identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
AB026FBBAFC45B74B458DB555D929ABB.text	AB026FBBAFC45B74B458DB555D929ABB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tryblionella calida (Grunow) Stroganov	<div><p>Tryblionella calida (Grunow) Stroganov</p><p>Figs 1, 2, 3, 4</p><p>Description.</p><p>LM (Fig. 1). Valves linear-lanceolate with wedge-shaped, slightly protruding apices. Valve dimensions (n = 17): length range 38–61 μm, width range 8.6–10.6 μm at the constricted middle part. Valve surfaces only slightly undulate from side to side, appearing almost flat. Axial sternum absent. Transapical ribs parallel, continuous across valve surface, 15–17 in 10 μm. Raphe system eccentric, located along one side of valve. Fibulae visible, short, rectangular to square (e. g., Fig. 1 E, H, arrows), 9–10 in 10 μm. Ratio of transapical-ribs density to fibula density ca. 1.6.</p><p>SEM (Figs 2, 3, 4). Transapical ribs elevated, continuously across valve surface (Fig. 2 A, B). Striae uniseriate, 34–36 in 10 μm (measured on SEM images, n = 3). Marginal ridge is a high standing silica lamella, composed of a bottom part with long slit-like fenestrae (labelled in Fig. 2 E, F) and a top part, smooth with a plain margin (Fig. 2). Distal mantle shallow, lacking ornamentations (Figs 2, 4 A, C, D). Central nodule present (Fig. 2 C, D, arrow; Fig. 4 B, arrow), distal raphe fissures curved (Fig. 2 E, F, curved arrows). A longitudinal row of short bar-like ribs is produced along the raphe canal wall from pole to pole (Fig. 2 D – F, double-headed arrows). Row of poroids is produced in the raphe canal wall on the side of valve surface (Fig. 2 E, F, arrows). Inner openings of areolae rounded, rimmed. Proximal mantle very deep so that it often hides the short fibulae (Fig. 3 A). Proximal mantle composed of two parts: advalvar part (composed of bar-like ribs and narrow, rectangular openings, labelled in Fig. 3 D), and abvalvar part (hyaline without ornamentations, labelled in Fig. 3 B). A row of poroids produced in the raphe canal wall on the side of the proximal mantle (Fig. 3 B – D, arrows). Inner central raphe fissure straight, central nodule helictoglossa-like (Fig. 4 B, arrow). Fibulae squat, some apically elongated (Fig. 4 C, D, arrows). Helictoglossae small (Fig. 4 C, D, curved arrows).</p><p>Comments.</p><p>The basionym of Tryblionella calida is Nitzschia calida Grunow (in Cleve and Grunow 1880). Grunow’s protologue, written in German, reads: “ N. calida Grunow. Klein, Schaalen linear länglich, meist in der Mitte schwach verengt, mit keilförmigen, schwach vorgezogenen Spitzen. Längsfalte breit, sehr flach, oft kaum sichtbar. Querstreifen 17–19 in 0.01 mm., Kielpunkte klein, rund, circa 10 in 0.01 mm. Länge 0.034–0.043 mm., Schaalenbreite 0.009–0.01 mm. In den Thermen von Ofen in Ungarn. […] ” (Cleve and Grunow 1880, p. 75). The specimens examined fit Grunow’s protologue in valve morphology (e. g., especially the almost flat valve face), overlapping valve sizes (38–61 μm vs. 34–43 μm in length, 8.6–10.6 μm vs. 9–10 μm in width), overlapping density of transapical ribs (15–17 in 10 μm vs. 17–19 in 10 μm), overlapping fibula density (9–10 in 10 μm vs. 10 in 10 μm), and their occurrence in freshwater habitats.</p><p>Outside Europe, Kociolek (2011) provided observations of T. calida based on specimens collected from America and provided the following description: “ […] A narrow, indistinct fold runs along the longitudinal axis of the valve. The fold is barely undulated or may even appear not to rise or taper from the main valve surface. […]. ” Thus, Kociolek also noted the almost flat valve faces in T. calida .</p><p>SEM observations of T. calida are available in Metzeltin et al. (2005, plate 109) and Bertolli et al. (2020, Fig. 5 D – G), but these studies provided only low-magnification images and did not show many details of the frustule ultrastructure. In this paper, the ultrastructure of its marginal ridge, proximal and distal mantles, and fibulae is described and illustrated in detail for the first time. Thus, based on the protologue and the observations presented here, the following diagnostic criteria are proposed for T. calida . These include a linear-lanceolate valve outline with wedge-shaped, slightly protruding apices, a very slightly undulate valve surface, elevated transapical ribs continuous across the valve face and short bar-like ribs connecting the raphe canal wall, uniseriate striae, squat and sometimes apically elongated fibulae, and a high marginal ridge composed of a bottom part with elongated slit-like fenestrae and a solid top part with a plain margin.</p><p>Stroganov (1927) made the valid transfer of Nitzschia calida Grunow (in Cleve and Grunow 1880) to Tryblionella before Mann (in Round et al. 1990). Therefore, the combination Tryblionella calida (Grunow) D. G. Mann should not be used.</p></div>	https://treatment.plazi.org/id/AB026FBBAFC45B74B458DB555D929ABB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ma, Ya-Lun;Yu, Can-Ping;Liu, Bing;Rioual, Patrick;Ma, Yi-Dan;Jiang, Xiao-Yu	Ma, Ya-Lun, Yu, Can-Ping, Liu, Bing, Rioual, Patrick, Ma, Yi-Dan, Jiang, Xiao-Yu (2026): Ultrastructure of three species of Tryblionella (Bacillariophyceae) found in freshwater habitats in Hunan Province, China, including descriptions of two new species. PhytoKeys 278: 259-277, DOI: 10.3897/phytokeys.278.204336
F32B08FA71E152C98764F400FD061EF6.text	F32B08FA71E152C98764F400FD061EF6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tryblionella dongtingensis Bing Liu & Rioual 2026	<div><p>Tryblionella dongtingensis Bing Liu &amp; Rioual sp. nov.</p><p>Figs 5, 6, 7</p><p>Holotype.</p><p>The specimen circled on the slide DIA 2026002, illustrated here as Fig. 5 A, deposited in the Herbarium of Jishou University (JIU), China. Registration: http://phycobank.org/106074 .</p><p>Type locality.</p><p>China • Hunan Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=112.79222&amp;materialsCitation.latitude=29.318611" title="Search Plazi for locations around (long 112.79222/lat 29.318611)">East Dongting Lake</a>. Sampling location (29°19'7"N, 112°47'32"E, 39 m a. s. l.) in the lake littoral zone, collected by Bing Liu, 5 March 2023.</p><p>Description.</p><p>LM (Fig. 5). Valves linear-lanceolate, slightly constricted in the middle on both sides, with wedge-shaped apices. Valve dimensions (n = 30): length range 33–65 μm, width range 10.3–12.1 μm at constricted middle parts. Valve surface undulated from side to side. Axial sternum absent. A longitudinal fold is present along the valve midline (e. g., Fig. 5 G, double-headed arrow), dividing the valve face into two non-coplanar halves. Transapical ribs more evident on one half of valve surface than on the other when observed at a given focal plane, 9–14 in 10 μm, parallel except approaching apices, where they become radiate. Raphe system eccentric, located along one side of valve only. Fibulae visible, short, rectangular (e. g., Fig. 5 E, F, arrows), 6–9 in 10 μm.</p><p>SEM (Figs 6, 7). Transapical ribs elevated, some run across the valve face continuously whereas others terminate at the middle longitudinal fold (Fig. 6 D, E, curved arrows). Striae uniseriate, 38–40 in 10 μm (measured on SEM images, n = 3). A longitudinal row of bar-like ribs produced along the raphe canal wall, from pole to pole (Fig. 6 C – F, arrows). Central nodule present (Fig. 6 C, curved arrow). Distal raphe fissures curved (Fig. 6 D). Marginal ridge is a high standing silica lamella, composed of a bottom part with long slit-like fenestrae (Fig. 6 F, two curved arrows) and a top part with a serrate margin (Fig. 6 C – E, double-headed arrows). Distal mantle very shallow, lacking ornamentations (Fig. 6 F). Inner openings of areolae rounded, rimmed (Fig. 7 D). Proximal mantle markedly deep so that it often hides the short fibulae (Fig. 7 A). Proximal mantle composed of two parts: advalvar part (composed of bar-like ribs and narrow, rectangular openings), and abvalvar part (hyaline without ornamentations). Fibulae squat, often hidden by the deep proximal mantle (Fig. 7 A, B, D, curved arrow), helictoglossa small (Fig. 7 D, arrow).</p><p>Etymology.</p><p>Named after Dongting Lake, where the species was found.</p><p>Vernacular name.</p><p>The Chinese name for Tryblionella dongtingensis is “洞庭湖盘杆藻.”</p><p>Ecology and distribution.</p><p>Tryblionella dongtingensis sp. nov. was collected from the surface sediments in the littoral zone of East Dongting Lake. Associated species included Craticula ambigua D. G. Mann, C. cuspidata D. G. Mann, Dorofeyukea kotschyi Kulikovskiy, Kociolek, Tusset &amp; T. Ludwig, Gyrosigma dongtingense Bing Liu &amp; Rioual, G. kuetzingii (Grunow) Cleve, Hantzschia amphioxys Grunow, Surirella angusta Kützing, S. suecica Grunow, Tryblionella calida, T. hungarica (Grunow) Frenguelli, Ulnaria rhombus D. M. Williams, and several species of Navicula Bory, Nitzschia Hassall, and Sellaphora Mereschkowsky. The following environmental parameters were measured in the field: conductivity was 494 ± 0.9 μS cm - 1, pH was 8.4 ± 0.06, and water temperature was 27.8 ± 0.3 ° C. The environmental variables and the composition of the diatom assemblage suggest alkaline and mesotrophic to eutrophic conditions. So far, T. dongtingensis has been found only at the type locality.</p><p>Comments.</p><p>Tryblionella dongtingensis sp. nov. is characterized by its linear-lanceolate valve outline with wedge-shaped apices, undulate valve surface from side to side, elevated transapical ribs and short bar-like ribs connecting the raphe canal wall, uniseriate striae, short fibulae, and a high marginal ridge composed of a bottom part with elongate slit-like fenestrae and a solid top part with an undulate margin. A review of the literature revealed few species similar to T. dongtingensis sp. nov. The most similar species to T. dongtingensis is T. calida, and they can coexist in the same habitat. The two species, however, differ in their morphometry. Tryblionella calida has narrower valves (8.6–10.6 μm vs. 10.3–12.1 μm), a higher density of transapical ribs (15–17 in 10 μm vs. 9–14 in 10 μm), and a higher density of fibulae (9–10 in 10 μm vs. 6–9 in 10 μm). In addition, T. calida has almost flat valve surfaces (Fig. 1), whereas T. dongtingensis has an undulate valve surface (Fig. 5). Furthermore, the marginal ridge in T. calida has a plain top margin (see Fig. 2), whereas in T. dongtingensis it has an undulate top margin (see Fig. 6).</p></div>	https://treatment.plazi.org/id/F32B08FA71E152C98764F400FD061EF6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ma, Ya-Lun;Yu, Can-Ping;Liu, Bing;Rioual, Patrick;Ma, Yi-Dan;Jiang, Xiao-Yu	Ma, Ya-Lun, Yu, Can-Ping, Liu, Bing, Rioual, Patrick, Ma, Yi-Dan, Jiang, Xiao-Yu (2026): Ultrastructure of three species of Tryblionella (Bacillariophyceae) found in freshwater habitats in Hunan Province, China, including descriptions of two new species. PhytoKeys 278: 259-277, DOI: 10.3897/phytokeys.278.204336
B874CEDBD961506BA8789FBFC975CA9E.text	B874CEDBD961506BA8789FBFC975CA9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tryblionella hunanensis Bing Liu & Rioual 2026	<div><p>Tryblionella hunanensis Bing Liu &amp; Rioual sp. nov.</p><p>Figs 8, 9, 10, 11, 12</p><p>Holotype.</p><p>The specimen circled on the slide DIA 2026003, illustrated here as Fig. 8 A, deposited in the Herbarium of Jishou University (JIU), China. Registration: http://phycobank.org/106075.</p><p>Type locality.</p><p>China • Hunan Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=113.59861&amp;materialsCitation.latitude=26.918888" title="Search Plazi for locations around (long 113.59861/lat 26.918888)">Yongle River (a tributary of the largest river, Xiang River, in Hunan Province, China)</a>. Sampling location (26°55'08"N, 113°35'55"E, 119 m a. s. l.) in a riffle, collected by Bing Liu, 24 March 2024.</p><p>Description.</p><p>LM (Fig. 8). Valves panduriform with cuneate to acute apices. Valve dimensions (n = 31): length range 37–53 μm, width range 8.9–13.2 μm at constricted middle parts. Valve surfaces slightly undulated from side to side. Axial sternum absent. Transapical ribs parallel, across valve face continuously, 9–12 in 10 μm. Raphe system eccentric, located only along one side of valve. Fibulae visible, rectangular to square (e. g., Fig. 8 D, two arrows), 7–10 in 10 μm.</p><p>SEM (Figs 9, 10, 11, 12). Valves panduriform with elevated transapical ribs continuously across valve surface (Figs 9, 10). Dense silica granules present on the valve surface, striae uniseriate, 34–36 in 10 μm (measured on SEM images, n = 3). A longitudinal row of bar-like ribs produced along the raphe canal wall, from pole to pole (Fig. 9 B – D, double-headed arrows). Marginal ridge is a standing silica lamella, composed of a bottom part with a row of pores produced along the junction between the distal mantle and the base of the marginal ridge, and a solid top part with a distinctly lobed margin (Figs 9, 10). Four to seven longitudinal grooves are produced in the raphe canal wall on the side of valve (Figs 9 A, 10 A, 10 B). Distal mantle shallow, lacking ornamentations (Fig. 10 C – F). Distal raphe fissures curved (Figs 10 C, 11 B, curved arrows). Proximal mantle markedly deep, composed of two parts: advalvar part (composed of bar-like ribs and narrow, rectangular openings), and abvalvar part (hyaline part without ornamentations) (Fig. 11). A row of poroids produced in the raphe canal wall on the side of the proximal mantle (Fig. 11 B – D, arrows). Inner openings of areolae rounded, rimmed. Fibulae squat, mostly apically elongated (Fig. 12), two median fibulae farther apart (Fig. 12 A, B, arrows). Helictoglossae small (Fig. 12 C, D, curved arrows).</p><p>Etymology.</p><p>Named after Hunan Province, where the species was found (specific locality is Yongle River).</p><p>Vernacular name.</p><p>The Chinese name for Tryblionella hunanensis is “湖南盘杆藻.”</p><p>Ecology and distribution.</p><p>Tryblionella hunanensis sp. nov. was collected from the mud sediments on the stone surfaces in the Yongle River. There are numerous associated species, with the most abundant being Ctenophora hunanensis Bing Liu &amp; Rioual, Cymbella turgidula Grunow, Grunowia tabellaria (Grunow) Rabenhorst, Gyrosigma kuetzingii (Grunow) Cleve, Hippodonta capitata (Ehrenberg) Lange-Bertalot, Metzeltin &amp; Witkowski, Nitzschia palea (Kützing) W. Smith, Tryblionella calida, Ulnaria fanjingensis Bing Liu, and several species of Cocconeis Ehrenberg, Fragilaria Lyngbye, Gomphonella Rabenhorst, Luticola D. G. Mann, Navicula Bory, Planothidium Round &amp; Bukhtiyarova, Sellaphora Mereschowsky, and Surirella Turpin. The following environmental parameters were measured in the field: conductivity was 111.7 ± 0.8 μS cm – 1, pH was 7.6 ± 0.04, and water temperature was 22.3 ± 0.3 ° C. The environmental variables and the composition of the diatom assemblage suggest alkaline and mesotrophic conditions. So far, T. hunanensis has been found at the type locality and in East Dongting Lake.</p><p>Comments.</p><p>Tryblionella hunanensis sp. nov. is characterized by its panduriform valve outline with cuneate to acute apices, slightly undulate valve surface, elevated transapical ribs continuous across the valve face and short bar-like ribs connecting the raphe canal wall, uniseriate striae, squat, mostly apically elongated fibulae, and a marginal ridge composed of a bottom part with a row of pores produced along the junction between the distal mantle and the base of the marginal ridge and a solid top part with a distinctly lobed margin. A review of the literature revealed few species similar to T. hunanensis sp. nov. The species most similar to T. hunanensis are T. calida and T. dongtingensis . All three species have similar valve outlines and overlapping morphometric ranges, and they can coexist in the same habitat in East Dongting Lake. However, T. calida has almost flat valve surfaces (Fig. 1), whereas T. hunanensis has a more undulate valve surface (Fig. 13). Furthermore, the solid top part of the marginal ridge in T. calida is straight (see Fig. 2), whereas in T. hunanensis it is distinctly lobed (see Fig. 14). Tryblionella hunanensis also differs from T. dongtingensis in the structure of the raphe canal wall on the valve side (in the former, it is grooved, whereas in the latter, it is smooth) and the marginal ridge (in the former, its bottom part has no slit-like fenestrae, whereas slit-like fenestrae are present in the latter).</p></div>	https://treatment.plazi.org/id/B874CEDBD961506BA8789FBFC975CA9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Ma, Ya-Lun;Yu, Can-Ping;Liu, Bing;Rioual, Patrick;Ma, Yi-Dan;Jiang, Xiao-Yu	Ma, Ya-Lun, Yu, Can-Ping, Liu, Bing, Rioual, Patrick, Ma, Yi-Dan, Jiang, Xiao-Yu (2026): Ultrastructure of three species of Tryblionella (Bacillariophyceae) found in freshwater habitats in Hunan Province, China, including descriptions of two new species. PhytoKeys 278: 259-277, DOI: 10.3897/phytokeys.278.204336
