in the Philippines. Mindanao (Surigao del Sur) (Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692362831FF14FB6FFCDCFA44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14FA57FCDCF9AC.text	3C6E87D692362831FF14FA57FCDCF9AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera fasciculata (Moser 1922)	<div><p>Maladera (sensu lato) fasciculata (Moser, 1922)</p><p>Autoserica fasciculata Moser, 1922b: 103 .</p><p>Type locality. “Luzon (Montalban).”</p><p>Distribution in the Philippines. Luzon (Rizal) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692362831FF14FA57FCDCF9AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362831FF14F93FFCDCF894.text	3C6E87D692362831FF14F93FFCDCF894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera iliganica (Moser 1917)	<div><p>Maladera (sensu lato) iliganica (Moser, 1917)</p><p>Autoserica iliganica Moser, 1917b: 54 .</p><p>Type locality. “ Mindanao (Iligan).”</p><p>Distribution in the Philippines. Mindanao (Lanao del Norte) (Moser 1917b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692362831FF14F93FFCDCF894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692362830FF14F8E7FCDCFF0C.text	3C6E87D692362830FF14F8E7FCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera isarogensis (Moser 1922)	<div><p>Maladera (sensu lato) isarogensis (Moser, 1922)</p><p>Autoserica isarogensis Moser, 1922b: 102 .</p><p>Type locality. “ Luzon (Mt. Isarog).”</p><p>Distribution in the Philippines. Luzon (Camarines Sur) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692362830FF14F8E7FCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FE9FFCDCFE74.text	3C6E87D692372830FF14FE9FFCDCFE74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera kasigurana (Moser 1922)	<div><p>Maladera (sensu lato) kasigurana (Moser, 1922)</p><p>Autoserica kasigurana Moser, 1922b: 102 .</p><p>Type locality. “Luzon (Kasiguran).”</p><p>Distribution in the Philippines. Luzon (Aurora) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692372830FF14FE9FFCDCFE74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FE47FCDCFD70.text	3C6E87D692372830FF14FE47FCDCFD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera nigrorubra (Brenske 1894)	<div><p>Maladera (sensu lato) nigrorubra (Brenske, 1894)</p><p>Serica nigrorubra Brenske, 1894a: 9; Brenske 1899: 208.</p><p>Autoserica nigrorubra: Dalla Torre 1912a: 24.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Brenske 1894a, 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692372830FF14FE47FCDCFD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FD4BFCDCFC7C.text	3C6E87D692372830FF14FD4BFCDCFC7C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera philippinensis (Blanchard 1851)	<div><p>Maladera (sensu lato) philippinensis (Blanchard, 1851)</p><p>Omaloplia philippinensis Blanchard, 1851: 77; Brenske 1899: 210.</p><p>Autoserica philippinensis: Dalla Torre 1912a: 25.</p><p>Type locality. “ Philippines.”</p><p>Distribution in the Philippines. Philippine archipelago (Blanchard 1851; Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692372830FF14FD4BFCDCFC7C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FC4FFCDCFB78.text	3C6E87D692372830FF14FC4FFCDCFB78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera philippinica (Brenske 1894)	<div><p>Maladera (sensu lato) philippinica (Brenske, 1894)</p><p>Serica philippinica Brenske, 1894a: 9; Brenske 1899: 206.</p><p>Autoserica philippinica: Dalla Torre 1912a: 25.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Luzon (Benguet) (Brenske 1894a, 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692372830FF14FC4FFCDCFB78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FB53FCDCFAA0.text	3C6E87D692372830FF14FB53FCDCFAA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera siargaoensis (Moser 1922)	<div><p>Maladera (sensu lato) siargaoensis (Moser, 1922)</p><p>Autoserica siargaoensis Moser, 1922b: 101 .</p><p>Type locality. “ Philippinen ( Siargao).”</p><p>Distribution in the Philippines. Mindanao (Surigao del Norte) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692372830FF14FB53FCDCFAA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14FA3BFCDCF988.text	3C6E87D692372830FF14FA3BFCDCF988.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera sparsesetosa (Moser 1922)	<div><p>Maladera (sensu lato) sparsesetosa (Moser, 1922)</p><p>Autoserica sparsesetosa Moser, 1922b: 101 .</p><p>Type locality. “Mindanao (Siargao).”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692372830FF14FA3BFCDCF988	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372830FF14F9E3FCDCF894.text	3C6E87D692372830FF14F9E3FCDCF894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera stolida (Brenske 1899)	<div><p>Maladera (sensu lato) stolida (Brenske, 1899)</p><p>Autoserica stolida Brenske, 1899: 210; Dalla Torre 1912a: 26.</p><p>Type locality. “ South Palawan.”</p><p>Distribution in the Philippines. Luzon (Manila and Palawan) and Mindanao (Zamboanga del Norte) (Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692372830FF14F9E3FCDCF894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692372833FF14F8E7FCDCFF0C.text	3C6E87D692372833FF14F8E7FCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera subaana (Moser 1922)	<div><p>Maladera (sensu lato) subaana (Moser, 1922)</p><p>Autoserica subaana Moser, 1922b: 102 .</p><p>Type locality. “Mindoro (Subaan).”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692372833FF14F8E7FCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FED7FCDCFE2C.text	3C6E87D692342833FF14FED7FCDCFE2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera (Omaladera) boettcheri (Moser 1926)	<div><p>Maladera (Omaladera) boettcheri (Moser, 1926)</p><p>Cephaloserica boettcheri Moser, 1926 .</p><p>Type locality. “ Luzon: Mt. Data. ”</p><p>Distribution in the Philippines. Luzon (Mountain Province) (Moser 1926).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692342833FF14FED7FCDCFE2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FDBFFCDCFD14.text	3C6E87D692342833FF14FDBFFCDCFD14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Maladera (Omaladera) longiclava (Moser 1926)	<div><p>Maladera (Omaladera) longiclava (Moser, 1926)</p><p>Cephaloserica longiclava Moser, 1926: 199 .</p><p>Type locality. “ Baguio.”</p><p>Distribution in the Philippines. Luzon (Baguio) (Moser 1926).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692342833FF14FDBFFCDCFD14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FCDFFCDCFC34.text	3C6E87D692342833FF14FCDFFCDCFC34.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica abbreviata Brenske 1899	<div><p>Microserica abbreviata Brenske, 1899</p><p>Microserica abbreviata Brenske, 1899: 216; Dalla Torre 1912a: 34.</p><p>Type locality. “ Philippinen, Mindanao, Bitaibitai.”</p><p>Distribution in the Philippines. Mindanao (Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692342833FF14FCDFFCDCFC34	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FB86FCDCFB1C.text	3C6E87D692342833FF14FB86FCDCFB1C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica arorogensis Moser 1922	<div><p>Microserica arorogensis Moser, 1922</p><p>Microserica arorogensis Moser, 1922b: 111 .</p><p>Type locality. “ Masbate ( Arorog).”</p><p>Distribution in the Philippines. Luzon (Masbate) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692342833FF14FB86FCDCFB1C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FB6EFCDCFA44.text	3C6E87D692342833FF14FB6EFCDCFA44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica balabacensis Moser 1922	<div><p>Microserica balabacensis Moser, 1922</p><p>Microserica balabacensis Moser, 1922b: 108 .</p><p>Type locality. “Balabac.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692342833FF14FB6EFCDCFA44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14FA56FCDCF9AC.text	3C6E87D692342833FF14FA56FCDCF9AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica balbalana Moser 1922	<div><p>Microserica balbalana Moser, 1922</p><p>Microserica balbalana Moser, 1922b: 109 .</p><p>Type locality. “Luzon (Balbalan).”</p><p>Distribution in the Philippines. Luzon (Kalinga) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692342833FF14FA56FCDCF9AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342833FF14F93EFCDCF894.text	3C6E87D692342833FF14F93EFCDCF894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica binaluana Moser 1922	<div><p>Microserica binaluana Moser, 1922</p><p>Microserica binaluana Moser, 1922b: 107 .</p><p>Type locality. “ Palawan ( Binaluan).”</p><p>Distribution in the Philippines. Luzon (Palawan) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692342833FF14F93EFCDCF894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692342832FF14F8E6FCDCFF0C.text	3C6E87D692342832FF14F8E6FCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica calapana Moser 1922	<div><p>Microserica calapana Moser, 1922</p><p>Microserica calapana Moser, 1922b: 109 .</p><p>Type locality. “Mindoro (Calapan).”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692342832FF14F8E6FCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FE9EFCDCFE74.text	3C6E87D692352832FF14FE9EFCDCFE74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica dansalana Moser 1922	<div><p>Microserica dansalana Moser, 1922</p><p>Microserica dansalana Moser, 1922b: 110 .</p><p>Type locality. “Mindanao (Dansalan).”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692352832FF14FE9EFCDCFE74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FE46FCDCFD5C.text	3C6E87D692352832FF14FE46FCDCFD5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica difficilis Moser 1922	<div><p>Microserica difficilis Moser, 1922</p><p>Microserica difficilis Moser, 1922b: 108 .</p><p>Type locality. “ Palawan ( Binaluan).”</p><p>Distribution in the Philippines. Luzon (Palawan) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692352832FF14FE46FCDCFD5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FD2EFCDCFC58.text	3C6E87D692352832FF14FD2EFCDCFC58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica fugax (Erichson 1834)	<div><p>Microserica fugax (Erichson, 1834)</p><p>Serica fugax Erichson, 1834: 239 .</p><p>Microserica fugax: Brenske 1899: 213; Dalla Torre 1912a: 34.</p><p>Type locality. “ Die Insel Luzon.”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Erichson 1834; Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692352832FF14FD2EFCDCFC58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FC32FCDCFB80.text	3C6E87D692352832FF14FC32FCDCFB80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica humilis Brenske 1899	<div><p>Microserica humilis Brenske, 1899</p><p>Microserica humilis Brenske, 1899: 218; Dalla Torre 1912a: 34.</p><p>Type locality. “ Philippinen, Luzon.”</p><p>Distribution in the Philippines. Luzon (Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692352832FF14FC32FCDCFB80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FB1AFCDCFAE8.text	3C6E87D692352832FF14FB1AFCDCFAE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica kolambugana Moser 1922	<div><p>Microserica kolambugana Moser, 1922</p><p>Microserica kolambugana Moser, 1922b: 110 .</p><p>Type locality. “Mindanao (Kolambugan).”</p><p>Distribution in the Philippines. Mindanao (Lanao del Norte) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692352832FF14FB1AFCDCFAE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14FAC2FCDCF9D0.text	3C6E87D692352832FF14FAC2FCDCF9D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica liangensis Brenske 1899	<div><p>Microserica liangensis Brenske, 1899</p><p>Microserica liangensis Brenske, 1899: 216; Dalla Torre 1912a: 34.</p><p>Type locality. “ Philippinen, Mindanao, Lianga.”</p><p>Distribution in the Philippines. Mindanao (Surigao del Sur) (Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692352832FF14FAC2FCDCF9D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14F9AAFCDCF938.text	3C6E87D692352832FF14F9AAFCDCF938.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica mindoroana Brenske 1899	<div><p>Microserica mindoroana Brenske, 1899</p><p>Microserica mindoroana Brenske, 1899: 214; Dalla Torre 1912a: 35; Moser 1922b: 111.</p><p>Type locality. “ Philippinen, Mindoro.”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Brenske 1899; Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692352832FF14F9AAFCDCF938	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692352832FF14F892FCDCF860.text	3C6E87D692352832FF14F892FCDCF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica negrosiana Brenske 1899	<div><p>Microserica negrosiana Brenske, 1899</p><p>Microserica negrosiana Brenske, 1899: 215; Dalla Torre 1912a: 35.</p><p>Type locality. “ Philippinen, Negros Isl. ”</p><p>Distribution in the Philippines. Visayas (Negros) (Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692352832FF14F892FCDCF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FF0BFCDCFEBD.text	3C6E87D692322835FF14FF0BFCDCFEBD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica oceana (Brenske 1894)	<div><p>Microserica oceana (Brenske, 1894)</p><p>Serica oceana Brenske, 1894a: 45 .</p><p>Microserica oceana: Brenske 1899: 218; Dalla Torre 1912a: 35.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Philippine archipelago (Brenske 1894a, 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692322835FF14FF0BFCDCFEBD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FE0FFCDCFDE5.text	3C6E87D692322835FF14FE0FFCDCFDE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica palawana Brenske 1899	<div><p>Microserica palawana Brenske, 1899</p><p>Microserica palawana Brenske, 1899: 214; Dalla Torre 1912a: 35.</p><p>Type locality. “ South Palawan.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692322835FF14FE0FFCDCFDE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FDF7FCDCFCCD.text	3C6E87D692322835FF14FDF7FCDCFCCD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica panayana Moser 1924	<div><p>Microserica panayana Moser, 1924</p><p>Microserica panayana Moser, 1924a: 139 .</p><p>Type locality. “ N. W. Panay. ”</p><p>Distribution in the Philippines. Visayas (Panay Island) (Moser 1924a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692322835FF14FDF7FCDCFCCD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FCDFFCDCFC35.text	3C6E87D692322835FF14FCDFFCDCFC35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica rufolutea Moser 1915	<div><p>Microserica rufolutea Moser, 1915</p><p>Microserica rufolutea Moser, 1915c: 185 .</p><p>Type locality. “ Philippinen ( Paete).”</p><p>Distribution in the Philippines. Luzon (Laguna) (Moser 1915c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692322835FF14FCDFFCDCFC35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FB87FCDCFB1D.text	3C6E87D692322835FF14FB87FCDCFB1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica samarana Brenske 1899	<div><p>Microserica samarana Brenske, 1899</p><p>Microserica samarana Brenske, 1899: 215; Dalla Torre 1912a: 35.</p><p>Type locality. “ Philippinen, Samar Isl.”</p><p>Distribution in the Philippines. Visayas (Samar) (Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692322835FF14FB87FCDCFB1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14FB6FFE13F9D1.text	3C6E87D692322835FF14FB6FFE13F9D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica semperi (Brenske 1894)	<div><p>Microserica semperi (Brenske, 1894)</p><p>Serica semperi Brenske, 1894: 12 .</p><p>Microserica Semperi: Brenske 1899: 217; Dalla Torre 1912a: 36.</p><p>Type locality. “ Philippinen.”</p><p>Distribution in the Philippines. Luzon (Brenske 1894a, 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks. Brenske observed morphological variations and gave a detailed description of this species (see Brenske 1899).</p></div>	https://treatment.plazi.org/id/3C6E87D692322835FF14FB6FFE13F9D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14F9ABFCDCF939.text	3C6E87D692322835FF14F9ABFCDCF939.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica sibuyana Moser 1922	<div><p>Microserica sibuyana Moser, 1922</p><p>Microserica sibuyana Moser, 1922b: 108 .</p><p>Type locality. “ Philippinen: Ins. Sibuyan.”</p><p>Distribution in the Philippines. Luzon (Sibuyan) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692322835FF14F9ABFCDCF939	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692322835FF14F893FCDCF861.text	3C6E87D692322835FF14F893FCDCF861.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica theodoroensis Moser 1922	<div><p>Microserica theodoroensis Moser, 1922</p><p>Microserica theodoroensis Moser, 1922b: 111 .</p><p>Type locality. “Mindoro (San Theodoro, Subaan).”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692322835FF14F893FCDCF861	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FF0BFCDCFE99.text	3C6E87D692332834FF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microserica viridifrons Moser 1922	<div><p>Microserica viridifrons Moser, 1922</p><p>Microserica viridifrons Moser, 1922b: 110 .</p><p>Type locality. “ Masbate (Arorog).</p><p>Distribution in the Philippines. Luzon (Masbate) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692332834FF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FB87FCDCFB1D.text	3C6E87D692332834FF14FB87FCDCFB1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica armata Moser 1922	<div><p>Neoserica (sensu lato) armata Moser, 1922</p><p>Neoserica armata Moser, 1922: 105 .</p><p>Type locality. “ Luzon (Mt. Banahao).”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692332834FF14FB87FCDCFB1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FB6FFCDCFA45.text	3C6E87D692332834FF14FB6FFCDCFA45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica balabaca Brenske 1899	<div><p>Neoserica (sensu lato) balabaca Brenske, 1899</p><p>Neoserica balabaca Brenske, 1899: 211; Dalla Torre 1912a: 30.</p><p>Type locality. “Balabac.”</p><p>Distribution in the Philippines. Luzon (Palawan) (Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692332834FF14FB6FFCDCFA45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FA57FCDCF9AD.text	3C6E87D692332834FF14FA57FCDCF9AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica butuana Moser 1922	<div><p>Neoserica (sensu lato) butuana Moser, 1922</p><p>Neoserica butuana Moser, 1922b: 105 .</p><p>Type locality. “ Mindanao (Butuan).”</p><p>Distribution in the Philippines. Mindanao (Agusan del Norte) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692332834FF14FA57FCDCF9AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14FE63FE0DFC7D.text	3C6E87D692332834FF14FE63FE0DFC7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica davaoensis Moser 1917	<div><p>Neoserica (sensu lato) davaoensis Moser, 1917</p><p>Autoserica davaoensis Moser, 1917b: 55 .</p><p>Type locality. “ Mindanao (Davao).”</p><p>Distribution in the Philippines. Mindanao (Davao) (Moser 1917b).</p><p>Overall distribution. Endemic species to the Philippines.</p><p>Remarks. This species is currently placed in the genus Neoserica and subgenus Autoserica . However, this placement is highly doubtful, as most species described under Autoserica [auctorum] by Brenske and Moser are now recognized as Maladera Mulsant, 1871 (Pope, 1960; Ahrens &amp; Fabrizi, 2016). The taxonomic status of all Philippine Sericinae requires a major revision, since most species are known only from their original descriptions. While several taxonomic reappraisals of numerous genera and subgenera have been made in other Oriental and Palearctic regions (e.g., by Ahrens and colleagues), the Philippine fauna remains largely unrevised.</p></div>	https://treatment.plazi.org/id/3C6E87D692332834FF14FE63FE0DFC7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332834FF14F93FFCDCF895.text	3C6E87D692332834FF14F93FFCDCF895.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica denticulata Moser 1922	<div><p>Neoserica (sensu lato) denticulata Moser, 1922</p><p>Neoserica denticulata Moser, 1922b: 105 .</p><p>Type locality. “ Philippinen: Bucas Grande (Socorro).”</p><p>Distribution in the Philippines. Mindanao (Surigao del Norte) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692332834FF14F93FFCDCF895	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692332837FF14F8E7FCDCFF0C.text	3C6E87D692332837FF14F8E7FCDCFF0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica fuscula Moser 1921	<div><p>Neoserica (sensu lato) fuscula Moser, 1921</p><p>Neoserica fuscula Moser, 1921a: 49 .</p><p>Type locality. “ Luzon (Mt. Makiling. ”</p><p>Distribution in the Philippines. Luzon (Laguna) (Moser 1921a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692332837FF14F8E7FCDCFF0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FE9EFCDCFE74.text	3C6E87D692302837FF14FE9EFCDCFE74.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica longiclava Moser 1916	<div><p>Neoserica (sensu lato) longiclava Moser, 1916</p><p>Neoserica longiclava Moser, 1916a: 156 .</p><p>Type locality. “Luzon (Bagnio).”</p><p>Distribution in the Philippines. Luzon (Baguio) (Moser 1916a).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692302837FF14FE9EFCDCFE74	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FE46FCDCFD70.text	3C6E87D692302837FF14FE46FCDCFD70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica lucifuga Brenske 1899	<div><p>Neoserica (sensu lato) lucifuga Brenske, 1899</p><p>Neoserica lucifuga Brenske, 1899: 212; Dalla Torre 1912a: 31.</p><p>Type locality. “ Philippinen, Mindanao, Dapitan.”</p><p>Distribution in the Philippines. Luzon (Catanduanes) and Mindanao (Zamboanga del Norte) (Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692302837FF14FE46FCDCFD70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FD4AFCDCFC58.text	3C6E87D692302837FF14FD4AFCDCFC58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica luzonica Moser 1915	<div><p>Neoserica (sensu lato) luzonica Moser, 1915</p><p>Neoserica luzonica Moser, 1915c: 170 .</p><p>Type locality. “ Luzon (Mt. Banahao).”</p><p>Distribution in the Philippines. Luzon (Mt. Banahao) (Moser 1915c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692302837FF14FD4AFCDCFC58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FC32FCDCFB80.text	3C6E87D692302837FF14FC32FCDCFB80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica makilingica Moser 1915	<div><p>Neoserica (sensu lato) makilingica Moser, 1915</p><p>Neoserica makilingica Moser, 1915c: 169 .</p><p>Type locality. “ Luzon (Mt. Makiling).”</p><p>Distribution in the Philippines. Luzon (Laguna) (Moser 1915c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692302837FF14FC32FCDCFB80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FB1AFCDCFAE8.text	3C6E87D692302837FF14FB1AFCDCFAE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica mucronota Moser 1922	<div><p>Neoserica (sensu lato) mucronota Moser, 1922</p><p>Neoserica mucronota Moser, 1922b: 106 .</p><p>Type locality. “ Masbate ( Arorog).”</p><p>Distribution in the Philippines. Luzon (Masbate) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692302837FF14FB1AFCDCFAE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14FAC2FCDCF9D0.text	3C6E87D692302837FF14FAC2FCDCF9D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica opacula Moser 1924	<div><p>Neoserica (sensu lato) opacula Moser, 1924</p><p>Neoserica opacula Moser, 1924b: 349 .</p><p>Type locality. “Sibuyan.”</p><p>Distribution in the Philippines. Luzon (Sibuyan) (Moser 1924b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692302837FF14FAC2FCDCF9D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14F9AAFCDCF938.text	3C6E87D692302837FF14F9AAFCDCF938.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica setiventris Moser 1913	<div><p>Neoserica (sensu lato) setiventris Moser, 1913</p><p>Neoserica setiventris Moser, 1913: 331 .</p><p>Type locality. “ Philippinen ( Los Banos).”</p><p>Distribution in the Philippines. Luzon (Laguna) (Moser 1913).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692302837FF14F9AAFCDCF938	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692302837FF14F892FCDCF860.text	3C6E87D692302837FF14F892FCDCF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica squamuligera Moser 1915	<div><p>Neoserica (sensu lato) squamuligera Moser, 1915</p><p>Neoserica squamuligera Moser, 1915c: 173 .</p><p>Type locality. “Mindanao (Dapitan).”</p><p>Distribution in the Philippines. Mindanao (Zamboanga del Norte) (Moser 1915c).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692302837FF14F892FCDCF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FF0BFCDCFE99.text	3C6E87D692312836FF14FF0BFCDCFE99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica surigaoana Moser 1917	<div><p>Neoserica (sensu lato) surigaoana Moser, 1917</p><p>Neoserica surigaoana Moser, 1917b: 59 .</p><p>Type locality. “Mindanao (Surigao).”</p><p>Distribution in the Philippines. Mindanao (Surigao) (Moser 1917b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692312836FF14FF0BFCDCFE99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FEF3FCDCFDC1.text	3C6E87D692312836FF14FEF3FCDCFDC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Neoserica uncinata Brenske 1899	<div><p>Neoserica (sensu lato) uncinata Brenske, 1899</p><p>Neoserica uncinata Brenske, 1899: 213; Dalla Torre 1912a: 33.</p><p>Type locality. “ North Luzon. ”</p><p>Distribution in the Philippines. Luzon (Brenske 1899).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692312836FF14FEF3FCDCFDC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FD13FCDCFCE1.text	3C6E87D692312836FF14FD13FCDCFCE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Periserica zamboangensis Moser 1917	<div><p>Periserica zamboangensis Moser, 1917</p><p>Periserica zamboangensis Moser, 1917b: 61 .</p><p>Type locality. “ Mindanao (Zamboanga).”</p><p>Distribution in the Philippines. Mindanao (Zamboanga) (Moser 1917b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692312836FF14FD13FCDCFCE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FC33FCDCFB81.text	3C6E87D692312836FF14FC33FCDCFB81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trioserica lepichaeta Moser 1922	<div><p>Trioserica lepichaeta Moser, 1922</p><p>Trioserica lepichaeta Moser, 1922b: 112 .</p><p>Type locality. “Mindoro (Subaan).”</p><p>Distribution in the Philippines. Luzon (Mindoro) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692312836FF14FC33FCDCFB81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FB1BFCDCFAE9.text	3C6E87D692312836FF14FB1BFCDCFAE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trioserica sparsesquamosa Moser 1922	<div><p>Trioserica sparsesquamosa Moser, 1922</p><p>Trioserica sparsesquamosa Moser, 1922b: 112 .</p><p>Type locality. “ Mindanao ( Surigao, Iligan).”</p><p>Distribution in the Philippines. Mindanao (Lanao del Norte and Surigao) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692312836FF14FB1BFCDCFAE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
3C6E87D692312836FF14FAC3FCDCF9D1.text	3C6E87D692312836FF14FAC3FCDCF9D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trioserica trifoliata Moser 1922	<div><p>Trioserica trifoliata Moser, 1922</p><p>Trioserica trifoliata Moser, 1922b: 112 .</p><p>Type locality. “ Mindanao (Butuan).”</p><p>Distribution in the Philippines. Mindanao (Agusan del Norte) (Moser 1922b).</p><p>Overall distribution. Endemic species to the Philippines.</p></div>	https://treatment.plazi.org/id/3C6E87D692312836FF14FAC3FCDCF9D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Magallanes, Joriemar G.;Nuñeza, Olga M.;Mondejar, Eddie P.	Magallanes, Joriemar G., Nuñeza, Olga M., Mondejar, Eddie P. (2026): First in 109 years: Catalogue of scarab beetles (Coleoptera: Scarabaeidae) of the Philippines. Zootaxa 5746 (1): 1-148, DOI: 10.11646/zootaxa.5746.1.1, URL: https://doi.org/10.11646/zootaxa.5746.1.1
