taxonID	type	description	language	source
2C2F878FFFA7C8127FCFCFB7B9216BDC.taxon	materials_examined	Type: — CHINA. Yunnan: Malipo, Xiajinchang Xiang, Yunling Cun, Liangzishang, 23 ° 0 ’ 22 ” N, 104 ° 49 ’ 39 ” E, elev. 1800 m, on wet rocks along a stream near road, 11 December 2015, Ke-Wang Xu, Long-Yuan Wang & Hui-Zhe Feng TTJ-YN- 030 (holotype SYS!, isotypes MO!, SYS!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA7C8127FCFCFB7B9216BDC.taxon	diagnosis	Diagnosis: — Hymenasplenium chingii is somewhat similar to H. hondoense (Murakami & Hatanaka 1988: 191; Nakaike 1992: 841) in having pinnae falcate or trapeziform and teeth at base of the pinnae often slightly retuse, but the former has pinna apex acute or acuminate and sori supra-medial to (sub) marginal, while the latter has pinna apex subacute to obtuse and sori medial.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA7C8127FCFCFB7B9216BDC.taxon	description	Plants perennial, evergreen, 20 – 38 cm tall. Rhizomes long creeping, ca. 2.5 mm in diam., apex scaly, scales dark brown, narrowly triangular or lanceolate, margins nearly entire, (0.1 –) 0.4 × 1 – 1.2 mm; root yellowish brown when dried, slender, up to 8 cm long, ca. 0.25 mm diam, densely covered with yellow brown woolly indument or subglabrous. Fronds remote, 10 – 12 mm apart, herbaceous; petioles shiny, dark brown, 6 – 15 cm long, ca. 1 mm in diam., base near rhizome sparsely covered with scales, scales narrowly triangular, dark brown, margins nearly entire, 0.2 × 0.5 – 0.8 mm, and densely covered with yellow brown woolly indument. Laminae narrowly ovate, 1 - pinnate, 12 – 22 × 3.5 – 6 cm, widest near base, gradually narrowing toward apex, apex acute; rachises 0.5 – 1 mm in diam., glabrous, shiny, dark brown, adaxial side grooved and with 2 green narrow wings. Pinnae shortly stalked to almost sessile at apex, 15 – 30 pairs, alternate, falcate or trapeziform, (middle pairs) 1.5 – 3 × 0.4 – 0.7 cm, about 3 – 6 mm apart, base asymmetrical, acroscopic side truncate and subparallel or forming an angle less than 15 ° with rachis, basiscopic side cuneate, with ca. 10 mm absent (calculated from the base to the first tooth appearing in the margin of basiscopic side), acroscopic margins serrate, teeth acute or obtuse, usually retuse at basal portion of pinnae, basiscopic margins entire at base, serrate in the middle and apex; pinna apex acuminate. Veins visible on both sides of pinnae, stramineous, rarely brown near pinnae petioles, lateral veins visible abaxially but obscure adaxially, forking and terminating in marginal teeth or below notches, 2 – 4 basal basiscopic veins lacking. Sori linear, 2 – 3 (– 4) mm long, medial, (1 –) 5 – 7 on acroscopic side and (1 –) 4 – 6 on distal basiscopic side of the pinnae in the middle part of the laminae, centers 1 – 2 (– 5) mm apart from one another; indusia brown, linear, membranous, entire, the indusium of the first sorus on the acroscopic side of pinnae opening toward rachis or absent, other indusia opening toward costa.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA7C8127FCFCFB7B9216BDC.taxon	distribution	Geographical distribution: — Hymenasplenium chingii is found in Guizhou and Yunnan, Southwest China.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA7C8127FCFCFB7B9216BDC.taxon	biology_ecology	Ecology: — Hymenasplenium chingii was observed to grow on wet rocks along streams with shady conditions under broad-leaved forest.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA7C8127FCFCFB7B9216BDC.taxon	etymology	Etymology: — The species epithet is in honor of the late professor Ren-Chang Ching based at PE for his great contributions to Chinese pteridology in general and to the taxonomy of Hymenasplenium in particular (Ching 1965, Ching & Liu 1983).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA7C8127FCFCFB7B9216BDC.taxon	vernacular_names	Vernacular name: — Àkůďäffǎ (qin shi mo ye tie jiao jue).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA7C8127FCFCFB7B9216BDC.taxon	discussion	Taxonomic notes: — Morphologically, Hymenasplenium chingii is similar to H. hondoense (see above). Phylogenetically, H. chingii was resolved as sister to a clade containing H. hondoense, H. wangpeishanii, and H. speluncicola (Xu et al. 2018 a). Our large sampling in the molecular analysis suggested that H. hondoense, with its type from Honshu, Japan, probably does not occur in China, contrasting the report of the latter in China by Lin & Viane (2013).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA7C8127FCFCFB7B9216BDC.taxon	materials_examined	Additional material examined: — CHINA. Yunnan: Gongshan, Cikai, Qiqigou, elev. 1550 m, 13 August 2011, Xiaohua Jin & Liang Zhang 11347 (CDBI!); Guizhou: Nayong, Yangchang Zhen, Qinglin Cun, Black Cave, 26 ° 41 ’ 3 ” N, 105 ° 10 ’ 27 ” E, elev. 1570 m, 23 August 2017, Li Bing Zhang, Yi-Fan Duan & Matthias Kropf 9780 (CDBI!); Guizhou: Libo, Xiaoqikong Zhen, Chuandong, 25 ° 43 ’ 20 ” N, 107 ° 56 ’ 46 ” E, elev. 950 m, 3 August 2017, Li Bing Zhang, Yi-Fan Duan & Matthias Kropf 9455 (CDBI!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA2C8107FCFCB00B8F56D7B.taxon	materials_examined	Type: — CHINA. Guizhou: Daozhen County, Baijishan, elev. 750 m, on wet palisades near streams, 16 October 15, Zhengyu Liu 16082 (holotype MO! isotype MO!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA2C8107FCFCB00B8F56D7B.taxon	diagnosis	Diagnosis: — Hymenasplenium denticulatum is most similar to H. murakami-hatanakae Nakaike (1992: 841) in having pinnae falcate to trapeziform and usually ascending, sori supramedial to (sub) marginal, but the former has plants 30 – 50 cm tall, marginal teeth of the pinnae shallow (0.5 – 1 mm), dense, and usually shallowly retuse, and veins usually terminating in the notches, or terminating in the marginal teeth, while the latter has plants 25 – 40 cm tall, marginal teeth of the pinnae deep (1 mm at base), sparse, acute, not retuse, and veins terminating in marginal teeth.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA2C8107FCFCB00B8F56D7B.taxon	description	Plants perennial, evergreen, 30 – 50 cm tall. Rhizomes long creeping, ca. 3 mm in diam., apex scaly, scales brown, narrowly triangular, margins nearly entire, 1 × 0.25 mm; root brown when dried, slender, up to 4 cm long, rhizome and root glabrous or covered with yellowish brown woolly indument. Fronds remote, 10 – 20 mm apart, herbaceous; petioles dark brown, glabrous, 15 – 25 cm long. Laminae narrowly ovate, 1 - pinnate, 15 – 25 × 4 – 6 cm, base deflexed, slightly reduced, widest near middle, gradually narrowing toward apex, apex acuminate; rachises 1 mm in diam., glabrous, adaxial side grooved and with 2 green narrow wings. Pinnae shortly stalked to almost sessile at apex, 20 – 30 pairs, subopposite or alternate, falcate to trapeziform, (middle pairs) 2 – 3.5 × 0.6 – 1 cm, up to 3 – 5 mm apart, base asymmetrical, acroscopic side truncate and subparallel or overlapping with rachis, with ca. 10 mm absent (calculated from the base to the first tooth appearing in the margin of basiscopic side), acroscopic margins serrate, teeth shallow and dense, usually retuse, basiscopic margins deep and sparse, teeth obtuse and not retuse, apex acute or acuminate. Veins visible on both sides of pinnae, stramineous, lateral veins visible on both sides of pinnae, green, forking and terminating in notches or marginal teeth, 2 – 3 basal basiscopic veins lacking. Sori linear, 1 – 3 mm long, supramedial to (sub) marginal, (2 –) 6 – 8 on acroscopic side and 4 – 6 (– 8) on distal basiscopic side of the pinnae in the middle part of the laminae, centers 1 – 2 mm apart from one another; indusia brown, linear, membranous, entire, opening toward costa.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA2C8107FCFCB00B8F56D7B.taxon	distribution	Geographical distribution: — Hymenasplenium denticulatum is currently only known from the adjacent Chongqing and northern Guizhou. It is likely endemic to Southwest China.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA2C8107FCFCB00B8F56D7B.taxon	biology_ecology	Ecology: — Hymenasplenium denticulatum was observed to grow on wet palisades near streams.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA2C8107FCFCB00B8F56D7B.taxon	etymology	Etymology: — The species epithet is based on the Latin adjective, denticulatum, denticulate, referring to the denticulate pinna margins of the new species.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA2C8107FCFCB00B8F56D7B.taxon	vernacular_names	Vernacular name: — àŵůďäffǎ (xi chi mo ye tie jiao jue).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA2C8107FCFCB00B8F56D7B.taxon	discussion	Taxonomic notes: — Hymenasplenium denticulatum can be easily distinguished by its shallow and dense marginal teeth, usually retuse at apex, and supramedial to (sub) marginal sori. In addition, it is likely to be endemic to Southwest China, while the morphologically similar species H. murakami-hatanakae is only distributed in Japan, Taiwan Island based on our large-scale phylogenetic study (Xu et al. 2018 a).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA2C8107FCFCB00B8F56D7B.taxon	materials_examined	Additional material examined: — CHINA. Chongqing: Nanchuan, Jinfoshan Mt., elev. 800 m, 5 April 1996, Zhengyu Liu 15536 (MO!); 14 October 1995, Sirong Yi 15799 (MO!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA0C81F7FCFCE5FB9056FF6.taxon	materials_examined	Type: — VIETNAM. Hoa Binh: Ngoc Son Ngo Luong Nature Reserve, 26 ° 25 ’ 53 ” N, 106 ° 2 ’ 30 ” E, elev. 1350 m, 20 November 2013, Li Bing Zhang, Liang Zhang & Lu Thi Ngan 6296 (holotype CDBI!, isotypes MO!, VNMN!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA0C81F7FCFCE5FB9056FF6.taxon	diagnosis	Diagnosis: — Hymenasplenium distans is most similar to H. apogamum (Murakami & Hatanaka 1988: 193; Nakaike 1992: 841) in having pinnae quadrangular-trapeziform in outline, entire on tooth margins, spreading or deflexed at base, obtuse at apex, usually crispate at base auricle, but the former has petioles black, fronds up to 15 mm apart, and the ratio of laminae to petioles near 0.5, while the latter has petioles brown, fronds 2 – 3 mm apart, and the ratio of laminae to petioles near 1.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA0C81F7FCFCE5FB9056FF6.taxon	description	Plants perennial, evergreen, 35 – 40 cm tall. Rhizomes short creeping, ca. 1 – 2 mm in diam., apex scaly, scales dark brown, narrowly triangular, margins nearly entire, 0.5 – 1 × 2.5 – 3.5 mm; root stramineous when dried, up to 8 cm long, ca. 1 mm in diam, with orange-brown woolly indument or subglabrous. Fronds remote, up to 15 mm apart, herbaceous; petioles shiny, black, 15 – 26 cm long, ca. 1.2 mm in diam., base sparsely covered with scales similar to those on rhizome, or subglabrous. Laminae ovate, 1 - pinnate, 14 – 18 × 4.5 – 8 cm, widest near base, gradually narrowing toward apex, apex subacute; rachises ca. 1 mm in diam., subglabrous, shiny and black abaxially, adaxial side grooved and with 2 grayish green narrow wings. Pinnae almost sessile to shortly stalked with rachis color extending onto the base of costa abaxially, 15 – 25 pairs, opposite or subopposite at the base of laminae, alternate in the middle and apex, lower pinnae deflexed toward stipe, quadrangular-trapeziform, (middle pairs) 3 – 3.5 × 0.7 – 1 cm, about 0 – 2 mm apart, base asymmetrical, acroscopic side truncate and subparallel or forming an angle less than 15 ° with rachis, subauriculate, basiscopic side cuneate, with 5 – 15 mm absent (calculated from the base to the first tooth appearing in the margin of basiscopic side), acroscopic margins sinuate to serrate, teeth acute, not retuse, basiscopic margins entire at base, sinuate to serrate in the middle and apex; pinna apex obtuse to subacute. Veins visible on both sides of pinnae, dark brown at base of abaxial side of pinnae, lateral veins visible abaxially but obscure adaxially, forking and terminating in marginal teeth, only 1 or 2 basal basiscopic veins lacking. Sori linear, (2 –) 4 – 6 mm long, medial, 8 – 15 on acroscopic side and 5 – 10 on distal basiscopic side of the pinnae in the middle part of the laminae, centers 1 – 3 (– 5) mm apart from one another; indusia brownish, linear, membranous, entire, the indusium of the second sorus on the acroscopic side of pinnae opening toward rachis or absent, other indusia opening toward costa.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA0C81F7FCFCE5FB9056FF6.taxon	distribution	Geographical distribution: — Hymenasplenium distans is currently only known from Hoa Binh and Thanh Hoa, northern Vietnam. It is likely endemic to Vietnam.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA0C81F7FCFCE5FB9056FF6.taxon	biology_ecology	Ecology: — Hymenasplenium distans was observed to grow on wet rock in seasonal broad-leaved closed lowland forest.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA0C81F7FCFCE5FB9056FF6.taxon	etymology	Etymology: — The species epithet is based on the Latin adjective, distans, distant, referring to the relatively distant space on the rhizome between fronds of the new species.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA0C81F7FCFCE5FB9056FF6.taxon	vernacular_names	Vernacular name: — ãďůďäffǎ (yuan ye mo ye tie jiao jue).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA0C81F7FCFCE5FB9056FF6.taxon	discussion	Taxonomic notes: — Hymenasplenium distans can be easily distinguished from other members of the H. unilaterale s. l. group by its quadrangular-trapeziform pinnae and long black stipe and 10 mm distance between neighboring fronds. The most similar species H. apogamum is widely distributed in Southeast Asia, while the new species is only found in North Vietnam. In our recent phylogeny H. distans formed a strongly supported clade with H. apogamum and H. ngheanense (described below).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA0C81F7FCFCE5FB9056FF6.taxon	materials_examined	Additional material examined: — VIETNAM. Thanh Hoa: Quan Hoa District, Phu Le Municipality, Hang village, 20 ° 31 ’ 33 ” N, 105 ° 05 ’ 06 ” E, elev. 350 – 500 m, seasonal broad-leaved closed lowland forest on very steep slopes of rocky ridge composed with crystalline marble-like highly eroded limestone, 30 September 2003, L. Averyanov, P. K. Loc, D. T. Doan & N. T. Vinh 3623 (MO!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC81F7FCFCF26BF2C6956.taxon	materials_examined	Type: — VIETNAM. Ngh ệ An: Con Cuong District, Chau Khe Commune, Khe Choang: Pu Mat National Park, 18 ° 57 ’ 21.70 ” N, 104 ° 38 ’ 39.62 ” E, elev. 400 m, 28 October 2014, Li Bing Zhang, Liang Zhang & Lu Thi Ngan 7163 (holotype CDBI!, isotypes MO!, VNMN!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC81F7FCFCF26BF2C6956.taxon	diagnosis	Diagnosis: — Hymenasplenium ngheanense is most similar to H. apogamum in having stipes brown and pinnae spreading or deflexed at base, entire at tooth margins, and usually crispate at base auricle, but the former has pinnae falcate or lanceolate, pinna apex acute or acuminate, while the latter has pinnae quadrangular-trapeziform, pinna apex obtuse to subacute.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC81F7FCFCF26BF2C6956.taxon	description	Plants perennial, evergreen, 22 – 30 cm tall. Rhizomes short creeping, ca. 1 – 1.5 mm in diam., apex scaly, scales dark brown, lanceolate, margins nearly entire, 0.2 – 0.5 × 1.5 – 1.8 mm; root yellowish brown when dried, up to 8 cm long, ca. 1 mm diam, with yellow woolly indument or subglabrous. Fronds remote, 5 – 8 mm apart, herbaceous; petioles shiny, dark brown, 10 – 15 cm long, ca. 1 mm in diam., base sparsely covered with scales, scales lanceolate, castaneous, margins nearly entire, 0.2 × 1 mm, and covered with yellow woolly indument, or subglabrous. Laminae narrowly ovate or rarely narrowly triangular, 1 - pinnate, 10 – 20 × 4.5 – 6 cm, widest near base, gradually narrowing toward apex, apex acuminate; rachises 0.5 – 1 mm in diam., subglabrous, shiny and castaneous abaxially, adaxial side grooved and with 2 green narrow wings. Pinnae almost sessile to shortly stalked, 12 – 20 pairs, alternate, lower pinnae deflexed toward stipe, falcate or lanceolate, (middle pairs) 2 – 3 × 0.4 – 0.8 cm, about 1 – 0.6 mm apart, base asymmetrical, acroscopic side truncate and subparallel or forming an angle less than 15 ° with rachis, subauriculate, basiscopic side cuneate, with 4 – 12 mm absent (calculated from the base to the first tooth appearing in the margin of basiscopic side), acroscopic margins sinuate to serrate, teeth shallow and obtuse, not retuse, basiscopic margins entire at base, sinuate to serrate in the middle and apex; pinnae apex acute or acuminate. Veins visible on both sides of pinnae, green, rarely brown near pinnae petioles, lateral veins visible abaxially but obscure adaxially, forking and terminating in marginal teeth, only 1 or 2 basal basiscopic veins lacking. Sori linear, 2 – 4 (– 5) mm long, medial, (1 –) 4 – 8 (– 10) on acroscopic side and (1 –) 3 – 6 on distal basiscopic side of the pinnae in the middle part of the laminae, centers 1.5 – 2 (– 5) mm apart from one another; indusia stramineous, linear, membranous, entire, the indusium of the second sorus on the acroscopic side of pinnae opening toward rachis or absent, other indusia opening toward costa.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC81F7FCFCF26BF2C6956.taxon	distribution	Geographical distribution: — Hymenasplenium ngheanense is currently only known from central and northern Vietnam.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC81F7FCFCF26BF2C6956.taxon	biology_ecology	Ecology: — Hymenasplenium ngheanense was observed to grow on wet rocks in humid and shady conditions of broad-leaved forest.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC81F7FCFCF26BF2C6956.taxon	etymology	Etymology: — The species epithet is based on the name of Vietnamese province Ngh ệ An, and the Latin suffix, - ense, of origin, referring to the type locality of the new species.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC81F7FCFCF26BF2C6956.taxon	vernacular_names	Vernacular name: — XȐůďäffǎ (yi an mo ye tie jiao jue).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC81F7FCFCF26BF2C6956.taxon	discussion	Taxonomic notes: — Hymenasplenium ngheanense, H. apogamum, and H. distans share some morphological similarities such as pinnae spreading or deflexed at base, entire at tooth margins, and usually crispate at base auricle. However, they are strongly supported as distinct lineages in our molecular study (Xu et al. 2018 a).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC81F7FCFCF26BF2C6956.taxon	materials_examined	Additional material examined: — VIETNAM. Phu Tho: Tan Son District, Xuan Son National Park, 26 November 2013, Li Bing Zhang, Liang Zhang & Lu Thi Ngan 6532 (CDBI!, MO!, and VNMN!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC8187FCFCA86BE446C0E.taxon	materials_examined	Type: — VIETNAM. Khánh Hòa: Hòn Bà Nature Reserve, trail at 32 nd km of main road toward Yersin’s house, 12 ° 01 ’ 45 ” – 12 ° 12 ’ 00 ” N, 102 ° 53 ’ 45 ” – 109 ° 02 ’ 34 ” E, elev. 1100 m, 26 October 2015, Li Bing Zhang, Xinmao Zhou, Thien Tam Luong, Minh Tri Dang, Bich Ha Duong 8819 (holotype CDBI!, isotypes MO!, PHH!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC8187FCFCA86BE446C0E.taxon	diagnosis	Diagnosis: — Hymenasplenium phamhoanghoi is most similar to H. unilaterale (Lamark 1786: 305; Hayata 1927: 712) in having pinna teeth entire, veins terminating in the marginal teeth and reaching nearly apex of teeth, but the former has pinnae rectangular or trapeziform, pinna apex obtuse or acute, and bases of acroscopic pinnae with auricles, while the latter has pinnae falcate or trapeziform, pinna apex acute or acuminate, and bases of acroscopic pinnae without auricles.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC8187FCFCA86BE446C0E.taxon	description	Plants perennial, evergreen, up to 40 cm tall. Rhizomes long creeping, ca. 4 mm in diam., apex scaly, scales dark brown, narrowly triangular or lanceolate, margins nearly entire, 1 – 3 × 0.3 – 0.5 mm; root dark brown when dried, slender, up to 9 cm long, ca. 0.2 mm in diam, rhizome and root often glabrous, rarely with sparsely brown woolly indument. Fronds remote, up to 20 mm apart, herbaceous; petioles shiny, dark brown, base near rhizome sparsely covered with scales, scales small, 0.1 × 0.4 mm, lanceolate, brown, margins nearly entire. Laminae narrowly ovate, 1 - pinnate, 15 – 25 × 4 – 6 cm, base slightly reduced, widest near base, gradually narrowing toward apex, apex acuminate; rachises 1 mm in diam., glabrous, shiny, dark brown, adaxial side grooved and with 2 green narrow wings. Pinnae shortly stalked to almost sessile at apex, 20 – 30 pairs, alternate, rectangular or trapeziform, (middle pairs) 2.5 – 3 × 0.5 – 0.8 cm, about 2 – 4 mm apart, base asymmetrical, acroscopic side truncate and subparallel or forming an angle less than 15 ° with rachis, basiscopic side cuneate, with 15 – 20 mm absent (calculated from the base to the first tooth appearing in the margin of basiscopic side), acroscopic margins serrate, teeth not retuse, acute or obtuse, basiscopic margins entire at base, serrate in the middle and apex; pinna apex acute or obtuse. Veins visible on both sides of pinnae, green, rarely brown near pinnae petioles, lateral veins visible on both sides of pinnae, green, forking and terminating in marginal teeth, 3 – 4 basal basiscopic veins lacking. Sori linear, 2 – 3 mm long, medial to supramedial, (1 –) 5 – 7 on acroscopic side and 1 – 3 on distal basiscopic side of the pinnae in the middle part of the laminae, centers 2 – 4 mm apart from one another; indusia brown, linear, membranous, entire, opening toward costa.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC8187FCFCA86BE446C0E.taxon	distribution	Geographical distribution: — Hymenasplenium phamhoanghoi is currently only known from Hòn Bà Nature Reserve, Khánh Hòa Province, southern Vietnam. It is likely endemic to Vietnam.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC8187FCFCA86BE446C0E.taxon	biology_ecology	Ecology: — Hymenasplenium phamhoanghoi was observed to grow in acidic soils under evergreen forest.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC8187FCFCA86BE446C0E.taxon	etymology	Etymology: — The species epithet is in honor of Prof. Ph ạm Hoàng H ộ for his great contributions to the fern flora of Vietnam (e. g., Pham 1991, 1999).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC8187FCFCA86BE446C0E.taxon	vernacular_names	Vernacular name: — ħflůďäffǎ (nan yue mo ye tie jiao jue).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFAFC8187FCFCA86BE446C0E.taxon	discussion	Taxonomic notes: — Morphologically, Hymenasplenium phamhoanghoi is different from any members of Hymenasplenium from Southeast Asia in having pinna apex obtuse, bases of acroscopic pinnae with auricles, pinna teeth entire, and veins terminating in the marginal teeth and reaching nearly apex of teeth. In our molecular analysis, H. phamhoanghoi formed a well supported clade with H. unilaterale and some unidentified material from Australia and Pacific islands. These relationships are supported by their similar characters of pinna teeth and veins.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA8C8067FCFCF68B9A46AD2.taxon	materials_examined	Type: — VIETNAM. Qu ảng Nam: Song Thanh Nature Reserve, Nam Giang District, Ta Bhing Community, Ta Ding Village, 15 ° 37 ’ 58.18 ” N, 107 ° 44 ’ 17.86 ” E, elev. 200 – 350 m, 4 December 2014, Li Bing Zhang, Liang Zhang & Ngan Thi Lu 7884 (holotype CDBI!, isotypes MO!, VNMN!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA8C8067FCFCF68B9A46AD2.taxon	diagnosis	Diagnosis: — Hymenasplenium quangnamense is most similar to H. apogamum in having rhizomes densely scaly, roots robust (ca. 1 mm in diam.), and pinnae quadrangular or trapeziform, but the former has marginal teeth at the base of pinnae often slightly retuse, veins terminating below notches, and sori supramedial to (sub) marginal, while the latter has marginal teeth at the base of pinnae entire, veins terminating in marginal teeth, and sori medial.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA8C8067FCFCF68B9A46AD2.taxon	description	Plants perennial, evergreen, 25 – 40 cm tall. Rhizomes creeping, ca. 4 mm in diam., apex densely scaly, scales black, narrowly triangular or lanceolate, margins nearly entire, 2 – 3.6 × 0.4 – 0.7 mm; root yellow or yellowish brown when dried, robust, up to 10 cm long, ca. 1 mm diam, rhizome and root sparsely with yellow woolly indument or glabrous. Fronds remote, up to 8 mm apart, herbaceous; petioles slender, shiny, dark brown, glabrous, 8 – 20 cm long, ca. 1 mm in diam., base near rhizome sparsely covered with scales, scales lanceolate, black, margins nearly entire, 0.1 – 0.2 × 2 – 3 mm. Laminae narrowly ovate, 1 - pinnate, 15 – 20 × 4.5 – 6.5 cm, base slightly reduced, widest near base, gradually narrowing toward apex, apex acute; rachises 0.5 – 1 mm in diam., glabrous, shiny, dark brown, adaxial side grooved and with 2 green narrow wings. Pinnae shortly stalked to almost sessile at apex, 15 – 20 pairs, alternate, quadrangular or trapeziform, (middle pairs) 2.5 – 3 × 0.5 – 0.8 cm, about 5 – 10 (– 15) mm apart, base asymmetrical, acroscopic side truncate and subparallel or forming an angle less than 15 ° with rachis, basiscopic side cuneate, with 10 – 15 mm absent (calculated from the base to the first tooth appearing in the margin of basiscopic side), acroscopic margins serrate, teeth not retuse, acute or obtuse, basiscopic margins entire at base, serrate in the middle and apex; pinna apex obtuse. Veins visible on both sides of pinnae, stramineous to green, rarely brown near pinnae petioles, lateral veins visible on both sides of pinnae, green, forking and terminating in marginal teeth, 2 – 3 basal basiscopic veins lacking. Sori linear, 2 – 5 mm long, supramedial to (sub) marginal, (1 –) 9 – 11 on acroscopic side and (2 –) 5 – 7 on distal basiscopic side of the pinnae in the middle part of the laminae, centers 1 – 2 (– 4) mm apart from one another; indusia brown, linear, membranous, entire, the indusium of the second sorus on the acroscopic side of pinnae opening toward rachis or absent, others opening toward costa.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA8C8067FCFCF68B9A46AD2.taxon	distribution	Geographical distribution: — Hymenasplenium quangnamense is currently only known from Qu ảng Nam Province, central Vietnam. It is likely endemic to Vietnam.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA8C8067FCFCF68B9A46AD2.taxon	biology_ecology	Ecology: — Hymenasplenium quangnamense was observed to grow on rocks cave at an elevation of ca. 300 m with humid and shady conditions.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA8C8067FCFCF68B9A46AD2.taxon	etymology	Etymology: — The species epithet is based on the name of Vietnamese province Qu ảng Nam, and the Latin suffix, - ense, of origin, referring to the type locality of the new species.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA8C8067FCFCF68B9A46AD2.taxon	vernacular_names	Vernacular name: — Ḟħůďäffǎ (guang nan mo ye tie jiao jue).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFA8C8067FCFCF68B9A46AD2.taxon	discussion	Taxonomic notes: — Hymenasplenium quangnamense is morphologically similar to H. apogamum. Our molecular analysis (Xu et al. 2018 a) did not well resolve its relationships but it had a very long branch length on the phylogenetic tree showing its isolated relationships in the genus.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB6C8057FCFC805BF6F6D4B.taxon	materials_examined	Type: — CHINA. Guizhou: Songtao Xian, Lengjiaba, near the confluence of the Xiaohe and Dahe rivers, Northeast Fanjingshan Mountain range, elev. 820 – 1120 m, on wet rocks and on rock ledges near river in forest, 5 – 9 October 1986, Sino-American Guizhou Botanical Expedition no. 2207 (holotype MO!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB6C8057FCFC805BF6F6D4B.taxon	diagnosis	Diagnosis: — Hymenasplenium sinense is most similar to H. obliquissimum (Hayata 1914: 230; Sugimoto 1966: 406) in having rhizome scales dense, fronds membranous and sub-translucent, and sori located in the middle or apex of pinnae, but the former has rhizome scales brown, pinnae 15 – 25 pairs, laminae ovate, and marginal teeth of the pinnae not retuse, and veins terminating in marginal teeth, while the latter has rhizome scales black, pinnae more than 25 pairs (often more than 30 pairs), laminae lanceolate, and marginal teeth of the pinnae retuse, and veins terminating below the tooth notches.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB6C8057FCFC805BF6F6D4B.taxon	description	Plants perennial, evergreen, 15 – 30 cm tall. Rhizomes long creeping, 1.5 – 2.5 mm in diam., apex densely scaly, scales grayish brown, narrowly triangular, margins nearly entire, 0.4 – 0.5 × 1 – 1.8 mm; root brown when dried, slender, up to 5 cm long, ca. 0.2 mm diam, root glabrous or covered with yellow woolly indumenta, phyllopodia distinct, ca. 2 mm tall, up to 10 mm apart. Fronds membranous, thin to translucent, brown-green when dry; petioles shiny, castaneous, glabrous. Laminae narrowly ovate, 1 - pinnate, 15 – 25 × 2 – 5 cm, base slightly reduced, widest near base, gradually narrowing toward apex, apex acute or acuminate; rachises 0.5 – 1 mm in diam., subglabrous, castaneous, adaxial side grooved and with 2 green narrow wings. Pinnae shortly stalked to almost sessile at apex, 15 – 25 pairs, subopposite or alternate, trapeziform or falcate, (middle pairs) 1.5 – 3 × 0.5 – 0.8 cm, up to 3 mm apart, base asymmetrical, acroscopic side truncate and subparallel or forming an angle less than 15 ° with rachis, basiscopic side cuneate, with ca. 10 mm absent (calculated from the base to the first tooth appearing in the margin of basiscopic side), acroscopic margins serrate, teeth not retuse, acute or obtuse, basiscopic margins crenate-sinuate, teeth obtuse and not retuse, apex subacute to obtuse. Veins visible on both sides of pinnae, stramineous, lateral veins visible on both sides of pinnae, green, forking and terminating in marginal teeth, 2 – 4 basal basiscopic veins lacking. Sori linear, 1 – 3 mm long, basal on subtending vein and appearing subcostal, 2 – 5 on acroscopic side and 2 – 3 on distal basiscopic side of the pinnae in the middle part of the laminae, centers 1 – 2 mm apart from one another; indusia brown, linear, membranous, entire, opening toward costa.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB6C8057FCFC805BF6F6D4B.taxon	distribution	Geographical distribution: — Hymenasplenium sinense is widely distributed in the central, southern, and southwestern China. It might also occur in Thailand. Further studies are needed for material from India, Indonesia, Japan, Nepal, and Vietnam.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB6C8057FCFC805BF6F6D4B.taxon	biology_ecology	Ecology: — Hymenasplenium sinense was observed to grow on wet rocks near or in streams in forests.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB6C8057FCFC805BF6F6D4B.taxon	etymology	Etymology: — The species epithet is based on the Latin, Sina, China, and the Latin suffix, - ense, of origin, referring to where the type was collected from and the current major known distribution of the species in China.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB6C8057FCFC805BF6F6D4B.taxon	vernacular_names	Vernacular name: — ϕṣůďäffǎ (zhong hua mo ye tie jiao jue).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB6C8057FCFC805BF6F6D4B.taxon	discussion	Taxonomic notes: — Due to the previous confusion with Hymenasplenium obliquissimum, H. sinense has not been known by modern pteridologists. Lin and Viane (2013) assigned material of this species to H. obliquissimum whose type is from Arisan, Taiwan. However, H. obliquissimum sensu Lin and Viane (2013) has different morphology from its type and should represent H. sinense in our delimitation.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB6C8057FCFC805BF6F6D4B.taxon	materials_examined	Additional material examined: — CHINA. Yunnan: Pingbian Xian, Daweishan, on wet rocks near stream, 29 October 2015, Xu PB 001 (SYS!); Guangxi: Rongshui Xian, Yuanbaoshan, 27 November 2015, Xu GX 018 (SYS!); Jiangxi: Suichuan Xian, Daijiapu Xiang, Qianmo Cun, 26 ° 17 ’ 39.57 ” N, 114 ° 3 ’ 45.24 ” E, 880 m, 17 May 2016, Kewang Xu 134 (SYS!); Jingangshan, Jingzhushan, 28 June 1965, Lai et al. 4281 (IBK!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB5C8027FCFCEB5BA5C6FD3.taxon	materials_examined	Type: — CHINA. Guangxi: Jingxi, Wuping Xiang, Wuping Street, Gongtong Big Cave, 23 ° 11 ’ 52.97 ” N, 106 ° 28 ’ 09.88 ” E, elev. 700 – 800 m, 31 October 2010, Li Bing Zhang, Hai He & Yu Wang 5494 (holotype CDBI!, isotypes CTC!, MO!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB5C8027FCFCEB5BA5C6FD3.taxon	diagnosis	Diagnosis: — Hymenasplenium speluncicola is morphologically similar to H. hondoense in having teeth at base of pinnae slightly retuse and sori medial, but the former has rhizomes densely covered with long yellowish brown woolly indumentum, petioles slender (ca. 1 mm in diam.), pinnae falcate or lanceolate and acuminate, while the latter has rhizomes densely covered with long yellow brown woolly indumentum, petioles thick (ca. 1.5 mm in diam.), pinnae trapeziform to falcate and subacute to obtuse at apex.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB5C8027FCFCEB5BA5C6FD3.taxon	description	Plants perennial, evergreen, 20 – 40 cm tall. Rhizomes long creeping, ca. 2 mm in diam., apex scaly, scales dark brown, narrowly triangular or lanceolate, margins nearly entire, 0.2 – 0.4 × 0.8 – 1.5 mm; root yellowish brown when dried, slender, up to 6 cm long, ca. 0.2 mm diam, rhizome and root densely covered with long yellow brown woolly indument. Fronds remote, 10 – 20 mm apart, herbaceous; petioles slender, shiny, dark brown, glabrous, 15 – 25 cm long, ca. 1 mm in diam. Laminae narrowly ovate, 1 - pinnate, 15 – 20 × 3.5 – 7 cm, base reduced, widest near base, gradually narrowing toward apex, apex acuminate; rachises 0.5 mm in diam., glabrous, shiny, dark brown, adaxial side grooved and with 2 green narrow wings. Pinnae shortly stalked to almost sessile at apex, 12 – 25 pairs, base nearly opposite, middle and apex alternate, falcate or lanceolate, (middle pairs) 2 – 4 × 0.4 – 0.6 cm, about 2 – 3 (– 5) mm apart, base asymmetrical, acroscopic side truncate and subparallel or forming an angle less than 15 ° with rachis, basiscopic side cuneate, with 10 – 15 mm absent (calculated from the base to the first tooth appearing in the margin of basiscopic side), acroscopic margins serrate, teeth small and dense, usually slightly retuse at the basal portion of pinnae, basiscopic margins entire at base, serrate in the middle and apex; pinna apex acuminate. Veins visible on both sides of pinnae, stramineous, often brown near pinnae petioles, lateral veins visible on both sides of pinnae, green, forking and terminating in marginal teeth or below notches, 2 – 3 basal basiscopic veins lacking. Sori linear, 1 – 2 mm long, medial, 3 – 5 (– 6) on acroscopic side and (1 –) 2 – 4 on distal basiscopic side of the pinnae in the middle part of the laminae, centers 1 – 2 (– 5) mm apart from one another; indusia brown, linear, membranous, entire, opening toward costa.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB5C8027FCFCEB5BA5C6FD3.taxon	distribution	Geographical distribution: — Hymenasplenium speluncicola is currently known from southeastern Guangxi and southern Guizhou, China.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB5C8027FCFCEB5BA5C6FD3.taxon	biology_ecology	Ecology: — Hymenasplenium speluncicola was observed to grow on rocks or soil at the entrance of limestone caves.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB5C8027FCFCEB5BA5C6FD3.taxon	etymology	Etymology: — The species epithet is based on the Latin noun, spelunca, cave, and the Latin suffix, - cola, dweller, referring to the cave-dwelling habitat of the species.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB5C8027FCFCEB5BA5C6FD3.taxon	vernacular_names	Vernacular name: — AE Ḱůďäffǎ (dong xue mo ye tie jiao jue).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB5C8027FCFCEB5BA5C6FD3.taxon	discussion	Taxonomic notes: — Morphologically, Hymenasplenium speluncicola is very special. It has rhizomes densely covered with long yellow brown woolly indumentum, petioles slender, fronds thin herbaceous, pinnae narrow, and pinna apex acuminate. It is likely that H. speluncicola only grows in limestone areas of Guangxi and Guizhou, southern China. Hymenasplenium speluncicola is another species in the genus discovered from limestone caves after H. hastifolium K. W. Xu, Li Bing Zhang & W. B. Liao (2018 b: 281). Similar cave endemics are common in some other fern genera such as Polystichum Roth (1800: 31; Dryopteridaceae; e. g., Han et al. 2016). Additional material examined: — CHINA. Guizhou: Luodian, Limu Xiang, La’ang Cun, Gaopo Zu, Gaopo Big Cave, 25 ° 38 ’ 03.40 ” N, 106 ° 43 ’ 00.95 ” E, elev. 1000 m, 7 November 2011, Li Bing Zhang, Hai He & Yu Wang 6035 (CDBI!, CTC!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB3C8007FCFCD81BF816B17.taxon	materials_examined	Type: — CHINA. Guizhou: Puding, Chengguan Zhen, Lianhua Dong, Dahua Dong, elev. 1250 m, 26 ° 20 ’ 45 ” N, 105 ° 47 ’ 27 ” E, in a cave, 7 August 2017, Li Bing Zhang, Yi-Fan Duan & Matthias Kropf 9610 (holotype CDBI!).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB3C8007FCFCD81BF816B17.taxon	diagnosis	Diagnosis: — Hymenasplenium wangpeishanii is most similar to H. murakami-hatanakae in having pinnae falcate to trapeziform, marginal teeth acute, but the former has sori near costa or submedial, while the latter has sori supramedial to (sub) marginal.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB3C8007FCFCD81BF816B17.taxon	description	Plants perennial, evergreen, up to 45 cm tall. Rhizomes long creeping, ca. 3 mm in diam., apex densely scaly, scales dark brown, narrowly triangular or lanceolate, margins nearly entire, 1 – 1.5 × 0.4 mm; root dark brown when dried, up to 5 cm long, ca. 0.4 mm diam, rhizome and root covered with yellowish brown woolly indument. Fronds remote, up to 12 mm apart, herbaceous; petioles dull brown, base sparsely covered with scales and yellowish brown woolly indument, scales 0.7 × 2 – 3 mm, lanceolate, brown, margins nearly entire. Laminae narrowly ovate, 1 - pinnate, 25 – 30 × 5 – 7 cm, base deflexed, slightly reduced, widest near middle, gradually narrowing toward apex, apex acuminate; rachises 1 mm in diam., subglabrous, brown, adaxial side grooved and with 2 green narrow wings. Pinnae shortly stalked to almost sessile at apex, 20 – 30 pairs, subopposite near base and alternate in the middle and apex, falcate to trapeziform, (middle pairs) 2 – 3.5 × 0.7 cm, about 3 mm apart, base asymmetrical, acroscopic side truncate and subparallel or forming an angle less than 15 ° with rachis, basiscopic side cuneate, with 10 mm absent (calculated from the base to the first tooth appearing in the margin of basiscopic side), acroscopic margins serrate, teeth not retuse, acute, basiscopic margins entire at base, serrate in the middle and apex; pinna apex acute. Veins visible on both sides of pinnae, stramineous, lateral veins visible on both sides of pinnae, green, forking and terminating in marginal teeth, 2 basal basiscopic veins lacking. Sori linear, 1 – 3 mm long, near costa or submedial, (2 –) 6 – 8 (– 10) on acroscopic side and 3 – 6 on distal basiscopic side of the pinnae in the middle part of the laminae, centers 1 – 2 mm apart from one another; indusia brown, linear, membranous, entire, indusium of the second sorus on the acroscopic side of pinnae opening toward rachis, others opening toward costa.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB3C8007FCFCD81BF816B17.taxon	distribution	Geographical distribution: — Hymenasplenium wangpeishanii is mainly only known from Guizhou and Sichuan, Southwest China.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB3C8007FCFCD81BF816B17.taxon	biology_ecology	Ecology: — Hymenasplenium wangpeishanii was observed to grow inside a limestone cave with humid and shady conditions.	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB3C8007FCFCD81BF816B17.taxon	etymology	Etymology: — The species epithet is in honor of Prof. Pei-Shan Wang for his great contributions to the pteridology in China, especially in Guizhou (e. g., Wang & Wang 2001).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB3C8007FCFCD81BF816B17.taxon	vernacular_names	Vernacular name: — áěůďäffǎ (pei shan mo ye tie jiao jue).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
2C2F878FFFB3C8007FCFCD81BF816B17.taxon	discussion	Taxonomic notes: — Hymenasplenium wangpeishanii is also similar to H. apogamum in the sorus distribution and the indusia opening direction, but the former has pinnae falcate to trapeziform and marginal teeth of the pinnae deep (1 mm), while the latter has pinnae quadrangular-trapeziform and marginal teeth of the pinnae shallow (0.3 – 0.7 mm). In addition to the differences from H. murakami-hatanakae and H. apogamum, the new species is also morphologically similar to and phylogenetically closely related to H. speluncicola in the limestone habitat, but the new species has petioles slender (ca. 1 mm in diam.), while H. speluncicola has petioles robust (ca. 2 mm in diam.).	en	Xu, Ke-Wang, Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, He, Hai, Luong, Thien Tam, Knapp, Ralf, Liao, Wen-Bo, Zhang, Li-Bing (2018): Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358 (1): 1-25, DOI: 10.11646/phytotaxa.358.1.1, URL: http://dx.doi.org/10.11646/phytotaxa.358.1.1
