taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
5AD52DCA3E58563CBAAD1D21F3756CF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582158	https://doi.org/10.3897/zookeys.1276.173213.figure2	Figure 2. Distributional maps of Leodamas species. Stars represent the type localities of the species.	Figure 2. Distributional maps of Leodamas species. Stars represent the type localities of the species.	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
5AD52DCA3E58563CBAAD1D21F3756CF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582159	https://doi.org/10.3897/zookeys.1276.173213.figure3	Figure 3. Leodamas chevalieri, type specimens, light microscopy, stereomicroscope; A – D. Lectotype, MNHN - IA-TYPE 0728; E. Paralectotype. A. General view from left side; B. Anterior part from left side, prostomium is not seen; C. Thorax, dorsal view; D. Middle thorax, lateral view; E. Abdomen, dorsal view. Scale bars: 1 mm (A, C); 200 µm (B, D); 500 µm (E).	Figure 3. Leodamas chevalieri, type specimens, light microscopy, stereomicroscope; A – D. Lectotype, MNHN - IA-TYPE 0728; E. Paralectotype. A. General view from left side; B. Anterior part from left side, prostomium is not seen; C. Thorax, dorsal view; D. Middle thorax, lateral view; E. Abdomen, dorsal view. Scale bars: 1 mm (A, C); 200 µm (B, D); 500 µm (E).	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
5AD52DCA3E58563CBAAD1D21F3756CF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582160	https://doi.org/10.3897/zookeys.1276.173213.figure4	Figure 4. Leodamas chevalieri, paralectotypes. A – C. Light microscopy, compound microscope; D – J. SEM. A, B. Thoracic neurochaetae; C. Abdominal parapodium; D. Neuropodium of chaetiger 2; E. Neuropodium of chaetiger 12; F. Neuropodium of chaetiger 16; G. Neuropodium of chaetiger 20; H. Neuropodium of chaetiger 21; I, J. Abdominal parapodia; K. Forked chaetae from abdominal notopodium. Scale bars: 100 µm (A, C); 50 µm (B, D – H, J); 150 µm (I); 6 µm (K).	Figure 4. Leodamas chevalieri, paralectotypes. A – C. Light microscopy, compound microscope; D – J. SEM. A, B. Thoracic neurochaetae; C. Abdominal parapodium; D. Neuropodium of chaetiger 2; E. Neuropodium of chaetiger 12; F. Neuropodium of chaetiger 16; G. Neuropodium of chaetiger 20; H. Neuropodium of chaetiger 21; I, J. Abdominal parapodia; K. Forked chaetae from abdominal notopodium. Scale bars: 100 µm (A, C); 50 µm (B, D – H, J); 150 µm (I); 6 µm (K).	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
5AD52DCA3E58563CBAAD1D21F3756CF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582161	https://doi.org/10.3897/zookeys.1276.173213.figure5	Figure 5. Leodamas chevalieri, non-type specimen, ZMBN 115302, Senegal, light microscopy. A. General view; B. Parapodium of chaetiger 8; C. Same, neuropodium; D. Abdominal parapodium. Scale bars: 1 mm (A); 100 µm (B – D).	Figure 5. Leodamas chevalieri, non-type specimen, ZMBN 115302, Senegal, light microscopy. A. General view; B. Parapodium of chaetiger 8; C. Same, neuropodium; D. Abdominal parapodium. Scale bars: 1 mm (A); 100 µm (B – D).	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
5AD52DCA3E58563CBAAD1D21F3756CF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582162	https://doi.org/10.3897/zookeys.1276.173213.figure6	Figure 6. Leodamas chevalieri, non-type specimen, ZMBN 115302, Senegal, SEM. A. Anterior end, dorso-lateral view; B. Neuropodium of chaetiger 7; C. Same, upper uncini; D. Neuropodia of chaetigers 8 and 9; E. Neuropodium of chaetiger 16; F. Same, lower uncini; G. Same lower uncinus; H. Abdominal neuropodia. Scale bars: 250 µm (A); 20 µm (B, F); 10 µm (G); 30 µm (C, D); 50 µm (E); 100 µm (H).	Figure 6. Leodamas chevalieri, non-type specimen, ZMBN 115302, Senegal, SEM. A. Anterior end, dorso-lateral view; B. Neuropodium of chaetiger 7; C. Same, upper uncini; D. Neuropodia of chaetigers 8 and 9; E. Neuropodium of chaetiger 16; F. Same, lower uncini; G. Same lower uncinus; H. Abdominal neuropodia. Scale bars: 250 µm (A); 20 µm (B, F); 10 µm (G); 30 µm (C, D); 50 µm (E); 100 µm (H).	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
5AD52DCA3E58563CBAAD1D21F3756CF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582166	https://doi.org/10.3897/zookeys.1276.173213.figure10	Figure 10. Leodamas edgari sp. nov., paratypes, ZMMU WS 8283, parapodia slides. A. Parapodium of chaetiger 7; B. Same, upper neurochaetae; C. Same, lower neurochaetae; D. Parapodium of chaetiger 24 (anterior abdomen); E. Same, neuropodium; F. Posterior abdominal parapodium with slightly curved aciculum; G. Same, neuropodium; H. Posterior abdominal notopodium, two aciculae and forked chaetae seen; I. Posterior abdominal parapodium with strongly hooked aciculum; J. Same, neuropodium. Scale bars: 100 µm (A, D, F, I); 50 µm (B, C, E, G, H, J).	Figure 10. Leodamas edgari sp. nov., paratypes, ZMMU WS 8283, parapodia slides. A. Parapodium of chaetiger 7; B. Same, upper neurochaetae; C. Same, lower neurochaetae; D. Parapodium of chaetiger 24 (anterior abdomen); E. Same, neuropodium; F. Posterior abdominal parapodium with slightly curved aciculum; G. Same, neuropodium; H. Posterior abdominal notopodium, two aciculae and forked chaetae seen; I. Posterior abdominal parapodium with strongly hooked aciculum; J. Same, neuropodium. Scale bars: 100 µm (A, D, F, I); 50 µm (B, C, E, G, H, J).	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
3F464663B94150C6815B68E976461DF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582158	https://doi.org/10.3897/zookeys.1276.173213.figure2	Figure 2. Distributional maps of Leodamas species. Stars represent the type localities of the species.	Figure 2. Distributional maps of Leodamas species. Stars represent the type localities of the species.	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
3F464663B94150C6815B68E976461DF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582163	https://doi.org/10.3897/zookeys.1276.173213.figure7	Figure 7. Leodamas edgari sp. nov., holotype, ZMMU WS 14151, light microscopy. A. Living specimen; B. Anterior end, dorso-lateral view. Scale bars: 5 mm (A); 1 mm (B).	Figure 7. Leodamas edgari sp. nov., holotype, ZMMU WS 14151, light microscopy. A. Living specimen; B. Anterior end, dorso-lateral view. Scale bars: 5 mm (A); 1 mm (B).	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
3F464663B94150C6815B68E976461DF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582165	https://doi.org/10.3897/zookeys.1276.173213.figure9	Figure 9. Leodamas edgari sp. nov., paratypes, light microscopy. A. ZMMU WS 8279, anterior end, lateral view; B. P 12378, dorsal view; C. Same as A., neuropodia of chaetigers 10–12, lateral view; D. ZMMU WS 8300, thorax-abdomen transition, lateral view; E. ZMMU WS 8281, posterior end with regenerating pygidium; F. ZMMU WS 8283, posterior end with pygidium. Scale bars: 1 mm (A, B); 200 µm (C, D); 500 µm (E, F).	Figure 9. Leodamas edgari sp. nov., paratypes, light microscopy. A. ZMMU WS 8279, anterior end, lateral view; B. P 12378, dorsal view; C. Same as A., neuropodia of chaetigers 10–12, lateral view; D. ZMMU WS 8300, thorax-abdomen transition, lateral view; E. ZMMU WS 8281, posterior end with regenerating pygidium; F. ZMMU WS 8283, posterior end with pygidium. Scale bars: 1 mm (A, B); 200 µm (C, D); 500 µm (E, F).	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
3F464663B94150C6815B68E976461DF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582167	https://doi.org/10.3897/zookeys.1276.173213.figure11	Figure 11. Leodamas edgari sp. nov., paratypes, SEM. A – I. ZMMU WS 8283, J – M. ZMMU WS 8279. A. General lateral view; B. Anterior end, lateral view; C. Thorax-abdomen transition, chaetigers 15–19; D. Neuropodium of chaetiger 4; E. Neuropodium of chaetiger 9; F. Upper uncini of chaetiger 11; G. Lower uncini of chaetiger 11; H. Abdominal parapodia; I. Abdominal notopodium and neuropodium with strongly hooked aciculum; J. Chaetigers 19–23, thorax-abdomen transition, lateral organs (lo) are seen; K. Forked chaetae; L. Abdominal segments, dorsal view; M. Dorsal organ. Scale bars: 300 µm (A); 100 µm (B, C); 30 µm (D, E, H, I); 10 µm (F, G); 250 µm (J, L); 10 µm (K); 50 µm (M).	Figure 11. Leodamas edgari sp. nov., paratypes, SEM. A – I. ZMMU WS 8283, J – M. ZMMU WS 8279. A. General lateral view; B. Anterior end, lateral view; C. Thorax-abdomen transition, chaetigers 15–19; D. Neuropodium of chaetiger 4; E. Neuropodium of chaetiger 9; F. Upper uncini of chaetiger 11; G. Lower uncini of chaetiger 11; H. Abdominal parapodia; I. Abdominal notopodium and neuropodium with strongly hooked aciculum; J. Chaetigers 19–23, thorax-abdomen transition, lateral organs (lo) are seen; K. Forked chaetae; L. Abdominal segments, dorsal view; M. Dorsal organ. Scale bars: 300 µm (A); 100 µm (B, C); 30 µm (D, E, H, I); 10 µm (F, G); 250 µm (J, L); 10 µm (K); 50 µm (M).	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
3F464663B94150C6815B68E976461DF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582164	https://doi.org/10.3897/zookeys.1276.173213.figure8	Figure 8. Leodamas edgari sp. nov., holotype, ZMMU WS 14151, light microscopy. A. Anterior end, lateral view; B. Chaetigers 10–16, dorso-lateral view; C. Neuropodia of chaetigers 10–12, lateral view; D. Thorax-abdomen transition, dorso-lateral view; E. Abdominal parapodia, chaetigers 82–84; F. Neuropodia of chaetiger 82; G. Neuropodia of chaetiger 87. Scale bars: 500 µm (A, D, E); 200 µm (B, C, F); 100 µm (G).	Figure 8. Leodamas edgari sp. nov., holotype, ZMMU WS 14151, light microscopy. A. Anterior end, lateral view; B. Chaetigers 10–16, dorso-lateral view; C. Neuropodia of chaetigers 10–12, lateral view; D. Thorax-abdomen transition, dorso-lateral view; E. Abdominal parapodia, chaetigers 82–84; F. Neuropodia of chaetiger 82; G. Neuropodia of chaetiger 87. Scale bars: 500 µm (A, D, E); 200 µm (B, C, F); 100 µm (G).	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
3F464663B94150C6815B68E976461DF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582166	https://doi.org/10.3897/zookeys.1276.173213.figure10	Figure 10. Leodamas edgari sp. nov., paratypes, ZMMU WS 8283, parapodia slides. A. Parapodium of chaetiger 7; B. Same, upper neurochaetae; C. Same, lower neurochaetae; D. Parapodium of chaetiger 24 (anterior abdomen); E. Same, neuropodium; F. Posterior abdominal parapodium with slightly curved aciculum; G. Same, neuropodium; H. Posterior abdominal notopodium, two aciculae and forked chaetae seen; I. Posterior abdominal parapodium with strongly hooked aciculum; J. Same, neuropodium. Scale bars: 100 µm (A, D, F, I); 50 µm (B, C, E, G, H, J).	Figure 10. Leodamas edgari sp. nov., paratypes, ZMMU WS 8283, parapodia slides. A. Parapodium of chaetiger 7; B. Same, upper neurochaetae; C. Same, lower neurochaetae; D. Parapodium of chaetiger 24 (anterior abdomen); E. Same, neuropodium; F. Posterior abdominal parapodium with slightly curved aciculum; G. Same, neuropodium; H. Posterior abdominal notopodium, two aciculae and forked chaetae seen; I. Posterior abdominal parapodium with strongly hooked aciculum; J. Same, neuropodium. Scale bars: 100 µm (A, D, F, I); 50 µm (B, C, E, G, H, J).	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
BE5CDFC6D527571783CF99E0F889DC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582158	https://doi.org/10.3897/zookeys.1276.173213.figure2	Figure 2. Distributional maps of Leodamas species. Stars represent the type localities of the species.	Figure 2. Distributional maps of Leodamas species. Stars represent the type localities of the species.	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
BE5CDFC6D527571783CF99E0F889DC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582168	https://doi.org/10.3897/zookeys.1276.173213.figure12	Figure 12. Leodamas gracilis, P 12379, light microscopy. A – C. Stereomicroscope; D – G. Compound microscope. A. General lateral view; B. Anterior end, dorsal view; C. Thorax-abdomen transition, lateral view; D. Thoracic parapodia; E. Same, neurochaetae; F. Abdominal segment; G. Abdominal parapodia. Scale bars: 1 mm (A); 500 µm (B, C); 200 µm (D, F); 50 µm (E); 100 µm (G).	Figure 12. Leodamas gracilis, P 12379, light microscopy. A – C. Stereomicroscope; D – G. Compound microscope. A. General lateral view; B. Anterior end, dorsal view; C. Thorax-abdomen transition, lateral view; D. Thoracic parapodia; E. Same, neurochaetae; F. Abdominal segment; G. Abdominal parapodia. Scale bars: 1 mm (A); 500 µm (B, C); 200 µm (D, F); 50 µm (E); 100 µm (G).	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
BE5CDFC6D527571783CF99E0F889DC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582169	https://doi.org/10.3897/zookeys.1276.173213.figure13	Figure 13. Leodamas gracilis, ZMMU WS 8293, SEM. A. Anterior end, lateral view; B. Anterior end, dorsal view; C. Neuropodia of chaetigers 4 and 5; D. Neuropodia of chaetiger 1, antero-dorsally; E. Chaetigers 5–9, dorsal view, beginning of branchiae and dorsal organs are seen; F. Posterior upper part of neuropodium of chaetiger 5; G. Neuropodium of chaetiger 9; H. Lower part of neuropodium of chaetiger 8; I. Modified spines of chaetiger 8; J. Thorax-abdomen junction, chaetigers 14–17; K. Neuropodium of chaetiger 10; L. Abdomen, dorsal view; M. Dorsal organ; N. Abdominal neuropodium; O. Forked chaeta from abdominal notopodium. Scale bars: 500 µm (A, B); 20 µm (C, G, K, M); 10 µm (D, F, H, I); 100 µm (E); 50 µm (J); 200 µm (L); 5 µm (N).	Figure 13. Leodamas gracilis, ZMMU WS 8293, SEM. A. Anterior end, lateral view; B. Anterior end, dorsal view; C. Neuropodia of chaetigers 4 and 5; D. Neuropodia of chaetiger 1, antero-dorsally; E. Chaetigers 5–9, dorsal view, beginning of branchiae and dorsal organs are seen; F. Posterior upper part of neuropodium of chaetiger 5; G. Neuropodium of chaetiger 9; H. Lower part of neuropodium of chaetiger 8; I. Modified spines of chaetiger 8; J. Thorax-abdomen junction, chaetigers 14–17; K. Neuropodium of chaetiger 10; L. Abdomen, dorsal view; M. Dorsal organ; N. Abdominal neuropodium; O. Forked chaeta from abdominal notopodium. Scale bars: 500 µm (A, B); 20 µm (C, G, K, M); 10 µm (D, F, H, I); 100 µm (E); 50 µm (J); 200 µm (L); 5 µm (N).	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
BE5CDFC6D527571783CF99E0F889DC82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1582157	https://doi.org/10.3897/zookeys.1276.173213.figure1	Figure 1. Bayesian inference (BI) based on the concatenated dataset of COI and ITS 2. Bayesian posterior probabilities are shown on the nodes. Species delimitation results inferred by DNA-based methods are indicated by four colored bars to the right of the BI tree with the lineage number. White bars indicate missing data.	Figure 1. Bayesian inference (BI) based on the concatenated dataset of COI and ITS 2. Bayesian posterior probabilities are shown on the nodes. Species delimitation results inferred by DNA-based methods are indicated by four colored bars to the right of the BI tree with the lineage number. White bars indicate missing data.	2026-04-03	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya		Zenodo	biologists	Meca, Miguel A.;Zhadan, Anna;Al-Kandari, Manal;Budaeva, Nataliya			
