identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
381F87A1FFAEFFC1FF41F9A858EF2B78.text	381F87A1FFAEFFC1FF41F9A858EF2B78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhynchoclaviger Wasmann 1891	<div><p>Key to genera of Rhynchoclaviger group</p><p>1 Antennae pentamerous; sides of pronotum completely non-dentate, lacking longitudinal median sulcus; elytra lacking striae.............................................................................. Rhynchoclaviger Wasmann</p><p>- Antennae tetramerous; sides of pronotum dentate, with longitudinal median sulcus; elytra with some striae................................................................................................. Paussiger Wasmann</p></div>	https://treatment.plazi.org/id/381F87A1FFAEFFC1FF41F9A858EF2B78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Raveloson, Ricca;Andrianaharison, Fumiko;Rakotonirina, Jean Claude;Hlaváč, Peter	Raveloson, Ricca, Andrianaharison, Fumiko, Rakotonirina, Jean Claude, Hlaváč, Peter (2025): The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. X. Revision of the myrmecophilous clavigerine genera Rhynchoclaviger Wasmann, 1891 and Paussiger Wasmann, 1893. Zootaxa 5673 (3): 399-410, DOI: 10.11646/zootaxa.5673.3.4, URL: https://doi.org/10.11646/zootaxa.5673.3.4
381F87A1FFADFFC3FF41FF405CE02DE6.text	381F87A1FFADFFC3FF41FF405CE02DE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhynchoclaviger Wasmann 1891	<div><p>Rhynchoclaviger Wasmann, 1891</p><p>(Figs 1, 5–9)</p><p>Rhynchoclaviger Wasmann, 1891: 4 . Types species: Rhynchoclaviger cremastogastris Wasmann, by monotypy.</p><p>Rhynchoclaviger Wasmann; Wasmann 1893a: 111 (in key); Raffray 1908: 420 (in key); Raffray 1908: 422 (redescription, catalogue); Jeannel 1954: 295 (in key); Jeannel 1954: 304 (redescription); Jeannel 1960: 191 (in key); Jeannel 1960: 201; Newton &amp; Chandler 1989: 66 (catalogue); Sabella, Viglianisi &amp; Petralia 2012: 432 (catalogue).</p><p>Diagnosis. Head capsule about twice as long as wide, lacking frontal and vertexal foveae, temples non-dentate, parallel-sided; head capsule delimited from neck region by straight occipital carina; posterior tentorial pits confluent in deep median impression; antenna pentamerous, scape minuscule, pedicel cylindrical, slightly transverse; terminal antennomere longer than 1–3 combined; pronotum non-dentate on sides, shorter than head capsule, with median antebasal foveae; lacking longitudinal sulcus or carinae; metaventrite in anterior part, with sharp, triangular median carina posteriorly bearing bunch of golden setae; elytra glabrous, lacking antebasal foveae, lacking striae, with lateral carinae, posterior margin of elytron lacking trichomes but with few long setae; first paratergite lacking lateral protrusion; mesotibia modified in both sexes; aedeagus almost symmetrical, about 2.8 times as long as wide, distal lobe with two long setae.</p><p>Redescription. Body (Fig. 1) more than twice as long as wide, more than 2.3 mm long; reddish-brown, shiny, head capsule and pronotum with rough surface, elytra and abdomen smooth.</p><p>Head capsule sub-rectangular, about twice as long as wide, widest at level of eyes; with rostral lateral carina; lacking frontal and vertexal foveae; clypeus partly visible on sides; rostrum at anterior margin triangular; eyes moderately prominent, situated at about midpoint along sides of head capsule; temples non-dentate, parallel-sided, slightly shorter than frontal rostrum; head capsule delimited from neck region by straight occipital carina, neck region almost entirely retracted into prothorax; gular plate with variable structure, lateral portions with rugose surface, median part smooth, posterior tentorial pits confluent in deep median impression; neck region ventrally demarcated by transverse carinae.</p><p>Antenna pentamerous, clearly longer than head capsule; scape minuscule, entirely hidden under rostrum; pedicel cylindrical, slightly transverse; antennomere 3 resembles mushroom of the genus Boletus, more than twice as long as pedicel; antennomere 4 pedunculate at base, about 1.6 times as long as wide; terminal antennomere longest, longer than 1–3 combined, slightly dilated from base, from middle to apex parallel-sided, its apex truncate, lacking concentric short setae.</p><p>Pronotum about as long as wide, non-dentate on sides, shorter than head capsule; lacking squamous setae; lateral margin slightly convergent posteriad, strongly convergent anteriad; with median antebasal foveae; lacking longitudinal sulcus or carinae.</p><p>Prosternum fused with hypomera, with median triangular, sharp elevation, procoxae contiguous. Mesoventrite distinctly shorter than metaventrite; mesocoxae narrowly separated, isthmus width about third of diameter of mesocoxal cavity; disc of metaventrite in anterior part convex, with sharp, triangular median carina bearing on its posterior part bunch of golden setae; metaventrite declining to posterior margin, lacking any projection; metacoxae widely separated by wide metaventral posterior process which is strongly concave in middle.</p><p>Elytra 1.4–1.5 times as wide as long, much shorter than abdomen, glabrous, lacking antebasal foveae, lacking striae, with lateral carinae; postero-lateral corner of elytron obliquely bevelled mesad, with minuscule projection; humeri rounded, with weakly-defined humeral protrusion, lacking spines; posterior margin of elytron lacking trichomes but with few long setae.</p><p>Abdomen with composite tergite convex, with wide impression in basal half; lacking keels, with two lateral small asetose foveae; first three visible paratergites well-developed along all length of composite tergite, narrow, first paratergite lacking lateral protrusion. First visible sternite (III) short, second (IV) with transversal impression and median longitudinal carina, long, longer than third visible sternite (V).</p><p>Legs short and stout, protibia (Fig. 5) with short, sharp preapical spine; mesofemur with sharp, long, basal spine, mesotibia folded in middle, apical part moderately dentate (Fig. 6).</p><p>Aedeagus (Figs 8, 9) almost symmetrical, elongate, about 2.8 times as long as wide, moderately slender; basal bulb well-separated from distal lobe, distal lobe about 1.2 times as long as basal bulb, bearing two long setae; endophallus with some different sclerites; dorsal diaphragm elliptical.</p><p>Sexual dimorphism. Females larger, more than 2.60 mm long, with shorter antennae, antennomere 4 only about 1.25 times as long as wide; mesofemora simple, mesotibiae with obtuse, preapical protrusion (Fig. 7).</p><p>Distribution. Madagascar.</p><p>Remarks. Rhynchoclaviger is similar in appearance to Paussiger, but can be readily separated by the combination of following characters: 1) pentamerous antennae (tetramerous in Paussiger), 2) temples and pronotum on sides non-dentate (strongly dentate in Paussiger), 3) posterior part of head capsule behind eyes lacking carinae (with various number of carinae in Paussiger), 4) lateral abdominal foveae of composite tergite asetose (setose in Paussiger) and 5) metaventrite with median carina with bunch of setae on its posterior part (glabrous, lacking bunch of setae in Paussiger).</p></div>	https://treatment.plazi.org/id/381F87A1FFADFFC3FF41FF405CE02DE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Raveloson, Ricca;Andrianaharison, Fumiko;Rakotonirina, Jean Claude;Hlaváč, Peter	Raveloson, Ricca, Andrianaharison, Fumiko, Rakotonirina, Jean Claude, Hlaváč, Peter (2025): The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. X. Revision of the myrmecophilous clavigerine genera Rhynchoclaviger Wasmann, 1891 and Paussiger Wasmann, 1893. Zootaxa 5673 (3): 399-410, DOI: 10.11646/zootaxa.5673.3.4, URL: https://doi.org/10.11646/zootaxa.5673.3.4
381F87A1FFACFFC3FF41FDC05C8729D5.text	381F87A1FFACFFC3FF41FDC05C8729D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhynchoclaviger cremastogastris Wasmann 1891	<div><p>Rhynchoclaviger cremastogastris Wasmann, 1891</p><p>Figs 1, 5–9</p><p>Rhynchoclaviger cremastogastris Wasmann, 1891: 4 (Type locality: Madagascar, Andrangoloaka. Type series: more syntypes deposited in MNHN (Jeannel 1954: 305).</p><p>Rhynchoclaviger cremastogastris Wasmann; Jeannel 1954: 305 (redescription); Jeannel 1956: 50 (distribution, host ants); Jeannel 1960: 201 (distribution, host ant).</p><p>Material examined. Holotype ♂, four labels: “ Mdgk. Sikora / Andrangoloaka “ [white, printed and handwritten], “TYPE“ [red, printed], “MUSEUM PARIS / 1952 / COLL. R. OBERTHÜR“ [blue, printed], “HOLOTYPE / Rhynchoclaviger / cremastogastris Wasmann / P. Hlaváč des. 2020“ [red, printed] (MNHN).</p><p>Other material examined. 1 ♀: two labels “ Madagascar / ( Sikora)“ [white, handwritten], “ Rhynchoclaviger / cremastogastris Wasm / P. Hlaváč det., 2017“ [white, printed] (MNHN). 1 ♀: three labels “MUSEUM PARIS / MADAGASCAR / GRANDIDIER 2855-90“ [white, printed], “ Rhynchoclaviger / cremastogastris / Raffray det. 1907 Wasmann“ [white, handwritten], “ Rhynchoclaviger / cremastogastris Wasm / P. Hlaváč det., 2017“ [white, printed] (MNHN). 2 ♂, 2 ♀: three labels “ Madagascar / La Mandrake “ [white, handwritten], “Robinson I. 59 / Crematogaster “ [white, handwritten], “ Rhynchoclaviger / cremastogastris Wasm. / P. Hlaváč det., 2017“ [white, printed] (MNHN). 1 ♀: four labels “ Périnet “ [white, printed], “ Rhynchoclaviger / cremastogastris Ws. “ [white, handwritten], “INSTITUT / SCIENTIFIQUE / MADAGASCAR “ [green, printed] “ Rhynchoclaviger / cremastogastris Wasm / P. Hlaváč det., 2017“ [white, printed] (MNHN).</p><p>Diagnosis. As for the genus.</p><p>Redescription. Body reddish-brown; males: 2.30–2.40 mm, females: 2.60–2.70 mm long. Head capsule 1.95– 2.00 times as long as wide, 1.15–1.25 as long as pronotum. Antenna long, length 0.65 mm in males, 0.55 mm in females, when bent backward reaching half of length of pronotum, antennomere 4 in males 1.6–1.7 times as long as wide, in females 1.2–1.3 times as long as wide. Aedeagus 0.43 mm long, apex of distal lobe sharply curved in lateral view, acute (Figs 8, 9).</p><p>Host ant. Crematogaster sp.</p><p>Distribution. Madagascar.</p></div>	https://treatment.plazi.org/id/381F87A1FFACFFC3FF41FDC05C8729D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Raveloson, Ricca;Andrianaharison, Fumiko;Rakotonirina, Jean Claude;Hlaváč, Peter	Raveloson, Ricca, Andrianaharison, Fumiko, Rakotonirina, Jean Claude, Hlaváč, Peter (2025): The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. X. Revision of the myrmecophilous clavigerine genera Rhynchoclaviger Wasmann, 1891 and Paussiger Wasmann, 1893. Zootaxa 5673 (3): 399-410, DOI: 10.11646/zootaxa.5673.3.4, URL: https://doi.org/10.11646/zootaxa.5673.3.4
381F87A1FFACFFC5FF41F9FD5E772BEB.text	381F87A1FFACFFC5FF41F9FD5E772BEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paussiger Wasmann 1893	<div><p>Paussiger Wasmann, 1893</p><p>(Figs 2–4, 10, 11)</p><p>Paussiger Wasmann, 1893b: 257 . Types species: Paussiger limicornisi Wasmann, by monotypy.</p><p>Paussiger Wasmann: Wasmann 1893a: 110 (in key); Raffray 1905: 452 (in key); Raffray 1908: 422 (in key); Raffray 1908: 444 (redescription, catalogue); Jeannel 1954: 295 (in key); Jeannel 1954: 303 (redescription); Jeannel 1960: 191 (in key); Jeannel 1960: 200; Newton &amp; Chandler 1989: 66 (catalogue); Besuchet 2008: 65 (diagnosis, synonymy of Marofusiger); Sabella, Viglianisi &amp; Petralia 2012: 432 (catalogue).</p><p>Sufifer Newton &amp; Chandler 1989: 66 . New name for Fusifer Raffray, preoccupation, not Dendy 1896. Syn. nov.</p><p>Sufifer Newton &amp; Chandler; Sabella, Viglianisi &amp; Petralia 2012: 433 (catalogue).</p><p>Fusifer Raffray, 1900: 524 . Types species: Commatoceropsis perrieri Fairmaire, by monotypy.</p><p>Fusifer Raffray; Raffray 1908: 444 (catalogue, as synonym to Paussiger); Jeannel 1954: 295 (in key); Jeannel 1954: 302 (redescription); Jeannel 1960: 192 (in key); Jeannel 1960: 199 (key to species).</p><p>Marofusiger Dajoz, 1982: 517 . Types species: Marofusiger grandis Dajoz, original designation; synonymy in Besuchet 2008: 65.</p><p>Diagnosis. Head capsule sub-rectangular, longer than wide, with various number of carina, lacking frontal and vertexal foveae; temples parallel-sided, strongly dentate, head capsule delimited from neck region by occipital carina; posterior tentorial pits small, well-separated; antenna tetramerous, scape cylindrical; antennomere 3 strongly transverse, trapezoidal, well-delimited or fused to terminal antennomere which is longer than 1–3 combined; pronotum slightly transverse, strongly dentate on sides; lateral margin slightly convergent or divergent posteriad; with shallow median longitudinal sulcus, with or without longitudinal carinae, lacking foveae; metaventrite convex in anterior part, declining to posterior margin, lacking any projection or setae; elytral disc glabrous or with small setae, lateral margin with few short setae; with four antebasal foveae; with some striae or lines of adjacent dots; first paratergite with or without round lateral protrusion bearing trichome; mesofemur and mesotibia in males bearing spines or teeth, unmodified in females; aedeagus elongate, moderately slender, clearly longer than wide.</p><p>Redescription. Body (Figs 2–4) more than twice as long as wide, 2.10–2.80 mm long; reddish-brown; head capsule and pronotum with rugose surface, elytra with variously structured surface, abdomen smooth, both shiny.</p><p>Head capsule sub-rectangular, longer than wide, with various number of carina, widest in level of eyes; lacking frontal and vertexal foveae; clypeus partly visible on sides; rostrum at anterior margin triangular or rounded (one undescribed species); eyes moderately prominent, situated at about midpoint along sides of head capsule; temples parallel-sided, strongly dentate, slightly shorter than frontal rostrum; head capsule delimited from neck region by occipital carina, neck region almost entirely retracted into prothorax; gular plate with variable structure, whole surface rough, lateral portions with transverse ribs, with large median sulcus; posterior tentorial pits small, well-separated, situated on posterior margin of sulcus; neck region ventrally demarcated by transverse impression.</p><p>Antenna tetramerous, short, at most as long as head capsule; scape cylindrical, almost entirely hidden under rostrum; pedicel resembles mushroom of the genus Boletus, longer than scape; antennomere 3 strongly transverse, trapezoidal, well-delimited or fused to antennomere 4; terminal antennomere longest, longer than 1–3 combined, slightly narrowed distad, its apex truncate, lacking concentric short setae.</p><p>Pronotum slightly wider than long, strongly dentate on sides, slightly shorter than head capsule; lacking squamous setae; lateral margin slightly convergent or divergent posteriad; with shallow median longitudinal sulcus, median and lateral foveae, median fovea sometimes ill-defined, with or without longitudinal carinae.</p><p>Prosternum fused with hypomera, with median triangular elevation, procoxae contiguous.Mesoventrite distinctly shorter than metaventrite; mesocoxae narrowly separated, isthmus about half of diameter of mesocoxal cavity; disc of metaventrite convex in anterior part, declining to posterior margin, lacking any projection; metaventral posterior process wide, strongly concave.</p><p>Elytra clearly longer than wide, clearly shorter than abdomen, disc glabrous or with small setae, lateral margin with few short setae; with four antebasal foveae; with some striae or lines of adjacent dots; postero-lateral corner of elytron obliquely bevelled mesad, lacking projection; humeri rounded, lacking spines; posterior margin of elytron lacking trichomes.</p><p>Abdomen with composite tergite with transverse basal impression; lacking keels, with two lateral large setose foveae; first three visible paratergites well-developed along almost all length of composite tergite, narrow, first paratergite with or without round lateral protrusion bearing trichome. First visible sternite long (III) with median carina, with lateral impressions, about as long as second visible sternite (IV); sternite (IV) clearly longer than third visible sternite (V), sternite (V) longer than (VI).</p><p>Legs short and stout; femora with interlocking ridges; mesofemur and mesotibia in males bearing spines or teeth, metatibia bent in antebasal part.</p><p>Aedeagus elongate, clearly longer than wide; basal bulb well-separated from distal lobe, distal lobe longer or shorter than basal bulb; endophallus lacking sclerites; dorsal diaphragm round or elliptical.</p><p>Sexual dimorphism. Females with mesofemora and mesotibia simple, lacking spines.</p><p>Host ant. The only Crematogaster hova -complex Blaimer, 2010 [as Crematogaster (Decacrema) schencki Forel, 1891] is so far known to host the genus Paussiger .</p><p>Distribution. Madagascar.</p><p>Remarks. The synonymy of monospecific Sufifer with Paussiger was already proposed by Raffray (1908) and this synonymy is confirmed here. Except for the different structure of antennae, some differences in external structures, and the different shape of the aedeagus, no relevant diagnostic character to support the separation of these two genera are apparent. Based on this synonymy and the description of one new species bellow, Paussiger has four species which can be identified using the following key</p></div>	https://treatment.plazi.org/id/381F87A1FFACFFC5FF41F9FD5E772BEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Raveloson, Ricca;Andrianaharison, Fumiko;Rakotonirina, Jean Claude;Hlaváč, Peter	Raveloson, Ricca, Andrianaharison, Fumiko, Rakotonirina, Jean Claude, Hlaváč, Peter (2025): The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. X. Revision of the myrmecophilous clavigerine genera Rhynchoclaviger Wasmann, 1891 and Paussiger Wasmann, 1893. Zootaxa 5673 (3): 399-410, DOI: 10.11646/zootaxa.5673.3.4, URL: https://doi.org/10.11646/zootaxa.5673.3.4
381F87A1FFA9FFC6FF41FF4058EE2EB1.text	381F87A1FFA9FFC6FF41FF4058EE2EB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paussiger Wasmann 1893	<div><p>Key to species of the genus Paussiger</p><p>1 Antennae short, antennomere 4 about 0.20 mm long, less than 1.9 times as long as wide, pronotum with well-defined median fovea; distal lobe of aedeagus more than 3 times shorter than basal bulb (Jeannel, 1954: 303, Fig. 163)............................................................................................ Paussiger perrieri (Fairmaire)</p><p>- Antennae long, antennomere 4 more than 0.25 mm long, more than twice as long as wide, pronotum with ill-defined median fovea; distal lobe of aedeagus longer than basal bulb......................................................... 2</p><p>2 First visible paratergite strongly protruding laterally, width here considerably wider than maximum width of elytra; antennomere 3 well delimited from 4; pronotum sub-elliptical, from mid length slightly convergent posteriad or parallel-sided......... 3</p><p>- First visible paratergite not protruding laterally, width here slightly less than maximum width of elytra; antennomere 3 confluent with 4; pronotum trapezoidal, evenly divergent posteriad........................... Paussiger limicornis (Wasmann)</p><p>3 Head capsule with two parallel carinae; antenna long, about as long as head capsule; antennomere 5 more than 2.50 times as long as wide, mesofemur with sharp, long, outwards curved basal spine..................... Paussiger grandis (Dajoz)</p><p>- Head capsule with 5 parallel carinae; antenna shorter than head capsule; antennomere 5 less than 2.20 times as long as wide, mesofemur with sharp, short, almost straight basal spine.................................. Paussiger fisheri sp. nov.</p></div>	https://treatment.plazi.org/id/381F87A1FFA9FFC6FF41FF4058EE2EB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Raveloson, Ricca;Andrianaharison, Fumiko;Rakotonirina, Jean Claude;Hlaváč, Peter	Raveloson, Ricca, Andrianaharison, Fumiko, Rakotonirina, Jean Claude, Hlaváč, Peter (2025): The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. X. Revision of the myrmecophilous clavigerine genera Rhynchoclaviger Wasmann, 1891 and Paussiger Wasmann, 1893. Zootaxa 5673 (3): 399-410, DOI: 10.11646/zootaxa.5673.3.4, URL: https://doi.org/10.11646/zootaxa.5673.3.4
381F87A1FFA9FFC7FF41FD185C872C86.text	381F87A1FFA9FFC7FF41FD185C872C86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paussiger limicornis (Wasmann 1893)	<div><p>Paussiger limicornis (Wasmann, 1893)</p><p>Paussiger limicornis Wasmann, 1893b: 257 (Type locality: Amparafaravantsiv [Amparafara, Moramanga prefecture, 6 km west of Perinet, 18.929°S, 48.394°E]. Type series: MNHN, probably 2 syntypes (Jeannel 1954: 304).</p><p>Paussiger limicornis Wasmann; Wasmann 1897: 264 (discussion); Raffray 1905: 466 (catalogue); Raffray 1908: 445 (catalogue); Raffray 1911: 178 (catalogue); Jeannel 1954: 304 (redescription); Jeannel 1960: 200 (distribution); Dajoz 1982: 514 (distribution).</p><p>Material studied. 2 ♀, one label: “VOH/ Aug.2012 /14 MADAGASCAR / Réserv. Expérimentale de VOHIMANA/ „ Circuit 3“; 23.viii.2012 / S18°55’02.3‘‘E48°30’21.2‘‘; 909m / sifting litter; Winkler app. extr.; L.S. / Rahanitriniaina &amp; E.M. Rabotoson lgt.“ [white, printed] (PCPH); 1 ♀, one label: “EAST MADAGASCAR / Tamatave [Toamasina] distr. / Andasibe, 17-30.XII.2001 / D. Hauck lgt.“ [white, printed] (PCPH); 1 ♀; one label: “ASB/ Nov 2011 MADAGASCAR / border of ANDASIBE N.P.; 27.xi.- / 3.xii.; FEON’NY ALA hotel surr. / sifting leaf litter, Winkler app. / extr.; L.S. Rahanitriniaina lgt.“ [white, printed] (PCPH).</p><p>Diagnosis. Antenna long, in females 0.40–0.45 mm long, terminal antennomere more than twice as long as wide; first visible paratergite not protruding laterally, distal lobe of aedeagus about 1.6 times as long as basal bulb.</p><p>Redescription. Body length about 2.7 mm. Head capsule rectangular, about 1.5 times as long as wide, slightly longer than pronotum, with four parallel carinae; lateral sides dentate, with about 5 small teeth located from under eyes to posterior margin of head capsule; anterior margin of rostrum triangular. Antenna of female 0.40–0.45 mm long, antennomere 3 confluent with 4, terminal antennomere about 0.3 mm long, slightly, evenly narrowed to truncate apex.</p><p>Pronotum about 1.3 times as wide as long, sides divergent posteriad, widest in posterior margin, with about 5 sharp teeth, with dentate median and lateral carinae; with wide, flat median sulcus bordered by discal carinae; lacking foveae.</p><p>Elytra 1.6–1.7 times as wide as long, shorter than abdomen, glabrous, with almost entire sutural striae, with three discal striae, adjacent to sutural striae, lacking humeral striae, lateral elytral margin non-dentate, with longitudinal impression; interstriae almost lacking punctures, very fine punctation only on anterior fourth.</p><p>Abdomen long, 1.6–1.7 times as long as elytra, anterior part of composite tergite with transverse impression, lateral foveae small, adjacent to posterior margin of elytra, with ill-defined trichome; first visible paratergite about as long as second, lacking lateral protrusion, with modest trichome, second visible paratergite parallel-sided, slightly longer than third visible paratergite, all paratergites slender.</p><p>Legs short and stout; mesofemur with sharp, long, outwards curved basal spine; mesotibia with apical, stout thorn (Jeannel 1954: 301, Fig. 159).</p><p>Aedeagus (Jeannel 1954: 303, Fig. 161) elongate, about 2.9 times as long as wide, apical lobe strongly convergent, about 1.6 times as long as basal bulb.</p><p>Sexual dimorphism. Females with mesofemora and mesotibia simple, lacking spines.</p><p>Host ant. Crematogaster hova -complex Blaimer, 2010 [as Crematogaster (Decacrema) schencki Forel, 1891].</p><p>Distribution. Madagascar.</p></div>	https://treatment.plazi.org/id/381F87A1FFA9FFC7FF41FD185C872C86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Raveloson, Ricca;Andrianaharison, Fumiko;Rakotonirina, Jean Claude;Hlaváč, Peter	Raveloson, Ricca, Andrianaharison, Fumiko, Rakotonirina, Jean Claude, Hlaváč, Peter (2025): The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. X. Revision of the myrmecophilous clavigerine genera Rhynchoclaviger Wasmann, 1891 and Paussiger Wasmann, 1893. Zootaxa 5673 (3): 399-410, DOI: 10.11646/zootaxa.5673.3.4, URL: https://doi.org/10.11646/zootaxa.5673.3.4
381F87A1FFA8FFC7FF41FED55C872AFD.text	381F87A1FFA8FFC7FF41FED55C872AFD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paussiger perrieri (Raveloson & Andrianaharison & Rakotonirina & Hlaváč 1900) Raveloson & Andrianaharison & Rakotonirina & Hlaváč 2025	<div><p>Paussiger perrieri (Fairmaire, 1898), comb. nov.</p><p>(Fig. 2)</p><p>Commatoceropsis perrieri Fairmaire, 1898: 344 (Type locality: Madagascar, Suberbieville [Maevatanana]. Type serie: no information in the original description, five syntypes should be located in MNHN (Jeannel 1954: 302).</p><p>Fusifer perrieri Fairmaire; Jeannel 1954: 302 (redescription); Jeannel 1960: 199 (in key, diagnosis, distribution).</p><p>Paussiger perrieri Fairmaire; Raffray 1908: 445 (catalogue).</p><p>Type material examined. LECTOTYPE: MADAGASCAR: ♂, here designated, seven labels: “Madag / illegible text“ [dirty white, handwritten], “ Commatoceropsis / Perrieri / illegible text Madag“ [dirty white, handwritten], “TYPE“ [red, printed], “ Fusifer / Perrieri F.“ [white, handwritten], “ Sufifer Newt. / = Fusifer Raffr. / perrieri Fairm. / Cl. Besuchet [white, handwritten and printed], “PHOTO (P. Krásenský) / P. Hlaváč, Prague“ [yellow, printed], “SYNTYPE / SUFIFER / perrieri (Fairmaire) / P. Hlaváč des., 2022“ [red, printed] (MNHN). Note. Lectotype with small red square label which means that a microscopic preparation of the aedeagus should be held in MNHN, it was not available for this study . PARALECTOTYPE: MADAGASCAR: 1 ♀, six labels: “Madagascar“ [white, handwritten], “ Commatoceropsis / Perrieri / illegible text Madag“ [dirty white, handwritten], “TYPE“ [red, printed], “ F. Perrieri / A. Raffray det.“ [white, handwritten and printed], “MUSÉUM PARIS / 1917 / COLL. A. RAFFRAY“ [bleu, printed], “SYNTYPE / SUFIFER / perrieri (Fairmaire) / P. Hlaváč des., 2022“ [red, printed] (MNHN) .</p><p>Diagnosis. Antenna short, about 0.35 mm long, terminal antennomere less than twice as long as wide; pronotum with ill-defined median sulcus, with well-defined median fovea; first visible paratergite strongly protruding laterally, distal lobe of aedeagus very short, more times shorter than basal bulb.</p><p>Redescription. Body length 2.1–2.2 mm. Head capsule sub-rectangular, about 1.2 times as long as wide, slightly longer than pronotum, with two sub-parallel carinae; lateral sides moderately dentate from under eyes to posterior margin of head capsule; anterior margin of rostrum triangular.Antenna 0.35 mm long, antennomere 3 confluent with 4, terminal antennomere about 0.2 mm long, from half of length slightly narrowed to truncate apex.</p><p>Pronotum about 1.15 times as wide as long, sides slightly convergent posteriad, widest in about midlength, with about 5 small teeth, lateral, short, slightly dentate carinae only in posterior half; lacking median carina, with ill-defined median sulcus, median fovea well-defined.</p><p>Elytra about 1.5 times as wide as long, shorter than abdomen, glabrous, lacking sutural striae, with eight carinae bearing setae, four sutural carinae reaching about half of elytral length, fifth reaching three fourth of elytral length and three lateral carinae entire, lateral elytral margin non-dentate, lacking longitudinal impression.</p><p>Abdomen long, about 1.6 times as long as elytra, anterior part of composite tergite with transverse impression, lateral foveae small with ill-defined trichome; first visible paratergite clearly longer than second, with well-defined lateral protrusion bearing well-defined trichome, second visible paratergite parallel-sided, about 1.3 times as long as third visible paratergite.</p><p>Legs short and stout; mesofemur with sharp, long, straight basal spine; mesotibia with small preapical spine.</p><p>Aedeagus known only from lateral view (Jeannel 1954: 303, Fig. 163), elongate, apical lobe strongly curved downwards, short; basal bulb about 3.3 times as long as apical lobe.</p><p>Sexual dimorphism. Females with mesofemora and mesotibiae simple, lacking spines.</p><p>Host ant. Crematogaster hova -complex Blaimer, 2010 [as Crematogaster (Decacrema) schencki Forel, 1891].</p><p>Distribution. Madagascar.</p></div>	https://treatment.plazi.org/id/381F87A1FFA8FFC7FF41FED55C872AFD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Raveloson, Ricca;Andrianaharison, Fumiko;Rakotonirina, Jean Claude;Hlaváč, Peter	Raveloson, Ricca, Andrianaharison, Fumiko, Rakotonirina, Jean Claude, Hlaváč, Peter (2025): The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. X. Revision of the myrmecophilous clavigerine genera Rhynchoclaviger Wasmann, 1891 and Paussiger Wasmann, 1893. Zootaxa 5673 (3): 399-410, DOI: 10.11646/zootaxa.5673.3.4, URL: https://doi.org/10.11646/zootaxa.5673.3.4
381F87A1FFA8FFC8FF41F8CA5C8728DF.text	381F87A1FFA8FFC8FF41F8CA5C8728DF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paussiger grandis (Raveloson & Andrianaharison & Rakotonirina & Hlaváč 1982) Raveloson & Andrianaharison & Rakotonirina & Hlaváč 2025	<div><p>Paussiger grandis (Dajoz, 1982), comb. nov.</p><p>(Fig. 3)</p><p>Marofustiger grandis Dajoz, 1982: 518 (Type locality: Madagascar Nord-Est, massif du Marojejy, 1450 m. Type serie: holotype ♂ in MNHN).</p><p>Type material examined. HOLOTYPE ♂, seven labels: “ Madagascar: Marojejy / 1450 m, 7/XII/1972 / J. M. Betsch leg.“ [blue, handwritten], “HOLOTYPE“ [red, printed], “ Muséum Paris“ [blue, printed], Marofustiger / grandis m. g., n. sp. / R. Dajoz det.1961“ [white, printed and handwritten], “ Paussiger / grandis Dajoz / Cl. Besuchet / dét. 2007“ [white, printed and handwritten], “PHOTO (P. Krásenský) / P. Hlaváč, Prague“ [yellow, printed], “ PAUSSIGER / = MAROFUSTIGER / grandis (Dajoz) / P. Hlaváč des., 2020“ [red, printed]. (MNHN).</p><p>Other material examined. 2 ♀, four labels: “ MADAGASCAR EST / Majorejy / Rés. Nat. Int. XII“ [white, printed], “1300 M / 8-XII-1972 / A. PEYRIERAS“ [white, printed] / FDHMA“ [white, printed], “ “ PAUSSIGER / = MAROFUSTIGER / grandis (Dajoz) / P. Hlaváč det., 2020“ [white, printed]. (MNHN) .</p><p>Diagnosis. Antenna long, about 0.44–0.46 mm long, terminal antennomere about 2.7 times as long as wide; pronotum with median sulcus, lacking median fovea; first visible paratergite strongly protruding laterally.</p><p>Redescription. Body length 2.65–2.80 mm. Head capsule rectangular, about 1.5 times as long as wide, slightly longer than pronotum, with two parallel lateral carinae; lateral sides dentate, with about eight sharp teeth located from under eyes to posterior margin of head capsule; anterior margin of rostrum triangular. Antenna 0.44-0.46 mm long, antennomere 3 well-separated from 4, terminal antennomere about 0.3 mm long, slightly, evenly narrowed to truncate apex.</p><p>Pronotum 1.15–1.20 times as wide as long, rounded on sides, with about 5–6 sharp teeth, with dentate median and lateral carinae; with wide, well-defined median sulcus bordered by discal carinae; lacking foveae.</p><p>Elytra 1.4–1.5 times as wide as long, shorter than abdomen, glabrous, with sutural striae reaching two third of elytral length, with five discal and one humeral striae, humeral stria and lateral elytral margin finely dentate; interstriae with small punctures, punctation especially dense close to striae.</p><p>Abdomen long, 1.6–1.7 times as long as elytra; anterior part of composite tergite with transverse impression; lateral foveae large, distant from posterior margin of elytra, with well-defined trichome; first visible paratergite longest, strongly protruding to side, protrusion rounded, with well-defined trichome, second visible paratergite longer than third visible paratergite, both narrowed distad.</p><p>Legs short and stout; mesofemur with sharp, long, outwards curved basal spine; mesotibia with apical, small spine.</p><p>Sexual dimorphism. Females with mesofemora and mesotibia simple, lacking spines.</p><p>Remarks. The aedeagus is still unknown, only one male, the holotype was not dissected to avoid its possible damage.</p><p>Host ant. Unknown, collected in ant nest on tree branch.</p><p>Distribution. Madagascar.</p></div>	https://treatment.plazi.org/id/381F87A1FFA8FFC8FF41F8CA5C8728DF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Raveloson, Ricca;Andrianaharison, Fumiko;Rakotonirina, Jean Claude;Hlaváč, Peter	Raveloson, Ricca, Andrianaharison, Fumiko, Rakotonirina, Jean Claude, Hlaváč, Peter (2025): The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. X. Revision of the myrmecophilous clavigerine genera Rhynchoclaviger Wasmann, 1891 and Paussiger Wasmann, 1893. Zootaxa 5673 (3): 399-410, DOI: 10.11646/zootaxa.5673.3.4, URL: https://doi.org/10.11646/zootaxa.5673.3.4
381F87A1FFA7FFC9FF41FAE75C872E9F.text	381F87A1FFA7FFC9FF41FAE75C872E9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paussiger fisheri Hlavac 2025	<div><p>Paussiger fisheri Hlaváč, new species</p><p>(Figs 4, 10, 11)</p><p>Material studied. HOLOTYPE: MADAGASCAR: ♂, two labels: “ MADAGASCAR, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=49.5&amp;materialsCitation.latitude=-14.75" title="Search Plazi for locations around (long 49.5/lat -14.75)">Befingotra</a> / (6.5 km SSW) <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=49.5&amp;materialsCitation.latitude=-14.75" title="Search Plazi for locations around (long 49.5/lat -14.75)">Rés. Anjanaharibe-Sud</a> / 14°45'S, 49°30'E, 875 m / rainforest, 19.X.1994 “ [white, printed], “FMNH #94-54, Winkler / extraction of sifted litter (leaf / mold, rotten wood), / B. L. / Fisher # 1070 (1-50) / FIELD MUS. NAT. HIST.“ [white, printed] (FMNH) . PARATYPES: MADAGASCAR: 1 ♂, 3 ♀, same data as for holotype (PCPH, NHMG); 2 ♂, two labels: “ MADAGASCAR: Antsiranana / <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=49.466667&amp;materialsCitation.latitude=-14.75" title="Search Plazi for locations around (long 49.466667/lat -14.75)">Befingotra</a> (9.2km WSW) <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=49.466667&amp;materialsCitation.latitude=-14.75" title="Search Plazi for locations around (long 49.466667/lat -14.75)">Rés. Anjanaharibe-Sud</a> / 14°45' S, 49°28'E, 1200m / 9.xi.1994 “ [white, printed], “FMHD #94-56, montane / rainforest, sifted litter / Winkler B. L. Fisher / BF#1990 / FIELD MUSEUM NAT.HIST“ [white, printed] (CASC, FMNH, MHNG, PHPC) .</p><p>Diagnosis. Antenna long, about 0.38–0.40 mm long, terminal antennomere about 2.1 times as long as wide; pronotum with well-defined median sulcus, median fovea ill-defined; first visible paratergite strongly protruding laterally, distal lobe of aedeagus slightly longer than basal bulb.</p><p>Redescription. Body length 2.45–2.65 mm. Head capsule rectangular, about 1.4–1.5 times as long as wide, slightly longer than pronotum, with five parallel lateral carinae; lateral sides dentate, with about 5-6 sharp teeth located from under eyes to posterior margin of head capsule; anterior margin of rostrum triangular. Antenna 0.38– 0.40 mm long, antennomere 3 well-separated from 4, terminal antennomere about 0.25 mm long, evenly narrowed to truncate apex.</p><p>Pronotum 1.15–1.20 times as wide as long, straight on sides, with about five to six sharp teeth, lacking median carinae, with dentate lateral carinae; with wide, well-defined median sulcus; lacking foveae.</p><p>Elytra 1.40–1.55 times as wide as long, shorter than abdomen, glabrous, with almost entire sutural striae, with five discal and one humeral stria, humeral stria and lateral elytral margin finely dentate; interstriae with small punctures, punctation evenly distributed between striae.</p><p>Abdomen long, 1.7–1.8 times as long as elytra; anterior part of composite tergite with transverse impression; lateral foveae large, distant from posterior margin of elytra, with well-defined trichome; first visible paratergite slightly shorter than second, strongly protruding to side, protrusion rounded, with well-defined trichome, second visible paratergite longest, clearly longer than third visible paratergite, both narrowed distad.</p><p>Legs short and stout; mesofemur with sharp, short, almost straight basal spine; mesotibia with small, apical spine.</p><p>Aedeagus (Figs 10, 11) elongate, about 0.62 mm long, about 2.9 times as long as wide, slightly asymmetrical; apical lobe slightly convergent distad, modestly rounded at apex, about 1.17 times as long as basal bulb; endophallus lacking internal sclerites, apical lobe laterally curved, with pointed apex.</p><p>Sexual dimorphism. Females with mesofemora and mesotibia simple, lacking spines.</p><p>Host ant. Unknown, all known specimens were collected by sifting of leaf, mold and rotten wood litter.</p><p>Distribution. Madagascar.</p></div>	https://treatment.plazi.org/id/381F87A1FFA7FFC9FF41FAE75C872E9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Raveloson, Ricca;Andrianaharison, Fumiko;Rakotonirina, Jean Claude;Hlaváč, Peter	Raveloson, Ricca, Andrianaharison, Fumiko, Rakotonirina, Jean Claude, Hlaváč, Peter (2025): The Pselaphinae (Coleoptera: Staphylinidae) of Madagascar. X. Revision of the myrmecophilous clavigerine genera Rhynchoclaviger Wasmann, 1891 and Paussiger Wasmann, 1893. Zootaxa 5673 (3): 399-410, DOI: 10.11646/zootaxa.5673.3.4, URL: https://doi.org/10.11646/zootaxa.5673.3.4
