identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
964BC0C7260550E39F55EAE90EADB66C.text	964BC0C7260550E39F55EAE90EADB66C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Niveomyces J. P. M. Araujo & de Bekker	<div><p>Niveomyces J. P. M. Araújo &amp; de Bekker, Persoonia 49: 186 (2022)</p><p>Type.</p><p>Niveomyces coronatus J. P. M. Araújo &amp; de Bekker, Persoonia 49: 188 (2022).</p><p>Description.</p><p>Parasitic on Ophiocordyceps sp. Sexual morph: Undetermined. Asexual morph: Niveomyces species are characterized by white to pale yellow, cottony to velvety mycelium that typically covers the host. The synnemata are enveloped by a hymenium-like layer of densely arranged conidiogenous cells, which are polyblastic, cylindrical to elongated, sometimes irregular or capitate, and bear multiple crowded denticles on the apical region, forming a distinctive crown-like apex. Conidia are produced singly on the denticles and are hyaline, smooth-walled, aseptate, and varying from globose or ovoid to ellipsoidal or cylindrical depending on the species.</p><p>Notes.</p><p>Species of Niveomyces share highly similar morphological characteristics, particularly in their conidiogenous cells, which bear multiple denticles along the phialides and form a crown-like apex, producing conidia singly (Araújo et al. 2022; Kobmoo et al. 2023). Their distinctions lie mainly in conidial size and shape, which remain the primary diagnostic characters separating species within the genus (Kobmoo et al. 2023). In this study, one new species, Niveomyces pseudoalbus, is introduced, supported by both phylogenetic evidence and detailed morphological observations.</p></div>	https://treatment.plazi.org/id/964BC0C7260550E39F55EAE90EADB66C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Xian;Peng, Xing-Can;Mapook, Ausana;Hyde, Kevin D.;Wei, De-Ping;Liu, Zheng-Hui;Saichana, Natsaran;Wen, Ting-Chi	Zhang, Xian, Peng, Xing-Can, Mapook, Ausana, Hyde, Kevin D., Wei, De-Ping, Liu, Zheng-Hui, Saichana, Natsaran, Wen, Ting-Chi (2026): Sorobiellomyces jilinensis gen. et sp. nov. (Polycephalomycetaceae) and Niveomyces pseudoalbus sp. nov. (Cordycipitaceae): Two hyperparasitic fungi associated with Ophiocordyceps. MycoKeys 134: 47-73, DOI: 10.3897/mycokeys.134.194187
9C90C5EA17FE5378AC0C765A569F95F9.text	9C90C5EA17FE5378AC0C765A569F95F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Niveomyces pseudoalbus X. Zhang, T. C. Wen & K. D. Hyde 2026	<div><p>Niveomyces pseudoalbus X. Zhang, T. C. Wen &amp; K. D. Hyde sp. nov.</p><p>Fig. 5</p><p>Etymology.</p><p>Reference to its close morphological resemblance to, but distinction from, its sister species Niveomyces albus .</p><p>Holotype.</p><p>China • Yunnan Province: Tengchong City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.264786&amp;materialsCitation.latitude=25.189608" title="Search Plazi for locations around (long 98.264786/lat 25.189608)">Houqiao Town</a>, 25°11'22.59"N, 98°15'53.24"E, alt. 2148 m, 18 October 2024, Xian Zhang, HQLSP 1 (HKAS 125706, holotype) .</p><p>Description.</p><p>The host is enveloped by dense, white, cottony mycelium that covers the stromata of Ophiocordyceps dipterigena sensu lato . Sexual morph: undetermined. Asexual morph: Hyphae are septate, hyaline, smooth-walled, and irregularly branched. Conidiophores are cylindrical, septate, simple, and either dichotomously or irregularly branched, with considerable variation in length. Conidiogenous cells arise directly from the hyphae, cylindrical, measuring 12.3–65.7 × 2–3.2 μm (x ̄ = 33.2 × 2.5 μm, n = 20), and bear an irregular, geniculate rachis. Conidia are produced singly on denticles, oval to ellipsoidal with an apiculus, hyaline, smooth-walled, aseptate, and measuring 6.4–10.4 × 1.8–3.4 μm (x ̄ = 7.6 × 2.7 μm, n = 40).</p><p>Habitat.</p><p>The Ophiocordyceps dipterigena sensu lato on the abaxial surface of leaves.</p><p>Additional material examined.</p><p>China • Yunnan Province: Tengchong City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.264786&amp;materialsCitation.latitude=25.189608" title="Search Plazi for locations around (long 98.264786/lat 25.189608)">Houqiao Town</a>, 25°11'22.59"N, 98°15'53.24"E, alt. 2148 m, on Ophiocordyceps dipterigena sensu lato, 18 October 2024, Xian Zhang, HQLSP 2 (HKAS 125705, paratype) .</p><p>Notes.</p><p>Based on multilocus phylogenetic analysis, Niveomyces pseudoalbus (HKAS 152706 and HKAS 152705) formed a highly supported clade with N. albus (100 % ML / 1.00 PP; Fig. 2). Although these two species share similar morphology, N. pseudoalbus possesses more slender conidia (6.4–10.4 × 1.8–3.4 μm) than those of N. albus (5–7 × 1–2 μm). The conidia of N. albus are obtuse at both ends, whereas those of N. pseudoalbus are distinctly acute at one end. In addition, N. pseudoalbus (HKAS 152706) differs from N. albus (BCC 83025) by 29 / 834 bp (3 %, 2 gaps) in LSU and 13 / 908 bp (1 %, 0 gaps) in tef 1 - α. Overall, phylogenetic evidence supports recognizing N. pseudoalbus as a new species following the guidelines of Jayawardena et al. (2021).</p></div>	https://treatment.plazi.org/id/9C90C5EA17FE5378AC0C765A569F95F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Xian;Peng, Xing-Can;Mapook, Ausana;Hyde, Kevin D.;Wei, De-Ping;Liu, Zheng-Hui;Saichana, Natsaran;Wen, Ting-Chi	Zhang, Xian, Peng, Xing-Can, Mapook, Ausana, Hyde, Kevin D., Wei, De-Ping, Liu, Zheng-Hui, Saichana, Natsaran, Wen, Ting-Chi (2026): Sorobiellomyces jilinensis gen. et sp. nov. (Polycephalomycetaceae) and Niveomyces pseudoalbus sp. nov. (Cordycipitaceae): Two hyperparasitic fungi associated with Ophiocordyceps. MycoKeys 134: 47-73, DOI: 10.3897/mycokeys.134.194187
74FA7FA4C8285DE1A911045DEA3583CF.text	74FA7FA4C8285DE1A911045DEA3583CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ophiocordyceps jilinensis Y. P. Xiao & Y. Yang	<div><p>Ophiocordyceps jilinensis Y. P. Xiao &amp; Y. Yang, in Cao et al., Fungal Diversity 132: 489 (2025)</p><p>Fig. 4</p><p>Description.</p><p>Parasitic on adult Orthoptera. Sexual morph: Undetermined. Asexual morph: Hyphomycetous. Hyphae hyaline, septate, with warts. Phialides monophialidic arise laterally or terminally from the hyphae. Phialides 22.3–29.0 × 1–4.5 µm (x ̄ = 26.3 × 2.4 µm, n = 30), with swollen base and slender neck, hyaline, cylindrical, and bear small warts. Conidia hyaline, smooth, ellipsoid to lemon shaped, arising from the apex of the neck singly, 4.5–7.1 × 4–5.1 μm (x ̄ = 6.2 × 4.5 μm, n = 30), each enclosed within a mucous sheath.</p><p>Habitat.</p><p>Adult Orthoptera on the branches.</p><p>Additional material examined.</p><p>China • Jilin Province: Baishan City, Fusong County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=127.9&amp;materialsCitation.latitude=42.55" title="Search Plazi for locations around (long 127.9/lat 42.55)">Lushuihe Village</a>, 42°33'N, 127°54'E, alt. 774 m, on adult Orthoptera, 19 August 2024, Xian Zhang, H 51 (HKAS 149968) .</p><p>Notes.</p><p>Based on a phylogenetic analysis of concatenated ITS, SSU, LSU, tef 1 - α, rpb 1, and rpb 2 sequences, sample HKAS 149968 grouped in a clade with the type species of Ophiocordyceps jilinensis (JL 2421), with strong statistical support (100 % ML / 1.00 PP; Fig. 1). Morphologically, phialides in Ophiocordyceps jilinensis (JL 2421) are variable, comprising both monophialidic and polyphialidic forms (Cao et al. 2025), whereas sample HKAS 149968 was observed to possess only monophialidic phialides. This difference is interpreted as intraspecific variation and does not contradict the molecular evidence.</p></div>	https://treatment.plazi.org/id/74FA7FA4C8285DE1A911045DEA3583CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Xian;Peng, Xing-Can;Mapook, Ausana;Hyde, Kevin D.;Wei, De-Ping;Liu, Zheng-Hui;Saichana, Natsaran;Wen, Ting-Chi	Zhang, Xian, Peng, Xing-Can, Mapook, Ausana, Hyde, Kevin D., Wei, De-Ping, Liu, Zheng-Hui, Saichana, Natsaran, Wen, Ting-Chi (2026): Sorobiellomyces jilinensis gen. et sp. nov. (Polycephalomycetaceae) and Niveomyces pseudoalbus sp. nov. (Cordycipitaceae): Two hyperparasitic fungi associated with Ophiocordyceps. MycoKeys 134: 47-73, DOI: 10.3897/mycokeys.134.194187
4D7D677A53FE50CFAEC961CCF3257DC2.text	4D7D677A53FE50CFAEC961CCF3257DC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ophiocordyceps Petch	<div><p>Ophiocordyceps Petch, Trans. Brit. Mycol. Soc. 16 (1): 74 (1931)</p><p>Type.</p><p>Ophiocordyceps blattae (Petch) Petch, Trans. Br. mycol. Soc. 16 (1): 74 (1931).</p><p>Description.</p><p>See Sung et al. (2007).</p><p>Notes.</p><p>Ophiocordyceps was established by Petch (1931) for species of Cordyceps bearing non-disarticulating ascospores, but it was subsequently treated as a subgenus by later authors (Kobayasi 1941; Mains 1958). The genus was revalidated and formally delimited through multigene phylogenetic analyses by Sung et al. (2007), who also proposed Ophiocordycipitaceae, with Ophiocordyceps as the type genus. A comprehensive revision of Ophiocordycipitaceae by Quandt et al. (2014) further refined the circumscription and phylogenetic placement of Ophiocordyceps within Hypocreales . Within this phylogenetic framework, the present study describes a new collection, O. jilinensis, supported by multilocus phylogenetic data and distinctive morphological characters.</p></div>	https://treatment.plazi.org/id/4D7D677A53FE50CFAEC961CCF3257DC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Xian;Peng, Xing-Can;Mapook, Ausana;Hyde, Kevin D.;Wei, De-Ping;Liu, Zheng-Hui;Saichana, Natsaran;Wen, Ting-Chi	Zhang, Xian, Peng, Xing-Can, Mapook, Ausana, Hyde, Kevin D., Wei, De-Ping, Liu, Zheng-Hui, Saichana, Natsaran, Wen, Ting-Chi (2026): Sorobiellomyces jilinensis gen. et sp. nov. (Polycephalomycetaceae) and Niveomyces pseudoalbus sp. nov. (Cordycipitaceae): Two hyperparasitic fungi associated with Ophiocordyceps. MycoKeys 134: 47-73, DOI: 10.3897/mycokeys.134.194187
99E3F16CCBFC52F5A01D2A22B3D7CC38.text	99E3F16CCBFC52F5A01D2A22B3D7CC38.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sorobiellomyces jilinensis X. Zhang, Q. F. Huang, T. C. Wen & K. D. Hyde 2026	<div><p>Sorobiellomyces jilinensis X. Zhang, Q. F. Huang, T. C. Wen &amp; K. D. Hyde, sp. nov.</p><p>Fig. 3</p><p>Holotype.</p><p>China • Jilin Province: Baishan City, Fusong County, Lushuihe Village, 42°51'N, 127°78'E, 19 August 2024, Xian Zhang, H 42-1 (HKAS 149967, holotype) .</p><p>Etymology.</p><p>Reference to Jilin Province, the locality where the type specimen was collected.</p><p>Description.</p><p>Parasitic on Ophiocordyceps jilinensis . Sexual morph: Undetermined. Asexual morph: Synnemata never observed. Loose white hyphae cover the fungal hyphal mat on the insect. Phialides hirsutella-like, arising from aerial hyphae, cylindrical to subulate, base slightly swollen to inconspicuous, tapering gradually towards a narrow apex, mono- to polyphialidic, smooth, hyaline, 12.5–25.9 × 1–2.4 μm (x ̄ = 18.7 × 1.8 μm, n = 30). Conidia 1.8–3 × 1.2–2.7 µm (x ̄ = 2.4 × 1.7 µm, n = 30), one-celled, hyaline, smooth-walled, ovoid.</p><p>Habitat.</p><p>On Ophiocordyceps jilinensis parasitizing an insect host.</p><p>Additional material examined.</p><p>China • Jilin Province: Baishan City, Fusong County, Lushuihe Village, 42°51'N, 127°78'E, on Ophiocordyceps jilinensis on Orthoptera, 19 August 2024, Xian Zhang, H 42-2 (HKAS 152704, paratype) .</p><p>Notes.</p><p>The newly proposed species, Sorobiellomyces jilinensis, represents a previously undescribed hirsutella-like fungus that forms a strongly supported sister lineage (100 % ML / 1.0 PP; Fig. 1) to Torrubiellomyces zombiae . Ecologically, S. jilinensis parasitizes Ophiocordyceps jilinensis, whereas the type species T. zombiae infects the entomopathogenic fungus Ophiocordyceps camponoti-floridani . Morphologically, S. jilinensis lacks a known sexual morph, and its asexual morph is produced directly on the host, forming a prominent, superficial, white to yellowish hyphal mat. In contrast, T. zombiae produces conspicuous, blackened perithecia on the host, and its asexual morph is primarily observed in culture. A key diagnostic difference lies in the phialides of S. jilinensis, which are consistently longer (12.5–25.9 μm compared to 5.5–12 μm in T. zombiae) and explicitly noted as branched (Araújo et al. 2022). Based on this distinct phylogenetic position and unique morphological traits, Sorobiellomyces jilinensis is proposed as the type species of the new genus Sorobiellomyces .</p></div>	https://treatment.plazi.org/id/99E3F16CCBFC52F5A01D2A22B3D7CC38	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Xian;Peng, Xing-Can;Mapook, Ausana;Hyde, Kevin D.;Wei, De-Ping;Liu, Zheng-Hui;Saichana, Natsaran;Wen, Ting-Chi	Zhang, Xian, Peng, Xing-Can, Mapook, Ausana, Hyde, Kevin D., Wei, De-Ping, Liu, Zheng-Hui, Saichana, Natsaran, Wen, Ting-Chi (2026): Sorobiellomyces jilinensis gen. et sp. nov. (Polycephalomycetaceae) and Niveomyces pseudoalbus sp. nov. (Cordycipitaceae): Two hyperparasitic fungi associated with Ophiocordyceps. MycoKeys 134: 47-73, DOI: 10.3897/mycokeys.134.194187
348383C98456583E971E83F64E522299.text	348383C98456583E971E83F64E522299.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sorobiellomyces X. Zhang, Q. F. Huang, T. C. Wen & K. D. Hyde 2026	<div><p>Sorobiellomyces X. Zhang, Q. F. Huang, T. C. Wen &amp; K. D. Hyde, gen. nov.</p><p>Type.</p><p>Sorobiellomyces jilinensis X. Zhang, Q. F. Huang, T. C. Wen &amp; K. D. Hyde, sp. nov.</p><p>Etymology.</p><p>Soro- (L.) refers to sister, indicating its phylogenetic position as the sister genus to Torrubiellomyces, - ubiella is derived from the name of the sister genus.</p><p>Description.</p><p>Hyperparasitic, parasitic on Ophiocordyceps jilinensis . Sexual morph: Undetermined. Asexual morph: The host is covered by a superficial, white to yellowish hyphal mat, with loose white hyphae present on the surface of the Ophiocordyceps mat. Phialides are cylindrical to subulate, frequently slightly curved, and taper gradually toward the apex, mostly branched. Conidia are unicellular, hyaline, smooth-walled, and ovoid.</p><p>Notes.</p><p>Sorobiellomyces and Torrubiellomyces are both monotypic genera with hosts belonging to Ophiocordyceps (Araújo et al. 2022) . Multigene phylogenetic analysis confirms that they form distinct monophyletic lineages, supporting their status as separate genera. Morphologically, Sorobiellomyces is distinguished from Torrubiellomyces by several key characteristics of its phialides. The phialides of Torrubiellomyces are cylindrical to subulate, usually slightly curved, and taper gradually toward the apex. In contrast, those of Sorobiellomyces are consistently longer, hirsutella-like, and branched (Araújo et al. 2022). In addition, the asexual morph of Sorobiellomyces has been observed directly on natural specimens, whereas in species of Torrubiellomyces, the asexual morph has so far been reported only from cultures.</p></div>	https://treatment.plazi.org/id/348383C98456583E971E83F64E522299	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Zhang, Xian;Peng, Xing-Can;Mapook, Ausana;Hyde, Kevin D.;Wei, De-Ping;Liu, Zheng-Hui;Saichana, Natsaran;Wen, Ting-Chi	Zhang, Xian, Peng, Xing-Can, Mapook, Ausana, Hyde, Kevin D., Wei, De-Ping, Liu, Zheng-Hui, Saichana, Natsaran, Wen, Ting-Chi (2026): Sorobiellomyces jilinensis gen. et sp. nov. (Polycephalomycetaceae) and Niveomyces pseudoalbus sp. nov. (Cordycipitaceae): Two hyperparasitic fungi associated with Ophiocordyceps. MycoKeys 134: 47-73, DOI: 10.3897/mycokeys.134.194187
