taxonID	type	description	language	source
F4F525B0C7DA59BEB72BAEABB0471088.taxon	type_taxon	Type species. Lachnodiopsis szemaoensis Borchsenius, by original designation.	en	Zarkani, Agustin, Reonaldi, Habib Al Ayubi, Fauzi, Ariffatchur, Apriyanto, Dwinardi, Yahumri,, Rosfiansyah,, Kaydan, Mehmet Bora (2025): Two new species of the mealybug genus Paraputo Laing (Hemiptera, Pseudococcidae) from Borneo Island, Indonesia. ZooKeys 1249: 49-68, DOI: 10.3897/zookeys.1249.159569
FCD7B2A81669511AA8899B7C3D6969E0.taxon	description	Figs 1, 2, 3	en	Zarkani, Agustin, Reonaldi, Habib Al Ayubi, Fauzi, Ariffatchur, Apriyanto, Dwinardi, Yahumri,, Rosfiansyah,, Kaydan, Mehmet Bora (2025): Two new species of the mealybug genus Paraputo Laing (Hemiptera, Pseudococcidae) from Borneo Island, Indonesia. ZooKeys 1249: 49-68, DOI: 10.3897/zookeys.1249.159569
FCD7B2A81669511AA8899B7C3D6969E0.taxon	description	Description of adult female. Appearance in life. Adult females produce a powdery white wax covering the dorsal surface of the body. Living on woody parts of its host plant (Fig. 1), commonly attended by ants of the genus Dolichoderus Lund (Formicidae) (not visible in Fig. 1). Slide-mounted adult female (values of holotype followed by range of 3 paratypes in parentheses) (Figs 2, 3). Body broadly oval, sometimes almost rotund, membranous, largest specimens 2.13 (2.13 – 2.25) mm long and 1.62 (1.62 – 1.73) mm wide. Anal lobes weakly developed, each ventral surface bearing a stout apical seta 80 (80 – 88) μm long arising from a large circular-to-oval sclerotized area, about 57 – 79 μm wide, on ventral margin of abdominal segment VIII; small sclerotized areas present also on ventral margins of segments VII and VI. Antennae each 340 (340 – 360) μm in total length, with 7 antennomeres; antennal setae mostly short. Individual antennal antennomere lengths (in μm) are as follows: antennomere I – VI, each 260 (260 – 270); and VII (apical antennomere), 80 (80 – 90) long and 40 (40 – 45) wide. Apical antennomere with 4 fleshy setae, each 30 – 38 μm long, and an apical seta 36 (36 – 90) μm long. Clypeolabral shield 258 (258 – 263) μm long and 245 (245 – 250) μm wide. Labium unusually long and pointed, longer than clypeolabral shield, 3 segmented, 280 (280 – 300) μm long, with a basal segment 150 (150 – 155) μm wide. Anterior spiracles each 110 (110 – 113) μm long and 58 (58 – 60) μm wide across the atrium, while the posterior spiracles are 110 (110 – 113) μm long and 65 (50 – 65) μm wide across the atrium. Legs well developed (Fig. 3 A); hind leg segments measuring (in μm): coxa, 213 (213 – 215); trochanter + femur, 378 (378 – 400); tibia + tarsus, 238 (238 – 253); and claw, 55 (55 – 70), without a denticle. Ratio of lengths of hind tibia + tarsus to trochanter + femur 0.69 (0.69 – 0.7): 1; ratio of lengths of the tibia to tarsus 1.5 (1.2 – 1.5): 1; and ratio of length of trochanter + femur to the greatest width of femur 3.1 (3.1 – 3.2): 1. Hind legs with only about 10 – 20 translucent pores, by anterior margin of coxa only (Fig. 3 B). Tarsal digitules setose, each 48 – 50 μm long, claw digitules also setose, each about 28 – 30 μm long. Anterior and posterior ostioles well developed, each containing a total of 62 (60 – 70) trilocular pores across both lips and 14 (14 – 25) setae. Circulus situated between abdominal segments III and IV, 140 (140 – 150) μm wide and divided by an intersegmental line. Anal ring about 108 (100 – 108) μm wide, situated on dorsum at about 1.0 – 1.5 times its length from apex of abdomen; with 2 rows of cells and bearing 6 setae, each 78 – 83 μm long. Cerarii numbering not distinct, with many intermediate conical setae, merged to form a marginal band of trilocular pores and enlarged conical setae. Anal lobe cerarii each normally containing 6 or 7 conical setae of different sizes, largest about 25 μm long and 7.5 μm wide at base, and a small compact group of trilocular pores. Anterior cerarii similar to anal lobe cerarii but with largest conical setae slenderer, and each cerarius often subdivided into 2 or 3 smaller cerarii: additional conical setae also present, so that cerarii appear to form a continuous band around margin. Dorsum with slender, stiff setae, each about 20 – 25 µm long, present around margins and across abdominal segments (Fig. 3 C). Setae flanking anal ring longer than other dorsal setae, longest 78 – 88 µm long but mostly each 45 – 50 µm long, each seta shorter than or same length as an anal ring seta (each ring seta about 75 – 88 µm long). Apparent cisanal and obanal setae stout, each 100 – 113 µm long, displaced onto dorsum, situated immediately posterior to anal ring. Multilocular disc pores absent. Trilocular pores abundant, evenly distributed. Discoidal pores fairly numerous, each only slightly smaller than a trilocular pore. Venter with normal flagellate setae present, mostly slender and each longer than a dorsal seta, each 33 – 58 µm long. Ventral sclerotized patches present on anal lobes and on margins of abdominal segments VI and VII (Fig. 3 D). Multilocular disc pores, each 7.5 – 8.8 μm in diameter, few, numbering only 6 – 10 by posterior medial edge of abdominal segment VII only (Fig. 3 E). Trilocular pores numerous but less abundant than on dorsum. Discoidal pores same size as on dorsum, scattered. Vulva becoming heavily sclerotized at maturity, 100 – 113 μm long and 88 – 100 μm wide. Trilocular pores same as those on dorsum, each 2.5 – 3.8 μm wide, scattered throughout. Oral collar tubular ducts of 2 sizes: larger ducts each 10 – 13 μm long and 3.8 μm wide, few, with 2 or 4 situated marginally on abdominal segments VII and VIII; smaller ducts each 7 – 8 μm long and 2.5 μm wide, intermingled with a few large ducts, forming a submarginal-to-marginal cluster of 10 – 14 ducts on each marginal area between segments VII and VIII, a few also present in medial area of abdominal segment VI.	en	Zarkani, Agustin, Reonaldi, Habib Al Ayubi, Fauzi, Ariffatchur, Apriyanto, Dwinardi, Yahumri,, Rosfiansyah,, Kaydan, Mehmet Bora (2025): Two new species of the mealybug genus Paraputo Laing (Hemiptera, Pseudococcidae) from Borneo Island, Indonesia. ZooKeys 1249: 49-68, DOI: 10.3897/zookeys.1249.159569
FCD7B2A81669511AA8899B7C3D6969E0.taxon	etymology	Etymology. The species is named after Prof. Dr. Edhi Martono, an Indonesian entomologist who has dedicated his work to Integrated Pest Management (IPM) and the toxicology of insecticides.	en	Zarkani, Agustin, Reonaldi, Habib Al Ayubi, Fauzi, Ariffatchur, Apriyanto, Dwinardi, Yahumri,, Rosfiansyah,, Kaydan, Mehmet Bora (2025): Two new species of the mealybug genus Paraputo Laing (Hemiptera, Pseudococcidae) from Borneo Island, Indonesia. ZooKeys 1249: 49-68, DOI: 10.3897/zookeys.1249.159569
FCD7B2A81669511AA8899B7C3D6969E0.taxon	distribution	Distribution. Indonesia (Kalimantan, East Kalimantan Province, Sepaku).	en	Zarkani, Agustin, Reonaldi, Habib Al Ayubi, Fauzi, Ariffatchur, Apriyanto, Dwinardi, Yahumri,, Rosfiansyah,, Kaydan, Mehmet Bora (2025): Two new species of the mealybug genus Paraputo Laing (Hemiptera, Pseudococcidae) from Borneo Island, Indonesia. ZooKeys 1249: 49-68, DOI: 10.3897/zookeys.1249.159569
AC2F415AD19F59969EC92D79F71F03DE.taxon	description	Figs 4, 5, 6	en	Zarkani, Agustin, Reonaldi, Habib Al Ayubi, Fauzi, Ariffatchur, Apriyanto, Dwinardi, Yahumri,, Rosfiansyah,, Kaydan, Mehmet Bora (2025): Two new species of the mealybug genus Paraputo Laing (Hemiptera, Pseudococcidae) from Borneo Island, Indonesia. ZooKeys 1249: 49-68, DOI: 10.3897/zookeys.1249.159569
AC2F415AD19F59969EC92D79F71F03DE.taxon	description	Description of adult female. Appearance in life (Fig. 4). The adult female secretes powdery white wax covering over the dorsal surface of the body. Living on woody parts of the host plant, commonly attended by ants of the genus Dolichoderus Lund (Formicidae). Slide-mounted adult female (values of holotype followed by range of 3 paratypes in parentheses) (Figs 5, 6). Body broadly oval, sometimes almost rotund, membranous, largest specimens 2.5 (2.5 – 2.9) mm long and 1.7 (1.7 – 2.2) mm wide. Anal lobes moderately developed, each ventral surface bearing a stout apical seta 84 (80 – 88) μm long arising from a large circular-to-oval sclerotized area, 76 (76 – 80) μm wide, on ventral margin of abdominal segment VIII, occupying much of lobe. Antennae each 390 (390 – 410) μm in total length, with 7 antennomeres; antennal setae mostly short. Individual antennal antennomere lengths (in μm) are as follows: antennomere I – VI, each 310 (310 – 320); and VII (apical antennomere), 90 (90 – 100) long and 30 (30 – 40) wide. Apical antennomere with 4 fleshy setae, each 50 – 55 μm long, and an apical seta 34 (34 – 38) μm long. Clypeolabral shield 345 (345 – 425) μm long and 388 (388 – 393) μm wide. Labium unusually long and pointed, longer than clypeolabral shield, 3 segmented, 210 (210 – 300) μm long, with basal segment 123 (123 – 128) μm wide. Anterior spiracles each 138 (138 – 143) μm long and 70 (70 – 75) μm wide across atrium; posterior spiracles each 138 (138 – 143) μm long and 70 (70 – 88) μm wide across atrium. Legs well developed; hind leg segments measuring (in μm): coxa, 307 (307 – 313); trochanter + femur, 378 (378 – 400); tibia + tarsus, 238 (238 – 253); and claw, 55 (55 – 70), without a denticle. Ratio of lengths of hind tibia + tarsus to trochanter + femur 0.63: 1; ratio of lengths of tibia to tarsus 1.4 (1.2 – 1.4): 1; and ratio of length of trochanter + femur to greatest width of femur 3.3 (3.3 – 3.8): 1. Hind leg with more than 300 translucent pores on coxa only (Fig. 6 A). Tarsal digitules setose, each 53 – 60 μm long; claw digitules each minutely dilated distally, about 28 – 38 μm long. Anterior and posterior ostioles well developed, each containing a total of 62 (60 – 100) trilocular pores across both lips and 7 (5 – 9) setae. Circulus situated between abdominal segments III and IV, 155 (150 – 193) μm wide and divided by an intersegmental line. Anal ring about 100 (100 – 120) μm wide, situated on dorsum about 1.0 – 1.5 times its length from apex of abdomen; with 2 rows of cells and bearing 6 setae, each 75 (75 – 80) μm long. Cerarii numbering not distinct, with many intermediate conical setae, tending to merge. Anal lobe cerarii each normally containing 6 or 7 conical setae of different sizes, largest about 27.5 μm long and 6.3 μm wide at base, and a small compact group of trilocular pores. Anterior cerarii similar to anal lobe cerarii but with largest conical setae slenderer, and each cerarius often subdivided into 2 or 3 smaller cerarii: additional conical setae also present, so that cerarii appear to be continuous on some parts of margin. Dorsum. Slender, stiff setae present, mostly each about 12.5 – 20.0 μm long; long setae flanking anal ring fairly numerous, each 77.5 – 87.5 µm long, similar length to or shorter than an anal ring seta (each ring seta 75 – 100 µm long). Two pairs of long stout setae present just to posterior to anal ring, each about 105 – 113 µm long, longer and wider than an anal ring seta (Fig. 6 B). Trilocular pores abundant. Discoidal pores fairly numerous, each slightly smaller than a trilocular pore. Venter. Normal flagellate setae present, mostly slender and each longer than a dorsal seta, 10 – 35 µm long. Ventral sclerotized patches present on anal lobes only. Cisanal and obanal setae slender, each 38 – 40 µm long, displaced to dorsal surface between anal ring and apex of abdomen. Multilocular disc pores each 10.0 – 12.5 μm in diameter, present on abdominal segments, distributed as follows (numbers): segment V: 3 – 5, VI: 31 – 38, VII: 30 – 32, and VIII: 8 – 10. Trilocular pores numerous but less abundant than on dorsum. Discoidal pores same size as on dorsum, scattered. Vulva becoming heavily sclerotized at maturity, 163 – 175 μm long and 88 – 100 μm wide; with a pair of noticeable sclerotized internal pockets, base of each 70 – 75 μm long and 30 – 38 μm wide (Fig. 6 D). Oral collar tubular ducts of 3 sizes: (i) largest ducts each 10.0 – 12.5 μm long and 7.5 μm wide, distributed on abdominal segments (numbers): VII: 6 – 8, and VIII: 2 – 4; (ii) slightly smaller ducts, each 7.5 – 10.0 μm long and about 5 μm wide, varying in number, present posterior to vulva and across medial posterior edges of abdominal segments V – VIII; and (iii) minute ducts, each 7.5 – 8.0 μm long and about 2.5 μm wide, few, with 5 or 6 situated medially on abdominal segments V and VI.	en	Zarkani, Agustin, Reonaldi, Habib Al Ayubi, Fauzi, Ariffatchur, Apriyanto, Dwinardi, Yahumri,, Rosfiansyah,, Kaydan, Mehmet Bora (2025): Two new species of the mealybug genus Paraputo Laing (Hemiptera, Pseudococcidae) from Borneo Island, Indonesia. ZooKeys 1249: 49-68, DOI: 10.3897/zookeys.1249.159569
AC2F415AD19F59969EC92D79F71F03DE.taxon	etymology	Etymology. The species is named after Prof. Dr. Aunu Rauf, an Indonesian entomologist who is interested in the conservation of native insects of Indonesia.	en	Zarkani, Agustin, Reonaldi, Habib Al Ayubi, Fauzi, Ariffatchur, Apriyanto, Dwinardi, Yahumri,, Rosfiansyah,, Kaydan, Mehmet Bora (2025): Two new species of the mealybug genus Paraputo Laing (Hemiptera, Pseudococcidae) from Borneo Island, Indonesia. ZooKeys 1249: 49-68, DOI: 10.3897/zookeys.1249.159569
AC2F415AD19F59969EC92D79F71F03DE.taxon	distribution	Distribution. Indonesia (Kalimantan, East Kalimantan Province, Sepaku).	en	Zarkani, Agustin, Reonaldi, Habib Al Ayubi, Fauzi, Ariffatchur, Apriyanto, Dwinardi, Yahumri,, Rosfiansyah,, Kaydan, Mehmet Bora (2025): Two new species of the mealybug genus Paraputo Laing (Hemiptera, Pseudococcidae) from Borneo Island, Indonesia. ZooKeys 1249: 49-68, DOI: 10.3897/zookeys.1249.159569
