identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
8DE70235C63A5418A03AB360F95EA606.text	8DE70235C63A5418A03AB360F95EA606.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ducanhia balansai (Morlet 1886)	<div><p>Ducanhia balansai (Morlet, 1886)</p><p>Figs 6, 7, 8, 9</p><p>Helix balansai Morlet, 1886: 7 .</p><p>Helix balansai: Morlet 1887: 258, 270, 271, pl. 12, figs 4, 4 a, 4 b.</p><p>Helix balansai: Dautzenberg and d’Hamonville 1887: 218. (Rochers de marble du Nuy-Dong-Nay [marble rocks of Dong Nai Province])</p><p>Helix (Trachia) balansai: Fischer 1891: 26.</p><p>Chloritis balansai: Fischer and Dautzenberg 1904: 11.</p><p>Helix (Chloritis) balansai var. cincta: Dautzenberg and Fischer: 1905: 90, pl. 3, figs 5–9.</p><p>Chloritis (Trichochloritis) balansai: Gude 1906: 116.</p><p>Chloritis (Trichochloritis) balansai var. cincta: Gude 1906: 116.</p><p>Chloritis balansai: Richardson 1985: 86.</p><p>Trachia balansai: Schileyko 2011: 45.</p><p>Type locality.</p><p>Not stated (Tonkin – from title) ( H. balansai); Ile Krieu, Archipel des Faï-Tsi-Long, Tonkin [Krieu Island, Ha Long Provincial, Quang Ninh Province, Vietnam] ( H. balansai var. cincta).</p><p>Type material examined.</p><p>Syntypes • MNHN -IM-2000-2078 (1 syntype of H. balansai) ; • MNHN -IM-2000-2077 (2 syntypes H. balansai var. cincta) .</p><p>Additional material.</p><p>• Tonkin, ex Balansa 1887 MNHN -IM-2013-49807 (1 shell) ; • Tonkin, coll. Blaise MNHN -IM-2013-49808 (9 shells); • Tonkin, Ile Krieu, coll. Blaise, MNHN -IM-2013-49809 (5 adult + 1 juvenile shells) ; • Tonkin, Ile Krieu, archipel des Fai-Tsi-Long, coll. Blaise, MNHN -IM-2013-49811 (s juvenile shells) ; • Tonkin, coll. Jousseaume MNHN -IM-2013-49810 (2 shells); • Tonkin, leg. Abbé Vathelet, MNHN -IM-2013-49812 (6 shells); • Baie d’Along, leg. Abbé Vathelet 1887, MNHN -IM-2013-49813 (3 shells) ; • Vietnam, Quang Ninh Prov., Halong Bay Area, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.3025&amp;materialsCitation.latitude=20.8715" title="Search Plazi for locations around (long 107.3025/lat 20.8715)">unnamed island 1.8 km W of S point Cong Tai Isl.</a>, steep limestone slope bordering beach, dense vegetation, handpicked + soil sample, 20°52.29N, 107°18.15E, J. J. Vermeulen &amp; A. J. Whitten leg., 3 Oct. 1998, ex coll. Vermeulen 6455, NHMUK 19991421 (2 complete + 1 damaged shells) “ Chloritis balansai cincta ” ; • Vietnam, Cat Ba Island, Cat Ba N. P., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.007&amp;materialsCitation.latitude=20.7969" title="Search Plazi for locations around (long 107.007/lat 20.7969)">Hai Phong City, near pass in front of May Bau</a>, 20.7969°N, 107.0070°E, ca 97 m, leg Otani, J. U., 22. 11. 2007, IEBR_LS_Ducanhia 001 H .</p><p>Diagnosis.</p><p>Shell large, thin, bright yellow, almost flat, with a subsutural furrow; hairs cover the complete shell, umbilicus partly to almost completely covered by hair scars or bristles.</p><p>Description.</p><p>The shell is large, thin, depressed, with only a slightly elevated spire and somewhat domed apical part. The body whorl is somewhat expanded with a clearly marked subsutural furrow visible on the whole last whorl. The shell colour is pale greyish to yellow, a red spiral band may run above the furrow. The protoconch consists of 1.75 – ~ 2 whorls, completely covered by dense pattern of hair scars. The teleoconch also beats hairs (in fresh shells) or hair scars (weathered shells). The 4.25–4.75 whorls are separated by a rather shallow suture. The hairs are short and tend to be permanent. The aperture is semilunar, the reflected peristome reinforced by a thick white or even reddish lip. The parietal region is with an inconspicuous additional layer which is not necessarily paler than the rest of the shell, but glossy. The umbilicus is slightly funnel-shaped and partly to almost completely concealed by the reflection of the columella.</p><p>Measurements.</p><p>D = 23.2–24.5 mm, H = 13.0– 13.6 mm (n = 2: MNHN -IM-2013-49808).</p><p>Description of the genital organs.</p><p>The right ommatophoral retractor crosses the penis and the vagina. The atrium is very short. The penis is relatively long, consists of a spindle-shaped distal and a thickened, globular proximal portion. The inner wall of the distal part has irregular longitudinal, thick folds; the thickest longitudinal fold bears some transversal wrinkles. The proximal penial portion internally has a strongly thickened sphincter-like structure with one side being convex, and the other concave. These two sides fit exactly to each other. Just above of this muscular sphincter-like part there is a small, papilla-like penial verge having central opening. The long epiphallus consists of a thicker distal, and a slender proximal portion. The boundary between these two parts is transitional, not abrupt. The thicker part internally with four longitudinal, slightly serrate folds. The thinner portion of the epiphallus internally with small rhomboid papillae. The retractor muscle attaches on the diaphragm and inserts near the distal end of the epiphallus. There is an additional, weaker muscle, which is divided into several branches inserting on the proximal part of the penis and the distal part of the epiphallus. The flagellum is very short, with a curved tip. The flagellum is internally with broad, low, ribbed longitudinal folds, and an elevated, slender longitudinal fold starting from the point where vas deferens originates. The vas-deferens is overall slender. The vagina is approximately as long as the penis, its distal end is the thinnest and it gradually tapers towards the spermoviduct. The distal part of the vagina (i. e., closer to the atrium) internally bears strongly serrated longitudinal folds. In proximal direction (i. e., towards the spermoviduct) these folds become less serrated and weaker. At the proximal end of the vagina there are free, elevated lobes internally. The bursa copulatrix is conspicuously short, thick, and with a rather pointed end. An amorphous spermatophore was found inside the bursa.</p><p>Remarks.</p><p>Inkhavilay et al. (2019: fig. 48 D) reported this species from Laos. However, the shell they figured has a wider umbilicus, and lacks the subsutural furrow characteristic for typical Helix balansai shells. Moreover, that shell had multiple reddish bands on the body whorl, suggesting that it may belong to Burmochloritis (see Páll-Gergely et al. 2023) instead of conspecific with H. balansai .</p><p>The red spiral may be present or absent even in the same populations, so it does not justify a separation of a subspecies. The degree of coverage of the umbilicus varies from only a very small proportion concealed to almost completely reflected.</p></div>	https://treatment.plazi.org/id/8DE70235C63A5418A03AB360F95EA606	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Dedov, Ivailo	Páll-Gergely, Barna, Gojšina, Vukašin, Dedov, Ivailo (2026): New taxa of Camaenidae from northern Vietnam (Gastropoda, Stylommatophora). ZooKeys 1280: 365-392, DOI: 10.3897/zookeys.1280.185160
4EDC29605E115A5EB90D3EA3A687B765.text	4EDC29605E115A5EB90D3EA3A687B765.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ducanhia Pall-Gergely 2026	<div><p>Ducanhia Páll-Gergely gen. nov.</p><p>Type species.</p><p>Helix balansai Morlet, 1886 .</p><p>Diagnosis.</p><p>Shell relatively large, dorsal side nearly flat, umbilicus only slightly open, entire shell covered with hairs. Small penial verge present with central opening; flagellum present, penial caecum absent; retractor muscle inserts near the distal end of the epiphallus; stalk of bursa copulatrix very short.</p><p>Differential diagnosis.</p><p>The anatomical traits differentiate this new genus from all Southeast Asian Chloritis - like genera. Bellatrachia has no penial verge and has a thickened base of the bursa copulatrix (Schileyko 2018; Páll-Gergely and Neubert 2019). Bouchetcamaena has a much more elongated, pointed flagellum, and lacks a penial verge (Páll-Gergely et al. 2022). Trichochloritis has a penial caecum (Páll-Gergely and Neubert 2019). Continental Chloritis species have a pointed flagellum and a less muscular, thinner walled penis (Sutcharit and Panha 2010; Páll-Gergely et al. 2020). Moreover, all mentioned genera have a longer stalk of the bursa copulatrix.</p><p>Etymology.</p><p>this new genus is dedicated to and named after Duc Anh Nguyen, specialist of millipedes, a much-valued friend of the first author. Genus feminine.</p><p>Remarks.</p><p>The key traits of this new genus are the short bursa copulatrix, and the complicated inner structure of the penis. Namely, distally (i. e., in direction of the atrium) from the small penial verge, there is a sphincter-like region between the penial verge and the most distal (basal) part of the penis. A similar penial structure was described in Planispira H. Beck, 1837 (see Schileyko 2003). However, the latter genus inhabits a different biogeographical region (New Guinea and Moluccas) and Planispira possesses a long stalk of the bursa copulatrix.</p><p>Recently, Helix balansai was classified in the genus Trachia Martens, 1860 . Unfortunately, the anatomy of type species of Trachia (i. e., Helix asperella L. Pfeiffer, 1846) is unknown. Helix asperella was described from India (type locality: “ Bithonia in ripa Gangis ”) and was subsequently reported from several localities in central and southern India (see Gude 1914; Ramakrishna Mitra and Dey 2010), but not from the eastern Himalayas, which would have biogeographical connections with Southeast Asia. Schileyko (2018) assigned several species to Trachia from India, Myanmar, Sri Lanka, and the Andaman Islands based on conchological similarities, including Trachia delibrata (W. H. Benson, 1836), the only species with known anatomical traits. The latter species, however, is known from the western Himalayas (type locality: “ North-East frontier of Bengal ”), and was classified in Bouchetcamaena based on anatomical similarities with other Bouchetcamaena species (Páll-Gergely et al. 2022). Therefore, it seems that Trachia is a genus restricted to central and southern India and maybe Sri Lanka and the Andaman Islands, but species east of the Himalayas would belong to other genera.</p></div>	https://treatment.plazi.org/id/4EDC29605E115A5EB90D3EA3A687B765	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Dedov, Ivailo	Páll-Gergely, Barna, Gojšina, Vukašin, Dedov, Ivailo (2026): New taxa of Camaenidae from northern Vietnam (Gastropoda, Stylommatophora). ZooKeys 1280: 365-392, DOI: 10.3897/zookeys.1280.185160
4E551E6AE89E5C9D83E840D5EE9C141C.text	4E551E6AE89E5C9D83E840D5EE9C141C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fansipanica gereti (Bavay & Dautzenberg 1900) Páll-Gergely & Gojšina & Dedov 2026	<div><p>Fansipanica gereti (Bavay &amp; Dautzenberg, 1900) comb. nov.</p><p>Fig. 5</p><p>Helix (Chloritis) gereti Bavay &amp; Dautzenberg, 1900: 112 .</p><p>Helix (Chloritis) gereti: Bavay and Dautzenberg 1900: 442, pl. IX, figs 7–9.</p><p>Chloritis gereti: Fischer and Dautzenberg 1904: 11.</p><p>Chloritis (Trichochloritis) gereti: Gude 1906: 116.</p><p>Chloritis gereti: Richardson 1985: 98.</p><p>Trichochloritis gereti: Schileyko 2011: 47.</p><p>Type locality.</p><p>“Bac-Kan et Phi-Mi” [Bãc Can: 22°08'N, 105°49'E, see Schileyko 2011, Phi Mi could not be found on the map].</p><p>Type material.</p><p>Syntype. • Bac Kan et Phi-Mi, leg. Messager, MNHN -IM-2000-1900 (1 specimen) Fig. 5 .</p><p>Additional material.</p><p>• Tonkin, Bac Kan, coll. Messager, MNHN -IM-2012-27100 (48 shells) (this is probably the original lot); • Bac-Kan, coll. Denis MNHN -IM-2012-27099 (7 shells); • Tonkin, Bac Kan, coll. Letellier MNHN -IM-2012-27101 (1 shell); • Tonkin, Bac-Kan, coll. Staadt MNHN -IM-2012-27102 (3 shells); • Tonkin, “ v. minor ”, MNHN -IM-2012-27103 (2 shells); • Tonkin, MNHN -IM-2012-27104 (13 shells); • Tonkin, Salisbury collection, “imaged by H. Taylor (NHM) for North Vietnamese Land Snail Guide Mollusca Section Neg. No. 2050”, NHMUK 20250272 (1 shell) .</p><p>Diagnosis.</p><p>A small species with globular shell, closed or nearly closed umbilicus, irregularly wrinkled shell bearing deep, regularly arranged hair scars.</p><p>Description.</p><p>The shell is small, globular, pale brown. The subsutural furrow is absent, the body whorl is rounded. The protoconch consists of 1.5 whorls, and is very finely, irregularly wrinkled and bears hair scars. The teleoconch is finely and irregularly wrinkled, and covered by widely spaced but rather regularly arranged deep hair scars. The hairs are probably not permanent (although no live collected specimens were available). The 4.75–5 whorls are separated by a moderately deep suture. The aperture is semilunar. The peristome is expanded but not reflexed, whitish. The parietal region is glossy, paler than the rest of the shell in some specimens. The umbilicus is very narrow, covered by the reflection of the columellar peristome, in some specimens covered entirely.</p><p>Anatomy of genital organs.</p><p>Unknown.</p><p>Measurements.</p><p>D = 13.7–16.4 mm, H = 10.9–13.0 mm (n = 4).</p><p>Remarks.</p><p>This species is included in Fansipanica gen. nov. due to its conchological similarities with its type species. The assignment of F. gereti to Fansipanica gen. nov. should be considered provisional and based on the only information available at the moment, i. e., the conchological similarity.</p></div>	https://treatment.plazi.org/id/4E551E6AE89E5C9D83E840D5EE9C141C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Dedov, Ivailo	Páll-Gergely, Barna, Gojšina, Vukašin, Dedov, Ivailo (2026): New taxa of Camaenidae from northern Vietnam (Gastropoda, Stylommatophora). ZooKeys 1280: 365-392, DOI: 10.3897/zookeys.1280.185160
A15685B371C85B0589F4D2F2D0EA4A36.text	A15685B371C85B0589F4D2F2D0EA4A36.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fansipanica milae Pall-Gergely & Dedov 2026	<div><p>Fansipanica milae Páll-Gergely &amp; Dedov sp. nov.</p><p>Figs 1, 2, 3, 4</p><p>Type material.</p><p>Holotype (D: 12.8 mm, H: 9.5 mm); • Vietnam, Northern, Lào Cai Province, Sa Pa district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.7759&amp;materialsCitation.latitude=22.3046" title="Search Plazi for locations around (long 103.7759/lat 22.3046)">Fansipan Peak</a>, bamboo forest below the peak, 22.3046°N, 103.7759°E, ca 3040 m, 24 X 2023, leg. I. Dedov, N. Simov, R. Bekchiev, M. Langurov, HNHM 105552 . Paratypes: • IBER Coll. No. 40619/2, same locality; IBER Coll. No. 40633/1, same locality, 22.3052°N, 103.7756°E, ca 3033 m, 09 X 2025, leg. N. Simov, K. Ivanov .</p><p>Description.</p><p>Body. Body surface granular, uniformly dark grey coloured, including long eye tentacles. Area around genital pore smoother and paler. The sole is also paler, whitish-yellowish.</p><p>Shell. The shell of the living animal is dark brown coloured with brown-reddish hue, in preserved specimens pale reddish brown. The shell is fragile, medium sized, globose, slightly wider than high, the subsutural furrow is absent, the body whorl is rounded. The 4.25–4.5 regularly increasing whorls are separated by a shallow suture. The protoconch is consisting of 1.25–1.5 whorls, and bears dense, oval scales. This sculpture abruptly changes to the teleoconch sculpture at the protoconch-teleoconch boundary. The teleoconch is evenly covered by widely spaced hairs (where absent: raised, papilliform hair scars). The hairs are permanent on the entire shell. The hair density is higher on the ventral side near the umbilical area than on the dorsal side and the periphery of the last whorl. The aperture is semilunar, slightly oblique to shell axis (prosocline). The parietal callus is absent, the parietal region is only indicated with an additional thin, transparent calcareous layer. The peristome is pinkish-reddish, slightly expanded and reflected only near the umbilical area. The umbilicus is extremely narrow, almost covered by the reflected peristome.</p><p>Measurements.</p><p>D = 11.9–12.8 mm, H = 9.4–9.8 mm (n = 3).</p><p>Genital organs.</p><p>The atrium is short. The penis is long, nearly cylindrical, its proximal (“ apical ”) third is covered with a thin penial sheath. The inner wall of penis is covered by irregular, slightly serrate longitudinal wrinkles, which gradually becoming stronger (more elevated) and more regular from the atrium towards the proximal end, where their number is five. The penial verge is small, conical, opening along a slit that ends close to its tip. The thick penial retractor muscle inserts at the penis-epiphallus junction. The epiphallus is slender, slightly shorter than the penis, internally with a single, elevated fold. A small, blunt flagellum marks the boundary between epiphallus and vas deferens. The vas deferens is only slightly more slender than the epiphallus. The vagina is slightly shorter than the penis, its distal part (closer to the atrium) is attached to the body wall with numerous fibres. The inner wall of the vagina bears numerous (~ 10) slender folds that are variable in width. The folds (mostly their distal parts) are slightly serrate, and the inner wall of the proximal part of the vagina (closer to the spermoviduct) is finely sculptured by a maze-like structure. The stalk of the bursa copulatrix is very long (&gt; 2 × longer than the penis), approximately as wide as the epiphallus. The bursa was torn off when the animal was pulled out of its shell, therefore its morphology could not be examined.</p><p>Differential diagnosis.</p><p>The most similar species is F. gereti (Bavay &amp; Dautzenberg, 1900), comb. nov. due to the similar size (although it is slightly larger), the shape of the shell and the (nearly) closed umbilicus. However, the new species has much denser hairs on the entire shell surface and has a darker peristome.</p><p>Etymology.</p><p>The species is named after Ivailo Dedov’s wife Mila Taseva, for her support over the years and patience during the long field trips.</p><p>Habitat.</p><p>Fansipan Mountain and the adjacent areas in Lao Cai Province (western part of the Hoang Lien Mountains) are a very unique region in northern Vietnam, where a large area of closed canopy primary forest in a mountain habitat persists. Nowadays, due to excessive utilisation, the tropical rainforest is only maintained in disconnected patches. The total number of species that can still be described makes the area rich (Sobey 1997). Hoang Lien National Park is located in the Hoang Lien Mountain Range with Fansipan Peak, which is considered the highest mountain in Vietnam, at 3143 m a. s. l. (Nguyen et al. 2022). The mollusc fauna of this region is almost unknown (Páll-Gergely et al. 2015).</p><p>Fansipanica milae Páll-Gergely &amp; Dedov, gen. et sp. nov. was found at an altitude of ca 3040 m. The topography here is diversified: the mountain chains alternate with steep slopes and some relatively flat areas appearing like soil hills. The forest in the area of peak was protected from human activity by the long and high slopes, so that it appeared in pristine condition. At this altitude, in addition to the species from the families of Fagaceae and Lauraceae, Ericaceae ( Rhododendron) was present as well as Theaceae ( Ternstroemia) and Platanaceae ( Platanus). On lower slopes and relatively flat places, Fagaceae, Lauraceae, Platanaceae, Sterculiaceae, Elaeocarpaceae, Magnoliaceae, Araliaceae, Aceraceae, Betulaceae, Ericaceae, Theaceae, Rhodoleiaceae, Cupressaceae (young trees), Verbenaceae, Rosaceae, Juglandaceae, and Aquifoliaceae dominate. One specific characteristic of this mountain assemblage is the bamboo forest (cf. Arundinaria) under the canopy. On the mountain chains and steep slopes, Theaceae, Ericaceae, Magnoliaceae, Rhodoleiaceae, Lauraceae, Cupressaceae, Aceraceae, Betulaceae, and Rosaceae families are present. The epiphytic plants include Orchidaceae, Lycopodiaceae, Polypodiaceae, Aspleniaceae, Davalliaceae, Hymenophyllaceae, Vittariaceae, Lepidopteridaceae, Gesneriaceae, Ericaceae, Bryophyta, and also Rosaceae and Araliaceae (Sobey 1997) .</p></div>	https://treatment.plazi.org/id/A15685B371C85B0589F4D2F2D0EA4A36	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Dedov, Ivailo	Páll-Gergely, Barna, Gojšina, Vukašin, Dedov, Ivailo (2026): New taxa of Camaenidae from northern Vietnam (Gastropoda, Stylommatophora). ZooKeys 1280: 365-392, DOI: 10.3897/zookeys.1280.185160
460A3FC51D1F50979A53EECC9EF96CC5.text	460A3FC51D1F50979A53EECC9EF96CC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fansipanica Pall-Gergely & Dedov 2026	<div><p>Fansipanica Páll-Gergely &amp; Dedov gen. nov.</p><p>Type species.</p><p>Fansipanica milae sp. nov.</p><p>Diagnosis.</p><p>This camaenid genus is characterised by its small (~ 1 cm), globular, brownish, hairy shell. Genitalia with a long a cylindrical penis whose apical part covered by a penial sheath, a conical penial verge, a slender epiphallus (as slender as the vas deferens) and a very small, rounded flagellum.</p><p>Differential diagnosis.</p><p>There are several genera in continental Southeast Asia possessing hairy / scaly / pitted protoconchs and mostly hairy shells: Bellatrachia Schileyko, 2018, Bouchetcamaena Thach, 2018, Burmochloritis Godwin-Austen, 1920, Chloritis H. Beck, 1837, and Trichochloritis Pilsbry, 1891 . They all differ from the new genus in their reproductive anatomy. Namely, Bellatrachia has a large, fleshy flagellum (Schileyko 2018; Páll-Gergely and Neubert 2019), Bouchetcamaena has a very long flagellum (Páll-Gergely et al. 2022), Burmochloritis has a penial caecum, a vermiform flagellum and a blind-ending organ deriving from the wall of vagina (Godwin-Austen 1920; Páll-Gergely et al. 2023), continental Chloritis species have a more strongly developed flagellum and lack a penial sheath (Sutcharit and Panha 2010; Páll-Gergely et al. 2020), and Trichochloritis has a penial caecum (Páll-Gergely and Neubert 2019). Furthermore, in all these genera the vas deferens is much thinner than the epiphallus, while in Fansipanica gen. nov. the vas deferens and the epiphallus have approximately the same diameter.</p><p>Etymology.</p><p>The new genus is named after the Fansipan Peak, the type locality of the new taxon. Genus feminine.</p><p>Remarks.</p><p>The key traits of this new genus are the penial sheath on the proximal (apical) part of the penis, and the epiphallus, which is as slender as the vas deferens. The epiphallus is much thicker than the vas deferens in all Camaenidae, and their boundary is clearly visible in the abrupt change of diameter (Schileyko 2003). In this new genus, however, only the small and blunt flagellum indicates the transition between the epiphallus and the vas deferens (besides differences of the inner surface, see Fig. 3).</p><p>The penial sheath is probably not homologous with that of other genera of Camaeninae, covering mostly the distal (basal) part of the penis. However, it may be homologous with the thin fibrous capsule covering the proximal part of the penis of Bouchetcamaena platytropis (Möllendorff, 1894) (see Páll-Gergely et al. 2022).</p></div>	https://treatment.plazi.org/id/460A3FC51D1F50979A53EECC9EF96CC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Dedov, Ivailo	Páll-Gergely, Barna, Gojšina, Vukašin, Dedov, Ivailo (2026): New taxa of Camaenidae from northern Vietnam (Gastropoda, Stylommatophora). ZooKeys 1280: 365-392, DOI: 10.3897/zookeys.1280.185160
38565B8C161350ADBE5F47287A89B3CE.text	38565B8C161350ADBE5F47287A89B3CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vinatachea delsaerdti (Thach & F. Huber 2018) Páll-Gergely & Gojšina & Dedov 2026	<div><p>Vinatachea delsaerdti (Thach &amp; F. Huber, 2018) comb. nov.</p><p>Camaena delsaerdti Thach &amp; F. Huber, in Thach, 2018: 67–68, figs 889–892.</p><p>Remarks.</p><p>This species and its subspecies Camaena delsaerdti melanica Thach &amp; F. Huber, 2018 are moved to Vinatachea due to their similarity with the type species of the genus, V. porcellana sp. nov. The holotype of the second subspecies, C. delsaerdti aurantia Thach &amp; F. Huber, 2020 is a juvenile, corroded shell, which lacks the important traits for classification; therefore, it is considered a taxon inquirendum.</p></div>	https://treatment.plazi.org/id/38565B8C161350ADBE5F47287A89B3CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Dedov, Ivailo	Páll-Gergely, Barna, Gojšina, Vukašin, Dedov, Ivailo (2026): New taxa of Camaenidae from northern Vietnam (Gastropoda, Stylommatophora). ZooKeys 1280: 365-392, DOI: 10.3897/zookeys.1280.185160
FA909BD6E6A359E8A13CF69CD632F1E5.text	FA909BD6E6A359E8A13CF69CD632F1E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vinatachea porcellana Pall-Gergely 2026	<div><p>Vinatachea (?) porcellana Páll-Gergely sp. nov.</p><p>Figs 10, 11, 12, 13</p><p>Type material.</p><p>Holotype • Vietnam, Nghe An Province, Con Cuong District, Anh Son District, Hoi Son commune, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.04384&amp;materialsCitation.latitude=18.951979" title="Search Plazi for locations around (long 105.04384/lat 18.951979)">Pu Huong Nature Reserve</a>, 18.951979°N, 105.043838°E, ca 28 m, J. U. Otani leg., 10 May 2008 (empty shell and body in ethanol), IEBR_LS_Ngheania 001 H .</p><p>Description.</p><p>Shell. The shell is large, thick walled, overall glossy, nearly smooth. The shell shape is helicoid, with a blunt keel on the periphery. The shell colour is complex. The basic colour is ochre, and there is a slender, dark brown belt running just above the keel. Below the brown belt, there is a white belt, which is ~ 2 × thicker than the brown one. On the dorsal side, there is a thin, white belt above the suture, and a thin brown belt below the suture. The base colour of the ventral side is slightly paler than that of the dorsal surface. There is a thick (~ 1 mm), dark brown band on the ventral side, with two much more slender brown bands in both directions. The inner side of umbilicus is chocolate brown. The protoconch consists of ~ 1.75 whorls, and is rather glossy, with irregular fine wrinkles and spiral grooves. The sculpture of the teleoconch is dominated by extremely fine spiral grooves and fine growth lines. The entire shell consists of 4.75 whorls separated by a shallow suture. The aperture is semilunar, the parietal side is nearly straight. The peristome is discontinuous, strongly expanded, not reflected, porcelain white in colour, although there are three chocolate brown spiral bands on the ventral side, while the umbilical colour is also visible. The parietal callus is indicated by a transparent, thin calcareous layer, making the surface matte. The umbilicus is narrow, shows all whorls, and is partly covered by the reflected peristome.</p><p>Measurements.</p><p>D = 31.3 mm, H = 20.7 mm (n = 1).</p><p>Description of the genital organs.</p><p>The right ommatophoral retractor crosses the penis and the vagina. The atrium is short and thick. The penis is pear-shaped, consists of a slimmer distal (= closer to the genital opening) and a thickened proximal portion. The latter bears a slight central depression. The internally widened part of penis has numerous elongated, small papillae (some of the filiform), which are arranged in longitudinal folds that are converging towards atrium. The penial verge and the flagellum are absent. A slender duct, probably homologous with the epiphallus, starts from thickened part of penis, just from the base of the long and slender retractor muscle. The duct from the penis to the spermoviduct (probably epiphallus + vas deferens) is long, its distal part is slender, tapers until the middle, when it reaches maximum diameter, and remains thick until reaching the spermoviduct. The vagina is strongly developed, overall thick, attached to body wall with several fibres, with externally identifiable, slightly thinner distal, and slightly thicker proximal portion. The thinner part internally bears finely reticulate sculpture, while the inner wall of the thicker part is ornamented by ~ 10 longitudinal folds. The bursa copulatrix has a very slender, long stalk, originating from the insertion point of vas deferens, and a thickened, oval-rounded bursa. The spermoviduct and the albumen gland are very large.</p><p>Differential diagnosis.</p><p>Vinatachea delsaerdti (Thach &amp; F. Huber, 2018), comb. nov. is probably the most similar species. However, it has a stronger keel, possesses a single, slender brown belt on its base, and has a much rougher sculpture, most importantly with much more widely-spaced and deeper spiral grooves. The other three species of Vinatachea ( V. anhi Thach, 2025, V. khoai Thach, 2026, and V. thienanae Thach, 2025) have much more and densely arranged, usually slender spiral bands. Neotrachia duporti (Bavay &amp; Dautzenberg, 1909) is larger with a blunter keel, and its colouration is also different (yellowish-greenish base with pale brown or reddish spiral lines). Camaena gabriellae (Dautzenberg &amp; d’Hamonville, 1887) has a rougher surface sculpture, a spirally striated, but also rather glossy protoconch, a rounded body whorl, a mostly unicoloured base and a narrower umbilicus. Camaena marmorivaga (Mabille, 1889) is larger, has more elevated spire, blunter keel, and open umbilicus. On the other hand, it also belongs to the new genus because it has a smooth, glossy protoconch. Camaena choboensis (Mabille, 1889) is much larger than the new species, and has different colouration (greenish base with pale brown stripes).</p><p>Etymology.</p><p>The specific epithet porcellana refers to the porcelain-like glossy surface of the shell.</p><p>Remarks.</p><p>Three Peninsular Malaysian endemic species have been classified in the genus Kenyirus Clements &amp; S. K. Tan, 2012 . The reproductive anatomy is not known in any of them. Kenyirus balingensis S. K. Tan, S. Y. Chan &amp; Foon, 2017 and K. sheema Foon, S. K. Tan &amp; Clements, 2015 differ from the type species, K. sodhii Clements &amp; S. K. Tan, 2012 considerably, suggesting that they may belong to different genera. Kenyirus sodhii is superficially similar to Vinatachea porcellana, but it differs in the strongly keeled body whorl, the subtriangular peristome, and the spout-like rostrum. Moreover, according to the original description, the protoconch of K. sodhii bears axial striae, while that of Vinatachea porcellana has spiral grooves.</p></div>	https://treatment.plazi.org/id/FA909BD6E6A359E8A13CF69CD632F1E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Dedov, Ivailo	Páll-Gergely, Barna, Gojšina, Vukašin, Dedov, Ivailo (2026): New taxa of Camaenidae from northern Vietnam (Gastropoda, Stylommatophora). ZooKeys 1280: 365-392, DOI: 10.3897/zookeys.1280.185160
2B2B067DF08D5CF9950BB06CF7396DF2.text	2B2B067DF08D5CF9950BB06CF7396DF2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vinatachea Thach 2025	<div><p>Vinatachea Thach, 2025</p><p>Vinatachea Thach, 2025: 83 .</p><p>Type species.</p><p>Vinatachea thienanae Thach, 2025 (by original designation).</p><p>Diagnosis.</p><p>The short, blunt penis, the epiphallus, which is very slender when connected to the penis, the absence of the flagellum and a penial caecum and the strongly thickened vas deferens distinguishes this genus from all other camaenid genera in Southeast Asia.</p><p>Remarks.</p><p>The anatomy of the type species is unknown. Vinatachea porcellana sp. nov. is classified in this genus provisionally, due to the similar shell size and shape, and the geographic proximity.</p></div>	https://treatment.plazi.org/id/2B2B067DF08D5CF9950BB06CF7396DF2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Páll-Gergely, Barna;Gojšina, Vukašin;Dedov, Ivailo	Páll-Gergely, Barna, Gojšina, Vukašin, Dedov, Ivailo (2026): New taxa of Camaenidae from northern Vietnam (Gastropoda, Stylommatophora). ZooKeys 1280: 365-392, DOI: 10.3897/zookeys.1280.185160
