taxonID	type	description	language	source
3170BF66FFBCFF8FFDD0FBDAFA11F977.taxon	description	urn: lsid: zoobank. org: act: 32 B 8 E 0 B 1 - F 821 - 4873 - ABCD- 68 D 0742 D 97 D 7 (Figs 1 ‒ 22; 43 ‒ 45)	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFBCFF8FFDD0FBDAFA11F977.taxon	type_taxon	Type species. Clerus crassus Newman, 1840, by present designation.	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFBCFF8FFDD0FBDAFA11F977.taxon	description	Gender. Masculine.	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFBCFF8FFDD0FBDAFA11F977.taxon	diagnosis	Differential diagnosis. The following character-state differences separate Megaxestodes from Zenithicola: Club (Figs 1, 16) large, strongly differentiated from flagellum, terminal segment squared-off (Zenithicola with club smaller and terminal segment rounded-off or weakly pointed, Fig. 26); terminal labial and maxillary palpomeres respectively securiform and sub-securiform (sub-securiform and sub-digitiform in Zenithicola); anterior half of elytral disc (Figs 1, 9, 11, 16 ‒ 18, 43 ‒ 45) heavily foveolate-punctate in rows, posterior half nitid, white setal tufts present (Zenithicola with non-linear sculpturing of irregular form and depth, white setal patches absent, Figs 23, 26); scutellum with dense tuft of white setae (setae differs in Zenithicola); metaventrite (Fig. 7) anteriorly, between mesocoxae, with a large wedge-shaped process strongly projecting ventrad (mesocoxae of Z enithicola with weaklyprojecting inter-coxal node-like process, Fig. 24); tibial spur formula 2 ‒ 2 ‒ 1 (males) or 2 ‒ 2 ‒ 2 (females) (2 ‒ 2 ‒ 2 in males and females of Zenithicola); hindwing crossvein CuA 1 absent or incomplete (Fig. 8) (complete in Zenithicola, Fig. 25); tegmen (Figs 2, 12, 19) narrow apically (parameroid region) and basally (apodeme), with middle section (tegminal arms) notably broader (tegmen of Zenithicola, Figs 27, 28, short and broad with the base splayed as in Phlogistomorpha and Phlogistus, Figs 29, 30).	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFBCFF8FFDD0FBDAFA11F977.taxon	description	Description. Stocky, robust in appearance (Figs 1, 9, 16 ‒ 18, 43 ‒ 45); length 8.5 ‒ 14.0 mm. Head: Eyes emarginate, finely-facetted, separated by approximately two eye widths; terminal labial palpomeres securiform (outer to inner edge ratio ~ 2.4: 1), terminal maxillary palpomeres sub-securiform (outer to inner edge ratio range 1.2 ‒ 1.5: 1); antennae 11 - segmented, about as long as width of pronotum, flagellomeres filiform, terminal three segments forming a large compact club (Figs 1, 16), terminal segment relatively quadrate in females, more elongated in males. Thorax: Subquadrate to slightly transverse (length to width ratio 0.9 ‒ 0.95: 1), procoxal cavities open posteriorly, about half closed by postcoxal hypomeral processes and distal expansion of prosternal process; anterior part of metaventrite with large wedge-shaped ventral projection between mesocoxae (Fig. 7); scutellum with dense tuft of white setae; elytra (Figs 1, 9, 11, 16 ‒ 18, 43 ‒ 45) compact (length to width ratio range 1.4 ‒ 1.6: 1), basal half deeply punctate, middle of base with conspicuous basal hump, apical half of disc mostly nitid, each elytron with two short transverse tufts of white setae at the posterior limit of punctation, a preapical tuft is typically present (not in all specimens); hindwing with CuA 1 crossvein absent or incomplete (Fig. 8 a), CuA 3 + 4 crossvein complete (Fig. 8 b) (when CuA 1 incomplete, both crossveins meet at same point along main CuA vein); tibiae longitudinally carinate, tibial spur formula 2 ‒ 2 ‒ 1 (males) or 2 ‒ 2 ‒ 2 (females), ventral tarsal pad formula 3 ‒ 3 ‒ 3, pretarsal claws simple. Abdomen. Pygidium (Fig. 6, 15, 22) with slender pygidial arms. Male Genitalia: Tegmen (Figs 2, 3, 12, 19) with tegminal arms not looped below point of fusion to phallobasic apodeme; median lobe (Fig. 4, 13, 20) with nodulate apico-medial process; spicular fork (Fig. 5, 14, 21) division near middle.	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFBCFF8FFDD0FBDAFA11F977.taxon	etymology	Etymology. The generic epithet Megaxestodes (from Greek megas, meaning ‘ large’ or ‘ great’, and Greek xestes, meaning ‘ scraper’ and the Latin suffix - odes meaning ‘ like’) refers to the large ventral projection of the metasternum which, with the outer margin serrated, has the appearance of a scraper-like structure. Speculatively, such a structure may facilitate adult emergence from a pupal cocoon or, having been recorded in association with termites (Moore 1973), to break through the outer walls of a termite nest.	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFBCFF8FFDD0FBDAFA11F977.taxon	discussion	Remarks. Photographs taken by the author of syntype specimens of Zenithicola funestus (Figs 9, 10), Clerus obesus (Fig. 11), Z. socius (Fig. 17) and Z. cribricollis (Fig. 18) (all in MNHN, Paris), facilitated the following observations: (a) the black pronotum of Clerus obesus suggests it to be a synonym of Z. funestus, rather than Z. crassus; (b) the pronotal structure and shape of the apical elytral setal tuft of Z. socius and Z. funestus differ significantly enough to revoke Schenkling’s (1906) synonymy of the former with the latter; (c) similarities between the types of Z. socius and Z. cribricollis (robust pronotum with prominent lateral setal tuft, non-linear shape of the apical elytral setal tuft) suggest synonymy of the two names. Blackburn’s (1891) suggestion of synonymy of Clerus obesus with Zenithicola crassus has been followed in catalogues since (Lohde, 1900, Schenkling 1910, Corporaal 1950). A MNHN specimen (Fig. 11) resembling Megaxestodes funestus (Chevrolat) comb. nov., with a black pronotum, and bearing two hand written labels, one in a style identical to many Westwood type specimens (pers. obs.; noting co-publication of Westwood and White, in White 1849), i. e., a square label with ‘ Clerus? obesus White’ plus the publication title and page and figure numbers, and another label stating ‘ Z. funestus Chv. Compared with type’, is taken here to be White’s Clerus obesus syntype, designated here as the Lectotype. Clerus obesus White, 1846 syn. nov. is therefore synonymised with Megaxestodes funestus (Chevrolat, 1874) comb. nov.. The Zenithicola funestus syntype (MNHN; Figs 9, 10), labelled ‘ Z. funestus Chv. ’, and positioned above a faded pink hand written label (Zenithicola funestus Chv. Rev. Zool 1874 - 49 New South Wales) often associated with type material in the Chevrolat / Sedillot Collection, is designated here as the Lectotype. Megaxestodes crassus (Newman) and M. funestus are strikingly similar morphologically, apparently differing only in pronotal colouration, orange and black respectively. Sexual dichromatism can be ruled out as males and females of both are known to the author. Whether the slightly broader tegmen (Fig. 12) and less elongate pygidium (Fig. 15) of M. funestus (compare Figs 2 and 6), or variability in the intensity of the metallic blue sheen observed in specimens with orange, and black pronota, are indications of species-level difference or interspecific variation, is yet to be investigated. Megaxestodes crassus and M. funestus are here retained as valid species, pending DNA sequencing evidence to the contrary. The robust pronotum with anterolateral setal tuft, and circular rather than linear (as in Z. funestus / crassus) preapical elytral setal tuft, are characteristics shared by the syntypes of Z. socius and Z. cribricollis indicating possible synonymy, with only size of the apical setal tuft (very small in Z. cribricollis, compare Figs 17 and 18), casting doubt over conspecificity of the two taxa. Two specimens collected together on Leptospermum 90 km north of Zanthus, Western Australia (28.11.92, D. Knowles, JSBC), a male with a large preapical setal tuft (resembling the Z. socius syntype) and a female with a smaller preapical setal tuft (as in the Z. cribricollis syntype), however, provide evidence for intraspecific variability in the size of the preapical setal tuft. Zenithicola cribricollis Pic, 1941 syn. nov. is therefore synonymised with Megaxestodes socius (Chevrolat, 1874) stat. rev., comb. nov.. The Zenithicola socius syntype (MNHN; Fig. 17), with a yellow label printed with ‘ 266 ’, and pinned through a faded pink hand-written label (Zenithicola sosius Chv. Rev. Zool 1874 - [illegible] Adelaidia) commonly associated with type material in the Chevrolat / Sedillot Collection (author’s obs.), is designated here as the Lectotype. The Zenithicola cribricollis syntype (MNHN; Fig. 18), labelled ‘ Australie / cribricollis mihi. ’, with a rectangular red TYPE label, is designated here as the Lectotype. Historical identifications: A photograph (Fig. 40, in Opitz 2003) published as Zenithicola funestus, shows a specimen that conforms morphologically to M. socius. Previous identifications of Megaxestodes taxa that, due to collecting locality, are likely to be correct, include those of Blackburn (1891 and 1892, for M. socius) and Elston (1921, for M. crassus and funestus).	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFBCFF8FFDD0FBDAFA11F977.taxon	distribution	Distribution. Megaxestodes socius is an arid-zone adapted species with specimens known from WA (south of Mount Magnet), SA (near Adelaide), NSW (a single record from Western NSW, see Maywald 2023) and Qld (one specimen from Normanton, in MNHN, spelled as Normantown). Megaxestodes crassus and M. funestus occur in eastern Australia from Vic to Qld (to south of Mount Carbine). The genus is absent from Tasmania. While specimens from Northern Territory are so far unknown, the specimen from Normanton, far north-west Qld, suggests occurrence in NT to be likely.	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFBBFF8DFDD0F89EFE14FEC7.taxon	description	In addition to differences between Zenithicola australis (the type species of Zenithicola) and species now assigned to Megaxestodes gen. nov., the present study also revealed that Z. australis and the genera Phlogistus and Phlogistomorpha (containing 48 and four species respectively, Bartlett 2018) share the same fundamental form of the male tegmen, i. e., short, with a broad splayed base (resembling a fishes caudal fin) in place of a defined apodeme, and with the phallobase seemingly reduced to a fine spicule (Figs 28, 30) or absent (Fig. 29) (obscured in Fig. 27). The monotypic genera Mimolesterus Gerstmeier, from Australia, and Balcus Sharp, from New Zealand, are proposed as putatively related phylogenetically to the above three genera, due to the very similar tegmen, which differs mainly by the base being broad and spatulate, not splayed. Preliminarily surveying morphological character states of the above five genera indicates possible polyphyly of Phlogistomorpha and Phlogistus that requires more focused testing. Notably significant is the incongruence observed in number of antennomeres between the type species of these genera and all other species presently assigned to them, specifically: Phlogistomorpha blackburni (Schenkling) with 11 antennal segments (not 10 as reported by Bartlett 2009) and its current congeners (all with 10 segmented antennae); Phlogistus instabilis (Newman) (10 segmented antennae) and all other species hitherto assigned to Phlogistus (11 segments). Notable variation observed in other characters (e. g., broadness of terminal labial palpomere, shape of terminal antennomere, shape and sculpturing of pronotal disc, presence / absence of tibial carination) should be further documented and taxonomically investigated. Similarity in general appearance between Mimolesterus ventralis (Westwood) and Phlogistus leucocosmus Elston, both with a pale setose elytral band (uncharacteristic of Phlogistus), and between Zenithicola australis and Phlogistus rotundopunctatus Hintz, with comparable crater-like elytral sculpturing, similarly rotund pronotum and sub-digitiform terminal maxillary palpomeres, reiterate the need to revisit genus assignment of species presently assigned to Phlogistus.	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFB9FF8DFDD0FE0FFA10F8A7.taxon	description	The identity of the type species of a genus is generally of high importance in terms of stability of genus-defining morphological characters. This is less crucial in the case of Zenithicola sensu novo (i. e., minus species transferred to Megaxestodes) as the genus is now monotypic and character states considered here of fundamental importance at the genus level are present in all ‘ australis - like’ specimens across the geographic range. An attempt has nevertheless been made to recognise true Z. australis. Boisduval (1835) described Clerus australis from material collected during the 1826 ‒ 1829 voyage of the French Fluyt Astrolabe, giving no indication of collecting locality other than New Holland (Il se trouve à la Nouvelle-Hollande). Museum specimens (AM, ANIC, ASCU, MV, QDPC, QM), and iNaturalist observations, presently identified as Z. australis are distributed between Maryborough in Queensland, and Sydney in New South Wales. Comparison of tegmina of males from Woy Woy (45 km north of Sydney) and Brisbane, though fundamentally similar in form, shows proportional and shape differences potentially indicating non-conspecificity (compare figs 27 and 28). Similarly, variability of external characters observed across the distributional range (e. g., body length, leg colour, tint of metallic sheen of elytra, setal vestiture, elytral sculpturing), though relatively subtle, suggest possibility that the pool of ‘ australis - like’ specimens may not constitute a singular species. The author is in possession of a digital photograph (courtesy Ganyang Yang, not reproduced here) of the entire case of insects containing the holotype of Zenithicola australis sensu Spinola, 1845 (a junior homonym of Clerus australis Boisduval, see Ekis 1975: 39) in MRSN. Zooming closely into the Z. australis holotype in that photograph, despite being slightly dark and lacking clarity in finer detail, shows an insect that undeniably has the bulky form and irregular elytral punctation distinctive of beetles universally labelled as Z. australis in museum collections. A possible Clerus australis syntype was located in MNHN (Fig. 23) but should be considered putative until Zenithicola specimens of MNHN and NHML are thoroughly checked again. In the absence of a definitive type, specimens considered representative of true Z. australis should (a) be collected from the same region as Boisduval’s missing type, and (b) conform morphologically to the original description. During its voyage around Australia, the Astrolabe moored only in Albany, WA (7 ‒ 25 October 1826), Western Port, Victoria (12 ‒ 19 November 1826), Jervis Bay (26 ‒ 29 November 1826) and Sydney (3 ‒ 17 December 1826) in NSW, and Hobart, Tasmania (20 December 1827 to 4 January 1828) (Rosenman 1992). Considering the abovementioned geographic range, the type locality of Clerus australis is therefore likely to be in the vicinity of Jervis Bay or Sydney, New South Wales. The dissected male specimen from Woy Woy (Fig 26), being collected only 45 km north of Sydney, and conforming to Boisduval’s (1835) description of Z. australis in all aspects (hairy, bronze, elytral base roughly puntate, antennae and legs redbrown), is considered here to be representative of Z. australis.	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFB9FF8DFDD0FE0FFA10F8A7.taxon	discussion	Synonymies and taxonomic notes Publication of new synonymies, and other taxonomic information, here was deemed constructive ahead of subsequent work on Zenithicola- complex taxa. Synonymies are based on examination of type material unless otherwise noted.	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFB6FF82FDD0FE0EFE3DFCFC.taxon	discussion	Synonymy of Aulicus corallipes Chevrolat, 1874 syn. nov. with Phlogistus castanipes (Westwood, 1849) stat. rev., comb. nov. is proposed despite not examining Chevrolat’s syntype as: a) corallipes appears to be a relatively distinct and well-known taxon, see Blackburn 1900, Hintz 1908, Elston 1921, all of whom ignored Schenkling’s (1899) synonymy of A. corallipes with Phlogistus foveicollis (Macleay), the latter taxon clearly unknown to Schenkling, the elytral punctation of type specimens being notably less uniform than corallipes / castanipes; b) an SDEI specimen determined by Schenkling as Phlogistus corallipes appears to be identical to Westwood’s Clerus castanipes syntype (Fig. 33), with uniformly-arranged, slightly transverse elytral punctation; and c) the type locality of both taxa is Tasmania.	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFB6FF82FDD0FC32FABDFA3C.taxon	discussion	Phlogistus eximius (White, 1849) was originally described from ‘ W. Australia’ with its location subsequently given as ‘ Hunter’s River, Australia Occidentali’ by Westwood (1853). Though there is a Hunter River in the Kimberly region of Western Australia, this author is only aware of specimens resembling P. eximius from eastern Australia (Qld, NSW). All other clerid species described from the ‘ Hunter River’ around the same time, i. e., Cleromorpha novemguttata (Westwood, 1853), Mimolesterus ventralis (Westwood, 1853), and Lemidia hilaris (Newman, 1850) (erroneously as Hunter River, New Zealand), now being known to occur only in eastern Australia (ALA 2026), provides reason to consider the type locality of eximius Hunter River in NSW (i. e., Newcastle), not Western Australia. Interestingly, photographs taken in Skennars Head, north of Ballina, NSW, and posted to iNaturalist (Taylor 2024; Fig. 46), show a typically eximius - like multichromatic female being mated by a monochromatic male somewhat resembling P. kolbei Hintz. The holotype of Phlogistus kolbei, described from Cape York, far north Qld, however, appears to be more conspicuously punctate at the elytral base than the less obviously punctate kolbei - like individuals in NSW and southeast Qld, and therefore should not be assumed conspecific with P. eximius. Any future synonymy of the two names should be based on molecular evidence. Nevertheless, Phlogistus eximius is a species exhibiting striking sexual dichromaticism, much in the same manner as P. schulzi Hintz and P. sculptus (see below).	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
3170BF66FFB4FF81FDD0F9B5FB2AFF77.taxon	discussion	The species identity and correct genus placement of Xylotretus scrobiculatus (Spinola, 1845), hitherto assigned to Zenithicola (following Blackburn 1900), is impossible to determine until type material can be located. Ekis (1975), who was unable to locate X. scrobiculatus type material in the Spinola collection (now in MRSN), suggested that Spinola types missing from MRSN should be found in Paris (MNHN). Two visits to MNHN (2008, 2011) by the present author did not yield discovery of X. scrobiculatus types, only one non-type specimen determined to be X. scrobiculatus by J. R. Mawdsley resembling a dark coloured Zenithicola australis, but not resembling particularly well the illustration published by Spinola (1845: pl. XV, fig. 4; wrongly labelled Xylotretus foveolatus Spinola, see Gorham 1876) which does not appear to be congeneric with Zenithicola australis, due to the clearly defined puncture rows and relatively long antennae, or with Megaxestodes crassus comb. nov., due to the lessbroad body form, absence of elytral setal patches, smallness of the eyes and elongation of the terminal antennal segment. Spinola’s (1845) description of a transverse fold at the posterior limit of the foveolate part of the elytra, to this author’s knowledge unknown in any Australian clerid, suggested to Blackburn (1900) the possibility that the Australian type locality of X. scrobiculatus was erroneous. Until discovery of type material provides elucidation, or discovery of an Australian-collected specimen undeniably resembling Spinola’s description and illustration, it is proposed that Xylotretus scrobiculatus Spinola, 1845 be classified as Clerinae incertae sedis.	en	Bartlett, Justin S. (2026): Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘ Zenithicola-complex’ including taxonomic notes and new synonymies. Zootaxa 5828 (2): 333-346, DOI: 10.11646/zootaxa.5828.2.8, URL: https://doi.org/10.11646/zootaxa.5828.2.8
