taxonID	type	description	language	source
983CC7D69DF85C61993ACBA3AEFC15CA.taxon	description	Figs 2 A, B, 3, 4	en	Tang, Song-Ming, Yu, Feng-Ming, Karunarathna, Samantha C., Luo, Zong-Long, Niu, Kai-Yang, Li, Rui-Yu, Li, Lin, Su, Xi-Jun, Li, Shu-Hong (2025): Morpho-phylogenetic analyses of two novel edible mushrooms from China and a mini review of Lyophyllum (Agaricales, Lyophyllaceae) cultivation and bioactivities. MycoKeys 112: 307-334, DOI: 10.3897/mycokeys.112.141615
983CC7D69DF85C61993ACBA3AEFC15CA.taxon	etymology	Etymology. The epithet “ edulis ” refers to the edibility of this species; locally it is considered a delicacy.	en	Tang, Song-Ming, Yu, Feng-Ming, Karunarathna, Samantha C., Luo, Zong-Long, Niu, Kai-Yang, Li, Rui-Yu, Li, Lin, Su, Xi-Jun, Li, Shu-Hong (2025): Morpho-phylogenetic analyses of two novel edible mushrooms from China and a mini review of Lyophyllum (Agaricales, Lyophyllaceae) cultivation and bioactivities. MycoKeys 112: 307-334, DOI: 10.3897/mycokeys.112.141615
983CC7D69DF85C61993ACBA3AEFC15CA.taxon	description	Description. Pileus 3 – 8 cm diameter, fleshy, fragile, hemispherical, becoming convex with age, smooth on the surface, dry, dark grayish orange (# 8 a 7971) on the center, soft orange (# e 9 c 7 a 7) with margin, subumbonate of center, inflexed of margin; pileus context thick, 0.2 – 0.3 cm wide, white (# fcfcfc). Lamellae moderately close together, arcuate, subdecurrent to decurrent, broad, white (# fcfcfc), unchanging color when injured, 3 – 4 tiers, 0.4 – 0.5 cm wide, edge even or entire. Stipe 3.7 – 6.9 × 0.8 – 1.4 cm, cylindrical, grayish orange (# d 9 cdc 2) in the upper, soft orange (# e 9 c 7 a 7) gradually downward, smooth; stipe context white (# fcfcfc), solid, unchanging in color when injured. The odor and taste were not distinctive. Basidiospores [84 / 2 / 2] 5.1 – 6.5 (– 8) × 4.6 – 6.6 μm, (Q = 1.0 – 1.2, Qm = 1.11 ± 0.05), av. 5.81 ± 0.28 × 5.47 ± 0.38 μm, globose, subglobose to broadly ellipsoid, hyaline, smooth. Basidia 25 – 39 × 8 – 11 μm (N = 20), av. 32.7 ± 5.1 × 9.7 ± 1.0 μm, mostly 4 - spored, rarely 2 - spored, sterigmata long 1.8 – 4.9 μm, sometimes with basal clamp connections, clavate, siderophilous granulations. Subhymenium is composed of moderately thin-walled hyphae, 40 – 55 μm thick, with 2 – 3 layers of ovoid, fusiform to narrowly cylindrical hyphae, and 6 – 8 × 3 – 5 μm. Hymenophoral trama regular, 120 – 150 μm wide, consisting of thin and hyaline hyphae, some with clamp connections, narrowly cylindrical hyphal elements, 6 – 12 μm wide. Cheilocystidia were 21 – 24 × 4 – 7 μm in size and av. 22.9 ± 1.3 × 6.4 ± 0.7 μm, narrowly cylindrical or narrowly clavate, thin-walled, and rarely mucronate or rostrate on the apex. Pleurocystidia 24 – 28 × 4 – 6 μm, av. 26.3 ± 1.6 × 5.3 ± 0.6 μm, narrowly cylindrical or narrowly clavate, thin-walled. Pileipellis colorless and hyaline in 5 % KOH solution, parallel, thin-walled, almost cylindrical to subcylindrical, filamentous hyphae 4 – 6 μm wide. Stipitipellis composed of appressed, parallel, thin-walled, hyphae 2 – 7 µm wide. Clamp connections are present at some septa in the pileipellis, lamellae, and stipitipellis.	en	Tang, Song-Ming, Yu, Feng-Ming, Karunarathna, Samantha C., Luo, Zong-Long, Niu, Kai-Yang, Li, Rui-Yu, Li, Lin, Su, Xi-Jun, Li, Shu-Hong (2025): Morpho-phylogenetic analyses of two novel edible mushrooms from China and a mini review of Lyophyllum (Agaricales, Lyophyllaceae) cultivation and bioactivities. MycoKeys 112: 307-334, DOI: 10.3897/mycokeys.112.141615
531101CA873851DAB0DF17D6DDC079FC.taxon	description	Figs 2 C – F, 5, 6	en	Tang, Song-Ming, Yu, Feng-Ming, Karunarathna, Samantha C., Luo, Zong-Long, Niu, Kai-Yang, Li, Rui-Yu, Li, Lin, Su, Xi-Jun, Li, Shu-Hong (2025): Morpho-phylogenetic analyses of two novel edible mushrooms from China and a mini review of Lyophyllum (Agaricales, Lyophyllaceae) cultivation and bioactivities. MycoKeys 112: 307-334, DOI: 10.3897/mycokeys.112.141615
531101CA873851DAB0DF17D6DDC079FC.taxon	description	Description. Pileus 2.0 – 6.0 cm diameter, fleshy, fragile, hemispherical, becoming convex with age, smooth on the surface, dry, grayish orange (# e 4 dfdb) on the center, soft orange (# dbcca 9) with margin, slightly depressed to papilla of center, deflexed to inflexed of margin; pileus context thick, 0.2 – 0.3 cm wide, white (# fcfcfc). Lamellae moderately close together, arcuate, subdecurrent to decurrent, broad, white (# fcfcfc), unchanging color when injured, 2 – 3 tiers, 0.2 – 0.3 cm wide, edge even or entire. Stipe 4 – 7 × 0.9 – 1.1 cm, wide bulbous at the base, smooth; stipe context white (# fcfcfc), 1.2 – 3.0 cm wide, bulbous at the base, smooth; stipe context white (# fcfcfc), unchanging in color when injured. The odor and taste were not distinctive. Basidiospores [73 / 2 / 2] 4.5 – 6.6 × 4.0 – 5.9 μm, (Q = 1.0 – 1.3, Qm = 1.11 ± 0.10), av. 5.38 ± 0.59 × 4.89 ± 0.61 μm, globose, subglobose to broadly ellipsoid, hyaline, smooth. Basidia 19 – 28 (– 35) × 10 – 15 μm (N = 20), av. 25.6 ± 4.1 × 12.1 ± 1.64 μm, mostly 2 - spored, rarely 4 - spored, sterigmata long 2.9 – 4.1 μm, sometimes with basal clamp connections, clavate, siderophilous granulations. Subhymenium is composed of moderately thin-walled hyphae, 15 – 20 μm thick, with 1 – 2 layers of ovoid, fusiform to narrowly cylindrical hyphae, 3 – 7 × 2 – 4 μm. Hymenophoral trama regular, 110 – 160 μm wide, consisting of thin and hyaline hyphae, some with clamp connections, narrowly cylindrical hyphal elements 2 – 5 μm wide. Cheilocystidia were 10 – 15 × 4 – 6 μm, av. 12.2 ± 1.8 × 5.0 ± 0.4 μm, narrowly cylindrical or narrowly clavate, rarely apex flexed, mostly narrowing with apex, thin-walled. Pleurocystidia were 12 – 18 × 4 – 6 μm in size and av. 14.8 ± 4.1 × 4.4 ± 1.1 μm, narrowly cylindrical or narrowly clavate, rarely apex flexed, mostly narrowing with apex, thin-walled. Pileipellis is an interwoven trichodermium composed of almost hyaline interwoven filamentous hyphae, terminal cells 2 – 5 μm wide, almost cylindrical to subcylindrical, occasional hyphal tips flexuous and sometimes inflate, and some with clamp connections. Stipitipellis composed of appressed, parallel, thin-walled, 2 – 4 µm wide, fusiform, thin-walled. Clamp connections are present at some septa in the pileipellis, lamellae, and stipitipellis.	en	Tang, Song-Ming, Yu, Feng-Ming, Karunarathna, Samantha C., Luo, Zong-Long, Niu, Kai-Yang, Li, Rui-Yu, Li, Lin, Su, Xi-Jun, Li, Shu-Hong (2025): Morpho-phylogenetic analyses of two novel edible mushrooms from China and a mini review of Lyophyllum (Agaricales, Lyophyllaceae) cultivation and bioactivities. MycoKeys 112: 307-334, DOI: 10.3897/mycokeys.112.141615
18303D58D9745E26A17D54C6D4C84675.taxon	description	Figs 7 C, D, 10, 11	en	Tang, Song-Ming, Yu, Feng-Ming, Karunarathna, Samantha C., Luo, Zong-Long, Niu, Kai-Yang, Li, Rui-Yu, Li, Lin, Su, Xi-Jun, Li, Shu-Hong (2025): Morpho-phylogenetic analyses of two novel edible mushrooms from China and a mini review of Lyophyllum (Agaricales, Lyophyllaceae) cultivation and bioactivities. MycoKeys 112: 307-334, DOI: 10.3897/mycokeys.112.141615
18303D58D9745E26A17D54C6D4C84675.taxon	description	Description. Pileus 2.0 – 3.0 cm diameter, fleshy, fragile, hemispherical, becoming convex with age, abundant black floccus on the surface, dry, dark orange (# 756450), slightly depressed of center, deflexed to inflexed of margin; pileus context thick, 0.5 – 0.7 cm wide, white (# fcfcfc). Lamellae moderately close together, arcuate, subdecurrent to decurrent, broad, white (# fcfcfc), unchanging color when injured, 3 – 4 tiers, 0.3 – 0.4 cm wide, edge even or entire. Stipe 3.0 – 5.1 × 1.0 – 1.4 cm, cylindrical to clavate, grayish yellow (# 89877 b) on the surface, tapering upwards, smooth; stipe context white (# fcfcfc), unchanging in color when injured. The odor and taste were not distinctive. Basidiospores [75 / 2 / 2] 5.4 – 7.3 × 4.6 – 6.6 μm, (Q = 1.0 – 1.3, Qm = 1.10 ± 0.19), av. 6.03 ± 0.38 × 5.55 ± 0.65 μm, globose, subglobose to broadly ellipsoid, smooth. Basidia 32 – 41 × 6 – 9 μm (N = 20), av. 36.2 ± 3.8 × 8.3 ± 1.15 μm, mostly 4 - spored, rarely 2 - spored, sterigmata long 3.1 – 4.5 μm, sometimes with basal clamp connections, clavate, siderophilous granulations. Subhymenium is composed of moderately thin-walled hyphae, 15 – 30 μm thick, with 2 – 3 layers of ovoid, fusiform to narrowly cylindrical hyphae, 5 – 8 × 3 – 4 μm. Hymenophoral trama regular, 120 – 180 μm wide, consisting of thin and hyaline hyphae, some with clamp connections, narrowly cylindrical hyphal elements 2 – 4 μm wide. Cheilocystidia 15 – 22 (– 26) × 3 – 5 μm, av. 20.6 ± 4.4 × 4.7 ± 1.1 μm, narrowly cylindrical or narrowly clavate, thin-walled. Pleurocystidia were 16 – 20 × 3 – 5 μm in size and av. 18.6 ± 3.7 × 3.7 ± 0.4 μm, narrowly cylindrical or narrowly clavate, thin-walled. Pileipellis is an interwoven trichodermium composed of almost hyaline interwoven filamentous hyphae, terminal cells 2 – 4 μm wide, almost cylindrical to subcylindrical, occasional hyphal tips flexuous and sometimes inflated, and some with clamp connections. Stipitipellis composed of appressed, parallel, thin-walled, 3 – 6 μm wide. Clamp connections are present at some septa in the pileipellis, lamellae, and stipitipellis.	en	Tang, Song-Ming, Yu, Feng-Ming, Karunarathna, Samantha C., Luo, Zong-Long, Niu, Kai-Yang, Li, Rui-Yu, Li, Lin, Su, Xi-Jun, Li, Shu-Hong (2025): Morpho-phylogenetic analyses of two novel edible mushrooms from China and a mini review of Lyophyllum (Agaricales, Lyophyllaceae) cultivation and bioactivities. MycoKeys 112: 307-334, DOI: 10.3897/mycokeys.112.141615
0B170E826D405461909A7F97218AE929.taxon	description	Figs 7 A, B, 8, 9	en	Tang, Song-Ming, Yu, Feng-Ming, Karunarathna, Samantha C., Luo, Zong-Long, Niu, Kai-Yang, Li, Rui-Yu, Li, Lin, Su, Xi-Jun, Li, Shu-Hong (2025): Morpho-phylogenetic analyses of two novel edible mushrooms from China and a mini review of Lyophyllum (Agaricales, Lyophyllaceae) cultivation and bioactivities. MycoKeys 112: 307-334, DOI: 10.3897/mycokeys.112.141615
0B170E826D405461909A7F97218AE929.taxon	etymology	Etymology. The epithet “ sinense ” refers to the country “ China, ” where this fungus was first discovered.	en	Tang, Song-Ming, Yu, Feng-Ming, Karunarathna, Samantha C., Luo, Zong-Long, Niu, Kai-Yang, Li, Rui-Yu, Li, Lin, Su, Xi-Jun, Li, Shu-Hong (2025): Morpho-phylogenetic analyses of two novel edible mushrooms from China and a mini review of Lyophyllum (Agaricales, Lyophyllaceae) cultivation and bioactivities. MycoKeys 112: 307-334, DOI: 10.3897/mycokeys.112.141615
0B170E826D405461909A7F97218AE929.taxon	description	Description. Pileus 2.0 – 3.0 cm diameter, fleshy, fragile, hemispherical, becoming convex with age, abundant black floccus on the surface, dry, dark grayish orange (# a 4 a 3 a 0) on the center, grayish yellow (# cac 4 b 0) with margin, slightly depressed of center, involute of margin; pileus context thick, 0.3 – 0.5 cm wide, white (# fcfcfc). Lamellae moderately close together, arcuate, subdecurrent to decurrent, broad, white (# fcfcfc), grey dark orange (# a 4 a 3 a 0) when injured, 3 – 4 tiers, 0.3 – 0.4 cm wide, edge even or entire. Stipe 3.0 – 4.0 × 0.9 – 1.8 cm, cylindrical to clavate, dark grayish orange (# a 4 a 3 a 0) points and lines on the surface, bulbous at the base, smooth; stipe context white (# fcfcfc), changing to grayish orange (# c 2 bbab) when injured. The odor and taste were not distinctive. Basidiospores [68 / 2 / 2] 6.1 – 8.6 × 5.5 – 7.1 μm, (Q = 1.0 – 1.3, Qm = 1.21 ± 0.12), av. 7.28 ± 0.68 × 6.07 ± 0.62 μm, quadrangular to very broadly fusiform, hyaline, smooth. Basidia 28 – 41 × 8 – 10 μm (N = 20), av. 34.6 ± 4.0 × 9.5 ± 0.53 μm, mostly 4 - spored, rarely 2 - spored, sterigmata long 2.2 – 3.9 μm, sometimes with basal clamp connections, clavate, siderophilous granulations. Subhymenium is composed of moderately thin-walled hyphae, 40 – 60 μm thick, with 2 – 3 layers of ovoid, fusiform to narrowly cylindrical hyphae, 5 – 7 × 2 – 4 μm. Hymenophoral trama regular, 130 – 180 μm wide, consisting of thin and hyaline hyphae, some with clamp connections, narrowly cylindrical hyphal elements, 4 – 7 μm wide. Cheilocystidia were 14 – 23 × 3 – 5 μm, av. 17.6 ± 2.4 × 4.1 ± 0.7 μm, narrowly cylindrical or narrowly clavate, thin-walled. Pleurocystidia were 10 – 25 × 3 – 6 μm in size and av. 17.2 ± 3.2 × 4.3 ± 1.1 μm, narrowly cylindrical or narrowly clavate, thin-walled. Pileipellis colorless and hyaline in 5 % KOH solution, parallel, thin-walled, almost cylindrical to subcylindrical, filamentous hyphae 2 – 3 μm wide. Stipitipellis composed of appressed, parallel, thin-walled, hyphae 2 – 4 µm wide. Clamp connections are present at some septa in the pileipellis, lamellae, and stipitipellis.	en	Tang, Song-Ming, Yu, Feng-Ming, Karunarathna, Samantha C., Luo, Zong-Long, Niu, Kai-Yang, Li, Rui-Yu, Li, Lin, Su, Xi-Jun, Li, Shu-Hong (2025): Morpho-phylogenetic analyses of two novel edible mushrooms from China and a mini review of Lyophyllum (Agaricales, Lyophyllaceae) cultivation and bioactivities. MycoKeys 112: 307-334, DOI: 10.3897/mycokeys.112.141615
