identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
8F02C7C803CC5B25B607F1847FA459A8.text	8F02C7C803CC5B25B607F1847FA459A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptoconops communis Gonzalez & Tanase 2026	<div><p>Leptoconops communis González &amp; Tanase sp. nov.</p><p>Fig. 7, Suppl. material 1: figs S 19 – S 25</p><p>Material examined.</p><p>Holotype. SPAIN • ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.11586&amp;materialsCitation.latitude=37.800274" title="Search Plazi for locations around (long -6.11586/lat 37.800274)">Almadén de la Plata</a>, Sevilla; 37.800273, -6.11586; alt. 398 m. 16 May 2024; M. A. González leg.; BGS trap; EBD I-016381 . Paratypes: SPAIN • 1 ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.751417&amp;materialsCitation.latitude=37.688618" title="Search Plazi for locations around (long -6.751417/lat 37.688618)">Zalamea la Real</a>, Huelva; 37.688618, -6.751417; alt. 289 m; 25 Apr. 2024; M. A. González leg.; BGS trap; EBD I-016382 - 3 ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.072888&amp;materialsCitation.latitude=37.564304" title="Search Plazi for locations around (long -6.072888/lat 37.564304)">Guillena</a>, Sevilla; 37.564304, -6.072888; alt. 85 m. 05 May 2023; M. A. González leg.; BGS trap; EBD I-016383; EBD I-016384 - 1 ♀; labelled as for the holotype except for NHMUK 014316792 - 1 ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.751417&amp;materialsCitation.latitude=37.688618" title="Search Plazi for locations around (long -6.751417/lat 37.688618)">Zalamea la Real</a>, Huelva; 37.688618, -6.751417; alt. 289 m; 25 Apr. 2024; M. A. González leg.; BGS trap; MNCN_Ent 283587 .</p><p>Diagnosis.</p><p>Under close magnification (20–40 ×), the third palp segment in L. communis sp. nov. is broadly oval and has abundant sensilla (&gt; 70), with the sensory pit occupying three-quarters of the segment. Spermathecae oval and very large (length&gt; 65 µm) with no apical pubescence on terminal end (Fig. 7 H). Armature of the anal cone with a small or inconspicuous protuberance in the middle (Fig. 7 I).</p><p>Description.</p><p>Females medium-sized (1.69 ± 0.08 mm). Thorax is pruinose and slightly shiny, with dark-green metallic tones; legs bicolored (foreleg uniformly dark brown, mid and hindlegs with mixed coloration) (Fig. 7 A, Suppl. material 1: fig. S 19 A).</p><p>Head. Subspherical, polished dark with metallic-green reflections on the clypeus, frons, and vertex (Suppl. material 1: fig. S 19 C). Head length 368.81 ± 20.38 µm; width 416.42 ± 26.97 µm (Suppl. material 1: fig. S 19 B). Head width 3.7 × (3.5–3.8) the interocular space (Suppl. material 1: fig. S 19 B). A single interocular seta on each side was observed in one specimen. Vertex slightly spiculated at 400 × magnification, with twelve supraorbital setae (Suppl. material 1: fig. S 19 D) arranged in a single a row and three setae situated in the upper interocular space. Eyes bare, separated by the diameter of 11 ommatidia (111.40 ± 5.29 µm).</p><p>Antenna. Pedicel has three setae; flagellomere I bears four setae arranged in a row (Suppl. material 1: fig. S 20 C). Lengths of flagellomeres I – XII are provided in Suppl. material 1: table S 2. Total antennal length 450.53 ± 28.67 µm; AR = 0.66. Proximal flagellomeres II – IV longitudinally flattened, V – VI subspherical, VII slightly spherical, and distal VIII – XI entirely spherical (Suppl. material 1: fig. S 20 A, B). Terminal flagellomere elongate, of similar length to the combined length of the preceding 2–2.5 segments, densely setose, with numerous minute hyaline ss and 5–7 sensilla chaetica (1 apical, 3 or 4 subapical and 1 or 2 basal) (Suppl. material 1: fig. S 20 D). Each flagellomere has 12–16 sensilla trichodea arranged in two or three semi-verticils on both faces (Fig. 7 B, Suppl. material 1: fig. S 20 E). Distinctive, completely straight, conspicuous sensilla chaetica are present on one face: four on flagellomere I, three or four on flagellomeres II – VII and two or three on flagellomeres VIII – XI (Fig. 7 B, Suppl. material 1: fig. S 20 E). Numerous minute hyaline ss are scattered over each flagellomere but are difficult to observe. One pair of hyaline ss on I – XI, thickened basally, thin beyond the middle, abruptly curved near the base (bull-horn shaped), and positioned subapically on one face, with very close insertions (Fig. 7 C). In proximal flagellomeres (II – V), the hyaline ss tips usually exceed by one-third to one-half the length of the following flagellomeres; in distal flagellomeres (VI – XI), the tips exceed by ¼ the length of the subsequent flagellomere.</p><p>Mouthparts. Blackish. Clypeus as described for the preceding species (Suppl. material 1: fig. S 19 D). Proboscis 224.47 ± 23.83 µm, shorter than head. Palpus not exceeding the proboscis tip; total palpus length 216.12 ± 8.44 µm; PR = 1.29 (1.17–1.29) (Suppl. material 1: fig. S 21 A). The first two palpal segments 36.56 ± 7.11 µm each bear three distinctive conspicuously long setae (usually exceeding 1 / 3 the length of the third palp segment). Third palp segment large (Fig. 7 D), markedly incrassate, with 11–13 setae (3 or 4 on outer margin, four on sensory area usually arranged in 1 or 2 rows, 3 between the margins on each side, and 3 on inner margin) (Suppl. material 1: fig. S 21 B, C). Third palpal segment moderately longer than wide (91.11 ± 2.5 µm and 71.08 ± 8.98 µm). Sensorium with 70–80 sensilla arranged in 8–11 irregular rows, occupying ¾ (Suppl. material 1: fig. S 21 C). Sensillar area: 1,918 ± 76.49 µm 2. Fourth segment cylindrical (length: 88.44 ± 3.28 µm and width: 22.57 ± 2.89 µm), with one or two median (usually 1), two subapical, and four apical setae (Suppl. material 1: fig. S 21 D). Lacinia with 21–22 apically curved teeth (Suppl. material 1: fig. S 21 E). Mandible armed with 21–23 teeth (Suppl. material 1: fig. S 21 D).</p><p>Thorax. Mesonotum pruinose, slightly shiny with dark-green metallic tones (depending on light), length 684 ± 43.49 µm and width 426 ± 33.04 µm (Suppl. material 1: fig. S 22 A – D). Three longitudinal rows of setae present (1 acrostical and 2 dorsocentral), with additional setae scattered antero- and postero-laterally; posterior dorsocentral setae longer (Suppl. material 1: fig. S 22 C). Three distinctive supraalar setae on each side. Humeral callus prominent, angular, with four bristles (Suppl. material 1: fig. S 22 C). Four small rounded black humeral pits aligned anteriorly (Suppl. material 1: fig. S 22 B); four additional distinctive black pits in a paired quadrate pattern in the prescutellar area (Suppl. material 1: fig. S 22 C). Anepisternum shiny, metallic green, without setae (Suppl. material 1: fig. S 22 B), with amber tones in the anepisternal cleft (Suppl. material 1: fig. S 22 E). Katepisternum blackish with metallic-green tones, shiny, with 10–12 scattered setae on each side (Suppl. material 1: fig. S 22 B). Scutellum pruinose with two setae on each side (1 apical and 1 basal), the basal one the shorter of the two (Suppl. material 1: fig. S 22 D). Post-scutellum slightly arched, pruinose, and bare. Halters entirely yellowish white (Suppl. material 1: fig. S 22 A).</p><p>Legs. Bicolored. Measurements in Suppl. material 1: table S 3. Foreleg (1,592 ± 99.43 µm) and midleg (1,773 ± 42.92 µm) shorter than hindleg (1,866.69 ± 50.45 µm). Femur and tibia pruinose, uniformly dark brown. Foreleg: first tarsomere evenly dark brown with light / dark coloration proximally, the remaining segments entirely dark brown (Fig. 7 E). Midleg: first tarsomere pale yellowish-brown becoming darker distally; remaining segments entirely dark brown (Fig. 7 F). Hindleg: first tarsomere entirely pale yellowish-brown with slightly darker apex; second tarsomere mixed (proximally yellowish brown, dark brown beyond the middle), remainder dark brown (Fig. 7 G). Coxa with five or six bristles arranged in a row on each side (Suppl. material 1: fig. S 23 D). Trochanter with 6–9 scattered bristles (Suppl. material 1: fig. 23 E). Proximal trochanter with twelve or thirteen chordotonal organs in two groups (5–7 + 6–7) (Suppl. material 1: fig. S 23 B); femur with twelve chordotonal organs in two rows (8 + 4) (Suppl. material 1: fig. S 23 C). Trochanter, femur, and tibia pilose; femur with three rows of bristles; tibia with three irregular proximal rows and four irregular distal rows of setae arranged longitudinally. Tibial comb with four spines (Fig. 7 J, Suppl. material 1: fig. S 23 A), the two median ones longer. Apical spines present on first (2 spines), second (1 spine), and third (1 spine) foreleg tarsomeres; on the first (2 spines), second (1 or 2 spines), and third (1 spine) midleg tarsomeres; and on first (2 or 3 spines), second, and third (1 or 2 spines) hindleg tarsomeres. Basitarsus densely setulose; the remaining tarsomere moderately setulose. Claws symmetrical, straight to slightly curved apically, each with a basal tooth longer than wide, and a small accessory bristle (Suppl. material 1: fig. S 24).</p><p>Wing. Length: 1.16 ± 0.03 mm; width: 0.43 ± 0.02 mm, and CR: 0.39 (0.38–0.41). Veins distinct, radial cells strongly sclerotized, with marked yellowish-brown tones (Suppl. material 1: fig. S 25 A, B).</p><p>Abdomen. Tergites and sternites polished, dorsally dark brown (Suppl. material 1: fig. S 25 C), ventrally slightly paler, with metallic-green reflections (depending on light) (Suppl. material 1: fig. S 25 D). Setae scattered dorsally and ventrally, abundant on segments 1–5 (&gt; 30), moderately abundant on segments 6 and 7 (16–30) but fewer on segments 8–10 (&lt;10). Sternites with larger setae than tergites. Tergites with two longitudinal rows of pits (one pair per tergite) (Suppl. material 1: fig. S 25 C). Sternites 6–8 with three central pits arranged in a triangular configuration (Suppl. material 1: fig. S 25 D). Two functional oval and elongated spermathecae (65.84 ± 3.93 × 45.31 ± 4.64 µm and 60.73 ± 4.62 × 45.95 ± 5.19 µm), highly sclerotized, lacking apical pubescence on terminal end, with a long, conspicuous, tronco-conical sclerotized neck (10.97 ± 1.36 µm) (Fig. 7 H, Suppl. material 1: fig. S 25 I). A third rudimentary spermatheca present.</p><p>Genitalia. Sternite VIII with the lateral arms of the ventral plate strongly sclerotized, projecting posteriorly (Suppl. material 1: fig. S 25 H). Genital atrium large and semicircular, with four curved bristles of similar size orientated posteriorly and a variable number of smaller marginal bristles (Suppl. material 1: fig. S 25 G). Behind the atrium, sternite IX has a pigmented, highly setulose structure with medial excavation, with three lateral bristles. Genital armature of the anal cone V-shape in ventral view, heavily sclerotized, with a small median protuberance (Suppl. material 1: fig. S 25 H). Genital lamella elongate (Fig. 7 I, Suppl. material 1: fig. S 25 E), light grey (Suppl. material 1: fig. S 25 F), (217.84 ± 10.87 µm × 66.31 ± 8.6 µm, length × width; index 3.31, 2.91–3.67), each with 40–50 long, thick, scattered setae, including one long apical seta on each lamella (Suppl. material 1: fig. S 25 E).</p><p>Male. Unknown.</p><p>Etymology.</p><p>The species epithet communis refers to its high abundance in the studied region.</p><p>Distribution.</p><p>This species is known from numerous locations in the three sampled provinces (Fig. 8).</p><p>Habitat.</p><p>Specimens were collected in large number (&gt; 1,000) from Mediterranean shrubs near arable fields (various crop types) and pastures (Fig. 9 A – E).</p></div>	https://treatment.plazi.org/id/8F02C7C803CC5B25B607F1847FA459A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Paun-Tanase, Dumitru Ionut;González, Mikel Alexander;Magallanes, Sergio;Naro, Giovanni;Epis, Sara;Figuerola, Jordi	Paun-Tanase, Dumitru Ionut, González, Mikel Alexander, Magallanes, Sergio, Naro, Giovanni, Epis, Sara, Figuerola, Jordi (2026): Hidden diversity in Europe: new species and a revised taxonomy of the subgenus Leptoconops (Leptoconops) (Diptera, Ceratopogonidae). ZooKeys 1279: 201-239, DOI: 10.3897/zookeys.1279.186179
D3BFF2BAEB50581F964172A91D4E7136.text	D3BFF2BAEB50581F964172A91D4E7136.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptoconops nigrithorax Gonzalez & Tanase 2026	<div><p>Leptoconops nigrithorax González &amp; Tanase sp. nov.</p><p>Fig. 1, Suppl. material 1: figs S 1 – S 6</p><p>Material examined.</p><p>Holotype: SPAIN • ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.098175&amp;materialsCitation.latitude=36.590805" title="Search Plazi for locations around (long -6.098175/lat 36.590805)">Las Pachecas</a>, Cádiz; 36.590805, -6.098175, alt. 61 m; 29 May 2023; M. A. González leg.; BGS trap; EBD I-016375 . Paratypes: SPAIN • 1 ♀; labelled as for the holotype, except for EBD I-016376 - 3 ♀; labelled as for the holotype, except for NHMUK 014316789 - NHMUK 014316791 - 2 ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.042228&amp;materialsCitation.latitude=37.14261" title="Search Plazi for locations around (long -6.042228/lat 37.14261)">Los Palacios</a>, Sevilla; 37.14261, -6.042228, alt. 1 m; 21 Jun 2023; M. A. González leg.; BGS trap; MNCN_Ent 283581 .</p><p>Diagnosis.</p><p>Leptoconops nigrithorax sp. nov. can be distinguished from other European Leptoconops species by the combination of the following characters: (i) shiny black thorax, visible both dorsally and laterally under a stereomicroscope (Fig. 1 A, B), (ii) uniformly dark brown legs, without pale coloration on tarsomeres (Fig. 1 F – H), and (iii) a higher number of lacinial teeth. In addition, the type and distribution of antennal sensilla also provide useful diagnostic features.</p><p>Description.</p><p>Females large (2.06 ± 0.14 mm). Specimens dark, with a shiny black thorax (Fig. 1 A, B, Suppl. material 1: fig. S 1 A, B), brownish abdomen, and uniformly dark-brown legs (Fig. 1 F – H).</p><p>Head. Subspherical, with dull metallic green tones on clypeus, frons, and vertex (Suppl. material 1: fig. S 1 C, D). Head length: 343.52 ± 7.59 µm; width: 377.61 ± 7.85 µm (Suppl. material 1: fig. S 1 D). Head width is 4.2-4.5 × the interocular distance. Two (occasionally 1) lateral interocular setae present, one each side (Suppl. material 1: fig. S 1 F). Vertex moderately spiculated (400 × magnification) and with 10–12 supraorbital setae arranged in a row (Suppl. material 1: fig. S 1 E). Eyes bare, moderately separated by a diameter of 7.28 ommatidia (85.89 ± 2.67 µm) (Suppl. material 1: fig. S 1 F).</p><p>Antenna. Pedicel with four setae; flagellomere I has five or six setae. Lengths of flagellomeres I – XII are provided in Suppl. material 1: table S 2. Total antennal length = 388.75 ± 29.39 µm; AR = 0.7. Proximal flagellomeres II – IV are longitudinally flattened, while V – VI vary from flattened to subspherical; distal flagellomeres VII – XI are spherical (Suppl. material 1: fig. S 2 A, B). Terminal flagellomere elongate and similar in length to the preceding three flagellomeres combined: densely setose with three or four sensilla chaetica on the anterior mid-section and one or two minute hyaline ss (Suppl. material 1: fig. S 2 C). Flagellomeres armed with abundant long sensilla trichodea that extend beyond the base of the next segment (sum of both faces): 6–10 on flagellomeres I – IV and 18–22 on flagellomeres V – XI. The sensilla trichodea are arranged in two or three semi-verticils on the external face and in a basal whorl on the internal face (Fig. 1 D). Straight and thick hyaline ss (one pair per flagellomere, II – XI) are diametrically opposite (facing each other), tips reach the base of the next flagellomere (Fig. 1 E). One minute hyaline ss located on one side of flagellomeres II – XI (sometimes variable and / or difficult to see). No sensilla chaetica on flagellomeres except on I (occasionally 1 on II).</p><p>Mouthparts. Dark brown (Suppl. material 1: fig. S 3 A). Clypeus with four frontoclypeal setae (similar length and thickness) arranged trapezoidally: the anterior pair closer but the posterior pair widely separated (Suppl. material 1: fig. S 1 G). Proboscis (216.98 ± 8.71 µm) shorter than the head. Palps slightly exceed the proboscis tip. Palpus length: 222.57 ± 7.31 µm. PR = 2.06 (1.80–2.32). The first two palpal segments (length: 44.17 ± 7.74 µm) bear three long setae. The third palpal segment (Fig. 1 C) is longer than wide (94.95 ± 4.78 µm and 46.41 ± 6.19 µm), slightly incrassate and with 8–10 setae (3 on outer margin, 2 or 3 in sensory area, and 3 or 4 on inner margin). Sensorium with 35–40 sensory pits arranged in 4–6 irregular rows, covering roughly three-quarters of the surface area. The sensillar area measures 1,500.74 ± 93.06 µm 2. The fourth segment (83.75 ± 1.81 µm) cylindrical, with one or two median setae (usually 1) and 3–5 (usually 4) apical / subapical setae. Lacinia with 22–30 evenly arranged teeth (Suppl. material 1: fig. S 3 B). Mandible is armed with 19–24 teeth (Suppl. material 1: fig. S 3 C).</p><p>Thorax. Mesonotum polished, shiny black, length 660.10 ± 34.10 µm, width 440.24 ± 10.00 µm (Fig. 1 B, Suppl. material 1: fig. S 4 A, B); setose, with three longitudinal straight rows of dark, distinct setae (1 acrostical and 2 dorsocentral, 14–16 setae per row) (Suppl. material 1: fig. S 4 C, D). Scapular setae are present and a pair of longer prescutellar setae also present. Anterolateral scutal margin with 15–22 scattered setae per side (Suppl. material 1: fig. S 4 D). Lateral thoracic sclerites polished and shiny, often with reddish mahogany tones, especially in clefts (Suppl. material 1: fig. S 4 B). Humeral pits absent or inconspicuous. Katepisternum shiny black, with 8–10 scattered setae on each side. Scutellum pruinose with two pairs of long medial setae (subapical and lateral) of similar length (Suppl. material 1: fig. S 4 E). Post-scutellum strongly arched and bare, with ornamentation and distinct metallic green tones (Suppl. material 1: fig. S 4 F). Halters white apically, with dark basally (Suppl. material 1: fig. S 4 F).</p><p>Legs. Uniformly dark brown, not shiny (Fig. 1 F – H). Measurements in Suppl. material 1: table S 3. Hindleg (1,896.39 ± 105.81 µm) longer than both foreleg (1,529 ± 50.15 µm) and midleg (1,689.31 ± 77.10 µm). Coxa with six bristles; distal trochanters, basal femur, and tibia possess chordotonal organs (Suppl. material 1: fig. S 5 D). Trochanter surface with 5–7 long, distinctive bristles plus two short straight bristles and 14–16 chordotonal organs (arranged in two groups, 5–7 + 8–10), femur with 12 bristles arranged in two lines (4 + 8), and tibia with three or four (Suppl. material 1: fig. S 5 D). Tarsomeres of legs covered with uniformly fine pilosity; femur and tibia with a variable number of long distinct bristles arranged in rows. Typically, at least two lines of bristles arranged in one row, with the other two or three lines mixed. Both anterior and posterior apical regions of the femur each with two or three additional long bristles. Tibial comb with four spines (outer spines shorter than inner ones) (Suppl. material 1: fig. S 5 A). Apical spines are present on the first (1 or 2, except hind tarsomere, which is variable, having 2–6 strong spines) and second (usually 2) tarsomeres but absent on the third and fourth (variable on the third of the hindlegs). The basitarsus is highly setose in all legs; claws symmetrical, each with a narrow, pointed, moderate-sized basal tooth (Suppl. material 1: fig. S 5 B, C); empodium with a branched bristle.</p><p>Wing. Length: 1.08 ± 0.01 mm; width: 0.50 ± 0.03 mm; CR: 0.39 (0.38–0.41). Veins are distinct, radial cells highly sclerotized, with yellow tones (Suppl. material 1: fig. S 5 E).</p><p>Abdomen. Tergites (Suppl. material 1: fig. S 6 A) and sternites (Suppl. material 1: fig. S 6 B) brown to dark brown, pleural membrane with a leathery texture. Ventral setae are abundant on segments 1–5 (&gt; 30), moderate on segments 6 and 7 (15–22), and scarce on segments 8–10 (&lt;15). Two small functional spermathecae of equal size (47.09 ± 3.82 µm × 28.76 ± 3.12 µm and 45.79 ± 4.78 µm × 27.65 ± 2.05 µm), oval, and highly sclerotized, with an inconspicuous uncolored area apically and a short, well-formed neck (5.65 ± 2.05 µm) (Fig. 1 I, J, Suppl. material 1: fig. S 6 D). Two small accessory spermathecae present, one bigger than the other (depending on how placed during preparation; the smallest may be difficult to observe) (Fig. 1 I, J).</p><p>Genitalia. On sternite VIII the extremities of the lateral arms of the ventral plate are sclerotized, pointed, and membranous at the base (with stretch marks). These arms are orientated horizontally outwards or slightly curved (Suppl. material 1: fig. S 6 C). Ventral extremities of segment IV are shaped in a sclerotized structure with three or four bristles. Genital atrium semicircular (slightly wider than long), with four large, thick, curved bristles (arising from a pediculate protuberance) on each side posteriorly and numerous weak bristles evenly arranged at the base (Suppl. material 1: fig. S 6 E). Behind the atrium lies a pigmented structure with an excavation in the middle section, sometimes featuring a tiny projection. Genital armature of anal cone large, V-shaped ventrally, with a trapezoidal and well-sclerotized upper part (Suppl. material 1: fig. S 6 C, E). Genital lamella dark, elongate (255.64 ± 7.32 µm × 69.37 ± 9.38 µm, length × width; index: 3.7, 3.10–4.49) bearing 55–70 setae (Fig. 1 I, Suppl. material 1: fig. S 6 C).</p><p>Male. Unknown.</p><p>Etymology.</p><p>The specific epithet nigrithorax refers to the shiny black thorax that characterizes this species.</p><p>Distribution.</p><p>This species was detected at numerous different sites across the three sampled provinces (Fig. 2).</p><p>Habitat.</p><p>Specimens were collected in large numbers (&lt;1,000) from Mediterranean shrubs near arable land (diverse crops), pastures, and small water bodies (e. g. creeks) (Fig. 9 A – D).</p><p>Notes.</p><p>Ethanol alters the coloration of some structures in fresh specimens. The shiny appearance is less evident in ethanol preserved specimens. Lighting conditions also influence the perception of colors and tones.</p></div>	https://treatment.plazi.org/id/D3BFF2BAEB50581F964172A91D4E7136	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Paun-Tanase, Dumitru Ionut;González, Mikel Alexander;Magallanes, Sergio;Naro, Giovanni;Epis, Sara;Figuerola, Jordi	Paun-Tanase, Dumitru Ionut, González, Mikel Alexander, Magallanes, Sergio, Naro, Giovanni, Epis, Sara, Figuerola, Jordi (2026): Hidden diversity in Europe: new species and a revised taxonomy of the subgenus Leptoconops (Leptoconops) (Diptera, Ceratopogonidae). ZooKeys 1279: 201-239, DOI: 10.3897/zookeys.1279.186179
F334D35D3AAC5BFA86F207CFD0667B57.text	F334D35D3AAC5BFA86F207CFD0667B57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptoconops pseudoirritans Gonzalez & Tanase 2026	<div><p>Leptoconops pseudoirritans González &amp; Tanase sp. nov.</p><p>Fig. 5, Suppl. material 1: figs S 13 – S 18</p><p>Material examined.</p><p>Holotype: SPAIN • ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.778454&amp;materialsCitation.latitude=37.28834" title="Search Plazi for locations around (long -5.778454/lat 37.28834)">Alcalá de Guadaíra</a>, Sevilla; 37.288340, -5.778454; alt. 45 m. 6 Jun. 2024; M. A. González leg.; BGS trap; EBD I-016379 . Paratypes: SPAIN • 1 ♀; labelled as for the holotype, except for EBD I-016380; 2 ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.630657&amp;materialsCitation.latitude=37.445282" title="Search Plazi for locations around (long -5.630657/lat 37.445282)">Carmona</a>, Sevilla; 37.445281, -5.630657; alt. 87 m; 09 Aug 2023; M. A. González leg.; BGS trap; NHMUK 014316795; NHMUK 014316796 - 1 ♀; Arahal, Sevilla; 37.26508016923277, -5.636044311686363; alt. 65 m; 06 Jun 2024; M. A. González leg.; BGS trap; NHMUK 014316794 - 1 ♀ labelled as previous paratype, except for NHMUK 014316793 - 1 ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.500349&amp;materialsCitation.latitude=37.370434" title="Search Plazi for locations around (long -5.500349/lat 37.370434)">El Palomar</a>, Sevilla; 37.370435, -5.500349; alt 113 m; 10 May 2023; M. A. González leg.; BGS trap; MNCN_Ent 283585 - 1 ♀; Arahal, Sevilla; 37.26508016923277, -5.636044311686363; alt. 65 m; 06 Jun 2024; M. A. González leg.; BGS trap; MNCN_Ent 283586 .</p><p>Diagnosis.</p><p>Claws with bristle-like teeth; proboscis length shorter (&lt;280 µm). Sensory pits with &lt;40 sensilla (Fig. 5 D). Spermathecae nearly as long as wide (Fig. 5 I). Flagellomere XII usually with four sensilla chaetica.</p><p>Description.</p><p>Females small (1.52 ± 0.06 mm). Thorax dorsally pruinose with metallic green reflections; abdomen pale cream to brown; legs bicolored (Fig. 5 A, Suppl. material 1: fig. S 13 A).</p><p>Head. Subspherical, dull pruinose to dark brown, with metallic-green tones (Suppl. material 1: fig. S 13 B – D). Head length 347.93 ± 11.51 µm, slightly exceeding width 337.44 ± 14.61 µm. Head width is 3.7 × (3.3–4.0) the interocular distance; two lateral setae are present, one on each side. Vertex with around ten supraorbital setae arranged in a single row; surface slightly spiculate at 400 × magnification (Suppl. material 1: fig. S 13 E). Eyes bare, separated by the diameter of 8.2 ommatidia (89.41 ± 6.54 µm) (Suppl. material 1: fig. S 13 D, E).</p><p>Antenna. Pedicel with 2–5 long bristles; flagellomere I with five (4 + 1) bristles. Lengths of flagellomeres I – XII are provided in Suppl. material 1: table S 2. Total antennal length = 340.20 ± 17.46 µm; AR = 0.64. Flagellomeres I – XI slightly flattened, subspherical (Suppl. material 1: fig. S 14 A). Terminal flagellomere (Suppl. material 1: fig. S 14 B) thick and club-shaped, markedly wider than the preceding segments, densely setose, similar in length to the previous two segments combined and with four sensilla chaetica on the distal third (3 beyond the central part, one subapical). Each of flagellomeres I – XI has a pair of short (similar in length to flagellomere) hyaline ss arranged in pairs (diametrically opposed) (Fig. 5 B, Suppl. material 1: fig. S 14 C). On flagellomeres II – XI, 10–14 sensilla trichodea are arranged in semi-verticils on one face (Fig. 5 B). Medium-length sensilla chaetica are present on one face of proximal flagellomeres II – VIII (2–4) and distal flagellomeres IX – XI (0–2) (Fig. 5 C). Flagellomeres have a variable number of minute hyaline ss scattered across their surface.</p><p>Mouthparts. Dark brown. Frontoclypeal setae as described in previous species. Proboscis length (265.04 ± 22.99 µm) shorter than head length (Suppl. material 1: fig. S 13 B, C). Palps reach the tip of the proboscis. Palpus length 314.52 ± 16.81 µm; PR = 2.28 (2.14–2.39). The first two palpal segments measure 68.29 ± 7.41 µm and each bear three long setae. The third segment (Fig. 5 D, Suppl. material 1: fig. S 15 A – C) is brown, slightly incrassate, with 3–7 setae (1–3 on the outer margin, 1–3 on the sensory area, and 1 or 2 on the inner margin), and is twice as long as wide (97.24 ± 4.76 µm and 42.71 ± 3.88 µm). Sensorial area with 20–30 pits arranged in four or five irregular rows, occupying ½ of the segment surface (Fig. 5 D, Suppl. material 1: fig. S 15 C) (area 987.79 ± 156.29 µm 2). The fourth palpal segment very elongated and narrow (length: 106.27 ± 10.20 µm and width: 17.14 ± 1.17 µm), with two or three median setae (usually 2), two or three subapical setae (usually 2), and 2–4 apical setae. Lacinia with 13–15 teeth (Suppl. material 1: fig. S 15 D): Mandible armed with 12–18 teeth (Suppl. material 1: fig. S 15 E).</p><p>Thorax. Mesonotum pruinose with green or greyish reflections that vary with light; length 530.20 ± 20.68 µm and width 340.10 ± 10.10 µm (Suppl. material 1: fig. S 16 A). It has indistinct brown setae arranged in three longitudinal rows (one acrostical and two dorsocentral), plus scattered antero-posterolaterally setae (~ 30). Present on each side of the mesonotum are two or three conspicuous supraalar setae. The humeral callus is slightly prominent and has angular margins. The humeral pits (3 or 4 circular black spots) are aligned anteriorly, with 2–4 small black pits in the prescutellar area (Suppl. material 1: fig. S 16 A – C). The anepisternum and katepisternum are similar in color to the mesonotum and have abundant inconspicuous scattered setae (~ 20). Anepisternum clefts have reddish or yellowish tones. The pruinose scutellum has two subapical and two lateral setae. Post-scutellum pruinose and bare. Halters whitish apically and dark basally (Suppl. material 1: fig. S 16 D).</p><p>Legs. Bicolored, not pruinose, with alternating dark and pale patterns. Measurements in Suppl. material 1: table S 3. Foreleg (1,368.39 ± 81.67 µm) and midleg (1,383 ± 68.37 µm) shorter than hindleg (1,691.755 ± 136.31 µm). Femur and tibia uniformly dark brown. Foreleg tarsomeres: first variable, usually with a darker basal portion (three-quarters dark, one-quarter pale); other tarsomeres uniformly dark (Fig. 5 E). Mid and hindleg tarsomeres: first and second pale except for dark apical tips; other segments uniformly dark (Fig. 5 F, G). Coxae and trochanters with 6–9 bristles each (Suppl. material 1: fig. S 17 B). Basal regions of trochanter, femur, and tibia have 1–12, 4 + 8, and 2–4 chordotonal organs, respectively. Surface of legs has fine pilosity; femur and tibia with three or four longitudinal rows of long setae (sometimes incomplete or partially merged) (Suppl. material 1: fig. S 17 C). Apical region of femur with supplementary setae, as in the two previously described species. Tibial comb with four spines, the outer ones shorter than the inner ones (Suppl. material 1: fig. S 17 A). Apical spines present on the first (1 or 2 spines) and second (1 spine) foreleg tarsomeres; on the first (2 spines), second (2 spines), and third (variable, usually 2 spines) midleg tarsomeres; and on the first, second, and third (2 spines each) hindleg tarsomeres. Basitarsus of all legs highly setulose, particularly the hind one. Claws symmetrical, each with a small basal tooth (sometimes inconspicuous) plus one associated bristle (Suppl. material 1: fig. S 17 D – F).</p><p>Wing. Length 1.02 ± 0.07 mm; width of 0.39 ± 0.02 mm, and CR: 0.46 (0.41–0.5). Veins distinct, radial cells highly sclerotized, with marked yellow tones (Suppl. material 1: fig. S 17 G).</p><p>Abdomen. Tergites and sternites pale fawn to pale brown. Tergites sometimes with faint, blurred dark spots on segments 1–2 (Suppl. material 1: fig. S 18 A). Ventral abdominal setae abundant on segments 1–5 (&gt; 60), moderate on segments 6 and 7 (25–35), and scarce on segments 8–10 (&lt;20). Two functional ovoid spermathecae of equal size (48.76 ± 3.11 µm × 41.33 ± 3.72 µm and 50.07 ± 4.49 µm × 41.21 ± 3.07 µm), moderately sclerotized, each with an indistinct discolored apical area, and a small, sclerotized neck (6.18 ± 1.22 µm) (Fig. 5 I). A small rudimentary third spermatheca present.</p><p>Genitalia. On sternite VIII the apices of the lateral arms of the ventral plate have differing orientations and degrees of development but are usually weakly developed (Suppl. material 1: fig. S 18 B). Ventral surface of extremities of segment IV with two fine bristles. Genital atrium large and semicircular (as tall as wide) with four curved bristles of similar size arising posteriorly, and a variable number of smaller bristles covering the margin (Suppl. material 1: fig. S 18 C, D). Behind the atrium lies a pigmented structure with a central excavation (Suppl. material 1: fig. S 18 D). Genital armature of the anal cone trapezoidal in ventral view, with the upper margin weakly sclerotized. Lamella long (241.58 ± 29.8 µm × 70.12 ± 6.15 µm, length × width; index 3.34, 3.07–3.91), slightly darker than abdominal coloration, with 55–65 scattered setae, somewhat longer in the apical region (Fig. 5 H, Suppl. material 1: fig. S 18 B).</p><p>Male. Unknown.</p><p>Etymology.</p><p>The specific epithet pseudoirritans refers to its close morphological similarity to the nominal species L. irritans .</p><p>Distribution.</p><p>This species is known from several locations in the three sampled provinces (Fig. 6).</p><p>Habitat.</p><p>Specimens were collected from Mediterranean scrub adjacent to arable land (diverse crops), although in relatively low numbers (&lt;100) (Fig. 9 A – D).</p></div>	https://treatment.plazi.org/id/F334D35D3AAC5BFA86F207CFD0667B57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Paun-Tanase, Dumitru Ionut;González, Mikel Alexander;Magallanes, Sergio;Naro, Giovanni;Epis, Sara;Figuerola, Jordi	Paun-Tanase, Dumitru Ionut, González, Mikel Alexander, Magallanes, Sergio, Naro, Giovanni, Epis, Sara, Figuerola, Jordi (2026): Hidden diversity in Europe: new species and a revised taxonomy of the subgenus Leptoconops (Leptoconops) (Diptera, Ceratopogonidae). ZooKeys 1279: 201-239, DOI: 10.3897/zookeys.1279.186179
E51EB5930E155D56B318C721398484A8.text	E51EB5930E155D56B318C721398484A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptoconops triangularis Gonzalez & Tanase 2026	<div><p>Leptoconops triangularis González &amp; Tanase sp. nov.</p><p>Fig. 3, Suppl. material 1: figs S 7 – S 12</p><p>Material examined.</p><p>Holotype: SPAIN • ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-6.090697&amp;materialsCitation.latitude=36.417118" title="Search Plazi for locations around (long -6.090697/lat 36.417118)">Pago del Humo</a>, Cádiz; 36.417118, -6.090697; alt. 11 m. 06 Jun. 2024; M. A. González leg.; BGS trap; EBD I-016377 . Paratypes: SPAIN • 1 ♀; labelled as for the holotype, except for EBD I-016378 - 1 ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.907928&amp;materialsCitation.latitude=36.425053" title="Search Plazi for locations around (long -5.907928/lat 36.425053)">La Canaleja</a>, Cádiz; 36.425053, -5.9079281; alt. 54 m; 6 Jun 2024; M. A. González leg.; BGS trap; MNCN_Ent 283582 - 1 ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.963067&amp;materialsCitation.latitude=36.28213" title="Search Plazi for locations around (long -5.963067/lat 36.28213)">Vejer de la Frontera</a>, Cádiz; 36.282130, -5.963067; alt. 59 m; 6 Jun 2024; M. A. González leg.; BGS trap; NHMUK 014316788 - 1 ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.1076546&amp;materialsCitation.latitude=37.44853" title="Search Plazi for locations around (long -5.1076546/lat 37.44853)">Écija</a>, Sevilla; 37.448530, -5.1076545; alt. 183 m; 18 Jun 2024; M. A. González leg.; BGS trap; NHMUK 014316787 - 1 ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.1076546&amp;materialsCitation.latitude=37.44853" title="Search Plazi for locations around (long -5.1076546/lat 37.44853)">Écija</a>, Sevilla; 37.448530, -5.1076545; alt. 183 m; 18 Jun 2024; M. A. González leg.; BGS trap; MNCN_Ent 283583 - 1 ♀; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.963067&amp;materialsCitation.latitude=36.28213" title="Search Plazi for locations around (long -5.963067/lat 36.28213)">Vejer de la Frontera</a>, Cádiz; 36.282130, -5.963067; alt. 59 m; 6 Jun 2024; M. A. González leg.; BGS trap; MNCN_Ent 283584 .</p><p>Diagnosis.</p><p>The abdominal terga possess characteristic sharply defined, black triangular spots that are large on segments I – IV but become progressively smaller and more diffuse towards the apex (segments V – VII) (Fig. 3 B). This character is unique among the Leptoconops species recorded in the Western Palearctic.</p><p>Description.</p><p>Females medium-sized (1.63 ± 0.10 mm). Thorax dorsally pruinose with green metallic reflections; abdomen pale cream to light brown (almost orange), with distinctive black triangular spots on the tergites; legs bicolored (Fig. 3 A, B, Suppl. material 1: fig. S 7 A, B).</p><p>Head. Subspherical, dull-pruinose dark with metallic-green tones on the clypeus, frons, and vertex (Suppl. material 1: fig. S 7 C, D). Head width (445.12 ± 43.02 µm) exceeds head length (372.30 ± 34.55 µm) (Suppl. material 1: fig. S 7 D). Head width 4.3 × (4.1–4.6) the interocular distance. Two lateral interocular setae present (one per side) that are sometimes hard to see (Suppl. material 1: fig. S 7 E). Vertex moderately spiculated under 400 x magnification and have 8–10 setae aligned in a row, four setae within the upper interocular space. Eyes bare, separated by the diameter of 8 ommatidia (102.68 ± 14.35 µm) (Suppl. material 1: fig. S 7 D, E).</p><p>Antenna. Pedicel usually with three setae; flagellomere I has six setae (4 + 2). Lengths of flagellomeres I – XII are provided in Suppl. material 1: table S 2. Total antennal length = 485.68 ± 17.92 µm; AR = 0.68. Proximal flagellomeres II – IV longitudinally flattened, V subspherical, and VI – XI becoming progressively spherical (Suppl. material 1: fig. S 8 A). Terminal flagellomere elongate, similar in length to the preceding 2–2.5 segments combined, densely setose, with various minute hyaline ss and four or five sensilla chaetica (1 apical, 1 basal, and 3 beyond the mid-section) (Fig. 3 D, Suppl. material 1: fig. S 8 B). Antenna with 8–14 sensilla trichodea arranged in two or three semi-verticils on both faces of each flagellomere (Suppl. material 1: fig. S 8 C). Distinctive and conspicuous sensilla chaetica present on one face: three or four long and curved on flagellomeres I – VIII, and one or two entirely straight on flagellomeres IX – XI (Suppl. material 1: fig. S 8 D). Minute hyaline ss arranged on the distal third in variable numbers (2–5 per flagellomere). A pair of thin hyaline ss present on flagellomeres I – XI, situated sub-laterally on one face, curved beyond mid-length, and proximate (Fig. 3 E, Suppl. material 1: fig. S 8 E).</p><p>Mouthparts. Dark brown. Clypeus setae with the same configuration as the previous species (Suppl. material 1: fig. S 7 F). Proboscis 204.27 ± 18.49 µm, shorter than head length. Palps slightly protrude beyond the proboscis tip; total palpal length 226.83 ± 19.83 µm; PR = 1.56 (1.40–1.70). First two palpal segments (length 44.36 ± 11.51 µm) each with three long setae. Third segment (Fig. 3 C) large and moderately incrassate, markedly longer than wide (97.25 ± 4.28 µm and 62.43 ± 5.01 µm), with 6–11 setae (2–4 on the outer margin, 2–4 in sensory area, and 2 or 3 on the inner margin). Sensorium with 70–85 sensilla arranged in 6–8 irregular rows, occupying ~ 3 / 4 of surface with sensory pits; sensillar area 1,755.57 ± 35.31 µm 2. Fourth segment (85.21 ± 8.31 µm) cylindrical, with one or two median setae (usually 1), one or two subapical setae (usually 1), and two or three apical setae (Fig. 3 C, Suppl. material 1: fig. S 9). Lacinia has 21–24 teeth, typically curved apically (Suppl. material 1: fig. S 9). Mandible armed with 15–20 teeth (Suppl. material 1: fig. S 9).</p><p>Thorax. Mesonotum pruinose with green or greyish reflections, depending on light, length 553.75 ± 30.08 µm, width 451.66 ± 35.16 µm (Suppl. material 1: fig. S 10 A). Mesonotum bears brown setae arranged in three longitudinal rows (one acrostical and two dorsocentral), and with scattered setae antero- and post-laterally. Three distinctive supraalar setae present on each side. Humeral callus prominent with angular edges (Suppl. material 1: fig. S 10 A). Humeral pits consisting of three or four small, rounded black spots aligned anteriorly, with an additional group of three or four black pits centrally in the prescutellar area (Suppl. material 1: fig. S 10 B, C). Anepisternum without setae; katepisternum pruinose with 8–10 scattered setae on each side. Scutellum pruinose with two (subapical and lateral) or three (1 subapical and 2 lateral, the outer lateral pair shorter) pairs of setae. Post-scutellum strongly arched, pruinose, and bare. Halters yellow apically and dark basally (Suppl. material 1: fig. S 10 B).</p><p>Legs. Bicolored, with alternating dark and pale patterns. Measurements in Suppl. material 1: table S 3. Foreleg (1,545.09 ± 58.79 µm) and midleg (1,628.80 ± 124.13 µm) shorter than hindleg (2,066 ± 98.10 µm). Femur and tibia uniformly dark brown. Foreleg tarsomeres: first bicolored (three-quarters dark, one quarter pale), others uniformly dark (Fig. 3 F). Midleg and hindleg tarsomeres: first and second pale except for dark apical tips; others uniformly dark (Fig. 3 G, H). Trochanter, femur, and tibia bear 12–14 (2 groups), 10–12 (basal region, usually 8 + 4), and three or four (basal region), respectively, chordotonal organs. Trochanter with eight bristles (6 distinctive and long, 2 weak, closely positioned, straight). Leg surface with fine pilosity and 3–5 rows of setae arranged in a row or irregularly. Apical regions of femur and tibia have additional long bristles. Tibial comb with four spines, the central two equal in size (largest), the outer two smaller and unequal (Fig. 3 K, Suppl. material 1: fig. S 11 A). Hind basitarsus highly setulose. Apical spines present on the first (1 spine) and second (1 spine) tarsomeres of foreleg, on first, second, third, and fourth (1 or 2 spines each) tarsomeres of midleg, and on first (2–4 spines), second (2 spines), third (1 or 2 spines), and fourth (variable) tarsomeres of hindlegs. Claws similar on all legs, slightly undulate in profile, symmetrical, each with a thick, pointed basal tooth and a bristle arising from its base (Suppl. material 1: fig. S 11 B, C). Empodium with a branched bristle.</p><p>Wing. Length 1.14 ± 0.09 mm; width of 0.46 ± 0.02 mm, CR: 0.43 (0.39–0.53). Veins distinct; radial cells highly sclerotized and yellowish (Suppl. material 1: fig. S 11 D, E).</p><p>Abdomen. Tergites and sternites pale fawn to pale brown. Tergites 1–4 with well-defined triangular black spots (Fig. 3 B). Ventral setae more abundant on segments 1–5 (&gt; 30), moderately numerous on segments 6 and 7 (15–25), and less abundant on segments 8–10 (&lt;16). The two functional elongate spermathecae are nearly equal in size (74.54 ± 6.3 × 22.29 ± 2.03 µm and 70.71 ± 3.65 × 45.76 ± 1.53 µm), highly sclerotized, and have a discolored apical region (minute yellow spots), and a distinct tronco-conical sclerotized neck (9.89 ± 1.26 µm) (Fig. 3 I, J, Suppl. material 1: fig. S 12 A, D). A small, rudimentary third spermatheca is present (Suppl. material 1: fig. S 12 A).</p><p>Genitalia. Sternite VIII and the extremities of the lateral arms of the ventral plates show basal striations and well-developed extremities with variable orientation (Suppl. material 1: fig. S 12 B, C). Ventral extremities of segment IV form a sclerotized structure with two or three fine bristles. The genital atrium is semicircular (wider than long), with four long, strongly curved bristles, arising from enlarged posterior bases, of similar size. Base with a moderate number of weaker bristles (Suppl. material 1: fig. S 12 B, C). Behind the atrium lies a pigmented structure with a concave medial region and sclerotized margins. Genital armature of the anal cone large, V-shaped in ventral view, and strongly sclerotized (Suppl. material 1: fig. S 12 A – C). Lamella dark greyish, not especially long (195.65 ± 15.45 µm × 71.50 ± 6.85 µm, length × width; index 2.7, 2.2–3.5), and bear 50–60 scattered setae, longer apically (Fig. 3 I, J, Suppl. material 1: fig. S 12 A).</p><p>Male. Unknown.</p><p>Etymology.</p><p>The specific epithet triangularis refers to the distinctive triangular black spots on the abdominal tergites that characterize this species.</p><p>Distribution.</p><p>This species is known from just a few locations in the three sampled provinces (Fig. 4).</p><p>Habitat.</p><p>Specimens were collected in low numbers (&lt;20) from Mediterranean shrub vegetation bordering arable land (Fig. 9 A, B, D).</p></div>	https://treatment.plazi.org/id/E51EB5930E155D56B318C721398484A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Paun-Tanase, Dumitru Ionut;González, Mikel Alexander;Magallanes, Sergio;Naro, Giovanni;Epis, Sara;Figuerola, Jordi	Paun-Tanase, Dumitru Ionut, González, Mikel Alexander, Magallanes, Sergio, Naro, Giovanni, Epis, Sara, Figuerola, Jordi (2026): Hidden diversity in Europe: new species and a revised taxonomy of the subgenus Leptoconops (Leptoconops) (Diptera, Ceratopogonidae). ZooKeys 1279: 201-239, DOI: 10.3897/zookeys.1279.186179
