identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
CE53F1FC7D085F48AB4E7E0B3EA754EF.text	CE53F1FC7D085F48AB4E7E0B3EA754EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyllena hermes Radea & Protopapas 2026	<div><p>Cyllena hermes Radea &amp; Protopapas, 2026 sp. nov.</p><p>Figs 4, 5, 6, 7, 8</p><p>Type material.</p><p>Holotype. Ethanol-fixed specimen, deposited in the National Museum of Natural History Goulandris (GNHM), accession number 51-01-045021 . Paratypes (from the same lot). One ethanol-fixed specimen deposited in the National Museum of Natural History Goulandris (GNHM), accession number 51-01-045022 . Four mature ethanol-fixed specimens dissected for anatomical study and one immature specimen coated for SEM in the personal collection of C. Radea deposited in the Section of Ecology and Systematics, Department of Biology, National and Kapodistrian University of Athens.</p><p>Diagnosis.</p><p>As for genus.</p><p>Description.</p><p>Shell (Figs 4 A – E, G – R, 5). Valvatiform to planorboid, minute (maximum height 0.76 mm, maximum width 1.46 mm) (Table 1) with up to 2.25 whorls, thin-walled, colourless, glossy, transparent when fresh; spire depressed, blunt; periostracum pale corneous; aperture roundish to slightly ovate, slightly prosocline adhering to the last whorl; peristome continuous, thickened and reflected at columellar margin, the outer margin simple; umbilicus open and extremely wide, revealing the inner (earlier) whorls; teleoconch smooth, bearing fine growth lines with the last portion of the dilated body whorl descending near aperture at an angle of approximately 30 °; protoconch of approximately one whorl, slightly elevated, with a broad nucleus; microsculpture of the embryonic shell consisting of shallow, circular to oval pits with concave sides; growth lines visible after the first embryonic whorl, while the pits gradually fade; protoconch – teleoconch transition indistinct.</p><p>Operculum (Fig. 4 F). Circular, paucispiral, thin, corneous, orange-colored; inner face weakly convex, without any peg; nucleus central.</p><p>Head (Figs 6 E, 8 A – J). White-creamy coloured, without dark pigments; eye spots absent; snout about as wide as long, non-tapered, with strong distal lobation; tentacles (in live specimens) parallel-sided and approximately ten times longer than wide.</p><p>Ctenidium – Osphradium. Absent.</p><p>Radula. Unknown</p><p>Rectum. Rectum forming a U-shaped bend, wider in females than in males. Faecal pellets ellipsoidal, light orange-colored.</p><p>Anterior female reproductive system (Figs 6 A, 6 B, 7 A). Albumen and capsule glands very small; bursa copulatrix pyriform with bursal duct thick and short (bursal duct length less than 50 % of bursa length); coiled oviduct unpigmented, thick, with irregular external surface, forming a single, open, circular loop, fully protruding from posterior end of albumen gland; a single seminal receptacle (proximal, rs 2) large and irregular in shape.</p><p>Penis (Figs 6 C – E, 7 B). Large relative to head size, elongate, dorso-ventrally flattened, with a proximal portion bearing distinct folds and a distal portion with blunt apex; a single elongate and blunt lobe at middle-length on the left side; penial duct weakly undulating at base and straight distally, near-centrally positioned, opening in an apical papilla.</p><p>Etymology.</p><p>The specific name, used in apposition, is derived from Greek mythology. Hermes (Ἑρμῆς in Greek) was one of the twelve Olympian gods, the messenger of the other Olympian deities, and was born in a cave on Mt Kyllini.</p><p>Distribution and habitat.</p><p>So far, the distribution of Cyllena hermes gen. et sp. nov. is restricted to the type locality. The area is characterized by carbonate bedrock, and owing to karstification of the limestone, groundwater emerges through karstic springs, forming a small stream flowing towards Lake Stymphalia (Figs 1, 2). In several sections, the stream bed is partially covered by Helosciadium nodiflorum (L.) W. D. J. Koch . The surrounding landscape is dominated by herbaceous vegetation, while the riparian zone along the stream is sparsely lined by Platanus sp. The streambed substrate consists of sand and predominantly angular to subangular clasts, ranging from gravel to boulder size, mainly of limestone origin and derived from the adjacent mountainous massif.</p><p>Associated freshwater gastropod species.</p><p>Radomaniola sp., Bythinella sp., Isimerope semele Radea &amp; Parmakelis, 2013 and Daphniola longipenia Grego &amp; Falniowski, 2021 were also recorded in the type locality.</p><p>Remarks.</p><p>Two female and two male specimens were dissected. One female bears a pseudopenis on the right side of the head.</p><p>In one male, the distal penial portion is somewhat dilated (Fig. 6 C).</p><p>Individuals of the species were observed performing circular movements while feeding inside the transport container during transfer from the sampling site to the laboratory (Fig. 8 A – J).</p><p>Specimens of Cyllena hermes gen. et sp. nov., Radomaniola sp., Bythinella sp., I. semele and D. longipenia were observed incorporated into larval cases of caddisflies ( Insecta, Trichoptera) (Fig. 8 K). The new locality record of the last two endemic hydrobiids extends their currently documented distribution in the Peloponnese.</p></div>	https://treatment.plazi.org/id/CE53F1FC7D085F48AB4E7E0B3EA754EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Radea, Canella;Protopapas, Dimitrios;Parmakelis, Aristeidis;Koskeridou, Efterpi	Radea, Canella, Protopapas, Dimitrios, Parmakelis, Aristeidis, Koskeridou, Efterpi (2026): From the dark to the light: A new genus and species of stygobiont hydrobiid (Caenogastropoda, Truncatelloidea) from southern Greece. Subterranean Biology 57: 1-21, DOI: 10.3897/subtbiol.57.189090
C52D896B9E585644A4DF97ACD591461B.text	C52D896B9E585644A4DF97ACD591461B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyllena Radea & Protopapas 2026	<div><p>Cyllena Radea &amp; Protopapas, 2026 gen. nov.</p><p>Type species.</p><p>Cyllena hermes sp. nov. by original designation.</p><p>Diagnosis.</p><p>Shell minute, valvatiform to planorboid; umbilicus wide, with earlier whorls visible; soft body unpigmented; eyes absent; bursa copulatrix pyriform; a single seminal receptacle [proximal, rs 2 (Radoman 1973)] large and irregular in shape; penis large, dorso-ventrally flattened, distally blunt with papilla and proximally folded, bearing a single, elongate, blunt, medial lobe.</p><p>Differential diagnosis.</p><p>Cyllena gen. nov. is distinguished from other stygobiont valvatiform and valvatiform – planorboid genera that possess a bursa copulatrix and a single seminal receptacle ( Arganiella Fo. Giusti &amp; Pezzoli, 1980; Hadziella Kuščer, 1932; Hauffenia Pollonera, 1898; Heraultiella Bodon, Manganelli &amp; Fo. Giusti, 2002; Kerkia Radoman, 1978; Zaumia Radoman, 1983). This separation is based primarily on the configuration of the female genital tract and the distinctive morphology of the penis.</p><p>Cyllena gen. nov. further differs from genera with unknown female genitalia ( Alagoniella Quiñonero-Salgado, López-Soriano, Raven &amp; Sánchez, 2025; Alfahariella Quiñonero-Salgado, López-Soriano, Raven, Glöer &amp; Prieto, 2024; Berkanella Taybi, Glöer &amp; Mabrouki, 2024; Maroccoarganiella Ghamizi &amp; Falniowski, 2024; Onubiella Martín Álvarez, Quiñonero-Salgado, López-Soriano, Raven &amp; Glöer, 2024; Robustiella Quiñonero-Salgado, López-Soriano &amp; García-Meseguer, 2025; Salaeniella Boeters, Quiñonero-Salgado &amp; Ruiz-Cobo, 2019; Siuraniella Quiñonero-Salgado, López-Soriano, Raven, Pascual, García, Solé &amp; Sánchez, 2025; and Vilertia López-Soriano, Quiñonero-Salgado, Suárez, Rolán &amp; Glöer, 2022) in the shape and lobation of the penis.</p><p>Finally, the new genus differs from Dalmatella Velkovrh, 1970 and Navalis Quiñonero-Salgado &amp; Rolán, 2017 in having a nearly smooth teleoconch lacking ornamentation.</p><p>In addition, Cyllena gen. nov. is distinguished from other stygobiont valvatiform and valvatiform – planorboid genera by several supplementary characters, including the absence of an operculum with a peg ( Bracenica Radoman, 1973; Hauffenia), a differently shaped rectum ( Arganiella; Corbellaria Callot-Girardi &amp; Boeters, 2012; Fissuria Boeters, 1981; Hadziella; Hauffenia; Heraultiella; Kerkia; Pezzolia Bodon &amp; Giusti, 1986; Salaeniella; Sardohoratia Manganelli, Bodon, Cianfanelli, Talenti &amp; Fo. Giusti, 1998), and the sculpture of the protoconch ( Fissuria and Pezzolia).</p><p>Type-locality.</p><p>Greece, Peloponnese: (37°52.06'N, 22°27.65'E), karstic spring situated at ca 610 m elevation a. s. l. on the <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=22.460833&amp;materialsCitation.latitude=37.86767" title="Search Plazi for locations around (long 22.460833/lat 37.86767)">foothills of Mt Kyllini</a>, 11 January 2026, D. Protopapas leg.</p><p>Etymology.</p><p>The generic name derives from the Greek mythology: Cyllena (Κυλλήνη in Greek) was a nymph of Mt Kyllini, Korinthia, Peloponnese, and the nurturer of the god Hermes. Gender feminine.</p></div>	https://treatment.plazi.org/id/C52D896B9E585644A4DF97ACD591461B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Radea, Canella;Protopapas, Dimitrios;Parmakelis, Aristeidis;Koskeridou, Efterpi	Radea, Canella, Protopapas, Dimitrios, Parmakelis, Aristeidis, Koskeridou, Efterpi (2026): From the dark to the light: A new genus and species of stygobiont hydrobiid (Caenogastropoda, Truncatelloidea) from southern Greece. Subterranean Biology 57: 1-21, DOI: 10.3897/subtbiol.57.189090
