identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9C12E3FC95BF56E0A67242AE8286C339.text	9C12E3FC95BF56E0A67242AE8286C339.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clitopilus fraxinicola T. Bau & Tian Y. Zhang 2026	<div><p>Clitopilus fraxinicola T. Bau &amp; Tian Y. Zhang sp. nov.</p><p>Figs 2 A, 2 B, 5</p><p>Etymology.</p><p>The specific epithet “ fraxinicola ” is derived from Latin fraxinus (ash tree) and - cola (dweller), referring to its growth on decaying branches of the Manchurian ash ( Fraxinus mandshurica Rupr.).</p><p>Holotype.</p><p>China. • Jilin Province, Jiaohe City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.71083&amp;materialsCitation.latitude=43.958057" title="Search Plazi for locations around (long 127.71083/lat 43.958057)">Qianjin Experimental Forest Farm</a>, 16 June 2025, 43°57'29"N, 127°42'39"E, alt. 347 m, Tolgor Bau and Long-Hao Li LH 2561602 (FJAU 75563) .</p><p>Diagnosis.</p><p>Basidiomata omphalinoid, signal white. Pileus centrally depressed; lamellae sparse; stipe central. Basidiospores subovoid to subfusiform with 6–8 longitudinal ridges. Gregarious on decaying branches of Fraxinus mandshurica in mixed forests.</p><p>Description.</p><p>Basidiomata omphalinoid. Pileus diameter 0.3–0.7 cm, initially hemispherical, later concave. When fresh, signal white (RAL 9003) to traffic white (RAL 9016); when dry, nearly white (RAL 1013). Pileus depressed at center, surface finely tomentose; margin entire, slightly incurved. Context thin, grayish white (RAL 9002); odor indistinct. Lamellae decurrent, white, sparse, unequally long. Stipe central, 0.7–2.1 cm long, 0.2 cm thick, subcylindrical, equal, solid, hyaline, signal white (RAL 9003), slightly curved in middle; surface smooth or finely tomentose; base with signal white (RAL 9003) mycelium.</p><p>Basidiospores.</p><p>[80 / 4 / 3] (5.9 –) 6.2–6.9 – 7.9 (– 8.3) × (3.2 –) 3.3–3.9 – 4.6 (– 4.9) µm, Q = 1.51–1.98, Q m = 1.75 (± 0.12), hyaline, subovoid to subfusiform or fusiform in profile and face view, obscurely angled in polar view with 6–8 facets produced by obscure longitudinal ridges (often indistinct under the light microscope), with minute transverse folds visible under SEM. Basidia 18–25 × 6–8 µm, clavate, 4 (2) - sterigmata. Cheilocystidia and pleurocystidia absent. Hymenophoral trama subregular, composed of subcylindrical hyphae 4–7 µm wide. Pileipellis a cutis, composed of interwoven, cylindrical, hyaline hyphae 5–8 µm in diameter, terminal cells 12–29 × 4–7 μm, subcylindrical, hyaline. Stipitipellis composed of repent, cylindrical hyphae 4–11 µm in diameter, interspersed with erect, hyaline hyphae 4–6 µm in diameter; terminal cells 15–39 × 5–8 μm, subcylindrical, hyaline, apex slightly acute or inflated. Clamp connections absent in all tissues.</p><p>Habitat.</p><p>Gregarious on dead wood or branches of Fraxinus mandshurica in mixed forests during summer.</p><p>Distribution.</p><p>Currently known only from Jilin Province, China.</p><p>Additional specimens examined.</p><p>China. • Jilin Province, Jiaohe City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.71083&amp;materialsCitation.latitude=43.958057" title="Search Plazi for locations around (long 127.71083/lat 43.958057)">Qianjin Experimental Forest Farm</a>, 29 June 2025, 43°57'29"N, 127°42'39"E, alt. 347 m, Hong Cheng C 25062905 (FJAU 75564), Tian-Yu Zhang ZTY 25062909 (FJAU 75565), Wei Sun SW 25062912 (FJAU 75567) .</p><p>Notes.</p><p>Based on its morphology and phylogenetic position, this species belongs to Clitopilus sect. Scyphoides . Morphologically, the present species resembles C. scyphoides (Fr.) Singer, but the latter possesses a larger pileus (0.5–3 cm) and longer basidiospores (6.5–8.5 × 3.5–5.0 µm) and typically grows on soil, allowing differentiation (Noordeloos 1984; Singer 1986; Bas et al. 1988). While sharing small basidiomata with C. subscyphoides W. Q. Deng et al. from Guangdong, China, the latter species differs in occurring on soil and being distributed in subtropical regions (Deng et al. 2013; Jian et al. 2023). Furthermore, it is distinguished from its five phylogenetically close relatives by key characteristics: from C. cretoalbus Izhar et al. by the latter’s possession of polymorphic cheilocystidia (subcylindrical, lageniform to filiform) and soil habitat (Izhar et al. 2023); from C. pirosylvanus Asif et al. by the latter’s pale brown pileus with a deep depression and a smaller spore Q value (Q m = 1.4) (Asif et al. 2025); from C. giovanellae (Bres.) Singer by the latter’s smooth basidiospores (Moreno et al. 2007); and from both C. zoysiae Kun L. Yang et al. and C. kamaka J. A. Cooper by their pleurotoid basidiomata with an eccentric or reduced stipe, with the latter species also recorded from Oceania (Cooper 2014; Yang et al. 2025).</p></div>	https://treatment.plazi.org/id/9C12E3FC95BF56E0A67242AE8286C339	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Tian-Yu;Bau, Tolgor	Zhang, Tian-Yu, Bau, Tolgor (2026): Seven new species of the Rhodocybe – Clitopilus clade (Entolomataceae, Agaricales) from Northeast China. MycoKeys 132: 273-306, DOI: 10.3897/mycokeys.132.181204
E721B6E9690C5E8A87C2891A786B2E27.text	E721B6E9690C5E8A87C2891A786B2E27.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lulesia variabilicolor T. Bau & Tian Y. Zhang 2026	<div><p>Lulesia variabilicolor T. Bau &amp; Tian Y. Zhang sp. nov.</p><p>Figs 2 C – E, 6</p><p>Etymology.</p><p>The epithet “ variabilicolor ” is a combination of Latin variabilis (variable) and color (color), alluding to the species’ highly variable pileus coloration.</p><p>Holotype.</p><p>China. • Jilin Province, Huadian City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.12&amp;materialsCitation.latitude=42.84889" title="Search Plazi for locations around (long 127.12/lat 42.84889)">Hongshi National Forest Park</a>, 16 August 2025, 42°50'56"N, 127°07'12"E, alt. 498 m, Tian-Yu Zhang, ZTY 2581617, (FJAU 75569) .</p><p>Diagnosis.</p><p>The pileus exhibits colors ranging from brownish gray to purple violet, is centrally slightly depressed. The stipe is central, cylindrical, and broadens downwards. The basidiospores are subglobose to broadly ellipsoid. The cheilocystidia are subfusiform with slightly curved apices. The hymenophoral trama is regular, and the pileipellis is a trichoderm.</p><p>Description.</p><p>Basidiomata collybioid. Pileus diameter 1.6–2.3 cm, initially hemispherical, later applanate. When fresh, center brown gray (RAL 7013), dark gray (RAL 7022), purple violet (RAL 4007) to blue-black (RAL 5004), margin yellow gray (RAL 7034), deep pigeon gray (RAL 7044), light gray (RAL 7035); when dry, light olive gray (RAL 7015). Pileus slightly depressed at center, covered in pruinose pubescence; margin entire, slightly incurved. Context thin, nearly white (RAL 1013), odor not distinctive. Lamellae decurrent, forking at stipe apex, gray white (RAL 9012), crowded, unequally long. Stipe central, 1.6–4.2 cm long, 0.7–1.1 cm thick, cylindrical, broadening downwards, hollow; apex concolorous with the lamellae, gradually turning to pearl light gray (RAL 9022) to pearl dark gray (RAL 9023), surface fibrillose, base slightly bulbous, with mycelium and rhizomorphs. Surfaces of dried basidiomata negative in KOH.</p><p>Basidiospores.</p><p>[60 / 3 / 2] (4.9 –) 5.1–5.4 – 5.8 (– 6.0) × (4.2 –) 4.3–4.6 – 4.9 (– 5.0) μm, Q = 1.06–1.28, Q m = 1.16 (± 0.06), hyaline, subglobose to broadly ellipsoid in profile view and face view, obscurely angled in polar view; ornamentation undulate-pustulate. Basidia 25–30 × 7–9 μm, clavate, 4 (1, 2) - sterigmata. Cheilocystidia 23–35 × 5–8 μm, subfusiform, apex slightly curved, rare. Hymenophoral trama regular, composed of tightly arranged, cylindrical hyphae 2–5 µm wide. Pileipellis a trichoderm, composed of cylindrical hyphae 3–6 μm in diameter; terminal cells 17–37 × 6–11 μm, clavate, bullet-shaped, fusoid, with pale brown pigment. Stipitipellis a compact layer of repent, cylindrical hyphae 3–7 μm in diameter, with pale grayish-brown pigment. Clamp connections absent in all tissues.</p><p>Habitat.</p><p>Growing singly or scattered in mixed forests during the summer and autumn seasons.</p><p>Distribution.</p><p>Currently known only from Jilin Province, China.</p><p>Additional specimens examined.</p><p>China. • Jilin Province, Huadian City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.12&amp;materialsCitation.latitude=42.84889" title="Search Plazi for locations around (long 127.12/lat 42.84889)">Hongshi National Forest Park</a>, 16 August 2024, 42°50'56"N, 127°07'12"E, alt. 498 m, Tolgor Bau and Ri-Hong Lin, LRH 24081621, (FJAU 75570) .</p><p>Notes.</p><p>This species was initially discovered in 2024, with basidiomata exhibiting purple tones and occurring singly. It was recollected in 2025, during which the basidiomata displayed gray tones. Micromorphological and DNA sequencing data confirmed the conspecificity of both collections. Morphologically, characterized by its pileus with gray to purple tones and subglobose to broadly ellipsoid basidiospores, this new species fits well within subg. Lulesia, which presently comprises 10 species (Vizzini et al. 2023 a). Although it shares a trichoderm pileipellis with L. densifolia (Singer) Singer, the latter is distinguished by its clitocyboid basidiomata, distinctly larger brown pileus (3–8 cm in diam.), and more slender stipe (2–8 × 0.5–0.8 cm) (Angelini and Contu 2012). While both the present species and L. colorata (L. Fan &amp; N. Mao) T. J. Baroni et al. show considerable variation in pileus color, L. colorata is characterized by a white to gray-brown pileus, an irregular hymenophoral trama, absent cystidia, and a cutis pileipellis (Mao et al. 2022). Phylogenetically, the present species is a sister taxon to L. umbrinomarginata Y. Q. Xiao et al.; however, the latter is clearly distinguished by its deeply depressed pileus center, a reaction of the dried basidiomata to KOH, an irregular lamellar trama, and its subtropical distribution (Xiao et al. 2024). Compared to L. nigrescens (Maire) Vizzini et al., the latter is clearly distinguished by its typically pleurotoid basidiomata, a cutis pileipellis, and a positive red reaction to KOH on dried specimens (Vizzini et al. 2023 b).</p></div>	https://treatment.plazi.org/id/E721B6E9690C5E8A87C2891A786B2E27	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Tian-Yu;Bau, Tolgor	Zhang, Tian-Yu, Bau, Tolgor (2026): Seven new species of the Rhodocybe – Clitopilus clade (Entolomataceae, Agaricales) from Northeast China. MycoKeys 132: 273-306, DOI: 10.3897/mycokeys.132.181204
2D0442D1DC2E5133A60AB7846358C943.text	2D0442D1DC2E5133A60AB7846358C943.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhodocybe eburneolutea T. Bau & T. Y. Zhang 2026	<div><p>Rhodocybe eburneolutea T. Bau &amp; T. Y. Zhang sp. nov.</p><p>Figs 3 J, 3 K, 10</p><p>Etymology.</p><p>The specific epithet “ eburneolutea ” refers to the ivory to yellowish coloration of the pileus, derived from the Latin ‘ eburneus ’ (ivory-colored) and ‘ luteus ’ (yellow).</p><p>Holotype.</p><p>China. • Jilin Province, Huadian City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.12&amp;materialsCitation.latitude=42.84889" title="Search Plazi for locations around (long 127.12/lat 42.84889)">Hongshi National Forest Park</a>, 23 August 2024, 42°50'56"N, 127°07'12"E, alt. 498 m, Tolgor Bau and Xian-Yan Zhou, Y 2482316 (FJAU 75556) .</p><p>Diagnosis.</p><p>The pileus is ivory to beige, centrally depressed, and brown beige. The pileus surface is smooth and translucent striate. The basidiospores are subellipsoid to obscurely angular, ornamented with bumpy undulate-pustulate. The pseudocystidia are abundant, predominantly subfusoid to rostrate, rarely branched, and non-dextrinoid. Both the hymenophoral trama and the pileipellis are non-dextrinoid.</p><p>Description.</p><p>Basidiomata clitocyboid. Pileus diameter 0.4–1.1 cm, initially hemispherical, later applanate, depressed at center; when fresh beige (RAL 1001) to ivory (RAL 1014), center brown beige (RAL 1011); when dry ochre brown (RAL 8001), clay brown (RAL 8003). Pileus surface smooth, translucent striate; margin entire. Context thin, off-white (RAL 1013); odor indistinct or not distinctive. Lamellae adnate to subdecurrent, off-white (RAL 1013) to bright ivory (RAL 1015), slightly crowded, unequally long, fragile. Stipe central, 0.7–1.4 cm long, 0.2 cm thick, cylindrical, equal, hollow, ivory (RAL 1014) to yellow ochre (RAL 1024), slightly flexuous; surface smooth.</p><p>Basidiospores.</p><p>[80 / 4 / 3] (5.3 –) 5.5–6.0 – 6.6 (– 6.7) × (3.9 –) 4.0–4.4 – 4.7 (– 5.0) μm, Q = 1.20–1.49, Q m = 1.37 (± 0.08), hyaline, subellipsoid and somewhat rounded angular in profile view, broadly ellipsoid in face view, obscurely angular in polar view, ornamentation strongly bumpy undulate-pustulate. Basidia 20–25 × 6–7 μm, clavate, 4 (2) - sterigmata. Cheilocystidia (pseudocystidia) 23–41 × 4–7 μm, subclavate, narrowly lageniform, with a rarely branched apex, with golden pearl contents, non-dextrinoid. Pleurocystidia (pseudocystidia) 18–36 × 4–7 μm, flexuous, subcylindrical, subfusiform, with golden pearl contents, non-dextrinoid. Hymenophoral trama regular, composed of cylindrical hyphae 4–10 μm in diameter, non-dextrinoid. Pileipellis a cutis, composed of cylindrical hyphae 5–7 μm in diameter, with pale yellowish-brown pigment, non-dextrinoid. Stipitipellis composed of repent, cylindrical hyphae 3–5 μm in diameter, with yellowish-brown pigment. Clamp connections absent in all tissues.</p><p>Habitat.</p><p>Gregarious on the ground in mixed forests during summer.</p><p>Distribution.</p><p>Currently known only from Jilin Province, China.</p><p>Additional specimens examined.</p><p>China. • Jilin Province, Huadian City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.12&amp;materialsCitation.latitude=42.84889" title="Search Plazi for locations around (long 127.12/lat 42.84889)">Hongshi National Forest Park</a>, 23 August 2024, 42°50'56"N, 127°07'12"E, alt. 498 m, Tian-Yu Zhang, ZTY 2482307 (FJAU 75557) ; • Same location, 15 September 2025, Yu-Fei Han, F 25915027 (FJAU 75571) .</p><p>Notes.</p><p>Morphologically, R. eburneolutea is similar to R. asyae E. Sesli &amp; Vizzini from Turkey, as both possess small, clitocyboid basidiomata with an ivory-colored pileus. However, the latter differs in lacking striations on the pileus surface, lacking pseudocystidia, and in its classification within sect. Rufobrunnea, allowing clear distinction (Sesli and Vizzini 2017). R. eburneolutea also shares a yellowish pileus and a slender stipe with R. zijinshanensis S. P. Jian &amp; X. Chen from Nanjing, China, but R. zijinshanensis can be distinguished by its complete absence of hymenial cystidia and its distinct ecology of growing solitarily on rotten wood in broad-leaved forests (Jian et al. 2025).</p><p>Phylogenetically, R. eburneolutea is sister to R. caelata (Fr.) Maire (rpb 2: KC 816934). However, Kluting et al. (2014) have demonstrated that R. caelata as presently understood is polyphyletic with ambiguous species boundaries. This study reveals significant morphological differences between the new species and the polyphyletic R. caelata: the latter has more robust basidiomata, a grayish pileus that is velutinous and lacks translucent striations, larger basidiospores (7.0–9.0 × 4.0–5.0 µm, Q m = 1.75), and pseudocystidia that are mostly cylindrical to clavate (Baroni 1981; Bas et al. 1988). R. eburneolutea is also phylogenetically close to R. pakistanica Z. Khan &amp; Khalid from Pakistan; nevertheless, R. pakistanica differs in its dark reddish-brown pileus, a stipe surface covered with white fibrils, and scarce pseudocystidia (Khan and Khalid 2024).</p></div>	https://treatment.plazi.org/id/2D0442D1DC2E5133A60AB7846358C943	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Tian-Yu;Bau, Tolgor	Zhang, Tian-Yu, Bau, Tolgor (2026): Seven new species of the Rhodocybe – Clitopilus clade (Entolomataceae, Agaricales) from Northeast China. MycoKeys 132: 273-306, DOI: 10.3897/mycokeys.132.181204
AEAD85330B635D58AB2D4D186EFA0661.text	AEAD85330B635D58AB2D4D186EFA0661.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhodocybe galericulata T. Bau & T. Y. Zhang 2026	<div><p>Rhodocybe galericulata T. Bau &amp; T. Y. Zhang sp. nov.</p><p>Figs 3 I, 9</p><p>Etymology.</p><p>The specific epithet “ galericulata ” is a Latin noun meaning “ a small hat ” or “ cap, ” referring to the characteristic bowler hat-like shape of the pileus, caused by its depressed center and raised surface.</p><p>Holotype.</p><p>China. • Heilongjiang Province, Harbin City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=124.69639&amp;materialsCitation.latitude=45.774166" title="Search Plazi for locations around (long 124.69639/lat 45.774166)">Guli Park</a>, 24 July 2023, 45°46'27"N, 124°41'47"E, alt. 128 m, Li-Yang Zhu, Z 23072405 (FJAU 75558) .</p><p>Diagnosis.</p><p>Pileus chocolate brown to deep brown, centrally depressed with raised disc, resembling a bowler hat. Lamellae adnate to subdecurrent. Basidiospores subellipsoid to amygdaliform with undulate-pustulate ornamentation. Pseudocystidia clavate, subcylindrical to undulate in outline, with curry contents.</p><p>Description.</p><p>Basidiomata collybioid. Pileus diameter 0.6–1.0 cm, initially hemispherical to convex, later slightly applanate, eventually depressed at center, with area around the disc raised resembling a bowler hat; when fresh deep brown (RAL 8011), chestnut brown (RAL 8015), chocolate brown (RAL 8017); when dry blackish brown (RAL 8022), light grayish brown (RAL 8028). Pileus surface smooth; margin entire, slightly incurved. Context thin, soil brown (RAL 8003); odor indistinct. Lamellae adnate to subdecurrent, nearly white (RAL 1013), slightly crowded, unequally long. Stipe central, 1.2–1.7 cm long, 0.2 cm thick, cylindrical, equal, hollow, grayish brown (RAL 7013) to light brown (RAL 8025); surface smooth, base with signal white (RAL 9003) mycelium.</p><p>Basidiospores.</p><p>[80 / 4 / 2] (5.4 –) 5.6–6.1 – 6.8 (– 7.0) × (3.9 –) 4.0–4.5 – 4.9 (– 5.0) μm, Q = 1.22–1.56, Q m = 1.36 (± 0.08), hyaline, subellipsoid to amygdaliform in profile view, broadly ellipsoid in face view, obscurely angular in polar view, with undulate-pustulate ornamentation. Basidia 24–30 × 6–8 μm, clavate, 4 (2) - sterigmata. Cheilocystidia (pseudocystidia) 20–45 × 3–8 μm, clavate, subcylindrical to undulate in outline, with curry contents, non-dextrinoid. Pleurocystidia (pseudocystidia) similar to cheilocystidia in morphology, dimensions, and contents. Hymenophoral trama subregular, composed of cylindrical hyphae 3–7 μm in diameter, non-dextrinoid. Pileipellis a cutis, composed of interwoven cylindrical hyphae 5–9 μm in diameter, with brown pigment, non-dextrinoid. Stipitipellis composed of repent, cylindrical hyphae 3–7 μm in diameter, with pale yellowish-brown pigment. Clamp connections absent in all tissues.</p><p>Habitat.</p><p>Gregarious on soil in broad-leaved forests during summer.</p><p>Distribution.</p><p>Currently known only from Heilongjiang Province, China.</p><p>Additional specimens examined.</p><p>China. • Heilongjiang Province, Harbin City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=124.69639&amp;materialsCitation.latitude=45.774166" title="Search Plazi for locations around (long 124.69639/lat 45.774166)">Guli Park</a>, 24 July 2023, 45°46'27"N, 124°41'47"E, alt. 128 m, Wei-Nan Hou, H 2307142 (FJAU 75559) .</p><p>Notes.</p><p>R. galericulata has pseudocystidia with curry contents and is morphologically classified within sect. Rhodocybe . While R. galericulata is macroscopically similar to R. brunnea Contu due to its brown pileus, the latter is distinguished by its larger basidiomata, a pileus ranging from pale yellowish to brown without a central depression, fusoid to lageniform pseudocystidia, and a gregarious habit on humus of Cupressus sempervirens L. (Contu 2006). R. galericulata also shares a brownish, centrally depressed pileus with R. fuliginea E. Horak; however, the latter differs in its densely crowded lamellae, a minutely velutinous to tomentose pileus surface, and more rounded, ovoid basidiospores (6.0–7.5 × 4.5–6.0 μm) (Horak 1979). Compared to its phylogenetically close relative R. collybioides Singer, the latter is distinguished by its slightly depressed pileus, solid stipe, and lanceolate-fusoid pseudocystidia with acute apices (Vrinda et al. 1999).</p></div>	https://treatment.plazi.org/id/AEAD85330B635D58AB2D4D186EFA0661	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Tian-Yu;Bau, Tolgor	Zhang, Tian-Yu, Bau, Tolgor (2026): Seven new species of the Rhodocybe – Clitopilus clade (Entolomataceae, Agaricales) from Northeast China. MycoKeys 132: 273-306, DOI: 10.3897/mycokeys.132.181204
5C921ADEACB955D0AFE6EC254B5037C5.text	5C921ADEACB955D0AFE6EC254B5037C5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhodocybe jilinensis T. Bau & Tian Y. Zhang 2026	<div><p>Rhodocybe jilinensis T. Bau &amp; Tian Y. Zhang sp. nov.</p><p>Figs 2 F, 7</p><p>Etymology.</p><p>The specific epithet “ jilinensis ” refers to Jilin Province, China, the origin of the type collection.</p><p>Holotype.</p><p>China. • Jilin Province, Ji’an City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.19195&amp;materialsCitation.latitude=41.138336" title="Search Plazi for locations around (long 126.19195/lat 41.138336)">Yushan Park</a>, 24 July 2025, 41°08'18"N, 126°11'31"E, alt. 216 m, Long-Hao Li, LH 2572405 (FJAU 75554) .</p><p>Diagnosis.</p><p>The basidiomata are omphalinoid. The pileus is light grayish brown to violet blue, centrally slightly depressed, with a velutinous surface. The lamellae are subdecurrent and white aluminium. The basidiospores are subellipsoid to subpyriform with strongly bumpy, undulate-pustulate ornamentation. The pseudocystidia are subcylindrical to subfusoid and non-dextrinoid.</p><p>Description.</p><p>Basidiomata omphalinoid. Pileus diameter 0.5–1.4 cm, initially hemispherical, later slightly applanate; when fresh violet blue (RAL 4009), dove gray (RAL 7032), platinum gray (RAL 7036); when dry chocolate brown (RAL 8017), light grayish brown (RAL 8028). Pileus slightly depressed at center, surface velutinous, non-striate; margin entire, slightly incurved. Context thin, deep brown (RAL 8011); odor indistinct or not distinctive. Lamellae subdecurrent, white aluminium (RAL 9006), pearl dark gray (RAL 9023), slightly crowded, unequally long. Stipe central, 1.4–2.9 cm long, 0.2–0.4 cm thick, cylindrical, equal or unequal, hollow, platinum gray (RAL 7036), slightly flexuous; surface fibrillose, base with signal white (RAL 9003) mycelium.</p><p>Basidiospores.</p><p>[60 / 3 / 2] (5.9 –) 6.1–6.5 – 7.0 (– 7.2) × (4.6 –) 4.9–5.0 – 5.2 (– 5.6) μm, Q = 1.16–1.36, Q m = 1.29 (± 0.05), hyaline, subellipsoid to subpyriform in profile view, broadly ellipsoid in face view, distinctly angular in polar view, ornamentation strongly bumpy undulate-pustulate. Basidia 26–33 × 7–9 μm, clavate, 4 (2) - sterigmata. Cheilocystidia (pseudocystidia) 27–43 × 4–7 μm, subcylindrical to subfusoid, with clay brown contents, non-dextrinoid. Pleurocystidia (pseudocystidia) similar to cheilocystidia in morphology, dimensions, and contents. Hymenophoral trama subregular, composed of cylindrical hyphae 5–7 μm in diameter, non-dextrinoid. Pileipellis a cutis with transitions to a trichoderm, composed of cylindrical hyphae 5–9 μm in diameter, with yellowish-brown to brown pigment, non-dextrinoid. Stipitipellis composed of repent, cylindrical hyphae 5–13 μm in diameter, with pale grayish-brown pigment. Clamp connections absent in all tissues.</p><p>Habitat.</p><p>Gregarious to scattered on the ground in mixed forests during summer.</p><p>Distribution.</p><p>Currently known only from Jilin Province, China.</p><p>Additional specimens examined.</p><p>China. • Jilin Province, Ji’an City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.19195&amp;materialsCitation.latitude=41.138336" title="Search Plazi for locations around (long 126.19195/lat 41.138336)">Yushan Park</a>, 24 July 2025, 41°08'18"N, 126°11'31"E, alt. 216 m, Tolgor Bau and Yu Wang, WY 2572413 (FJAU 75555) .</p><p>Notes.</p><p>R. jilinensis has abundant pseudocystidia and is classified in Rhodocybe sect. Rhodocybe . R. caelata (Fr.) Maire, the type species of the genus, shares a grayish pileus and abundant pseudocystidia with R. jilinensis . However, R. caelata is distinguished by its pileus, which often becomes concentrically areolate-rimose, and by its larger basidiospores (7.0–9.0 × 4.0–5.0 µm, Q m = 1.75), which are ellipsoid to oblong (Baroni 1981; Bas et al. 1988). R. jilinensis is macroscopically similar to R. griseonigrella (Vila, Contu, F. Caball. &amp; Ribes) Vizzini et al. and can be easily confused since both species share small basidiomata with gray pileus and lamellae; however, the latter differs in subglobose basidiospores and a dextrinoid reaction in both the hymenium and pileipellis in Melzer’s reagent (Vila et al. 2009; Vizzini et al. 2016). Rhodocybe brunnescens T. J. Baroni &amp; E. Horak is another comparable species, but it can be readily separated from R. jilinensis by its lamellae, which turn brown on bruising, and by its larger basidiospores (6.5–9.5 × 4.5–5.5 µm) (Baroni and Horak 1994).</p><p>Phylogenetically, R. jilinensis is closely related to R. spongiosa T. J. Baroni et al.; however, the latter is distinguished by its pleurotoid basidiomata, white coloration, and the absence of pigment in the pileipellis hyphae (Henkel et al. 2010). Compared to R. fuliginea E. Horak, the latter possesses a deeply umbilicate pileus, densely crowded lamellae, basidiospores that are distinctly rugulose to subangular, and acute-fusoid pseudocystidia (Horak 1979).</p></div>	https://treatment.plazi.org/id/5C921ADEACB955D0AFE6EC254B5037C5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Tian-Yu;Bau, Tolgor	Zhang, Tian-Yu, Bau, Tolgor (2026): Seven new species of the Rhodocybe – Clitopilus clade (Entolomataceae, Agaricales) from Northeast China. MycoKeys 132: 273-306, DOI: 10.3897/mycokeys.132.181204
67C6E162D421565FB6F13D3D45F2AB1F.text	67C6E162D421565FB6F13D3D45F2AB1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhodocybe subnuciolens T. Bau & Tian Y. Zhang 2026	<div><p>Rhodocybe subnuciolens T. Bau &amp; Tian Y. Zhang sp. nov.</p><p>Figs 3 G, 3 H, 8</p><p>Etymology.</p><p>The specific epithet “ subnuciolens ” refers to its morphological similarity to R. nuciolens .</p><p>Holotype.</p><p>China. • Jilin Province, Huadian City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.12&amp;materialsCitation.latitude=42.84889" title="Search Plazi for locations around (long 127.12/lat 42.84889)">Hongshi National Forest Park</a>, 15 August 2025, 42°50'56"N, 127°07'12"E, alt. 498 m, Tian-Yu Zhang, ZTY 2581517, (FJAU 75553) .</p><p>Diagnosis.</p><p>Morphologically similar to R. nuciolens, but distinguished by its smaller and more slender basidiomata, smaller basidiospores, and more crowded lamellae.</p><p>Description.</p><p>Basidiomata collybioid. Pileus diameter 1.2–3.4 cm, initially convex, later applanate; when fresh grayish white (RAL 9012); when dry beige (RAL 1001). Pileus centrally slightly depressed or non-depressed; surface smooth, sometimes hygrophanous, non-striate; margin entire. Context slightly thin, sand yellow (RAL 1002); odor indistinct. Lamellae adnate to subdecurrent, off-white (RAL 1013), slightly crowded, unequally long. Stipe central, 1.7–2.9 cm long, 0.2–0.4 cm thick, cylindrical, equal, solid, grayish white (RAL 9012); surface smooth, basal signal white (RAL 9003), with mycelium and rhizomorphs.</p><p>Basidiospores.</p><p>[100 / 5 / 3] (4.8 –) 5.4–5.6 – 6.3 (– 6.5) × (3.3 –) 3.5–3.9 – 4.2 (– 4.5) μm, Q = 1.26–1.59, Q m = 1.45 (± 0.08), hyaline, subellipsoid or subamygdaliform in profile view, broadly ellipsoid in face view, minutely angular in polar view, undulate-pustulate in all views. Basidia 19–25 × 5–7 μm, clavate, 4 (2) - sterigmata. Pseudocystidia absent. Cheilocystidia 17–33 × 3–5 μm, subcylindrical, often with bent or rarely branched apex, irregular, non-dextrinoid. Pleurocystidia absent. Hymenophoral trama regular, composed of cylindrical hyphae 7–12 μm in diameter, non-dextrinoid. Pileipellis a cutis, composed of thin-walled, hyaline, interwoven hyphae 6–9 μm in diameter, non-dextrinoid. Stipitipellis composed of repent, cylindrical, hyaline hyphae 5–8 μm in diameter. Clamp connections absent in all tissues.</p><p>Habitat.</p><p>Solitary or scattered on the ground in mixed forests during summer.</p><p>Distribution.</p><p>Currently known only from Jilin Province, China.</p><p>Additional specimens examined.</p><p>China. • Jilin Province, Huadian City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.12&amp;materialsCitation.latitude=42.84889" title="Search Plazi for locations around (long 127.12/lat 42.84889)">Hongshi National Forest Park</a>, 16 August 2025, 42°50'56"N, 127°07'12"E, alt. 498 m, Tian-Yu Zhang, ZTY 2581616, (FJAU 75551) ; • Same location, 31 July 2025, Tian-Yu Zhang, ZTY 2573118, (FJAU 75552) .</p><p>Notes.</p><p>Rhodocybe subnuciolens belongs to sect. Rufobrunnea. Compared with this species, R. nuciolens has larger basidiomata with a pileus diameter of 3.5–8 cm, more distant lamellae, larger basidiospores (5.5–8 × 4–5 μm), and is currently known only from North America (Murrill 1913). The basidiomata color of R. subnuciolens is close to that of R. subasyae T. Bau &amp; Y. L. Sun, but the latter has a reddish tinge on the pileus, sinuate lamellae, and a two-layered pileipellis composed of short-cylindrical and inflated cylindrical hyphae. Furthermore, a significant genetic distance separates the two species, providing a clear basis for distinction (Sun and Bau 2023).</p><p>Phylogenetically, R. subnuciolens is a sister taxon to R. pseudopiperita T. J. Baroni &amp; G. M. Gates, but the latter is distinguished by its lack of hymenial cystidia and its distribution in Oceania (Baroni and Gates 2006). The other three close relatives ( R. alutacea Singer, R. dominicana T. J. Baroni &amp; Angelini, and R. pseudoalutacea T. J. Baroni et al.) are all distributed in North America (Baroni et al. 2020). Among them, R. alutacea also possesses a pale pileus, but hymenial cystidia are absent; R. dominicana not only lacks hymenial cystidia but also has a two-layered pileipellis composed of cylindrical and subglobose hyphae; and R. pseudoalutacea has a brownish-orange pileus, lacks hymenial cystidia, and possesses caulocystidia.</p></div>	https://treatment.plazi.org/id/67C6E162D421565FB6F13D3D45F2AB1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Tian-Yu;Bau, Tolgor	Zhang, Tian-Yu, Bau, Tolgor (2026): Seven new species of the Rhodocybe – Clitopilus clade (Entolomataceae, Agaricales) from Northeast China. MycoKeys 132: 273-306, DOI: 10.3897/mycokeys.132.181204
C0ED5D1A4ADE5E058A68888577746604.text	C0ED5D1A4ADE5E058A68888577746604.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhodophana hongshiensis T. Bau & Tian Y. Zhang 2026	<div><p>Rhodophana hongshiensis T. Bau &amp; Tian Y. Zhang sp. nov.</p><p>Figs 3 L, 11</p><p>Etymology.</p><p>The specific epithet “ hongshiensis ” refers to Hongshi National Forest Park, the origin of the type collection.</p><p>Holotype.</p><p>China. • Jilin Province, Huadian City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.12&amp;materialsCitation.latitude=42.84889" title="Search Plazi for locations around (long 127.12/lat 42.84889)">Hongshi National Forest Park</a>, 22 September 2024, 42°50'56"N, 127°07'12"E, alt. 498 m, Wei-Nan Hou, H 2492218, (FJAU 75561) .</p><p>Diagnosis.</p><p>The pileus light orange-yellow to broom yellow, centrally depressed, smooth, with translucent striations at margin. Lamellae adnexed, slightly crowded. Stipe smooth. Basidiospores ellipsoid to amygdaliform with pronounced undulate-pustulate ornamentation.</p><p>Description.</p><p>Basidiomata collybioid. Pileus diameter 0.4–0.9 cm, initially hemispherical, later convex. When fresh, light orange-yellow (RAL 1028), broom yellow (RAL 1032); when dry, ochre-yellow (RAL 1024). Pileus depressed at center, surface smooth; margin with translucent striations, slightly recurved. Context thin, ivory (RAL 1014); odor indistinct. Lamellae adnexed, sandy yellow (RAL 1002), ventricose, slightly crowded, unequally long. Stipe central, 1.6–2.7 cm long, 0.2–0.3 cm thick, subcylindrical, soil-brown (RAL 8003), slightly curved, solid; surface smooth.</p><p>Basidiospores.</p><p>[40 / 2 / 2] (6.3 –) 6.4–6.7 – 7.5 (– 7.7) × (3.9 –) 4.1–4.3 – 4.6 (– 4.7) μm, Q = 1.35–1.77, Q m = 1.57 (± 0.10), hyaline, ellipsoid to amygdaliform in profile view, ellipsoid in face view, obscurely angular in polar view, with undulate-pustulate surface ornamentation. Basidia 22–28 × 8–9 μm, clavate, 4 (2) - sterigmata. Cheilocystidia and pleurocystidia absent. Hymenophoral trama regular, composed of cylindrical hyphae 9–18 μm in diameter. Pileipellis a cutis, composed of cylindrical hyphae 9–16 μm in diameter, with pale yellowish pigment; terminal cells 38–77 × 12–23 μm, cylindrical, subfusoid. Stipitipellis composed of repent, cylindrical hyphae 6–11 μm in diameter, with yellowish-brown pigment. Clamp connections present in all tissues.</p><p>Habitat.</p><p>Solitary or scattered on the ground in mixed forests during summer.</p><p>Distribution.</p><p>Currently known only from Jilin Province, China.</p><p>Additional specimens examined.</p><p>China. • Jilin Province, Huadian City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.12&amp;materialsCitation.latitude=42.84889" title="Search Plazi for locations around (long 127.12/lat 42.84889)">Hongshi National Forest Park</a>, 21 September 2024, 42°50'56"N, 127°07'12"E, alt. 498 m, Tolgor Bau and Tian-Yu Zhang, ZTY 2492107, (FJAU 75562) .</p><p>Notes.</p><p>Morphologically, Rho. hongshiensis closely resembles Rho. nitellina (Fr.) Papetti in its orange-toned pileus with translucent striations. However, the latter can be distinguished by its significantly larger pileus (40–50 mm diam.) and larger basidiospores measuring 7.5–10 × 4.5–5.6 μm (Bas et al. 1988). Within the Rho. nitellina complex, a small-sized taxon, Rho. nitellina var. minor (Fr.) Papetti, has been documented. According to the taxonomic revision by Papetti, this entity was not elevated to species rank but was transferred from its previous placements in Agaricus and Collybia into Rhodophana, establishing the new combination Rho. nitellina var. minor (Fr.) Papetti. Compared to Rho. hongshiensis, Rho. nitellina var. minor possesses more robust basidiomata, a pileus with a more reddish-brown tinge, and grows gregariously on the ground in coniferous forests, allowing clear differentiation (Papetti 2014).</p><p>Rho. hongshiensis is phylogenetically closely related to Rho . almae - lunae Papetti; however, the latter can be distinguished by its basidiospores with rounded pustules and its primary distribution in coniferous forests (on litter of Pinus or Cedrus) (Papetti 2024). Compared to Rho. melleopallens (P. D. Orton) Kluting et al., the latter has larger basidiomata and smaller basidiospores (4.5–7.0 × 3–4 μm, Q = 1.3–1.4) (Bas et al. 1988). Rho. hongshiensis and Rho . aershanensis J. Z. Xu from the Inner Mongolia Autonomous Region of China both possess small basidiomata. However, the latter has a reddish-brown pileus and smaller basidiospores (2.7–5.8 × 2.2–5.1 μm, Q m = 1.1) with slightly undulate-pustulate surfaces (Xu et al. 2023).</p></div>	https://treatment.plazi.org/id/C0ED5D1A4ADE5E058A68888577746604	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Tian-Yu;Bau, Tolgor	Zhang, Tian-Yu, Bau, Tolgor (2026): Seven new species of the Rhodocybe – Clitopilus clade (Entolomataceae, Agaricales) from Northeast China. MycoKeys 132: 273-306, DOI: 10.3897/mycokeys.132.181204
