identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
48B437F33CCE5BB0A674916CF4BD1D5C.text	48B437F33CCE5BB0A674916CF4BD1D5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eurycanthablatta Fritzsche and Zompro 2008	<div><p>3.2. 5. Eurycanthablatta Fritzsche and Zompro, 2008 (incertae sedis)</p><p>Remarks.</p><p>Fritzsche &amp; Zompro established that Eurycanthablatta genitalia were, in part, similar to both Cryptocercus and Lamproblatta . Unfortunately, due to the lack of a more detailed description, or imaging of these characters, they are not systematically useful. Based on the data in the original paper, we think it is reasonable to assume this taxon is Blattoidea and indeed could be closely related to Xylophagodea. Given that it is so morphologically different from both Cryptocercus, and the other Lamproblattidae, we think it is possible that Eurycanthablatta could represent a new deep lineage. Yet, without material for examination or further details about the types of Eurycanthablatta pugionata, we cannot make systematic progress on this taxon. We consider it to be incertae sedis in Kittrickea based on the limited information available.</p></div>	https://treatment.plazi.org/id/48B437F33CCE5BB0A674916CF4BD1D5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Medina-Espinoza, Emmy Fiorella;Mendieta Yáñez, Stephany Karla;Drager, Kim;Swichtenberg, Kali L.;Gile, Gillian;Kang, Yifei;Hromádka, Jiří;Vanker, Katharine;Sanchez-Herrera, Melissa	Evangelista, Dominic A., Medina-Espinoza, Emmy Fiorella, Mendieta Yáñez, Stephany Karla, Drager, Kim, Swichtenberg, Kali L., Gile, Gillian, Kang, Yifei, Hromádka, Jiří, Vanker, Katharine, Sanchez-Herrera, Melissa (2026): Xylomycophagy in a close relative of termites: Lamproblattidae (Blattodea) systematics, ecology, and diet. Arthropod Systematics & Phylogeny 84: 309-342, DOI: 10.3897/asp.84.e144506
C1132434F3475179A39C1C6DE4E0F87C.text	C1132434F3475179A39C1C6DE4E0F87C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lamproblatta flavomaculata Princis 1946	<div><p>3.2. 3. Lamproblatta flavomaculata Princis, 1946</p><p>Remarks.</p><p>By Princis (1946) ’ s description, and our examination of photos of types, we can say that this species is indeed similar to other Lamproblatta spp., but with two main departures. First and most notable, the coloration (bearing yellow stripes and spots laterally on the dorsum, and deep mahogany brown body instead of black) is entirely unique among Lamproblattidae and appears more like some Eurycotis spp. (e. g., E. abdominalis). Second, the styli are noticeably longer than other Lamproblatta spp., although perhaps not as elongated as in Eurycotis (Fig. 4). Otherwise, all other external features appear similar to Lamproblatta . We have obtained images of the type specimen from Lund University Biological Museum (available on https://cockroach.speciesfile.org/otus/863559).</p><p>Our cladistic analyses had L. flavomaculata in various places, but it was always within Lamproblattidae s. s. (Fig. 7).</p></div>	https://treatment.plazi.org/id/C1132434F3475179A39C1C6DE4E0F87C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Medina-Espinoza, Emmy Fiorella;Mendieta Yáñez, Stephany Karla;Drager, Kim;Swichtenberg, Kali L.;Gile, Gillian;Kang, Yifei;Hromádka, Jiří;Vanker, Katharine;Sanchez-Herrera, Melissa	Evangelista, Dominic A., Medina-Espinoza, Emmy Fiorella, Mendieta Yáñez, Stephany Karla, Drager, Kim, Swichtenberg, Kali L., Gile, Gillian, Kang, Yifei, Hromádka, Jiří, Vanker, Katharine, Sanchez-Herrera, Melissa (2026): Xylomycophagy in a close relative of termites: Lamproblattidae (Blattodea) systematics, ecology, and diet. Arthropod Systematics & Phylogeny 84: 309-342, DOI: 10.3897/asp.84.e144506
EBC6A7F851335BE198B8B95A7D44230C.text	EBC6A7F851335BE198B8B95A7D44230C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lamproblatta Hebard 1919	<div><p>3.2. 2. Lamproblatta Hebard, 1919</p><p>Taxonomic scope.</p><p>Lamproblatta albipalpus Hebard, 1919; L. ancistroides Rehn, 1930; L. flavomaculata Princis, 1946; L. gorgonis Rehn, 1930; L. meridionalis (Bruner, 1906); L. mimadelfi sp. nov.; L. mimetes Rehn, 1930 = L. mimetis [sic] Rehn, 1930; L. romani Rehn, 1930; L. zamorensis (Giglio-Tos, 1898); L. antoni Evangelista, Kotyková Varadínová and Jůna 2019; L. neuque Sánchez Herrera, De Martino, Realpe Sanabria, Realpe Rebolledo, 2025 .</p><p>Diagnosis.</p><p>(modified from Roth 2003 a) Apterous. Anteroventral margin of profemur with a row of spines of about equal length, the proximal ones more widely spaced than the distal ones, terminating in 2 or 3 larger spines; 4 proximal tarsomeres with pulvilli present, tarsal claws symmetrical, simple, arolia present. SG plate symmetrical or subsymmetrical, with two cylindrical styli pointing posteriorly, or slightly angled medially. Maxillary palps mostly white (or slightly buff) and largely lacking dark pigmentation. SA plate lacking pigmentation posteriorly (on edge or throughout posterior half). Female: Same as the male, but SG plate valvular and SA plate usually narrower and without whitish edge.</p><p>Remarks.</p><p>Rehn (1930) separated the genus into two groups depending on the shape of the maxillary palps. Roth assigned names to the two groups and applied Rehn’s description to them. We give the description below with since-described species added as appropriate.</p><p>I. Zamorensis-species-group: Maxillary palps robust, penultimate palpomere moderately narrowed proximad, the antepenultimate palpomere rather stout. Male abdominal tergites unspecialized. Species: albipalpus Hebard; mimadelfi sp. nov.; mimetes Rehn; zamorensis Giglio-Tos.</p><p>II. Meridionalis-species-group: Maxillary palps slenderer, penultimate palpomere narrower than the above. Species: ancistroides Rehn; flavomaculata Princis (1946); gorgonis Rehn; meridionalis Bruner; romani Rehn; neuque Sánchez Herrera, De Martino, Realpe Sanabria, Realpe Rebolledo, 2025; antoni Evangelista, Varadinova and Juna, 2019 .</p><p>Despite Rehn’s observations and Roth’s lack of disagreement, we see too little variation in the maxillary palps (Fig. 3) to uphold these species groupings. Genitalia or genetics will be needed to revise the systematics of the genus. Parsimony analysis of morphological data (Fig. 7) shows that neither group is monophyletic, and there is generally insufficient character-state information to resolve the phylogeny of Lamproblatta spp.</p><p>Male genitalia are fairly variable within the genus, although many species have not yet had their genitalia characterized. The shapes of L 3, L 2 d, and R 3 are consistent in overall shape with only minor differences. R 1 F, R 1 J, and L 4 are moderately variable, and L 2 v, R 2 are strongly variable in shape. Unfortunately, many of the sclerites, particularly R 2 and L 2 v, have quite complicated shapes, so it is difficult to articulate the differences between them. Even in photographs of the same specimen, slightly different angles yield very different perceptions of the same shape.</p><p>Lamproglandifera Roth, 2003 was differentiated from Lamproblatta by the presence of a tergal gland on the SA plate with associated modifications on preceding terga, as opposed to Lamproblatta, which lack a visible external tergal gland. In all other aspects, Lamproglandifera and Lamproblatta are nearly identical. Herrera et al. (2026) described a new Lamproblattidae species, L. neuque . Herrera et al. (2026) showed that L. neuque had a tergal gland on the SA plate and a differently shaped L 4 T + L 2 d from some Lamproblatta, but that it was perhaps more genetically similar to Lamproblatta spp. than Lamproglandifera flavoglandis . Indeed, numerous cockroach genera have variable tergal gland presentation (e. g., Periplaneta, Melanozosteria, Polyzosteria), suggesting that these characteristics are often evolutionarily labile. A wide taxonomic sampling of Lamproblatta and Lamproglandifera species in a molecular phylogeny could therefore reveal non-monophyly of Lamproblatta with respect to this other genus. Our morphological phylogeny places Lamproglandifera flavoglandis and Lamproblatta neuque with other Lamproblatta spp. (Fig. 7). While it is likely that Lamproglandifera should be synonymized with Lamproblatta, there is currently little evidence to support either claim (Fig. 7). While Roth (2003) should perhaps not have established Lamproglandifera, we refrain from reversing his decision until more robust data are collected.</p><p>More work still must be done to resolve the systematics of Lamproblatta spp. For instance, iNaturalist photos show some previously undocumented populations have similar male SA plate morphology to highly geographically disjunct populations. In particular, populations in mainland South America as far south as Brasilia are most morphologically similar to L. meridionalis, which was described only from Trinidad. Considering that Lamproblattidae are apterous, we consider this distribution unlikely, and these are probably separate species. In short, a revision of Lamproblattidae using molecular data is likely the only way its systematics can be resolved. Considering the limited number of variable characteristics and difficulty in coding them, genitalia characters may still be unable to resolve their relationships.</p></div>	https://treatment.plazi.org/id/EBC6A7F851335BE198B8B95A7D44230C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Medina-Espinoza, Emmy Fiorella;Mendieta Yáñez, Stephany Karla;Drager, Kim;Swichtenberg, Kali L.;Gile, Gillian;Kang, Yifei;Hromádka, Jiří;Vanker, Katharine;Sanchez-Herrera, Melissa	Evangelista, Dominic A., Medina-Espinoza, Emmy Fiorella, Mendieta Yáñez, Stephany Karla, Drager, Kim, Swichtenberg, Kali L., Gile, Gillian, Kang, Yifei, Hromádka, Jiří, Vanker, Katharine, Sanchez-Herrera, Melissa (2026): Xylomycophagy in a close relative of termites: Lamproblattidae (Blattodea) systematics, ecology, and diet. Arthropod Systematics & Phylogeny 84: 309-342, DOI: 10.3897/asp.84.e144506
454357C3D83A5CD390EC10E8B9C5B508.text	454357C3D83A5CD390EC10E8B9C5B508.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lamproblatta mimadelfi Evangelista, Medina Espinoza, Vanker 2026	<div><p>3.2. 4. Lamproblatta mimadelfi Evangelista, Medina Espinoza, Vanker sp. nov.</p><p>Specimens examined.</p><p>Holotype: • 1 ♂ (in UIUC Dept. of Entomology to be deposited in the Museo de Historia Natural of Universidad Nacional Mayor de San Marcos); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.107666&amp;materialsCitation.latitude=-12.559556" title="Search Plazi for locations around (long -70.107666/lat -12.559556)">Los Amigos Research Station, Trail 17 &amp; 2</a>; 12°33'34.4"S 70°06'27.6"W; 6 Jul. 2021; E. Medina Espinoza, D. Evangelista-Huaman, I. Evangelista-Huaman; Hand collected at night; AUDE-PE-14-32 . — Paratypes: • 1 ♀ adult; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.10036&amp;materialsCitation.latitude=-12.569139" title="Search Plazi for locations around (long -70.10036/lat -12.569139)">Los Amigos Research Station</a>; 12°34'08.9"S 70°06'01.3"W; 2 Jul. 2021; same collectors as holotype; collected in and around buildings at night; AUDE-PE-14-24 . • 1 ♀ adult; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.09312&amp;materialsCitation.latitude=-12.571722" title="Search Plazi for locations around (long -70.09312/lat -12.571722)">Los Amigos Research Station, Trail 8</a>; 12°34'18.2"S 70°05'35.2"W; 1 Jul. 2021; collected by coleopterists; AUDE-PE-14-17 . • 1 ♀ adult; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-70.10036&amp;materialsCitation.latitude=-12.569139" title="Search Plazi for locations around (long -70.10036/lat -12.569139)">Los Amigos Research Station</a>; 12°34'08.9"S 70°06'01.3"W; Jul. 2021; same collectors as holotype; collected in and around buildings at night; AUDE-PE-14-34 . • 1 ♂ adult; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.11444&amp;materialsCitation.latitude=-12.228611" title="Search Plazi for locations around (long -69.11444/lat -12.228611)">Finca Las Piedras Research Station</a>; 12°13'43"S 69°06'52"W; 12 Jul. 2024; E. Medina Espinoza, D. Evangelista-Huaman, J. Schwartz, J. Martin; Trails in forest, near station at night; UIRB-PE-22-89 . • 1 ♂ adult;, Finca Las Piedras Research Station; coordinates same as previous; 13 Jul. 2024; same collectors as previous; Trails in forest, near station at night; UIRB-PE-22-98 . • 1 ♂ adult; Finca Las Piedras Research Station, Anaconda and Tapir trail; coordinates same as previous; 15 Jul. 2024; same collectors as previous; UIRB-PE-25-26 . • 1 ♀ adult; same information as previous; UIRB-PE-25-25 . • 1 ♀ adult; Finca Las Piedras Research Station; coordinates same as previous; 11 Jul. 2024; same collectors as previous; UIRB-PE-25-48 . • 1 ♀ adult; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.015&amp;materialsCitation.latitude=-12.526944" title="Search Plazi for locations around (long -69.015/lat -12.526944)">Kawsay Biological Station</a>; 12°31'37"S 69°00'54"W; 18 Jul. 2024; same collectors as previous; collected in wood with Passalidae within concession, near station, at night; UIRB-PE-23-52 . • 1 ♀ adult; same locality and coordinates as previous; 22 Jul. 2024; same collectors as previous; UIRB-PE-23-56 .</p><p>Differential diagnosis.</p><p>Differs from L. mimetes, its closest known relative (Fig. 7), based primarily on the shape of the interstylar region of the SG plate, with minor differences in hind-leg tarsi. L. mimadelfi has the SG plate truncate between the styli ( L. mimetes has the region broadly rounded and expanded posteriorly), and the hind-leg tarsus of a similar length throughout ( L. mimetes hind tarsus narrows noticeably at the base). Comparisons of the male and female genitalia may reveal new characters, but genitalia in the types of L. mimetes have not been examined by us.</p><p>L. mimadelfi sp. nov. differs from all other Lamproblattidae by the lack of specialization on SA plate (as opposed to specialized SA plate in L. neuque and Lamproglandifera), solid black coloration throughout body (as opposed to bearing yellow stripes and spots laterally on the dorsum, and deep mahogany brown body instead of black in L. flavomaculata), the inflated basal tarsomere on hindleg (as opposed to not inflated or bulbously inflated), and a medial notch on the wide SA plate’s posterior edge (as opposed to no notch on the narrow edge in L. antoni).</p><p>Description holotype.</p><p>(male; AUDE-PE-14-32) Head. Frons and clypeus near uniformly black and shining, with some sparse hairs. Inter-ocular distance greater than inter-antennal distance. Ocelli yellowish, medium sized and positioned against antennal pit. Maxillary palps whitish or buffy, with distal palpomere (and distal-ventral portion of 2 nd to last palpomere) dark brown. Shape of maxillary penultimate maxillary palpomere similar to L. zamorensis (Fig. 3). — Thorax. Anterior-ventral margin of profemur with 15 large mostly equally sized spines, and two larger apical spines. Proleg without genicular spine. Pulvilli present on all four proleg tarsomeres. Other legs damaged. — Ventral abdomen. Entirely shining black, except for posterior edge of SG plate, which is slightly lighter. Styli small and positioned just medial to cerci. Posterior edge of SG plate forming a rounded-square like projection that is slightly notched medially. — Dorsal abdomen. SA plate wide, subtrapezoidal, with a slightly notched posterior end, forming a W-like shape with rounded peaks; posterior edge whitish. Terga otherwise unmodified, and no tergal gland is visible externally. — Overall, body is shining black with little undertones of brown or red. Some sparse hairs dorsally, mostly near edges of segments. Pronotum is sub-semi-circular and almost covers the head. — Genitalia as in Fig. 8 E – H. Male genital apomorphies for this species are the unique shapes of R 2, R 1 J, and other sclerites.</p><p>Description paratype.</p><p>(male; UIRB-PE-22-89) Same as the holotype with the following differences. Maxillary palps colored as in holotype but also with small spots of brown at the base of the palpomeres. Clypeus lighter in coloration (light coffee brown). SA plate with sharper peaks and deeper notch on W-shape. Hind metatarsus not narrowing at base as much as in L. mimetes, but otherwise more similar to L. mimetes than L. albipalpus .</p><p>Description paratype.</p><p>(female; AUDE-PE-14-34) Same as male except in the following. Anterior-ventral margin of profemur with 14 (right) or 16 (left) large mostly equally sized spines, and two larger apical spines. Abdomen entirely shining black with undertones of red, most apparent at the base of the segments. SG plate valvate. SA plate sub trapezoidal, with tip narrow and slightly concave. Ootheca as in Fig. 9.</p><p>Remarks.</p><p>Juveniles of this species appear the same as adults, but the coxae have large areas of stark white and / or brown.</p><p>Rehn (1930) differentiated L. mimetes from L. albipalpus based on the relative width of the SA plate (wider in L. mimetes) and interstylar region of the SG plate, and the angle of the hook on L 3 (acute in L. mimetes, rounded in L. albipalpus). Lacking molecular or further morphological data from L. mimetes, we resort to utilizing external differences established by Rehn (1930) and Hebard (1919) as described above. Our cladistic analysis places L. mimadelfi as sister to L. mimetes .</p><p>Individuals from two (Los Amigos, Puerto Maldonado) of the four localities studied are practically identical, and specimens from the other localities are not much different than one another (i. e., compare the holotype and paratype male, which are from distant localities). Thus, we expect that all four populations studied here would have low genetic differentiation and could be considered the same genetic species. This is speculation though, and our species concept for L. mimadelfi is based solely on the morphological characters mentioned above.</p><p>Range.</p><p>The species is currently only known from the Madre de Dios region of Peru but is widespread in that region.</p><p>Etymology.</p><p>The specific epithetic combines Greek derived mimetes (imposter) with adelfi (sister). Also, this specimen was first examined at Adelphi University, which also provided partial support for this research.</p></div>	https://treatment.plazi.org/id/454357C3D83A5CD390EC10E8B9C5B508	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Medina-Espinoza, Emmy Fiorella;Mendieta Yáñez, Stephany Karla;Drager, Kim;Swichtenberg, Kali L.;Gile, Gillian;Kang, Yifei;Hromádka, Jiří;Vanker, Katharine;Sanchez-Herrera, Melissa	Evangelista, Dominic A., Medina-Espinoza, Emmy Fiorella, Mendieta Yáñez, Stephany Karla, Drager, Kim, Swichtenberg, Kali L., Gile, Gillian, Kang, Yifei, Hromádka, Jiří, Vanker, Katharine, Sanchez-Herrera, Melissa (2026): Xylomycophagy in a close relative of termites: Lamproblattidae (Blattodea) systematics, ecology, and diet. Arthropod Systematics & Phylogeny 84: 309-342, DOI: 10.3897/asp.84.e144506
7F95B891D96250BDB3CDA51838A06354.text	7F95B891D96250BDB3CDA51838A06354.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lamproblattidae McKittrick 1964	<div><p>3.2. 1. Lamproblattidae McKittrick, 1964</p><p>Taxonomic scope.</p><p>Lamproblatta Hebard, 1919, Lamproglandifera Roth, 2003</p><p>History.</p><p>Hebard (1919) proposed that Lamproblatta was most closely related to Eurycotis, but differentiated it based on the lack of forewings. Indeed, Lamproblatta is more superficially similar to Eurycotis than any other local genus known at this time. McKittrick (1964) identified the unique morphology of Lamproblatta ’ s internal structures and established it as a new subfamily, which Klass and Meier (2006) elevated to family status. Since then, Sanger data (Legendre et al. 2015; Malem et al. 2023), mitochondrial genomes (Bourguignon et al. 2018; Deng et al. 2023), transcriptomic data (Evangelista et al. 2019 b; Liu et al. 2023; Evangelista et al. 2024), and phenotypic data (Fig. 7; also, see supplementary material in Evangelista et al. 2019 b) has established a relationship with Xylophagodea (but see below for alternative hypotheses). Upon the description of Lamproglandifera Roth, 2003, Lamproblattidae was no longer monogeneric. Lamproglandifera is externally identical to Lamproblatta except for the presence of a visible tergal gland in the former. A few years later, Fritzsche &amp; Zompro (2008) placed Eurycanthablatta in Lamproblattidae but gave little concrete evidence for it. We do not consider Eurycanthablatta as part of Lamproblattidae (see section below).</p><p>Diagnosis.</p><p>The following is revised from McKittrick (1964), who based her comparisons on Lamproblatta albipalpus and other non- Lamproblattidae (mostly Cryptocercus punctulatus). To her comparisons, we add data from our morphological analysis. (i) Female genitalia. Compared to Cryptocercus, adult female Lamproblattidae genitalia are more heavily pigmented, valvifers more elongate, paratergites much shorter, laterosternites IX are “ enormously expanded ”, second valvifer ring larger and heavier, and paratergo-mediosternal muscle IX is longer. Compared to Cryptocercus, the intersternal musculature of Lamproblattidae have the intersternal muscle VII – VIII divided into three pairs, and segmental muscles not continuous. (ii) Proventriculus. Greatly expanded armarium, heavily sculpted primary teeth (large and triangular in transverse section, rather than bladelike), projections of denticles more variable than in Cryptocercidae (i. e. with a stronger bilateral component), intercalary sclerites are noticeably shorter than the interdentaries, primary pulvilli are shorter than in Cryptocercus . Klass and Meier (2006) noted that Lamproblattidae is unique in having the tips of all six teeth inclined counterclockwise when viewed anteriorly. (iii) Male genitalia. Sclerite regions R 1 v and R 1 d simplified and forming a single sclerite R 1 J and with a medial spine-like projection, designated “ sra ” by Klass (1997) (in Cryptocercus the spine is missing, and R 1 is wider). L 1 large, flat, and boot-shaped (as opposed to a hollow lobe, or in the shape of hollow spines). L 2 c (base of McKittrick’s L 2 d) moderately narrow, nodule-like (in Cryptocercus, L 2 is broader and more rounded), process pda (sclerotized by region L 4 l) a highly elongate curved spine (pda differently shaped and not a spine in Cryptocercus, but spine-like in some Blattidae). Hook hla of left phallomere (with sclerite L 3) highly elongated, shaped like a curved cane (stout and robust in Cryptocercus) (Klass 1997). (iv) Other characteristics (Fig. 2). Maxillary palps whitish (but can have some weak pigmentation) as opposed to strongly pigmented, posterior end of SA plate whitish and lacking pigmentation to various degrees, SA plate edge slightly notched medially (but deeply notched in a few species).</p><p>Remarks.</p><p>Within Lamproblattidae, phylogenetic relationships are largely unknown, and morphological differences used historically (e. g., morphology of the penultimate maxillary palpomere; Rehn 1930) may not be informative when a robust species concept is applied. The shape of the penultimate maxillary palpomere was homoplasious in most cladistic analyses here (e. g., Fig. 7). Regardless, we find this character to be extremely subtle and perhaps subjective, even under quantitative analysis (e. g., if quantifying it as the basal angle of the palpomere, it is not always clear where exactly to measure the angle from). Eurycanthablatta, at least, is clearly different from Lamproblattidae species, and perhaps molecular systematics would also reveal them to be a deeper lineage. Lamproglandifera and Lamproblatta are less distinct from one another. Visible tergal gland modifications are highly volatile among Blattodea, often being variable between otherwise similar species (Roth 1969). Lamproblatta neuque Sánchez Herrera, De Martino, Realpe Sanabria, Realpe Rebolledo, 2025 also has a visible tergal gland modification (Herrera et al. 2026).</p><p>The type of tergal gland modification (a patch of hairs on a modified SA plate) is, in itself, interesting because it is also present in Anaplectidae (Deng et al. 2026) . Recent phylogenetic studies have shown some molecular support for Anaplectidae as sister to Lamproblattidae (Bourguignon et al. 2018; Li 2022; Deng et al. 2023; Liu et al. 2023; Evangelista et al. 2024) and we recover it here on the basis of other characters (character 14 – tergal gland modification – was usually assigned as a homoplasy). However, statistical tests in molecular analyses have not been able to rule out other possible placements (Evangelista et al. 2024).</p></div>	https://treatment.plazi.org/id/7F95B891D96250BDB3CDA51838A06354	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Evangelista, Dominic A.;Medina-Espinoza, Emmy Fiorella;Mendieta Yáñez, Stephany Karla;Drager, Kim;Swichtenberg, Kali L.;Gile, Gillian;Kang, Yifei;Hromádka, Jiří;Vanker, Katharine;Sanchez-Herrera, Melissa	Evangelista, Dominic A., Medina-Espinoza, Emmy Fiorella, Mendieta Yáñez, Stephany Karla, Drager, Kim, Swichtenberg, Kali L., Gile, Gillian, Kang, Yifei, Hromádka, Jiří, Vanker, Katharine, Sanchez-Herrera, Melissa (2026): Xylomycophagy in a close relative of termites: Lamproblattidae (Blattodea) systematics, ecology, and diet. Arthropod Systematics & Phylogeny 84: 309-342, DOI: 10.3897/asp.84.e144506
