identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9C909A0914A255C9BFB990B8B8A91CCD.text	9C909A0914A255C9BFB990B8B8A91CCD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deltochilum abdominale Martinez 1947	<div><p>Deltochilum abdominale Martínez, 1947</p><p>Figs 3 B, 3 G, 4 A, 5, 6, 7 A (blue circle)</p><p>Deltochilum (D.) abdominalis Martínez, 1947: 269–274 . Lámina VIII (description).</p><p>Deltochilum (Deltohyboma) abdominalis Martínez 1954: 47 (cited from Colombia, error; see D. ventripuncturatussp. nov.).</p><p>Deltochilum (Deltohyboma) abdominale Vulcano &amp; Pereira, 1964: 652 (catalogue).</p><p>Deltochilum (Deltohyboma) abdominalis Blanco 1987: 43 (distribution, Venezuela).</p><p>Deltochilum abdominale Medina et al. 2001: 136 (cited from Guaviare (gv?) Colombia with doubts).</p><p>Deltochilum (Deltohyboma) abdominalis González-Alvarado &amp; Vaz-de-Mello, 2021: 62, 74 (species group).</p><p>Deltochilum abdominale Lozano De La Rosa et al. 2024: 72 (declination of the species epithet in neuter, distribution, Venezuela).</p><p>Type material examined.</p><p>Holotype ♂ Venezuela, D. [istrito] F. [ederal], 720 mts alt., Mo. del Naiguatá, AGO. 941. R. Lichy. Legit. Coll Martínez. [white label] HOLOTIPO. [red label] Deltochilum abdominalis [male symbol] sp. nov. A. Martínez-DET. 946. [white label] FICHADO. [white label] MACN-En 790. (MACN) .</p><p>Additional material examined</p><p>(81 specimens): Colombia: Norte de Santander: Toledo, IP Santa María, Alto de Herrera, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.21667&amp;materialsCitation.latitude=7.116667" title="Search Plazi for locations around (long -72.21667/lat 7.116667)">Vda. Diamante, Fca. La Primavera. PNN Tamá</a>, 7°7'N, 72°13'W, 1000 m, 1 ♀, 2 ♂, 1999. ix. 22, González E., T. Exc. Hum (IAvH) , Toledo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.311005&amp;materialsCitation.latitude=7.1857777" title="Search Plazi for locations around (long -72.311005/lat 7.1857777)">Vda. San Alberto</a>. Bosque, 07°11'08.8"N, 072°18'39.6"W, 823 m, 2 ♂, 11. iv. 2019, Ramírez, M. &amp; Lopera, A., T. Exc. H # SAF 1 (24 h) (1 ♂ IAvH, 1 ♂, CALT-ECC) , Toledo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.30786&amp;materialsCitation.latitude=7.1875" title="Search Plazi for locations around (long -72.30786/lat 7.1875)">Vda. San Alberto</a>. Bosque, 07°11'15.0"N, 072°18'28.3"W, 1104 m, 1 ♀, 11. iv. 2019, Ramírez, M. &amp; Lopera, A., T. Exc. H # SAF 3 (24 h) (IAvH) . Venezuela: Chichiriviche (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.27782&amp;materialsCitation.latitude=10.933753" title="Search Plazi for locations around (long -68.27782/lat 10.933753)">Litoral, D. F</a>), [10°56'1.51"N], [68°16'40.14"W], 1 ♀, 27. i. 1977, Bordón coll. (BDGC) ; Aragua: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.69556&amp;materialsCitation.latitude=10.445" title="Search Plazi for locations around (long -67.69556/lat 10.445)">Cuyagua</a>, [10°26'42"N], [67°41'44"W], 800 m, 2 ♀, 1 ♂, x. 1972, Martínez coll. (CMNC) , Quebrada Coirral da Piedra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.65&amp;materialsCitation.latitude=10.316667" title="Search Plazi for locations around (long -67.65/lat 10.316667)">El Limón</a>, [10°19'N], [67°39'W], 700 m, 1 ♂, 7. vi. 1973, Bordón coll. (CMNC) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.542786&amp;materialsCitation.latitude=10.067056" title="Search Plazi for locations around (long -67.542786/lat 10.067056)">Racho Grande</a>, [10°4'1.40"N], [67°32'34.02"W], 1200 m, 1 ♂, 27–30. viii. 1983, B Gill coll. (BDGC) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.542786&amp;materialsCitation.latitude=10.067056" title="Search Plazi for locations around (long -67.542786/lat 10.067056)">Racho Grande</a>, [10°4'1.40"N], [67°32'34.02"W], 1150 m, 1 ♂, 4–9. vii. 1986, B Gill coll. (BDGC) ; Miranda: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.49472&amp;materialsCitation.latitude=10.093611" title="Search Plazi for locations around (long -66.49472/lat 10.093611)">Parque Nacional Guatopo</a>, [10°5'37"N], [66°29'41"W], 6 ♀, 6 ♂, 3–8. iii. 1988, F. Génier coll., pitfall trap, human feces (CMNC) , Parque Nacional Guatopo, 50 km SE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.48333&amp;materialsCitation.latitude=10.083333" title="Search Plazi for locations around (long -66.48333/lat 10.083333)">Caracas</a>. forest, [10°5'N], [66°29'W], 400 m, 2 ♀, 1 ♂, 5–6. iii. 1971, S. Peck coll., pitfall trap, human dung (CMNC) , Santa Crucita, 33 km N Altagracia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.01667&amp;materialsCitation.latitude=10.233334" title="Search Plazi for locations around (long -67.01667/lat 10.233334)">Parque Nacional Guatopo</a>. forest, [10°14'N], [67°1'W], 400 m, 1 ♂, 7–10. vi. 1987, S. &amp; J. Peck coll., dung trap (CMNC) ; Monagas: Cueva Guácharo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-63.541668&amp;materialsCitation.latitude=10.183333" title="Search Plazi for locations around (long -63.541668/lat 10.183333)">Caripe</a>. forest over coffee, [10°11'N], [63°32'30"W], 750 m, 1 ♀, 1 ♂, 20–31. vii. 1987, S. &amp; J. Peck coll. (CMNC) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-63.483334&amp;materialsCitation.latitude=10.183333" title="Search Plazi for locations around (long -63.483334/lat 10.183333)">Teresén</a>, [10°11'N], [63°29'W], 800 m, 3 ♂, 12. iv. 1963, R. Hernandez coll. (CMNC) ; Táchira: 20 km NE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.08417&amp;materialsCitation.latitude=7.959444" title="Search Plazi for locations around (long -72.08417/lat 7.959444)">San Cristobal</a>, [7°57'34"N], [72°5'3"W], 3000 m, 1 ♂, 20. v. 1974, S. Peck coll. (CMNC) , 20 km NE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.08417&amp;materialsCitation.latitude=7.959444" title="Search Plazi for locations around (long -72.08417/lat 7.959444)">San Cristobal</a>, [7°57'34"N], [72°5'3"W], 1200 m, 1 ♂, 20–22. v. 1974, S. Peck coll. (CMNC) , La Trampita, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.723335&amp;materialsCitation.latitude=7.9336114" title="Search Plazi for locations around (long -71.723335/lat 7.9336114)">Presa La Honda</a>, [7°56'1"N], [71°43'24"W], 1100 m, 1 ♀, 17. xii. 1988, D. Havranek coll., mini-dung cup traps (CMNC) , Las Trampitas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.76056&amp;materialsCitation.latitude=8.011111" title="Search Plazi for locations around (long -71.76056/lat 8.011111)">Pregonero</a>. sub-mountain forest, [8°0'40"N], [71°45'38"W], 1240 m, 10 ♀, 11 ♂, 9–14. vii. 1989, S. &amp; J. Peck coll., carrion traps (CMNC) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.76363&amp;materialsCitation.latitude=7.996914" title="Search Plazi for locations around (long -71.76363/lat 7.996914)">Pregonero Las Trampitas</a>, [7°59'48.89"N], [71°45'49.08"W], 1240 m, 2 ♀, 4 ♂, 9–14. vii. 1989, J Peck &amp; S Peck coll. (BDGC) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.1093&amp;materialsCitation.latitude=7.8436418" title="Search Plazi for locations around (long -72.1093/lat 7.8436418)">Rio Frio</a>, [7°50'37.11"N]?, [72°6'33.48"W]?, 500 m, 3 ♀, 3 ♂, 11–18. viii. 1983, B Gill coll. (BDGC) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.20573&amp;materialsCitation.latitude=7.7586083" title="Search Plazi for locations around (long -72.20573/lat 7.7586083)">San Cristobal</a>, [7°45'30.99"N], [72°12'20.62"W], 1200 m, 3 ♀, 6 ♂, 10–18. viii. 1983, B Gill coll. (BDGC) ; Yaracuy: Bolívar. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-68.83492&amp;materialsCitation.latitude=10.33965" title="Search Plazi for locations around (long -68.83492/lat 10.33965)">Aroa</a>, 10°20'22.74"N, 68°50'5.71"W, 1372 m, 1 ♀, 1 ♂, 19. VII. 2009, Asmüssen M, Colmenares P., Martínez H., Cebo Heces humanas (CEMT) .</p><p>Diagnosis.</p><p>This species can be distinguished from all other species within the plebejum species group by having the smallest interstrial punctures (Fig. 5 C). These punctures are nearly the same size as the interstrial shiny points; in all other species of the group, these shiny points are distinctly smaller than the interstrial punctures. Furthermore, the males possess a denticle on the mesofemur (Fig. 6 A, G), a character shared with D. tyba sp. nov. (Fig. 9 G), but the two species can be separated by the size of the interstrial punctures previously described.</p><p>Redescription.</p><p>Minor male. Size 10 mm. Color. Dark brown with some blue reflections dorsally. Dark blue ventrally. Head (Fig. 5 A). Interocular distance 8 × the width of one eye. Punctures on frons separated by one or less than one diameter of each puncture and subequal in size to head discal punctures. Disc punctures separated by one or less than one diameter of each puncture. Punctures from disc towards anterior area successively of almost with the same size. Genal punctures subequal in size to the discal punctures and separated by one diameter of each puncture. Pronotum (Fig. 5 B). Margin between anterior and medial-lateral angle almost straight. Medial angle of pronotum rounded. Margin between medial-lateral angle and posterior-lateral angle almost straight. Disc punctures half size to anterior-lateral ones and separated by more than one diameter of each puncture. Basal punctures are smaller than anterior-lateral ones and separated by approximately one diameter of each puncture. Hypomera. With ocellated and setigerous punctures on the anterior part (Fig. 3 B, white arrow). Most of punctures incompletely closed posteriorly and without setae (Fig. 3 B, green arrow). Anterior punctures denser and larger than posterior ones. Elytra (Fig. 5 C). Carina of the ninth interstria almost reaching middle of elytral length. Elytral apex on interstriae III, V – VII with tubercles. Striae I – VII almost inconspicuous. Width of third stria on disc ~ 1 / 55 of the distance between striae II and III. Strial punctures ~ 2 × wider than the stria. First stria as wider as second stria. Stria VIII conspicuous apically, laterally almost inconspicuous. Punctures on interstriae approximately separated by two diameters of each puncture and the same size or slightly smaller than strial punctures. Punctures of the third interstria on disc occupying ~ 1 / 18 of the distance between striae II and III. Interstriae with shiny points mixed with the punctures. Points almost the same size as the interstrial punctures. Metaventrite (Fig. 5 D). Disc with few deep excavation, occupying metaventral basal third. Anterolateral areas of metaventral process with punctures completely closed and separated by less than one diameter of each puncture. Anterocentral area of metaventral process with few and small punctures, subequal in size to the discal punctures. Punctures on the lateral lobules not completely closed (some of them) and separated by one or less than one diameter of each puncture. Abdomen (Fig. 5 E). Ventrite I expanded posteriorly, expansion reaching the fifth ventrite; on ventrite III, the expansion is as wide as the distance between mesocoxae. Basal part of the ventrite I, between the metacoxae not modified. Legs (Fig. 6 A, B). Protibia strongly broadens toward the apex. Profemora with punctures separated by less than one diameter of each puncture. Mesofemur with punctures separated by 2 × the diameter of each puncture. Mesofemur with a basal denticle on the posterior edge. Metafemur regularly broadens toward the apex. Metafemur with punctures separated by 2 × the diameter of each puncture. Metatibial spur insertion not elongated. Metatibial spur clearly distinct from the metatibia. Internal margin of metatibia with tubercles on the basal half of the metatibial length. Pygidium (Fig. 6 C). Pygidial punctures almost uniform in size and density. Punctures separated by one or less than the diameter of each puncture. Discal punctures, on the center of the pygidium, small, each puncture occupying ~ 1 / 45 of the distance between the lateral pygidial margins. Genitalia (Fig. 6 D – F). Paramera smaller than the phallobase. Apex of paramera with a large denticle in lateral view. Medial area of endophallus with two endophallites, the right ~ 4 × longer than the other.</p><p>Variation.</p><p>The female exhibits the differences described here under the sexual dimorphism section of the species group redescription. Medium-sized males have both the basal denticle of the metafemur and the metatibial tubercles (Fig. 6 J, white arrow) that are larger than those found in the minor male. The metaspur is articulated. Major males: the metatibial basal carina is irregular and strongly interrupted by setae, giving it the appearance of tubercles (Fig. 6, white arrow). This carina is significantly reduced in the mid-length of the metatibial; distal to this reduction, the carina rises again (giving the appearance of having both tubercles and carina). Other major males possess this tuberculate basal part, but the reduction is less pronounced, and the tubercles are continuous (Fig. 6 I, white arrow). The basal denticle of the metafemur is larger and more strongly curved (Fig. 6 G). The metatibia is also more strongly curved (Fig. 6 H, I). Metaspur fused.</p><p>Known distribution</p><p>(Fig. 7 A blue circle). Colombia, Norte de Santander. Venezuela, Aragua, Miranda, Monagas, Táchira, Yaracuy.</p><p>Remarks.</p><p>This species has been collected in forest, submontane forest and few specimens in forest mixed with coffee. This species is one of the members of the plebejum species group that is widely distributed, possessing a broad altitudinal range from 400 to probably 1,400 m above sea level.</p></div>	https://treatment.plazi.org/id/9C909A0914A255C9BFB990B8B8A91CCD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	González-Alvarado, Arturo;Neita, Jhon Cesar	González-Alvarado, Arturo, Neita, Jhon Cesar (2026): Eight new species of Deltochilum Eschscholtz, 1822 (subgenus D. Deltohyboma Lane, 1946) from the Eastern Cordillera of Colombia and Venezuelan Andes (Coleoptera, Scarabaeinae, plebejum species group). ZooKeys 1280: 171-229, DOI: 10.3897/zookeys.1280.182114
34656E6E64CD5FE0BA2FA05B7F679BCA.text	34656E6E64CD5FE0BA2FA05B7F679BCA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deltochilum altiventris González-Alvarado & Neita 2026	<div><p>Deltochilum altiventris sp. nov.</p><p>Figs 3 A, 3 C, 3 H, 4 H, 7 A (yellow triangles), 18, 19</p><p>Type material.</p><p>Holotype: ♂, Colombia: Cundinamarca: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.4345&amp;materialsCitation.latitude=4.516361" title="Search Plazi for locations around (long -73.4345/lat 4.516361)">Medina, Vda. Periquito, Fca. Miramar, Campamento 2. Bosque subandino, Bosque entresacado</a>, 04°30'58.9"N, 73°26'04.2"W, 1212 m, 2018. iii. 15–17, Castro M. I., Neita J. C. &amp; Torres E., T. Exc. H. T 2 T 9 (48 h) (IAvH) [IAvH -E-203206] . Paratypes (108 specimens): Colombia: Arauca: Tame. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.86583&amp;materialsCitation.latitude=6.523611" title="Search Plazi for locations around (long -71.86583/lat 6.523611)">Brisas el Cravo</a>. Bosque, 06°31'25"N, 71°51'57"W, 1500 m, 1 ♂, 15–19 / II / 08, F. Alvarado, E. H. (CALT-ECC) ; Boyacá: Cusiana <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.68806&amp;materialsCitation.latitude=5.3941665" title="Search Plazi for locations around (long -72.68806/lat 5.3941665)">Cerca a Pajarito</a>, 5°23'39"N, 72°41'17"W, 1250 m, 1 ♀, vi. 1997, F. Escobar coll. (CMNC) , Cusiana <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.68806&amp;materialsCitation.latitude=5.3941665" title="Search Plazi for locations around (long -72.68806/lat 5.3941665)">Cerca a Pajarito</a>, 5°23'39"N, 72°41'17"W, 1350 m, 1 ♂, vi. 1997, F. Escobar coll. (CMNC) , Cusiana, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.68806&amp;materialsCitation.latitude=5.3941665" title="Search Plazi for locations around (long -72.68806/lat 5.3941665)">Cerca a Pajarito</a>. Bosque, 5°23'39"N, 72°41'17"W, 1250 m, 2 ♀, 3 ♂, 1997. vi, Escobar F., T. Exc. H. (IAvH) , Cusiana, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.666664&amp;materialsCitation.latitude=5.2833333" title="Search Plazi for locations around (long -72.666664/lat 5.2833333)">Cerca de Pajarito, Fca. Guayabetal</a>, el paisano. Bosque, 5°17'N, 72°40'W, 1300 m, 1 ♂, 1997. vi. 1, Escobar F., T. Exc. H. 74 (IAvH) , Cusiana, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.683334&amp;materialsCitation.latitude=5.4333334" title="Search Plazi for locations around (long -72.683334/lat 5.4333334)">Cerca de Pajarito, Fca. Guayabetal, el paisano. Potrero</a>, 5°26'N, 72°41'W, 1000 m, 1 ♂, 1997. vi. 1, Escobar F., T. Exc. H. 73 (IAvH) , Pajarito. Corinto. Vereda La Sabana, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.72478&amp;materialsCitation.latitude=5.4006057" title="Search Plazi for locations around (long -72.72478/lat 5.4006057)">Corregimiento Corinto, Finca El Mayor. Borde de bosque</a>, 5°24'02.18"N, 72°43'29.20"W, 1542 m, 4 Unsexed, 2 / 04 / 2021, Andrés Felipe Morales Alba | David Camilo Martínez Dueñas, Trampa pitfall 4. 24 H (UPTC-CE) , Paya. Vda. Bocas de Monte. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.46547&amp;materialsCitation.latitude=5.5893335" title="Search Plazi for locations around (long -72.46547/lat 5.5893335)">Finca La Piedrota</a>. Bosque, 5°35'21.6"N, 72°27'55.7"W, 1577 m, 1 Unsexed, 9 / 08 / 2021, Andrés Felipe Morales Alba | David Camilo Martínez Dueñas, Trampa pitfall 1. 24 H (UPTC-CE) , Santa María, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.255905&amp;materialsCitation.latitude=4.869803" title="Search Plazi for locations around (long -73.255905/lat 4.869803)">La Almenará</a>, [4°52'11.29"N], [73°15'21.26"W], 800–1000 m, 1 ♀, Ago- 2008, G. Amat &amp; R. Sarmiento (ICN) , Santa María, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.255905&amp;materialsCitation.latitude=4.869803" title="Search Plazi for locations around (long -73.255905/lat 4.869803)">La Almenará</a>, [4°52'11.29"N], [73°15'21.26"W], 880 m, 1 ♂, Octubre 2008, G. Amat &amp; Sarmiento R. (ICN) , Santa María, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.2754&amp;materialsCitation.latitude=4.849286" title="Search Plazi for locations around (long -73.2754/lat 4.849286)">Vda. Caño Negro</a>. Bosque, [4°50'57.43"N], [73°16'31.43"W], 1 ♀, 2 ♂, 17–19 / 11 / 03, Pitfall. M. 26 54 (ICN) , Santa María. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.25347&amp;materialsCitation.latitude=4.8800836" title="Search Plazi for locations around (long -73.25347/lat 4.8800836)">RN La Almenara. Vda. Sta Cecilia</a>. Bosque, 04°52'48.3"N, 073°15'12.5"W, 1191 m, 1 ♀, 1 ♂, 16. vii. 2018, Martínez, D., T. Ecx. H. # ALF 2 (72 h) (IAvH) , Santa María. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.28819&amp;materialsCitation.latitude=4.8395834" title="Search Plazi for locations around (long -73.28819/lat 4.8395834)">Vda. Caño Negro. Fca. Santa María</a>. Bosque, 04°50'22.5"N, 073°17'17.5"W, 1470 m, 2 ♀, 2 ♂, 11. vii. 2018, Martínez, D., T. Ecx. H. # SMF 1 R 3 (72 h) (IAvH) , Sta. Maria, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.28333&amp;materialsCitation.latitude=4.883333" title="Search Plazi for locations around (long -73.28333/lat 4.883333)">Sendero Ecológico</a>, [4°53'N], [73°17'W], 1 ♀, 2 ♂, 17–04 – 97, G. Amat (ICN) ; Comijoque. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.683334&amp;materialsCitation.latitude=5.4166665" title="Search Plazi for locations around (long -72.683334/lat 5.4166665)">Transecto Cusiana</a>. Bosque, 5°25'N, 72°41'W, 2000 m, 1 ♀, 1997. iv. 6, Escobar F. (UPTC-CE) , Cusiana cerca de Pajarito. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.683334&amp;materialsCitation.latitude=5.383333" title="Search Plazi for locations around (long -72.683334/lat 5.383333)">Fca Guayabetal. El Paisano</a>. Bosque, 5°23'N, 72°41'W, 1250 m, 1 ♀, 1 ♂, 1997. iii. 6, Escobar F. (UPTC-CE) , Cusiana cerca de Pajarito. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.683334&amp;materialsCitation.latitude=5.4333334" title="Search Plazi for locations around (long -72.683334/lat 5.4333334)">Fca Guayabetal. El Paisano</a>. Bosque, 5°26'N, 72°41'W, 1000 m, 1 ♀, 1997. v. 6, Escobar F. (UPTC-CE) , 1 ♂, 1997. vi. 6, Escobar F. (UPTC-CE); Casanare: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.683334&amp;materialsCitation.latitude=5.383333" title="Search Plazi for locations around (long -72.683334/lat 5.383333)">Cusiana</a>. Bosque, 5°23'N, 72°41'W, 1200 m, 2 ♂, 1997. vi, Escobar F., T. Exc. H. (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.683334&amp;materialsCitation.latitude=5.383333" title="Search Plazi for locations around (long -72.683334/lat 5.383333)">Cusiana</a>. Bosque, 5°23'N, 72°41'W, 1000 m, 1 ♂, 1997. vi, Escobar F., T. Exc. H. (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.683334&amp;materialsCitation.latitude=5.383333" title="Search Plazi for locations around (long -72.683334/lat 5.383333)">Cusiana</a>. Bosque, 5°23'N, 72°41'W, 1200 m, 1 ♂, 1997. vi, Escobar F., T. Exc. H. (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.683334&amp;materialsCitation.latitude=5.383333" title="Search Plazi for locations around (long -72.683334/lat 5.383333)">Cusiana</a>. Bosque, 5°23'N, 72°41'W, 1000 m, 1 ♂, 1997. vi, Escobar F., T. Exc. H. (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.683334&amp;materialsCitation.latitude=5.383333" title="Search Plazi for locations around (long -72.683334/lat 5.383333)">Cusiana</a>. Bosque, 5°23'N, 72°41'W, 1200 m, 2 ♂, 1997. vi, Escobar F., T. Exc. H. (IAvH) , Tauramena. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.85551&amp;materialsCitation.latitude=5.069943" title="Search Plazi for locations around (long -72.85551/lat 5.069943)">Monserrate Alto</a>. Bosque, 5.069943, -72.855504, 1130 m, 1 ♀, IV / 2018, P. Triviño, PF exc hum (CALT-ECC) , Tauramena. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.84851&amp;materialsCitation.latitude=5.064737" title="Search Plazi for locations around (long -72.84851/lat 5.064737)">Monserrate Alto</a>. Bosque, 5.064737, -72.84851, 1311 m, 1 ♀, IV / 2018, P. Triviño, PF exc hum (CALT-ECC) , Tauramena. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.85516&amp;materialsCitation.latitude=5.069697" title="Search Plazi for locations around (long -72.85516/lat 5.069697)">Monserrate Alto</a>. Bosque, 5.069697, -72.855163, 1120 m, 3 ♂, IV / 2018, P. Triviño, PF exc hum (CALT-ECC) , Tauramena. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.85551&amp;materialsCitation.latitude=5.069943" title="Search Plazi for locations around (long -72.85551/lat 5.069943)">Monserrate Alto</a>. Bosque, 5.069943, -72.855504, 1130 m, 4 ♂, IV / 2018, P. Triviño, PF exc hum (CALT-ECC) , Tauramena. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.85743&amp;materialsCitation.latitude=5.07047" title="Search Plazi for locations around (long -72.85743/lat 5.07047)">Monserrate Alto</a>. Bosque, 5.07047, -72.85743, 1169 m, 1 ♂, IV / 2018, P. Triviño, PF exc hum (CALT-ECC) , Tauramena. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.848526&amp;materialsCitation.latitude=5.064961" title="Search Plazi for locations around (long -72.848526/lat 5.064961)">Monserrate Alto</a>. Bosque, 5.064961, -72.848524, 1271 m, 1 ♂, IV / 2018, P. Triviño, PF exc hum (CALT-ECC) , Tauramena. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.84847&amp;materialsCitation.latitude=5.064435" title="Search Plazi for locations around (long -72.84847/lat 5.064435)">Monserrate Alto</a>. Bosque, 5.064435, -72.848473, 1290 m, 2 ♂, IV / 2018, P. Triviño, PF exc hum (CALT-ECC) , Tauramena. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.84743&amp;materialsCitation.latitude=5.063548" title="Search Plazi for locations around (long -72.84743/lat 5.063548)">Monserrate Alto</a>. Bosque, 5.063548, -72.847425, 1339 m, 3 ♂, IV / 2018, P. Triviño, PF exc hum (CALT-ECC) , Tauramena. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.84743&amp;materialsCitation.latitude=5.063548" title="Search Plazi for locations around (long -72.84743/lat 5.063548)">Vda. El Palmar</a>. Bosque, 5.063548, -72.847425, 1339 m, 1 ♂, IV / 2018, P. Triviño, PF exc hum (CALT-ECC) ; Cundinamarca: Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.388054&amp;materialsCitation.latitude=4.5925" title="Search Plazi for locations around (long -73.388054/lat 4.5925)">Miralindo</a>, 4°35'33"N, 73°23'17"W, 1500 m, 1 ♀, 1997. ii-iii, Escobar F., T. Exc. H. (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.388054&amp;materialsCitation.latitude=4.5925" title="Search Plazi for locations around (long -73.388054/lat 4.5925)">Miralindo</a>, 4°35'33"N, 73°23'17"W, 1250 m, 1 ♀, 1997. ii-iii, Escobar F., T. Exc. H. (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.388054&amp;materialsCitation.latitude=4.5925" title="Search Plazi for locations around (long -73.388054/lat 4.5925)">Miralindo</a>, 4°35'33"N, 73°23'17"W, 1000 m, 2 ♀, 1997. ii-iii, Escobar F., T. Exc. H. (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.388054&amp;materialsCitation.latitude=4.5925" title="Search Plazi for locations around (long -73.388054/lat 4.5925)">Miralindo</a>, 4°35'33"N, 73°23'17"W, 1500 m, 1 ♀, 1997. ii-iii, Escobar F., T. Exc. H. (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.388054&amp;materialsCitation.latitude=4.5925" title="Search Plazi for locations around (long -73.388054/lat 4.5925)">Miralindo</a>, 4°35'33"N, 73°23'17"W, 1000 m, 2 ♂, 1997. ii-iii, Escobar F., T. Exc. H. (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.388054&amp;materialsCitation.latitude=4.5925" title="Search Plazi for locations around (long -73.388054/lat 4.5925)">Miralindo</a>, 4°35'33"N, 73°23'17"W, 1500 m, 1 ♂, 1997. ii-iii, Escobar F., T. Exc. H. (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.388054&amp;materialsCitation.latitude=4.5925" title="Search Plazi for locations around (long -73.388054/lat 4.5925)">Miralindo</a>, 4°35'33"N, 73°23'17"W, 1000 m, 1 ♂, 1997. ii-iii, Escobar F., T. Exc. H. (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.388054&amp;materialsCitation.latitude=4.5925" title="Search Plazi for locations around (long -73.388054/lat 4.5925)">Miralindo</a>, 4°35'33"N, 73°23'17"W, 1250 m, 1 ♂, 1997. ii-iii, Escobar F., T. Exc. H. (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.388054&amp;materialsCitation.latitude=4.5925" title="Search Plazi for locations around (long -73.388054/lat 4.5925)">Miralindo</a>, 4°35'33"N, 73°23'17"W, 1000 m, 4 ♂, 1997. ii-iii, Escobar F., T. Exc. H. (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.43592&amp;materialsCitation.latitude=4.516111" title="Search Plazi for locations around (long -73.43592/lat 4.516111)">Medina, Vda. Periquito, Fca. Miramar, Campamento 2. Bosque subandino, Bosque entresacado</a>, 04°30'58.0"N, 73°26'09.3"W, 1277 m, 1 ♀, 2 ♂, 2018. iii. 15–16, Castro M. I., Neita J. C. &amp; Torres E., T. Exc. H. T 2 T 6 (24 h) (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.434975&amp;materialsCitation.latitude=4.516222" title="Search Plazi for locations around (long -73.434975/lat 4.516222)">Medina, Vda. Periquito, Fca. Miramar, Campamento 2. Bosque subandino, Bosque entresacado</a>, 04°30'58.4"N, 73°26'05.9"W, 1229 m, 1 ♀, 2018. iii. 15–16, Castro M. I., Neita J. C. &amp; Torres E., T. Exc. H. T 2 T 8 (24 h) (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.436386&amp;materialsCitation.latitude=4.5157776" title="Search Plazi for locations around (long -73.436386/lat 4.5157776)">Medina, Vda. Periquito, Fca. Miramar, Campamento 2. Bosque subandino, Bosque entresacado</a>, 04°30'56.8"N, 73°26'11.0"W, 1289 m, 1 ♂, 2018. iii. 15–17, Castro M. I., Neita J. C. &amp; Torres E., T. Exc. H. T 2 T 5 (48 h) (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.4355&amp;materialsCitation.latitude=4.5164165" title="Search Plazi for locations around (long -73.4355/lat 4.5164165)">Medina, Vda. Periquito, Fca. Miramar, Campamento 2. Bosque subandino, Bosque entresacado</a>, 04°30'59.1"N, 73°26'07.8"W, 1246 m, 1 ♂, 2018. iii. 15–17, Castro M. I., Neita J. C. &amp; Torres E., T. Exc. H. T 2 T 7 (48 h) (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.442276&amp;materialsCitation.latitude=4.5145555" title="Search Plazi for locations around (long -73.442276/lat 4.5145555)">Vda. Periquito, Fca. Miramar, Campamento 2. Bosque subandino, Bosque primario</a>, 04°30'52.4"N, 73°26'32.2"W, 1510 m, 1 ♀, 2018. iii. 14–15, Castro M. I., Neita J. C. &amp; Torres E., T. Exc. H. T 1 T 2 (24 h) (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.44164&amp;materialsCitation.latitude=4.515222" title="Search Plazi for locations around (long -73.44164/lat 4.515222)">Vda. Periquito, Fca. Miramar, Campamento 2. Bosque subandino, Bosque primario</a>, 04°30'54.8"N, 73°26'29.9"W, 1507 m, 1 ♂, 2018. iii. 14–15, Castro M. I., Neita J. C. &amp; Torres E., T. Exc. H. T 1 T 4 (24 h) (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.44239&amp;materialsCitation.latitude=4.5140834" title="Search Plazi for locations around (long -73.44239/lat 4.5140834)">Vda. Periquito, Fca. Miramar, Campamento 2. Bosque subandino, Bosque primario</a>, 04°30'50.7"N, 73°26'32.6"W, 1515 m, 1 ♀, 2 ♂, 2018. iii. 14–16, Castro M. I., Neita J. C. &amp; Torres E., T. Exc. H. T 1 T 1 (48 h) (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.44192&amp;materialsCitation.latitude=4.5148335" title="Search Plazi for locations around (long -73.44192/lat 4.5148335)">Vda. Periquito, Fca. Miramar, Campamento 2. Bosque subandino, Bosque primario</a>, 04°30'53.4"N, 73°26'30.9"W, 1509 m, 1 ♂, 2018. iii. 14–16, Castro M. I., Neita J. C. &amp; Torres E., T. Exc. H. T 1 T 3 (48 h) (IAvH) , Medina, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.44125&amp;materialsCitation.latitude=4.5155277" title="Search Plazi for locations around (long -73.44125/lat 4.5155277)">Vda. Periquito, Fca. Miramar, Campamento 2. Bosque subandino, Bosque primario</a>, 04°30'55.9"N, 73°26'28.5"W, 1527 m, 1 ♀, 2018. iii. 14–16, Castro M. I., Neita J. C. &amp; Torres E., T. Exc. H. T 1 T 5 (48 h) (IAvH) , Medina. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.388054&amp;materialsCitation.latitude=4.5925" title="Search Plazi for locations around (long -73.388054/lat 4.5925)">Miralindo</a>, 4°35'33"N, 73°23'17"W, 550 m, 1 ♂, Fev. Mar / 1997, F. Escobar, Ex. Hum. (CALT-ECC) , Miralindo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.388054&amp;materialsCitation.latitude=4.5925" title="Search Plazi for locations around (long -73.388054/lat 4.5925)">Medina</a>, 4°35'33"N, 73°23'17"W, 1000 m, 1 ♀, 1 ♂, ii-iii. 1997, F. Escobar coll. (CMNC) ; Meta: Cubarral. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.88944&amp;materialsCitation.latitude=3.8274999" title="Search Plazi for locations around (long -73.88944/lat 3.8274999)">F. La Cabaña. B. Montano</a>, 3°49'39"N, 73°53'22.00"W, 1000 m, 1 ♀, 1 ♀, IV / 2015, M. Ramírez, PF Exc Hum (CALT-ECC) , Cubarral. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.88944&amp;materialsCitation.latitude=3.8274999" title="Search Plazi for locations around (long -73.88944/lat 3.8274999)">F. La Cabaña. B. Montano</a>, 3°49'39"N, 73°53'22.00"W, 530 m, 2 ♀, IV / 2015, M. Ramírez, PF Exc Hum (CALT-ECC) , Cubarral. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.88944&amp;materialsCitation.latitude=3.8274999" title="Search Plazi for locations around (long -73.88944/lat 3.8274999)">F. La Cabaña. B. Montano</a>, 3°49'39"N, 73°53'22.00"W, 1000 m, 2 ♀, IV / 2015, M. Ramírez, PF Exc Hum (CALT-ECC) , Cubarral. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.88944&amp;materialsCitation.latitude=3.8274999" title="Search Plazi for locations around (long -73.88944/lat 3.8274999)">F. La Cabaña. B. Montano</a>, 3°49'39"N, 73°53'22.00"W, 530 m, 2 h? #, IV / 2015, M. Ramírez, PF Exc Hum (CALT-ECC) , Cubarral. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.88944&amp;materialsCitation.latitude=3.8274999" title="Search Plazi for locations around (long -73.88944/lat 3.8274999)">F. La Cabaña. B. Montano</a>, 3°49'39"N, 73°53'22.00"W, 1000 m, 4 ♂, IV / 2015, M. Ramírez, PF Exc Hum (CALT-ECC) , Cubarral. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.88944&amp;materialsCitation.latitude=3.8274999" title="Search Plazi for locations around (long -73.88944/lat 3.8274999)">F. La Cabaña. B. Montano</a>, 3°49'39"N, 73°53'22.00"W, 530 m, 1 ♂, IV / 2015, M. Ramírez, PF Exc Hum (CALT-ECC) , Cubarral. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.88944&amp;materialsCitation.latitude=3.8274999" title="Search Plazi for locations around (long -73.88944/lat 3.8274999)">F. La Cabaña. B. Montano</a>, 3°49'39"N, 73°53'22.00"W, 1000 m, 2 ♂, IV / 2015, M. Ramírez, PF Exc Hum (CALT-ECC) , Cubarral. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.88944&amp;materialsCitation.latitude=3.8274999" title="Search Plazi for locations around (long -73.88944/lat 3.8274999)">F. La Cabaña. B. Montano</a>, 3°49'39"N, 73°53'22.00"W, 530 m, 1 ♂, IV / 2015, M. Ramírez, PF Exc Hum (CALT-ECC) , Cubarral. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.88944&amp;materialsCitation.latitude=3.8274999" title="Search Plazi for locations around (long -73.88944/lat 3.8274999)">F. La Cabaña. B. Montano</a>, 3°49'39"N, 73°53'22.00"W, 1000 m, 1 ♂, IV / 2015, M. Ramírez, PF Exc Hum (CALT-ECC) , Cubarral. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.88944&amp;materialsCitation.latitude=3.8274999" title="Search Plazi for locations around (long -73.88944/lat 3.8274999)">F. La Cabaña. B. Montano</a>, 3°49'39"N, 73°53'22.00"W, 530 m, 1 ♂, IV / 2015, M. Ramírez, PF Exc Hum (CALT-ECC) , Cubarral. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.887695&amp;materialsCitation.latitude=3.8442776" title="Search Plazi for locations around (long -73.887695/lat 3.8442776)">Vda. Alto Vergel. RN Las Palmeras. Bosque premontano</a>, 03°50'39.4"N, 73°53'15.7"W, 1500 m, 2 ♂, Abril 2015, M. Ramírez, Exc H PF (CALT-ECC) .</p><p>Diagnosis.</p><p>This species can be distinguished from all other species within the plebejum species group by having incompletely closed punctures on the anterolateral areas of the metaventral process (as Fig. 3 E), in combination with the elevated first ventrite in males (Fig. 18 E) and the distribution (II – VII) of the tubercles on the elytral apex (Fig. 19 C).</p><p>Deltochilum pauxi sp. nov. also has these punctures incompletely closed, but its first ventrite is not elevated (Fig. 16 E), it has a salient medial angle on the pronotum (Fig. 17 G) (which is rounded in D. altiventris sp. nov. (Fig. 18 B)), and the striae are broad (Fig. 16 C) (narrow in D. altiventris sp. nov. (Fig. 18 C)).</p><p>The narrow striae and the elevated first ventrite in males may lead to confusion with D. plebejum Balthasar, 1939 (Fig. 24 C) and D. nobile sp. nov. (Fig. 22 C), but D. altiventris sp. nov. can be differentiated by the apical tubercles of the elytra: on interstriae III, V, and VII in those species (Figs 23 C, 24 E) versus interstriae II – VII in D. altiventris sp. nov. (Fig. 19 C).</p><p>Description.</p><p>Major male. Size 10.30 mm. Color. Dark brown dorsally. Dark greenish-blue ventrally. Head (Fig. 18 A). Interocular distance 8 × the width of one eye. Punctures on frons separated by one or more than one diameter of each puncture and subequal or larger in size than discal punctures. Disc punctures separated by two or more diameter of each puncture. Punctures from disc towards anterior area successively slightly enlarged. Genal punctures almost with the same density and slightly larger than the discal punctures. Pronotum (Fig. 18 B). Margin between anterior and medial-lateral angle almost straight. Medial angle of pronotum rounded. Margin between medial-lateral angle and posterior-lateral angle almost straight. Disc punctures three times smaller than the anterior-lateral ones and separated by more than one diameter of each puncture. Basal punctures smaller than anterior-lateral ones and separated by one or more diameters of each puncture. Hypomera. With most of punctures incompletely closed and setigerous (Fig. 3 A, white and green arrows). Anterior punctures denser and larger than posterior ones. Elytra (Fig. 18 C). Carina of the ninth interstria almost reaching middle of elytral length. Elytral apex on interstriae II – VII with tubercles. Striae I – VII conspicuous. Width of third stria on disc ~ 1 / 16 of the distance between striae II and III. Strial punctures ~ 2 × wider than the striae. First stria slightly wider and deeper than second stria. Stria VIII conspicuous apically, laterally almost inconspicuous. Punctures on interstriae approximately separated by two diameters and almost with the same size as strial punctures. Punctures of the third interstria on disc occupying ~ 1 / 9 of the distance between striae II and III. Interstriae with shiny points mixed with the punctures. Points smaller than interstrial punctures, ~ 4 × smaller. Metaventrite (Fig. 18 D). Disc shallowly excavated, occupying the metaventral basal third. Anterolateral areas of metaventral process with punctures not completely closed and separated by one diameter of each puncture. Anterocentral area of metaventral process with few and small punctures, slightly larger in size than the discal punctures. Punctures on the lateral lobules most of them not completely closed, separated by less than one diameter of each puncture, dispersed towards the metaventral disc. Legs (Fig. 19 A, B). Protibia strongly broadens toward the apex. Profemora with punctures separated by one or less than one diameter of each puncture. Mesofemur with punctures separated by ≥ 2 × the diameter of each puncture. Mesofemur unarmed on the posterior edge. Metafemur strongly constricted basally and broadens toward the apex. Metafemur with punctures separated by ≥ 2 × the diameter of each puncture. Metatibial spur insertion elongated, with the length subequal to the basal two tasomeres. Metatibial spur fused with the metatibia. Internal margin of metatibia with a basal irregular carina. Carina reduced in the middle of the metatibial length; following this reduction, the carina rises again as an irregular carina (giving the appearance of having two carinae). Abdomen (Fig. 18 E). Ventrite I expanded posteriorly, expansion reaching the fourth ventrite; on ventrite III, the expansion narrower than the distance between mesocoxae. Basal part of the ventrite I, between the metacoxae elevated. Pygidium (Fig. 19 C). Pygidial punctures almost uniform in size and density. Punctures separated by one or less than the diameter of each puncture. Some lateral-basal punctures not completely closed. Discal punctures, on the center of the pygidium, medium sized, each puncture occupying ~ 1 / 36 of the distance between the lateral pygidial margins. Genitalia (Fig. 19 D – F). Paramera smaller than the phallobase. Apex of paramera with a denticle in lateral view. Medial area of endophallus with one endophallite.</p><p>Variation.</p><p>The female exhibits the differences described here under the sexual dimorphism section of the redescription of the species group. In females the punctures of the anterior-lateral areas of the metaventral process extend more towards the anterior-central area than in males. Some specimens have the punctures on the anterior part of the head more dispersed and / or smaller than the holotype; others a little larger and denser. In worn specimens, the striae are very difficult to see.</p><p>The minor males have the metatibial basal carina smaller. The first ventrite is less elevated. The metafemur is less broadened towards the apex. Metaspur articulate. In medium-sized males, the metatibial basal carina is larger, the first ventrite is more elevated and the metafemur is more broadened towards the apex than those of the minor males. Metaspur articulate. In the largest major males, the metatibial spur insertion is broadened apically, and the spur has a hook-like (as Fig. 1 A) and the protibia is broader in the apical half (Fig. 3 H).</p><p>Etymology.</p><p>The name, altiventris, is an adjective formed from the Latin altus meaning elevated, and the Latin ventris meaning abdomen. The name refers to the distinctly elevated first ventrite characteristic (although not exclusive) of this species.</p><p>Known distribution</p><p>(Fig. 7 A yellow triangles). Type locality: Colombia: Cundinamarca: Medina, Vda [Vereda]. Periquito, Fca [Finca]. Miramar, Campamento 2. Bosque subandino, Bosque entresacado, 04°30'58.9"N, 73°26'04.2"W, 1212 m. This species also is known from Colombia, Arauca, Boyacá, Casanare, and Meta.</p><p>Remarks.</p><p>This species has been collected in forest, sub-montane and montane forest, primary and secondary forest and one specimen was collected in pasture. This species is one of the members of the plebejum species group that is widely distributed, possessing a broad altitudinal range from 500 to probably 1,500 m.</p></div>	https://treatment.plazi.org/id/34656E6E64CD5FE0BA2FA05B7F679BCA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	González-Alvarado, Arturo;Neita, Jhon Cesar	González-Alvarado, Arturo, Neita, Jhon Cesar (2026): Eight new species of Deltochilum Eschscholtz, 1822 (subgenus D. Deltohyboma Lane, 1946) from the Eastern Cordillera of Colombia and Venezuelan Andes (Coleoptera, Scarabaeinae, plebejum species group). ZooKeys 1280: 171-229, DOI: 10.3897/zookeys.1280.182114
19CA152DFF1755AA8E5ED04CF976C30E.text	19CA152DFF1755AA8E5ED04CF976C30E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deltochilum nobile González-Alvarado & Neita 2026	<div><p>Deltochilum nobile sp. nov.</p><p>Figs 4 J, 7 A (white star), 22, 23</p><p>Type material.</p><p>Holotype: ♂, Colombia: Cesar: Agustín Codazzi, Buenos Aires. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.10529&amp;materialsCitation.latitude=10.022979" title="Search Plazi for locations around (long -73.10529/lat 10.022979)">Bosque</a>, 10.022979, -73.105296, 1200–1300 m, 14–15. VII. 2025, Chaverra, D., T. Exc. H # G 2 P 2 (ICN) . Paratypes (22 specimens): Colombia: Cesar: Agustín Codazzi, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.10529&amp;materialsCitation.latitude=10.022979" title="Search Plazi for locations around (long -73.10529/lat 10.022979)">Buenos Aires</a>. Bosque, 10.022979, -73.105296, 1200–1300 m, 3 ♀, 12 ♂, 14–15. VII. 2025, Chaverra, D., T. Exc. H # G 2 P 2 (ICN) ; La Guajira: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.67083&amp;materialsCitation.latitude=10.838472" title="Search Plazi for locations around (long -72.67083/lat 10.838472)">Barrancas. Las Pavas</a>, 10°50'18.5"N, 72°40'15"W, 1700 m, 1 ♂, 16 / 05 / 2016, A. Redondo, T. Exc. H (CBUMAG) , Barrancas. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.66964&amp;materialsCitation.latitude=10.840777" title="Search Plazi for locations around (long -72.66964/lat 10.840777)">Las Pavas</a>, 10°50'26.8"N, 72°40'10.7"W, 1708 m, 1 ♂, 16 / 05 / 2016, A. Redondo, T. Caida-Carroña (IAvH) , Barrancas. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.67789&amp;materialsCitation.latitude=10.854806" title="Search Plazi for locations around (long -72.67789/lat 10.854806)">Surimena</a>, 10°51'17.3"N, 72°40'40.4"W, 1053 m, 1 ♀, 30 / 10 / 2015, A. Redondo (CBUMAG) , Barrancas. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.67091&amp;materialsCitation.latitude=10.852973" title="Search Plazi for locations around (long -72.67091/lat 10.852973)">Surimena</a>, 10°51'10.7"N, 72°40'15.3"W, 1027 m, 1 ♀, 17 / 05 / 2016, A. Redondo, T. Caida-Carroña (IAvH) , Urumita. [Vereda Los Medios] Serranía del Perijá. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.96327&amp;materialsCitation.latitude=10.460333" title="Search Plazi for locations around (long -72.96327/lat 10.460333)">Tres Picos</a>, [10°27'37.2"N], [72°57'47.8"W], [1800 m], 2 ♀, 1 ♂, 5 / 11 / 2009, J. Cárdenas (UPTC-CE) .</p><p>Diagnosis.</p><p>This species can be distinguished from all other species within the plebejum species group by having the largest punctures on the anterior part of the head, which are larger than the punctures on the head disc (Fig. 22 A). In D. altiventris sp. nov., these punctures are only slightly larger than the punctures of the head disc (Fig. 18 A). These species can also be separated by the apical tubercles of the elytra: tubercles on interstriae III, V, and VII in D. nobile sp. nov. (Fig. 23 C) versus interstriae II – VII in D. altiventris sp. nov. (Fig. 19 C).</p><p>Description.</p><p>Medium-sized male. Size 10.40 mm. Color. Pale greenish-brown dorsally. Dark greenish-blue ventrally. Head (Fig. 22 A). Interocular distance 8 × the width of one eye. Punctures on frons separated by one or less than one diameter of each puncture and subequal in size to head discal punctures. Disc punctures separated by one or more than one diameter of each puncture. Punctures from disc towards anterior area successively larger. Genal punctures larger and denser than the discal punctures. Pronotum (Fig. 22 B). Margin between anterior and medial-lateral angle almost straight. Medial angle of pronotum rounded. Margin between medial-lateral angle and posterior-lateral angle almost straight. Disc punctures half size to anterior-lateral ones and separated by more than one diameter of each puncture. Basal punctures smaller than anterior-lateral ones and separated by approximately two diameters of each puncture. Hypomera. With ocellated and setigerous punctures on the anterior part (as Fig. 3 B, white arrow). Most of punctures incompletely closed posteriorly and without setae. Anterior punctures denser and larger than posterior ones. Elytra (Fig. 22 C). Carina of the ninth interstria almost reaching middle of elytral length. Elytral apex on interstriae III, V – VII with tubercles. Striae I – VII almost inconspicuous. Width of third stria on disc ~ 1 / 30 of the distance between striae II and III. Strial punctures ~ 2 × wider than the stria. First stria slightly wider and deeper than second stria. Stria VIII conspicuous apically and laterally and reaching carina of the ninth interstria. Punctures on interstriae approximately separated by two diameters, and are almost the same size or smaller than strial punctures. Punctures of the third interstria on disc occupying ~ 1 / 10 of the distance between striae II and III. Interstriae with shiny points mixed with the punctures. Points several times smaller than interstrial punctures. Metaventrite (Fig. 22 D). Disc shallowly excavated, occupying the metaventral basal third. Anterolateral areas of metaventral process with punctures completely closed and separated by less than one diameter of each puncture. Anterocentral area of metaventral process with few and small punctures, subequal in size to the disc punctures. Punctures on the lateral lobules not completely closed (some of them) and variable in density, dispersed towards the metaventral disc. Legs (Fig. 23 A, B). Protibia regularly broadens toward the apex. Profemora with punctures separated by less than one diameter of each puncture. Mesofemur with punctures separated by ≥ 2 × the diameter of each puncture. Mesofemur unarmed on the posterior edge. Metafemur strongly constricted basally and broadens toward the apex. Metafemur with punctures separated by ≥ 2 × the diameter of each puncture. Metatibial spur insertion almost with the length of the two basal tarsomeres. Metatibial spur clearly distinct from the metatibia. Internal margin of metatibia with a basal irregular carina. Carina reduced in the middle of the metatibial length; following this reduction, the carina rises again as an irregular carina (giving the appearance of having two carinae). Abdomen (Fig. 22 E). Ventrite I expanded posteriorly, expansion reaching the fifth ventrite; on ventrite III, the expansion narrower than the distance between mesocoxae. Basal part of the ventrite I, between the metacoxae elevated. Pygidium (Fig. 23 C). Pygidial punctures almost uniform in size and density. Punctures separated by one or less than the diameter of each puncture. Some lateral-basal punctures are not completely closed. Discal punctures on the center of the pygidium large, each puncture occupying ~ 1 / 25 of the distance between the lateral pygidial margins. Genitalia (Fig. 23 D – F). Paramera smaller than the phallobase. Apex of paramera with a denticle in lateral view. Medial area of endophallus with one endophallite.</p><p>Variation.</p><p>The female exhibits the differences described here under the sexual dimorphism section of the redescription of the species group. Minor males have the first ventrite less elevated. The metafemur is regularly broadened toward the apex, and the carina of the internal margin of the metatibia is very small and only slightly reduced in the middle. Metaspur articulate. In major males, the first ventrite is more elevated, the metafemur is strongly broadened towards the apex, and the carina of the internal margin of the metatibia is stronger. In largest males, the metatibial spur insertion is broadened apically, and the spur has a hook-like (as Fig. 1 A)</p><p>Etymology.</p><p>The name nobile is derived from the Latin nobilis meaning noble. The name is given in direct contrast to the closely related and morphologically similar species, Deltochilum plebejum (meaning common or plebeian).</p><p>Known distribution</p><p>(7 A white star). Type locality: Colombia: Cesar: Agustín Codazzi, Buenos Aires. Bosque, 10.022979, -73.105296, 1200–1300 m. This species is known from Cesar and La Guajira, Colombia.</p></div>	https://treatment.plazi.org/id/19CA152DFF1755AA8E5ED04CF976C30E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	González-Alvarado, Arturo;Neita, Jhon Cesar	González-Alvarado, Arturo, Neita, Jhon Cesar (2026): Eight new species of Deltochilum Eschscholtz, 1822 (subgenus D. Deltohyboma Lane, 1946) from the Eastern Cordillera of Colombia and Venezuelan Andes (Coleoptera, Scarabaeinae, plebejum species group). ZooKeys 1280: 171-229, DOI: 10.3897/zookeys.1280.182114
33C9C9DB076451CE9983C0C628F35C97.text	33C9C9DB076451CE9983C0C628F35C97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deltochilum parapseudoabdominale González-Alvarado & Neita 2026	<div><p>Deltochilum parapseudoabdominale sp. nov.</p><p>Figs 3 F, 4 E, 7 B (red diamonds), 12, 13</p><p>Type material.</p><p>Holotype: ♂, VEN. [EZUELA]: Edo. Aragua: Portachuela, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.6875&amp;materialsCitation.latitude=10.3475" title="Search Plazi for locations around (long -67.6875/lat 10.3475)">Rancho Grande</a>, [10°20'51"N], [67°41'15"W], 1100 m, 21–24. ii. 1971, S. Peck, limburg (CMNC) . Paratypes (33 specimens): Venezuela: Aragua: Portachuelo Pass, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.6875&amp;materialsCitation.latitude=10.3475" title="Search Plazi for locations around (long -67.6875/lat 10.3475)">Parque Nacional Henri Pittier</a>, [10°20'51"N], [67°41'15"W], 1200 m, 1 ♀, 7–13. vi. 1999, Ratcliffe, Jameson, Smith, Villatoro coll., flight interception trap (CMNC) , Portachuelo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.6875&amp;materialsCitation.latitude=10.3475" title="Search Plazi for locations around (long -67.6875/lat 10.3475)">Rancho Grande</a>, [10°20'51"N], [67°41'15"W], 1100 m, 2 ♀, 19–21. ii. 1971, S. Peck coll., human dung (CMNC) , 4 ♀, 6 ♂, 21–24. ii. 1971, S. Peck coll., Limburg (CMNC), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.542786&amp;materialsCitation.latitude=10.067056" title="Search Plazi for locations around (long -67.542786/lat 10.067056)">Racho Grande</a>, [10°4'1.40"N], [67°32'34.02"W], 1200 m, 2 ♀, 3 ♂, 27–30. viii. 1983, B Gill coll. (BDGC) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.542786&amp;materialsCitation.latitude=10.067056" title="Search Plazi for locations around (long -67.542786/lat 10.067056)">Racho Grande</a>, [10°4'1.40"N], [67°32'34.02"W], 1460 m, 2 ♀, 3 ♂, 4–14. vii. 1983, B Gill coll. (BDGC) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.68194&amp;materialsCitation.latitude=10.349722" title="Search Plazi for locations around (long -67.68194/lat 10.349722)">Rancho Grande [site 2]</a>, [10°20'59"N], [67°40'55"W], 1100 m, 1 ♀, 22–23. ii. 1971, H. &amp; A. Howden coll. (CMNC) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.67555&amp;materialsCitation.latitude=10.359445" title="Search Plazi for locations around (long -67.67555/lat 10.359445)">Rancho Grande [site 5]</a>, [10°21'34"N], [67°40'32"W], 1500 m, 1 ♀, 2 ♂, 21–25. ii. 1971, S. Peck coll., carrion trap (CMNC) , 3 ♀, 21–25. ii. 1971, S. Peck coll., human dung trap (CMNC), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.683334&amp;materialsCitation.latitude=10.333333" title="Search Plazi for locations around (long -67.683334/lat 10.333333)">Rancho Grande [site 7]</a>, [10°20'N], [67°41'W], 1460 m, 1 ♀, 1 ♂, 4–14. vii. 1986, B. Gill coll. (CMNC) , Tiara, 50 km SW <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-67.155556&amp;materialsCitation.latitude=10.129723" title="Search Plazi for locations around (long -67.155556/lat 10.129723)">Caracas</a>. forest, [10°7'47"N], [67°9'20"W], 2000 m, 1 ♂, 22–25. ii. 1971, S. Peck coll., human dung (CMNC) .</p><p>Diagnosis.</p><p>This species is very similar to D. pseudoabdominale sp. nov., D. picachos sp. nov., and D. pauxi sp. nov. by having the striae wide (Figs 10 C, 12 C, 14 C, 16 C), and the males with the first ventrite not elevated (Figs 10 E, 12 E, 14 E, 16 E) and the mesofemur not modified (Figs 11 A, 13 A, 15 A, 17 A). It can be separated from these species by having the male first ventrite wide, on the third ventrite, the width of the first ventrite is as wide as the intermesocoxal distance (Fig. 12 E), in all other species the width of the first ventrite is narrower than the intermesocoxal distance (Figs 10 E, 14 E, 16 E). Additionally, it can be differentiated from D. pseudoabdominale sp. nov. and D. picachos sp. nov. by the punctures on the anterior part of the head: they are slightly larger than the punctures of the head disc in D. parapseudoabdominale sp. nov. (Fig. 12 A), but are nearly the same size in the other two species ( D. pseudoabdominale sp. nov. and D. picachos sp. nov.) (Figs 10 A, 14 A). Furthermore, from D. pauxi sp. nov., this species can be distinguished by the medial angle of the pronotum, which is rounded in D. parapseudoabdominale sp. nov. (Fig. 12 B) and salient in D. pauxi sp. nov. (Fig. 17 G)</p><p>Description.</p><p>Minor male. Size 9.70 mm. Color. Dark brown with some blue reflections dorsally. Dark blue ventrally. Head (Fig. 12 A). Interocular distance 8 × the width of one eye. Punctures on frons separated by one or less than one diameter of each puncture and slightly larger than the head discal punctures. Disc punctures separated by one or more than one diameter of each puncture. Punctures from disc towards anterior area successively slightly enlarged. Genal punctures subequal in size and density to the discal punctures. Pronotum (Fig. 12 B). Margin between anterior and medial-lateral angle almost straight. Medial angle of pronotum rounded. Margin between medial-lateral angle and posterior-lateral angle almost straight. Disc punctures half size to anterior-lateral ones and separated by more than one diameter of each puncture. Basal punctures smaller than anterior-lateral ones and separated by one or more than one diameter of each puncture. Hypomera. With most of punctures incompletely closed and setigerous (as Fig. 3 A, white and green arrows). Anterior punctures denser and larger than posterior ones. Elytra (Fig. 12 C). Carina of the ninth interstria almost reaching middle of elytral length. Elytral apex on interstriae III, V – VII with tubercles; interstria IV with a small elevation with the same brightness as the tubercles. Striae I – VII conspicuous. Width of third stria on disc ~ 1 / 25 of the distance between striae II and III. Strial punctures almost the same size as interstrial punctures. First stria slightly wider and deeper than second stria. Stria VIII almost inconspicuous apical and laterally. Punctures on interstriae approximately separated by two diameters and the same size or slightly larger than strial punctures. Punctures of the third interstria on disc occupying ~ 1 / 13 of the distance between striae II and III. Interstriae with shiny points mixed with the punctures. Points slightly smaller than interstrial punctures. Metaventrite (Fig. 12 D). Disc shallowly excavated, occupying the metaventral basal third. On the middle with a small elevation. Anterolateral areas of metaventral process with punctures with few punctures completely closed and separated by one diameter of each puncture. Anterocentral area of metaventral process with few and small punctures, subequal in size to the discal punctures. Punctures on the lateral lobules closed and separated by one or more than one diameter of each puncture. Dispersed towards the metaventral disc. Legs (Fig. 13 A, B). Protibia regularly broadens toward the apex. Profemora with punctures separated by less than one diameter of each puncture. Mesofemur with punctures separated by ≥ 1 × the diameter of each puncture. Mesofemur unarmed on the posterior edge. Metafemur regularly broadens toward the apex. Metafemur with punctures separated by ≥ 1 × the diameter of each puncture. Metatibial spur insertion not elongated. Metatibial spur clearly distinct from the metatibia. Internal margin of metatibia with a very small basal carina. Abdomen (Fig. 12 E). Ventrite I expanded posteriorly, expansion reaching the fifth ventrite; on ventrite III, the expansion almost as wide as the distance between mesocoxae. Basal part of the ventrite I, between the metacoxae not modified. Pygidium (Fig. 13 C). Pygidial punctures almost uniform in size and density. Punctures separated by one or less than the diameter of each puncture. Discal punctures, on the center of the pygidium, medium, each puncture occupying ~ 1 / 35 of the distance between the lateral pygidial margins. Genitalia (Fig. 13 D – F). Paramera smaller than the phallobase. Apex of paramera with a small denticle in lateral view. Medial area of endophallus with one endophallite.</p><p>Variation.</p><p>The female exhibits the differences described here under the sexual dimorphism section of the redescription of the species group. The medium-sized and major males are unknown.</p><p>Etymology.</p><p>The name parapseudoabdominale is an adjective formed from the Greek prefix pará -, meaning beside or allied to, and pseudoabdominale. This name is given to denote the strong morphological similarity and close affinity to the previously described species Deltochilum pseudoabdominale sp. nov.</p><p>Known distribution</p><p>(Fig. 7 B red diamonds). Type locality: Venezuela: Aragua: Portachuelo, Rancho Grande, [10°20'51"N], [67°41'15"W], 1100 m. This species is known from Venezuela, Aragua.</p><p>Remarks.</p><p>This species appears to have a restricted distribution. This species is found in sympatry with D. abdominale . Deltochilum parapseudoabdominale sp. nov. appears to have a more restricted altitudinal range (from 1,000 to probably 2,000 m) than D. abdominale .</p></div>	https://treatment.plazi.org/id/33C9C9DB076451CE9983C0C628F35C97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	González-Alvarado, Arturo;Neita, Jhon Cesar	González-Alvarado, Arturo, Neita, Jhon Cesar (2026): Eight new species of Deltochilum Eschscholtz, 1822 (subgenus D. Deltohyboma Lane, 1946) from the Eastern Cordillera of Colombia and Venezuelan Andes (Coleoptera, Scarabaeinae, plebejum species group). ZooKeys 1280: 171-229, DOI: 10.3897/zookeys.1280.182114
28DE431D40255542986A53A8C6F2EA00.text	28DE431D40255542986A53A8C6F2EA00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deltochilum pauxi González-Alvarado & Neita 2026	<div><p>Deltochilum pauxi sp. nov.</p><p>Figs 1 C, 3 E, 3 I, 4 G, 7 B (yellow triangles), 16, 17</p><p>Type material.</p><p>Holotype: ♂, Colombia: Santander: Betulia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.44231&amp;materialsCitation.latitude=6.9705834" title="Search Plazi for locations around (long -73.44231/lat 6.9705834)">RN Pauxi pauxi</a>. Bosque, 06°58'14.1"N, 073°26'32.3"W, 1124 m, 06. iii. 2019, Ramírez, M., T. Exc. H # PPF 2 (24 h) (IAvH) [IAvH -E-270466] . Paratypes: Colombia (6 specimens): Santander: Betulia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.44111&amp;materialsCitation.latitude=6.969139" title="Search Plazi for locations around (long -73.44111/lat 6.969139)">RN Pauxi pauxi</a>. Bosque, 06°58'08.9"N, 073°26'28.0"W, 1295 m, 1 ♂, 06. iii. 2019, Ramírez, M., T. Exc. H # PPF 1 (24 h) (IAvH) , Betulia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.443306&amp;materialsCitation.latitude=6.972111" title="Search Plazi for locations around (long -73.443306/lat 6.972111)">RN Pauxi pauxi</a>. Bosque, 06°58'19.6"N, 073°26'35.9"W, 1134 m, 1 ♀, 06. iii. 2019, Ramírez, M., T. Exc. H # PPF 3 (24 h) (IAvH) , Betulia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.44111&amp;materialsCitation.latitude=6.969139" title="Search Plazi for locations around (long -73.44111/lat 6.969139)">RN Pauxi pauxi</a>. Bosque, 06°58'08.9"N, 073°26'28.0"W, 1295 m, 1 ♀, 07. iii. 2019, Ramírez, M., T. Exc. H # PPF 1 (48 h) (IAvH) , Betulia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.44622&amp;materialsCitation.latitude=6.9793887" title="Search Plazi for locations around (long -73.44622/lat 6.9793887)">RN Pauxi pauxi</a>. Bosque, 06°58'45.8"N, 073°26'46.4"W, 1096 m, 2 ♀, 09. iii. 2019, Ramírez, M., T. Exc. H # PPF 6 (96 h) (IAvH) , El Carmen de Chucurí, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.55222&amp;materialsCitation.latitude=6.557167" title="Search Plazi for locations around (long -73.55222/lat 6.557167)">Vereda La Belleza. Bosque subandino Potrero</a>, 6°33'25.8"N, 73°33'8.0"W, 1115 m, 1 ♂, 2018.2. 18 – 19, Castro M. I., Neita J. C. &amp; Torres E., T. Exc. H. (IAvH) .</p><p>Diagnosis.</p><p>This species can be distinguished from all other species within the plebejum species group by having the punctures on the anterolateral areas of the metaventral process incompletely closed (Fig. 3 E), in combination with a salient medial angle on the pronotum (Fig. 17 G) (this angle is rounded in all other species). Deltochilum altiventris sp. nov. also exhibits these incompletely closed punctures, but it differs in several ways: the male first ventrite is elevated (Fig. 18 E) (not elevated in D. pauxi sp. nov. (Fig. 16 E)), and the striae are narrow (Fig. 18 C) (broad in D. pauxi sp. nov. (Fig. 16 C)).</p><p>Description.</p><p>Medium-sized male. Size 11 mm. Color. Pale greenish-brown dorsally. Dark greenish-blue ventrally. Head (Fig. 16 A). Interocular distance 9 × the width of one eye. Punctures on frons separated by one or less than one diameter of each puncture and slightly larger than the head discal punctures. Disc punctures separated by one or more than one diameter of each puncture. Punctures from disc towards anterior area successively slightly enlarged. Genal punctures more dispersed and slightly larger than the discal punctures. Pronotum (Fig. 16 B). Margin between anterior and medial-lateral angle almost straight. Medial angle of pronotum salient. Margin between medial-lateral angle and posterior-lateral angle subsinuate. Disc punctures half size to anterior-lateral ones and separated by more than one diameter of each puncture. Basal punctures smaller than anterior-lateral ones and separated by approximately one diameter of each puncture. Hypomera. With most of punctures incompletely closed and setigerous (as Fig. 3 A, white and green arrows). Anterior punctures denser and larger than posterior ones. Elytra (Fig. 16 C). Carina of the ninth interstria almost reaching middle of elytral length. Elytral apex on interstriae III, V – VII with tubercles; interstriae I and IV with a small elevation with the same brightness as the tubercles. Striae I – VII conspicuous. Width of third stria on disc ~ 1 / 20 of the distance between striae II and III. Strial punctures ~ 2 × wider than the stria. First stria slightly wider and deeper than second stria. Stria VIII conspicuous apically and laterally and reaching carina of the ninth interstria. Punctures on interstriae approximately separated by one diameter and almost with the same size or slightly larger than strial punctures. Punctures of the third interstria on disc occupying ~ 1 / 9 of the distance between striae II and III. Interstriae with shiny points mixed with the punctures. Points smaller than interstrial punctures, ~ 4 × smaller. Metaventrite (Figs 3 E, 16 D). Disc shallowly excavated, occupying the metaventral basal third. Anterolateral areas of metaventral process with punctures not completely closed and separated by less than one diameter of each puncture (Fig. 3 E). Anterocentral area of metaventral process with few and small punctures, subequal in size to the discal punctures. Punctures on the lateral lobules most of them not completely closed, separated by less than one diameter of each puncture, dispersed towards the metaventral disc. Legs (Figs 1 C, 3 I, 17 A, 17 B). Protibia regularly broadens toward the apex (Fig. 3 I). Profemora with punctures separated by less than one diameter of each puncture. Mesofemur with punctures separated by ≥ 2 × the diameter of each puncture. Mesofemur unarmed on the posterior edge. Metafemur regularly broadens toward the apex. Metafemur with punctures separated by ≥ 2 × the diameter of each puncture. Metatibial spur insertion elongated, with the length subequal to the basal two tarsomeres (Fig. 1 C). Metatibial spur clearly distinct from the metatibia (Fig. 1 C). Internal margin of metatibia with a basal irregular carina; this carina is interrupted by setae giving it the appearance of tubercles. Carina reduced in the middle of the metatibial length; following this reduction, the carina rises again as an irregular carina (giving the appearance of having tubercles). Abdomen (Fig. 16 E). Ventrite I expanded posteriorly, expansion reaching the fifth ventrite; on ventrite III, the expansion narrower than the distance between mesocoxae. Basal part of the ventrite I, between the metacoxae not modified. Pygidium (Fig. 17 C). Pygidial punctures almost uniform in size and density. Punctures separated by one or more than the diameter of each puncture. Some lateral-basal punctures not completely closed. Discal punctures, on the center of the pygidium, small, each puncture occupying ~ 1 / 40 of the distance between the lateral pygidial margins. Genitalia (Fig. 17 D – F). Paramera smaller than the phallobase. Apex of paramera with a denticle in lateral view. Medial area of endophallus with two endophallites, the right ~ 5 × longer than the other.</p><p>Variation.</p><p>The female exhibits the differences described here under the sexual dimorphism section of the redescription of the species group. The major male (Fig. 17 H) has the basal metatibia carina larger. The medial angle of the pronotum is more salient (Fig. 17 G). The first ventrite is wider. The metafemur is strongly widened toward the apex. The metaspur is fused. Minor males are unknown.</p><p>Etymology.</p><p>The name refers to the type locality, RN Pauxi pauxi, noun in apposition.</p><p>Known distribution</p><p>(7 B yellow triangles). Type locality: Colombia: Santander: Betulia, RN Pauxi pauxi. Bosque, 06°58'14.1"N, 073°26'32.3"W, 1124 m. Besides the type locality, this species is known from El Carmen de Chucurí, Vereda La Belleza, Santander Colombia.</p><p>Remarks.</p><p>This species is currently known from two localities: the type locality and El Carmen de Chucurí, Vereda La Belleza, Santander, Colombia. The type locality represents a forested habitat, while the second appears to be situated in a transition zone between forest and pasture.</p></div>	https://treatment.plazi.org/id/28DE431D40255542986A53A8C6F2EA00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	González-Alvarado, Arturo;Neita, Jhon Cesar	González-Alvarado, Arturo, Neita, Jhon Cesar (2026): Eight new species of Deltochilum Eschscholtz, 1822 (subgenus D. Deltohyboma Lane, 1946) from the Eastern Cordillera of Colombia and Venezuelan Andes (Coleoptera, Scarabaeinae, plebejum species group). ZooKeys 1280: 171-229, DOI: 10.3897/zookeys.1280.182114
2C2A6A7FC0885663821575EED5B6A1D6.text	2C2A6A7FC0885663821575EED5B6A1D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deltochilum picachos González-Alvarado & Neita 2026	<div><p>Deltochilum picachos sp. nov.</p><p>Figs 1 D, 3 D, 4 F, 7 B (purple squares), 14, 15</p><p>Type material.</p><p>Holotype: ♂, Colombia: Caquetá: San Vicente del Caguán, Alto del río Pato, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.88333&amp;materialsCitation.latitude=2.7333333" title="Search Plazi for locations around (long -74.88333/lat 2.7333333)">Fca. Andalucía. PNN Los Picachos</a>, 02°44'N, 74°53'W, 1560 m, 1997. ix. 23, Escobar F., T. Exc. H. 14 (IAvH) [IAvH -E-209544] . Paratypes (33 specimens): Colombia: Caquetá: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.666664&amp;materialsCitation.latitude=2.8" title="Search Plazi for locations around (long -74.666664/lat 2.8)">PNN Los Picachos</a>, 02°48'N, 74°40'W, 1560 m, 1 ♀ [with aedeagus], 1998. ix, Escobar F., T. Exc. H. (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.666664&amp;materialsCitation.latitude=2.8" title="Search Plazi for locations around (long -74.666664/lat 2.8)">PNN Los Picachos</a>, 02°48'N, 74°40'W, 1780 m, 1 ♂, 1998. ix, Escobar F., T. Exc. H. (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.666664&amp;materialsCitation.latitude=2.8" title="Search Plazi for locations around (long -74.666664/lat 2.8)">PNN Los Picachos</a>, 02°48'N, 74°40'W, 1560 m, 1 ♂, 1998. ix, Escobar F., T. Exc. H. (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.666664&amp;materialsCitation.latitude=2.8" title="Search Plazi for locations around (long -74.666664/lat 2.8)">PNN Los Picachos</a>, 02°48'N, 74°40'W, 1780 m, 1 ♂, 1998. ix, Escobar F., T. Exc. H. (IAvH) , PNN Los Picachos . Bosque, 02°47'51'N, 74°51'18"W, 1770 m, 1 ♀, 1 ♂, 1997. xi – xii, Escobar F. (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.666664&amp;materialsCitation.latitude=2.8" title="Search Plazi for locations around (long -74.666664/lat 2.8)">PNN Los Picachos</a>. Bosque, 02°48'N, 74°40'W, 1780 m, 1 ♀, 1998. ix, Escobar F., T. Exc. Hum (IAvH) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.666664&amp;materialsCitation.latitude=2.8" title="Search Plazi for locations around (long -74.666664/lat 2.8)">San Vicente del Caguán – PICACHOS</a>. Bosque, [02°48'N], [74°40'W], 1800–2000 m, 3 Unsexed, Nov - 1997, F. Escobar, Hongos (CALT-ECC) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.666664&amp;materialsCitation.latitude=2.8" title="Search Plazi for locations around (long -74.666664/lat 2.8)">San Vicente del Caguán – PICACHOS</a>. Bosque, [02°48'N], [74°40'W], 1780 m, 2 Unsexed, Nov – Dec 1997, F. Escobar, T. 2 (CALT-ECC) , San Vicente del Caguán, Alto del río Pato, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.88333&amp;materialsCitation.latitude=2.7333333" title="Search Plazi for locations around (long -74.88333/lat 2.7333333)">Fca. Andalucía. PNN Los Picachos</a>, 02°44'N, 74°53'W, 1560 m, 1 ♂, 1997. ix. 23, Escobar F., T. Exc. H. 14 (IAvH) , San Vicente del Caguán, Alto del río Pato, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.88333&amp;materialsCitation.latitude=2.7333333" title="Search Plazi for locations around (long -74.88333/lat 2.7333333)">Fca. Andalucía. PNN Los Picachos</a>, 02°44'N, 74°53'W, 1780 m, 1 ♀, 1997. xi, Escobar F., T. Exc. H. 21 (IAvH) , San Vicente del Caguán, Alto del río Pato, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.88333&amp;materialsCitation.latitude=2.7333333" title="Search Plazi for locations around (long -74.88333/lat 2.7333333)">Fca. Andalucía. PNN Los Picachos</a>, 02°44'N, 74°53'W, 1560 m, 1 ♀ / ♂, 1 ♂ / ♀, 1997. xi, Escobar F., Trampa con Hongos 20 (IAvH) , San Vicente del Caguán, Alto del río Pato, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.88333&amp;materialsCitation.latitude=2.7333333" title="Search Plazi for locations around (long -74.88333/lat 2.7333333)">Fca. Andalucía. PNN Los Picachos</a>. Bosque, 02°44'N, 74°53'W, 1780 m, 1 Female / Male #, 1997. xi, Escobar F., T. Exc. H. 21 (IAvH) , San Vicente del Caguán, IP Guayabal, Alto del río Pato, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.854996&amp;materialsCitation.latitude=2.7975" title="Search Plazi for locations around (long -74.854996/lat 2.7975)">Fca. Andalucia. PNN Los Picachos</a>, 02°47'51"N, 74°51'18"W, 1560 m, 1 ♂ [without aedeagus], 1997. xi – xii, Escobar F., T. Exc. H. (IAvH) , San Vicente del Caguán, IP Guayabal, Alto del río Pato, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.666664&amp;materialsCitation.latitude=2.8" title="Search Plazi for locations around (long -74.666664/lat 2.8)">Fca. Andalucia. PNN Los Picachos</a>, 02°48'N, 74°40'W, 1560 m, 1 ♂, 1998. ix, Escobar F., T. Exc. H. (IAvH) , San Vicente del Caguán. Guayabal, PNN Los Picachos, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.85&amp;materialsCitation.latitude=2.7833333" title="Search Plazi for locations around (long -74.85/lat 2.7833333)">Alto del Río Pato, Fca. Andalucía</a>, 02°47'N, 74°51'W, 1780 m, 8 Unsexed, 1997. xi – xii, Escobar F., Trampa de caída (UPTC-CE) , San Vicente del Caguán. Guayabal, PNN Los Picachos, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.88333&amp;materialsCitation.latitude=2.7333333" title="Search Plazi for locations around (long -74.88333/lat 2.7333333)">Alto del Río Pato, Fca. Andalucía</a>, 02°44'N, 74°53'W, 1560 m, 2 Unsexed, 1997. xi – xii, Escobar F., Trampa de intercepción de vuelo (UPTC-CE) , San Vicente del Caguán. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.666664&amp;materialsCitation.latitude=2.8" title="Search Plazi for locations around (long -74.666664/lat 2.8)">PNN Los Picachos</a>. Bosque, [02°48'N], [74°40'W], 1800 m, 2 Unsexed, 1997. xi, Escobar F., Trampa de intercepción de vuelo (UPTC-CE) ; Meta: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-73.88944&amp;materialsCitation.latitude=3.8274999" title="Search Plazi for locations around (long -73.88944/lat 3.8274999)">Cubarral. F. La Cabaña. B. Montano</a>, 3°49'39"N, 73°53'22.00"W, 1000 m, 2 ♂, IV / 2015, M. Ramírez, PF Exc Hum (CALT-ECC) .</p><p>Diagnosis.</p><p>This species is very similar to D. pseudoabdominale sp. nov., D. parapseudoabdominale sp. nov., and D. pauxi sp. nov. by having the striae wide (Figs 10 C, 12 C, 14 C, 16 C), and the males with the first ventrite not elevated (Figs 10 E, 12 E, 14 E, 16 E) and the mesofemur not modified (Figs 11 A, 13 A, 15 A, 17 A). This species can be separated from D. parapseudoabdominale sp. nov. by the width of the male first ventrite: at the level of the third ventrite, the width of the first ventrite is narrower than the intermesocoxal distance in this species (Fig. 14 E), whereas it is equal to the intermesocoxal distance in D. parapseudoabdominale sp. nov. (Fig. 12 E). Additionally, they are separated by the punctures on the anterior part of the head: these punctures are slightly larger than the punctures of the head disc in D. parapseudoabdominale sp. nov. (Fig. 12 A), but are nearly the same size in D. picachos sp. nov. (Fig. 14 A).</p><p>From D. pauxi sp. nov., this species can be distinguished by the medial angle of the pronotum, which is rounded in D. picachos sp. nov. (Fig. 14 B) and salient in D. pauxi sp. nov. (Fig. 17 G). Furthermore, this species can be separated from D. pseudoabdominale sp. nov. by the apical tubercles of the elytra: these tubercles are present on interstriae III, V, and VII in Deltochilum pseudoabdominale sp. nov. (Fig. 11 C), but are present on interstriae III through VII in D. picachos sp. nov. (15 C).</p><p>Description.</p><p>Medium-sized male. Size 10 mm. Color. Dark brown. Dark greenish-blue ventrally. Head (Fig. 14 A). Interocular distance 8 × the width of one eye. Punctures on frons separated by one or less than one diameter of each puncture and larger in size than discal punctures. Disc punctures separated by two diameters of each puncture. Punctures from disc towards anterior area successively of almost with the same size. Genal punctures subequal in size and density to the discal punctures. Pronotum (Fig. 14 B). Margin between anterior and medial-lateral angle almost straight. Medial angle of pronotum rounded. Margin between medial-lateral angle and posterior-lateral angle subsinuate. Disc punctures three times smaller than the anterior-lateral ones and separated by more than one diameter of each puncture. Basal punctures smaller than anterior-lateral ones and separated by approximately one diameter of each puncture. Hypomera. With most of punctures incompletely closed and setigerous (as Fig. 3 A, white and green arrows). Anterior punctures denser and larger than posterior ones. Elytra (Fig. 14 C). Carina of the ninth interstria almost reaching middle of elytral length. Elytral apex on interstriae III – VII with tubercles. Striae I – VII conspicuous. Width of third stria on disc ~ 1 / 30 of the distance between striae II and III. Strial punctures ~ 1.5 × wider than the stria. First stria as wide as second stria. Stria VIII conspicuous apical and laterally and reaching carina of the ninth interstria. Punctures on interstriae approximately separated by two or slightly more than two diameters of each puncture and slightly larger than strial punctures. Punctures of the third interstria on disc occupying ~ 1 / 12 of the distance between striae II and III. Interstriae with shiny points mixed with the punctures. Points smaller than interstrial punctures, ~ 3 × smaller. Metaventrite (Figs 3 D, 14 D). Disc shallowly excavated, occupying the metaventral basal third. On the middle with a small elevation. Anterolateral areas of metaventral process with punctures with few punctures completely closed and separated by less than one diameter of each puncture. Anterocentral area of metaventral process with few and small punctures, slightly larger in size than the discal punctures. Punctures on the lateral lobules not completely closed (some of them) and variable in density, dispersed towards the metaventral disc. Legs (Fig. 15 A, B). Protibia regularly broadens toward the apex. Profemora with punctures separated by one or less than one diameter of each puncture. Mesofemur with punctures separated by ≥ 1 × the diameter of each puncture. Mesofemur unarmed on the posterior edge. Metafemur regularly broadens toward the apex. Metafemur with punctures separated by ≥ 1 × the diameter of each puncture. Metatibial spur insertion elongated, with the length subequal to the first tarsomere. Metatibial spur clearly distinct from the metatibia. Internal margin of metatibia with a small basal irregular carina, this carina is interrupted by setae giving it the appearance of small tubercles. Abdomen (Fig. 14 E). Ventrite I expanded posteriorly, expansion reaching the fifth ventrite; on ventrite III, the expansion narrower than the distance between mesocoxae. Basal part of the ventrite I, between the metacoxae not modified. Pygidium (Fig. 15 C). Pygidial punctures almost uniform in size and density. Punctures separated by one or less than the diameter of each puncture. Discal punctures, on the center of the pygidium, small, each puncture occupying ~ 1 / 40 of the distance between the lateral pygidial margins. Genitalia (Fig. 15 D – F). Paramera smaller than the phallobase. Apex of paramera with a denticle in lateral view. Medial area of endophallus with one endophallite.</p><p>Variation.</p><p>The female exhibits the differences described here under the sexual dimorphism section of the redescription of the species group. In the minor males, the metafemur is regularly broadened toward the apex, and the carina of the internal margin is very small and only slightly reduced in the middle. Metaspur articulate (Fig. 1 D). In the major males, the metatibia has a small basal carina. This carina is strongly reduced in the middle, giving the appearance that it is composed of two carinae. Metaspur fused.</p><p>Etymology.</p><p>The name refers to the type locality, PNN Los Picachos, noun in apposition.</p><p>Known distribution</p><p>(Fig. 7 B purple squares). Type locality: Colombia: Caquetá: San Vicente del Caguán, Alto del río Pato, Fca. Andalucía. PNN Los Picachos, 02°44'N, 74°53'W, 1560 m. Besides the type locality, this species is known from Meta, Colombia.</p><p>Remarks.</p><p>Three paratype specimens housed in the IAvH - E are chimeras, consisting of the head and pronotum of one individual joined to the meso- and metathorax and abdomen of another. For these specimens, we added labels indicating the sex of the head-pronotum and the sex of the remaining part. In the same collection one male paratype lacked the aedeagus, and one female paratype had an aedeagus glued in a triangle pinned with the specimen.</p><p>This species has been collected with traps baited with dung and fungi in forest and montane forest. Its altitudinal range extends from 1,000 to 2,000 m.</p></div>	https://treatment.plazi.org/id/2C2A6A7FC0885663821575EED5B6A1D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	González-Alvarado, Arturo;Neita, Jhon Cesar	González-Alvarado, Arturo, Neita, Jhon Cesar (2026): Eight new species of Deltochilum Eschscholtz, 1822 (subgenus D. Deltohyboma Lane, 1946) from the Eastern Cordillera of Colombia and Venezuelan Andes (Coleoptera, Scarabaeinae, plebejum species group). ZooKeys 1280: 171-229, DOI: 10.3897/zookeys.1280.182114
BEE117808E485F1C94750BFFE7AF4509.text	BEE117808E485F1C94750BFFE7AF4509.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deltochilum plebejum Balthasar 1939	<div><p>Deltochilum plebejum Balthasar, 1939</p><p>Figs 4 K – L, 7 A (red cross), 24, 25</p><p>Deltochilum (s. str.) plebejum Balthasar, 1939: 14 .</p><p>Deltochilum plebejum: Génier 2001: 6.</p><p>Type material examined.</p><p>Lectotype (here designated) ♂ Venezuela: Maracaibo Basin, [10.455446]?, [- 71.787763]? E. Gevaerts / coll Valck Lucassen / Museum Leiden verz FT Valck Lucassen / Deltochilum (s. str.) plebejum sp. nov. Dr. V. Balthasar det / [red label] TYPUS / [red label] LECTOTYPE [male symbol] Deltochilum plebejum Balth. des. F. Z. Vaz-de-Mello, 2014 (RMNH) . Paralectotype: ♀ Venezuela: Maracaibo Basin, [10.455446]?, [- 71.787763]? E. Gevaerts / coll Valck Lucassen / ex coll. V. Balthasar National Museum Prague, Czech Republic / Deltochilum spec. ? det. Boucomont / [red label] TYPUS / [green label] plebejum m. / [yellow label] PARALECTOTYPE [male symbol] Deltochilum plebejum Balth. des. F. Z. Vaz-de-Mello, 2014 (NMPC) .</p><p>Diagnosis.</p><p>This species shares the narrow striae and the apical tubercles of the elytra (on interstriae III, V, and VII) with D. nobile sp. nov. (Figs 23 C, 24 E), but it can be differentiated by the head punctures. The punctures on the anterior part of the head are nearly the same size as the head disc punctures in D. plebejum (Fig. 24 A), whereas they are larger in D. nobile sp. nov. (Fig. 22 A).</p><p>Redescription.</p><p>Female. Size 9.50 mm. Color. Pale reddish-brown dorsally. Dark greenish-blue ventrally. Head (Fig. 24 A). Interocular distance eight and a half times the width of one eye. Punctures on frons separated by less than one diameter of each puncture and slightly larger than the head discal punctures. Disc punctures separated by one or more than one diameter of each puncture. Punctures from disc towards anterior area successively almost the same size. Genal punctures subequal in size and density to the discal punctures. Pronotum (Fig. 24 B). Margin between anterior and medial-lateral angle almost straight. Medial angle of pronotum rounded. Margin between medial-lateral angle and posterior-lateral angle almost straight. Disc punctures half size to anterior-lateral ones and separated by more than one diameter of each puncture. Basal punctures smaller than anterior-lateral ones and separated by approximately one diameter of each puncture. Hypomera. With ocellated and setigerous punctures on the anterior part (as Fig. 3 B, white arrow). Most of punctures incompletely closed posteriorly and with setae (as Fig. 3 B, green arrow). Anterior punctures denser and larger than posterior ones. Elytra (Fig. 24 C). Carina of the ninth interstria not reaching middle of elytral length. Elytral apex on interstriae III, V – VII with tubercles. Striae I – VII almost inconspicuous. Width of third stria on disc ~ 1 / 40 of the distance between striae II and III. Strial punctures variable in size, ≥ 2 × wider than the stria. First stria slightly wider and deeper than second stria. Stria VIII conspicuous apically and laterally and reaching carina of the ninth interstria. Punctures on interstriae approximately separated by two diameters. Punctures of the third interstria on disc occupying ~ 1 / 7 of the distance between striae II and III. Interstriae with shiny points mixed with the punctures. Points smaller than interstrial punctures, ~ 4 × smaller. Metaventrite. Anterolateral areas of metaventral process with punctures not completely closed and separated by less than one diameter of each puncture. Anterocentral area of metaventral process with few and small punctures, slightly larger in size than the discal punctures. Punctures on the lateral lobules most of them not completely closed, separated by less than one diameter of each puncture. Legs. Protibia regularly broadens toward the apex. Profemora with punctures separated by less than one diameter of each puncture. Mesofemur with punctures separated by ≥ 2 × the diameter of each puncture. Mesofemur with unarmed on the posterior edge. Metafemur with punctures separated by ≥ 1 × the diameter of each puncture. Metatibial spur insertion not elongated. Metatibial spur clearly distinct from the metatibia. Abdomen (Fig. 24 D). Ventrite I regular, not expanded posteriorly. Pygidium (Fig. 24 E). Pygidial punctures variable in size and density. Punctures separated by one or less than the diameter of each puncture on the disc, more disperse towards the sides. Discal punctures, on the center of the pygidium, small, each puncture occupying ~ 1 / 38 of the distance between the lateral pygidial margins.</p><p>Variation.</p><p>Medium-sized male: Protibia regularly broadens toward the apex. Mesofemur unarmed on the posterior edge. Metafemur strongly constricted basally and broadens toward the apex. Metatibial spur insertion elongated, with the length subequal to the basal two tarsomeres. Metatibial spur fused with the metatibia. Internal margin of metatibia with a basal irregular carina. Carina reduced in the middle of the metatibial length; following this reduction, the carina rises again as a irregular carina (giving the appearance of having two carinae). Ventrite I expanded posteriorly, expansion reaching the fourth ventrite; on ventrite III, the expansion narrower than the distance between mesocoxae. Apparently, the basal part of the ventrite I, between the metacoxae is elevated (Fig. 25 B – C).</p><p>Known distribution</p><p>(7 A red cross). This species is only known from the type locality: the Maracaibo Basin, Venezuela. As this locality is highly unspecific, the red cross is intended only as a graphical representation of the possible distribution and, for this reason, is highlighted with a question mark (?).</p><p>Remarks.</p><p>The lectotype is designated following the Article 74 of the International Code of Zoological Nomenclature (ICZN 1999) and the recommendations herein, in order to stabilize the nomenclature.</p><p>This species is known to us only from the type series: a male lectotype (RMNH) and a female paralectotype (NMPC) (although this specimen bears a male symbol in the paralectotype label). The now-designated lectotype was originally housed in the Valck Lucassen collection in Vorden (Holland) (Balthasar 1939). After Mr. F. T. Valck Lucassen’s death in September 1939, his collection was transported to the Leiden Museum (De Jong 1970). The paralectotype was originally housed in Balthasar collection and is now housed in NMPC (Génier 2001). The paralectotype was reviewed in person and the redescription here was made using that specimen. The lectotype was examined by a photographs kindly provided by Dr. Fernando Vaz de Mello (Fig. 4 K) and Dr. Olivier Montreuil (Fig. 25) It is not clearly visible, but apparently the first ventrite is elevated (Fig. 25 B, C).</p><p>We consider this species to be closely related to D. nobile sp. nov. based on the size of the striae, the size and density of the interstrial punctures, the number and distribution of the tubercles on the elytral apex, the shape of the internal carina of the metatibia, the apparent elevation of the first ventrite, and the distribution.</p></div>	https://treatment.plazi.org/id/BEE117808E485F1C94750BFFE7AF4509	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	González-Alvarado, Arturo;Neita, Jhon Cesar	González-Alvarado, Arturo, Neita, Jhon Cesar (2026): Eight new species of Deltochilum Eschscholtz, 1822 (subgenus D. Deltohyboma Lane, 1946) from the Eastern Cordillera of Colombia and Venezuelan Andes (Coleoptera, Scarabaeinae, plebejum species group). ZooKeys 1280: 171-229, DOI: 10.3897/zookeys.1280.182114
6BB1BD39F3085A8B8EFD5947EA6F1268.text	6BB1BD39F3085A8B8EFD5947EA6F1268.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deltochilum pseudoabdominale González-Alvarado & Neita 2026	<div><p>Deltochilum pseudoabdominale sp. nov.</p><p>Figs 1 B, 4 D, 7 B (blue circles), 10, 11</p><p>Type material.</p><p>Holotype: ♂, VEN. [EZUELA]: Mérida: ULA Biological Reserve, La Carbonerra [Carbonera?], 20 km SE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.409164&amp;materialsCitation.latitude=8.656388" title="Search Plazi for locations around (long -71.409164/lat 8.656388)">Azulita</a>. Podocarpus forest, [8°39'23"N], [71°24'33"W], 2300 m, 28. vi- 3. viii. [19] 89, S. &amp; J. Peck, malaise (CMNC) [WSD 00039446] . Paratypes (48 specimens): Colombia: Arauca: Tame. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.86583&amp;materialsCitation.latitude=6.523611" title="Search Plazi for locations around (long -71.86583/lat 6.523611)">Brisas el Cravo</a>. Bosque, 06°31'25"N, 71°51'57"W, 2000 m, 1 ♀, 14–19 / II / 08, F. Alvarado, E. H. (CALT-ECC) , Tame. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.52945&amp;materialsCitation.latitude=6.5169444" title="Search Plazi for locations around (long -71.52945/lat 6.5169444)">Brisas el Cravo</a>. Bosque, 06°31'01"N, 71°31'46"W, 1000 m, 1 ♀, 20–25 / XI / 07, F. Alvarado, E. H. (CALT-ECC) ; Norte de Santander: Toledo, IP Santa María, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.21667&amp;materialsCitation.latitude=7.116667" title="Search Plazi for locations around (long -72.21667/lat 7.116667)">Alto de Herrera, Vda. Diamante, Fca. La Primavera. PNN Tamá</a>, 7°7'N, 72°13'W, 1450 m, 4 ♀, 5 ♂, 1999. ix, González E., T. Exc. H. (IAvH) , Toledo, IP Santa María, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.21667&amp;materialsCitation.latitude=7.1" title="Search Plazi for locations around (long -72.21667/lat 7.1)">Alto de Herrera, Vda. Diamante, Fca. La Primavera. PNN Tamá</a>, 7°6'N, 72°13'W, 1000 m, 1 ♀, 1999. ix. 22, González E., T. Exc. H. (IAvH) , Toledo, IP Santa María, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.21667&amp;materialsCitation.latitude=7.1" title="Search Plazi for locations around (long -72.21667/lat 7.1)">Alto de Herrera, Vda. Diamante, Fca. La Primavera. PNN Tamá</a>, 7°6'N, 72°13'W, 1250 m, 1 ♂, 1999. ix. 30, González E., T. Exc. H. (IAvH) , Toledo, IP Santa María, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.21667&amp;materialsCitation.latitude=7.116667" title="Search Plazi for locations around (long -72.21667/lat 7.116667)">Alto la Herrera, Vda. Diamante, Fca. La Primavera. PNN Tamá. Bosque secundario conservado</a>, 7°7'N, 72°13'W, 1450 m, 2 ♂, 1999. ix. 29, González E., T. Exc. H. 10 (IAvH) , Toledo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.30786&amp;materialsCitation.latitude=7.1875" title="Search Plazi for locations around (long -72.30786/lat 7.1875)">Vda. San Alberto</a>. Bosque, 07°11'15.0"N, 072°18'28.3"W, 1104 m, 1 ♂, 14. iv. 2019, Ramírez, M. &amp; Lopera, A., T. Exc. H # SAF 3 (96 h) (IAvH) , Toledo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.303696&amp;materialsCitation.latitude=7.192528" title="Search Plazi for locations around (long -72.303696/lat 7.192528)">Vda. San Alberto</a>. Bosque, 07°11'33.1"N, 072°18'13.3"W, 1358 m, 1 ♂, 14. iv. 2019, Ramírez, M. &amp; Lopera, A., T. Exc. H # SAF 5 (96 h) (IAvH) , Toledo. PNN Tamá, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.21667&amp;materialsCitation.latitude=7.116667" title="Search Plazi for locations around (long -72.21667/lat 7.116667)">Santa María Alto de Herrera, Vda. El Diamante, Finca La Primavera</a>, 7°7'N, 72°13'W, 1000 m, 2 ♀, 1 ♂, 1999. x. 25, González, E. Colecta Manual (UPTC-CE) , Toledo. PNN Tamá, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.21667&amp;materialsCitation.latitude=7.116667" title="Search Plazi for locations around (long -72.21667/lat 7.116667)">Santa María Alto de Herrera, Vda. El Diamante, Finca La Primavera</a>. Bosque, 7°7'N, 72°13'W, 1000 m, 2 ♀, 1999. x. 09, González, E. Trampa de excremento humano (UPTC-CE) . Venezuela: Lara: Yacambu N. P. 14 km SE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.566666&amp;materialsCitation.latitude=9.7" title="Search Plazi for locations around (long -69.566666/lat 9.7)">Sinaré</a>, 9°42'N, 69°34'W, 1650 m, 2 ♀, 1 ♂, v. 1998, J. Ashe, R Brooks &amp; R Hanley coll., F. I. T. (BDGC) , Yacambu N. P. 17 km SE <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.566666&amp;materialsCitation.latitude=9.7" title="Search Plazi for locations around (long -69.566666/lat 9.7)">Sinaré</a>, 9°42'N, 69°34'W, 1510 m, 2 ♀, 18. v. 1998, J. Ashe, R Brooks &amp; R Hanley coll., F. I. T. (BDGC) ; Táchira: La Idea, Campamento Siberia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.76667&amp;materialsCitation.latitude=8.016666" title="Search Plazi for locations around (long -71.76667/lat 8.016666)">Pregonero</a>. rainforest, [8°1'N], [71°46'W], 1200 m, 1 ♂, 10–31. vii. 1989, S. &amp; J. Peck coll., flight interception trap (CMNC) , <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.20573&amp;materialsCitation.latitude=7.7586083" title="Search Plazi for locations around (long -72.20573/lat 7.7586083)">San Cristóbal</a>, [7°45'30.99"N], [72°12'20.62"W], 1200 m, 12 ♀, 7 ♂, 10–18. viii. 1983, B Gill coll. (BDGC) , Via Chorro del Indio, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.19195&amp;materialsCitation.latitude=7.7222223" title="Search Plazi for locations around (long -72.19195/lat 7.7222223)">San Cristóbal</a>, [7°43'20"N], [72°11'31"W], 1 ♀, iv. 1987, D. Havranek coll. (CMNC) .</p><p>Diagnosis.</p><p>This species is very similar to D. parapseudoabdominale sp. nov., D. picachos sp. nov. and D. pauxi sp. nov. by having the striae wide (Figs 10 C, 12 C, 14 C, 16 C), and the males with the first ventrite not elevated (Figs 10 E, 12 E, 14 E, 16 E) and the mesofemur not modified (Figs 11 A, 13 A, 15 A, 17 A). This species can be separated from D. parapseudoabdominale sp. nov. by the width of the male first ventrite: at the level of the third ventrite, the width of the first ventrite is narrower than the intermesocoxal distance in this species (Fig. 10 E), whereas it is equal to the intermesocoxal distance in D. parapseudoabdominale sp. nov. (Fig. 12 E). Additionally, they are separated by the punctures on the anterior part of the head: these punctures are slightly larger than the punctures of the head disc in D. parapseudoabdominale sp. nov. (Fig. 12 A), but are nearly the same size in D. pseudoabdominale sp. nov. (Fig. 10 A).</p><p>From D. pauxi sp. nov., this species can be distinguished by the medial angle of the pronotum, which is rounded in D. pseudoabdominale sp. nov. (Fig. 10 B) and salient in D. pauxi sp. nov. (Fig. 17 G). Furthermore, this species can be separated from D. picachos sp. nov. by the apical tubercles of the elytra: tubercles on interstriae III, V, and VII in D. pseudoabdominale sp. nov. (Fig. 11 C), whereas are present on interstriae III – VII in D. picachos sp. nov. (Fig. 15 C).</p><p>Description.</p><p>Minor male. Size 10.30 mm. Color. Dark brown dorsally. Dark blue ventrally. Head (Fig. 10 A). Interocular distance 8 × the width of one eye. Punctures on frons separated by one or less than one diameter of each puncture and slightly larger in size than discal punctures. Disc punctures separated by more than one diameter of each puncture. Punctures from disc towards anterior area successively of almost with the same size. Genal punctures subequal in size and density to the discal punctures. Pronotum (Fig. 10 B). Margin between anterior and medial-lateral angle almost straight. Medial angle of pronotum rounded. Margin between medial-lateral angle and posterior-lateral angle subsinuate. Disc punctures half size to anterior-lateral ones and separated by more than one diameter of each puncture. Basal punctures smaller than anterior-lateral ones and separated by approximately two diameters of each puncture. Hypomera. With ocellated and setigerous punctures on the anterior part (as Fig. 3 B, white arrow). Most of punctures incompletely closed posteriorly and without setae. Anterior punctures denser and larger than posterior ones. Elytra (Fig. 10 C). Carina of the ninth interstria almost reaching middle of elytral length. Elytral apex on interstriae III, V – VII with tubercles. Striae I – VII conspicuous. Width of third stria on disc ~ 1 / 24 of the distance between striae II and III. Strial punctures ~ 2 × wider than the stria. First stria slightly wider and deeper than second stria. Stria VIII almost inconspicuous apically and laterally and reaching carina of the ninth interstria. Punctures on interstriae approximately separated by two diameters and the same size or slightly larger than strial punctures. Punctures of the third interstria on disc occupying ~ 1 / 12 of the distance between striae II and III. Interstriae with shiny points mixed with the punctures. Points smaller than interstrial punctures, almost half the size. Metaventrite (Fig. 10 D). Disc shallowly excavated, occupying the metaventral basal third. Anterolateral areas of metaventral process with punctures completely closed and separated by less than one diameter of each puncture. Anterocentral area of metaventral process with few and small punctures, subequal in size to the discal punctures. Punctures on the lateral lobules closed and variable in density, dispersed towards the metaventral disc. Legs (Fig. 11 A, B). Protibia regularly broadens toward the apex. Profemora with punctures separated by less than one diameter of each puncture. Mesofemur with punctures separated by 2 × the diameter of each puncture. Mesofemur unarmed on the posterior edge. Metafemur regularly broadens toward the apex. Metafemur with punctures separated by 2 × the diameter of each puncture. Internal margin of metatibia with a small carina on the basal third. Metatibial spur insertion not elongated. Metatibial spur clearly distinct from the metatibia. Abdomen (Fig. 10 E). Ventrite I expanded posteriorly, expansion reaching the fifth ventrite; on ventrite III, the expansion narrower than the distance between mesocoxae. Basal part of the ventrite I, between the metacoxae not modified. Pygidium (Fig. 11 C). Pygidial punctures variable in size and density. Punctures separated by one or less than the diameter of each puncture on the disc, more disperse towards the apex. Discal punctures, on the center of the pygidium, medium, each puncture occupying ~ 1 / 35 of the distance between the lateral pygidial margins. Genitalia (Fig. 11 D – F). Paramera smaller than the phallobase. Apex of paramera with a denticle in lateral view. Medial area of endophallus with one endophallite.</p><p>Variation.</p><p>The female exhibits the differences described here under the sexual dimorphism section of the redescription of the species group. Medium-sized male: the metatibial basal carina is stronger. This carina is significantly reduced in the middle of the metatibial length; following this reduction, the carina rises again (giving the appearance of having two carinae) (Fig. 11 G). Metafemur strongly broadened towards the apex. Metaspur articulate. Major males: the metatibial basal carina stronger. This carina is significantly reduced in the middle of the metatibial length; following this reduction, the carina rises again (giving the appearance of having two carinae). Metafemur strongly broadened toward the apex. Metaspur fused (Fig. 1 B).</p><p>Etymology.</p><p>The name, pseudoabdominale, is an adjective derived from the Greek prefix pseudēs -, meaning false or resembling, and abdominale. This name is given in reference to the sympatry of this new species with Deltochilum abdominale .</p><p>Known distribution</p><p>(Fig. 7 B blue circles). Type locality: Venezuela: Mérida: ULA Biological Reserve, La Carbonerra [Carbonera?], 20 km SE Azulita. Podocarpus forest, [8°39'23"N], [71°24'33"W], 2300 m. This species in known from Colombia, Arauca, Norte de Santander and Venezuela, Lara, Mérida y Táchira.</p><p>Remarks.</p><p>This species has been collected in forest, conserved secondary forest, rainforest, and in Podocarpus forest. This species is one of the members of the plebejum species group that is widely distributed alongside D. abdominale . These two species are found in sympatry. Deltochilum pseudoabdominale sp. nov. appears to have a more restricted altitudinal range (from 1,000 to probably 1,400 m) than D. abdominale .</p></div>	https://treatment.plazi.org/id/6BB1BD39F3085A8B8EFD5947EA6F1268	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	González-Alvarado, Arturo;Neita, Jhon Cesar	González-Alvarado, Arturo, Neita, Jhon Cesar (2026): Eight new species of Deltochilum Eschscholtz, 1822 (subgenus D. Deltohyboma Lane, 1946) from the Eastern Cordillera of Colombia and Venezuelan Andes (Coleoptera, Scarabaeinae, plebejum species group). ZooKeys 1280: 171-229, DOI: 10.3897/zookeys.1280.182114
6526FD6131595BA99C054F407091639B.text	6526FD6131595BA99C054F407091639B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deltochilum tyba González-Alvarado & Neita 2026	<div><p>Deltochilum tyba sp. nov.</p><p>Figs 4 B, 4 C, 7 A (red diamonds), 8, 9</p><p>Type material.</p><p>Holotype: ♂, Colombia: Santander: Capitanejo. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.68831&amp;materialsCitation.latitude=6.6098056" title="Search Plazi for locations around (long -72.68831/lat 6.6098056)">Bosque Ribera</a>, 6°36'35.3"N, 72°41'17.9"W, 1256 m, 9 / 3 / 15, N. J. Beltran, Pitfall Exc Hum (Ex- CALT-ECC) [IAvH -E-268920] . Paratype: ♂, Colombia: Santander: Capitanejo. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-72.68831&amp;materialsCitation.latitude=6.6098056" title="Search Plazi for locations around (long -72.68831/lat 6.6098056)">Bosque Ribera</a>, 6°36'35.3"N, 72°41'17.9"W, 1256 m, 9 / 3 / 15, N. J. Beltran, Pitfall Exc Hum (CALT-ECC) .</p><p>Diagnosis.</p><p>This species (Fig. 9 G) and D. abdominale Martínez, 1947 (Fig. 6 A, G), are the only two species in which males possess a denticle on the mesofemur. They can be distinguished by the interstrial punctures: these punctures are larger than the interstrial shiny points in D. tyba sp. nov. (Fig. 8 C), and are nearly the same size as the interstrial shiny points in D. abdominale (Fig. 5 C).</p><p>Description.</p><p>Minor male. Size 10 mm. Color. Pale green dorsally. Brown ventrally. Head (Fig. 8 A). Interocular distance 10 × the width of one eye. Punctures on frons separated by one or more than one diameter of each puncture and subequal in size than discal punctures. Disc punctures separated by one or more than one diameter of each puncture. Punctures from disc towards anterior area successively slightly enlarged. Genal punctures subequal in size and density to the discal punctures. Pronotum (Fig. 8 B). Margin between anterior and medial-lateral angle almost straight. Medial angle of pronotum rounded. Margin between medial-lateral angle and posterior-lateral angle almost straight. Disc punctures half size to anterior-lateral ones and separated by more than one diameter of each puncture. Basal punctures smaller than anterior-lateral ones and separated by approximately one diameter of each puncture. Hypomera. With ocellated and setigerous punctures on the anterior part (as Fig. 3 B, white arrow). Most of punctures incompletely closed posteriorly and with setae. Anterior punctures denser and larger than posterior ones. Elytra (Fig. 8 C). Carina of the ninth interstria not reaching middle of elytral length. Elytral apex on interstriae II – VII with tubercles. Striae I – VII almost inconspicuous. Width of third stria on disc ~ 1 / 32 of the distance between striae II and III. Strial punctures variable in size, ≥ 2 × wider than the stria. First stria slightly wider and deeper than second stria. Stria VIII conspicuous apically and laterally, and reaching carina of the ninth interstria. Punctures on interstriae approximately separated by two diameters. Punctures of the third interstria on disc occupying ~ 1 / 10 of the distance between striae II and III. Interstriae with shiny points mixed with the punctures. Points smaller than interstrial punctures, ~ 3 × smaller. Metaventrite (Fig. 8 D). Disc shallowly excavated, occupying the metaventral basal third. Anterolateral areas of metaventral process with punctures completely closed and separated by one diameter of each puncture. Anterocentral area of metaventral process with few and small punctures, slightly larger in size than the discal punctures. Punctures on the lateral lobules not completely closed (some of them) and variable in density, dispersed towards the metaventral disc. Legs (Fig. 9 A, B). Protibia regularly broadens toward the apex. Profemora with punctures separated by less than one diameter of each puncture. Mesofemur with punctures separated by ≥ 2 × the diameter of each puncture. Mesofemur with a basal denticle on the posterior edge. Metafemur regularly broadens toward the apex. Metafemur with punctures separated by ≥ 2 × the diameter of each puncture. Metatibial spur insertion elongated, with the length subequal to the basal two tarsomeres. Metatibial spur clearly distinct from the metatibia. Internal margin of metatibia with tubercles on the basal half of the metatibial length. Abdomen (Fig. 8 E). Ventrite I expanded posteriorly, expansion reaching the fourth ventrite; on ventrite III, the expansion narrower than the distance between mesocoxae. Basal part of the ventrite I, between the metacoxae elevated. Pygidium (Fig. 9 C). Pygidial punctures almost uniform in size and density. Punctures separated by one or more than the diameter of each puncture. Discal punctures on the center of the pygidium, medium, each puncture occupying ~ 1 / 36 of the distance between the lateral pygidial margins. Genitalia (Fig. 9 D – F). Paramera smaller than the phallobase. Apex of paramera with a denticle in lateral view. Medial area of endophallus with two endophallites, the right ~ 7 × longer than the other.</p><p>Variation.</p><p>The medium-sized paratype (Figs 4 C, 9 J) is a teneral and exhibits the following characteristics: the denticle of the mesofemur larger and more curved (Fig. 9 G, white arrow). The metatibial tubercles are larger (Fig. 9 I, white arrow). The first ventrite is more elevated (Fig. 9 H). Metaspur articulate. The major male and female are unknown.</p><p>Etymology.</p><p>The name tyba is a noun in apposition, derived from the Muisca (Chibcha) indigenous word ‘ tyba’ or ‘ tybá’, meaning ‘ Great Captain’ or ‘ Cacique’ (chief). This name is given to reference the type locality, Capitanejo is the diminutive of Capitán, used in allusion to a local indigenous leader.</p><p>Known distribution</p><p>(Fig. 7 A red diamonds). This species is only known from the type locality: Colombia: Santander: Capitanejo. Bosque Ribera, 6°36'35.3"N, 72°41'17.9"W, 1256 m.</p></div>	https://treatment.plazi.org/id/6526FD6131595BA99C054F407091639B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	González-Alvarado, Arturo;Neita, Jhon Cesar	González-Alvarado, Arturo, Neita, Jhon Cesar (2026): Eight new species of Deltochilum Eschscholtz, 1822 (subgenus D. Deltohyboma Lane, 1946) from the Eastern Cordillera of Colombia and Venezuelan Andes (Coleoptera, Scarabaeinae, plebejum species group). ZooKeys 1280: 171-229, DOI: 10.3897/zookeys.1280.182114
24AC097D44A95F378D915C0917D419C9.text	24AC097D44A95F378D915C0917D419C9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Deltochilum ventripuncturatus González-Alvarado & Neita 2026	<div><p>Deltochilum ventripuncturatus sp. nov.</p><p>Figs 1 A, 4 I, 7 A (purple squares), 20, 21</p><p>Deltochilum (Deltohyboma) abdominalis Martínez 1954: 47 (cited from Colombia, error).</p><p>Type material.</p><p>Holotype: ♂, Colombia: Magdalena: Santa Marta, Minca, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.03189&amp;materialsCitation.latitude=11.084694" title="Search Plazi for locations around (long -74.03189/lat 11.084694)">Cerro Kennedy</a>. Bosque, 11°05'04.9"N, 074°01'54.8"W, 1908 m, 2018. i. 29, Martínez D., T. Exc. H. SLF 4 (72 h) (IAvH) [IAvH -E-209518] . Paratypes (28 specimens): Colombia: Mag. [Magdalena]: Sra. Nev. Sta. Marta Cerro Kennedy – <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.0212&amp;materialsCitation.latitude=11.081828" title="Search Plazi for locations around (long -74.0212/lat 11.081828)">Vda. Minas, B. de sombra con cafetal</a>, [11°4'54.58"N], [74°1'16.32"W], 1280 m, 2 ♀, 1 ♂, Enero 5 / 1999, J. A. Noriega, T. pitfall Ex. Hum. Gradiente Altit. (CEMT) , Sra. Nev. Sta. Marta Cerro Kennedy – <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.0212&amp;materialsCitation.latitude=11.081828" title="Search Plazi for locations around (long -74.0212/lat 11.081828)">Vda. Minas, B. maduro con cafetal</a>, [11°4'54.58"N], [74°1'16.32"W], 1670 m, 1 ♀, Enero 2 / 1999, J. A. Noriega, T. pitfall Ex. Hum. Gradiente Altit. (CALT-ECC) ; Magdalena: Santa Marta, Minca, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.03189&amp;materialsCitation.latitude=11.084694" title="Search Plazi for locations around (long -74.03189/lat 11.084694)">Cerro Kennedy</a>. Bosque, 11°05'04.9"N, 074°01'54.8"W, 1908 m, 5 ♀, 5 ♂, 2018. i. 29, Martínez D., T. Exc. H. SLF 4 (72 h) (IAvH) , Santa Marta, Minca, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.03347&amp;materialsCitation.latitude=11.083777" title="Search Plazi for locations around (long -74.03347/lat 11.083777)">Cerro Kennedy</a>. Bosque, 11°05'01.6"N, 074°02'00.5"W, 1914 m, 1 ♂, 2018. i. 29, Martínez D., T. Exc. H. SLF 5 (48 h) (IAvH) , 1 ♀, 1 ♂, 2018. i. 29, Martínez D., T. Exc. H. SLF 5 (72 h) (IAvH), Santa Marta, Minca, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.03189&amp;materialsCitation.latitude=11.084694" title="Search Plazi for locations around (long -74.03189/lat 11.084694)">Cerro Kennedy</a>. Bosque, 11°05'04.9"N, 074°01'54.8"W, 1908 m, 2 ♀, 3 ♂, 2018. i. 30, Martínez D., T. Exc. H. SLF 4 (96 h) (IAvH) , Santa Marta, Minca, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.03347&amp;materialsCitation.latitude=11.083777" title="Search Plazi for locations around (long -74.03347/lat 11.083777)">Cerro Kennedy</a>. Bosque, 11°05'01.6"N, 074°02'00.5"W, 1914 m, 3 ♂, 2018. i. 30, Martínez D., T. Exc. H. SLF 5 (96 h) (IAvH) , Santa Marta, Minca, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.03347&amp;materialsCitation.latitude=11.085667" title="Search Plazi for locations around (long -74.03347/lat 11.085667)">Cerro Kennedy</a>. Bosque, 11°05'08.4"N, 074°02'00.5"W, 1973 m, 1 ♀, 1 ♂, 2018. i. 30, Martínez D., T. Exc. H. SLF 6 (96 h) (CBUMAG) , Santa Marta, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.082405&amp;materialsCitation.latitude=11.072969" title="Search Plazi for locations around (long -74.082405/lat 11.072969)">Mts. Vista Nieva</a>, [11°4'22.69"N], [74°4'56.66"W], 5400 ft. m, 1 ♂, 26. vi. 1920, F Saige &amp; M coll. (BDGC) .</p><p>Diagnosis.</p><p>This species can be distinguished from all other species within the plebejum species group by the unique combination of characters in males: the mesofemur possesses a basal protuberance (Fig. 21 A, G), the first ventrite is elevated and with small punctures (Figs 20 E, 21 I, 21 J). Also, it can be separated from all other species of the plebejum species group by the head punctures, the punctures on the anterior part of the head are smaller than the punctures of the head disc (Fig. 20 A), in the other species of plebejum species group, the punctures on the anterior part of the head are equal to or larger than the punctures of the head disc.</p><p>Description.</p><p>Medium-sized male. Size 10.70 mm. Color. Pale greenish-brown dorsally. Dark copperish-green ventrally. Head (Fig. 20 A). Interocular distance 8 × the width of one eye. Punctures on frons separated by one or less than one diameter of each puncture and subequal in size than discal punctures. Disc punctures separated by two diameters of each puncture. Punctures from disc towards anterior area successively smaller. Genal punctures larger and more dispersed than the discal punctures. Pronotum (Fig. 20 B). Margin between anterior and medial-lateral angle almost straight. Medial angle of pronotum rounded. Margin between medial-lateral angle and posterior-lateral angle almost straight. Disc punctures slightly smaller than anterior-lateral ones and separated by more than one diameter of each puncture. Basal punctures smaller than anterior-lateral ones and separated by approximately two diameters of each puncture. Hypomera. With most of punctures incompletely closed and setigerous (as Fig. 3 A, white and green arrows). Anterior punctures denser and larger than posterior ones. Elytra (Fig. 20 C). Carina of the ninth interstria almost reaching middle of elytral length. Elytral apex on interstriae II – VII with tubercles. Striae I – VII almost inconspicuous. Width of third stria on disc ~ 1 / 30 of the distance between striae II and III. Strial punctures variable in size, ≥ 3 × wider than the stria. First stria as wider as second stria. Stria VIII conspicuous apically and laterally and reaching carina of the ninth interstria. Punctures on interstriae approximately separated by two diameters and almost with the same size or smaller than strial punctures. Punctures of the third interstria on disc occupying ~ 1 / 15 of the distance between striae II and III. Interstriae with shiny points mixed with the punctures. Points 4 × smaller than interstrial punctures. Metaventrite (Fig. 20 D). Disc shallowly excavated, occupying the metaventral basal third. Anterolateral areas of metaventral process with punctures completely closed and separated by one diameter of each puncture. Anterocentral area of metaventral process with few and small punctures, larger in size to the discal punctures. Punctures on the lateral lobules not completely closed (some of them) and variable in density, separated by one or more than one diameter of each puncture. Legs (Fig. 21 A, B). Protibia strongly broadens toward the apex. Profemora with punctures separated by one or less than one diameter of each puncture. Mesofemur with punctures separated by ≥ 2 × the diameter of each puncture. Mesofemur with a basal protuberance in the basal third of the posterior edge. Metafemur regularly broadens toward the apex. Metafemur with punctures separated by ≥ 2 × the diameter of each puncture. Metatibial spur insertion elongated, with the length subequal to the first tarsomere. Metatibial spur clearly distinct from the metatibia. Internal margin of metatibia with large and irregular tubercles reaching the basal half of the metatibial length. Abdomen (Fig. 20 E). Ventrite I expanded posteriorly, expansion reaching the fourth ventrite; on ventrite III, the expansion narrower than the distance between mesocoxae. Basal part of the ventrite I, between the metacoxae elevated and with small and dense punctures. Pygidium (Fig. 21 C). Pygidial punctures almost uniform in size and density. Punctures separated by several times the diameter of each puncture. Some lateral-basal punctures not completely closed. Discal punctures on the center of the pygidium, medium, each puncture occupying ~ 1 / 33 of the distance between the lateral pygidial margins. Genitalia (Fig. 21 D – F). Paramera smaller than the phallobase. Apex of paramera with a denticle in lateral view. Medial area of endophallus with a large endophallite.</p><p>Variation.</p><p>The female exhibits the differences described here under the sexual dimorphism section of the redescription of the species group. Furthermore, in females, the medial part of the first ventrite bears punctures, but these are the same size as the other punctures of that ventrite. These medial punctures are not as small and dense as those found in males. The clypeal teeth are closer to one another in females than in males.</p><p>In minor males, the mesofemur bears a smaller basal protuberance. The tubercles on the metatibia are smaller and more regular in size. The first ventrite is less elevated. The small punctures of the first ventrite are less conspicuous. Metaspur articulate. In major males, the mesofemur bears strongest basal protuberance (Fig. 21 G). The metatibia is more strongly curved (Fig. 21 K). The metatibial basal carina is strongly interrupted by setae, giving it the appearance of large tubercles (Fig. 21 K, white arrow). This carina is significantly reduced in the middle of the metatibial length; following this reduction, the carina rises again (giving the appearance of having both tubercles and carina) (Fig. 21 K, white arrow). The first ventrite is more elevated (Fig. 21 I – J). Metaspur fused. In the largest males, the metatibial spur insertion is broadened apically, and the spur has a hook-like shape (Fig. 1 A).</p><p>Etymology.</p><p>The name, ventripuncturatus, is an adjective derived from the Latin ventris meaning abdomen and puncturatus meaning punctured. The name refers to the presence of small and dense punctures on the first abdominal ventrite, a unique diagnostic character of this species.</p><p>Known distribution</p><p>(Fig. 7 A purple squares). Type locality: Colombia: Magdalena: Santa Marta, Minca, Cerro Kennedy. Bosque, 11°05'04.9"N, 074°01'54.8"W, 1908 m. This species is only known from the Sierra Nevada de Santa Marta, Colombia.</p><p>Remarks.</p><p>We were able to examine a male specimen deposited in the BDGC collection that shares the exact collecting data (locality, date, collector) of the two specimens reported by Martínez (1954) from Santa Marta, Colombia, under the name D. abdominale . Upon revision, this specimen does not correspond to D. abdominale; it is definitively D. ventripuncturatus sp. nov. This finding suggests that the historical record of D. abdominale from Santa Marta, Colombia, is likely based on a misidentified specimen of D. ventripuncturatus sp. nov.</p><p>This species appears to be restricted to the Sierra Nevada de Santa Marta (Colombia). It has been collected from 1,200 to ~ 2,000 m.</p></div>	https://treatment.plazi.org/id/24AC097D44A95F378D915C0917D419C9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	González-Alvarado, Arturo;Neita, Jhon Cesar	González-Alvarado, Arturo, Neita, Jhon Cesar (2026): Eight new species of Deltochilum Eschscholtz, 1822 (subgenus D. Deltohyboma Lane, 1946) from the Eastern Cordillera of Colombia and Venezuelan Andes (Coleoptera, Scarabaeinae, plebejum species group). ZooKeys 1280: 171-229, DOI: 10.3897/zookeys.1280.182114
