taxonID	type	description	language	source
30736F332005386A4BC6FA827EDFFA8B.taxon	diagnosis	Diagnosis. Cells cup, r 5 and m 3 closed and petiolate, petioles with variable size. M 2 vein variable, complete or incomplete. Wing coloration variable: hyaline or smoky without spots in the crossveins; hyaline or smoky with faint spots or well-marked spots in the crossveins or with characteristic macule or patch patterns of yellow or yellowish coloration. Thoracic ornamentation variable, absent, inconspicuous longitudinal stripes or well-marked longitudinal stripes. Abdomen completely without ornamentation or with bands and / or tufts of contrasting setulae. Clypeus usually protuberant, markedly developed in some species (clypeal index between 0.5 – 1.2). Proboscis long to very long, between two and six times the height of the frons, extremely long in some species (approximately 12 times the height of the frons). Ocellar triangle usually not very prominent. Frontal callus absent, some species with a well-marked or indistinct patch on the frons. Frons generally slightly divergent below or with parallel sides.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33200D38744BC6FA0D7F4DFC53.taxon	description	(Fig. 6 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33200D38744BC6FA0D7F4DFC53.taxon	materials_examined	Type locality. Panama, Cerro Campana.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33200D38744BC6FA0D7F4DFC53.taxon	diagnosis	Diagnosis. Body predominantly brown. Thorax with inconspicuous stripes. Wing smoky brown. Thorax and abdomen with black setulae dense short. Legs yellow.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33200D38744BC6FA0D7F4DFC53.taxon	description	Redescription. Female. Body length: 11.4 mm, wing length: 12 mm. Head. Frons slightly divergent below (divergence index 0.8), frontal index 2.2. Frons without patch. Vertex dark brown with slightly dense orange pruinosity and long, dense black setulae, prominent ocellar triangle, orange ocelli. Frons brown and subcallus with slightly dense orange pruinosity, frons with black setulae that are not very long and not very dense. Parafacial brown with whitish pruinosity and dense, long black setulae. Gena covered with white pruinosity and dense, long, whitish-yellow setulae. Clypeus brown with whitish pruinosity and sparse black setulae, clypeal index 0.8. Antenna orange. Scape and pedicel with yellowish pruinosity and dense long black setulae. Palpus dark brown, first segment with dense long yellowish setulae. Proboscis shiny dark brown, about 2 x as high as frons. Thorax. Scutum and scutellum brown with slightly dense grayish pruinosity and dense black setulae. Thorax with inconspicuous stripes. Notopleuron slightly lighter in relation to the scutum and with a tuft of dense, long black setulae. Anepisternum, katepisternum and anepimeron covered with white pruinosity and dense long yellow setulae. Postalar callus with tuft of yellowish-brown setulae. Legs yellow. Femora with long dense black and yellow setulae. Tibiae and tarsi with short dense black and yellow setulae. Wing smoky brown, brown veins. Abdomen. Tergites yellowish-brown with short dense black setulae, last tergites darkened. Sternum yellow with small circular black integument areas and sparse black and yellowish setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33200D38744BC6FA0D7F4DFC53.taxon	distribution	Distribution. Panama, Peru *.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33200D38744BC6FA0D7F4DFC53.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the MCZ (examined through photographs). Additional material examined. 1 ♀, “ Peru, Cusco, Quincemil, 13 º 13 ’ 03.4 ” S 70 º 43 ’ 40 ” W, 633 m, 23 – 31. viii. 2012, Malaise, JA Rafael, RR Cavichioli, DM Takiya / Scione sp. 1 ” (INPA).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33200D38744BC6FA0D7F4DFC53.taxon	discussion	Comments. In the original description, Fairchild stated that by running the key of Kröber (1930) this species would key out to S. rufipes, whose type is lost, and the original description is quite succinct. However, according to Fairchild, S. ablusus differs from S. rufipes in being darker, with mainly black setulae, and without golden setae on tergites 1, 6 and 7. Kröber compares S. rufipes to S. huancabambae, which was also described by him and whose type is lost. Fairchild argues that S. ablusus differs from S. huancabambae in having narrower frons. Thus, according to Fairchild, the only species similar to S. ablusus is S. rufescens, from which it differs in smaller size, wider frons, different antennae and palpus. Scione rufescens has dark brown scape and pedicel, an orange-brown flagellum, and very tapered dark brown palpus with black setulae on both segments. In S. ablusus, the antenna is orange and the palpus is dark brown with dense yellowish setulae on the first segment. Furthermore, S. rufescens has dense coppery setulae on the abdomen, which are not present in S. ablusus.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201338764BC6FC2A79E5FD3B.taxon	description	(Fig. 7 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201338764BC6FC2A79E5FD3B.taxon	materials_examined	Type locality. Peru, Huánuco, Hacienda Éxito, 1150 m.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201338764BC6FC2A79E5FD3B.taxon	diagnosis	Diagnosis. Body yellowish-brown. Scutum with three white longitudinal stripes. Wing with a characteristic spot pattern, smoky brown at the apex and anterior region, with yellowish spots in cells r 1, r 2 + 3 and r 5 and a hyaline posterior region. Veins brown with faint brown spots on crossveins.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201338764BC6FC2A79E5FD3B.taxon	description	Redescription. Female. Body length: 11.1 mm, wing length: 11.3 mm. Head. Frons slightly divergent below (divergence index 0.9), frontal index 2.0. Dark brown patch in the frons, almost touching eyes. Vertex dark brown with long, dense black setulae, slightly prominent ocellar triangle, brown ocelli. Frons brown with whitish pruinosity and dense, long black setulae. Subcallus brown with orange pruinosity. Parafacial and gena brown with whitish pruinosity. Gena with dense long white setulae and some black setulae. Clypeus brown with whitish pruinosity and long, sparse black setulae, clypeal index 0.6. Scape and pedicel orange-brown with long dense black setulae. Flagellum orange, last flagellomeres slightly darkened. Palpus dark brown quite tapered; first segment with dense long white setulae in the proximal portion and dense, long black setulae in the distal portion; second orange segment on the dorsal margin and with short, sparse black setulae. Proboscis shiny dark brown, about three times the height of the frons. Thorax. Scutum brown with dense black setulae that not very long and some orange setulae and with three white longitudinal stripes; sublateral stripes wide and quite evident, the median stripe is narrow and less evident. Scutellum brown with long sparse black setulae on the posterior margin. Notopleuron with tuft of dense long black and orange setulae. Anepisternum with a tuft of dense, long black and white setulae. Katepisternum and anepimeron with tuft of dense long white setulae. Postalar callus with a tuft of dense long white setulae and some equally long black setulae. Legs dark brown. Fore and middle tibiae slightly orange. Femora with long, dense black setulae. Tibiae and tarsi with short, dense black setulae. Wing smoky brown at the apex and anterior region, with yellowish spots in the middle of cells r 1 and r 2 + 3 (“ window ”); posterior region hyaline. Veins brown with faint brown spots on crossveins. Abdomen. Tergites yellowish-brown with short, dense black setulae, last tergites darkened. Central triangles of whitish setulae on the posterior margin of tergites 1 to 6. Lateral margin of tergites with dense yellowish setulae. Sternites with whitish and black setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201338764BC6FC2A79E5FD3B.taxon	distribution	Distribution. Colombia, Peru (Huánuco).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201338764BC6FC2A79E5FD3B.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the CAS (examined through photographs). Additional material examined. 1 ♀, “ Peru, Dpto. [Departamento] Pasco: Camino a Cacazú, VIII: 21: 68 (5; 800.), Pedro Hocking ” (UNMSM).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201338764BC6FC2A79E5FD3B.taxon	discussion	Comments. Scione acris is similar to S. albifasciata, S. incompleta and S. maculipennis (Fig. 8). It differs from S. albifasciata by the spot pattern on the wings, this species has smoky brown wing with brown veins and faint brown spots in the crossveins, whereas S. acris has smoky brown wing at the apex and on the anterior region, with yellowish spots on the r 1, r 2 + 3 and r 5 cells and the posterior region is hyaline. It differs from S. incompleta because this species has the thorax with five slightly evident yellowish longitudinal stripes, whereas in S. acris the longitudinal stripes are white and more evident. It differs from S. maculipennis in that this species has unspotted frons and orange brown femora with long, dense black setulae, yellowish brown tibiae and tarsi with short, dense black setulae, and darkened hind tibiae and tarsi, whereas S. acris has slightly divergent frons below, a dark brown patch on the frons, almost touching the eyes, and dark brown legs.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201138774BC6F8E1795EF8DD.taxon	description	(Fig. 9 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201138774BC6F8E1795EF8DD.taxon	materials_examined	Type locality. “ Colombia ” (as Nouvelle Grenade).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201138774BC6F8E1795EF8DD.taxon	diagnosis	Diagnosis. Body predominantly brown. Thorax with white longitudinal stripes. Wing smoky brown with yellowish spots in the middle of cells r 1 and r 2 + 3 (“ window ”) and lighter areas in the middle of cells m 1 and m 2.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201138774BC6F8E1795EF8DD.taxon	description	Redescription. Female. Body length: 11 – 12.6 mm (X = 11.9 mm, N = 3), wing length: 11.6 – 12.5 mm (X = 12.2 mm, N = 3). Head. Frons slightly divergent below, divergence index 0.8 – 0.9 (X = 0.8, N = 3), frontal index 1.7 – 2.0 (X = 1.9 mm, N = 3). Dark brown patch on the frons, almost touching the eyes and extending to the vertex. Vertex dark brown with long dense black setulae, slightly prominent ocellar triangle, dark brown ocelli. Frons brown with yellowish pruinosity and long dense black setulae and some sparse orange setulae. Subcallus brown with yellowish pruinosity and some sparse black setulae. Parafacial brown with some sparse black setulae. Gena covered with white pruinosity and dense long white setulae. Clypeus brown with sparse black setulae and whitish pruinosity on the sides, clypeal index 0.7 – 0.9 (X = 0.8, N = 3). Scape and pedicel orange brown with short black setulae. Flagellum orange, last flagellomeres slightly darkened. Palpus dark brown, quite tapered; first segment with dense long white and black setulae; second segment with sparse short black setulae. Proboscis dark brown, about 3 x the height of the frons. Thorax. Scutum and scutellum brown. Scutum with dense not very long black setulae and some orange setulae. Scutellum with long black setulae. Scutum with white longitudinal stripes. The median stripe is narrow and inconspicuous, the sublateral stripes slightly wider and well-marked. The sublateral and lateral stripes are connected by a narrow white stripe at the transverse suture. Notopleuron with tuft of black and orange setulae. Anepisternum with a tuft of dense, long, black and white setulae. Katepisternum and anepimeron with tuft of dense, long white setulae. Postalar callus with a tuft of dense long white and yellowish setulae. Coxae and trochanters orange brown with whitish pruinosity and dense long black setulae. Fore and mid femora and tibiae yellowish brown, hind femora and tibiae orange brown. Femora with long dense black setulae, tibiae with short dense black setulae. Tarsi brown with short, dense black setulae. Wing smoky brown with yellowish spots in the middle of cells r 1 and r 2 + 3 (“ window ”) and lighter areas in the middle of cells m 1 and m 2. Abdomen. Tergites 1 and 2 yellowish-brown, other tergites darkened, tergites 2 – 5 with central triangles of whitish setulae on the central posterior margin. All tergites with short dense black setulae. Sternites with short dense whitish setulae. Sternites 1 and 2 yellowish, other sternites darkened.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201138774BC6F8E1795EF8DD.taxon	distribution	Distribution. Panama *, Colombia, Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201138774BC6F8E1795EF8DD.taxon	materials_examined	Type material examined. Syntype, ♀, deposited at the NHMUK (examined through photographs). Additional material examined. 1 ♀, “ Panama, Chiriqui, Alto Lino, 4000, 25 june 1965, H. G. Real / Herman G. Real Collection / Cornelius B. Philip Collection, Donated to California Academy of Sciences / same data but 25 june — II frons shorter palp / same data — II front. short – er [shorter] snout / Scione albifasciata (Macq.) Det. C. B. Philip 72 ” (CAS); 1 ♀, “ Las Nubes (4500 ft.), Dept. Magdalena, Colombia, S. A. / Dec. / H. H. Smith Acc. No. 1740 / Cornelius B. Philip Collection Donated to California Academy of Sciences / Comp. to Type Scione albifasciata C. B. Philip Ag. 53 Macq. Agrees exc (aver), minor diffs. see notes / Scione albifasciata (Macquart) det. Krolow, T. K., 2012 ” (CAS); 1 ♀, “ Colombia 77.96, Dpt. Cauca aprox. 40 km Popayan Monchique 3000 m, 19 - 20 July 1997, M. A. Tidwell / Scione albifasciata Macq., det. Fairchild 1988 / MPEG DIP 12006462 ” (MPEG); 1 ♀, “ Tambo to Starosa, Cascho Crespejo, Ecuador, VIII- 1 - 85 / Pres. by Mrs. Y. Mexia Collector / Cornelius B. Philip Collection, Donated to California Academy of Sciences / ♀ Ven. little this [?] is [illegible word] coll / Scione albifasciata (Macq.) Det. C. B. Philip ” (CAS).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201138774BC6F8E1795EF8DD.taxon	discussion	Comments. See S. flavescens.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201638714BC6FF4A7A4FFC27.taxon	materials_examined	Type locality. Peru, Urubamba, 3000 m.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201638714BC6FF4A7A4FFC27.taxon	description	Original description. Länge 10 mm, Fühler 1.5 mm, Rüssel 6.6 mm, Flügellänge 11 mm, Flügelbreite 3 mm. Gleicht flavohirta ausserordentlich. Stirn schwarz, oben zimtbraun, unten hellgrau tomentiert, lang zart schwarz behaart, am Scheitel lang vornübergeneigt. Ocellen dunkel. Stirndreieck weiss grau mit 2 bräunlichen Makeln. Untergesicht bräunlich olive, in der Mitte schwarz, etwas glänzend, zart weiss tomentiert, die Mitte zart schwarz behaart. 1. und 2. Fühlerglied gelbbraun, lang abstehend zart schwarz behaart, der Rest brennend rotgelb, die Spitze schwarz. Augenbehaarung lang, oben bräunlich, unten weiss. Bart dicht, schneeweiss. Hinterkopf weissgrau, weiss behaart. Thorax weissgrau mit 4 braunen Striemen. Behaarung lang zart abstehend schwarz und anliegend weiss. An den Seiten lang weiss behaart. Schildchen weissgrau, auffallend lang weiss behaart. Brustseiten und Hüften weiss tomentiert, dicht weiss behaart, p bleich rotgelb, f lang seidig weiss behaart, t kurz schwarz. Schwinger und Schüppchen gelblich, die Flocke weiss. Flügel ganz hyalin, Adern zart, gelblich und bräunlich. Hinterleib wie bei flavohirta, aber alle helle Behaarung weiss.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201638714BC6FF4A7A4FFC27.taxon	distribution	Distribution. Peru.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201638714BC6FF4A7A4FFC27.taxon	materials_examined	Holotype ♀. Destroyed (Kröber Collection).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201638714BC6FF4A7A4FFC27.taxon	discussion	Comments. According to the original description, S. albohirta is similar to S. flavohirta. However, S. albohirta has yellowish red flagellum with black last flagellomeres, whereas in S. flavohirta, the flagellum is orange with slightly darkened last flagellomere. In S. albohirta the gena setulae is white, in S. flavohirta the gena setulae is yellowish.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201638714BC6FBFB7BB7F95C.taxon	description	(Fig. 10 A – D)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201638714BC6FBFB7BB7F95C.taxon	materials_examined	Type locality. Costa Rica, Cartago, Monumento Nacional Guayabo, Turrialba, 1100 m.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201638714BC6FBFB7BB7F95C.taxon	diagnosis	Diagnosis. Body brown. Thorax with two very evident yellowish sublateral stripes and a slightly conspicuous median stripe. Dense white setulae on the gena. Tufts of white setulae on the gena, pleura and postalar callus contrasting with the dark body. Wing smoky. Abdomen dark brown; tergites with median triangles of yellowish setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201638714BC6FBFB7BB7F95C.taxon	distribution	Distribution. Costa Rica (Cartago).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201638714BC6FBFB7BB7F95C.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the MNCR (examined through photographs).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201638714BC6FBFB7BB7F95C.taxon	discussion	Comments. The type of S. albopilosa, as reported in the catalog of Coscarón & Papavero (2009), was originally deposited at the INBio (Instituto Nacional de Biodiversidad), but is currently at the MNCR (Museo Nacional de Costa Rica). According to the original description, this species resembles S. costaricana, from which it differs, in having much longer and narrower second segment of the palpus, snow-white setulae on the head and thorax, dark brown scutellum with black setulae, grayish hyaline wing, abdominal tergites with less distinct yellowish patches.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201438784BC6FF4A782FFE7F.taxon	description	(Figs 11 A – D, 12 A – F, 13 A – E, 14 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201438784BC6FF4A782FFE7F.taxon	materials_examined	Type locality. Mexico, Oaxaca.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201438784BC6FF4A782FFE7F.taxon	diagnosis	Diagnosis. Body yellowish-brown. Thorax with two inconspicuous yellowish-brown sublateral stripes. Wing hyaline with orange brown veins and no spots on crossveins.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201438784BC6FF4A782FFE7F.taxon	description	Redescription. Female. Body length: 9.5 – 10.6 mm (X = 10.1 mm, N = 6), wing length: 10.1 – 10.9 mm (X = 10.5 mm, N = 6). Head. Frons slightly divergent below, divergence index 0.7 – 0.8 (X = 0.8, N = 5), frontal index 2.3 – 3.2 (X = 2.6, N = 5). Frons without patch. Vertex brown with orange pruinosity and long, not very dense black setulae, slightly prominent ocellar triangle, orange-brown ocelli. Frons and subcallus brown with orange pruinosity, frons with sparse black setulae. Parafacial and gena orange, gena with whitish pruinosity and dense and long whitish setulae. Clypeus orange with sparse black setulae, clypeal index 0.7 – 0.9 (X = 0.8, N = 5). Scape and pedicel orange brown with black setulae, flagellum orange. Palpus orange; first segment with long dense setulae on the anterior margin; second segment tapered and with some short orange setulae. Proboscis shiny dark brown, about 4 x the height of the frons. Thorax. Scutum and scutellum brown with orange pruinosity and sparse black and orange setulae. Scutum with two slightly evident yellowish-brown sublateral stripes, from each of which an equally inconspicuous stripe departs in the region of the transverse suture. Notopleuron with a small tuft of black and whitish setulae. Anepisternum, katepisternum and anepimeron covered with white pruinosity and not very dense whitish setulae. Postalar callus with tuft of orange setulae. Wing hyaline with orange brown veins. Legs yellowish, femora with sparse yellowish and black setulae, tibiae and tarsi with short dense black setulae and some not so dense yellowish setulae. Abdomen. Tergites yellowish-brown with sparse black and yellow setulae. Sternites yellowish-brown with yellowish setulae. Male (Fig. 12). Like the female, except for the holoptic condition, straight palpus and longer, denser body setulae. Clypeus less developed than that of the female, clypeal index 0.5.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201438784BC6FF4A782FFE7F.taxon	distribution	Distribution. Southern Mexico to Costa Rica, Venezuela, Colombia, Peru.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201438784BC6FF4A782FFE7F.taxon	materials_examined	Type material examined. Lectotype of S. aurulans, ♀, deposited at the MNHUB (examined through photographs); Lectotype of S. lurida, ♀, deposited at the MNHUB (examined through photographs); Syntype of Diclisa misera, ♀ deposited at the NHMUK (examined through photographs). Additional material examined. 1 ♀, “ Mexico, Veracruz, 5 km, NNW. of Ocotal Chico, Rain Forest, 975 m, 24 - III- 1965, G. N. Ross, coll. / Cornelius B. Philip Collection, Donated to California Academy of Sciences / Scione aurulans (Wiedemann), det. C. B. Philip, 1977 / Scione aurulans (Wiedemann), det. Krolow, T. K., 2012 ” (CAS); 3 ♀, “ Mexico, Veracruz, 5 km, NNW. of Ocotal Chico, Rain Forest, 3110 m, 8. IV. 1965, G. N. Ross, coll. / Cornelius B. Philip Collection, Donated to California Academy of Sciences / Scione aurulans (Wiedemann), det. Krolow, T. K., 2012 ” (CAS); 1 ♀, “ Mexico, Veracruz, 1.6 km N. Sotepan 426 m, 29 - III- 1965, G. N. Ross, coll., Secondary rain For. / Cornelius B. Philip Collection, Donated to California Academy of Sciences / Scione aurulans (Wiedemann), det. Krolow, T. K., 2012 ” (CAS); 1 ♀, “ Palenque, Chis, Mex. 28 Ma. 51 / G. B. Fairchild det. / Scione aurulans Wied., det. Fairchild 1988 / MPEG DIP 12006463 ” (MPEG); 1 ♀, “ El Limon, AR, Venezuela, 450 m, 22 - xi- 60 / Em besia / M. Telbez. col / Propiedad del MIZA Fac. Agronomia UCV Maracay, Venezuela ” (MPEG); 1 ♀, “ E. E. Cataurito Villa de Cura, Venezuela Aragraa, 26 - x- 19 [.] 78 / J. L. Garcia col. / Propiedad del MIZA Fac. Agronomia UCV Maracay, Venezuela ” (MPEG); 1 ♂, “ Cortada del Guayaba Miranda, Venezuela, 28 - x- 68 / D. Villasmil J. B. Teran / Propiedad del MIZA Fac. Agronomia UCV Maracay, Venezuela [photographed specimen] ” (MPEG); 1 ♂, “ Serrania El Avila, 1400 ms., D. F. Venezuela, 3. x [.] 1958 / Los Venador [?] / C. J. Rosales col. / Propiedad del MIZA Fac. Agronomia UCV Maracay, Venezuela ” (MPEG); 1 ♂, “ Rancho Grande AR. Venezuela, 1100 m., 7. xii. 51 / F. Fernandez Y. col. / Propiedad del MIZA Fac. Agronomia UCV Maracay, Venezuela ” (MPEG); 1 ♂, “ Peru, Cusco, Tito, 2000 M, 13 º 31 ’ 35.5 ’’ S 70 º 53 ’ 55 ’’ W, 25 – 30. viii. 2012 / Arm. Malaise, J. A. Rafael; R. R. Cavichioli & D. M. Takiya / Scione sp. Det. Henriques, 2016 ” (INPA).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201438784BC6FF4A782FFE7F.taxon	discussion	Comments. The subspecies Scione lurida was used to refer to specimens collected in South America (Colombia and Venezuela), whereas S. aurulans was restricted to Central America. However, comparing the photos of the S. aurulans and S. lurida types, we did not see any striking differences, except for the body coloration and clypeus structure. S. lurida appears to be darker than S. aurulans and has less developed clypeus (Fig. 13), both common variations within the same species. Kröber (1934) synonymized S. lurida under S. aurulans; however, Fairchild (1966) and later works reverted to treating it as valid subspecies. We agree with the synonymy proposed by Kröber (1934) and consider S. lurida a revised synonym of S. aurulans.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201F38784BC6FE077E62F83B.taxon	description	(Fig. 15 A – D)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201F38784BC6FE077E62F83B.taxon	materials_examined	Type locality. Ecuador, Morona-Santiago, Limón, 900 m.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201F38784BC6FE077E62F83B.taxon	diagnosis	Diagnosis. Body yellowish-brown. Thorax with two conspicuous yellowish sublateral stripes. Wing slightly smoky with brown veins and a slightly conspicuous yellowish patch on the anterior margin. Light yellow marginated veins and brown spots on crossveins. Legs bicolored, dark brown femora, yellow tibiae and tarsi.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201F38784BC6FE077E62F83B.taxon	description	Redescription. Female. Body length: 12.0 – 12.8 mm (X = 12.3 mm, N = 6), wing length: 12.0 – 12.5 mm (X = 12.2 mm, N = 6). Head. Frons divergent below, divergence index 0.7 – 0.9 (X = 0.8, N = 6), frontal index 1.8 – 2.1 (X = 1.9, N = 6). Brown patch on the frons, almost touching the eyes and extending to the vertex. Vertex dark brown with slightly dense yellowish pruinosity and long, dense black setulae, slightly prominent ocellar triangle, dark brown ocelli. Frons and subcallus brown with yellowish pruinosity, frons with dense, long black setulae. Parafacial brown with yellowish pruinosity and long, sparse black setulae. Gena covered with yellowish pruinosity and dense, long, whitish yellow setulae. Clypeus brown with whitish pruinosity on the sides and long, not very dense black setulae, clypeal index 0.7 - 0.9 (X = 0.8, N = 6). Scape and pedicel brown with whitish pruinosity and dense, long black setulae. Flagellum orange, last segment brown. Palpus dark brown; first segment with dense, long black and yellowish setulae; second segment orange on the dorsal margin. Proboscis shiny dark brown, about 2 x the height of the frons. Thorax. Scutum and scutellum dark brown. Scutum with dense long orange setulae and two conspicuous yellowish sublateral stripes that extend to the region of the transverse suture where they become slightly longer. Posterior margin of the scutum with two yellowish patches. Scutellum with dense, long black setulae on the posterior margin. Notopleuron yellowish with tuft of dense, long, black and yellowish setulae. Anepisternum and katepisternum with tuft of dense long black setulae. Anepimeron with a tuft of dense long yellowish setulae. Postalar callus with a tuft of yellowish white setulae. Legs bicolored. Femora dark brown with long, dense black setulae. Tibiae and tarsi yellow with short, dense yellow setulae and some black setulae, tibiae and hind tarsi slightly darkened. Wing slightly smoky with brown veins. Light yellow marginated veins and brown spots on crossveins. Yellowish patch on the anterior region, in the region of cells r 1 and r 2 + 3. Short appendix at the fork of vein R 4 + 5. Abdomen. Tergites 1 to 3 yellow with black central patch. Tergites 4 to 7 dark brown. Tergites 1 to 6 with posterior band of pruinosity and yellowish setulae and with tufts of yellowish-white setulae on the central posterior margin. Lateral margin of tergites with orange setulae. Sternites yellowish-brown with dense, not very long black and yellowish setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201F38784BC6FE077E62F83B.taxon	distribution	Distribution. eastern Andes of Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201F38784BC6FE077E62F83B.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the AMNH (examined through photographs); Paratype, ♀ (specimen physically examined): “ Limon, 900 m, E. Ecuador, II- 1948, Z Muller / Cornelius B. Philip Collection Donated to California Academy of Sciences / Paratype ♀ Scione bilineata n. sp., Det. C. B. Philip ” (CAS). Additional material examined. 4 ♀, “ E. Loja, Ecuador, 2800 m., XI. 1970, L. E. Peña col. / MZSP ” (MZUSP); 1 ♀, “ Ecuad. [or] Azuay, Maylas 3192 m., Octubre 09, J. Buestán / MECN-EM-DIP 646 / INABIO / Donation INABIO 2019 / Scione equatoriensis ” (INPA).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201F38784BC6FE077E62F83B.taxon	discussion	Comments. Scione bilineata is similar to S. distincta. It differs in that S. distincta has thorax with five faint yellowish longitudinal stripes, whereas S. bilineata has thorax with two more conspicuous yellowish longitudinal stripes. In addition, S. bilineata has central black patch on tergites 1 to 3, which is absent in S. distincta.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201D387A4BC6FF4A7AB6F986.taxon	description	(Fig. 16 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201D387A4BC6FF4A7AB6F986.taxon	materials_examined	Type locality. Colombia, Valle, Lago Calima, 8 km below the Calima Lake Dam.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201D387A4BC6FF4A7AB6F986.taxon	diagnosis	Diagnosis. Body yellowish-brown. Median thoracic stripe and sublateral stripes connected at the posterior margin of the scutum and forming a central yellowish patch. Wing smoky brown with yellowish veins and very faint brown spots on crossveins. Fore legs and mid legs bicolored, femora brown and tibiae and tarsi yellow.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201D387A4BC6FF4A7AB6F986.taxon	description	Redescription. Female. Body length: 11.6 – 12.1 mm (X = 11.8 mm, N = 4), wing length: 11.4 – 12.5 mm (X = 11.7 mm, N = 4). Head. Frons slightly divergent below, divergence index 0.8 – 0.9 (X = 0.9, N = 4), frontal index 2.2 – 2.3 (X = 2.2, N = 4). Dark brown patch on frons, almost touching eyes. Vertex dark brown with slightly dense yellowish pruinosity and long, dense black setulae, slightly prominent ocellar triangle, dark brown ocelli. Frons brown with dense yellowish pruinosity and dense, long black setulae. Subcallus brown with dense yellowish pruinosity on the sides. Parafacial brown with whitish pruinosity and dense, long black and yellowish setulae. Gena covered with whitish pruinosity and dense, long yellowish setulae. Clypeus brown with slightly dense yellowish pruinosity and dense, long black setulae, clypeal index 0.7 – 0.8 (X = 0.7, N = 4). Scape and pedicel brown with yellowish pruinosity and long, dense black setulae. Flagellum orange-brown, last flagellomeres darkened. Palpus dark brown; first segment with dense, long yellowish setulae and some black setulae; second segment quite tapered and with a slightly orange dorsal margin. Proboscis shiny black, about 2 x the height of the frons. Thorax. Scutum orange brown with dense black and yellowish setulae and conspicuous longitudinal stripes. The median stripe and sublateral stripes connect at the posterior margin of the scutum and form a central yellowish patch. The sublateral and lateral stripes connected by a less evident yellow stripe at the transverse suture. Scutellum black with long dense black setulae. Notopleuron with tuft of dense long black and yellowish setulae. Anepisternum with a tuft of dense long black and yellowish setulae. Katepisternum and anepimeron with a tuft of dense long yellowish setulae. Postalar callus with a tuft of dense long yellowish setulae. Bicolored legs. Coxae, trochanters and femora dark brown with long, dense black setulae. Yellow fore and mid tibiae and tarsi, tibiae with short, dense yellow setulae and tarsi with short, dense black and yellow setulae. Hind tibiae and tarsi yellowish-brown with short, dense black and yellow setulae. Wing smoky brown with yellowish veins, very faint brown spots on crossveins and yellowish spots on the r 1 and r 2 + 3 cells. Abdomen. All tergites with short, dense black setulae. Tergites 1 and 2 yellowish and with posterior band of yellowish setulae. Tergites 1 to 5 with tufts of yellowish setulae on the central posterior margin. Sternites yellowish-brown with long, dense black and yellowish setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201D387A4BC6FF4A7AB6F986.taxon	distribution	Distribution. Colombia (Antioquia *, Valle).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201D387A4BC6FF4A7AB6F986.taxon	materials_examined	Type material examined: Holotype, ♀, deposited at the FSCA (examined through photographs). Additional material examined. 1 ♀, “ Colombia: Medelin, no date, Balfour coll. / pres. Welcome Trust BM. 1990 - 107 / Scione? sp. nov. nr. brevibeccus det. J. E. Chainey, 1992 / NHMUK 014428010 ” (NHMUK); 1 ♀, “ Colombia: Medelin, no date, Balfour coll. / pres. Welcome Trust BM. 1990 - 107 / NHMUK 014428011 ” (NHMUK); 1 ♀, “ idem / NHMUK 014428012 ” (NHMUK); 1 ♀, “ idem / NHMUK 014428013 ” (NHMUK).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33201D387A4BC6FF4A7AB6F986.taxon	discussion	Comments. See S. youngi.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202338454BC6FF4A7926FE7F.taxon	description	(Fig. 17 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202338454BC6FF4A7926FE7F.taxon	materials_examined	Type locality. Ecuador, Baños.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202338454BC6FF4A7926FE7F.taxon	diagnosis	Diagnosis. Body yellowish-brown. Antennae yellow, last flagellomeres black. Thorax with two conspicuous yellow sublateral stripes and a barely visible yellowish median stripe. Posterior region of the scutum with a central yellowish patch. Legs yellow. Smoky wing with brown spots on crossveins. Thorax and abdomen with dense coppery setulae. Tergites 3 to 5 with triangles of white setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202338454BC6FF4A7926FE7F.taxon	distribution	Distribution. Ecuador, Peru, Bolivia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202338454BC6FF4A7926FE7F.taxon	materials_examined	Type material examined. Holotype, ♂, deposited at the MNHUB (examined through photographs).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202338454BC6FF4A7926FE7F.taxon	discussion	Comments. See S. obscurefemorata.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238454BC6FE067865FBCF.taxon	description	(Fig. 18 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238454BC6FE067865FBCF.taxon	materials_examined	Type locality. Peru, Cuzco, Callanga.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238454BC6FE067865FBCF.taxon	diagnosis	Diagnosis. Body yellowish-brown. Antenna, palpus and frons brown. Thorax brown with yellowish longitudinal stripes. Gena and pleura with whitish hairiness. Wing smoky brown. Legs yellowish-brown. Tergite 1 yellow with dense black setulae. Tergite 2 yellow with dense yellow setulae. Other tergites yellowish-brown with dense black setulae, tergites 3 to 5 with triangles of yellow setulae on the central posterior margin.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238454BC6FE067865FBCF.taxon	distribution	Distribution. Peru (Cuzco).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238454BC6FE067865FBCF.taxon	materials_examined	Type material examined. Lectotype, ♀, deposited at the MNHUB (examined through photographs).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238454BC6FE067865FBCF.taxon	discussion	Comments. Scione cingulata is quite distinguishable from other species by having the first two yellow abdominal tergites contrasting with the other dark tergites.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238414BC6FB567E0AFEEB.taxon	description	(Figs 19 A – F, 20 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238414BC6FB567E0AFEEB.taxon	materials_examined	Type locality. Peru, Callanga.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238414BC6FB567E0AFEEB.taxon	diagnosis	Diagnosis. Body yellowish-brown. Thorax with conspicuous white longitudinal stripes. Wing hyaline with yellow orange pterostigma and no spots on crossveins.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238414BC6FB567E0AFEEB.taxon	description	Redescription. Female. Body length: 10.0 – 10.1 mm, wing length: 9.5 – 9.6 mm. Head. Frons divergent below (divergence index 0.7 – 0.8), frontal index 1.7 – 1.8. Frons with triangular dark brown patch not extending to the apex. Posterior region of the scutum with central yellowish patch. Vertex dark brown with dense black setulae, ocellar triangle prominent, ocelli dark brown. Frons and subcallus brown with whitish yellow pruinosity, frons with sparse black setulae. Parafacial and gena brown with yellowish pruinosity, gena with dense and long white setulae. Clypeus brown with whitish pruinosity and sparse black setulae, clypeal index 0.6 – 0.7. Scape and pedicel orange brown with yellowish pruinosity and not very dense black setulae, flagellum orange brown with the last flagellomeres darkened. First segment of palpus dark brown with dense white setulae and some black setulae, second segment orange brown with sparse black setulae. Proboscis shiny dark brown, about 2 x the height of the frons. Thorax. Scutum and scutellum brown with dense black setulae. Scutum with evident longitudinal white stripes, the sublateral and lateral stripes connected by a white stripe at the transverse suture, the median stripe narrower and less evident than the others. Notopleuron with tuft of white and black setulae. Anepisternum with tuft of black setulae, katepisternum and anepimeron covered with pruinosity and white setulae. Postalar callus with tuft of white setulae. Coxae dark brown with grayish pruinosity and long black setulae. Trochanters and femora dark brown with long, dense black setulae. Fore and mid tibiae orange brown with dense black setulae. Hind tibiae brown with dense black setulae. Tarsi brown with dense black setulae. Wing hyaline with orange-brown veins. Pterostigma yellow orange. Abdomen. Tergites yellowish-brown with dense black setulae. Tergites 1 to 6 with posterior band of pruinosity and white setulae. Tufts of pruinosity and white setulae on the central margin of tergites 1 to 6. Sternites with sparse black and yellowish setulae, sternites 1 to 6 with a narrow posterior band of pruinosity and white setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238414BC6FB567E0AFEEB.taxon	distribution	Distribution. Panama, Guyana, Ecuador, Peru, Bolivia, Argentina (Formosa).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238414BC6FB567E0AFEEB.taxon	materials_examined	Syntype ♀. Destroyed (MTMB). Type material examined. Holotype, ♀ (Rhinotriclista brevipalpis), deposited at the MNHUB (examined through photographs). Additional material examined. 2 ♀, “ Quincemil, Cuzco, Peru, VIII. 1962, L. Pena., 740 m. / Scione claripennis ” (INPA).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202238414BC6FB567E0AFEEB.taxon	discussion	Comments. Scione claripennis is similar to S. maculipennis, differing in that it has hyaline wings without spots in the crossveins and in the presence of well-marked dark brown patch with a triangular shape on the frons.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202538434BC6FF4A7818FF0F.taxon	description	(Figs 21 A – F, 22 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202538434BC6FF4A7818FF0F.taxon	materials_examined	Type locality. Peru, Lima, Matucana.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202538434BC6FF4A7818FF0F.taxon	description	Description and discussion. See Oliveira et al. (2024).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202538434BC6FF4A7818FF0F.taxon	distribution	Distribution. Peru (Lima).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202538434BC6FF4A7818FF0F.taxon	materials_examined	Holotype ♀. In good condition, deposited at the NHMUK.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332024384D4BC6FE977FCBFCAE.taxon	description	(Fig. 23 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332024384D4BC6FE977FCBFCAE.taxon	materials_examined	Type locality. Costa Rica, Suiza de Turrialba.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332024384D4BC6FE977FCBFCAE.taxon	diagnosis	Diagnosis. Body yellowish-brown. Scutum with yellowish longitudinal stripes. Wings hyaline without spots on crossveins. Legs dark brown.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332024384D4BC6FE977FCBFCAE.taxon	description	Redescription. Female. Body length: 12 – 12.7 mm (N = 2), wing length: 10.8 – 11.5 mm (N = 3, X = 11.1 mm). Head. Frons nearly parallel, divergence index 0.8 – 1.0 (N = 3, X = 0.9), frontal index 2.0 – 2.2 (N = 3, X = 2.1). Dark brown patch on the frons, almost touching the eyes and extending to the vertex. Vertex dark brown with long, dense black setulae, slightly prominent ocellar triangle, orange-brown ocelli. Frons and subcallus brown. Frons with dense, not very long black setulae and yellowish pruinosity on the sides. Parafacial and gena brown with slightly dense whitish pruinosity. Parafacial with sparse, not very dense black setulae. Gena with dense long yellowish setulae and some black setulae. Clypeus brown with slightly dense yellowish pruinosity and sparse black setulae, clypeal index 0.6 – 0.7 (N = 3, X = 0.6). Scape and pedicel brown with slightly dense orange pruinosity and long, dense black setulae. Flagellum orange-brown, last flagellomeres darkened. Palpus brown; first segment with long dense black setulae; second segment orange on the dorsal margin and with short sparse black setulae. Proboscis shiny dark brown, about 2 x the height of the frons. Thorax. Scutum and scutellum brown. Scutum with dense black setulae and some yellowish setulae. Scutellum with dense long yellow setulae. Scutum with yellowish longitudinal stripes. The median stripe is narrow and less evident. The sublateral stripes wider and more evident and connected to the lateral stripes by a yellowish stripe at the transverse suture. Notopleuron with tuft of yellowish setulae. Anepisternum and katepisternum with tuft of dense, long black setulae. Anepimeron with tuft of yellowish setulae. Postalar callus with tuft of yellowish setulae. Legs dark brown. Femora with long, dense black setulae. Tibiae with short dense black setulae. Wing hyaline with yellowish veins. Pterostigma yellowish. Abdomen. Tergites 1 and 2 yellowish with dense, short black setulae and a posterior band of pruinosity and white setulae. Tergites 3 to 7 dark brown. Tergites 3 to 5 with tufts of whitish setulae on the central posterior margin and short, dense brownish setulae. Sternites 1 and 2 yellowish with short, dense black setulae and a posterior band of pruinosity and white setulae. Other sternites dark brown with short dense black setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332024384D4BC6FE977FCBFCAE.taxon	distribution	Distribution. Costa Rica, Panama.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332024384D4BC6FE977FCBFCAE.taxon	materials_examined	Holotype ♀. Destroyed (MTMB). Neotype ♀. “ Est. Pittier, Prov. Punta, Costa Rica, 1670 m, 6 – 18 ENE 1995. M. lobo, L S 339900 577400 # 5316 / Costa Rica INBIO CRI 002 234631 / Scione costaricana (Szilády) det. Krolow, T. K., 2012 [photographed specimen] ” (MNCR). Additional material examined. 1 ♀, “ Costa Rica, Cent. Am. / Volcano Arazu / February 23 - 8 - 02 / M. A. CarrikerJr. Coll. / Cornelius B. Philip Collection Donated to California Academy of Sciences / [illegible word] / Scione claripennis Ric / Comp. w. Type Rh. brevipalpis End. Not [illegible word] CBP. / Scione costaricana Szil. CB Philip 73 / Scione costaricana Szilady det. C. B. Philip 1975 ” (CAS); 1 ♀, “ Est. Pittier, PILA-ACLA, Prov. Punta, Costa Rica, 1670 m, 5 – 18 ENE 1995. R. Villalobos, L _ N _ 330900 _ 577400 # 4437 / Costa Rica INBIO CRI 002 1833390 / Scione costaricana (Szilády) det. Krolow, T. K., 2012 ” (MNCR); 1 ♀, “ Rio Changena Bocas del Toro, 2400 ft. / Panama, 1 Sept 1961 / Scione costaricana Szilady det. Fairchild 1988 / MPEG DIP 12006464 ” (MPEG).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332024384D4BC6FE977FCBFCAE.taxon	discussion	Comments. In two of the specimens, setulae on the gena are whitish and the thoracic stripes are yellowish and less evident. S. costaricana is similar to S. maculipennis and S. claripennis. It differs from S. maculipennis in that this species has smoky brown wing with yellow spots in the middle of cells r 1 and r 2 + 3 and without lighter areas in the middle of cells m 1 and m 2, whereas S. costaricana has hyaline wing. It differs from S. claripennis in that this species has more evident thoracic stripes and the patch on the frons is dark brown and triangular, whereas in S. costaricana the thoracic stripes are less marked and the patch on the frons is irregularly shaped and extends to the apex. Typological note. The holotype of this species was lost in 1956 when Soviet Union invaded Hungary and bombed Budapest (Tyler et al. 2023). Following Chapter 16, Article 75 of the International Code of Zoological Nomenclature (ICZN 1999), we designate here a neotype for S. costaricana. The identification of the specimens and subsequent determination of the neotype was possible due to the comments made by Fairchild (1986) about this species. According to this author, specimens of S. costaricana from Panama vary greatly in setulae color. The setulae of the gena can be almost white, yellow, orange or practically black, and the tufts of setulae on each side of the scutum show the same variation of colors, although the scutum itself always appears reddish. Fairchild also argues that in the original description of S. costaricana, Szilády (1926) mentions that the species has smoky wings with dark spots on the crossveins, however in the key provided by him, S. costaricana is placed in the group of species that have hyaline wings without spots. According to Fairchild, it is possible that Szilády had a specimen of S. maculipennis with a weakly spotted wing, which would explain the ambiguity between his key and the description, since the species are quite similar except for the wings. In accordance with Article 75, the neotype was selected from material collected near the type locality, the species was redescribed and photographed, making its identification possible in later studies, and the neotype was deposited in a scientific collection (MNCR).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202A384D4BC6FC74792FF9AC.taxon	materials_examined	Type locality. Peru, Cagón.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202A384D4BC6FC74792FF9AC.taxon	description	Original description. Front broader, index 21 / 2: 1, slightly narrower above, darker in middle. Palpus and antennae reddish-brown, distal part of latter and hair of basal joints brown. Thorax brown with three narrow, yellowish longitudinal stripes, lateral ones with lateral branches along suture; pectoral hair and beard whitish. Abdomen brown, two basal segments reddish-yellow without any markings.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202A384D4BC6FC74792FF9AC.taxon	distribution	Distribution. Peru, Bolivia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202A384D4BC6FC74792FF9AC.taxon	materials_examined	Holotype ♀. Destroyed (MTMB).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202A384D4BC6FC74792FF9AC.taxon	discussion	Comments. According to the original description, this species resembles Diclisa misera, a synonym of S. aurulans. It differs in that S. aurulans has orange flagellum, thorax with two yellowish-brown longitudinal stripes, and yellowish-brown abdomen. Whereas S. crassa has reddish flagellum and the last flagellomeres brown, thorax with three yellowish longitudinal stripes, and brown abdomen. Chainey et al. (1994) recognize this species for Bolivia. However, we did not have access to this material, and their work does not provide a diagnosis, redescription or photographs. Therefore, it was not possible to confirm the identity of S. crassa and, for this reason, the species was not included in the identification key.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202A384F4BC6F9717F30FEEB.taxon	description	(Fig. 24 A – D)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202A384F4BC6F9717F30FEEB.taxon	materials_examined	Type locality. Colombia, Cauca, Cerro de Munchique, 2450 m.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202A384F4BC6F9717F30FEEB.taxon	diagnosis	Diagnosis. Body yellowish-brown densely covered with coppery setulae. Thorax with two yellow sublateral stripes that do not extend to the posterior margin of the scutum and a less evident yellowish median stripe. Wing yellowish on the anterior margin and in the middle of the cells r 1 and r 2 + 3, the rest smoky. Legs unicolored yellowish-brown.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202A384F4BC6F9717F30FEEB.taxon	distribution	Distribution. Colombia (Cauca).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202A384F4BC6F9717F30FEEB.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the FSCA (examined through photographs).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202A384F4BC6F9717F30FEEB.taxon	discussion	Comments. Scione cuprea is known only from the holotype, there are no paratypes. Among the specimens we saw, we did not find any specimen whose characteristics were consistent with S. cuprea.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332028384F4BC6FEB37913F89F.taxon	description	(Fig. 25 A – D)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332028384F4BC6FEB37913F89F.taxon	materials_examined	Type locality. “ South America ” (Venezuela).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332028384F4BC6FEB37913F89F.taxon	diagnosis	Diagnosis. Body yellowish-brown. Thorax with three yellowish longitudinal stripes. Wing slightly smoky, brown veins with faint brown spots on crossveins. Femora brown, tibiae and tarsi orange.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332028384F4BC6FEB37913F89F.taxon	description	Redescription. Female. Body length: 10 mm, wing length: 10.7 mm. Head. Frons divergent below (divergence index 0.6), frontal index 1.7. Frons without patch. Vertex dark brown with dense and long black setulae, ocellar triangle not very prominent, ocelli dark brown. Frons brown and undercoat with yellowish pruinosity, frons with dense long black setulae. Parafacial brown with yellowish pruinosity. Gena covered with yellowish pruinosity and dense long yellowish setulae. Clypeus yellowish brown with yellowish pruinosity and long dense black setulae and some yellowish setulae, clypeal index 0.6. Antenna yellow-orange. Scape and pedicel with yellowish pruinosity and long dense black setulae. Flagellum with last segment darkened. First segment of the palpus dark brown with dense and long yellowish and black setulae; second segment orange on the posterior margin and with short black setulae. Proboscis shiny dark brown, about 2 x the height of the frons. Thorax. Scutum and scutellum black. Scutum with short dense yellow and black setulae. Scutellum with long dense black and yellow setulae. Scutum with yellowish longitudinal stripes. The median stripe is not very evident and does not extend to the posterior margin of the scutum. Sublateral and lateral stripes connected by a yellowish stripe at the transverse suture. The sublateral stripes extend to the posterior margin of the scutum, where they become wider. Notopleuron with tuft of long dense black setulae. Anepisternum, katepisternum and anepimeron with long dense yellowish setulae and some black setulae. Postalar callus with a tuft of dense long yellowish setulae. Coxae and trochanters dark brown with long dense black setulae. Femora with long dense black and yellowish setulae, proximal portion dark brown and distal portion orange. Tibiae and tarsi orange. Tibiae with short dense yellow setulae. Tarsi with short dense yellow setulae in the proximal portion and short dense black setulae in the distal portion. Wing slightly smoky, brown veins with faint brown spots on crossveins. Abdomen. Tergites yellow with sparse black and yellowish setulae, denser black setulae on tergites 6 and 7. Tergite 1 with brown anterior margin. Tergite 2 with an integument median dark brown patch. Tergites 1 to 6 with slightly dense posterior bands of whitish-yellow setulae and a tuft of whitish-yellow setulae on the middle of posterior margin. Sternites with sparse, long yellowish setulae. Sternites 1 to 6 with slightly dense posterior bands of whitish setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332028384F4BC6FEB37913F89F.taxon	distribution	Distribution. Venezuela, Colombia, Peru *.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332028384F4BC6FEB37913F89F.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the NMW (examined through photographs). Additional material examined. 1 ♀, “ Peru, CA [jamarca], San Ignacio, Tabaconas, 2549 m, 5 º 11 ’ 56.7 ” S, 79 º 20 ’ 21.1 ” O, 13 – 15. x. 2017, P. Sánchez ” (UNMSM).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332028384F4BC6FEB37913F89F.taxon	discussion	Comments. In the specimen examined the frons is a little more divergent than in the type and the thoracic stripes are more evident, and besides that, the setulae of the abdomen of the holotype are denser than in the specimen seen here, which has lost many setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202E38494BC6FF4A7F94F82A.taxon	description	(Figs 26 A – F, 27 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202E38494BC6FF4A7F94F82A.taxon	materials_examined	Type locality. Ecuador, Pinullar.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202E38494BC6FF4A7F94F82A.taxon	diagnosis	Diagnosis. Body yellowish-brown. Scutum with five slightly evident whitish longitudinal stripes. Wing slightly smoky with brown veins and faint brown spots on crossveins, usually with a short appendix at the fork of vein R 4 + 5.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202E38494BC6FF4A7F94F82A.taxon	description	Redescription. Female. Body length 9.0 – 11.2 mm (X = 10.3 mm, N = 6), wing length 10.5 – 11.1 mm (X = 10.7 mm, N = 6). Head. Frons slightly divergent below, divergence index 0.6 – 0.8 (X = 0.7, N = 5), frontal index 1.6 – 1.9 (X = 1.7, N = 5). Frons with triangular-shaped brown patch. Vertex dark brown with dense and long black setulae, ocellar triangle prominent, ocelli yellowish-brown. Frons brown and subcallus with whitish pruinosity. Frons with long black setulae. Parafacial brown with whitish pruinosity and some black setulae. Gena covered with white pruinosity and dense long yellowish-white setulae. Clypeus brown with whitish pruinosity and long black setulae, clypeal index 0.7 – 0.8 (X = 0.8, N = 6). Scape and pedicel brown with whitish pruinosity and black setulae. Flagellum orange. Palpus brown; first segment with long black and yellowish setulae; second segment tapered and with short black setulae. Proboscis shiny black, about 4 x the height of the frons. Thorax. Scutum and scutellum dark brown. Scutum with dense short black and yellowish setulae, slightly evident whitish longitudinal stripes. Sublateral and lateral stripes connected by a narrower stripe at the transverse suture. Scutellum with brownish pruinosity and long yellowish-black setulae. Notopleuron with tuft of black setulae. Anepisternum, katepisternum and anepimeron with dense and long yellowish-white setulae. Postalar callus with tuft of whitish setulae. Wing slightly smoky with brown veins and faint brown spots on crossveins. Short appendix at the fork of vein R 4 + 5. Legs yellowish-brown. Femora with long, dense black and whitish setulae. Tibiae and tarsi darkened with short, dense black setulae. Abdomen. Tergite 1 with brown anterior margin and orange-yellowish posterior margin and with white and yellowish setulae. Tergite 2 orange-yellowish with black setulae and a central quadrangular dark brown patch, posterior margin with white setulae. Other tergites darkened and with black and white setulae. Sternites yellowish with black patch and short dense white setulae, some black setulae on sternites 7 and 8. Male (Fig. 27). Like the female, except for the holoptic condition, porrect palpus and longer and denser body setulae. In addition, the thoracic stripes are less evident in the male.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202E38494BC6FF4A7F94F82A.taxon	distribution	Distribution. Ecuador, Peru *.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202E38494BC6FF4A7F94F82A.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the MNHNP (examined through photographs). Additional material examined. 1 ♀, “ Equador, Pichincha, Prov., 9 km. E. Pifo, 16 – 20 - V- 1976, 3600 M. Elev. / Cornelius B. Philip Collection, Donated to California Academy of Sciences / Netted G. B. Fairchild / Scione equatoriensis Surc., Det. 1978, G. B. Fairchild / Scione equatoriensis Surc. det. Krolow, T. K., 2012 ” (CAS); 1 ♀, “ San Isidro, Carchi, Ecuador, elev. 2500 m, 23. VI. 65, L. Pena / Scione equatoriensis Surc. det. Krolow, T. K., 2012 ” (INPA); 1 ♀, “ Equador, Pichincha, Prov. Tinalandia, E. Alluriquin, 4 – 8 - v- 1976, 700 M. elev. / Netted G. B. Fairchild ” (UNMSM); 1 ♂, “ Ecuador, Pich., 33 km, E. Quito, 3200 m, 28. II. 76, G. E. Shewell / Scione equatoriensis [photographed specimen] ” (INPA); 2 ♀, “ Peru, Dept. Amazonas, Cerros Calla-Calla, 18 km, W. Leimebamba, 3100 m, 14 - X- 1964 / P. C. Hutchison and J. K. Wright Collectors / Scione sp. aff. equatoriensis Det. 1981, G. B. Fairchild ” (CAS).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33202E38494BC6FF4A7F94F82A.taxon	discussion	Comments. The patch on the frons reaches the vertex in some specimens while is barely visible in others. The appendix at the fork of vein R 4 + 5 is absent in three of the specimens. Scione equatoriensis is similar to S. incompleta and S. distincta. It differs from S. incompleta mainly in the pattern of spots on the wing of this species, yellowish-brown on the anterior margin and smoky on the posterior margin and with brown veins and brown spots on the crossveins. In S. equatoriensis the wing is slightly smoky with brown veins and faint brown spots on crossveins. This species differs from S. distincta in that this species has yellowish thoracic stripes that are slightly more evident than in S. equatoriensis and bicolored femora, dark brown proximally and orange-brown distally.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203338544BC6FF4A7860F94F.taxon	description	(Fig. 28 A – D)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203338544BC6FF4A7860F94F.taxon	materials_examined	Type locality. Colombia, Valle, Peñas Blancas, 1750 m, 10 km west of Cali.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203338544BC6FF4A7860F94F.taxon	diagnosis	Diagnosis. Body dark brown. Thorax with white longitudinal stripes. The thickest median stripe is on the posterior margin of the scutum, where it connects to the sublateral stripes. Wing smoky with yellowish spots on cells r 1, r 2 + 3, r 5, radial basal, medial basal and discal cells. Legs dark brown.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203338544BC6FF4A7860F94F.taxon	description	Redescription. Female. Body length: 11.8 – 12 mm, wing length: 12.4 – 12.7 mm. Head. Frons slightly divergent below (divergence index 0.7 – 0.8), frontal index 1.7 – 2.0. Frons with dark brown patch almost touching the eyes and extending to the vertex. Vertex dark brown with dense long black setulae, slightly prominent ocellar triangle, brown ocelli. Frons brown with whitish pruinosity and dense, long black setulae. Subcallus dark brown with slightly dense yellowish pruinosity. Parafacial brown with whitish pruinosity and sparse black setulae. Gena covered with whitish pruinosity and long dense yellowish-white setulae. Clypeus brown with not very long, dense setulae and dense whitish pruinosity on the sides, clypeal index between 0.8 and 1.0. Scape and pedicel brown with long dense black setulae. Flagellum orange-brown, last flagellomeres dark. Palpus brown; first segment with dense long black and white setulae; second segment quite tapered and with an orange dorsal margin. Proboscis shiny black, 3 x the height of the frons. Thorax. Scutum brown with dense black and yellowish setulae, not very long and with white longitudinal stripes. The median stripe and sublateral stripes connected at the posterior margin of the scutum. The sublateral and lateral stripes connected by a white stripe at the transverse suture. Scutellum brown with long dense black setulae. Notopleuron with tuft of dense long black setulae. Anepisternum with a tuft of long dense black and white setulae. Katepisternum and anepimeron with a tuft of long dense white setulae. Callus after speaking with a tuft of dense long yellowish-white setulae. Legs dark brown. Femora with long, dense black setulae. Tibiae and tarsi with short dense black setulae. Wing smoky with yellowish spots on cells r 1, r 2 + 3, r 5, radial basal, medial basal and on the discal cell. Abdomen. Tergites 1 and 2 yellowish-brown, remaining tergites dark brown. All tergites with short dense black setulae. Tergites 1 to 6 with tufts of yellowish-white setulae on the central posterior margin. Lateral margin of tergites with dense and long yellowish-white setulae. Sternites yellowish-brown with short dense black setulae. Sternites 2 to 6 with posterior bands of pruinosity and whitish setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203338544BC6FF4A7860F94F.taxon	distribution	Distribution. Western Colombia (Cauca, Valle), Ecuador (Pichincha), Peru *.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203338544BC6FF4A7860F94F.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the FSCA (examined through photographs). Additional material examined. 1 ♀, “ Peru: Dept. Amazonas, 3 km N. Pomacocha on road to Rioja 2000 m, 7 – 8. X. 1964, P. C. Hutchison & J. K. Wright / Scione equivexans Wilk. Det. 1981 G. B. Fairchild / Scione equivexans Wilkerson det. Krolow, T. K. 2012 (CAS) ”; 1 ♀, “ Tinajillas, 2.100 m, Morona-San. Prov., J. Buestán, 20. XI- 87 / MECN-EM-DIP 648 / INABIO / Donation INABio 2019 / Scione equivexans (INPA) ”.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203338544BC6FF4A7860F94F.taxon	discussion	Comments. Scione equivexans is similar to S. albifasciata and S. maculipennis. It differs from S. albifasciata in that this species has smoky brown wing with yellowish spots in the central region of cells r 1 and r 2 + 3, whereas S. equivexans has smoky wing with yellowish spots in cells r 1, r 2 + 3, r 5, basal radial, basal medial and in the discal cell. It differs from S. maculipennis in that this species has unspotted frons and legs with orange-brown femora and yellowish-brown tibiae and tarsi, whereas S. equivexans has frons with dark brown patch almost touching the eyes and extending to the vertex and dark brown legs (Fig. 29).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203138514BC6FA0078EDFA37.taxon	description	(Fig. 30 A – G)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203138514BC6FA0078EDFA37.taxon	materials_examined	Type locality. Ecuador, Tinajillas.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203138514BC6FA0078EDFA37.taxon	diagnosis	Diagnosis. Body orange-brown. Thorax brown with three yellow longitudinal stripes. Wing slightly smoky with faint brown spots on crossveins. Legs bicolored, femora black and tibiae and tarsi yellow.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203138514BC6FA0078EDFA37.taxon	etymology	Etymology. The species name refers to its similarity to S. bilineata; from the Latin falsus (false) + bilineata.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203138514BC6FA0078EDFA37.taxon	description	Description, Holotype ♀. Body length: 13.2 mm, wing length: 13 mm. Head. Frons slightly divergent below, divergence index 0.8, frontal index 2.2. Vertex dark brown with short dense black setulae, slightly prominent ocellar triangle, brown ocelli. Frons covered with yellow pruinosity and short dense black setulae. Brown frontal patch with a rounded shape from which a brown line starts and extends to the vertex. Subcallus brown with yellow pruinosity. Parafacial brown and gena with dense yellowish pruinosity. Gena with dense and long yellow setulae. Clypeus brown with yellowish pruinosity and long dense black setulae, clypeal index 0.6. Scape and pedicel brown with yellowish pruinosity and long, dense black setulae. Flagellum orange, last flagellomere slightly darkened. Palpus long and lanceolate; first segment of palpus brown with long dense yellow setulae; second segment of palpus orange-brown with short sparse black setulae. Proboscis shiny black, about 2 x the height of the frons. Thorax. Scutum brown with short dense black setulae and some yellow setulae on the lateral margins and posterior margin. Scutum with three yellow longitudinal stripes, from each of the sublateral stripes a narrower yellow stripe starts in the region of the transverse suture. Median stripe wider than sublateral stripes. Yellowish patch on posterior margin of scutum. Brown scutellum with long, dense black setulae. Notopleuron with yellow tufts of setulae. Anepisternum, katepisternum and anepimeron with tufts of long dense black setulae. Postalar callus with tuft of yellow setulae. Legs bicolored. Femora black with long dense black setulae. Tibiae and fore and mid tarsi yellow with short dense black setulae. Tibiae and hind tarsi brownish with short dense black setulae. Wing slightly smoky with faint brown spots on crossveins. Yellowish patch on the anterior region, in the region of cells r 1 and r 2 + 3. Abdomen. Tergites 1 and 2 orangish covered with short sparsely black setulae. Tergite 2 with dark integumental patch in middle and posterior band of yellow setulae. Tergite 3 yellowish brown with a posterior band of yellow pilosity and a small tuft of yellow hairs on the central posterior margin. Tergites 4 to 7 black with tuft of yellow setulae on central posterior margin. Sternites black with short black setulae. Male. Unknown. Variations of paratypes. Body length: 12 – 13.8 mm (X = 12.7 mm, N = 9), wing length: 11.4 – 13.5 mm (X = 12.5 mm, N = 9), frontal index: 2.0 – 2.5 (X = 2.3, N = 9), divergence index: 0.7 – 0.9 (X = 0.8, N = 9), clipeal index: 0.5 – 0.7 (X = 0.6, N = 9). The patch on the frons is absent in one specimen and slightly more marked in others. Hind tibiae and tarsi are yellow in some specimens. In some specimens, the abdominal tergite 3 is entirely orange with a central brown patch.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203138514BC6FA0078EDFA37.taxon	distribution	Distribution. Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203138514BC6FA0078EDFA37.taxon	materials_examined	Type material. Holotype, ♀, “ Tinajillas, 2.100 m, Morona-San. [tiago] Prov., J. Buestán, oct. 02 [October 2002?] / 13 / CJB ” (MECN). Paratypes: 4 ♀ (in good condition, one of the specimens with missing pedicels and flagella and another with the right mid leg missing), “ Tinajillas, 2.100 m, Morona-San. Prov. J. Buestán, oct. 02 [October 2002?] / 13 / CJB ” (MECN); 1 ♀ (in good condition, but hind tarsi missing) “ idem, except oct [tober] 2012 ” (MECN); 1 ♀ (in good condition, but pedicel and left flagellum missing), “ Tinajillas, 2.100 m, Morona-San. Prov. J. Buestán, 18 - ix- 87 / CJB- 2329 ” (MECN); 1 ♀ (in good condition, but left pedicel and flagellum missing and proboscis broken), “ Tinajillas, 2.100 m, Morona-San. Prov. J. Buestán, 10 - x- 87 / CJB- 2336 ” (MECN); 1 ♀, (in good condition, but left pedicel and flagellum missing, left posterior tarsus missing), “ Tinajillas, 2.100 m, Morona-San. Prov. J. Buestán, 10 - x- 87 / CJB- 2338 ” (INPA); 1 ♀ (in good condition, but pedicels and flagella missing, proboscis missing), “ Tinajillas, 2.100 m, Morona-San. Prov. J. Buestán, 10 - x- 87 / CJB- 2339 ” (MECN).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203138514BC6FA0078EDFA37.taxon	discussion	Comments. At the MZUSP there is an unlabeled specimen. This specimen was not included in the type series but is well preserved, except for the missing right hind tarsus and left hind tibia. In the holotype and in some paratypes there is a large amount of pollen, mainly on the head and legs. Scione falsibilineata sp. nov. is similar to S. bilineata. The former has thorax with three yellow longitudinal stripes, whereas S. bilineata has thorax with two yellow longitudinal stripes. Furthermore, S. bilineata has a darker frons and wing spots and dense orange setulae on the scutum which are absent in Scione falsibilineata sp. nov. (Fig. 31).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332036385C4BC6F9CE7AD6FA6B.taxon	description	(Figs 32 A – G, 33 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332036385C4BC6F9CE7AD6FA6B.taxon	materials_examined	Type locality. Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332036385C4BC6F9CE7AD6FA6B.taxon	diagnosis	Diagnosis. Body yellowish-brown. Clypeus markedly developed. Thorax with conspicuous white longitudinal stripes. Wing smoky with brown spots in the middle of cells r 1, r 2 + 3 and r 5 and spots brown on crossveins.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332036385C4BC6F9CE7AD6FA6B.taxon	description	Redescription. Female. Body length: 10.6 – 11.7 mm (X = 11.2 mm, N = 3), wing length: 11.5 – 11.8 mm (X = 11.7 mm, N = 4). Head. Frons slightly divergent below, divergence index 0.7 – 0.8 (X = 0.8, N = 4), frontal index 1.7 – 1.9 (X = 1.8, N = 4). Frons with a brown quadrangular patch, almost touching the eyes. Vertex dark brown with dense long black setulae, prominent ocellar triangle, orange-brown ocelli. Frons brown covered with yellowish and whitish pruinosity and dense long black setulae. Subcallus brown with slightly dense yellowish pruinosity. Parafacial brown with slightly dense orange pruinosity and sparse black setulae. Gena brown with dense whitish pruinosity and dense long whitish setulae. Clypeus brown with sparse black setulae and slightly dense yellowish pruinosity on the sides, markedly developed, clypeal index 1.0 – 1.2 (X = 1.1, N = 4). Scape and pedicel yellowish-brown with yellowish pruinosity and long, dense black setulae. Flagellum orange. First segment of the palpus is brown with long dense orange setulae in the proximal portion and short sparsely dense black setulae in the distal portion; second segment orange with short sparse black setulae. Proboscis dark brown, about 4 x the height of the frons. Thorax. Scutum and scutellum orange-brown. Scutellum with sparse black and yellowish setulae and white longitudinal stripes. The sublateral stripes quite evident and extend to the posterior margin of the scutum where they connect. The median stripe is less evident than the sublateral stripes. The sublateral and lateral stripes connected by a narrower white stripe at the transverse suture. Notopleuron with tuft of black and yellowish setulae. Anepisternum with tuft of black, yellowish and whitish setulae. Katepisternum and anepimeron with tuft of whitish setulae. Postalar callus with tuft of yellowish-white setulae. Legs yellowish-brown, femora with long, dense black and yellowish setulae, tibiae with short dense black and yellowish setulae. Wing smoky with yellowish spots in cells r 1, r 2 + 3 and r 5. Brown veins with faint brown spots on crossveins. Abdomen. Tergites yellowish-brown with sparse black and yellowish setulae. Tergites 1 to 6 with tuft of yellowish-white setulae on the central posterior margin. Sternites slightly darkened and sparse black and whitish setulae with small middorsal patches of shiny yellow brown hairs on tergites 1 to 6.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332036385C4BC6F9CE7AD6FA6B.taxon	distribution	Distribution. Colombia, Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332036385C4BC6F9CE7AD6FA6B.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the MNHUB (examined through photographs). Additional material examined. 1 ♀, “ Tambo to Starosa, Cascho Crespejo, Ecuador, VIII- 1 - 35 / Pres. by Mrs. Y. Mexia Collector [photographed specimen] ” (CAS); 1 ♀, “ Los Olivos, Carchi, Ecuador, VII- 11 - 1935, 3200 M. / Pres. by Mrs. Y. Mexia Collector ” (CAS); 1 ♀, “ Los Olivos, Carchi, Ecuador, VII- 11 - 1935, 3200 M. / Pres. by Mrs. Y. Mexia Collector / Cornelius B. Philip Collection, Donated to California Academy of Sciences / Snout to long / Compared with type ♀ F. albifasciata Macq. C. B. Philip 58, Agrees but type c [abbreviation of Latin cum, with] straw yel [low] bd [body] + [and] chest / Scione albifasciata ♀ (Macq.) 56, Det. C. B. Philip / Scione albifasciata (Macquart) det. Krolow, T. K., 2012 ” (CAS); 1 ♀, “ Los Olivos, Carchi, Ecuador, VII- 11 - 1935, 3200 M. [without abdomen] / Pres. by Mrs. Y. Mexia Collector / CAS / Scione albifasciata ♀ (Macq.) 56, Det. C. B. Philip ” (CAS).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332036385C4BC6F9CE7AD6FA6B.taxon	discussion	Comments. The thorax and abdomen of the holotype are more densely covered with yellowish setulae, which appear to be missing in the specimens examined; these are not in good condition of preservation. On the abdomen of the holotype the tuft of yellowish-white setulae is present on the central posterior margin of tergites 2 to 6 and is much denser than in the specimens seen by us. Scione flavescens is similar to S. albifasciata and S. maculipennis. However, this species has a markedly developed clypeus, unlike the other two species. Furthermore, S. albifasciata has more pronounced patch on the frons and this touches the vertex, unlike S. flavescens which has less evident patch on the frons. Wing of S. maculipennis has smoky brown coloration with yellow spots in the middle of cells r 1 and r 2 + 3 and no lighter areas in the middle of cells m 1 and m 2. Whereas in S. flavescens the yellowish spots are restricted to cells r 1, r 2 + 3 and r 5. Wilkerson (1979) provides a redescription for S. flavescens that agrees very well with the specimens seen here, however the specimens studied by Wilkerson vary mainly in body length, measuring between 9 and 11 mm.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203B385E4BC6FA327A0AF887.taxon	description	(Figs 34 A – F, 35 A – E, 36 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203B385E4BC6FA327A0AF887.taxon	materials_examined	Type locality. Bolivia, Cillunticara.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203B385E4BC6FA327A0AF887.taxon	diagnosis	Diagnosis. Body orange-brown. Flagellum orange. Scutum with yellowish stripes that not very evident. Wings hyaline without spots on crossveins. Abdomen brown, tergite 2 orange with a central black patch.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203B385E4BC6FA327A0AF887.taxon	description	Redescription. Female. Body length: 8.3 – 12.5 mm (X = 10.3 mm, N = 15), wing length 9.2 – 12.1 mm (X = 10.6 mm, N = 16). Head. Frons divergent below, divergence index 0.6 – 0.9 (X = 0.7, N = 11), frontal index 1.5 – 2.1 (X = 1.9, N = 11). Frons unspotted. Vertex dark brown, ocellar triangle prominent, ocelli yellowish-brown. Frons and subcallus brown covered with yellowish pruinosity, frons with a few long black setulae. Parafacial brown with sparse whitish pruinosity and some sparse black setulae. Gena covered with whitish pruinosity and dense, long yellowish setulae. Clypeus brown with sparse black setulae and whitish pruinosity on the sides, clypeal index 0.6 – 0.9 (X = 0.8, N = 16). Scape and pedicel orange-brown with whitish pruinosity and long, dense black setulae. Flagellum orange, last flagellomere slightly darkened. Palpus brown, first segment with dense, long yellowish setulae in the proximal portion and sparse, less long, black setulae in the distal portion; second segment of the palpus with a yellowish dorsal margin and short sparse black setulae. The two segments of the palpus of equal or subequal size. Proboscis shiny, dark brown, about 7 x the height of the frons. Thorax. Scutum and scutellum dark brown with sparse black and yellowish setulae. Scutellum with yellowish longitudinal stripes. The barely visible median stripe does not extend to the posterior margin of the scutum. The more evident sublateral stripes extend to the posterior margin and connected to the lateral stripes by a narrow and barely visible yellowish stripe at the transverse suture. Posterior margin of scutum with dense yellowish pruinosity, scutellum with less dense yellowish pruinosity on anterior and posterior margin. Notopleuron with tuft of dense and long black and yellowish setulae. Anepisternum, katepisternum and anepimeron covered with whitish pruinosity and a tuft of yellowish whitish setulae. Postalar callus with a tuft of yellowish-whitish setulae. Legs yellowish-brown. Femora with long dense yellowish-black setulae. Tibiae and tarsi with short dense black setulae. Wing hyaline with yellowish-brown veins. Abdomen. Tergite 1 with dark brown anterior margin and orange posterior margin. Tergite 2 orange with central black patch. Tergite 3 orange brown. Other tergites darkened. All tergites with black setulae and some yellowish setulae. Sternites yellowish-brown with whitish setulae. Male (Fig. 36). Like the female, except for the holoptic condition, porrect palpus, longer and denser body setulae and less evident thoracic dorsal stripes.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203B385E4BC6FA327A0AF887.taxon	distribution	Distribution. Ecuador, Peru, Bolivia, Argentina (Salta, Jujuy, Tucumán, Catamarca).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203B385E4BC6FA327A0AF887.taxon	materials_examined	Type material examined. Syntype, ♀, deposited at the NHMUK (examined through photographs). Additional material examined. 1 ♂, “ Vale, Azuay, Ecuad. VI 1947, Coll. Z. Muller / Cornelius B. Philip Collection, Donated to California Academy of Sciences / Scione fulva? Ric. det C. B. Philip 49 / Comp. to type Scione flavohirta Ric. C. B. Philip Ag. 53, Agrees exe. sex [illegible word], diffs. type has ots lines on thox [thorax?] see notes ” (CAS); 1 ♂, “ Vale, Azuay, Ecuad. VI 1947, Coll. Z. Muller / / Scione fulva? Ric. det C. B. Philip / Cornelius B. Philip Collection, Donated to California Academy of Sciences ” (CAS); 1 ♀, “ Ecuador, Bolivar, Las Guardias 2179, 16 Jul 09, J. Buestan / CJB 10242 / 15 / CJB ” (MECN); 1 ♀, “ idem, except CJB 10250 ” (MECN); 1 ♀, “ idem, except CJB 10252 ” (MECN); 1 ♀, “ idem, except CJB 10254 [photographed specimen] ” (MECN); 1 ♀, “ idem, except 17 Jul 09 / CJB 10243 ” (MECN); 1 ♀, “ idem, except CJB 10289 ” (MECN); 1 ♂, “ Ecuad. Chimborazo km. 5 N. Pallatanga, 11 Ago 09, J. Buestan / CJB 10295 / CJB ” (MECN); 1 ♂, “ idem, except CJB 10296 ” (MECN); 1 ♂, “ idem, except 13 Ago 09 / CJB 10297 [photographed specimen] ” (MECN); 1 ♂, “ Ducur, Rio Cañar, 9. vi. 85 / CJB- 2555 / 15 ” (MECN); 1 ♀, “ Peru, Rio Pacharcacaca, 5 - II- 95, coll: F. Monros / from Jucuman / Cornelius B. Philip Collection Donated to California Academy of Sciences / Scione flavohirta Ric, 65, Det. C. B. Philip ” (CAS); 1 ♀, “ Peru, Ayacucho, 8 - II- [1] 952, coll: Monros / Cornelius B. Philip Collection Donated to California Academy of Sciences ” (CAS); 1 ♀, “ 40 Mi. E. of Abancay, Peru, III- 5 - 51 / Ross and Michelbacher Collectors / Scione flavohirta ♀ Ric, 56, Det. C. B. Philip ” (CAS); 1 ♀, “ 35 Mi. E. of Abancay, Peru, III- 5 - 51 / Cactus mesquite zone / Scione flavohirta ♀ Ric, 56, Det. C. B. Philip ” (CAS); 1 ♀, “ 5 Mi. N. of, Andahuaylas, Peru, III- 7 - 51 / Ross and Michelbacher Collectors / Cornelius B. Philip Collection Donated to California Academy of Sciences ” (CAS); 1 ♀, “ 40 Mi. W. of Cusco, Peru, III- 4 - 51 (Subtropical cyn.) / Ross and Michelbacher Collectors / Cornelius B. Philip Collection Donated to California Academy of Sciences ” (CAS); 2 ♀, “ 40 Mi. E. of Abancay, Peru, III- 5 - 51 / Ross and Michelbacher Collectors / Cornelius B. Philip Collection Donated to California Academy of Sciences ” (CAS); 1 ♀, “ 5 Mi. N. of Andahuaylas, Peru, III- 7 - 51 / Ross and Michelbacher Collectors ” (CAS); 1 ♀, “ Abancay, Peru, III- 6 - 51 / Ross and Michelbacher Collectors / Cornelius B. Philip Collection Donated to California Academy of Sciences ” (CAS); 1 ♂, “ 5 Mi. N. of Andahuaylas, Peru, III- 7 - 51 / Ross and Michelbacher Collectors ” (CAS); 1 ♂, “ 5 Mi. N. of Andahuaylas, Peru, III- 7 - 51 / Ross and Michelbacher Collectors / Scione flavohirta ♂ Ric. 56 Det. C. B. Philip ” (CAS); 1 ♂, “ 5 Mi. N. of Andahuaylas, Peru, III- 7 - 51 / Ross and Michelbacher Collectors / Cornelius B. Philip Collection, Donated to California Academy of Sciences ” (CAS); 1 ♀, “ Tucuman, S. Pedro d., Colalao, Col. Duver, J. 54 / MZSP ” (MZUSP).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203B385E4BC6FA327A0AF887.taxon	discussion	Comments. The specimens from the CAS are not in good condition, some have been exposed to an oily substance, and the thoracic stripes are not visible. The length of the proboscis varies among the specimens examined, measuring between four and six times the height of the frons. The thorax and scutellum have dense black setulae in two specimens analyzed, in the others much of the setulae are missing. The males are in better condition and it is possible to observe, as in the female type, that the abdomen has tergites 1 to 5 with posterior bands of dense yellowish setulae that are not very long, tergites 2 to 4 have a tuft of dense and long yellowish setulae on the central posterior margin and dense black setulae that are not very long on tergites 3 to 7. In males, the setulae of the thorax are much denser and yellowish. The original description was based on a male, with a brief comment regarding the female. The catalogue by Coscarón & Papavero (2009) reports that the type that was deposited at the MTMB, and which was lost, was a female. However, the type that was deposited in that museum was the male described by Ricardo (1902).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203E38594BC6FF4A796CF8A0.taxon	description	(Figs 37 A – E, 38 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203E38594BC6FF4A796CF8A0.taxon	materials_examined	Type locality. Bolivia, Songo.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203E38594BC6FF4A796CF8A0.taxon	diagnosis	Diagnosis. Body orange-brown. Flagellum orange yellow with the last four flagellomeres black. Tergites 2 and 3 yellow with a central dark brown patch.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203E38594BC6FF4A796CF8A0.taxon	description	Redescription. Male. Body length: 11.2 mm (N = 1), wing length: 10.2 – 12.2 mm (X = 11.5 mm, N = 4). Head. Vertex dark brown with not very long black setulae, slightly prominent ocellar triangle, orange brown ocelli. Subcallus brown with whitish pruinosity. Gena and parafacial covered with whitish pruinosity. Gena with long not very dense black setulae. Parafacial with dense long black and yellowish setulae. Clypeus brown with whitish pruinosity and dense long black setulae, clypeal index 0.7 – 0.8 (X = 0.7, N = 4). Scape and pedicel brown with whitish pruinosity and long dense black setulae. Orange flagellum with the last four black flagellomeres. Palpus dark brown; first segment with long dense yellowish setulae in the proximal portion and long dense black setulae in the distal portion; second segment with dense black setulae slightly shorter than in the first segment and with an orange dorsal margin. Proboscis shiny black, about 6 x the height of the frons. Thorax. Scutum and scutellum dark brown. Scutum with dense not very long black setulae and some yellowish setulae and slightly evident whitish longitudinal stripes that extend to the posterior margin of the scutum. The sublateral and lateral stripes connected by a narrow, barely visible whitish stripe at the transverse suture. Scutellum with long dense black setulae and some yellowish setulae. Notopleuron with a tuft of long, dense black and yellowish setulae. Anepisternum with tuft of black and yellow setulae. Katepisternum with tufts and black setulae. Anepimeron with tuft of yellow setulae. Postalar callus with tuft of black setulae and some yellowish setulae. Femora brown in the proximal portion and yellowish in the distal portion with long dense black setulae. Tibiae and tarsi yellow with short, dense black setulae. Wing hyaline with yellowish-brown veins. Abdomen. Tergite 1 dark brown with yellow posterior margin and with long dense yellowish-black setulae. Tergite 2 yellow with long dense yellow setulae and a central dark brown patch. Tergite 3 yellow with long, dense, black setulae, with a central dark brown patch and a small tuft of yellow setulae on the central posterior margin. Other tergites yellowish-brown with long dense black setulae and a posterior band of yellowish pruinosity and setulae. Sternites 1 to 4 yellow with short dense yellow setulae. Sternites 5 to 7 yellowish-brown. Sternites 5 and 6 with short dense yellow setulae. Sternite 7 with long, dense black setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203E38594BC6FF4A796CF8A0.taxon	distribution	Distribution. Peru, Bolivia, Argentina.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203E38594BC6FF4A796CF8A0.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the NHMUK (examined through photographs). Additional material examined. 1 ♂, “ Termas de Reyes Jujuy [Argentina], 22. I. 48, Monros y Willink / Cornelius B. Philip Collection, Donated to California Academy of Sciences [photographed specimen] ” (CAS); 1 ♂, “ R. A. Jujuy [Argentina], 13 - II- 1951, coll: Monros y Willink / Scione fulva / 16 / MZSP ” (MZUSP); 1 ♂, “ R. A. Jujuy [Argentina], 13 - II- 1951, coll: Monros y Willink / S. fulva / Scione sp.? ♂ det. Krolow, T. K., 2012 / MZSP ” (MZUSP); 1 ♂, “ Rio Cochuna, Tucuman [Argentina], 6. V. 1948, Cola. Willink. / Scione flavohirta? / Scione flavohirta? Ricardo Det. T. K. Krolow, 2011 / MZSP ” (MZUSP).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33203E38594BC6FF4A796CF8A0.taxon	discussion	Comments. Three of the specimens examined had their genitalia removed and therefore the body length was not measured, as part of the abdomen is missing. However, all appear to be approximately 11 mm long. In the female holotype, the thoracic stripes are more evident than in the male specimens examined. Scione fulva and S. flavohirta are quite similar, they differ in the structure of the palpus, in S. fulva the palpus is not so sharp and the second segment is shorter than the first. The palpus of S. flavohirta female is tapered and the two segments are nearly equal in length. Furthermore, in S. flavohirta the flagellum is orange and only the last flagellomere is slightly darkened, whereas in S. fulva the flagellum is yellowish-orange with the last four flagellomeres black, this character can be observed in both females and males.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204338244BC6FF4A7E73FB0A.taxon	description	(Figs 39 A – F, 40 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204338244BC6FF4A7E73FB0A.taxon	materials_examined	Type locality. Panama, Darien (as Colombia).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204338244BC6FF4A7E73FB0A.taxon	diagnosis	Diagnosis and redescription. See Scione rufescens.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204338244BC6FF4A7E73FB0A.taxon	distribution	Distribution. Costa Rica to western Andes of Colombia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204338244BC6FF4A7E73FB0A.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the NHMUK (examined through photographs); Holotype of S. aureopygia, ♀, deposited at the MCZ (examined through photographs). Additional material examined. 1 ♀, “ Panama, Darien Prov. Tacarcuna, Yellow Fever Camp / Scione fulvosericea Krob. det Fairchild 1988 / MPEG DIP 12006468 / Scione fulvosericea ” (MPEG); 1 ♀, “ Pacific Canal Zone, 6 - 6 - 1972 / Scione rufescens Ric. det. G. B. Fairchild 1981 ” (INPA); 1 ♀, “ Pacific Canal Zone, 6 - 6 - 1972 / Scione rufescens (Ricardo) det. Krolow, T. K., 2012 ” (INPA); 1 ♀, “ Almirante Camp., 10 Oct. [19] 51, 6 - 7 P. M. / Cornelius B. Philip Collection Donated to California Academy of Sciences / Scione aureopygia Fchld. det. Fairchild 1951 / Scione rufescens (Ricardo) det. C. B. Philip / Scione rufescens (Ricardo) det. Krolow, T. K., 2012 ” (CAS); “ Panama: Darien Prov. Tacarcuna, Yellow Fever Camp / Shannon trap, 6 – 8 P. M., 21 - VI- 1963 / Scione rufescens Ric. det. G. B. Fairchild 1981 / Scione rufescens (Ricardo) det. Krolow, T. K., 2012 ” (INPA). 1 ♀, Pezo Azul, Costa Rica, C. F. Underwood, May-June, 1902, per Janson. / C. J. Wainwright Collection, B. M. 1948 - 488. / Cornelius B. Philip Collection, Donated to California Academy of Sciences / Compared with type ♀ Scione rufescens Ric. Agrees (over) C. B. P. 1953 / Compared with type ♀ Fidena fulvosericea Krb CB Philip 1958 agrees but darker / Scione rufescens (Ricardo) det. Krolow, T. K., 2012 ” (CAS).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204338244BC6FF4A7E73FB0A.taxon	discussion	Comments. Scione fulvosericea is very similar to S. rufescens. It differs in smaller size, the specimens of S. fulvosericea examined here measure between 14 – 16 mm, whereas those of S. rufescens measure between 17 – 17.4 mm and by the absence of the pronounced prominence at the apex of the clypeus, present in S. rufescens.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204338244BC6FA937917F864.taxon	description	(Fig. 41 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204338244BC6FA937917F864.taxon	materials_examined	Type locality. Bolivia, Cordilleras, 3000 m.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204338244BC6FA937917F864.taxon	diagnosis	Diagnosis. Body yellowish-brown. Scape and pedicel brown, flagellum orange with last flagellomeres brown. Palpus quite tapered, first segment brown; second segment orange brown. Thorax with whitish longitudinal stripes. Wing smoky brown. Abdomen yellowish-brown with dense yellow setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204338244BC6FA937917F864.taxon	distribution	Distribution. Peru, Bolivia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204338244BC6FA937917F864.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the MLUH (examined through photographs).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204338244BC6FA937917F864.taxon	discussion	Comments. Scione fumipennis is similar to S. fulva. It differs in that S. fulva has orange yellow flagellum with the last four flagellomeres black, whereas S. fumipennis has orange flagellum with the last four flagellomeres brown. In S. fulva the second segment of the palpus is smaller than the first, in S. fumipennis the two segments of the palpus are practically the same size.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204038204BC6F9037A1CFE27.taxon	description	(Fig. 42 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204038204BC6F9037A1CFE27.taxon	materials_examined	Type locality. Bolivia, Songo.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204038204BC6F9037A1CFE27.taxon	diagnosis	Diagnosis. Body yellowish-brown. Antennae yellow with greenish tones on the scape, pedicel and last flagellomeres. Legs yellow with greenish tones on the femora, brownish tarsi. Wing smoky brown with distinct yellow macule covering part of cells r 1, r 2 + 3, r 5 and m 3 and the entire discal cell.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204038204BC6F9037A1CFE27.taxon	distribution	Distribution. Peru, Bolivia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204038204BC6F9037A1CFE27.taxon	materials_examined	Holotype ♀. Destroyed (MTMB). Material examined. 1 ♀, “ Peru, Dep. Cuzco, Callanga i. Thale, des Pinipini 1500 m, Garlepp S. V., 1900 / Rhinotriclista fusca, (Ric.) ♀, Dr. Enderlein det. 1922 [examined through photographs] ” (MNHNP).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204038204BC6F9037A1CFE27.taxon	discussion	Comments. Although the type has been destroyed, this species is easily identifiable due to the wing spot pattern mentioned above.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204638224BC6FF4A7818F8A3.taxon	description	(Figs 43 A – E, 44 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204638224BC6FF4A7818F8A3.taxon	materials_examined	Type locality. Ecuador, Tungurahua, above Baños.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204638224BC6FF4A7818F8A3.taxon	description	Description and discussion. See Oliveira et al. (2024).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204638224BC6FF4A7818F8A3.taxon	distribution	Distribution. Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204638224BC6FF4A7818F8A3.taxon	materials_examined	Holotype ♀. In good condition, deposited at the NHMUK.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204438234BC6FF4A7998FCDC.taxon	description	(Fig. 45 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204438234BC6FF4A7998FCDC.taxon	materials_examined	Type locality. Peru, Rio Pionía, Puerto Bermúdez.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204438234BC6FF4A7998FCDC.taxon	diagnosis	Diagnosis. Predominantly black with dense and long yellow setulae on the parafacial, gena, anepimeron, katepisternum, anepisternum, notopleuron and postalar callus. Frons and subcallus whitish, frons with long yellow setulae. Wing black with translucent fenestrae in cells br, bm, d, CuA 1, m 2, m 3 and on the wing apex (in part of cells r 2 + 3 and r 4).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204438234BC6FF4A7998FCDC.taxon	distribution	Distribution. Peru.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204438234BC6FF4A7998FCDC.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the MCZ (examined through photographs).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204438234BC6FF4A7998FCDC.taxon	discussion	Comments. This species is known only from the holotype and its classification as Scione is uncertain. Although it has closed and petiolate r 5 and m 3 cells, characteristic of the genus, the shape of the cells, especially the discal cell, m 3 and cua 1, is quite different from that in other species of Scione. The unusually narrow frons, very robust proboscis, very long and narrow palpus and clypeus without protuberance also differ from other species of the genus. For all these reasons, S. grandis is treated here as incertae sedis. For a correct placement of the species, a complete examination of the specimen is necessary.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204438234BC6FCA67941F98C.taxon	materials_examined	Type locality. Peru, Huancabamba, 3000 m.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204438234BC6FCA67941F98C.taxon	description	Original description. Länge 13.6 – 14 mm, Fühler? mm, Rüssel 5 – 5.9 mm, Flügellänge 12 – 14.5 mm, Flügelbreite 4 – 4.5 mm. Schlecht erhalten, aber durch die Grösse und Ähnlichkeit mit Osca - Arten leicht erkennbar. Stirn breit, ca. 2 mal so hoch als unten breit, oben verschmälert, dunkelrotbraun, dicht kurz schwarz behaart. Ocellen blutrot. Der gesamte Körper einschliesslich der p ist eigentlich rotbraun. 1. und 2. Fühlerglied dunkel rotbraun, schwarz behaart, der Rest fehlt. 2. Tasterglied messerklingenförmig. Bart gelblich. Hinterkopf grauweiss, weisslich behaart. Augen sehr dicht gelblich behaart. Thorax, wenn denudiert mit 2 breiten schwarzen Seitenstriemen und unscharfer, hinten verkürzter Mittelstrieme. Behaarungsreste vorn schwarz, seitlich und vor dem Schildchen gelblich. Schildchen schwarz. Behaarung der Brustseiten gelb und schwarz, p goldrot und schwarz behaart, Tarsen fast schwarz. Flügel hyalin, Vorderrand und Queradern etwas braun, fastgesäumterscheinend. 1. Basalzelle grösstenteils braun. 1. – 2. Tergit heller rotbraun. Mitte des 1., 5. und 6. Tergits mit einem gelblichen Haardreieck. Die äussersten Seiten der Tergite am Hinterrand gelblich behaart; sonst ist die Behaarung schwarz. Bauch braungelb, graulich bestäubt, sehr zart schwarz und gelb behaart.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204438234BC6FCA67941F98C.taxon	distribution	Distribution. Peru.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204438234BC6FCA67941F98C.taxon	materials_examined	Holotype ♀. Destroyed (Kröber Collection).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204438234BC6FCA67941F98C.taxon	discussion	Comments. According to the original description, this species is easily recognizable due to its size and similarity to the genus Osca Walker, 1850.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204A382D4BC6FF4A79E5F930.taxon	description	(Figs 46 A – F, 47 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204A382D4BC6FF4A79E5F930.taxon	materials_examined	Type locality. Venezuela, Monte Roraima.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204A382D4BC6FF4A79E5F930.taxon	diagnosis	Diagnosis. Body dark brown. Thorax with yellowish longitudinal stripes. Wing slightly smoky with orange brown veins, without spots and with yellowish pterostigma.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204A382D4BC6FF4A79E5F930.taxon	description	Redescription. Female. Body length: 9.5 – 10.2 mm (X = 9.9 mm, N = 5), wing length 9.8 – 10.8 mm (X = 10.3 mm, N = 5). Head. Frons nearly parallel, divergence index 0.9 (in all examined specimens), frontal index 1.9 – 2.0 (X = 1.9, N = 5). Frons unspotted. Vertex brown with long dense black setulae, ocellar triangle prominent, ocelli yellowish-brown. Frons and subcallus brown with slightly dense orange pruinosity, frons with long black setulae that not very dense. Parafacial brown with slightly dense orange pruinosity and dense, long black setulae near the gena. Gena brown with dense yellowish pruinosity and dense long whitish setulae. Clypeus dark brown with sparse whitish pruinosity and sparse long black setulae, slightly protuberant, clypeal index 0.8 – 09 (X = 0.8, N = 5). Scape and pedicel brown with dense, long black setulae. First flagellomere brown in the proximal portion and orange in the distal portion, remaining flagellomeres orange, the last ones slightly darkened. First segment of the palpus dark brown with long dense black setulae; second segment orange with sparse short black setulae. Proboscis dark brown shiny, about 3 x the height of the frons. Thorax. Scutum orange brown with dense short black setulae and yellowish longitudinal stripes. Median stripe narrow and not very conspicuous. Sublateral stripes slightly wider and more evident. The sublateral stripes become wider in the region of the transverse suture, from where a narrow and not very evident stripe starts towards notopleuron. Scutellum orange brown with long dense black setulae. Notopleuron yellowish with a tuft of long dense black setulae and some yellowish setulae. Anepisternum and katepisternum with tuft of long dense black setulae. Anepimeron with tuft of long dense whitish setulae. Postalar callus with tuft of yellowish setulae. Legs brown. Coxae, trochanters and femora with long dense black setulae. Tibiae and tarsi with short dense black setulae. Wing slightly smoky with orange-brown veins, without spots and with yellowish pterostigma. Abdomen. Tergites yellowish-brown with short dense black setulae; tergites 3 to 7 darkened. Tergites 1, 4 and 5 with a small tuft of yellowish setulae on the central posterior margin. Sternites yellowish-brown with short dense black setulae, sternites 2 to 4 with a whitish posterior margin and dense yellowish setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204A382D4BC6FF4A79E5F930.taxon	distribution	Distribution. Venezuela, Brazil (Roraima) *.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204A382D4BC6FF4A79E5F930.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the NHMUK (examined through photographs). Additional material examined. 5 ♀, “ Brasil, Roraima, P. N. Monte Roraima, 05 º 07 ’ 24 ” N, 60 º 35 ’ 50 ” W, 1.000 m em floresta [in forest], xii. 2019 / malaise grande [Malaise Gressitt & Gressitt], F. Xavier & S. Lima col. / ♀ Scione immaculata (Kröber) Det. Henriques, 2022 ” (CEUFT); 12 ♀, “ idem (INPA) ”; 6 ♀, “ Brasil, Roraima, Monte Caboraí, Marco Divisor BV 11 A, 5 º 16 ’ 19.60 ” N, 60 º 12 ’ 43.29 ” W, Mata Super Umida 1450 m, Isca-Humana, 19 a 31 - XII- 2000 / Monte Caboraí, RR [Roraima], J. B. F. da Silva; B. C. Souza; S. P. do Nascimento & J. A. Schiwinden / Scione immaculata (Kröber) det: L. P. Oliveira, 2024 ” (MPEG).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204A382D4BC6FF4A79E5F930.taxon	discussion	Comments. The tuft of yellowish setulae on the posterior margin of tergites 1, 4 and 5 is not very dense and appears to be present on the other tergites as well. In the holotype, the ocelli are black, whereas in the specimens examined here the ocelli are yellowish- brown.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204F382A4BC6FF4A7E4AFA8E.taxon	description	(Figs 48 A – D, 49 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204F382A4BC6FF4A7E4AFA8E.taxon	materials_examined	Type locality. Colombia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204F382A4BC6FF4A7E4AFA8E.taxon	diagnosis	Diagnosis. Body yellowish-brown. Scutum with slightly evident yellowish longitudinal stripes. Wing yellowish-brown on the anterior region and smoky on the posterior region, brown veins and brown spots on crossveins.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204F382A4BC6FF4A7E4AFA8E.taxon	description	Redescription. Female. Body length: 9.2 – 9.6 mm, wing length: 9.4 – 9.8 mm. Head. Frons divergent below, divergence index 0.7 (X = 2), frontal index 1.8 – 1.9. Frons with irregular dark brown patch, almost touching the eyes and extending to the vertex. Vertex dark brown with long black setulae, prominent ocellar triangle, dark brown ocelli. Frons and subcallus brown with yellowish pruinosity, frons with long black setulae. Parafacial brown with yellowish pruinosity and some black setulae. Gena with yellowish white pruinosity and dense, long whitish setulae. Clypeus brown with yellowish pruinosity and dense long black setulae, slightly protuberant, clypeal index 0.7 – 0.8. Scape and pedicel brown with yellowish pruinosity and long, dense black setulae. Flagellum orange, last flagellomeres slightly darkened. Palpus brown, first segment with dense and long yellowish and black setulae; second segment with short black setulae. Proboscis dark brown, about 3 x the height of the frons. Thorax. Scutum and scutellum dark brown. Scutellum with long black setulae. Scutum with shorter black setulae and slightly evident yellowish longitudinal stripes, the sublateral and lateral stripes connected by an even less evident yellowish stripe on transverse suture. Notopleuron with tuft of long black setulae. Anepisternum with tuft of long dense black and yellowish setulae, katepisternum and anepimeron with tuft of long dense yellowish and white setulae. Postalar callus with tuft of whitish setulae. Legs yellowish. Femora with long dense black and yellowish setulae. Tibiae and tarsi with short dense black setulae. Wing yellowish-brown on the anterior region and smoky on the posterior region brown veins, brown spots on crossveins and yellowish patch on cells r 1, r 2 + 3 and in part of the radial basal cell. Abdomen. Tergites yellowish with dense short black setulae and some yellowish setulae. Tergites 3 and 4 with a small tuft of whitish setulae on the central posterior margin. Sternites with sparse black and whitish setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204F382A4BC6FF4A7E4AFA8E.taxon	distribution	Distribution. Colombia, Peru.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204F382A4BC6FF4A7E4AFA8E.taxon	materials_examined	Type material examined. Syntype, ♀, deposited at the MNHNP (examined through photographs). Additional material examined. 1 ♀, “ Santa Fé, Bogotá / Colombia / Cornelius B. Philip Collection Donated to California Academy of Sciences / Scione equatoriensis Surc. / Scione equatoriensis Surcouf, det. C. B. Philip / Topotype, Cenotype, Scione / Compared with type ♀ P. incompleta Macq by C. B. Philip Aug. 53, in almost exact agreem. [agreement] / Scione incompleta? (Macq.) det. Krolow, T. K., 2012 [photographed specimen] ” (CAS); 1 ♀, “ Bogota, Colombia / BGuevara coll / Cornelius B. Philip Collection Donated to California Academy of Sciences / minor Macq. / Compared with type ♀ Fidena minor Macq., C. B. Philip 58 Agrees but palp [us] little longer and more slender ” (CAS); 1 ♀, “ Cordill. v. Columbien, terra templada, Thieme S. / Scione equatoriensis (Macq.) ♀ Dr. Enderlein det. 1915 [examined through photographs] ” (MNHUB).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204F382A4BC6FF4A7E4AFA8E.taxon	discussion	Comments. Both physically examined specimens and those examined by means of photographs show spots on crossveins slightly darker than on the type. The tuft of whitish setulae on the central posterior margin of tergites 3 and 4 appeared to be present also on the other tergites, whose setulae was lost. The photographs of the S. incompleta specimen provided by Lessard (2014) agree well with the specimens seen here and with the holotype.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204D382A4BC6FA33786BF8E8.taxon	description	(Fig. 50 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204D382A4BC6FA33786BF8E8.taxon	materials_examined	Type locality. Peru, Amazonas, Olmos-Pomacocha.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204D382A4BC6FA33786BF8E8.taxon	description	Description and discussion. See Oliveira et al. (2024).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204D382A4BC6FA33786BF8E8.taxon	distribution	Distribution. Peru.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33204D382A4BC6FA33786BF8E8.taxon	materials_examined	Holotype ♀. In good condition, deposited at the CAS.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238354BC6FF4A79E5FD07.taxon	description	(Fig. 51 A – D)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238354BC6FF4A79E5FD07.taxon	materials_examined	Type locality. Brazil, Mato Grosso.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238354BC6FF4A79E5FD07.taxon	diagnosis	Diagnosis. Body yellowish-brown. Antenna orange with the last flagellomeres darkened, scape and pedicel with long quite dense black setulae. Scutum and scutellum dark brown. Scutum with five slightly evident whitish longitudinal stripes. Wing slightly smoky with dark brown veins edged with brown. Tergites 1 and 2 yellowish and with dense long yellowish setulae on the sides. Tergite 1 with brown anterior region, tergite 2 with a central dark brown quadrangular spot in the middle. Other tergites dark brown with dense black setulae. Tergites 3 to 5 with tuft of yellowish setulae on the central posterior margin.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238354BC6FF4A79E5FD07.taxon	distribution	Distribution. Peru, Bolivia, Brazil (Mato Grosso).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238354BC6FF4A79E5FD07.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the MNHUB (examined through photographs).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238354BC6FF4A79E5FD07.taxon	discussion	Comments. This species is easily identifiable due to the wing veins being markedly dark brown and the presence of a quadrangular dark brown spot on tergite 2.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238374BC6FC9D7A59FE39.taxon	description	(Fig. 52 A – G)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238374BC6FC9D7A59FE39.taxon	materials_examined	Type locality. Peru, Cajamarca.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238374BC6FC9D7A59FE39.taxon	diagnosis	Diagnosis. Body yellowish-brown. Clypeus markedly developed, clypeal index greater than or equal to one. Antennae, palpus and legs orange. Thorax with five yellow longitudinal stripes. Wing hyaline with brown veins marginated with yellowish-brown. Abdomen yellow, tergite 3 to 7 darkened.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238374BC6FC9D7A59FE39.taxon	etymology	Etymology. From Latin, longibeccus (long beak). In reference to the very prominent clypeus.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238374BC6FC9D7A59FE39.taxon	description	Description, Holotype ♀. Body length: 12.7 mm, wing length: 13.7 mm. Head. Frons divergent below, divergence index 0.6, frontal index 1.6. Vertex dark brown with sparsely dense black and yellowish setulae, slightly prominent ocellar triangle, yellowish-brown ocelli. Frons and subcallus brown with orange pruinosity, frons with sparse black and yellow setulae. Parafacial brown with yellow pruinosity. Gena covered with yellow pruinosity and long, dense yellowish setulae. Clypeus brown with sparse short black setulae, markedly developed, clypeal index 1.1. Antenna orange. Scape and pedicel with yellowish pruinosity and short, dense black setulae. Palpus orange, long and lanceolate; first segment with dense, long yellowish setulae and some sparse black setulae; second segment with sparse black setulae on the ventral margin. Proboscis black, quite long, about 12 x the height of the frons. Thorax. Scutum and scutellum brown with sparse yellow setulae, slightly longer and denser on the posterior margin of the scutum and on the posterior margin of the scutellum. Scutum with yellow longitudinal stripes, the median stripe wider than the sublateral stripes. From each of the sublateral stripes a narrrower yellow band departs in the region of the transverse suture. Yellowish patch on the posterior margin of the scutum. Notopleuron with tuft of dense yellow setulae. Anepisternum with tuft of dense and long yellow and white setulae. Katepisternum and anepimeron with tuft of dense yellow setulae. Postalar callus with tuft of white and yellowish setulae. Legs yellow. Femora with long, dense yellow and black setulae. Tibiae with short, dense yellow setulae. Tarsi with short, dense yellow and black setulae. Wing hyaline with brown veins marginated with yellowish-brown. Abdomen. Tergites yellow. Tergites 1 and 2 with short, sparse yellow and black setulae. Tergite 1 with a small tuft of yellow setulae on the central posterior margin. Tergite 2 with a central brown patch of integument. Tergites 3 to 7 darkened with short, dense black and yellow setulae. Male. Unknown. Paratype variations. Body length: 11.5 – 12.5 mm, wing length: 12.6 – 13.8 mm, frontal index: 1.5 – 1.7, clypeal index: 1.0 – 1.2. The median thoracic stripe is less evident in both paratypes.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238374BC6FC9D7A59FE39.taxon	distribution	Distribution. Peru.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238374BC6FC9D7A59FE39.taxon	materials_examined	Material type. Holotype, ♀, “ Peru: Dept. Cajamarca, Prov. Jaén. Rio Huancabamba. Acapulco at km 81 Olmos-Marañón rd. 1100 m, 1 - x- 1964 / P. C. Hutchison & J. K. Wright AT Coleman lantern / Scione sp. aff. flavescens End. DET. 1981 G. B. Fairchild ” (CAS). Paratypes: 1 ♀ (in good condition, but left pedicel and flagellum missing, second segment of left palpus missing, tibia of right hind leg missing), “ Peru: Dpto. Huánuco: Cerca de Bosque Taprag, vii: 15: 68 (8,500 p.), Pedro Hocking ” (UNMSM); 1 ♀ (in good condition, but last tarsomeres of right hind leg missing, tibia of left hind leg missing), “ Peru: Dpto. Huánuco, Molinos (Panao), 23: vii: 84 (2,300 mt.), Pedro Hocking ” (UNMSM). Additional material. 1 ♀, “ Peru: HU. Dist. Quisqui, 25. vi. 2019, 9 º 54 ’ 3.43 ’’ S 76 º 22 ’ 37.45 ’’ W, 2532 m, I. Medina ” (UNMSM).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205238374BC6FC9D7A59FE39.taxon	discussion	Comments. The specimen mentioned in the additional material was identified as Scione longibeccus sp. nov., but was not included in the type series because, in addition to not being well preserved, it differs from the other specimens in having more marked brown patch on the frons, less evident thoracic stripes, slightly smoky wings and darker legs.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738304BC6FF4A7F15FCD2.taxon	description	(Fig. 53 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738304BC6FF4A7F15FCD2.taxon	materials_examined	Type locality. Peru, Junín, Chanchamayo.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738304BC6FF4A7F15FCD2.taxon	diagnosis	Diagnosis. Body predominantly yellow. Palpus and clypeus brown. Gena with dense and long yellowish setulae. Scutum and scutellum yellowish-brown. Scutellum with conspicuous yellow longitudinal stripes, the sublateral stripe wider and quite evident, the median stripe narrower and less evident. Wing slightly smoky, with faint brown spots on crossveins and yellowish patch (“ window ”) in the middle of cells r 1 and r 2 + 3. Legs dark brown. Abdomen yellow.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738304BC6FF4A7F15FCD2.taxon	distribution	Distribution. Peru.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738304BC6FF4A7F15FCD2.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the MACN (examined through photographs).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738304BC6FF4A7F15FCD2.taxon	discussion	Comments. The catalogues of Fairchild (1971), Moucha (1976), Fairchild & Burger (1994) and Coscarón & Papavero (2009) report that the holotype of S. longirostris was deposited at the MHNL (currently UNMSM) and had been lost. However, the holotype is deposited at the MACN (Bachmann 2012). This description of the species was originally described in French in 1920, and was later republished in 1937, in Spanish.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738314BC6FCAB780FF8F3.taxon	description	(Figs 54 A – D, 55 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738314BC6FCAB780FF8F3.taxon	materials_examined	Type locality. “ South America ” (Venezuela).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738314BC6FCAB780FF8F3.taxon	diagnosis	Diagnosis. Body orange brown. Thorax with conspicuous whitish longitudinal stripes. Wing smoky brown with yellow patch in the middle of cells r 1 and r 2 + 3.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738314BC6FCAB780FF8F3.taxon	description	Redescription. Female. Body length: 11.2 – 13.4 mm (X = 12.2 mm, N = 13), wing length: 11 – 13.2 mm (X = 12.1 mm, N = 14). Head. Frons nearly parallel, divergence index 0.8 – 0.9 (X = 0.9, N = 14), frontal index 2.0 – 2.4 (X = 2.2, N = 14). Vertex dark brown with long, dense black setulae, ocellar triangle little prominent, ocelli yellowish-brown. Frons and subcallus brown with yellowish pruinosity, frons with long, dense black setulae. Parafacial brown with sparse black setulae. Gena with white pruinosity and long, dense white setulae. Clypeus brown with sparse black setulae, clypeal index 0.7 – 0.9 (X = 0.8, N = 14). Antenna orange brown. Scape and pedicel with yellowish pruinosity and short, dense black setulae. Palpus orange brown; first segment with black and yellowish setulae; second segment with sparse black setulae and quite tapered at the tip. Proboscis dark brown, about 3 x the height of the frons. Thorax. Scutum and scutellum orange brown with dense coppery setulae and some black setulae. Scutum with conspicuous whitish longitudinal stripes. Median stripe narrow and not very evident. Sublateral stripes wider than the median and quite evident. The sublateral and lateral stripes connected by a narrow white stripe at the transverse suture. The sublateral stripes connect at 2 / 3 of the posterior margin of the scutum and form a rectangular whitish patch. Notopleuron with tuft of black and coppery setulae. Anepisternum, katepisternum and anepimeron with dense and long white and coppery setulae. Postalar callus with tuft of yellowish white setulae. Veins yellowish-brown with faint brown spots on crossveins. Femora orange brown with long, dense black setulae. Tibiae and tarsi yellowish-brown with short dense black setulae, hind tibiae and tarsi darkened. Wing smoky brown with yellow patch in the middle of cells r 1 and r 2 + 3 and no lighter areas in the middle of cells m 1 and m 2. Abdomen. Tergites orange brown with black and coppery setulae. Central posterior margin of tergites 2 to 5 with triangles of setulae and white pruinosity. Lateral margin of tergites with white setulae. Sternites with coppery and white setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738314BC6FCAB780FF8F3.taxon	distribution	Distribution. Costa Rica to Venezuela, Ecuador (Morona Santiago).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738314BC6FCAB780FF8F3.taxon	materials_examined	Type material examined. Syntype, ♀, deposited at the NMW (examined through photographs). Additional material examined. 1 ♀, “ 14 km. N. Puerto, Quepos, San Jose Prov. Costa Rica, 1 - 14 - 63, R. Casebeer / LACM ENT 286749 / Scione maculipennis ” (CEUFT); 1 ♀, “ Costa Rica, Puntarenas, La Amistad Nat. Park, 8 º 57 ’ N, 82 º 50 ’ W, 23 Feb 1991, 1500 m, Norman D. Penny Cal. Acad. Sci. Coll. ” (CAS); 3 ♀ “ Costa Rica, Puntarenas, La Amistad Nat. Park, 8 º 57 ’ N, 82 º 50 ’ W, 1500 m, 18 February 1991, Norman D. Penny Cal. Acad. Sci. Coll. ” (CAS); 1 ♀, Volcan Cacao, 1 km W. del Cerro Cacao, Prov. Guana, Costa Rica, 1400 m., 7 FEB 1995, F. Alvarado, L N 323700 # 4453 / Costa Rica INBIO CRI 002, 196112 / Scione maculipennis (Schiner) det. J. Burger 2000 / Scione maculipennis (Schiner) det. Krolow, T. K., 2012 ” (MNCR); 1 ♀, “ Panama, Chiriqui Prov., 2 km NE. of Boquete, 24 - IV- 1976, D. C. Rentz, M. S. Carter and C. L. Mullinex / Scione maculipennis Schin, Det. 1981, G. B. Fairchild / Scione maculipennis (Schiner) det. Krolow, T. K., 2012 ” (CAS); 1 ♀, “ Panama, Provincia de Chiriqui, S. slope of Volcan de Chiriqui, (Barú) 1000 m, 28. I. 1987, Edward S. Ross / Scione maculipennis Schin., Det. G. B. Fairchild 19 [?] ” (CAS); 1 ♀, “ Boquete, Panama, Chiriqui Prov., Marc. 26. 1923, F. M. Gaige, 293 / Cornelius B. Philip Collection, Donated to California Academy Sciences / Scione maculipennis Schin., det. C. B. Philip / eyes unbanded / Compared with type ♀ Scione maculipennis Schin, C. B. Philip 53, agrees (over), some of cotyp have yellower beards [on the back of the label] / Genotype Rhinotriclista End. (= Scione Wik.) / Scione maculipennis (Schiner), det. Krolow, T. K., 2012 ” (CAS); 1 ♀, “ Rio Chanjena, Boca del Toro, 2400 ff. / Panama, 1 sept. 1961 / Scione maculipennis Schin. det. G. B. Fairchild 1981 / Scione maculipennis ” (INPA); 1 ♀, “ Potrerillos, Panama, Elev. 2000, VI- 14 - 35 / I. W. Mac Swain coll. / Cornelius B. Philip Collection, Donated to California Academy Sciences / Scione maculipennis Macq. ♀ det. C. B. Philip 41 ” (CAS); 1 ♀, “ Potrerillos, Panama, Elev. 2000, VI- 14 - 35 / I. W. Mac Swain coll. / T. Aitken Collectin / Scione maculipennis Macq. ♀ det. C. B. Philip 41 ” (CAS); 1 ♀, “ Panama, Chiriqui Prov. 2 km NE. of Boquete, 24 - IV- 1976, D. C. Rentz, M. S. Carter and C. L. Mullinex ” (CAS); 1 ♀, “ Panama, Chiriqui Prov.: 45 Km N. of Chiriqui, Fortuna water shed, 20 June 1992, Jean & Keve Ribarbo ” (CAS); 1 ♀, “ Panama, Chiriqui, Alto Lino, 4000, 25 june 1965, H. G. Real / Herman G. Real Collection / Scione albifasciata ♀ (Macq) Det. C. B. Philip 72 ” (CAS); 1 ♀, “ Rio Pomeca, 2500 — 2600 m, Colombia, 18 - 21 - VII- 1948 / L. Richter coll., Frank Johnson Donor / Cornelius B. Philip Collection, Donated to California Academy of Sciences ” (CAS); 1 ♀, “ E. Loja, Ecuador, 2800 m, XI. 1970, L. E. Peña col. / MZSP ” (MZUSP).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205738314BC6FCAB780FF8F3.taxon	discussion	Comments. As mentioned in the original description, the boundaries of the wing spots are nowhere sharp, giving the wing surface a variegated, cloudy appearance that is readily visible to the naked eye. Triangles of setulae and white pruinosity extend across the entire abdominal tergites in some specimens. This species resembles S. albifasciata, from which it differs by the absence of patch on the frons, almost parallel frons and the pattern of spots on the wings. S. maculipennis has yellow spots scattered across more regions of the wing, whereas in S. albifasciata the yellow spots are restricted to cells r 1 and r 2 + 3. Wilkerson (1979) provided a redescription for S. maculipennis, the specimens examined here differs mainly in length and general body coloration. The specimens examined by Wilkerson measured between 9 and 11 mm, the specimens examined here measure between 11.2 and 13.4 mm, furthermore, according to the author, the species has a yellowish-brown body, but when examining the photographs of the types and the specimens on loan we saw that the general coloration of the body is orange brown. This species was redescribed because we examined a relatively large series of specimens and because S. maculipennis has many similar species (see comments on S. acris, S. claripennis, S. costaricana, and S. flavescens), thus an updated redescription may aid in distinguishing these species.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205B383D4BC6FF4A7800F81C.taxon	description	(Figs 56 A – F, 57 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205B383D4BC6FF4A7800F81C.taxon	materials_examined	Type locality. Ecuador, Azuay, Guasipampa.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205B383D4BC6FF4A7800F81C.taxon	description	Description and discussion. See Oliveira et al. (2024).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205B383D4BC6FF4A7800F81C.taxon	distribution	Distribution. Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205B383D4BC6FF4A7800F81C.taxon	materials_examined	Holotype ♀. In good condition, deposited at the MECN.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332059383E4BC6FF4A7F7CF8D4.taxon	description	(Fig. 58 A – D)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332059383E4BC6FF4A7F7CF8D4.taxon	materials_examined	Type locality.? America.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332059383E4BC6FF4A7F7CF8D4.taxon	diagnosis	Diagnosis. Body yellowish-brown. Thorax with yellowish-white longitudinal stripes, the median stripe narrow and not very evident and the sublateral stripes a little wider and quite evident. Wing brownish on the anterior region and smoky on the posterior region. Veins brown and crossveins with brown spots.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332059383E4BC6FF4A7F7CF8D4.taxon	description	Redescription. Female. Body length: 10.5 mm, wing length: 10.8 mm. Head. Frons divergent below, divergence index 0.7 and frontal index 1.7. Frons with dark brown patch almost touching the eyes and extending to the vertex. Vertex dark brown with dense and long black setulae, ocellar triangle barely noticeable, brown ocelli barely evident. Frons brown with dense yellowish pruinosity and long dense black setulae. Subcallus brown with slightly dense yellowish pruinosity. Parafacial brown with whitish pruinosity and sparse long black setulae. Gena with whitish pruinosity and long, dense, yellowish-white setulae. Clypeus brown with long dense black setulae and whitish pruinosity on the sides, clypeal index 0.6. Scape and pedicel brown with slightly dense yellowish pruinosity and long dense black setulae. Palpus brown, slightly tapered; first segment with long dense yellowish setulae and some black setulae; second segment orange on the dorsal margin and with short, dense black setulae. Proboscis shiny dark brown, about 2 x the height of the frons. Thorax. Scutum dark brown with long dense black setulae and some orange setulae. Scutum with yellowish longitudinal stripes extending to the posterior margin. Median stripe narrow and not very evident. Sublateral stripes slightly wider and quite evident. Sublateral and lateral stripes connected by a narrow yellow-white stripe at the transverse suture. Scutellum brown with central orange patch and dense long black setulae on the posterior margin. Notopleuron and anepisternum with tuft of dense long black setulae. Katepisternum and anepimeron with tuft of dense long whitish setulae. Postalar callus with tuft of yellowish setulae. Legs orange. Femora with long, dense black setulae and some yellowish setulae. Tibiae and tarsi with short dense black setulae. Wing brownish on the anterior region and smoky on the posterior region. Brown veins, brown spots on crossveins and yellowish spots on cells r 1 and r 2 + 3. Abdomen. Tergite 1 yellow-orange with a small tuft of yellowish setulae on the central posterior margin. Tergite 2 yellow-orange with a central black patch. Other tergites dark brown with an orange posterior margin, tergites 3 to 6 with a tuft of yellowish setulae on the central posterior margin. All tergites with short dense black setulae. Sternites yellowish-brown with short dense black and white setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332059383E4BC6FF4A7F7CF8D4.taxon	distribution	Distribution. Colombia, Venezuela.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332059383E4BC6FF4A7F7CF8D4.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the NHMUK (examined through photographs). Additional material examined. 1 ♀, “ Monte Redondo, Meta, 1300 m, Rastrojo, Colombia, 19 - III- 1948 / L. Richter coll. Frank Johnson Donor / Cornelius B. Philip Collection Donated to California Academy of Sciences / Scione minor ♀ Macq., Det. C. B. Philip 61 [?] ” (CAS).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332059383E4BC6FF4A7F7CF8D4.taxon	discussion	Comments. The flagellum of the examined specimen is missing, but examination of the holotype photograph shows that the species has orange flagellum. The original description of S. minor is succinct, and the holotype is in very poor conditions of preservation, but the shape of the patch on the frons, the coloration of the palpus, legs and abdomen are consistent with the specimen examined. Scione minor is very similar to S. incompleta, differing in that S. incompleta has predominantly yellowish abdomen and the thoracic stripes that are not very evident, whereas S. minor has dark brown last tergites and the sublateral thoracic stripes clearly visible.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F38384BC6FF4A78ADFD2A.taxon	materials_examined	Type locality. Colombia, San Lorenzo.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F38384BC6FF4A78ADFD2A.taxon	description	Original description. Closely allied to S. claripennis Ric. only being distinguished by black antennae and palpi, by brown middle part of antennal segment, by 2 grayish tomented cross-bands on face brown front, by incomplete second posterior veinlet, wide yellowish segmental borders on abdomen and by its smaller size, the type from S. Lorenzo, Columbia, being 9 mm long.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F38384BC6FF4A78ADFD2A.taxon	distribution	Distribution. Colombia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F38384BC6FF4A78ADFD2A.taxon	materials_examined	Holotype ♀. Destroyed (MTMB).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F38384BC6FF4A78ADFD2A.taxon	discussion	Comments. According to the original description, which is succinct, this species is similar to S. claripennis, differing from it in the coloration of the antenna and palpus. Scione claripennis has orange-brown scape and pedicel and orange-brown flagellum with the last flagellomeres darkened, the first segment of the palpus is dark brown and the second orange-brown, whereas S. minuta has black antennae and palpus.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F383A4BC6FCF37B87FE93.taxon	description	(Fig. 59 A – G)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F383A4BC6FCF37B87FE93.taxon	materials_examined	Type locality. Colombia, Magdalena.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F383A4BC6FCF37B87FE93.taxon	diagnosis	Diagnosis. Body dark brown with contrasting dense yellow setulae on the gena. Thorax with yellow longitudinal stripes that do not extend to the posterior margin of the scutum. Fore and mid legs markedly bicolored, with black femora and yellow tibiae. Abdomen dark brown with very dense and short black setulae. Tergites 1 and 2 with orange lateral margin. Tergites 2 to 6 with small tuft of yellow setulae on the central posterior margin.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F383A4BC6FCF37B87FE93.taxon	etymology	Etymology. From Latin, nigellus (black), in reference to the predominantly dark brown color of the species.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F383A4BC6FCF37B87FE93.taxon	description	Description, Holotype ♀. Body length: 11.1 mm, wing length: 12.4 mm. Head. Frons nearly parallel, divergence index 0.9, frontal index 2.4. Vertex dark brown with long dense black setulae, ocellar triangle not very prominent, ocelli brown. Frons and subcallus brown with dense yellow pruinosity. Frons with very dense and long black setulae and a rounded black patch. A narrow black line extends from the patch to the vertex. Parafacial brown with white pruinosity and some black setulae. Gena covered with white pruinosity and dense long yellow setulae. Clypeus brown with white pruinosity and dense long black setulae, clypeal index 0.7. Antenna orange. Scape and pedicel with long dense black setulae. Palpus dark brown; first segment with dense long yellow setulae in the proximal portion and dense long black setulae in the distal portion; second segment with orange dorsal margin and sparse black setulae. Proboscis shiny black, about 1.5 x the height of the frons. Thorax. Scutum dark brown with dense, short black setulae and yellow longitudinal stripes that do not extend to the posterior margin of the scutum. Scutellum black with dense, long black setulae. Notopleuron with tuft of long dense black and yellow setulae. Anepisternum, katepisternum and anepimeron with tuft of long dense black setulae. Postalar callus with tuft of long dense black setulae. Wing smoky with brown veins and faint brown spots on crossveins. Femora black with long dense black setulae. Fore and mid tibiae yellow with short dense yellow setulae and some black setulae. Hind tibiae brown with short dense yellow setulae on the ventral region and short dense black setulae on the dorsal region. Fore tarsus yellow with short dense yellow and black setulae. Mid and hind tarsus black with short dense black setulae. Abdomen. Tergites dark brown with very dense and short black setulae. Tergites 1 and 2 with orange lateral margin. Tergites 2 to 6 with a small tuft of yellow setulae on the central posterior margin. Male. Unknown.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F383A4BC6FCF37B87FE93.taxon	distribution	Distribution. Colombia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F383A4BC6FCF37B87FE93.taxon	materials_examined	Type material. Holotype, ♀, “ Colombia, Magdalena, Sierra Nevada de Santa Marta, cloud forest above San Pedro de la Sierra, 18.12.76, M. Cooper / M. Cooper Neotropical Tabanidae BMNH (E) 2004 - 182 / Scione? sp. nov., det. J. E. Chainey, 2004 / NHMUK 014428018 ” (NHMUK).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205F383A4BC6FCF37B87FE93.taxon	discussion	Comments. Scione nigella sp. nov. is similar to S. immaculata and S. ablusus. It differs by having a gena covered with long dense yellow setulae, markedly bicolored fore and mid legs, with black femora and yellow tibiae, and smoky wings with faint brown spots on crossveins. Whereas S. immaculata has gena covered with long, dense whitish setulae, orange-brown legs, and slightly smoky wings without spots and with yellowish pterostigma. Scione ablusus has gena covered with long, dense whitish setulae, yellow legs and smoky brown wings without spots (Fig. 60).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D383A4BC6FC297F00F9C2.taxon	description	(Fig. 61 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D383A4BC6FC297F00F9C2.taxon	materials_examined	Type locality. “ New Granada ” (Colombia).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D383A4BC6FC297F00F9C2.taxon	diagnosis	Diagnosis. Body yellowish-brown. Frons broad covered with white pruinosity. Palpus dark brown, quite sharp. black, about twice the height of the frons. Gena with dense, long whitish setulae. Thorax brown with two incomplete yellowish sublateral stripes. Legs dark brown. Wing smoky yellow. Abdominal tergites 1 and 2 yellowish-brown, the first a little darker. Tergite 3 brown with yellowish posterior margin. Other tergites dark brown.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D383A4BC6FC297F00F9C2.taxon	distribution	Distribution. Colombia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D383A4BC6FC297F00F9C2.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the MLUH (examined through photographs).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D383A4BC6FC297F00F9C2.taxon	discussion	Comments. Scione nigripes is similar to S. cingulata. It differs in that S. cingulata has brown thorax with five yellowish longitudinal stripes, dark brown frons and is less wide than in S. nigripes.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D38064BC6F9587992FAA2.taxon	description	(Figs 62 A – F, 63 A – G)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D38064BC6F9587992FAA2.taxon	materials_examined	Type locality. “ Colombia, Peru ”.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D38064BC6F9587992FAA2.taxon	diagnosis	Diagnosis. Body yellowish-brown. Thorax and scutellum dark brown, scutum with three yellow longitudinal stripes. Wing smoky with brown spots on crossveins. Bicolored legs. Femora dark brown, tibiae and tarsi yellow.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D38064BC6F9587992FAA2.taxon	description	Redescription. Female. Body length: 9.5 – 11 mm (N = 2), wing length: 10.4 – 10.9 mm (N = 2). Head. Frons divergent below, divergence index 0.7 – 0.8 (N = 2), frontal index 1.9 – 2.0 (N = 2). Vertex dark brown with long dense black setulae, prominent ocellar triangle, yellow ocelli. Frons with dense yellow pruinosity and short dense black setulae, well-marked central brown patch. Subcallus brown with dense yellow pruinosity on the sides. Parafacial brown with yellowish pruinosity and short sparse black setulae. Gena with yellow pruinosity and long dense yellow setulae. Clypeus dark brown with short dense black setulae and sparse whitish pruinosity on the sides, clypeal index 0.7 (in both specimens). Scape and pedicel brown with long dense black setulae. Flagellum orange, last flagellomere slightly darkened. Palpus brown; first segment brown with long dense black and yellow setulae; second segment with orange dorsal margin. Proboscis shiny dark brown, about 2 x the height of the frons. Thorax. Scutum and scutellum black. Scutum with dense short black and yellow setulae and three yellowish longitudinal stripes. The median stripe is wide and less marked. The sublateral stripes narrow and quite evident, wider on the posterior margin of the scutum, forming a quadrangular orange patch. Scutellum with dense and long black setulae on the posterior margin. Notopleuron with tuft of long dense black and yellow setulae. Anepisternum with tuft of long dense black and yellow setulae. Katepisternum and anepimeron with tuft of long dense yellow setulae. Postalar callus with tuft of long dense white setulae. Femora dark brown with long, dense black setulae. Tibiae and tarsi yellowish-brown with short dense black setulae. Wing smoky with brown veins and brown spots on crossveins, yellowish spot on the anterior region, in the region of cells r 1 and r 2 + 3. Abdomen. Tergites 1 – 3 yellow with brown central median region and tuft of yellow setulae on the central posterior margin. Tergite 1 with a central median brown patch of tegument and a tuft of yellow setulae on the central posterior margin. Tergites 2 and 3 with a central median brown patch, tergite 3 with sparse black setulae. Tergites 4 to 7 darker with dense, short black setulae and tufts of yellowish setulae on the central posterior margin.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D38064BC6F9587992FAA2.taxon	distribution	Distribution. Colombia, Ecuador, Peru.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D38064BC6F9587992FAA2.taxon	materials_examined	Type material examined. Syntype, ♀, deposited at the MNHUB (examined through photographs). Additional material examined. 1 ♀, “ Peru, CA [Cajamarca], San Ignacio, Namballe, 2095 m, 5 º 5 ’ 59.8 ’’ S / 79 º 14 ’ 15.3 ’’ O, 08 – 10. x. 2017, P. Sánchez ” (UNMSM); 1 ♀, “ Peru, CA, San Ignacio, Tabaconas, 2549 m, 5 º 11 ’ 56.7 ’’ S / 79 º 20 ’ 21.1 ’’ O, 13 – 15. x. 2017, P. Sánchez [photographed specimen] ” (UNMSM).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33205D38064BC6F9587992FAA2.taxon	discussion	Comments. Scione obscurefemorata is similar to S. brevistriga. It differs in that S. brevistriga has yellow, unicolorous legs and the spots on crossveins are weaker. Wilkerson (1979) provided a complete redescription of the species. The redescription is consistent with the characters that can be seen in the type, which is not well-preserved, and in the specimens examined here. In his redescription, Wilkerson reported that tergites 1 to 6 have tufts of light setulae on the central posterior margin. In the specimens seen by us, many of the setulae are missing, and it is only possible to observe the tuft of setulae on tergite 1. According to the author, the species appears to exhibit considerable variation in the structure of the palpus, which can be quite short, intermediate as in the type, or longer and more pointed as in the specimens seen by him.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206138024BC6FA7B7A43FC77.taxon	description	(Figs 64 A – G, 65 A – G)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206138024BC6FA7B7A43FC77.taxon	materials_examined	Type locality. Ecuador, San Vicente.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206138024BC6FA7B7A43FC77.taxon	diagnosis	Diagnosis. Body yellowish-brown. Thorax dark brown to black with five whitish longitudinal stripes. Abdomen yellow, tergites 1 to 7 with tuft of white setulae on the central posterior margin. Wing smoky brown on the anterior region and hyaline on the posterior region. Yellowish spots on cells r 1, r 2 + 3, r 4, r 5, radial basal and in part of the medial basal cell (“ windows ”).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206138024BC6FA7B7A43FC77.taxon	etymology	Etymology. The specific epithet “ philipi ” is a tribute to Cornelius Becker Philip for his contributions to the knowledge of tabanids.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206138024BC6FA7B7A43FC77.taxon	description	Description, Holotype ♀. Body length: 9.2 mm, wing length: 11 mm. Head. Frons parallel, divergence index 1.0, frontal index 2.0. Vertex dark brown with long, dense black setulae, ocellar triangle not very prominent, ocelli brown. Frons covered with white pruinosity and a dark brown patch that extends to the vertex, dense and long black setulae along the entire length of the patch. Subcallus brown with white pruinosity on the sides. Parafacial brown with dense white pruinosity and long sparse black setulae. Gena with white pruinosity and dense long white setulae. Clypeus brown with slightly dense white pruinosity and long dense black setulae, clypeal index 0.7. Scape and pedicel brown with yellowish pruinosity and long dense black setulae. Flagellum orange, last flagellomeres dark. Palpus dark brown, short and rounded; first segment with dense long white setulae in the proximal portion and dense long black setulae in the distal portion; second segment with dense long black setulae and an orange dorsal margin. Proboscis shiny black, about 2 x the height of the frons. Thorax. Scutum brown with short dense black setulae and whitish longitudinal stripes. The median stripe is narrow and less evident. The sublateral and lateral stripes wider and more evident and connected by a white stripe at the transverse suture. Scutellum dark brown with long dense black setulae on the posterior margin. Notopleuron with tuft of long dense black setulae. Anepisternum with tuft of long dense black setulae. Katepisternum and anepimeron with tufts of long dense white setulae. Postalar callus with tufts of long dense white setulae. Femora brown with long dense black and white setulae. Tibiae and tarsi slightly lighter than femora and with short dense black setulae and some yellow setulae. Wing smoky brown on the anterior region and hyaline on the posterior region. Yellowish spots on cells r 1, r 2 + 3, r 4, r 5, radial basal and in part of the medial basal cell (“ window ”). Veins marginated with brown and brown spots on crossveins. Abdomen. Tergites yellow with short, dense black setulae. Tergites 1 to 7 with a tuft of white setulae on the central posterior margin. Tergite 1 with brown anterior margin. Tergites 2 with an irregular central median brown patch. Tergites 4 to 7 dark. Sternites yellow with short dense white setulae. Male (Fig. 65). Like the female, except for the holoptic condition, porrect palpus and longer and denser body setulae. In addition, the clypeus of males is less developed than in females. Variations of paratypes. Body length: 9.6 – 11.5 mm (X = 10.6 mm, N = 10), wing length 9.8 – 11.1 mm (X = 10.5 mm, N = 10), frontal index 1.6 – 2.0 (X = 1.7, N = 5), divergence index 0.7 – 0.9 (X = 0.8, N = 5), clipeal index 0.5 – 0.7 (X = 0.6, N = 10). The dark pruinosity on the frons and the spots on the wing are lighter in some specimens and more marked in others.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206138024BC6FA7B7A43FC77.taxon	distribution	Distribution. Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206138024BC6FA7B7A43FC77.taxon	materials_examined	Type material: Holotype, ♀, “ San Vicente, 2.770 m, Morona-San [tiago] prov., J. Buestán, 11. x. 87 / CJB- 3404 ” (MECN). Paratypes: 1 ♀ (in good condition, but pedicel and left flagellum are missing), “ idem, except CJB- 3405 ” (MECN); 1 ♀, (in good condition, but the upper edge of right wing and lower edge of left wing with a small broken area), “ Tinajillas, 2.100 m, Morona-San. Prov., Jaime Buestán, 22. xi. 83 / CJB- 3350 / 6 ” (MECN); 1 ♀, (in good condition, but the upper edge of left wing with a small broken area), “ Potrerillo [s], 3.040 m, 23 - xi- 85 / CJB- 3351 / 6 ” (MECN); 1 ♀ (in good condition, but right mid leg is missing), “ idem, except CJB- 3352 ” (INPA); 1 ♂ (in good condition), “ San Vicente, 2.770 m, Morona-San prov., J. Buestán, 11. x. 87 / CJB- 3447 ” (MECN); 1 ♂ (in good condition, but left hind leg and missing right hind tarsus are missing), “ idem, except CJB- 3448 ” (MECN); 1 ♂ (photographed specimen; in good condition), “ idem, except 18. ix. 87, CJB- 3440 ” (MECN); 1 ♂ (in good condition, but left hind leg is missing), “ idem, except CJB- 3437 ” (MECN); 1 ♂ (in good condition, but the upper margin of right wing with a small broken area, left hind tarsus are missing), “ idem, except CJB- 3438 ” (MECN); 1 ♀ (in good condition, but right pedicel and flagellum are missing, upper margin of left wing with a small broken area), “ Ecuador, Tungurahua, Banos, 20. xi. 1978, M. Cooper / Cloud forest, c. 2,500 m. / M. Cooper Neotropical Tabanidae BMNH (E) 2004 - 182 / Scione det. J. E. Chainey, 2004 / NHMUK 014428028 ” (NHMUK); 1 ♀ (in good condition), “ Ecuador, Pichincha, km 20 Aloag-StoDomingo Hda. Bomboli, 8 - 11 / vii / 2006, Col. J. Buestan / CJB- 7918 ” (MECN); 1 ♀ (in good condition, but crushed abdomen), “ idem, except CJB- 7919 ” (MECN); 1 ♀ (in good condition, but left mid leg is missing), “ Ecuador, Loja, Cord. Sabanilla (2450 m) 4 y 11 Oct 2006, J. Buestan & M. Mejia / CJB- 8454 ” (MECN); 1 ♀ (in good condition, but distal margin of both wings with a small broken area), “ Ecuador, Pichincha, km 20 Aloag-StoDomingo Hda. Bomboli, 8 – 11 / vii / 2006, Col. J. Buestan / CJB- 7920 ” (MECN); 1 ♀, (in good condition, but some of the setulae on the abdomen are missing), “ Ecuador, Loja, Cord. Sabanilla (2450 m) 4 y 11 Oct 2006, J. Buestan & M. Mejia / CJB- 8453 ” (MECN); 1 ♀ (in good condition, but some of the setulae on the abdomen are missing), “ idem, except CJB- 8455 ” (MECN); 1 ♀ (in good condition, but right pedicel and flagellum are missing, right wing with apex broken), “ Tinajillas, 2.100 m, Morona-San. Prov. J. Buestán, 10. x. 87 / CJB- 4311 ” (MECN); 1 ♀ (in good condition), “ idem, except CJB- 4314 ” (MECN); 1 ♀ (in good condition), “ idem, except CJB- 4315 ” (MECN); 1 ♀ (in good condition, but right wing apex broken), “ idem, except CJB- 4316 ” (MECN); 1 ♀ (in good condition, but left wing apex broken), “ idem, except CJB- 4310 ” (MECN); 1 ♀ (in good condition, but right wing apex broken), “ idem, except CJB- 4312 ” (MECN); 1 ♀ (in good condition, but left wing apex broken), “ idem, except CJB- 4313 ” (MECN); 1 ♀ (in good condition), “ idem, except 26. x. 85 / CJB- 4264 ” (MECN); 1 ♀ (in good condition), “ idem, except CJB- 4267 ” (MECN); 1 ♀ (in good condition), “ idem, except CJB- 4265 ” (MECN); 1 ♀ (in good condition, but left pedicel and flagellum, and left fore leg are missing), “ Ecuador, Loja, Cor. Sabanilla (2450 m), 23. [illegible number]. 2007, J. Buestán / CJB- 9010 ” (MECN). Additional material. 1 ♀, “ Ecuador, Tungurahua, Pastaza valley at Banos. c. 2000 m, 10. x. 1978, M. Cooper / M. Cooper Neotropical Tabanidae BMNH (E) 2004 - 182 / NHMUK 014428034 ” (NHMUK); 1 ♂, “ Ecuador, Tungurahua, above Banos, c. 2,500 m, 12. x. 1978, M. Cooper / M. Cooper Neotropical Tabanidae BMNH (E) 2004 - 182 / NHMUK 014428031 ” (NHMUK); 1 ♂, “ idem, except NHMUK 014428030 ” (NHMUK); 1 ♂, “ idem, except NHMUK 014428029 ” (NHMUK); 1 ♂, “ idem, except NHMUK 014428032 ” (NHMUK).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206138024BC6FA7B7A43FC77.taxon	discussion	Comments. The palpus of S. philipi sp. nov. varies among the specimens examined and may be short and rounded or slightly longer and tapered. The spots on crossveins are darker in some specimens. The specimens mentioned in the additional material were identified as S. philipi sp. nov., but were not included in the type series because they are not in good condition of preservation. Therefore, it is not possible to fully observe the diagnostic characteristics of the species. S. philipi sp. nov. is similar to S. merianae, differing in that S. merianae has wider and more marked thoracic stripes, dark brown antennae, wings with very evident brown spots on crossveins and without yellowish patch, and abdomen with very evident pruinosity and setulae bands white on the tergites. Whereas S. philipi sp. nov. has evident but less wide thoracic stripes, orange antennae, wings with very evident brown spots on the crossveins and lacking a yellowish patch, and abdomen without pruinosity and setulae bands on the tergites (Fig. 66).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332065380E4BC6FC0E78B1FDFD.taxon	description	(Figs 67 A – G, 68 A – G)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332065380E4BC6FC0E78B1FDFD.taxon	materials_examined	Type locality. Peru, Lima.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332065380E4BC6FC0E78B1FDFD.taxon	diagnosis	Diagnosis. Body yellowish-brown. Thorax dark brown with slightly evident white longitudinal stripes. Wing hyaline with brown veins and faint brown spots on crossveins. Abdomen predominantly yellow.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332065380E4BC6FC0E78B1FDFD.taxon	etymology	Etymology. The specific epithet “ peruviana ” is a reference to the country in which the specimens were collected (Peru).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332065380E4BC6FC0E78B1FDFD.taxon	description	Description, Holotype ♀. Body length: 12 mm, wing length: 12.4 mm. Head. Frons divergent below, divergence index 0.7, frontal index 1.5. Vertex brown with gray pruinosity and long dense black setulae, prominent ocellar triangle, brown ocelli. Frons and subcallus brown with dense gray pruinosity, frons with long sparse black setulae. Parafacial brown with dense gray pruinosity and some black setulae. Gena with gray pruinosity and dense long yellow setulae. Clypeus brown with slightly dense gray pruinosity and long sparse black setulae, clypeal index 0.9. Scape and pedicel brown with long dense black setulae. Flagellum orange, last flagellomeres brown. Palpus brown, long and lanceolate; first segment with long dense yellow setulae and some black setulae; second segment with orange dorsal margin and short sparse black setulae. Proboscis shiny black, about 6 x the height of the frons. Thorax. Scutum dark brown with sparse black and yellow setulae and with slightly evident whitish longitudinal stripes. From each of the sublateral stripes a narrower white stripe starts at the transverse suture. Scutellum dark brown with yellowish pruinosity on the posterior margin and long sparse black setulae. Notopleuron with tuft of long dense black and yellow setulae. Anepisternum, katepisternum and anepimeron with dense gray pruinosity and long dense yellow setulae. Postalar callus with tuft of dense long yellow setulae. Femora with long dense black setulae, proximal half of femora brown, distal portion yellow. Tibiae and tarsi yellow with short dense black setulae. Wing hyaline with brown veins and faint brown spots on crossveins. Small appendix at the fork of vein R 4 + 5. Abdomen. Tergite 1 with brown anterior margin and yellow posterior margin and with a posterior band of yellow setulae. Tergites 2 yellow with dense short yellow setulae and a central brown patch. Tergites 3 to 7 with brown anterior margin and yellowish posterior margin. Tergites 3 and 4 with short sparse black setulae. Tergites 5 to 7 with short dense yellow and black setulae. Sternites brown with short dense black setulae and a yellowish posterior margin. Male (Fig. 68). Like the female, except for the holoptic condition, porrect palpus and longer and denser body setulae. In addition, the clypeus of males is less developed than in females. Variations of paratypes. Body length: 11.5 – 13 mm (X = 12.4 mm, N = 3), wing length 10 – 11.5 mm (X = 10.9 mm, N = 3), clypeal index 0.5 (in all three specimens). The tibiae and tarsi of males are darker than those of females.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332065380E4BC6FC0E78B1FDFD.taxon	distribution	Distribution. Peru (Cajamarca, Lima).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332065380E4BC6FC0E78B1FDFD.taxon	materials_examined	Type material: Holotype, ♀, “ Peru, Dpto. Lima, Cerro de Surco (3,000 mt.) 21: iv: 73, P. Hocking ” (UNMSM). Paratypes: 1 ♂ (photographed specimen; in good condition, but last tarsomeres of left hind leg are missing), “ Peru, Dpto. Cajamarca, Prov. Cascascas [Cascas?], San Benito, 12: iii: 87 (1,000 mt.), P. Hocking ” (UNMSM); 2 ♂, (in good condition) “ Peru, Dpto. Cajamarca, Prov. Cascas, San Benito, 12: iii: 87 (1,000 mt.), P. Hocking ” (UNMSM).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332065380E4BC6FC0E78B1FDFD.taxon	discussion	Comments. In males, the body setulae are better preserved and it is possible to observe the presence of dense white setulae on the sternites, which have apparently been lost in the female.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332069380E4BC6FD8679A8FAE8.taxon	materials_examined	Type locality. Bolivia, San Antonio.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332069380E4BC6FD8679A8FAE8.taxon	description	Original description. A reddish-yellow species with black and whitish marks. Head brownish-yellow with whitish-yellow beard, lighter parts around bronze brown eyes and brown, club shaped spot in mid of front. Antennae and palpi reddish, basal joints of antennae whitish pollinose with yellow hair. Haustellum black, stilettos reddish-yellow. Thorax grayish yellow with reddish-brown and (around wings) gray hair, scutum with 5 dark brown spots, a median one with lineal extension to scutellum of same color, 2 anterior, smaller and 2 posterior larger lateral ones all with their apices directed towards scutellum and separated from one another by 2 Y-shaped whitish-gray stripes. Legs and halteres reddish-yellow, with golden yellow hair, coxae, claws and basal setulae of femora black. Wings smoky, median and basal parts yellowish translucent, all cross veins with dark brown borders. Abdomen reddish-yellow, covered with long, appressed, glistening rufous hair; front borders of tergites with larger brown, hind borders with small white spots, the latter being composed only of white hair.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332069380E4BC6FD8679A8FAE8.taxon	distribution	Distribution. Bolivia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332069380E4BC6FD8679A8FAE8.taxon	materials_examined	Holotype ♀. Destroyed (MTMB).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332069380E4BC6FD8679A8FAE8.taxon	discussion	Comments. In Szilády’s key (1926) the closest species to S. picta is S. incompleta, differing because S. incompleta has brown palpus and yellowish-brown wing on the anterior margin and smoky on the posterior margin, whereas S. picta has reddish palpus and smoky wing.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332069380E4BC6FAB57E3FF80F.taxon	materials_examined	Type locality. Colombia, Sierra San Lorenzo.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332069380E4BC6FAB57E3FF80F.taxon	description	Original description. Almost unicolorous pale brown with grayish hair which is lighter on ventral surface; spurs and claws of legs brown. Head darker brown; front parallel, index: 21 / 4: 1; antennal segment and oblong median portion of front elevated, slightly shining; vertex with brown hair; proboscis dark brown, as long as head. Eyes purplish brown with yellow hair. Thorax with two narrow, faint, whitish stripes anteriorly ending in a triangular patch near middle of scutum. Wings clear, all longitudinal veins yellow, all cross veins black with narrow, brown border; second posterior veinlet complete. Abdomen with indistinct brownish bands on hind tergites.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332069380E4BC6FAB57E3FF80F.taxon	distribution	Distribution. Colombia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332069380E4BC6FAB57E3FF80F.taxon	materials_examined	Holotype ♀. Destroyed (MTMB).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332069380E4BC6FAB57E3FF80F.taxon	discussion	Comments. The species cannot be compared with any other due to lack of information in the original description. The record made in Ecuador by Cardenas & Vieira (2005) should not be validated. The authors mention that the species was recorded in Ecuador by Fairchild & Léon (1986), however, in this work the authors claim not to have seen specimens from Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F38084BC6FF4A7F9BFCF3.taxon	description	(Fig. 69 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F38084BC6FF4A7F9BFCF3.taxon	materials_examined	Type locality. Colombia, Valle, Peñas Blancas, 1750 m, 10 km west of Cali.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F38084BC6FF4A7F9BFCF3.taxon	diagnosis	Diagnosis. Body predominantly brown. Thorax with narrow and inconspicuous yellowish longitudinal stripes. Wing smoky with yellowish patch in the middle of cells r 1 and r 2 + 3 (“ window ”). Legs yellowish-brown. Proboscis dark brown with small tufts of setulae on the sides, just above the lip. Tergites 1 to 6 with tuft of yellowish setulae on the central posterior margin.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F38084BC6FF4A7F9BFCF3.taxon	distribution	Distribution. Colombia (Valle).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F38084BC6FF4A7F9BFCF3.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the FSCA (examined through photographs).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F38084BC6FF4A7F9BFCF3.taxon	discussion	Comments. Scione rhinothrix is similar to S. equivexans. It differs in having more developed clypeus than S. equivexans, which is almost as long as the height of the frons. The frons of S. rhinothrix is nearly parallel, whereas in S. equivexans the frons diverges below. In S. rhinothrix the yellow spots on the wing are restricted to cells r 1 and r 2 + 3, whereas in S. equivexans the yellow spots are present in cells r 1, r 2 + 3, r 5, basal radial, basal medial and the discal cell. Furthermore, S. equivexans has dark brown legs and S. rhinothrix has yellowish-brown legs.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F380B4BC6FC8A7FC1FD72.taxon	description	(Figs 70 A – F, 71 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F380B4BC6FC8A7FC1FD72.taxon	materials_examined	Type locality. Peru, Callanga.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F380B4BC6FC8A7FC1FD72.taxon	diagnosis	Diagnosis. Body predominantly brown. Clypeus marked prominence at its apex. Thorax without stripes, wings slightly smoky without spots on crossveins. Abdomen with dense coppery setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F380B4BC6FC8A7FC1FD72.taxon	description	Redescription. Female. Body length: 17 – 17.4 mm (X = 17.2 mm, N = 3), wing length: 17.5 – 18.1 mm (X = 17.8 mm, N = 3). Head. Frons nearly parallel, divergence index 09 – 1.1 (X = 1.0, N = 3), frontal index 2.5 – 2.9 (X = 2.7, N = 3). Frons unspotted. Vertex dark brown with orange pruinosity and not very dense black setulae, ocellar triangle prominent, ocelli orange brown. Frons and subcallus dark brown; frons with sparse black setulae. Parafacial and gena dark brown with orange pruinosity; gena with dense and long black and orange setulae. Clypeus dark brown with some black setulae, marked prominence at its apex, clypeal index 0.7 – 0.9 (X = 0.8, N = 3). Scape and pedicel dark brown with black setulae, flagellum orange brown. Palpus dark brown with black setulae on both segments; second segment larger than the first and quite tapered towards the apex. Proboscis bright orange brown, about 3 x the height of the frons, lip short dark brown. Thorax. Scutum dark brown with dense black and orange setulae. Scutellum orange brown with dense black and orange setulae. Notopleuron with a small tuft of black and orange setulae. Anepisternum, katepisternum and anepimeron with tufts of long dense black and orange setulae. Postalar callus with long, dense orange setulae. Coxae and trochanters dark brown with dense black setulae. Femora orange brown with long dense black setulae. Tibiae and tarsi orange brown with short dense black and orange setulae. Wing slightly smoky with brown veins. Abdomen. Tergites dark brown with dense coppery setulae on the dorsal margin and on the lateral margin of the tergites. Sternites dark and with sparse black and coppery setulae.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F380B4BC6FC8A7FC1FD72.taxon	distribution	Distribution. Eastern Andes of Ecuador, Bolivia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F380B4BC6FC8A7FC1FD72.taxon	materials_examined	Type material examined. Paratype, ♀, deposited at the NHMUK (examined through photographs); Lectotype of Rhinotriclista generosa, ♀, deposited at the MNHUB (examined through photographs). Additional material examined. 3 ♀, “ Peru, Cuzco, Cosñipata Valley Quebra. San Pedro, 13 º 03 ’ S, 71 º 30 ’ W, 1400 m, 1 – 3 - II- 2013, Mielke Leg. / Scione sp. det. Henriques, 2016 (INPA) ”.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206F380B4BC6FC8A7FC1FD72.taxon	discussion	Comments. The paratype studied has two faint lighter stripes on the scutum, a characteristic not observed in any of the specimens examined. In addition, the coppery setulae on the sternites of the paratype are denser than in the specimens examined. Scione rufescens and S. fulvosericea have been considered closely related species for five decades. Fairchild (1956) was the first to synonymize S. fulvosericea under S. rufescens. Moucha (1976) and Wilkerson (1979) continued to treat S. fulvosericea as a synonym, but Fairchild & Burger (1994) once again treated it as a valid species. The authors mentioned that specimens of S. fulvosericea collected west of the Andes are consistently different from those of S. rufescens collected east of the Andes. They further noted that John E. Chainey (NHMUK) compared the specimens with the London types and confirmed that they were different species. Given the above, and taking into account that each of these species currently has its own synonym Rhinotriclista generosa Enderlein, 1925, for S. rufescens and S. aureopygia Fairchild, 1942, for S. fulvosericea and that we examined only a small amount of material, we decided to retain both species as valid. However, examination of the type photographs and those of the synonyms of both species revealed no marked differences between them.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206C380B4BC6FD0B7A21FA1B.taxon	materials_examined	Type locality. “ Venezuela ”.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206C380B4BC6FD0B7A21FA1B.taxon	description	Original Description. Gleicht huancabambae bis auf folgende Punkte. Ocellen hell. Fühler hellrotgelb, 1. und 2. Glied dunkler; goldrot behaart. Kopf heller rotbraun. Bart mehr rotgelb. Thorax und Schildchen ohne Zeichnung, vorn goldrot, hinten rotbraun behaart. Brustseiten rotbraun, goldrot behaart, p satt rotgelb, inklusive Tarsen; goldrot behaart. Flügel braun tingiert, ohne jede Säumung der Queradern. Schüppchen und Schwinger bräunlich. Hinterleib plump, breit. 1. Tergit heller rotbraun, 2. – 5. düster rotbraun, 6. – 7. wieder heller. Behaarung am 1., 6. – 7. goldrot, am 2. – 5. schwarz. Bauch rotbraun, dicht goldrot behaart.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206C380B4BC6FD0B7A21FA1B.taxon	distribution	Distribution. Venezuela, Colombia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206C380B4BC6FD0B7A21FA1B.taxon	materials_examined	Holotype ♀. Destroyed (Kröber Collection).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206C380B4BC6FD0B7A21FA1B.taxon	discussion	Comments. According to the original description, this species is similar to S. huancabambae, which was described in the same year by the same author and which type specimen is lost. It differs from S. huancabambae because this species has a thorax with two broad black sublateral stripes and hyaline wings, whereas S. rufipes has thorax without stripes and smoky brown wings. Bequaert & Renjifo-Salcedo (1946) recognize this species for Colombia, however, we did not have access to this material and their work does not provide diagnosis, redescription or illustration, so it was not possible to identify S. rufipes, which is why the species was not included in the identification key.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206C380B4BC6F9E27A83F80C.taxon	description	(Fig. 72 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206C380B4BC6F9E27A83F80C.taxon	materials_examined	Type locality. Colombia, Valle, “ Rio Bravo-Playa Rica ”, 15 km below the Calima Lake dam.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206C380B4BC6F9E27A83F80C.taxon	diagnosis	Diagnosis. Body predominantly brown with bright yellow setulae on tergites 2 to 7. Palpus and antennae brown. Flagellum serrated. Thorax brown without stripes. Wing slightly smoky without spots.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206C380B4BC6F9E27A83F80C.taxon	distribution	Distribution. Western Colombia.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206C380B4BC6F9E27A83F80C.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the FSCA (examined through photographs).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33206C380B4BC6F9E27A83F80C.taxon	discussion	Comments. This species is easily identifiable by its serrated flagellum, a characteristic not observed in any other species of the genus.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207238154BC6FF4A7F55F962.taxon	description	(Fig. 73 A – G)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207238154BC6FF4A7F55F962.taxon	materials_examined	Type locality. Ecuador, Morona-Santiago, Cordillera de Cutucú, 6 km east of Macas.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207238154BC6FF4A7F55F962.taxon	diagnosis	Diagnosis. Body yellowish-brown. Frons covered with yellow pruinosity. Antenna orange. Palpus orange, long and quite sharp. Scutum brown with white longitudinal stripes, the sublateral stripes wide and quite evident and the median stripe and barely visible. Wing hyaline with brown veins. Femora yellowish-brown, tibiae and tarsi brown. Abdominal tergites 1 and 2 yellow, tergites 3 to 7 brown.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207238154BC6FF4A7F55F962.taxon	etymology	Etymology. The specific epithet “ smithi ” is a tribute to Stephen M. Smith (the teacher Steve), a researcher at the University of Waterloo, for his contributions to Entomology.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207238154BC6FF4A7F55F962.taxon	description	Description, Holotype ♀. Body length: 11.7 mm, wing length: 12.2 mm. Head. Frons slightly divergent below, divergence index 0.9, frontal index 2.1. Vertex dark brown with long dense black setulae, prominent ocellar triangle, yellowish-brown ocelli. Frons and subcallus covered with yellow pruinosity. Frons with sparse black setulae and a slightly marked central brown patch of tegument. Parafacial brown with yellow pruinosity and some black setulae. Gena covered with yellow pruinosity and long dense yellow setulae. Clypeus brown with sparse yellow pruinosity and some short sparse black setulae, clypeal index 0.9. Antenna orange. Scape and pedicel with yellow pruinosity and long dense black setulae. Palpus orange, long and quite tapered; first segment slightly darker and with long, dense yellow setulae on the proximal margin and long, dense black setulae on the distal margin; second segment short, sparse black. Proboscis shiny, dark brown, about 6 x the height of the frons. Thorax. Scutum and scutellum brown with sparse black setulae. Scutum with white longitudinal stripes. The sublateral stripes wide and quite evident, connected at the posterior margin of the scutum. The median stripe is narrow and barely visible. The sublateral stripes connected to the lateral stripes by a white stripe at the transverse suture. Notopleuron with tuft of long dense yellow and black setulae. Anepisternum and katepisternum with tuft of long dense black setulae. Anepimeron with tuft of long dense yellow setulae. Postalar callus with tuft of long dense yellow setulae. Femora yellowish-brown with long dense black setulae. Tibiae and tarsi brown with short, dense black setulae. Wing hyaline with brown veins. Abdomen. Tergites 1 and 2 yellow with sparse black setulae. Tergite 1 with a tuft of white setulae on the central posterior margin. Tergites 3 to 7 brown. Tergite 3 with a posterior band of yellow setulae. Tergites 4 to 7 with dense short black setulae. Tergites 4 and 5 with a posterior band of yellow setulae and a tuft of yellow setulae on the central posterior margin. Male. Unknown.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207238154BC6FF4A7F55F962.taxon	distribution	Distribution. Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207238154BC6FF4A7F55F962.taxon	materials_examined	Type material: Holotype, ♀, “ Ecuador, Morona-Santiago, Cord. de Cutucu 6 K. e. of Macas 1100 m, 29. xii. 1981, M. Cooper / M. Cooper Neotropical Tabanidae BMNH (E) 2004 - 182 / NHMUK 014428017 / Scione det. J. E. Chainey, 2004 ” (NHMUK).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207238154BC6FF4A7F55F962.taxon	discussion	Comments. Scione smithi sp. nov. is similar to S. longirostris, S. flavescens, and S. longibeccus sp. nov. However, S. smithi sp. nov. differs from S. longirostris in having darker body, less wide thoracic stripes and the absence of the well-marked rectangular yellowish patch on the posterior margin of the scutum, present in S. longirostris. S. smithi sp. nov. differs from S. flavescens in having slightly smoky wing without spots around the crossveins and brown legs, whereas S. flavescens has smoky wing with brown spots on the crossveins and light (yellowish-brown) legs. S. smithi sp. nov. differs from S. longibeccus sp. nov. in having frons and subcallus covered with yellow pruinosity, dark legs and more marked thoracic stripes, whereas S. longibeccus sp. nov. has a brown frons and subcallus with orange pruinosity, yellow legs and the thoracic stripes are less evident (Fig. 74).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207038114BC6FA2E7BB6FC77.taxon	description	(Fig. 75 A – E)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207038114BC6FA2E7BB6FC77.taxon	materials_examined	Type locality. Ecuador, Santa Inés.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207038114BC6FA2E7BB6FC77.taxon	diagnosis	Diagnosis. Body orange brown. Frons with well-marked triangular dark brown patch. Thorax with quite evident white longitudinal stripes. Wing hyaline without spot on crossveins, veins orange.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207038114BC6FA2E7BB6FC77.taxon	description	Redescription. Female. Body length: 10.6 – 11.8 mm (X = 11.2 mm, N = 3), wing length: 11.2 – 11.5 mm (X = 11.4 mm, N = 3). Head. Frons slightly divergent below, divergence index 0.8 – 0.9 (X = 0.8, N = 3), frontal index 1.6 – 1.8 (X = 1.7, N = 3). Frons with dark brown triangular patch, extending to the apex and almost touching the eyes at its base, slightly narrower towards the apex. Apex dark brown with long black setulae, prominent ocellar triangle, orange brown ocelli. Subcallus covered with whitish pruinosity. Subcallus orange brown near the insertion of antennae. Parafacial brown covered with whitish pruinosity. Gena covered with whitish pruinosity and dense long white setulae. Clypeus brown with whitish pruinosity and sparse black and yellowish setulae, clypeal index 0.6 – 0.8 (X = 0.7, N = 3). Scape and pedicel brown with sparse black and yellowish setulae. Flagellum orange-brown, last flagellomeres dark. First segment of palpus orange brown with long black and yellowish setulae; second segment orange with a few black setulae. Proboscis dark brown shiny, about 3 x the height of the frons. Thorax. Scutum orange brown with black setulae and some sparse orange setulae and quite evident white longitudinal stripes. Sublateral and lateral stripes connected by a white stripe at the transverse suture. Median stripe less evident and extending to the posterior margin of the scutum. Scutellum orange brown with whitish pruinosity and dense long yellowish setulae. Notopleuron with tuft of dense black and yellowish setulae. Anepisternum and katepisternum with tufts of black setulae. Anepimeron with black and white setulae. Postalar callus with tuft of long dense whitish-yellow setulae. Pterostigma brownish. Legs dark brown with long dense black setulae on coxae, trochanters and femora and short dense black setulae on tibiae and tarsi. Wing hyaline with brown veins. Abdomen. Tergites orangish-brown with dense black setulae. Tergites 2 and 3 with posterior band of whitish pruinosity and some white setulae. Tergites 4 to 6 with dense white setulae. Lateral margin of tergites 2 to 6 with dense white setulae. Sternites with black and white setulae and a narrow posterior band of grayish pruinosity.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207038114BC6FA2E7BB6FC77.taxon	distribution	Distribution. Ecuador, Peru.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207038114BC6FA2E7BB6FC77.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the MNHUB (examined through photographs). Additional material examined. 1 ♀, “ Ecuador, Azuay, Guarumales, (2462 m), (2 º 36.64 ’’ S 78 º 31.35 ’’), 29 Oct — 1 Nov 2007, J. Buestan / CJB 9089 ” (MECN); 1 ♀, “ idem / CJB 9090 ” (MECN); 1 ♀, “ idem / CJB 9088 ” (MECN).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207038114BC6FA2E7BB6FC77.taxon	discussion	Comments. This species resembles S. claripennis, from which it differs mainly in the structure of the patch on the frons. In S. claripennis the patch is a central triangle that is not very wide and does not extend to the vertex. In S. strigata the patch is more marked and extends to the vertex. Furthermore, the thoracic stripes are wider in S. strigata.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207638134BC6FC0E7B8FFCE2.taxon	description	(Fig. 76 A – H)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207638134BC6FC0E7B8FFCE2.taxon	materials_examined	Type locality. Ecuador, Tinajillas.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207638134BC6FC0E7B8FFCE2.taxon	diagnosis	Diagnosis. Body yellowish-brown. Thorax dark brown with two whitish sublateral stripes. Wing hyaline with brown veins and faint brown spots on crossveins. Femora brown, tibiae and tarsi yellow. Abdominal tergites 1 to 3 predominantly yellow, tergites 4 to 7 brown.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207638134BC6FC0E7B8FFCE2.taxon	etymology	Etymology. The specific epithet is a reference to the type locality, the Tinajillas Mountains in Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207638134BC6FC0E7B8FFCE2.taxon	description	Description, Holotype ♀. Body length: 9.0 mm, wing length: 9.7 mm. Head. Frons divergent below, divergence index 0.8, frontal index 2.2. Vertex dark brown with long dense black setulae, ocellar triangle prominent, ocelli yellowish-brown. Frons and subcallus brown with white pruinosity, frons with long dense black setulae. Parafacial brown with white pruinosity and sparse long black setulae. Gena with white pruinosity and long dense yellowish-white setulae. Clypeus brown with sparsely dense whitish pruinosity and long dense black setulae, clypeal index 0.7. Scape and pedicel brown with whitish pruinosity and long dense black setulae. Flagellum yellow, last flagellomeres slightly darkened. Palpus brown; first segment with long dense whitish-yellow setulae; second segment orange on the dorsal margin and with short sparse black setulae. Proboscis shiny black, about 2.5 x the height of the frons. Thorax. Scutum dark brown with dense short black and yellow setulae and two whitish sublateral stripes. From each of the sublateral stripes there is a narrower whitish stripe in the region of the transverse suture. Scutellum dark brown with dense long black setulae on the posterior margin. Notopleuron with tuft of dense long white setulae. Anepisternum, katepisternum and anepimeron with tufts of yellow setulae. Postalar callus with tuft of long dense white setulae. Femora brown with long dense black and yellow setulae. Tibiae and tarsi yellow with short dense black and yellow setulae. Wing hyaline with brown veins and faint brown spots on crossveins. Abdomen. Tergite 1 with brown anterior margin, yellow posterior margin. Tergites 2 and 3 yellow. Tergite 3 with short dense black setulae. Tergites 2 and 3 with a central brown integument patch. Tergites 4 to 7 brown with short dense black setulae. All tergites with posterior band of yellowish setulae and with tuft of whitish setulae on the central median posterior margin. Sternites with short dense yellowish setulae. Sternites 1 to 3 yellow, remaining sternites brown. Male. Unknown. Paratype variations. Body length: 9.2 – 10.3 mm (X = 9.7 mm, N = 9), wing length: 10 – 11 mm (X = 10.6 mm, N = 9), frontal index 1.6 – 2.0 (X = 1.8, N = 9), divergence index 0.7 – 0.9 (X = 0.8, N = 9), clypeal index 0.7 – 0.8 (X = 0.7, N = 9).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207638134BC6FC0E7B8FFCE2.taxon	distribution	Distribution. Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207638134BC6FC0E7B8FFCE2.taxon	materials_examined	Type material: Holotype, ♀, “ Tinajillas, 16. xi. 84 / CJB- 2616 ” (MECN). Paratypes: 1 ♀ (in good condition, but both wings have small broken areas), “ idem, except CJB- 2612 / 16 ” (MECN); 1 ♀ (in good condition, but part of the right hind tibia and left hind tarsus are missing; lower margin of left wing with a broken piece), “ idem, except CJB- 2610 ” (MECN); 1 ♀, (in good condition, but right wing with proximal and distal margins damaged; left wing with proximal margin damaged), “ idem, except CJB- 2615 ” (MECN); 1 ♀ (in good condition, but left hind tibia is missing), “ idem, except CJB- 2614 ” (MECN); 1 ♀ (in good condition), “ idem, except CJB- 2613 ” (MECN); 1 ♀ (in good condition, but right pedicel and flagellum, part of the right mid tibia, are part of hind tibia, are missing), “ idem, except CJB- 2609 ” (MECN); 1 ♀ (in good condition, but abdominal tergites 2 and 3 and anterior margin of right wing, damaged), “ idem, except CJB- 2611 ” (MECN); 1 ♀ (in good condition, but proximal margin of both wings with small broken area), “ idem, except CJB- 2608 ” (INPA); 1 ♀, (in good condition, but left hind leg with last tarsomeres missing, right wing with the distal lower margin damaged, and a crushed abdomen), “ idem, except CJB- 2618 ” (MECN).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207638134BC6FC0E7B8FFCE2.taxon	discussion	Comments. In the holotype and some paratypes there is a large amount of pollen, mainly on the head and legs.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332074381D4BC6FCBB785CF829.taxon	description	(Figs 77 A – F, 78 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332074381D4BC6FCBB785CF829.taxon	materials_examined	Type locality. Peru, Ancash, cerro Huaylas.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332074381D4BC6FCBB785CF829.taxon	diagnosis	Diagnosis. Body yellowish-brown. Scutum brown with slightly evident whitish longitudinal stripes. Wing hyaline with brown veins and faint brown spots on crossveins. Abdominal tergites 1 and 2 predominantly yellow, other tergites dark brown.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332074381D4BC6FCBB785CF829.taxon	etymology	Etymology. The specific epithet “ walkeri ” is a tribute to Francis Walker, author of the genus Scione, for his contributions to the knowledge of tabanids.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332074381D4BC6FCBB785CF829.taxon	description	Description, Holotype ♀: body length: 10.6 mm, wing length: 11.6 mm. Head. Frons divergent below, divergence index 0.7, frontal index 1.7. Vertex dark brown with long dense black setulae, ocellar triangle not very prominent, ocelli brown. Frons and subcallus brown with grey pruinosity, frons with long dense black setulae. Parafacial brown with grey pruinosity and sparse black setulae. Gena with grey pruinosity and long dense yellow setulae. Clypeus brown with gray pruinosity and short dense black setulae, clypeal index 0.9. Scape and pedicel brown with gray pruinosity and long dense black setulae. Flagellum orange, last flagellomeres black. Palpus brown, long and lanceolate; first segment with long, dense yellow setulae; second segment with orange dorsal margin and some sparse black setulae. Proboscis shiny black, about 5 x the height of the frons. Thorax. Scutum brown with sparse black and yellow setulae and slightly evident whitish longitudinal stripes. A narrower white stripe extends from each of the sublateral stripes at the transverse suture. Scutellum brown with a few yellow setulae and with dense long black setulae on the dorsal margin. Notopleuron with a tuft of dense long black and yellow setulae. Anepisternum with dense long yellow setulae and some black setulae. Katepisternum and anepimeron with dense long yellow setulae. Postalar callus with a tuft of dense long yellow setulae and some black setulae. Femora with long dense black setulae, proximal half brown and distal half yellow. Tibiae and tarsi yellow with short dense black setulae. Wing hyaline with brown veins and faint brown spots on crossveins. Small appendix at the fork of vein R 4 + 5. Abdomen. Tergite 1 with brown anterior margin and yellow posterior margin and with a posterior band of yellow setulae. Tergites 2 yellow with dense black setulae, with a central brown patch and with posterior band of yellow setulae. Tergites 3 to 7 dark brown with dense black setulae. Tergite 3 with a tuft of yellow setulae on central posterior margin. Tergites 4 to 6 with a posterior band of yellow setulae. Lateral margin of tergites with dense and long yellow setulae. Sternites brown with dense and long black setulae. Sternites 3 to 5 with posterior band of yellow integument and setulae. Male (Fig. 78). Like the female, except for the holoptic condition, straight palpus and longer, denser body setulae. Paratype variations. Body length: 9.6 – 12 mm (X = 11.1 mm, N = 8), wing length 10.7 – 11.9 mm (X = 10.1 mm, N = 8), frontal index 1.6 – 1.7, divergence index 0.7 (in both females), clypeal index 0.6 – 0.9 (X = 0.6, N = 9). The last flagellomeres lighter in some specimens. The appendix at the fork of vein R 4 + 5 is absent in specimens from NHMUK and in other specimens the appendix is barely visible.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332074381D4BC6FCBB785CF829.taxon	distribution	Distribution. Peru (Ancash, Lima).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332074381D4BC6FCBB785CF829.taxon	materials_examined	Type material: Holotype, ♀, “ Peru: Dpto. Ancash, Huaylas (Cerro), 3: v: 84 (3,600 mt.), Pedro Hocking ” (UNMSM). Paratypes: 2 ♀ (poorly preserved, part of the setulae and coloration lost), “ idem, (UNMSM) ”; 3 ♂ (in good condition), “ idem, (UNMSM) ”; 1 ♂ (in good condition, but left hind leg with missing femur), “ PE: Ancash, Quitacocha, 8 - 5 - 79 (11,000 ft), P. Hocking ” (UNMSM); 1 ♂ (in good condition, but left pedicel and flagellum and left posterior tibia and tarsus are missing), “ Peru, Lima, Matucana, 2.600 m, 6. v. 1984, M. Cooper / M. Cooper Neotropical Tabanidae BMNH (E) 2004 - 182 / NHMUK 014428041 ” (NHMUK); 1 ♂ (in good condition, but left wing is missing), “ idem, except NHMUK 014428042 ” (NHMUK).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F332074381D4BC6FCBB785CF829.taxon	discussion	Comments. Scione walkeri sp. nov. is similar to S. peruviana sp. nov. It differs in that S. walkeri sp. nov. has tergites 3 to 7 dark brown with dense black setulae, tergite 3 with tuft of yellow setulae on the central posterior margin, and tergites 4 to 6 with posterior band of yellow setulae. In contrast, S. peruviana sp. nov. has tergites 3 to 7 with brown anterior margin and yellowish posterior margin, tergites 3 and 4 with short sparse black setulae and tergites 5 to 7 with short dense yellow and black setulae. Both species resemble S. cooperi, differing in that S. cooperi has black abdomen with tapered posterior end, tergites 1 and 2 orange with posterior band of yellowish integument and setulae, tergites 3 to 7 black, tergites 3 and 4 each with small central tuft of yellowish setulae, and tergite 5 with posterior band of yellowish setula (Fig. 79).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207938194BC6FA037A41F9DB.taxon	description	(Figs. 80 A – G, 81 A – G)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207938194BC6FA037A41F9DB.taxon	materials_examined	Type locality. Ecuador, Tinajillas.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207938194BC6FA037A41F9DB.taxon	diagnosis	Diagnosis. Body yellowish-brown. Thorax brown with yellowish longitudinal stripes. Wing smoky on the anterior region, with yellow spots in the middle of cells r 1 and r 2 + 3 and hyaline on the posterior region. Well-marked brown spots on crossveins. Legs yellow.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207938194BC6FA037A41F9DB.taxon	etymology	Etymology. The specific epithet “ wilkersoni ” is a tribute to Richard Charles Wilkerson, for his contributions to the knowledge of tabanids.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207938194BC6FA037A41F9DB.taxon	description	Description, Holotype ♀. Body length: 9.6 mm, wing length: 10.5 mm. Head. Frons slightly divergent below, divergence index 0.8, frontal index 2.0. Vertex dark brown with long dense black setulae, ocellar triangle slightly prominent, ocelli brown. Frons and subcallus brown with yellowish pruinosity, frons with long dense black setulae. Parafacial brown with yellowish pruinosity and long sparse black setulae. Gena covered with yellow pruinosity and dense long yellow setulae. Clypeus brown with slightly dense yellowish pruinosity and dense long black setulae, clypeal index 0.7. Antenna orange, scape and pedicel with long dense black setulae. Palpus brown; first segment with long, dense yellow setulae on the proximal margin and long, dense black setulae on the distal margin; second segment orange on the dorsal margin and with long, dense black setulae. Proboscis shiny black, about 2 x the height of the frons. Thorax. Scutum brown with dense black setulae and yellowish longitudinal stripes. The median stripe is wider and less evident. The sublateral stripes quite evident and extend to the posterior margin of the scutum, where they connect. The sublateral and lateral stripes connected by a less narrow whitish stripe in the region of the transverse suture. Scutellum brown with long dense black setulae in the central region and on the posterior margin. Notopleuron with tuft of long dense orange and black setulae. Anepisternum, katepisternum and anepimeron with tufts of long dense black and orange setulae. Postalar callus with tuft of long dense yellow and orange setulae. Legs yellow. Femora with long dense black setulae. Tibiae and tarsi slightly lighter than the femora and with short dense black and yellow setulae. Wing smoky on the anterior region, with yellow spots in the middle of cells r 1 and r 2 + 3 (“ window ”) and hyaline on the posterior region. Well-marked brown spots on crossveins. Abdomen. Tergites 1 to 3 yellow. Tergites 2 with short dense black setulae. Tergites 2 to 7 with a posterior band of yellow setulae and with a tuft of yellow setulae on the central posterior margin. Tergites 4 to 7 darkened and with short dense yellow setulae. Sternites with short sparse black and yellow setulae. Sternites 1 to 3 yellow. Sternites 4 to 7 darkened. Male (Fig. 81). Like the female, except for the holoptic condition, porrect palpus and longer and denser body setulae. In addition, the clypeus of males is less developed than in females. Paratype variations. Body length: 9.0 – 10 mm (X = 9.6 mm, N = 9), wing length 9.5 – 10.8 mm (X = 10.1 mm, N = 9), frontal index (X = 1.9, N = 6), divergence index (X = 0.7, N = 6), clypeal index (X = 0.7, N = 9). The brown spots on crossveins are less evident in some specimens.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207938194BC6FA037A41F9DB.taxon	distribution	Distribution. Ecuador.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207938194BC6FA037A41F9DB.taxon	materials_examined	Type material: Holotype, ♀, “ Tinajillas 2100 m, Morona-San. Prov. J. Buestán, 26 - x- 85 / CJB- 3306 / 5 a ” (MECN). Paratypes: 1 ♀ (in good condition), “ idem, except CJB- 3304 / 5 a ” (MECN); 1 ♀, (in good condition, but left hind tibia and tarsus are missing), “ idem, except CJB- 3301 / 5 a ” (MECN); 1 ♀ (in good condition, but with a small broken area on the distal margin of the right wing), “ idem, except CJB- 3312 / 5 a ” (MECN); 1 ♀ (in good condition, but right hind tibia and tarsus are missing), “ idem, except CJB- 3307 / 5 a ” (MECN); 1 ♀ (in good condition, but left wing is missing), “ idem, except CJB- 3302 / 5 a ” (INPA); 1 ♂ (specimen photographed; in good condition, but left mid leg is missing), “ idem, except 18. ix. 87 / CJB- 3341 ” (MECN); 1 ♂ (in good condition, but left pedicel and flagellum, and left mid leg, are missing), “ idem, except CJB- 3342 ” (MECN); 1 ♂ (in good condition, but abdomen is crushed, dorsal margin of right wing is broken), “ idem, except 2. xi. 84 / CJB- 3340 ” (MECN); 1 ♀ (in good condition, but with small broken areas on the distal margin of both wings), “ San Vicente 2770 m, Morona-San. Prov. J. Buestán, 11. x. 87 / CJB- 3339 ” (MECN).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207938194BC6FA037A41F9DB.taxon	discussion	Comments. The posterior band of yellow setulae and the tuft of yellow setulae on the central posterior margin of the tergites are missing in some specimens and are barely visible in others. S. wilkersoni sp. nov. is similar to S. tinajillas sp. nov. It differs in that S. wilkersoni sp. nov. has slightly tapered palpus, brown thorax with dense black setulae and five yellowish-white longitudinal stripes and smoky wings with brown veins and well-marked brown spots on the crossveins. In contrast, S. tinajillas sp. nov. has tapered palpus, dark brown thorax with two whitish lateral longitudinal stripes and wing hyaline with yellowish veins and faint brown spots on crossveins (Fig. 82).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207E381A4BC6F9A27F8EFE5B.taxon	description	(Fig. 83 A – F)	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207E381A4BC6F9A27F8EFE5B.taxon	materials_examined	Type locality. Colombia, Valle, Peñas Blancas, 10 km west of Cali.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207E381A4BC6F9A27F8EFE5B.taxon	diagnosis	Diagnosis. Body predominantly brown. Antenna black. Palpus dark brown. Thorax with white longitudinal stripes. The median stripe is barely visible, the sublateral stripes are well marked. Wing smoky with yellowish patch on cells r 1, r 2 + 3, r 5 and on the discal cell and with faint brown spots on crossveins. Femora dark brown, fore and mid tibiae yellowish-brown, hind tibiae dark brown.	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207E381A4BC6F9A27F8EFE5B.taxon	distribution	Distribution. Colombia (Valle).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207E381A4BC6F9A27F8EFE5B.taxon	materials_examined	Type material examined. Holotype, ♀, deposited at the FSCA (examined through photographs).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
30736F33207E381A4BC6F9A27F8EFE5B.taxon	discussion	Comments. Scione youngi is similar to S. brevibeccus and S. cuprea. It differs from S. brevibeccus in that this species has markedly bicolored fore and mid legs, orange brown antenna and smoky brown wing with yellowish veins and very faint brown spots on crossveins. Whereas in S. youngi the fore and mid legs are not markedly bicolored, the antenna is black, and the wing is smoky with yellowish spots on the r 1, r 2 + 3, r 5 cells and on the disc cell. It differs from S. cuprea in that this species has orange antenna and palpus and yellow gena setulae, whereas in S. youngi the antenna is black, the palpus is dark brown and the gena setulae are white (Fig. 84).	en	Oliveira, Lia Pereira, Henriques, Augusto Loureiro, Buestán, Jaime, Krolow, Tiago Kütter (2026): Taxonomic revision of the Neotropical genus Scione Walker, 1850 (Diptera: Tabanidae). Zootaxa 5761 (1): 1-133, DOI: 10.11646/zootaxa.5761.1.1, URL: https://doi.org/10.11646/zootaxa.5761.1.1
