taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
917FE7E338745D0D8D2ECC9B93261F83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601909	https://doi.org/10.3897/asp.84.e182696.figure3	Figure 3. Acanthocephala spp., dorsal habitus of type species and labels. A A. (Acanthocephala) latipes (Drury, 1782), male syntype, photographed by Laurence Livermore (© Zoologisk museum i København), original photo cropped and contrast adjusted, made available by Natural History Museum of Denmark Zoological Museum of Copenhagen under Creative Commons Attribution 3.0, International Public License, CC BY-NC 3.0; B A. (Spinipedia) heissi Brailovsky, 2006, male paratype; C A. (Contrastata) thomasii (Uhler, 1872), male syntype; D A. (Pronoptera) alata (Burmeister, 1835), male syntype, photographed by Laurence Livermore, original photo cropped and contrast adjusted; E A. (Metapodiessa) femorata (Fabricius, 1775), male syntype, original photo cropped and contrast adjusted (© The Trustees of the Natural History Museum, London (Banks collection )). Scale bar: 10 mm (A – E).	Figure 3. Acanthocephala spp., dorsal habitus of type species and labels. A A. (Acanthocephala) latipes (Drury, 1782), male syntype, photographed by Laurence Livermore (© Zoologisk museum i København), original photo cropped and contrast adjusted, made available by Natural History Museum of Denmark Zoological Museum of Copenhagen under Creative Commons Attribution 3.0, International Public License, CC BY-NC 3.0; B A. (Spinipedia) heissi Brailovsky, 2006, male paratype; C A. (Contrastata) thomasii (Uhler, 1872), male syntype; D A. (Pronoptera) alata (Burmeister, 1835), male syntype, photographed by Laurence Livermore, original photo cropped and contrast adjusted; E A. (Metapodiessa) femorata (Fabricius, 1775), male syntype, original photo cropped and contrast adjusted (© The Trustees of the Natural History Museum, London (Banks collection )). Scale bar: 10 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
917FE7E338745D0D8D2ECC9B93261F83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601910	https://doi.org/10.3897/asp.84.e182696.figure4	Figure 4. Acanthocephala spp., head. A Dorsal view, A. (Acanthocephala) latipes (Drury, 1782); B – F Lateral view. B A. (Acanthocephala) latipes; C A. (Spinipedia) heissi Brailovsky, 2006; D A. (Contrastata) thomasii (Uhler, 1872); E A. (Pronoptera) alata (Burmeister, 1835); F A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: bu, buccula; ju, juga; mxp, maxillary plate; ty, tylus. Asterisks indicate the apex of the tylus above (char. 26: 1) and below (char. 26: 0) to the longitudinal axis of the head in distal view. Scale bar: 1 mm (A – F).	Figure 4. Acanthocephala spp., head. A Dorsal view, A. (Acanthocephala) latipes (Drury, 1782); B – F Lateral view. B A. (Acanthocephala) latipes; C A. (Spinipedia) heissi Brailovsky, 2006; D A. (Contrastata) thomasii (Uhler, 1872); E A. (Pronoptera) alata (Burmeister, 1835); F A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: bu, buccula; ju, juga; mxp, maxillary plate; ty, tylus. Asterisks indicate the apex of the tylus above (char. 26: 1) and below (char. 26: 0) to the longitudinal axis of the head in distal view. Scale bar: 1 mm (A – F).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
917FE7E338745D0D8D2ECC9B93261F83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601911	https://doi.org/10.3897/asp.84.e182696.figure5	Figure 5. Acanthocephalini spp. A – D Dorsal view of pronotum. A Ctenomelynthus coxalis Breddin, 1903; B Acanthocephala (Acanthocephala) latipes (Drury, 1782); C A. (Pronoptera) panamensis Distant, 1881; D A. (Metapodiessa) femorata (Fabricius, 1775); E Caudal view of pronotum, A. (Pronoptera) panamensis; F Spines of the costal margin of hemelytra, A. (Pronoptera) panamensis. Abbreviations: alm, anterolateral margin of pronotum; cm, costal margin of hemelytra; ha, humeral angles of pronotum; has, spine of humeral angles of pronotum; plm, posterolateral margin of pronotum. Scale bars: 1 mm (A); 2 mm (B – F).	Figure 5. Acanthocephalini spp. A – D Dorsal view of pronotum. A Ctenomelynthus coxalis Breddin, 1903; B Acanthocephala (Acanthocephala) latipes (Drury, 1782); C A. (Pronoptera) panamensis Distant, 1881; D A. (Metapodiessa) femorata (Fabricius, 1775); E Caudal view of pronotum, A. (Pronoptera) panamensis; F Spines of the costal margin of hemelytra, A. (Pronoptera) panamensis. Abbreviations: alm, anterolateral margin of pronotum; cm, costal margin of hemelytra; ha, humeral angles of pronotum; has, spine of humeral angles of pronotum; plm, posterolateral margin of pronotum. Scale bars: 1 mm (A); 2 mm (B – F).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
917FE7E338745D0D8D2ECC9B93261F83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601912	https://doi.org/10.3897/asp.84.e182696.figure6	Figure 6. Acanthocephala spp., male and female metafemur. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E. A. (Metapodiessa) femorata (Fabricius, 1775). Scale bar: 5 mm (A – E).	Figure 6. Acanthocephala spp., male and female metafemur. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E. A. (Metapodiessa) femorata (Fabricius, 1775). Scale bar: 5 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
917FE7E338745D0D8D2ECC9B93261F83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601913	https://doi.org/10.3897/asp.84.e182696.figure7	Figure 7. Acanthocephala spp., male and female metatibia. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: de, dorsal expansion of the metatibia; ve, ventral expansion of the metatibia. Scale bar: 5 mm (A – E).	Figure 7. Acanthocephala spp., male and female metatibia. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: de, dorsal expansion of the metatibia; ve, ventral expansion of the metatibia. Scale bar: 5 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
917FE7E338745D0D8D2ECC9B93261F83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601914	https://doi.org/10.3897/asp.84.e182696.figure8	Figure 8. Acanthocephala spp., male abdominal expansion. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: II, abdominal sternite II; III, abdominal sternite III; IV, abdominal sternite IV; V, abdominal sternite V; VI, abdominal sternite VI. Arrows indicate the expansion of the abdominal segments. Scale bar: 2 mm (A – E).	Figure 8. Acanthocephala spp., male abdominal expansion. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: II, abdominal sternite II; III, abdominal sternite III; IV, abdominal sternite IV; V, abdominal sternite V; VI, abdominal sternite VI. Arrows indicate the expansion of the abdominal segments. Scale bar: 2 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
917FE7E338745D0D8D2ECC9B93261F83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601915	https://doi.org/10.3897/asp.84.e182696.figure9	Figure 9. Acanthocephala spp., male genitalia. A, B Pygophore, A. (Spinipedia) heissi Brailovsky, 2006. A Lateral view; B dorsal view; C – E A. (Acanthocephala) latipes (Drury, 1782). C Pygophore, dorsal view; D Right paramere, lateral views; E Aedeagus dorsal and ventral views. Abbreviations: ar, arm of paramere; bsh, basal shank of paramere; dap 1, first dorsal appendage of conjunctiva; dap 2, second dorsal appendage of conjunctiva; dm, dorsal margin of pygophore; do, dorsal opening of pygophore; ds, dorsal sac of conjunctiva; ejd, ejaculatory duct; end, endosoma; gp, gonoporal process; im, inner margin of paramere; om, outer margin of paramere; pdm, posterodorsal margin of pygophore; pha, phallobase; vap, ventral appendage of conjunctiva; vs, vesica. Scale bar: 0.50 mm (A – E).	Figure 9. Acanthocephala spp., male genitalia. A, B Pygophore, A. (Spinipedia) heissi Brailovsky, 2006. A Lateral view; B dorsal view; C – E A. (Acanthocephala) latipes (Drury, 1782). C Pygophore, dorsal view; D Right paramere, lateral views; E Aedeagus dorsal and ventral views. Abbreviations: ar, arm of paramere; bsh, basal shank of paramere; dap 1, first dorsal appendage of conjunctiva; dap 2, second dorsal appendage of conjunctiva; dm, dorsal margin of pygophore; do, dorsal opening of pygophore; ds, dorsal sac of conjunctiva; ejd, ejaculatory duct; end, endosoma; gp, gonoporal process; im, inner margin of paramere; om, outer margin of paramere; pdm, posterodorsal margin of pygophore; pha, phallobase; vap, ventral appendage of conjunctiva; vs, vesica. Scale bar: 0.50 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
917FE7E338745D0D8D2ECC9B93261F83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601916	https://doi.org/10.3897/asp.84.e182696.figure10	Figure 10. Acanthocephala spp., female genitalia. A – D A. (Acanthocephala) latipes (Drury, 1782). A Abdomen ventral view, details of pregenital and genital segments; B First gonocoxa and gonapophysis; C Second gonocoxa and gonapophysis; D Spermatheca; E A. (Spinipedia) heissi Brailovsky, 2006, spermatheca. Abbreviations: da, dorsal apodeme of first gonocoxa; dd, distal duct of spermatheca; df, distal flange; di, dilation of spermathecal duct; fi, fisura; fz, flexible zone; gpI, first gonapophysis; gpII, second gonapophysis; gxI, first gonocoxa; gxII, second gonocoxa; pd, proximal duct of spermatheca; pf, proximal flange; pl, plica; pt 8, paratergite VIII; pt 9, paratergite IX; raII, second ramus; sr, seminal receptacle; VI, abdominal sternite VI; VII, abdominal sternite VII. Scale bars: 1 mm (A); 0.50 mm (B – E).	Figure 10. Acanthocephala spp., female genitalia. A – D A. (Acanthocephala) latipes (Drury, 1782). A Abdomen ventral view, details of pregenital and genital segments; B First gonocoxa and gonapophysis; C Second gonocoxa and gonapophysis; D Spermatheca; E A. (Spinipedia) heissi Brailovsky, 2006, spermatheca. Abbreviations: da, dorsal apodeme of first gonocoxa; dd, distal duct of spermatheca; df, distal flange; di, dilation of spermathecal duct; fi, fisura; fz, flexible zone; gpI, first gonapophysis; gpII, second gonapophysis; gxI, first gonocoxa; gxII, second gonocoxa; pd, proximal duct of spermatheca; pf, proximal flange; pl, plica; pt 8, paratergite VIII; pt 9, paratergite IX; raII, second ramus; sr, seminal receptacle; VI, abdominal sternite VI; VII, abdominal sternite VII. Scale bars: 1 mm (A); 0.50 mm (B – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
917FE7E338745D0D8D2ECC9B93261F83.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601917	https://doi.org/10.3897/asp.84.e182696.figure11	Figure 11. Acanthocephalini spp. A – F Texture of the dorsal surface of pronotum. A Acanthocephala (Metapodiessa) granulosa (Dallas, 1852); B A. (Pronoptera) macrotuberculata Olivera et al., 2024; C A. (Metapodiessa) bicoloripes (Stål, 1855); D A. (Pronoptera) declivis (Say, 1832); E A. (Acanthocephala) rufa Olivera et al., 2024; F A. (Acanthocephala) carioca Olivera et al., 2024; G Male metathoracic acetabulum, Spilopleura parensis (Dallas, 1852); H Detail of the lateral margin of the male abdominal segments, Cervantistellus guerrerensis Brailovsky and Barrera, 2005; I Male metathoracic acetabulum, Petalops distinctus Montandon, 1895. Abbreviations: mtac, metathoracic acetabulum. Arrows indicate the spines and tubercles of the connexival segments. Scale bar: 1 mm (B – I).	Figure 11. Acanthocephalini spp. A – F Texture of the dorsal surface of pronotum. A Acanthocephala (Metapodiessa) granulosa (Dallas, 1852); B A. (Pronoptera) macrotuberculata Olivera et al., 2024; C A. (Metapodiessa) bicoloripes (Stål, 1855); D A. (Pronoptera) declivis (Say, 1832); E A. (Acanthocephala) rufa Olivera et al., 2024; F A. (Acanthocephala) carioca Olivera et al., 2024; G Male metathoracic acetabulum, Spilopleura parensis (Dallas, 1852); H Detail of the lateral margin of the male abdominal segments, Cervantistellus guerrerensis Brailovsky and Barrera, 2005; I Male metathoracic acetabulum, Petalops distinctus Montandon, 1895. Abbreviations: mtac, metathoracic acetabulum. Arrows indicate the spines and tubercles of the connexival segments. Scale bar: 1 mm (B – I).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
EEA42CBF6C1B5258BD7FB7F7F303EB73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601909	https://doi.org/10.3897/asp.84.e182696.figure3	Figure 3. Acanthocephala spp., dorsal habitus of type species and labels. A A. (Acanthocephala) latipes (Drury, 1782), male syntype, photographed by Laurence Livermore (© Zoologisk museum i København), original photo cropped and contrast adjusted, made available by Natural History Museum of Denmark Zoological Museum of Copenhagen under Creative Commons Attribution 3.0, International Public License, CC BY-NC 3.0; B A. (Spinipedia) heissi Brailovsky, 2006, male paratype; C A. (Contrastata) thomasii (Uhler, 1872), male syntype; D A. (Pronoptera) alata (Burmeister, 1835), male syntype, photographed by Laurence Livermore, original photo cropped and contrast adjusted; E A. (Metapodiessa) femorata (Fabricius, 1775), male syntype, original photo cropped and contrast adjusted (© The Trustees of the Natural History Museum, London (Banks collection )). Scale bar: 10 mm (A – E).	Figure 3. Acanthocephala spp., dorsal habitus of type species and labels. A A. (Acanthocephala) latipes (Drury, 1782), male syntype, photographed by Laurence Livermore (© Zoologisk museum i København), original photo cropped and contrast adjusted, made available by Natural History Museum of Denmark Zoological Museum of Copenhagen under Creative Commons Attribution 3.0, International Public License, CC BY-NC 3.0; B A. (Spinipedia) heissi Brailovsky, 2006, male paratype; C A. (Contrastata) thomasii (Uhler, 1872), male syntype; D A. (Pronoptera) alata (Burmeister, 1835), male syntype, photographed by Laurence Livermore, original photo cropped and contrast adjusted; E A. (Metapodiessa) femorata (Fabricius, 1775), male syntype, original photo cropped and contrast adjusted (© The Trustees of the Natural History Museum, London (Banks collection )). Scale bar: 10 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
EEA42CBF6C1B5258BD7FB7F7F303EB73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601910	https://doi.org/10.3897/asp.84.e182696.figure4	Figure 4. Acanthocephala spp., head. A Dorsal view, A. (Acanthocephala) latipes (Drury, 1782); B – F Lateral view. B A. (Acanthocephala) latipes; C A. (Spinipedia) heissi Brailovsky, 2006; D A. (Contrastata) thomasii (Uhler, 1872); E A. (Pronoptera) alata (Burmeister, 1835); F A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: bu, buccula; ju, juga; mxp, maxillary plate; ty, tylus. Asterisks indicate the apex of the tylus above (char. 26: 1) and below (char. 26: 0) to the longitudinal axis of the head in distal view. Scale bar: 1 mm (A – F).	Figure 4. Acanthocephala spp., head. A Dorsal view, A. (Acanthocephala) latipes (Drury, 1782); B – F Lateral view. B A. (Acanthocephala) latipes; C A. (Spinipedia) heissi Brailovsky, 2006; D A. (Contrastata) thomasii (Uhler, 1872); E A. (Pronoptera) alata (Burmeister, 1835); F A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: bu, buccula; ju, juga; mxp, maxillary plate; ty, tylus. Asterisks indicate the apex of the tylus above (char. 26: 1) and below (char. 26: 0) to the longitudinal axis of the head in distal view. Scale bar: 1 mm (A – F).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
EEA42CBF6C1B5258BD7FB7F7F303EB73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601912	https://doi.org/10.3897/asp.84.e182696.figure6	Figure 6. Acanthocephala spp., male and female metafemur. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E. A. (Metapodiessa) femorata (Fabricius, 1775). Scale bar: 5 mm (A – E).	Figure 6. Acanthocephala spp., male and female metafemur. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E. A. (Metapodiessa) femorata (Fabricius, 1775). Scale bar: 5 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
EEA42CBF6C1B5258BD7FB7F7F303EB73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601913	https://doi.org/10.3897/asp.84.e182696.figure7	Figure 7. Acanthocephala spp., male and female metatibia. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: de, dorsal expansion of the metatibia; ve, ventral expansion of the metatibia. Scale bar: 5 mm (A – E).	Figure 7. Acanthocephala spp., male and female metatibia. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: de, dorsal expansion of the metatibia; ve, ventral expansion of the metatibia. Scale bar: 5 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
EEA42CBF6C1B5258BD7FB7F7F303EB73.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601914	https://doi.org/10.3897/asp.84.e182696.figure8	Figure 8. Acanthocephala spp., male abdominal expansion. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: II, abdominal sternite II; III, abdominal sternite III; IV, abdominal sternite IV; V, abdominal sternite V; VI, abdominal sternite VI. Arrows indicate the expansion of the abdominal segments. Scale bar: 2 mm (A – E).	Figure 8. Acanthocephala spp., male abdominal expansion. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: II, abdominal sternite II; III, abdominal sternite III; IV, abdominal sternite IV; V, abdominal sternite V; VI, abdominal sternite VI. Arrows indicate the expansion of the abdominal segments. Scale bar: 2 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
78F569D19E2F5B7B8120504F672690DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601909	https://doi.org/10.3897/asp.84.e182696.figure3	Figure 3. Acanthocephala spp., dorsal habitus of type species and labels. A A. (Acanthocephala) latipes (Drury, 1782), male syntype, photographed by Laurence Livermore (© Zoologisk museum i København), original photo cropped and contrast adjusted, made available by Natural History Museum of Denmark Zoological Museum of Copenhagen under Creative Commons Attribution 3.0, International Public License, CC BY-NC 3.0; B A. (Spinipedia) heissi Brailovsky, 2006, male paratype; C A. (Contrastata) thomasii (Uhler, 1872), male syntype; D A. (Pronoptera) alata (Burmeister, 1835), male syntype, photographed by Laurence Livermore, original photo cropped and contrast adjusted; E A. (Metapodiessa) femorata (Fabricius, 1775), male syntype, original photo cropped and contrast adjusted (© The Trustees of the Natural History Museum, London (Banks collection )). Scale bar: 10 mm (A – E).	Figure 3. Acanthocephala spp., dorsal habitus of type species and labels. A A. (Acanthocephala) latipes (Drury, 1782), male syntype, photographed by Laurence Livermore (© Zoologisk museum i København), original photo cropped and contrast adjusted, made available by Natural History Museum of Denmark Zoological Museum of Copenhagen under Creative Commons Attribution 3.0, International Public License, CC BY-NC 3.0; B A. (Spinipedia) heissi Brailovsky, 2006, male paratype; C A. (Contrastata) thomasii (Uhler, 1872), male syntype; D A. (Pronoptera) alata (Burmeister, 1835), male syntype, photographed by Laurence Livermore, original photo cropped and contrast adjusted; E A. (Metapodiessa) femorata (Fabricius, 1775), male syntype, original photo cropped and contrast adjusted (© The Trustees of the Natural History Museum, London (Banks collection )). Scale bar: 10 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
78F569D19E2F5B7B8120504F672690DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601910	https://doi.org/10.3897/asp.84.e182696.figure4	Figure 4. Acanthocephala spp., head. A Dorsal view, A. (Acanthocephala) latipes (Drury, 1782); B – F Lateral view. B A. (Acanthocephala) latipes; C A. (Spinipedia) heissi Brailovsky, 2006; D A. (Contrastata) thomasii (Uhler, 1872); E A. (Pronoptera) alata (Burmeister, 1835); F A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: bu, buccula; ju, juga; mxp, maxillary plate; ty, tylus. Asterisks indicate the apex of the tylus above (char. 26: 1) and below (char. 26: 0) to the longitudinal axis of the head in distal view. Scale bar: 1 mm (A – F).	Figure 4. Acanthocephala spp., head. A Dorsal view, A. (Acanthocephala) latipes (Drury, 1782); B – F Lateral view. B A. (Acanthocephala) latipes; C A. (Spinipedia) heissi Brailovsky, 2006; D A. (Contrastata) thomasii (Uhler, 1872); E A. (Pronoptera) alata (Burmeister, 1835); F A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: bu, buccula; ju, juga; mxp, maxillary plate; ty, tylus. Asterisks indicate the apex of the tylus above (char. 26: 1) and below (char. 26: 0) to the longitudinal axis of the head in distal view. Scale bar: 1 mm (A – F).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
78F569D19E2F5B7B8120504F672690DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601911	https://doi.org/10.3897/asp.84.e182696.figure5	Figure 5. Acanthocephalini spp. A – D Dorsal view of pronotum. A Ctenomelynthus coxalis Breddin, 1903; B Acanthocephala (Acanthocephala) latipes (Drury, 1782); C A. (Pronoptera) panamensis Distant, 1881; D A. (Metapodiessa) femorata (Fabricius, 1775); E Caudal view of pronotum, A. (Pronoptera) panamensis; F Spines of the costal margin of hemelytra, A. (Pronoptera) panamensis. Abbreviations: alm, anterolateral margin of pronotum; cm, costal margin of hemelytra; ha, humeral angles of pronotum; has, spine of humeral angles of pronotum; plm, posterolateral margin of pronotum. Scale bars: 1 mm (A); 2 mm (B – F).	Figure 5. Acanthocephalini spp. A – D Dorsal view of pronotum. A Ctenomelynthus coxalis Breddin, 1903; B Acanthocephala (Acanthocephala) latipes (Drury, 1782); C A. (Pronoptera) panamensis Distant, 1881; D A. (Metapodiessa) femorata (Fabricius, 1775); E Caudal view of pronotum, A. (Pronoptera) panamensis; F Spines of the costal margin of hemelytra, A. (Pronoptera) panamensis. Abbreviations: alm, anterolateral margin of pronotum; cm, costal margin of hemelytra; ha, humeral angles of pronotum; has, spine of humeral angles of pronotum; plm, posterolateral margin of pronotum. Scale bars: 1 mm (A); 2 mm (B – F).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
78F569D19E2F5B7B8120504F672690DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601912	https://doi.org/10.3897/asp.84.e182696.figure6	Figure 6. Acanthocephala spp., male and female metafemur. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E. A. (Metapodiessa) femorata (Fabricius, 1775). Scale bar: 5 mm (A – E).	Figure 6. Acanthocephala spp., male and female metafemur. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E. A. (Metapodiessa) femorata (Fabricius, 1775). Scale bar: 5 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
78F569D19E2F5B7B8120504F672690DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601913	https://doi.org/10.3897/asp.84.e182696.figure7	Figure 7. Acanthocephala spp., male and female metatibia. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: de, dorsal expansion of the metatibia; ve, ventral expansion of the metatibia. Scale bar: 5 mm (A – E).	Figure 7. Acanthocephala spp., male and female metatibia. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: de, dorsal expansion of the metatibia; ve, ventral expansion of the metatibia. Scale bar: 5 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
78F569D19E2F5B7B8120504F672690DF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601914	https://doi.org/10.3897/asp.84.e182696.figure8	Figure 8. Acanthocephala spp., male abdominal expansion. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: II, abdominal sternite II; III, abdominal sternite III; IV, abdominal sternite IV; V, abdominal sternite V; VI, abdominal sternite VI. Arrows indicate the expansion of the abdominal segments. Scale bar: 2 mm (A – E).	Figure 8. Acanthocephala spp., male abdominal expansion. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: II, abdominal sternite II; III, abdominal sternite III; IV, abdominal sternite IV; V, abdominal sternite V; VI, abdominal sternite VI. Arrows indicate the expansion of the abdominal segments. Scale bar: 2 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
0E630DD617785DEB91628BE2C9FC8A88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601909	https://doi.org/10.3897/asp.84.e182696.figure3	Figure 3. Acanthocephala spp., dorsal habitus of type species and labels. A A. (Acanthocephala) latipes (Drury, 1782), male syntype, photographed by Laurence Livermore (© Zoologisk museum i København), original photo cropped and contrast adjusted, made available by Natural History Museum of Denmark Zoological Museum of Copenhagen under Creative Commons Attribution 3.0, International Public License, CC BY-NC 3.0; B A. (Spinipedia) heissi Brailovsky, 2006, male paratype; C A. (Contrastata) thomasii (Uhler, 1872), male syntype; D A. (Pronoptera) alata (Burmeister, 1835), male syntype, photographed by Laurence Livermore, original photo cropped and contrast adjusted; E A. (Metapodiessa) femorata (Fabricius, 1775), male syntype, original photo cropped and contrast adjusted (© The Trustees of the Natural History Museum, London (Banks collection )). Scale bar: 10 mm (A – E).	Figure 3. Acanthocephala spp., dorsal habitus of type species and labels. A A. (Acanthocephala) latipes (Drury, 1782), male syntype, photographed by Laurence Livermore (© Zoologisk museum i København), original photo cropped and contrast adjusted, made available by Natural History Museum of Denmark Zoological Museum of Copenhagen under Creative Commons Attribution 3.0, International Public License, CC BY-NC 3.0; B A. (Spinipedia) heissi Brailovsky, 2006, male paratype; C A. (Contrastata) thomasii (Uhler, 1872), male syntype; D A. (Pronoptera) alata (Burmeister, 1835), male syntype, photographed by Laurence Livermore, original photo cropped and contrast adjusted; E A. (Metapodiessa) femorata (Fabricius, 1775), male syntype, original photo cropped and contrast adjusted (© The Trustees of the Natural History Museum, London (Banks collection )). Scale bar: 10 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
0E630DD617785DEB91628BE2C9FC8A88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601910	https://doi.org/10.3897/asp.84.e182696.figure4	Figure 4. Acanthocephala spp., head. A Dorsal view, A. (Acanthocephala) latipes (Drury, 1782); B – F Lateral view. B A. (Acanthocephala) latipes; C A. (Spinipedia) heissi Brailovsky, 2006; D A. (Contrastata) thomasii (Uhler, 1872); E A. (Pronoptera) alata (Burmeister, 1835); F A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: bu, buccula; ju, juga; mxp, maxillary plate; ty, tylus. Asterisks indicate the apex of the tylus above (char. 26: 1) and below (char. 26: 0) to the longitudinal axis of the head in distal view. Scale bar: 1 mm (A – F).	Figure 4. Acanthocephala spp., head. A Dorsal view, A. (Acanthocephala) latipes (Drury, 1782); B – F Lateral view. B A. (Acanthocephala) latipes; C A. (Spinipedia) heissi Brailovsky, 2006; D A. (Contrastata) thomasii (Uhler, 1872); E A. (Pronoptera) alata (Burmeister, 1835); F A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: bu, buccula; ju, juga; mxp, maxillary plate; ty, tylus. Asterisks indicate the apex of the tylus above (char. 26: 1) and below (char. 26: 0) to the longitudinal axis of the head in distal view. Scale bar: 1 mm (A – F).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
0E630DD617785DEB91628BE2C9FC8A88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601911	https://doi.org/10.3897/asp.84.e182696.figure5	Figure 5. Acanthocephalini spp. A – D Dorsal view of pronotum. A Ctenomelynthus coxalis Breddin, 1903; B Acanthocephala (Acanthocephala) latipes (Drury, 1782); C A. (Pronoptera) panamensis Distant, 1881; D A. (Metapodiessa) femorata (Fabricius, 1775); E Caudal view of pronotum, A. (Pronoptera) panamensis; F Spines of the costal margin of hemelytra, A. (Pronoptera) panamensis. Abbreviations: alm, anterolateral margin of pronotum; cm, costal margin of hemelytra; ha, humeral angles of pronotum; has, spine of humeral angles of pronotum; plm, posterolateral margin of pronotum. Scale bars: 1 mm (A); 2 mm (B – F).	Figure 5. Acanthocephalini spp. A – D Dorsal view of pronotum. A Ctenomelynthus coxalis Breddin, 1903; B Acanthocephala (Acanthocephala) latipes (Drury, 1782); C A. (Pronoptera) panamensis Distant, 1881; D A. (Metapodiessa) femorata (Fabricius, 1775); E Caudal view of pronotum, A. (Pronoptera) panamensis; F Spines of the costal margin of hemelytra, A. (Pronoptera) panamensis. Abbreviations: alm, anterolateral margin of pronotum; cm, costal margin of hemelytra; ha, humeral angles of pronotum; has, spine of humeral angles of pronotum; plm, posterolateral margin of pronotum. Scale bars: 1 mm (A); 2 mm (B – F).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
0E630DD617785DEB91628BE2C9FC8A88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601912	https://doi.org/10.3897/asp.84.e182696.figure6	Figure 6. Acanthocephala spp., male and female metafemur. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E. A. (Metapodiessa) femorata (Fabricius, 1775). Scale bar: 5 mm (A – E).	Figure 6. Acanthocephala spp., male and female metafemur. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E. A. (Metapodiessa) femorata (Fabricius, 1775). Scale bar: 5 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
0E630DD617785DEB91628BE2C9FC8A88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601913	https://doi.org/10.3897/asp.84.e182696.figure7	Figure 7. Acanthocephala spp., male and female metatibia. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: de, dorsal expansion of the metatibia; ve, ventral expansion of the metatibia. Scale bar: 5 mm (A – E).	Figure 7. Acanthocephala spp., male and female metatibia. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: de, dorsal expansion of the metatibia; ve, ventral expansion of the metatibia. Scale bar: 5 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
0E630DD617785DEB91628BE2C9FC8A88.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601914	https://doi.org/10.3897/asp.84.e182696.figure8	Figure 8. Acanthocephala spp., male abdominal expansion. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: II, abdominal sternite II; III, abdominal sternite III; IV, abdominal sternite IV; V, abdominal sternite V; VI, abdominal sternite VI. Arrows indicate the expansion of the abdominal segments. Scale bar: 2 mm (A – E).	Figure 8. Acanthocephala spp., male abdominal expansion. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: II, abdominal sternite II; III, abdominal sternite III; IV, abdominal sternite IV; V, abdominal sternite V; VI, abdominal sternite VI. Arrows indicate the expansion of the abdominal segments. Scale bar: 2 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
4BCA99D22A18530C9C37E5C8B76A8FE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601909	https://doi.org/10.3897/asp.84.e182696.figure3	Figure 3. Acanthocephala spp., dorsal habitus of type species and labels. A A. (Acanthocephala) latipes (Drury, 1782), male syntype, photographed by Laurence Livermore (© Zoologisk museum i København), original photo cropped and contrast adjusted, made available by Natural History Museum of Denmark Zoological Museum of Copenhagen under Creative Commons Attribution 3.0, International Public License, CC BY-NC 3.0; B A. (Spinipedia) heissi Brailovsky, 2006, male paratype; C A. (Contrastata) thomasii (Uhler, 1872), male syntype; D A. (Pronoptera) alata (Burmeister, 1835), male syntype, photographed by Laurence Livermore, original photo cropped and contrast adjusted; E A. (Metapodiessa) femorata (Fabricius, 1775), male syntype, original photo cropped and contrast adjusted (© The Trustees of the Natural History Museum, London (Banks collection )). Scale bar: 10 mm (A – E).	Figure 3. Acanthocephala spp., dorsal habitus of type species and labels. A A. (Acanthocephala) latipes (Drury, 1782), male syntype, photographed by Laurence Livermore (© Zoologisk museum i København), original photo cropped and contrast adjusted, made available by Natural History Museum of Denmark Zoological Museum of Copenhagen under Creative Commons Attribution 3.0, International Public License, CC BY-NC 3.0; B A. (Spinipedia) heissi Brailovsky, 2006, male paratype; C A. (Contrastata) thomasii (Uhler, 1872), male syntype; D A. (Pronoptera) alata (Burmeister, 1835), male syntype, photographed by Laurence Livermore, original photo cropped and contrast adjusted; E A. (Metapodiessa) femorata (Fabricius, 1775), male syntype, original photo cropped and contrast adjusted (© The Trustees of the Natural History Museum, London (Banks collection )). Scale bar: 10 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
4BCA99D22A18530C9C37E5C8B76A8FE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601910	https://doi.org/10.3897/asp.84.e182696.figure4	Figure 4. Acanthocephala spp., head. A Dorsal view, A. (Acanthocephala) latipes (Drury, 1782); B – F Lateral view. B A. (Acanthocephala) latipes; C A. (Spinipedia) heissi Brailovsky, 2006; D A. (Contrastata) thomasii (Uhler, 1872); E A. (Pronoptera) alata (Burmeister, 1835); F A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: bu, buccula; ju, juga; mxp, maxillary plate; ty, tylus. Asterisks indicate the apex of the tylus above (char. 26: 1) and below (char. 26: 0) to the longitudinal axis of the head in distal view. Scale bar: 1 mm (A – F).	Figure 4. Acanthocephala spp., head. A Dorsal view, A. (Acanthocephala) latipes (Drury, 1782); B – F Lateral view. B A. (Acanthocephala) latipes; C A. (Spinipedia) heissi Brailovsky, 2006; D A. (Contrastata) thomasii (Uhler, 1872); E A. (Pronoptera) alata (Burmeister, 1835); F A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: bu, buccula; ju, juga; mxp, maxillary plate; ty, tylus. Asterisks indicate the apex of the tylus above (char. 26: 1) and below (char. 26: 0) to the longitudinal axis of the head in distal view. Scale bar: 1 mm (A – F).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
4BCA99D22A18530C9C37E5C8B76A8FE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601912	https://doi.org/10.3897/asp.84.e182696.figure6	Figure 6. Acanthocephala spp., male and female metafemur. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E. A. (Metapodiessa) femorata (Fabricius, 1775). Scale bar: 5 mm (A – E).	Figure 6. Acanthocephala spp., male and female metafemur. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E. A. (Metapodiessa) femorata (Fabricius, 1775). Scale bar: 5 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
4BCA99D22A18530C9C37E5C8B76A8FE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601913	https://doi.org/10.3897/asp.84.e182696.figure7	Figure 7. Acanthocephala spp., male and female metatibia. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: de, dorsal expansion of the metatibia; ve, ventral expansion of the metatibia. Scale bar: 5 mm (A – E).	Figure 7. Acanthocephala spp., male and female metatibia. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: de, dorsal expansion of the metatibia; ve, ventral expansion of the metatibia. Scale bar: 5 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
4BCA99D22A18530C9C37E5C8B76A8FE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601914	https://doi.org/10.3897/asp.84.e182696.figure8	Figure 8. Acanthocephala spp., male abdominal expansion. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: II, abdominal sternite II; III, abdominal sternite III; IV, abdominal sternite IV; V, abdominal sternite V; VI, abdominal sternite VI. Arrows indicate the expansion of the abdominal segments. Scale bar: 2 mm (A – E).	Figure 8. Acanthocephala spp., male abdominal expansion. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: II, abdominal sternite II; III, abdominal sternite III; IV, abdominal sternite IV; V, abdominal sternite V; VI, abdominal sternite VI. Arrows indicate the expansion of the abdominal segments. Scale bar: 2 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
4BCA99D22A18530C9C37E5C8B76A8FE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601915	https://doi.org/10.3897/asp.84.e182696.figure9	Figure 9. Acanthocephala spp., male genitalia. A, B Pygophore, A. (Spinipedia) heissi Brailovsky, 2006. A Lateral view; B dorsal view; C – E A. (Acanthocephala) latipes (Drury, 1782). C Pygophore, dorsal view; D Right paramere, lateral views; E Aedeagus dorsal and ventral views. Abbreviations: ar, arm of paramere; bsh, basal shank of paramere; dap 1, first dorsal appendage of conjunctiva; dap 2, second dorsal appendage of conjunctiva; dm, dorsal margin of pygophore; do, dorsal opening of pygophore; ds, dorsal sac of conjunctiva; ejd, ejaculatory duct; end, endosoma; gp, gonoporal process; im, inner margin of paramere; om, outer margin of paramere; pdm, posterodorsal margin of pygophore; pha, phallobase; vap, ventral appendage of conjunctiva; vs, vesica. Scale bar: 0.50 mm (A – E).	Figure 9. Acanthocephala spp., male genitalia. A, B Pygophore, A. (Spinipedia) heissi Brailovsky, 2006. A Lateral view; B dorsal view; C – E A. (Acanthocephala) latipes (Drury, 1782). C Pygophore, dorsal view; D Right paramere, lateral views; E Aedeagus dorsal and ventral views. Abbreviations: ar, arm of paramere; bsh, basal shank of paramere; dap 1, first dorsal appendage of conjunctiva; dap 2, second dorsal appendage of conjunctiva; dm, dorsal margin of pygophore; do, dorsal opening of pygophore; ds, dorsal sac of conjunctiva; ejd, ejaculatory duct; end, endosoma; gp, gonoporal process; im, inner margin of paramere; om, outer margin of paramere; pdm, posterodorsal margin of pygophore; pha, phallobase; vap, ventral appendage of conjunctiva; vs, vesica. Scale bar: 0.50 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
4BCA99D22A18530C9C37E5C8B76A8FE0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601916	https://doi.org/10.3897/asp.84.e182696.figure10	Figure 10. Acanthocephala spp., female genitalia. A – D A. (Acanthocephala) latipes (Drury, 1782). A Abdomen ventral view, details of pregenital and genital segments; B First gonocoxa and gonapophysis; C Second gonocoxa and gonapophysis; D Spermatheca; E A. (Spinipedia) heissi Brailovsky, 2006, spermatheca. Abbreviations: da, dorsal apodeme of first gonocoxa; dd, distal duct of spermatheca; df, distal flange; di, dilation of spermathecal duct; fi, fisura; fz, flexible zone; gpI, first gonapophysis; gpII, second gonapophysis; gxI, first gonocoxa; gxII, second gonocoxa; pd, proximal duct of spermatheca; pf, proximal flange; pl, plica; pt 8, paratergite VIII; pt 9, paratergite IX; raII, second ramus; sr, seminal receptacle; VI, abdominal sternite VI; VII, abdominal sternite VII. Scale bars: 1 mm (A); 0.50 mm (B – E).	Figure 10. Acanthocephala spp., female genitalia. A – D A. (Acanthocephala) latipes (Drury, 1782). A Abdomen ventral view, details of pregenital and genital segments; B First gonocoxa and gonapophysis; C Second gonocoxa and gonapophysis; D Spermatheca; E A. (Spinipedia) heissi Brailovsky, 2006, spermatheca. Abbreviations: da, dorsal apodeme of first gonocoxa; dd, distal duct of spermatheca; df, distal flange; di, dilation of spermathecal duct; fi, fisura; fz, flexible zone; gpI, first gonapophysis; gpII, second gonapophysis; gxI, first gonocoxa; gxII, second gonocoxa; pd, proximal duct of spermatheca; pf, proximal flange; pl, plica; pt 8, paratergite VIII; pt 9, paratergite IX; raII, second ramus; sr, seminal receptacle; VI, abdominal sternite VI; VII, abdominal sternite VII. Scale bars: 1 mm (A); 0.50 mm (B – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
D3347EF585335248901BA7004C609740.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601909	https://doi.org/10.3897/asp.84.e182696.figure3	Figure 3. Acanthocephala spp., dorsal habitus of type species and labels. A A. (Acanthocephala) latipes (Drury, 1782), male syntype, photographed by Laurence Livermore (© Zoologisk museum i København), original photo cropped and contrast adjusted, made available by Natural History Museum of Denmark Zoological Museum of Copenhagen under Creative Commons Attribution 3.0, International Public License, CC BY-NC 3.0; B A. (Spinipedia) heissi Brailovsky, 2006, male paratype; C A. (Contrastata) thomasii (Uhler, 1872), male syntype; D A. (Pronoptera) alata (Burmeister, 1835), male syntype, photographed by Laurence Livermore, original photo cropped and contrast adjusted; E A. (Metapodiessa) femorata (Fabricius, 1775), male syntype, original photo cropped and contrast adjusted (© The Trustees of the Natural History Museum, London (Banks collection )). Scale bar: 10 mm (A – E).	Figure 3. Acanthocephala spp., dorsal habitus of type species and labels. A A. (Acanthocephala) latipes (Drury, 1782), male syntype, photographed by Laurence Livermore (© Zoologisk museum i København), original photo cropped and contrast adjusted, made available by Natural History Museum of Denmark Zoological Museum of Copenhagen under Creative Commons Attribution 3.0, International Public License, CC BY-NC 3.0; B A. (Spinipedia) heissi Brailovsky, 2006, male paratype; C A. (Contrastata) thomasii (Uhler, 1872), male syntype; D A. (Pronoptera) alata (Burmeister, 1835), male syntype, photographed by Laurence Livermore, original photo cropped and contrast adjusted; E A. (Metapodiessa) femorata (Fabricius, 1775), male syntype, original photo cropped and contrast adjusted (© The Trustees of the Natural History Museum, London (Banks collection )). Scale bar: 10 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
D3347EF585335248901BA7004C609740.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601910	https://doi.org/10.3897/asp.84.e182696.figure4	Figure 4. Acanthocephala spp., head. A Dorsal view, A. (Acanthocephala) latipes (Drury, 1782); B – F Lateral view. B A. (Acanthocephala) latipes; C A. (Spinipedia) heissi Brailovsky, 2006; D A. (Contrastata) thomasii (Uhler, 1872); E A. (Pronoptera) alata (Burmeister, 1835); F A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: bu, buccula; ju, juga; mxp, maxillary plate; ty, tylus. Asterisks indicate the apex of the tylus above (char. 26: 1) and below (char. 26: 0) to the longitudinal axis of the head in distal view. Scale bar: 1 mm (A – F).	Figure 4. Acanthocephala spp., head. A Dorsal view, A. (Acanthocephala) latipes (Drury, 1782); B – F Lateral view. B A. (Acanthocephala) latipes; C A. (Spinipedia) heissi Brailovsky, 2006; D A. (Contrastata) thomasii (Uhler, 1872); E A. (Pronoptera) alata (Burmeister, 1835); F A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: bu, buccula; ju, juga; mxp, maxillary plate; ty, tylus. Asterisks indicate the apex of the tylus above (char. 26: 1) and below (char. 26: 0) to the longitudinal axis of the head in distal view. Scale bar: 1 mm (A – F).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
D3347EF585335248901BA7004C609740.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601911	https://doi.org/10.3897/asp.84.e182696.figure5	Figure 5. Acanthocephalini spp. A – D Dorsal view of pronotum. A Ctenomelynthus coxalis Breddin, 1903; B Acanthocephala (Acanthocephala) latipes (Drury, 1782); C A. (Pronoptera) panamensis Distant, 1881; D A. (Metapodiessa) femorata (Fabricius, 1775); E Caudal view of pronotum, A. (Pronoptera) panamensis; F Spines of the costal margin of hemelytra, A. (Pronoptera) panamensis. Abbreviations: alm, anterolateral margin of pronotum; cm, costal margin of hemelytra; ha, humeral angles of pronotum; has, spine of humeral angles of pronotum; plm, posterolateral margin of pronotum. Scale bars: 1 mm (A); 2 mm (B – F).	Figure 5. Acanthocephalini spp. A – D Dorsal view of pronotum. A Ctenomelynthus coxalis Breddin, 1903; B Acanthocephala (Acanthocephala) latipes (Drury, 1782); C A. (Pronoptera) panamensis Distant, 1881; D A. (Metapodiessa) femorata (Fabricius, 1775); E Caudal view of pronotum, A. (Pronoptera) panamensis; F Spines of the costal margin of hemelytra, A. (Pronoptera) panamensis. Abbreviations: alm, anterolateral margin of pronotum; cm, costal margin of hemelytra; ha, humeral angles of pronotum; has, spine of humeral angles of pronotum; plm, posterolateral margin of pronotum. Scale bars: 1 mm (A); 2 mm (B – F).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
D3347EF585335248901BA7004C609740.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601912	https://doi.org/10.3897/asp.84.e182696.figure6	Figure 6. Acanthocephala spp., male and female metafemur. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E. A. (Metapodiessa) femorata (Fabricius, 1775). Scale bar: 5 mm (A – E).	Figure 6. Acanthocephala spp., male and female metafemur. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E. A. (Metapodiessa) femorata (Fabricius, 1775). Scale bar: 5 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
D3347EF585335248901BA7004C609740.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601913	https://doi.org/10.3897/asp.84.e182696.figure7	Figure 7. Acanthocephala spp., male and female metatibia. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: de, dorsal expansion of the metatibia; ve, ventral expansion of the metatibia. Scale bar: 5 mm (A – E).	Figure 7. Acanthocephala spp., male and female metatibia. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: de, dorsal expansion of the metatibia; ve, ventral expansion of the metatibia. Scale bar: 5 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
D3347EF585335248901BA7004C609740.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601914	https://doi.org/10.3897/asp.84.e182696.figure8	Figure 8. Acanthocephala spp., male abdominal expansion. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: II, abdominal sternite II; III, abdominal sternite III; IV, abdominal sternite IV; V, abdominal sternite V; VI, abdominal sternite VI. Arrows indicate the expansion of the abdominal segments. Scale bar: 2 mm (A – E).	Figure 8. Acanthocephala spp., male abdominal expansion. A A. (Acanthocephala) latipes (Drury, 1782); B A. (Spinipedia) heissi Brailovsky, 2006; C A. (Contrastata) thomasii (Uhler, 1872); D A. (Pronoptera) alata (Burmeister, 1835); E A. (Metapodiessa) femorata (Fabricius, 1775). Abbreviations: II, abdominal sternite II; III, abdominal sternite III; IV, abdominal sternite IV; V, abdominal sternite V; VI, abdominal sternite VI. Arrows indicate the expansion of the abdominal segments. Scale bar: 2 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
D3347EF585335248901BA7004C609740.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601915	https://doi.org/10.3897/asp.84.e182696.figure9	Figure 9. Acanthocephala spp., male genitalia. A, B Pygophore, A. (Spinipedia) heissi Brailovsky, 2006. A Lateral view; B dorsal view; C – E A. (Acanthocephala) latipes (Drury, 1782). C Pygophore, dorsal view; D Right paramere, lateral views; E Aedeagus dorsal and ventral views. Abbreviations: ar, arm of paramere; bsh, basal shank of paramere; dap 1, first dorsal appendage of conjunctiva; dap 2, second dorsal appendage of conjunctiva; dm, dorsal margin of pygophore; do, dorsal opening of pygophore; ds, dorsal sac of conjunctiva; ejd, ejaculatory duct; end, endosoma; gp, gonoporal process; im, inner margin of paramere; om, outer margin of paramere; pdm, posterodorsal margin of pygophore; pha, phallobase; vap, ventral appendage of conjunctiva; vs, vesica. Scale bar: 0.50 mm (A – E).	Figure 9. Acanthocephala spp., male genitalia. A, B Pygophore, A. (Spinipedia) heissi Brailovsky, 2006. A Lateral view; B dorsal view; C – E A. (Acanthocephala) latipes (Drury, 1782). C Pygophore, dorsal view; D Right paramere, lateral views; E Aedeagus dorsal and ventral views. Abbreviations: ar, arm of paramere; bsh, basal shank of paramere; dap 1, first dorsal appendage of conjunctiva; dap 2, second dorsal appendage of conjunctiva; dm, dorsal margin of pygophore; do, dorsal opening of pygophore; ds, dorsal sac of conjunctiva; ejd, ejaculatory duct; end, endosoma; gp, gonoporal process; im, inner margin of paramere; om, outer margin of paramere; pdm, posterodorsal margin of pygophore; pha, phallobase; vap, ventral appendage of conjunctiva; vs, vesica. Scale bar: 0.50 mm (A – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
D3347EF585335248901BA7004C609740.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601916	https://doi.org/10.3897/asp.84.e182696.figure10	Figure 10. Acanthocephala spp., female genitalia. A – D A. (Acanthocephala) latipes (Drury, 1782). A Abdomen ventral view, details of pregenital and genital segments; B First gonocoxa and gonapophysis; C Second gonocoxa and gonapophysis; D Spermatheca; E A. (Spinipedia) heissi Brailovsky, 2006, spermatheca. Abbreviations: da, dorsal apodeme of first gonocoxa; dd, distal duct of spermatheca; df, distal flange; di, dilation of spermathecal duct; fi, fisura; fz, flexible zone; gpI, first gonapophysis; gpII, second gonapophysis; gxI, first gonocoxa; gxII, second gonocoxa; pd, proximal duct of spermatheca; pf, proximal flange; pl, plica; pt 8, paratergite VIII; pt 9, paratergite IX; raII, second ramus; sr, seminal receptacle; VI, abdominal sternite VI; VII, abdominal sternite VII. Scale bars: 1 mm (A); 0.50 mm (B – E).	Figure 10. Acanthocephala spp., female genitalia. A – D A. (Acanthocephala) latipes (Drury, 1782). A Abdomen ventral view, details of pregenital and genital segments; B First gonocoxa and gonapophysis; C Second gonocoxa and gonapophysis; D Spermatheca; E A. (Spinipedia) heissi Brailovsky, 2006, spermatheca. Abbreviations: da, dorsal apodeme of first gonocoxa; dd, distal duct of spermatheca; df, distal flange; di, dilation of spermathecal duct; fi, fisura; fz, flexible zone; gpI, first gonapophysis; gpII, second gonapophysis; gxI, first gonocoxa; gxII, second gonocoxa; pd, proximal duct of spermatheca; pf, proximal flange; pl, plica; pt 8, paratergite VIII; pt 9, paratergite IX; raII, second ramus; sr, seminal receptacle; VI, abdominal sternite VI; VII, abdominal sternite VII. Scale bars: 1 mm (A); 0.50 mm (B – E).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
D3347EF585335248901BA7004C609740.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1601917	https://doi.org/10.3897/asp.84.e182696.figure11	Figure 11. Acanthocephalini spp. A – F Texture of the dorsal surface of pronotum. A Acanthocephala (Metapodiessa) granulosa (Dallas, 1852); B A. (Pronoptera) macrotuberculata Olivera et al., 2024; C A. (Metapodiessa) bicoloripes (Stål, 1855); D A. (Pronoptera) declivis (Say, 1832); E A. (Acanthocephala) rufa Olivera et al., 2024; F A. (Acanthocephala) carioca Olivera et al., 2024; G Male metathoracic acetabulum, Spilopleura parensis (Dallas, 1852); H Detail of the lateral margin of the male abdominal segments, Cervantistellus guerrerensis Brailovsky and Barrera, 2005; I Male metathoracic acetabulum, Petalops distinctus Montandon, 1895. Abbreviations: mtac, metathoracic acetabulum. Arrows indicate the spines and tubercles of the connexival segments. Scale bar: 1 mm (B – I).	Figure 11. Acanthocephalini spp. A – F Texture of the dorsal surface of pronotum. A Acanthocephala (Metapodiessa) granulosa (Dallas, 1852); B A. (Pronoptera) macrotuberculata Olivera et al., 2024; C A. (Metapodiessa) bicoloripes (Stål, 1855); D A. (Pronoptera) declivis (Say, 1832); E A. (Acanthocephala) rufa Olivera et al., 2024; F A. (Acanthocephala) carioca Olivera et al., 2024; G Male metathoracic acetabulum, Spilopleura parensis (Dallas, 1852); H Detail of the lateral margin of the male abdominal segments, Cervantistellus guerrerensis Brailovsky and Barrera, 2005; I Male metathoracic acetabulum, Petalops distinctus Montandon, 1895. Abbreviations: mtac, metathoracic acetabulum. Arrows indicate the spines and tubercles of the connexival segments. Scale bar: 1 mm (B – I).	2026-04-21	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías		Zenodo	biologists	Olivera, Leonela;Melo, María Cecilia;Dellapé, Pablo Matías			
