identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
918210A95A885C0C86A4BB0E77A376EC.text	918210A95A885C0C86A4BB0E77A376EC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Acizzia uncatoides (Ferris & Klyver 1932)	<div><p>Acizzia uncatoides (Ferris &amp; Klyver, 1932)</p><p>First detection / collection in HI.</p><p>1966 (Funasaki 1968).</p><p>Distribution in HI.</p><p>Kaua‘i, O‘ahu, Moloka‘i, Lāna‘i, Maui, Hawai‘i.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 11 adults; O‘ahu, southern Wai‘anae Mts, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.0968&amp;materialsCitation.latitude=21.4098" title="Search Plazi for locations around (long -158.0968/lat 21.4098)">Palikea</a>; 21.4098°N, 158.0968°W; 824 m; 08 Apr. 2010; spec. PKSP 11274; P. D. Krushelnycky leg.; sweeping; PDK collection · 3 adults; O‘ahu, northern Wai‘anae Mtns., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.194&amp;materialsCitation.latitude=21.5408" title="Search Plazi for locations around (long -158.194/lat 21.5408)">Kahanahāiki Valley</a>; 21.5408°N, 158.1940°W; 585 m; 2 May 2009; spec. PKSP 3324; P. D. Krushelnycky leg.; ex Pipturus albidus; PDK collection · 1 ♂, 3 ♀; O‘ahu, northern Ko‘olau Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.0031&amp;materialsCitation.latitude=21.6419" title="Search Plazi for locations around (long -158.0031/lat 21.6419)">Pūpūkea</a>; 21.6419°N, 158.0031°W; 430 m; 08 Jul. 2014; ex Metrosideros; coll. Hi 78-14, D. M. Percy leg; DMPC · 1 adult; O‘ahu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.2745&amp;materialsCitation.latitude=21.5745" title="Search Plazi for locations around (long -158.2745/lat 21.5745)">Ka‘ena Point</a>; 21.5745°N, 158.2745°W; 30 m; 16 Apr. 2015; spec. PKSP 47458; P. D. Krushelnycky leg.; sweeping; UHIM · 1 adult; O‘ahu, central Wai‘anae Mtns, ‘ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.0935&amp;materialsCitation.latitude=21.439" title="Search Plazi for locations around (long -158.0935/lat 21.439)">Ēkahanui</a>; 21.4390°N, 158.0935°W; 658 m; 15 Dec. 2015; spec. COSP 3692; C. S. Ogura-Yamada leg.; pitfall trap; PDK collection · 8 adults; Maui, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-156.2354&amp;materialsCitation.latitude=20.7351" title="Search Plazi for locations around (long -156.2354/lat 20.7351)">Haleakalā NP; Kalahaku</a>; 20.7351°N, 156.2354°W; 2852 m; 20 Jun. 2003; spec. DS 04192; P. D. Krushelnycky leg.; ex Leptecophylla tameiameiae; PDK collection · 2 adults; Maui, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-156.2469&amp;materialsCitation.latitude=20.7466" title="Search Plazi for locations around (long -156.2469/lat 20.7466)">Haleakalā NP; Pu‘u o ‘ Ili</a>; 20.7466°N, 156.2469°W; 2390 m; 10 Jul. 2003; spec. DS 05143; P. D. Krushelnycky leg.; ex Sophora chrysophylla; PDK collection · 1 adult; Maui, Ko‘olau Forest Reserve, Flume Rd .; 20.48. 606 ° N, 156.14. 975 ° W; 20 Jun. 2006; L. Leblanc leg.; ex methyl eugenol trap; UHIM 2015.22504 · 3 adults; Maui, Haleakalā NP; 3049 m; 03 Jul. 1968; J. A. Tenorio leg.; UHIM 2015.22488 · 1 adult; Kaua‘i, Koke‘e; 16 Nov. 1968; J. A. Tenorio leg.; ex light; UHIM 2015.22482 · 1 adult; Hawai‘i, Mauna Kea, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-155.4534&amp;materialsCitation.latitude=19.7599" title="Search Plazi for locations around (long -155.4534/lat 19.7599)">Hale Pōhaku</a>; 19.7599°N, 155.4534°W; 13 Mar. 2015; J. Eiben leg.; ex Leptecophylla tameiameiae; UHIM · 2 ♀; Moloka‘i, Pālā‘au State Park; 1500 ’; 2–16 Sept. 1994; W. D. Perreira leg.; yellow sticky board trap; HDAB .</p><p>Host plants in HI.</p><p>Acacia koa, A. melanoxylon, and A. confusa ( Fabaceae).</p><p>Additional distribution and host plant records.</p><p>Native to Australia; adventive worldwide, including continental USA; Acacia spp., Albizia spp. ( Fabaceae) (Halbert and Burckhardt 2020; Suh and Choi 2020; Ouvrard 2023).</p><p>Remarks.</p><p>This species was first recorded in HI in 1966 and had spread to most of the islands within a year (Chong and Funasaki 1968). It can be found in abundance on introduced Acacia spp. and is a highly invasive pest of endemic Acacia koa . Immatures are free-living.</p><p>Genetic resources.</p><p>GenBank MG 988634 (COI), MG 988909 (cytB) [Hawaiian specimen]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/918210A95A885C0C86A4BB0E77A376EC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
069185A37B9E5270B4ADCCC4B247132D.text	069185A37B9E5270B4ADCCC4B247132D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aphalara rumicis Mally 1894	<div><p>Aphalara rumicis Mally, 1894</p><p>First detection / collection in HI.</p><p>1947 (Tuthill 1948).</p><p>Distribution in HI.</p><p>? O‘ahu (see Remarks).</p><p>Material examined.</p><p>No material located.</p><p>Host plant in HI.</p><p>Unknown.</p><p>Additional distribution and host plant records.</p><p>Continental USA, Canada; Rumex altissimus ( Polygonaceae) (Burckhardt and Lauterer 1997).</p><p>Remarks.</p><p>Tuthill (1948) reported taking a single specimen “ on a ridge overlooking Nuuanu Valley, Oahu on March 30, 1947 ”. Zimmerman (1957) referenced Tuthill’s find, listing Aphalara rumicis as the first introduced psyllid species in HI. Listings of A. rumicis for HI in Hodkinson (1983), Burckhardt and Lauterer (1997) and Nishida (2002) almost certainly cite the record from Tuthill. Tuthill’s single specimen could not be located. No other published records or specimens collected in HI could be found; hence, establishment is highly unlikely. Notably, there are several native Rumex species in HI, but no psyllids have been recorded from these.</p><p>Genetic resources.</p><p>None.</p></div>	https://treatment.plazi.org/id/069185A37B9E5270B4ADCCC4B247132D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
6F298CD612E85A6A8ABD44EDDDCC448A.text	6F298CD612E85A6A8ABD44EDDDCC448A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Australopsylla exotica Matsunaga & Percy 2026	<div><p>Australopsylla exotica Matsunaga &amp; Percy sp. nov.</p><p>Figs 7 C – H, 8 E – L, 9 A – K, 11 A – E, 12 A – K, 13 A – F</p><p>Type locality.</p><p>USA, Hawaiian Islands, O‘ahu, Hale‘iwa; 21.6440°N, 158.0355°W.</p><p>Type material.</p><p>Holotype: USA – Hawaiian Islands • ♂; O‘ahu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.0355&amp;materialsCitation.latitude=21.644" title="Search Plazi for locations around (long -158.0355/lat 21.644)">Hale‘iwa</a>; 21.6440°N, 158.0355°W; 29 Mar. 2019; A. Tateno, J. Yalemar &amp; D. Cho leg.; ex Eucalyptus robusta; coll. 2019-056; slide-mounted; BPBM . Paratypes: USA – Hawaiian Islands • 1 ♂, 3 immatures; as for holotype; slide-mounted; BPBM • 1 ♂, 4 ♀; O‘ahu; Wai‘anae Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.1602&amp;materialsCitation.latitude=21.5414" title="Search Plazi for locations around (long -158.1602/lat 21.5414)">Ka‘ala Rd., 3 rd Gate</a>; 21.5414°N, 158.1602°W; 01 Jun. 2021; K. N. Magnacca leg; ex Eucalyptus sp.; coll. 2021-125; slide-mounted; BPBM • 1 ♀; O‘ahu; southern Wai‘anae Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.1024&amp;materialsCitation.latitude=21.3954" title="Search Plazi for locations around (long -158.1024/lat 21.3954)">Pālehua</a>; 21.3954°N, 158.1024°W; 770 m; 23 Sep. 2021; K. N. Magnacca leg.; ex Eucalyptus robusta; coll. 2021-202 b; slide-mounted; BPBM ; • 2 ♀; O‘ahu; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.09808&amp;materialsCitation.latitude=21.404556" title="Search Plazi for locations around (long -158.09808/lat 21.404556)">Palikea Trail</a>; 21.404557°N, 158.098087°W; 23 Sep. 2021; J. N. Matsunaga &amp; K. N. Magnacca leg; ex Eucalyptus robusta; coll. 2021-202 a; slide mounted; BPBM .</p><p>Other material examined.</p><p>USA – Hawaiian Islands • 1 ♂, 1 ♀; O‘ahu, southern Wai‘anae Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.1024&amp;materialsCitation.latitude=21.3954" title="Search Plazi for locations around (long -158.1024/lat 21.3954)">Pālehua</a>; 21.3954°N, 158.1024°W; 770 m; 23 Sep. 2021; K. N. Magnacca leg.; ex Eucalyptus robusta; coll. 2021-202 b; BPBM • 20 immatures, O‘ahu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-157.8004&amp;materialsCitation.latitude=21.4133" title="Search Plazi for locations around (long -157.8004/lat 21.4133)">Kāne‘ohe</a>; 21.4133°N, 157.8004°W; 01 Jan. 2025; J. N. Matsunaga leg.; ex Eucalyptus robusta; BPBM • 2 ♂; O‘ahu, Wahiawā, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.05&amp;materialsCitation.latitude=21.4923" title="Search Plazi for locations around (long -158.05/lat 21.4923)">Schofield Barracks Army Base</a>; 21.4923°N, 158.0500°W; 12 Mar. 2019; A. Tateno, J. Yalemar &amp; D. Niide leg.; ex Eucalyptus robusta; coll. 2019-043; BPBM • 2 ♀; Kaua‘i, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-159.6665&amp;materialsCitation.latitude=22.038" title="Search Plazi for locations around (long -159.6665/lat 22.038)">Kōke‘e Road</a>; 22.0380°N, 159.6665°W; 760 m; 28 Apr 2025; K. N. Magnacca leg.; sweeping Eucalyptus sp.; BPBM .</p><p>Diagnosis.</p><p>Morphologically, A. exotica can be distinguished from A. carinata by the presence of sexual dimorphism in forewing coloration, and from A. revoluta, which shares sexual forewing dimorphism with A. exotica, by the type and extent of pattern in the apical portion of female forewings (Fig. 7 A – D). Additionally, A. exotica is a smaller species than the other two (Table 2). The structure of head, thorax, legs, and female terminalia are generally similar in all three species, with species-specific differences highlighted in the diagnostic key for Australopsylla adults (provided at the end of the checklist), as well as in Table 2 and the Systematics section for A. exotica (below).</p><p>Description.</p><p>Adult color and structure. General color light to mid-green, with some orange to orange-brown on the dorsum of thorax and head, and darker brown on the apical portion of the abdomen and terminalia of the female; forewing veins light brown, membrane ochreous, uniformly so in males, but with a distinctly darker brown apical third of forewing in females consisting of brown spots and a brown band around the apical margin (Fig. 13 A, B). Head narrower than, or equal to, width of thorax, deflexed downwards ca 50–60 ° from longitudinal axis of body (Figs 8 K, 13 A); coronal suture distinct; occipital sclerite developed into a preocular tubercule (Fig. 8 E); genal processes well developed and contiguous for most of the length, shorter than vertex length, broad and bluntly rounded in lateral view, in dorsal view somewhat rectangular and angular laterally (Fig. 8 E, J). Distal proboscis segment extremely short (Fig. 8 I). Antenna 10 - segmented, longer than head width (AL: HW 1.2–1.4) (Fig. 8 F), segment 3 longest, length of segments 4–9 subequal, segment 10 ~ 0.5 × length of segment 9, a single subapical rhinarium on each of segments 4, 6, 8, and 9, and one or two small rhinaria apically on segment 10 (and possibly segment 5); segment 10 with a single terminal seta as long as segment, and a short blunt seta subapically (Fig. 8 G, H). Thorax well arched (Figs 8 K, 13 A). Legs relatively short and stout (Fig. 9 A); hind leg coxa swollen into a rectangular projection with small blunt tubercle, but without distinct meracanthus (Fig. 9 A, B); metafemur short, slightly inflated sub-basally (Fig. 9 A); metatibia length subequal to head width, longer than metafemur, with prominent basal genual spine (Fig. 9 C), inflated apically with a crown of 6 (7) sclerotized spurs arranged in two groups (Fig. 9 D), one spur often more petiolate and separated from other spurs by unsclerotized setae (Fig. 9 E); metatarsal segments subequal in length, proximal segment with two lateral spurs (Fig. 9 D). Forewing length 2.7–3.7 × head width, more or less parallel-sided, slightly wider apically, apex broadly rounded (Fig. 7 C, D); costal break present, pterostigma well developed (length ~ 0.4 × forewing length); costal margin with fringe of short setae (Fig. 7 E) and minute setae bordering veins (Fig. 7 H), veins with transverse ridges (Fig. 7 G); vein Rs length ~ 0.7 × wing length, slightly curved medially towards fore margin, vein M longer than its branches, veins M + Cu and Cu subequal in length, both cells cu 1 and m 1 elongate and narrow, vein Cu 1 a strongly arched, vein Cu 1 b termination curved to wing margin (Fig. 7 C, D); membrane with surface spinules present in all cells, densely distributed throughout cells at apex (Fig. 7 F), more sparsely distributed anteriorly with spinule-free bands along veins (Fig. 7 H), lacking radular spine clusters. Hindwing broad, length ~ 0.8 × forewing length, costal margin slightly sinuate, venation prominent, apex broadly rounded (Fig. 7 C below). Male terminalia as in Fig. 11 A – E; proctiger two-segmented, total length 1.5 × length of paramere, basal segment, in lateral view more or less parallel-sided, hardly produced posteriorly, apical segment short, tubular, length ~ 0.3 × length of basal segment; subgenital plate rounded, in lateral view with dorsal margin sinuous (Fig. 11 A); paramere, in lateral view, wedge-shaped, narrowest at the base, expanding evenly and gradually to apex, exterior surface convex, interior surface concave with a row of sclerotized pegs along the apical margin, exterior surface with short simple setae, interior surface with longer, sometimes stout, simple setae, in posterior view, curving outwards in basal portion and inwards towards apex (Fig. 11 B – D). Aedeagus distal segment short, pear-shaped, expanding towards apex, apex rounded bearing a recurved spine dorsally (Fig. 11 E). Female terminalia as in Fig. 9 F, short; proctiger 0.6 × head width, dorsal surface with two raised humps anterior to and either side of anal ring, and a small, irregularly rounded pore in depression between humps anterior of anal ring, apex bluntly rounded (Fig. 9 F – I); anal ring relatively large (length 0.3 × proctiger length), heart-shaped, composed of a double row of cells that are discontinuous for a short, mid-anterior section (Fig. 9 I); subgenital plate very short (length 0.5 × proctiger length), apex blunt and slightly cleft medially (Fig. 9 K); ovipositor dorsal valves apices with a series of ridges (Fig. 9 L).</p><p>Immature color and structure. Fifth instar yellow-green, younger instars yellow-orange, all instars with dark brown to black sclerites, wing pads, legs, and terminal antennal segments (Fig. 13 C, D). Fifth instar: body ~ 1.2–1.5 × as long as wide, abdomen wider than head; wing pads protruding, lacking distinct humeral lobes; scattered, simple, short to medium-length setae throughout; thoracic and abdominal sclerites reduced but with larger, irregular shaped sclerites anterior to anus (Fig. 12 I – K). Antenna length subequal to head width, 10 - segmented with a single subapical rhinarium on each of segments 4, 6, 8, and 9, and two smaller rhinarium on segment 10 towards the apex, segment 3 longest, 6–10 subequal with 4 and 5 shorter, apex of segment 10 acute and bearing two short, unequal length setae (Fig. 12 C). Several notable appendages found on wing pads and sclerites of thorax and abdomen as follows: two digitate appendages on wing pads, one on exterior of forewing pad and one interior at junction of forewing and hindwing pads; thorax with a pair of tubercles on sternites at base of middle legs and two well developed appendages ventrally on sternites near base of hind legs; abdomen with a pair of tubercles dorsally on tergites of first abdominal segment (Fig. 12 E). Tarsi with no visible arolium, possibly a small knob-like unguitractor, claws small but well developed (Fig. 12 H). Abdomen with 3 + 3 lateral pore fields ventrally on segments 3 and 4, consisting of two uneven lines of rounded pores, anterior pores mostly on tubular stalks and posterior pores on surface (Fig. 12 A, K), third pore field smaller and adjacent to anal sclerite (Fig. 12 K). Anus terminal, acute, surrounded by stout setae with fringed apices ventrally (Fig. 12 K) and a fringed circular orifice dorsally (Fig. 12 I, J).</p><p>The lateral pore fields and most appendages described for 5 th instar are clearly visible in 1 st instar onwards (Fig. 12 G); exceptions are those on thoracic sternites, which are evident by 2 nd instar (though minute), and the exterior forewing pad appendage, which is evident by 3 rd instar. Antennae of 1 st instar 7 - segmented, 2 nd – 3 rd instars 9 - segmented and 4 th instar 10 - segmented.</p><p>Egg color and structure. Narrowly ovoid, dark brown with honeycomb surface sculpturing, short basal pedicel, apparently lacking apical filament (Fig. 9 M, N).</p><p>Measurements (mm) and ratios.</p><p>Adults (3 males, 6 females) WL 2.25–3.03; WW 0.90–1.20; HW 0.74–0.88; AL 0.98–1.16; PB 0.06–0.07; WL: WW 2.43–2.63; WL: RsL 1.30–1.43; CUR 1.14–1.43; MR 0.42–0.52; HW: VW 1.66–1.75; VW: VL 1.81–2.05; VL: GC 1.62–2.08; WL: HW 2.71–3.65; AL: HW 1.26–1.40; HW: HT 1.48–1.68; HT: HF 1.47–1.92. Male terminalia: MP 0.25–0.27; PL 0.18; AEL 0.20; PL: HW 0.24; MP: PL 1.50; PL: SH 0.90. Female terminalia: FP 0.48–0.56; FSP 0.23–0.29; RL 0.14–0.17; OVH 0.06;; FP: RL 3.25–3.43; FP: HW 0.58–0.64; FP: SP 1.93–2.22.</p><p>Immatures (5 th instar, n = 3): BL 1.84–2.27; BW 1.49–1.50; WPL 0.70–0.74; RW (orifice) 0.05–0.06; HW 0.82–0.90; AL 0.77–0.81; BL: BW 1.23–1.52; AL: HW 0.90–0.98.</p><p>Eggs (dissected from abdomen, 4 females): EL 0.23–0.28; EW 0.13–0.15; EL: EW 1.77–1.93.</p><p>Etymology.</p><p>Named for the exotic status in the Hawaiian Islands where the species was recorded first. Adjective in the nominative singular.</p><p>Distribution.</p><p>Hawaiian Islands, Kaua‘i and O‘ahu; specimens first collected on O‘ahu in 2018, and the first record on Kaua‘i in 2025. Australopsylla exotica sp. nov. is Australian in origin, with known distribution in Queensland (see Systematics below). Australopsylla exotica was first reported infesting Eucalyptus robusta trees on a military base in Wahiawā by arborists and has since been found throughout E. robusta plantings in the Wai‘anae Mountain Range (JNM observations) as well as discovered in a residential area of Kāne‘ohe on a single E. robusta tree in 2023.</p><p>Host plant and biology.</p><p>Adults and immatures were collected from Eucalyptus robusta; some adults were also collected from E. camaldulensis and unidentified Eucalyptus spp. but E. robusta appears to be the preferred host. Eggs are laid at the base of leaf bud petioles (Fig. 13 E). Emerging immatures settle on young shoots and produce lerps on the petioles and developing leaf buds (Fig. 13 E), with aggregations often in quantities that cause shoot senescence (Fig. 13 F). The lerps are brown, more or less smooth surfaced, rounded and shaped like a bivalve shell; as they enlarge they become clustered and overlapping (Fig. 13 E, F).</p><p>The two Australopsylla species described to date from Australia are recorded producing lerps and leaf roll on the leaf blade of young and mature leaves of Eucalyptus spp. (Froggatt 1900). The lerps of A. exotica appear similar to the shell-like lerps described for these species and particularly similar in shape to those of A. revoluta (figured in Froggatt 1900), although the darker brown coloration appears more similar to that described for A. carinata, while A. revoluta lerps are described as white (Froggatt 1900).</p><p>Systematics.</p><p>The mitogenomic data presented by Percy et al. (2018) places Australopsylla as sister to Creiis / Lasiopsylla, and these in turn as sister to Cardiaspina, which confirms the assessment by Tuthill and Taylor (1955) that Australopsylla “ lies between Creiis and Cardiaspina ” (Tuthill and Taylor 1955). Although not included in the study of Percy et al. (2018), the genus Blepharocosta is another endemic Australian Eucalyptus - feeding genus (Martoni et al. 2024) that is likely close to Australopsylla (Taylor 1992 a) . Since discovery of Australopsylla exotica sp. nov. in the Hawaiian Islands in 2018, a light trapping survey in Queensland, Australia listed “ Blepharocosta sp. A ” but without any morphological details (Martoni et al. 2020). Subsequent sequences for this taxon (F. Martoni pers. comm.) on GenBank (MT 375255 -57) were corrected by the original authors to “ Australopsylla sp. A ”. We have compared DNA data for “ Australopsylla sp. A ” with A. exotica and confirm these samples are conspecific, which therefore confirms the origin for A. exotica as Australia with native distribution in Queensland. Maximum genetic divergence using COI sequences of these GenBank samples from Queensland and the Hawaiian Islands is 0.6 %. Analysis of just COI data, as well as placement of A. exotica (using COI and cytB data) in the mitogenome tree using the RAxML backbone constraint analysis, shows that A. exotica is more closely related to A. revoluta, with A. carinata sister to these taxa (Fig. 15). Maximum genetic distance between A. exotica and A. revoluta (the type species of Australopsylla) is 15.3 %, and these in turn to A. carinata show 16.1–18.9 % genetic divergence. The shared sexual forewing dimorphism in A. exotica and A. revoluta also supports a closer relationship between these two species than either to A. carinata; however the overall forewing shape is shorter and broader in A. exotica, moreover A. carinata and A. revoluta share an Rs vein more curved medially towards the fore margin (only slightly so in A. exotica), longer and narrower cell m 1 (shorter and wider in A. exotica) and gradually tapering pterostigma (abruptly narrowing at apex in A. exotica) (Fig. 7). The shape of the head is similar in all three species (all have a distinct preocular tubercule), but the antennae are comparatively longer in A. revoluta (Fig. 8 A – F, Table 2). The female terminalia are similarly short in all three species, which share a proctiger with distinct anterior humps and rounded pore anterior to anal ring, but in A. carinata the anal ring does not have a discontinuous mid-anterior section (which is short in A. revoluta, longer in A. exotica), and does not extend outwards and upwards on the humps as in A. exotica and A. revoluta; in addition, the female subgenital plate has a deeper cleft apically in A. revoluta (Fig. 10 A, C); all three species have similarly serrated dorsal ovipositor valves (Figs 9, 10).</p><p>Genetic resources.</p><p>GenBank PX 507502 – PX 507503 (COI), PX 527527 – PX 527528 (cytB) [Hawaiian specimens] (Table 3); MT 375255 – MT 375257 [Australian specimens].</p></div>	https://treatment.plazi.org/id/6F298CD612E85A6A8ABD44EDDDCC448A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
DD1B2FC19724534FAA8CF4A2A48AB0AA.text	DD1B2FC19724534FAA8CF4A2A48AB0AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Australopsylla Tuthill & Taylor 1955	<div><p>Australopsylla Tuthill &amp; Taylor, 1955</p><p>Comment and diagnosis.</p><p>Australopsylla is a genus native to Australia that has had little taxonomic treatment since the original descriptions by Froggatt (1900). Froggatt described two species, Aphalara carinata Froggatt, 1900 and Rhinocola revoluta Froggatt, 1900, which were subsequently transferred to a new genus, Australopsylla, by Tuthill and Taylor (1955). One species of Australopsylla is found in the Hawaiian Islands and is described here as a new species. All species are native to Australia and occur on Eucalyptus spp. (Hollis 2004; Martoni et al. 2024), but neither of the previously described species has been reported from Eucalyptus robusta, which is considered the primary host of A. exotica Matsunaga &amp; Percy, sp. nov. (at least in the Hawaiian Islands). Australopsylla belongs to a clade that forms a well- supported group within Spondyliaspidinae (Percy et al. 2018; Fig. 15). The shape of the head in Australopsylla, with broadly truncate genal processes that are almost rectangular and a distinct preocular tubercule, is similar to Blepharocosta Taylor, Cardiaspina Crawford, Creiis Scott, and Lasiopsylla Froggatt (Tuthill and Taylor 1955; Taylor 1992 a, 1992 b). The shape of the male proctiger, including the much shorter apical portion, resembles Blepharocosta (Taylor 1992 a); the paramere that is wide apically, often with a row of blunt peg-like teeth along the apical margin, is also present in some Creiis, Dasypsylla Froggatt, Lasiopsylla and Cardiaspina (Froggatt 1900; Taylor 1992 b). All of these groups have immatures that produce lerps variously resembling bivalve-type shells. However, Australopsylla is unique among these groups in having sexually dimorphic forewings in some species, where pattern is present in females and absent in males ( Blepharocosta species may have pattern difference in male and female forewings, but patterns are either present or absent in both sexes; Taylor 1992 a).</p><p>Note.</p><p>Descriptions by Froggatt (1900) and Tuthill and Taylor (1955) lack many diagnostic characters required for reliable species identification, and no species key has previously been published. We therefore provide a key to the three described species of Australopsylla at the end of the checklist.</p></div>	https://treatment.plazi.org/id/DD1B2FC19724534FAA8CF4A2A48AB0AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
C3B8D28C24F65FD3BA14C5F191341029.text	C3B8D28C24F65FD3BA14C5F191341029.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Blastopsylla occidentalis Taylor 1985	<div><p>Blastopsylla occidentalis Taylor, 1985</p><p>First detection / collection in HI.</p><p>1993 (Beardsley and Uchida 2001).</p><p>Distribution in HI.</p><p>Kaua‘i, O‘ahu, Lāna‘i (new island record), Maui (probable new island record – see Remarks).</p><p>Material examined.</p><p>USA – Hawaiian Islands · 1 ♂, 1 ♀; Kaua‘i, Upper Wailua River, State Forestry Arboretum; 3 Jun. 1993; J. Beardsley &amp; G. Uchida leg.; ex Eucalyptus grandis; coll. 1993-234; HDAB · 8 adults; O‘ahu, Honolulu, UH Mānoa campus; 06 Jun. 1993; J. W. Beardsley leg.; ex Corymbia citriodora; UHIM · 2 adults, 2 immatures; Lāna‘i, Lāna‘i City, Koele Lodge; 01 May 1995; C. McGrath &amp; G. K. Uchida leg.; ex Eucalyptus sp.; HDAB · 1 adult; O‘ahu, southern Wai‘anae Mtns, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.0999&amp;materialsCitation.latitude=21.4134" title="Search Plazi for locations around (long -158.0999/lat 21.4134)">Palikea</a>; 21.4134°N, 158.0999°W; 859 m; 08 Apr. 2010; spec. PKSP 32083; P. D. Krushelnycky leg.; sweeping; PDK collection · 1 adult; O‘ahu, northern Wai‘anae Mtns., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.19409&amp;materialsCitation.latitude=21.54119" title="Search Plazi for locations around (long -158.19409/lat 21.54119)">Kahanahāiki Valley</a>; 21.54119°N, 158.19409°W; 580 m; 10 May 2012; spec. PKSP 26258; P. D. Krushelnycky leg.; sweeping; PDK collection .</p><p>Host plants in HI.</p><p>Eucalyptus camaldulensis, E. deglupta, E. grandis, E. robusta, Eucalyptus sp., Corymbia citriodora ( Myrtaceae).</p><p>Additional distribution and host plant records.</p><p>Native to Australia and adventive worldwide, including continental USA; Eucalyptus spp. ( Myrtaceae) (Halbert and Burckhardt 2020; Duarte et al. 2025; Martoni et al. 2025).</p><p>Remarks.</p><p>Previously recorded from Kaua‘i and O‘ahu (Beardsley and Uchida 2001). The Maui record (1994) appears in HDOA unpublished data; however, specimens could not be tracked. Immatures are free-living.</p><p>Genetic resources.</p><p>GenBank e. g., MG 988657 (COI), MG 988950 (cytB) [Norfolk Island specimen]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/C3B8D28C24F65FD3BA14C5F191341029	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
64272AB3FDD65D03A29F17483B56844D.text	64272AB3FDD65D03A29F17483B56844D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cacopsylla tobirae (Miyatake 1964)	<div><p>Cacopsylla tobirae (Miyatake, 1964)</p><p>First detection / collection in HI.</p><p>2013 (Matsunaga et al. 2019).</p><p>Distribution in HI.</p><p>O‘ahu.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 7 ♂, 7 ♀, 10 immatures; O‘ahu, Mānoa, UH; 8 Jul. 2013; S. Nelson leg.; ex Pittosporum tobira; coll. 2013-211; HDAB, USNM · 1 ♂, 1 ♀; O‘ahu, Pāwa‘a; 9 Jul. 2013; M. Tauyan leg.; ex Pittosporum tobira; coll. 2013-208; HDAB · 2 ♂, 20 immatures; O‘ahu, Mānoa, UH, East West Center; 31 May 2017; J. N. Matsunaga, K. Wong leg.; ex Pittosporum tobira; coll. 2017-126; HDAB .</p><p>Host plant in HI.</p><p>Pittosporum tobira ( Pittosporaceae).</p><p>Additional distribution and host plant records.</p><p>Native to Asia (Japan, South Korea, Taiwan); adventive in California, North and South Carolina (Miyatake 1964; Percy et al. 2012; Bertone 2016), intercepted but not considered established in Florida (Halbert and Burckhardt 2020). The host plant in HI is the only recorded host.</p><p>Remarks.</p><p>First collected in 2013, heavily infesting Pittosporum tobira from Pāwa‘a through Mānoa, Honolulu, O‘ahu areas (Matsunaga et al. 2019). Possibly now under control by parasitoids, although active removal of many of the host plants may also have impacted survival. Immatures are free-living.</p><p>Genetic resources.</p><p>GenBank e. g., OR 074490 [Florida specimen]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/64272AB3FDD65D03A29F17483B56844D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
F4CBD3AB83345004B2928726AD5E0249.text	F4CBD3AB83345004B2928726AD5E0249.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calophya rubra (Blanchard 1852)	<div><p>Calophya rubra (Blanchard, 1852)</p><p>First detection / collection in HI.</p><p>1989 (Conant et al. 1992).</p><p>Distribution in HI.</p><p>Maui.</p><p>Material examined.</p><p>No material was located in HI collections.</p><p>Host plant in HI.</p><p>Schinus molle ( Anacardiaceae).</p><p>Additional distribution and host plant records.</p><p>South America; Schinus spp. ( Anacardiaceae), at least four different species (Burckhardt and Basset 2000; Ouvrard 2023).</p><p>Remarks.</p><p>Collected throughout the Kula area from Ulupalakua to Makawao and in Wailuku (Conant et al. 1992). Immatures are found in closed galls on twigs and branchlets (Burckhardt and Basset 2000).</p><p>Genetic resources.</p><p>None.</p></div>	https://treatment.plazi.org/id/F4CBD3AB83345004B2928726AD5E0249	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
2325DDB45C955A419E45157091B75DF6.text	2325DDB45C955A419E45157091B75DF6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cryptoneossa triangula Taylor 1990	<div><p>Cryptoneossa triangula Taylor, 1990</p><p>First detection / collection in HI.</p><p>2018 (new state record; see Remarks).</p><p>Distribution in HI.</p><p>O‘ahu, Hawai‘i.</p><p>Material examined.</p><p>USA – Hawaiian Islands • 2 ♂, 1 immature; Hawai‘i, Keāhole, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-156.041&amp;materialsCitation.latitude=19.7311" title="Search Plazi for locations around (long -156.041/lat 19.7311)">Kona International Airport</a>; 19.7311°N, 156.0410°W; 22 May 2018; S. Chun &amp; M. Fukada leg.; ex Corymbia citriodora; coll. 2018-129; USNM • 2 ♂, 1 ♀, 2 immatures; same data as previous; HDAB • 2 ♀, immatures; Hawai‘i, Keāhole, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-156.041&amp;materialsCitation.latitude=19.7311" title="Search Plazi for locations around (long -156.041/lat 19.7311)">Kona International Airport</a>; 19.7311°N, 156.0410°W; 30 May 2018; C. Okamoto leg.; ex Corymbia citriodora; coll. 2018-195 USNM • 1 ♂, 1 ♀, 1 immature; same info as previous; HDAB • 3 ♂, 2 ♀; O‘ahu, ‘ Aiea Loop Trail; 16 Feb 2025, K. N. Magnacca leg.; sweeping Corymbia citriodora, BPBM .</p><p>Host plant in HI.</p><p>Corymbia citriodora ( Myrtaceae).</p><p>Additional distribution and host plant records.</p><p>Native to Australia, adventive in other Pacific regions, including western USA; Corymbia spp. ( Myrtaceae) (Percy et al. 2012; Castillo Carrillo et al. 2016; Martoni et al. 2025).</p><p>Remarks.</p><p>First discovered on lemon-scented gum trees planted around the Kona International Airport (island of Hawai‘i) while examining lerps of another species. Immatures are free-living, but often colonize the lerps of other species and were found under empty Eucalyptolyma maideni lerps. Identification confirmed by Cheryle O’Donnell (USNM). First discovered on O‘ahu in 2025.</p><p>Genetic resources.</p><p>GenBank e. g., MG 195336 (COI) [Australian specimen]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/2325DDB45C955A419E45157091B75DF6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
0E0008B0F92F50CC9674198737183469.text	0E0008B0F92F50CC9674198737183469.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ctenarytaina eucalypti (Maskell 1890)	<div><p>Ctenarytaina eucalypti (Maskell, 1890)</p><p>First detection / collection in HI.</p><p>1993 (Kumashiro et al. 2002).</p><p>Distribution in HI.</p><p>Maui, Hawai‘i.</p><p>Material examined.</p><p>USA – Hawaiian Islands • ~ 20 adults; Maui, East Maui, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-156.2766&amp;materialsCitation.latitude=20.8037" title="Search Plazi for locations around (long -156.2766/lat 20.8037)">Makawao, start of Olinda Road</a>; 20.8037°N, 156.2766°W; 830 m; 03 Jul. 2014; D. M. Percy leg.; ex Eucalyptus sp.; coll. Hi 500-14; DMPC • 1 ♂, 1 ♀; Hawai‘i, Kamuela, DLNR tree nursery; 12 Jul. 1993; J. Lum leg.; ex Eucalyptus sp.; coll. 1993-277; HDAB • 1 ♂, 1 ♀; Hawai‘i, Kamuela; 04 Aug. 1994; S. Matayoshi leg.; ex Eucalyptus globulus; coll. 1994-545; HDAB .</p><p>Host plant in HI.</p><p>Eucalyptus globulus ( Myrtaceae).</p><p>Additional distribution and host plant records.</p><p>Native to Australia, adventive in western USA, and many other locations worldwide; Eucalyptus spp. ( Myrtaceae) (Burckhardt et al. 1999; Castillo Carrillo et al. 2016; Halbert and Burckhardt 2020; Duarte et al. 2025).</p><p>Remarks.</p><p>Collected in several locations in East Maui and Kamuela, Hawai‘i. Immatures are free-living.</p><p>Genetic resources.</p><p>GenBank PX 507506 – PX 507507 (COI), PX 527531 – PX 527532 (cytB) (Table 3), SRX 1277825 (transcriptome) [Hawaiian specimens]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/0E0008B0F92F50CC9674198737183469	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
AA1D96054BC8562DA15A8BECDE0086FB.text	AA1D96054BC8562DA15A8BECDE0086FB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ctenarytaina longicauda Taylor 1987	<div><p>Ctenarytaina longicauda Taylor, 1987</p><p>First detection / collection in HI.</p><p>2010 (new state record; see Remarks).</p><p>Distribution in HI.</p><p>O‘ahu.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 1 ♂; O‘ahu, southern Wai‘anae Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.0983&amp;materialsCitation.latitude=21.4107" title="Search Plazi for locations around (long -158.0983/lat 21.4107)">Palikea</a>; 21.4107°N, 158.0983°W; 822 m; 08 Apr. 2010; spec. PKSP 12326; P. D. Krushelnycky leg.; sweeping; DMPC · 1 ♀; O‘ahu, northern Ko‘olau Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.012&amp;materialsCitation.latitude=21.6375" title="Search Plazi for locations around (long -158.012/lat 21.6375)">Pūpūkea</a>; 21.6375°N, 158.0120°W; 315 m; 08 Jul. 2014; D. M. Percy leg; ex Metrosideros; coll. Hi 76-14; DMPC · 3 ♂, 3 ♀; O‘ahu, southern Wai‘anae Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.09799&amp;materialsCitation.latitude=21.403475" title="Search Plazi for locations around (long -158.09799/lat 21.403475)">Palikea</a>; 21.403474°N, 158.097985°W; 23 Sept. 2021; J. Matsunaga &amp; K. Magnacca leg.; ex Lophostemon confertus; coll. 2021-200; HDAB, DMPC · 2 adults; O‘ahu, Wai‘anae Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.0931&amp;materialsCitation.latitude=21.4635" title="Search Plazi for locations around (long -158.0931/lat 21.4635)">Honouliuli Forest Reserve</a>; 21.4635°N, 158.0931°W; 04 Oct. 2021; K. Magnacca leg.; ex Lophostemon confertus; coll. 2021-220; USNM .</p><p>Host plant in HI.</p><p>Lophostemon confertus ( Myrtaceae).</p><p>Additional distribution and host plant records.</p><p>Native to Australia, adventive in New Zealand and California; Lophostemon confertus and possibly L. suaveolens ( Myrtaceae) (Taylor 1987 b; Hollis 2004; Percy et al. 2012; Martoni et al. 2016).</p><p>Remarks.</p><p>Collected in the northern Ko‘olau Mountains and southern Wai‘anae Mountains, O‘ahu. PDK identified specimens collected from the Palikea area in 2010, and identifications were confirmed by Cheryle O’Donnell (USNM). Immatures are free-living.</p><p>Genetic resources.</p><p>GenBank PP 218312 – PP 218313, PX 507508 (COI), PP 235413 – PP 235414, PX 527533 (cytB) [Hawaiian specimens] (Table 3). A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/AA1D96054BC8562DA15A8BECDE0086FB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
3A8CEAEB3A035C76890DDB17242A4805.text	3A8CEAEB3A035C76890DDB17242A4805.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ctenarytaina spatulata Taylor 1997	<div><p>Ctenarytaina spatulata Taylor, 1997</p><p>First detection / collection in HI.</p><p>2010 (new state record; see Remarks).</p><p>Distribution in HI.</p><p>Kaua‘i, O‘ahu, Hawai‘i.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 1 ♀; O‘ahu, southern Wai‘anae Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.0993&amp;materialsCitation.latitude=21.4142" title="Search Plazi for locations around (long -158.0993/lat 21.4142)">TNC Honouliuli Preserve, Palikea Trail</a>; 21.4142°N, 158.0993°W; 900 m; 19–20 Jul. 2010; spec. PKSP 31984; P. D. Krushelnycky leg.; sweeping; DMPC · 1 ♂, 2 ♀; O‘ahu, southern Ko‘olau Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-157.8048&amp;materialsCitation.latitude=21.3452" title="Search Plazi for locations around (long -157.8048/lat 21.3452)">Mānoa Cliff Trail</a>; 21.3452°N, 157.8048°W; 565 m; 10 Jul. 2014; D. M. Percy leg.; ex Metrosideros; coll. Hi 84-14; DMPC · 1 ♂; O‘ahu, southern Wai‘anae Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.0981&amp;materialsCitation.latitude=21.4046" title="Search Plazi for locations around (long -158.0981/lat 21.4046)">Palikea Trail</a>; 21.4046°N, 158.0981°W; 23 Sep. 2021; J. N. Matsunaga &amp; K. Magnacca leg.; ex Eucalyptus robusta; coll. 2021-201; HDAB · 3 adults; O‘ahu, southern Wai‘anae Mts., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.098&amp;materialsCitation.latitude=21.4046" title="Search Plazi for locations around (long -158.098/lat 21.4046)">Mauna Kapū, Palikea Trail</a>; 21.4046°N, 158.0980°W; 23 Sep. 2021; K. Magnacca leg.; ex Eucalyptus robusta; coll. 2021-221; USNM · 1 ♂, 2 ♀; Hawai‘i, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-155.65&amp;materialsCitation.latitude=19.859" title="Search Plazi for locations around (long -155.65/lat 19.859)">Waiki‘i</a>; 19.8590°N, 155.6500°W; 27 Jun. 2018; S. Chun leg.; ex Eucalyptus camaldulensis; coll. 2018-192; HDAB · 1 ♂, 3 ♀; Hawai‘i, Volcano; 13 May 2025; K. Magnacca leg.; sweeping Eucalyptus camaldulensis; BPBM · 1 ♀; Kaua‘i, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-159.67&amp;materialsCitation.latitude=22.114" title="Search Plazi for locations around (long -159.67/lat 22.114)">Kōke‘e State Park</a>; 22.114°N, 159.670°W; 23 Apr. 2021; K. Magnacca leg.; sweeping Dodonaea viscosa; coll. 2022-166; HDAB .</p><p>Host plant in HI.</p><p>Adults collected from Eucalyptus robusta and E. camaldulensis ( Myrtaceae).</p><p>Additional distribution and host plant records.</p><p>Native to Australia, adventive in Europe, New Zealand, South America, USA; several different Eucalyptus spp., including E. camaldulensis, E. globulus, E. grandis, E. parvifolia, E. viminalis ( Myrtaceae) (Percy et al. 2012; Halbert and Burckhardt 2020; Duarte et al. 2025).</p><p>Remarks.</p><p>Collected in southern Ko‘olau Mountains and southern Wai‘anae Mountains (O‘ahu), Waiki‘i (Hawai‘i), and Kōke‘e State Park (Kaua‘i). Identification confirmed by Cheryle O’Donnell (USNM). Immatures are free-living.</p><p>Genetic resources.</p><p>GenBank PX 507509 (COI), PX 527534 (cytB) [Hawaiian specimen] (Table 3). A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/3A8CEAEB3A035C76890DDB17242A4805	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
B5F45327EE24586A963AFA11DB49C80E.text	B5F45327EE24586A963AFA11DB49C80E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diaphorina citri Kuwayama 1908	<div><p>Diaphorina citri Kuwayama, 1908</p><p>First detection / collection in HI.</p><p>2006 (Conant et al. 2006).</p><p>Distribution in HI.</p><p>Kaua‘i, O‘ahu, Moloka‘i, Lāna‘i, Maui, Hawai‘i.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 8 ♂, 8 ♀; O‘ahu, Pāwa‘a, [HDOA]; 16 Aug. 2007; B. Kumashiro leg.; ex Murraya paniculata; coll. 2007-099; HDAB · 3 ♀; O‘ahu, Waipahu, Kunia; 4 Nov. 2008; J. N. Matsunaga leg.; ex Citrus sinensis; coll. 2008-183; HDAB</p><p>Host plant in HI.</p><p>Citrus spp. and Murraya paniculata are major hosts ( Rutaceae).</p><p>Additional distribution and host plant records.</p><p>Native to southeast Asia; adventive worldwide; Citrus spp., Murraya spp., and closely related genera ( Rutaceae) (Percy et al. 2012; Halbert and Burckhardt 2020; Ouvrard 2023).</p><p>Remarks.</p><p>This species was first detected on the island of Hawai‘i, and rapidly spread to all the other main islands (Conant et al. 2006; Matsunaga 2014). Oddly, it had been listed by Nishida (2002) but without any island distribution, and we can find no earlier records. Considered a serious pest of Citrus, particularly as the vector of citrus greening disease (huanglongbing) (Ghosh et al. 2022; Higgins et al. 2022). Currently, the disease agents, Candidatus Liberibacter spp., are not known to be present in HI. Two known parasitoids of this species, the eulophid Tamarixia cf. radiata (Waterston) and encyrtid Diaphorencyrtus aligarhensis (Shafee, Alam, &amp; Agarwal), also arrived in HI as fortuitous biocontrol agents (Matsunaga 2014). Immatures are free-living.</p><p>Genetic resources.</p><p>GenBank e. g., MG 988723 (COI), MF 426268 (complete mitochondrion) [Taiwan specimen]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/B5F45327EE24586A963AFA11DB49C80E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
1455AA6602C75CE49CC72F72CC09E877.text	1455AA6602C75CE49CC72F72CC09E877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eucalyptolyma maideni Froggatt 1901	<div><p>Eucalyptolyma maideni Froggatt, 1901</p><p>First detection / collection in HI.</p><p>2018 (new state record; see Remarks).</p><p>Distribution in HI.</p><p>O‘ahu,? Hawai‘i (see Remarks).</p><p>Material examined.</p><p>USA – Hawaiian Islands · Lerps only; Hawai‘i, Keāhole, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-156.041&amp;materialsCitation.latitude=19.7311" title="Search Plazi for locations around (long -156.041/lat 19.7311)">Kona International Airport</a>; 19.7311°N, 156.0410°W; 22 May 2018; S. Chun &amp; M. Fukada leg.; ex Corymbia citriodora; coll. 2018-249; HDAB, USNM · Lerps only; Hawai‘i, Keāhole, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-156.041&amp;materialsCitation.latitude=19.7311" title="Search Plazi for locations around (long -156.041/lat 19.7311)">Kona International Airport</a>; 19.7311°N, 156.0410°W; 30 May 2018; C. Okamoto leg.; ex Corymbia citriodora; coll. 2018-250; HDAB · 2 ♂; O‘ahu, ‘ Aiea Loop Trail; 16 Feb. 2025; K. N. Magnacca leg.; sweeping Corymbia citriodora; BPBM · 1 ♂, 2 ♀; O‘ahu, Makakilo Park; 24 May 2025; K. N. Magnacca leg.; sweeping Corymbia citriodora; BPBM .</p><p>Host plant in HI.</p><p>Corymbia citriodora ( Myrtaceae).</p><p>Additional distribution and host plant records.</p><p>Native to Australia, adventive in California; Corymbia spp. ( Myrtaceae) (Percy et al. 2012; Martoni et al. 2024).</p><p>Remarks.</p><p>In 2018, characteristic waxy lerps on host tree leaves were observed on Hawai‘i (Fig. 14). However, only immature exuviae, predatory insects, and parasitized immatures were found. No adults or live immatures were collected. The lerps were comparable with those of Eucalyptolyma maideni (Taylor 1987 a; Martoni et al. 2024). Then, in February and May 2025, adults were collected on O‘ahu. Additional sweeping of trees on which lerps were found on Hawai‘i did not locate specimens, and no lerps were visible upon re-inspection in August 2025. Immatures develop under lerps.</p><p>Genetic resources.</p><p>GenBank e. g., KY 923934, MG 132459 [Australian and New Zealand specimens]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/1455AA6602C75CE49CC72F72CC09E877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
1FB5BAAE423A5BB191138DD0E96CD3D3.text	1FB5BAAE423A5BB191138DD0E96CD3D3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euceropsylla orizabensis (Crawford 1914)	<div><p>Euceropsylla orizabensis (Crawford, 1914)</p><p>First detection / collection in HI.</p><p>2017 (Henry et al. 2018).</p><p>Distribution in HI.</p><p>O‘ahu.</p><p>Material examined.</p><p>None. Material has not been deposited at BPBM or HDAB as indicated in Henry et al. (2018) but may be deposited at USNM.</p><p>Host plant in HI.</p><p>Pithecellobium dulce ( Fabaceae).</p><p>Additional distribution and host plant records.</p><p>Native to Central America; Inga spp. ( Fabaceae) (Brown and Hodkinson 1988).</p><p>Remarks.</p><p>This is an unusual instance where an adventive species recorded for HI, as well as the host plant association (Henry et al. 2018), are not otherwise known outside the native range. As several Euceropsylla species groups are recognized that include highly similar species, and no type material was examined in the determination of HI specimens, presence of this species should be confirmed with additional collections. Immatures are free-living.</p><p>Genetic resources.</p><p>None.</p></div>	https://treatment.plazi.org/id/1FB5BAAE423A5BB191138DD0E96CD3D3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
14C12883F309565B9B6FE5CEAEAD8C50.text	14C12883F309565B9B6FE5CEAEAD8C50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glycaspis brimblecombei Moore 1964	<div><p>Glycaspis brimblecombei Moore, 1964</p><p>First detection / collection in HI.</p><p>2001 (Nagamine and Heu 2001; Matsunaga et al. 2019).</p><p>Distribution in HI.</p><p>O‘ahu, Maui, Hawai‘i.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 35 adults, immatures; Maui, Ulupalakua; 26 Apr. 2001; W. Nagamine leg.; ex Eucalyptus camaldulensis; coll. 2001-037; HDAB · 1 adult; O‘ahu, southern Wai‘anae Mountains, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.0994&amp;materialsCitation.latitude=21.4143" title="Search Plazi for locations around (long -158.0994/lat 21.4143)">Palikea</a>; 21.4143°N, 158.0994°W; 900 m; 18 Jul. 2012; spec. PKSP 36031; P. D. Krushelnycky leg.; sweeping; PDK collection · 1 adult; Maui, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-156.2477&amp;materialsCitation.latitude=20.7504" title="Search Plazi for locations around (long -156.2477/lat 20.7504)">Haleakalā NP</a>; 20.7504°N, 156.2477°W; 2302 m; 10 Jul. 2003; spec. DS 05040; P. D. Krushelnycky leg.; ex Sophora chrysophylla; PDK collection · 5 adults, nymphs; Hawai‘i, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-155.6499&amp;materialsCitation.latitude=19.859" title="Search Plazi for locations around (long -155.6499/lat 19.859)">Waiki‘i</a>; 19.8590°N, 155.6499°W; 27 Jun. 2018; S. Chun leg.; ex Eucalyptus camaldulensis; coll. 2018-191; HDAB .</p><p>Host plants in HI.</p><p>Eucalyptus camaldulensis, E. robusta, E. cf. robusta ( Myrtaceae).</p><p>Additional distribution and host plant records.</p><p>Native to Australia, adventive in other regions of the world including Africa, North, Central, and South America, and Europe; Eucalyptus spp. ( Myrtaceae) (Percy et al. 2012; Halbert and Burckhardt 2020; Martoni et al. 2024; Duarte et al. 2025).</p><p>Remarks.</p><p>Immatures develop under a whitish to greyish hemispherical lerp resembling an armored scale insect.</p><p>Genetic resources.</p><p>GenBank e. g., MW 535997, OR 068451 [European specimens]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/14C12883F309565B9B6FE5CEAEAD8C50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
03BA5A6F968C511AB6BA2301D85BBF8A.text	03BA5A6F968C511AB6BA2301D85BBF8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteropsylla cubana Crawford 1914	<div><p>Heteropsylla cubana Crawford, 1914</p><p>First detection / collection in HI.</p><p>1984 (Nakahara 1986 a; Beardsley and Uchida 1990; see Remarks).</p><p>Distribution in HI.</p><p>Kaua‘i, O‘ahu, Moloka‘i, Lāna‘i, Maui, Hawai‘i.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 1 ♂, 2 ♀; Kaua’i, Waimea; 31 Mar. 1995; R. Oyama leg.; ex corn; HDAB · 1 ♂, 2 ♀; Lāna‘i, Lāna‘i City; 28 Jun. 1984; W. Nagamine leg.; ex Leucaena leucocephala; HDAB · ~ 20 adults; Hawai‘i, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-155.6762&amp;materialsCitation.latitude=20.0108" title="Search Plazi for locations around (long -155.6762/lat 20.0108)">south of Waimea, Hwy 190</a>; 20.0108°N, 155.6762°W; 820 m; 17 Jul. 2013; D. M. Percy leg.; ex Leucaena leucocephala; coll. Hi 36-13; DMPC · 6 ♂, 10 ♀, 2 immatures; O‘ahu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.2749&amp;materialsCitation.latitude=21.5732" title="Search Plazi for locations around (long -158.2749/lat 21.5732)">Ka‘ena Point</a>; 21.5732°N, 158.2749°W; 26 m; 16 Apr. 2015; spec. PKSP 44454; P. D. Krushelnycky leg.; beating; PDK collection · 1 ♂, 1 ♀; O‘ahu, southern Wai‘anae Mtns, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.0994&amp;materialsCitation.latitude=21.4142" title="Search Plazi for locations around (long -158.0994/lat 21.4142)">Palikea</a>; 21.4142°N, 158.0994°W; 900 m; 16 Mar. 2011; spec. PKSP 16277; P. D. Krushelnycky leg.; sweeping; PDK collection · 1 ♀; O‘ahu, northern Wai‘anae Mtns., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.194&amp;materialsCitation.latitude=21.5406" title="Search Plazi for locations around (long -158.194/lat 21.5406)">Kahanahāiki Valley</a>; 21.5406°N, 158.1940°W; 588 m; 14 Dec. 2009; spec. PKSP 10937; P. D. Krushelnycky leg.; ex Pipturus albidus; PDK collection · 1 ♂; O‘ahu, northern Wai‘anae Mtns., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.1882&amp;materialsCitation.latitude=21.5408" title="Search Plazi for locations around (long -158.1882/lat 21.5408)">Pahole Natural Area Reserve</a>; 21.5408°N, 158.1882°W; 509 m; 15 Dec. 2009; spec. PKSP 7028; P. D. Krushelnycky leg.; pitfall trap; PDK collection .</p><p>Host plants in HI.</p><p>Leucaena leucocephala, Samanea saman ( Fabaceae).</p><p>Additional distribution and host plant records.</p><p>Native to Central and South America but adventive and now widely distributed in the Pacific and other tropical and subtropical regions; adventive on native Schleinitzia insularum ( Fabaceae) in the South Pacific (Muddiman et al. 1992; Claridge et al. 2014; Halbert and Burckhardt 2020; Martoni et al. 2025).</p><p>Remarks.</p><p>Recorded for HI by Beardsley and Uchida (1990) and also previously reported as Heteropsylla sp. nr. fusca (Crawford) and H. sp. poss. incisa (Šulc) (Nakahara 1986 a, 1986 b, 1988; Nagamine et al. 1992). Beardsley and Uchida (1990) reported four parasitoid species raised from immatures. Immatures are free-living.</p><p>Genetic resources.</p><p>GenBank PX 507510 (COI), PX 527535 (cytB) (Table 3), SRR 1821923 (transcriptome) [Hawaiian specimens]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/03BA5A6F968C511AB6BA2301D85BBF8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
6411BEAD37E45F7D8587390F99831668.text	6411BEAD37E45F7D8587390F99831668.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteropsylla fusca Crawford 1914	<div><p>Heteropsylla fusca Crawford, 1914</p><p>First detection / collection in HI.</p><p>1986 (Nakahara 1988).</p><p>Distribution in HI.</p><p>O‘ahu.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 1 ♂, 1 ♀; O‘ahu, St. Louis Hts.; 4 Jan. 1986; ex Acacia [= Vachellia] farnesiana; HDAB .</p><p>Host plant in HI.</p><p>Vachellia farnesiana ( Fabaceae).</p><p>Additional distribution and host plant records.</p><p>North, Central and South America; Acacia spp., Haemaloxylon campechianum, H. brasiletto ( Fabaceae) (Muddiman et al. 1992; Halbert and Burckhardt 2020; Horton et al. 2021; Ouvrard 2023).</p><p>Remarks.</p><p>First published for HI in 1988 (Nakahara 1988). Immatures are free-living.</p><p>Genetic resources.</p><p>None.</p></div>	https://treatment.plazi.org/id/6411BEAD37E45F7D8587390F99831668	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
C82C93CC37455814913DE1AC656B284E.text	C82C93CC37455814913DE1AC656B284E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteropsylla huasachae Caldwell 1941	<div><p>Heteropsylla huasachae Caldwell, 1941</p><p>First detection / collection in HI.</p><p>1975 (Beardsley 1977; Nagamine et al. 1992; see Remarks).</p><p>Distribution in HI.</p><p>Kaua‘i, O‘ahu.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 1 ♂; O‘ahu, Ewa; 24 Mar. 1977; J. W. Beardsley leg.; ex Desmanthus virgatus; UHIM 2015.22520 · 1 ♀; O‘ahu, Honolulu International Airport; 08 Feb. 1977; J. W. Beardsley leg.; ex light trap; UHIM 2015.22528 · 7 ♂; ‘ Ewa; 9 Apr. 1985; K. Murai leg.; ex Desmanthus; coll. Oa 85-77; HDAB .</p><p>Host plants in HI.</p><p>Acacia koa, Desmanthus virgatus, Samanea saman ( Fabaceae).</p><p>Additional distribution and host plant records.</p><p>Widespread in Central America and Caribbean islands, occurs as adventive in South and North America; Albizia spp. ( Fabaceae) (Brown and Hodkinson 1988; Muddiman et al. 1992; Ouvrard 2023).</p><p>Remarks.</p><p>Previously reported as Heteropsylla nr. fusca, H. sp. poss. mimosae, H. sp. prob. mimosae, H. sp. 2, H. sp. 3 (Beardsley 1977; Nakahara 1980, 1986 b, 1988; Nagamine et al. 1992). Lab tests by Nagamine et al. (1992) showed Acacia koa to be a weak, or sub-optimal, host plant. Immatures are free-living.</p><p>Genetic resources.</p><p>None.</p></div>	https://treatment.plazi.org/id/C82C93CC37455814913DE1AC656B284E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
2BC19E8293C75FEE93CFCE44CDDDD067.text	2BC19E8293C75FEE93CFCE44CDDDD067.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heteropsylla texana Crawford 1914	<div><p>Heteropsylla texana Crawford, 1914</p><p>First detection / collection in HI.</p><p>1999 (new state record; see Remarks).</p><p>Distribution in HI.</p><p>Kaua‘i, O‘ahu.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 1 ♂, 1 ♀; Kaua‘i, Nohili Dunes, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-159.7833&amp;materialsCitation.latitude=22.0621" title="Search Plazi for locations around (long -159.7833/lat 22.0621)">Barking Sands, Pacific Missile Range Facility</a>; 22.0621°N, 159.7833°W; 24 Mar. 2021; yellow pan traps placed below Prosopis [= Neltuma] pallida; K. Magnacca &amp; J. Preble leg.; coll. 2022-165; HDAB · 2 ♂; O‘ahu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.8871&amp;materialsCitation.latitude=21.3094" title="Search Plazi for locations around (long -57.8871/lat 21.3094)">Sand Island</a>; 21.3094°N, 57.8871°W; 30 Jun. 2022; M. Ramadan leg.; ex Neltuma pallida; coll 2022-176; HDAB .</p><p>Host plant in HI.</p><p>Neltuma pallida ( Fabaceae).</p><p>Additional distribution and host plant records.</p><p>Widespread in southern continental USA and Mexico; several species of Neltuma (= Prosopis) and also Pithecellobium sp. ( Fabaceae) (Muddiman et al. 1992; Percy et al. 2012; Ouvrard 2023).</p><p>Remarks.</p><p>In 2022, two individuals (1 ♂, 1 ♀) were collected in a yellow pan trap placed beneath Neltuma pallida trees at Barking Sands Beach, Pacific Missile Range Facility, Kaua‘i. Identification was confirmed by Cheryle O’Donnell (USNM). Initially thought to be a new state record until an unpublished HDOA report came to light with a citing of H. texana collected at Ke‘ehi Lagoon, O‘ahu in 1999. The report stated “ Surveys disclose Prosopis [= Neltuma] (kiawe), Pithecellobium (opiuma), Chrysopsis, Tamarix, and Acacia (koa) as hosts. ” However, no voucher specimens could be located to verify this, and it seems that the record was never publicly published. A survey of the Ke‘ehi Lagoon in 2022 confirmed establishment of H. texana on N. pallida . For this reason, we report Heteropsylla texana as a new state record here. Immatures are free-living.</p><p>Genetic resources.</p><p>GenBank e. g., MG 988760 (COI), MG 989069 (cytB) [California specimen]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/2BC19E8293C75FEE93CFCE44CDDDD067	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
6BD4AB384B695ECDBEECF6DF03ACB688.text	6BD4AB384B695ECDBEECF6DF03ACB688.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heterotrioza chenopodii (Reuter 1876)	<div><p>Heterotrioza chenopodii (Reuter, 1876)</p><p>First detection / collection in HI.</p><p>2010 (Percy et al. 2020).</p><p>Distribution in HI.</p><p>O‘ahu.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 1 ♂, 2 ♀; O‘ahu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-158.2797&amp;materialsCitation.latitude=21.5726" title="Search Plazi for locations around (long -158.2797/lat 21.5726)">Ka‘ena Point</a>; 21.5726°N, 158.2797°W; 9–19 m; 16 Apr. 2015; spec. PKSP 46529, PKSP 45882, PKSP 44470; P. D. Krushelnycky leg.; ex various Chenopodiaceae including Atriplex suberecta; coll. T 4 - PDK 980-15; BPBM, DMPC, HDAB, UHIM .</p><p>Host plants in HI.</p><p>A likely host plant is Atriplex suberecta, and possibly A. semibaccata and Chenopodium oahuense ( Amaranthaceae).</p><p>Additional distribution and host plant records.</p><p>Widespread Palearctic species known from North America since 1980 s; in addition to Atriplex spp. and Chenopodium spp. other host genera recorded include Beta and Spinacia ( Amaranthaceae) (Horton et al. 2018; Duarte et al. 2025).</p><p>Remarks.</p><p>First published records for HI in 2020 (Percy et al. 2020). Immatures are free-living.</p><p>Genetic resources.</p><p>GenBank MT 162454 – MT 162455 (COI), MT 176418 – MT 176419 (cytB) [Hawaiian specimens]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/6BD4AB384B695ECDBEECF6DF03ACB688	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
2CE791C9825B579198D59C92CB61A884.text	2CE791C9825B579198D59C92CB61A884.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptynoptera sulfurea Crawford 1919	<div><p>Leptynoptera sulfurea Crawford, 1919</p><p>First detection / collection in HI.</p><p>1977 (Beardsley 1981).</p><p>Distribution in HI.</p><p>Kaua‘i, O‘ahu, Moloka‘i, Maui, Hawai‘i.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 1 ♀; O‘ahu, Hickam AFB; 23 Nov. 1977; J. W. Beardsley leg.; ex Calophyllum; UHIM 2015.22572 · 2 ♂, 2 ♀, 20 immatures; same data as previous; HDAB · 1 ♂, 3 ♀, 10 immatures; Kāne‘ohe, Ho‘omaluhia Botanical Garden; 24 Jul. 2013; W. Nagamine leg.; ex Calophyllum brasiliense; coll. 2013-225; HDAB · 6 ♂, 4 ♀; Maui, Lahaina, Lahainaluna School; 30 Mar. 1993; R. Sher leg.; ex Calophyllum inophyllum; coll. 1993-109; HDAB · 5 ♂, 5 ♀; Kaua‘i, Līhu‘e Airport; 25 Mar. 1993; C. Campbell leg.; ex Calophyllum inophyllum; coll. 1993-107; HDAB · 1 ♀; Moloka‘i, Halawa Valley; 61 m; 03 Mar. 1995; W. D. Perreira leg.; yellow sticky trap; UHIM 2015.22578 .</p><p>Host plants in HI.</p><p>Calophyllum inophyllum, and possibly C. brasiliense ( Calophyllaceae).</p><p>Additional distribution and host plant records.</p><p>Broadly distributed in Asia-Pacific region; Calophyllum spp. ( Calophyllaceae) (Martin and Hollis 1992; Martoni and Brown 2018).</p><p>Remarks.</p><p>First reported for HI by Beardsley (1981). Although this species is widespread, and in many Pacific regions, such as French Polynesia and the Cook Islands, it is considered likely native – the Hawaiian occurrence is considered adventive due to the absence of HI records prior to 1970 s. Immatures are free-living.</p><p>Genetic resources.</p><p>GenBank e. g., KY 293709 (COI), MG 989229 (complete mitochondrion) [New Caledonia specimen]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/2CE791C9825B579198D59C92CB61A884	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
A42BF994A8DB5B4BAC874F54A334A289.text	A42BF994A8DB5B4BAC874F54A334A289.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macrohomotoma gladiata Kuwayama 1908	<div><p>Macrohomotoma gladiata Kuwayama, 1908</p><p>First detection / collection in HI.</p><p>2022 (Matsunaga 2022).</p><p>Distribution in HI.</p><p>O‘ahu.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 10 ♂, 10 ♀, 50 immatures; O‘ahu, Honolulu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-157.8975&amp;materialsCitation.latitude=21.3376" title="Search Plazi for locations around (long -157.8975/lat 21.3376)">Māpunapuna</a>; 21.3376°N, 157.8975°W; 3 Nov. 2002; M. Ramadan leg.; ex Ficus microcarpa; coll. 2022-231; HDAB, USNM · 2 ♂, 30 immatures; O‘ahu, Honolulu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-157.864&amp;materialsCitation.latitude=21.3289" title="Search Plazi for locations around (long -157.864/lat 21.3289)">Kapālama</a>; 21.3289°N, 157.8640°W; 4 Nov. 2022; D. Cho, J. Yalemar leg.; ex Ficus microcarpa; coll. 2022-234; HDAB · 1 ♂, O‘ahu, UH Mānoa campus; 11 May 2025; K. N. Magnacca leg.; on Ficus benjamina; BPBM · Immatures; O‘ahu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-157.803&amp;materialsCitation.latitude=21.4067" title="Search Plazi for locations around (long -157.803/lat 21.4067)">Kāne‘ohe</a>; 21.4067°N, 157.8030°W; 5 Sep. 2025; J. Matsunaga leg.; ex Ficus microcarpa; JNM 2025-01 .</p><p>Host plant in HI.</p><p>Ficus microcarpa and possibly F. benjamina (only adults collected) ( Moraceae).</p><p>Additional distribution and host plant records.</p><p>Native to China, India, Indonesia, Japan, Malaysia, and Taiwan; adventive in California and Mediterranean countries in southern Europe and North Africa; Ficus spp. ( Moraceae) (Yang 1984; Mifsud and Porcelli 2012; Rung 2016; Ouvrard 2023).</p><p>Remarks.</p><p>First recorded for HI as Pauropsylla apsylloides by Crawford (1919), but only based on a single damaged specimen (possibly dead on arrival off a recently imported plant), and the taxon was not recorded thereafter as established. However, subsequent authors continued to cite the original record (e. g., Hodkinson 1983, 1986; Hollis and Broomfield 1989; Nishida 2002; Burckhardt et al. 2018). Subsequently placed in Macrohomotoma and later synonymized with M. gladiata by Li (2011). The first true establishment of M. gladiata was in 2022 when a heavily infested F. microcarpa tree was reported to HDAB (Matsunaga 2022). Immatures are free-living but often produce abundant white flocculence.</p><p>Genetic resources.</p><p>GenBank e. g., MG 988795 (COI), MG 989108 (cytB) [Taiwan specimen]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/A42BF994A8DB5B4BAC874F54A334A289	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
2DBBDA7478B9507F9EBAF60BEA8C6D9B.text	2DBBDA7478B9507F9EBAF60BEA8C6D9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mesohomotoma hibisci (Froggatt 1901)	<div><p>Mesohomotoma hibisci (Froggatt, 1901)</p><p>First detection / collection in HI.</p><p>2005 (Matsunaga et al. 2019).</p><p>Distribution in HI.</p><p>O‘ahu, Maui.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 12 ♂, 8 ♀, 32 immatures; O‘ahu, Makiki; 20 May 2005; W. Nagamine leg.; ex Hibiscus tiliaceus; coll. 2005-044; HDAB, USNM · 9 ♂, 10 ♀, 30 immatures; Maui, Kahului, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-156.4812&amp;materialsCitation.latitude=20.891" title="Search Plazi for locations around (long -156.4812/lat 20.891)">UH Maui College Campus</a>; 20.8910°N, 156.4812°W; 3 Jan. 2020; M. Fukada leg.; ex Hibiscus tiliaceus; coll. 2020-025; HDAB .</p><p>Host plant in HI.</p><p>Hibiscus tiliaceus ( Malvaceae).</p><p>Additional distribution and host plant records.</p><p>Native to Australia and Pacific Islands; known from Africa, Asia; Hibiscus spp. ( Malvaceae) (Hollis 1987; Martoni et al. 2025).</p><p>Remarks.</p><p>Immatures are free-living but often produce abundant white flocculence.</p><p>Genetic resources.</p><p>GenBank e. g., MG 989231 (complete mitochondrion), KY 294172 [French Polynesia specimens]. A number of other accession numbers for this species can be found on GenBank.</p></div>	https://treatment.plazi.org/id/2DBBDA7478B9507F9EBAF60BEA8C6D9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
2E50B2CA403B556C9310F5D86C8E5F22.text	2E50B2CA403B556C9310F5D86C8E5F22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paurocephala wilderi Crawford 1927	<div><p>Paurocephala cf. wilderi Crawford, 1927</p><p>First detection / collection in HI.</p><p>1925 (Crawford 1927 b).</p><p>Distribution in HI.</p><p>Moloka‘i.</p><p>Material examined.</p><p>USA – Hawaiian Islands · 1 ♂; Moloka‘i, Kamiloloa; 3200 ft; 20 Dec. 1925; E. H. Bryan leg.; collected on “ pokeweed ”; BPBM .</p><p>Host plant in HI.</p><p>Uncertain (see remarks).</p><p>Additional distribution and host plant records.</p><p>Endemic to Samoa; adults only, no immatures, but all collections are from Ficus ( Moraceae) (Mifsud and Burckhardt 2002).</p><p>Remarks.</p><p>This HI record has previously been referred to as Paurocephala sp. with native status uncertain (Crawford 1927 b; Zimmerman 1948; Percy 2017 a). Here we consider P. cf. wilderi to be adventive in HI. On re-examination of the single male specimen, we find the species belongs in the psylloptera - group sensu Mifsud and Burckhardt (2002). Characters, including only a shallow horn on metascutellum, forewing venation including a wide cell cu 1 and long pterostigma, and reduced number (only 2–3) of stout spurs laterally along the metatibia, would assign this taxon to P. wilderi . However, as only a single male specimen was collected in 1925 with no additional specimens recorded for HI, slide preparation of the genitalia (not clearly visible in the dry specimen) would be needed to confirm species designation. Notably, Crawford, who described P. wilderi (in Crawford 1927 a), did not place the Hawaiian specimen in that taxon when he examined it (Crawford 1927 b; Zimmerman 1948). This may be partly due to the obscured genitalia, but also to the brown apical coloration on the forewing that characterizes P. wilderi being extremely faint in the HI specimen; additionally, the forewing shape in the HI specimen is wider apically than material from Samoa (as shown in Mifsud and Burckhardt 2002). Although the plant from which the specimen was collected is recorded as “ pokeweed ”, it is unclear whether this refers to the native pokeweed, Phytolacca sandwicensis ( Phytolaccaceae) or the introduced species P. octandra; both co-occur on Moloka‘i (Wagner et al. 1999). Regardless, Phytolacca is unlikely to be a host plant; the psylloptera - group are only known from plants in the urticoid Rosales with all adults of P. wilderi collected on Ficus in Samoa (Mifsud and Burckhardt 2002). As there are no native species of Ficus in HI, but many introduced taxa, Ficus was the most likely vehicle of introduction into HI. Immature biology unknown.</p><p>Genetic resources.</p><p>None.</p></div>	https://treatment.plazi.org/id/2E50B2CA403B556C9310F5D86C8E5F22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Matsunaga, Janis N.;Magnacca, Karl N.;Krushelnycky, Paul D.;Percy, Diana M.	Matsunaga, Janis N., Magnacca, Karl N., Krushelnycky, Paul D., Percy, Diana M. (2026): Checklist of the adventive psyllids (Hemiptera, Psylloidea) of the Hawaiian Islands, including new state records, identification key, and description of a new species of Australopsylla. ZooKeys 1279: 1-56, DOI: 10.3897/zookeys.1279.178071
