taxonID	type	description	language	source
2B718793FFFA1417FF5EE1DD055358A0.taxon	type_taxon	Type species: Frischianus rufulus (Kraatz, 1859), fixed here by original designation.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFFA1417FF5EE1DD055358A0.taxon	diagnosis	Diagnosis: Until now, four previously described species herein assigned to Frischianus had been included in Scopaeus. The two genera are distinguished as follows. Frischianus lacks the stridular file of the metaventrite and plectral ridges of the mesofemur that, together, comprise the stridulum, a defining feature of Scopaeus (Herman 2023: Figs 189 – 191, and many other illustrations). The metakatepisternal processes of Frischianus are small, apically rounded, narrowly separated lobes on the posterior margin of the metaventrite between the metacoxae (Fig. 26); for Scopaeus they are large, long, and tapered to acute apices (Herman 2023: Figs 89, 97, among many examples). The dorsal surface of the aedeagal median lobe of Frischianus is divided from the base to the ostium by a midlongitudinal membrane (Figs 36 – 38, 42) or there is an elongate, oval membrane separated from the ostium by sclerotization (Figs 39 – 41); the dorsal aedeagal surface is fused in Scopaeus (Herman 2023: Fig. 158). Three characters set Frischianus apart from all other genera of the Scopaeina: the nearly equal length and width of the squarish prothorax (Figs 1 – 4, 43 – 60); the reduction of the metakatepisternal processes to small, apically rounded lobes (Fig. 26); and the single gonocoxal plate of the females (Figs 29, 248 – 269). Among the other five scopaeine genera, the prothorax is distinctly longer than wide, the metakatepisternal processes are moderately large to large and apically rounded or pointed (Herman 2023: Figs 251, 270, 300, 335), and females have a pair of gonocoxal plates (Herman 2023: Figs 78, 80, 264, 290, 332). If the ventral side of segment IX is exposed and the genital sclerites visible, the females are easily recognized as Frischianus without dissection. The following are additional characters that separate Frischianus from other scopaeine genera. The neck of Frischianus is about a sixth to a fifth as wide as the head (Figs 1 – 4, 18, 43 – 60); the neck of Trisunius, most species of Orus, and most of the New World species of Micranops is wider, about a third of the width of the head. A few species of Orus and most of the Old World species of Micranops (Herman 2023: 131, 132, Table 3; Frisch, pers. comm.) have a narrow neck, about a fifth to a quarter of the width of the head, but are separated from Frischianus, not only by the larger, metakatepisternal processes, but also by the position of the trichobothrium. The trichobothrium of Frischianus is near the middorsal margin of the eye (Figs 5 – 9), for Orus the trichobothrium is adjacent to the posterodorsal margin of the eye (Herman 2023: Figs 297, 309), and for Micranops in a cavity behind the eye (Herman 2023: Figs 266, 267, 282). Frischianus and Hyperscopaeus, both with narrow necks, are separated, not only by the differential size of their metakatepisternal processes, but also by the widely separated gular sutures of the former (Fig. 18) and narrowly separated to contiguous gular sutures of the latter (Herman 2023: Fig. 244). Ancillary characters that help define Frischianus include a well-developed, almost straight, knife-edged epigenal ridge extending from the trichobothrium to the anterior margin of the antennal hump (Figs 5 – 9); the dorsoposterior side of the ridge has a contiguous groove with a polished surface (Figs 5 – 9). Some species of Hyperscopaeus have a poorly developed epigenal ridge that lacks the contiguous groove, is steeply curved ventrad, and usually does not reach the trichobothrium. Some species of Trisunius have a short epigenal ridge that is confined to the antennal hump, lacks a contiguous groove, and does not reach the eye. The metatibia of Frischianus has a small, apical comb on the outer surface (Figs 30 – 33). Only a few species of Hyperscopaeus have an outer, apical, metatibial comb (Fig. 35). An outer metatibial comb is also known in Trisunius (Fig. 34).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFFA1417FF5EE1DD055358A0.taxon	description	Description: Length from apex of labral denticles to posterior margin of abdomen 2.4 – 4.9 mm; forebody length from apex of labral denticles to posterior end of elytral suture 1.5 – 2.3 mm. Head with postocular lateral margin straight (e. g. Figs 3, 43, 52) to shallowly and broadly rounded (e. g. Figs 46, 48); basal angles distinct, strongly rounded, and well developed (e. g. Figs 43 – 45); basal margin truncate (e. g. Fig. 43, 45, 48) to broadly and shallowly (e. g. Figs 1, 44, 54) to moderately deeply emarginate (e. g. Figs 3, 46, 49, 53, 56); basal margin with median, shallow (rarely almost imperceptible) to moderately deep, narrow to wide, vertical groove (e. g. Figs 4, 49, 53); posteroventral surface with or without small tubercle laterad of neck. Dorsal cephalic surface with dense, setate punctation; punctures small and fine to moderately large to large and coarse; punctures discrete and separated from each other (e. g. Figs 3 a, 4 a, 57, 58) or some punctures partially confluent and arranged in short linear clusters (e. g. Figs 2 a, 44); macrosetae few, mostly confined to or near margins, dorsum with one on each side of midlongitudinal line about halfway between clypeal and posterior margins. Neck petiolate, nuchal groove deep and strongly constricted (Fig. 18), and base of neck abruptly expanded; neck across nuchal constriction one fifth to one sixth as wide as greatest postocular width of head (Fig. 18); nuchal ridge strongly developed, present dorsally and laterally. Trichobothrium contiguous with dorsal margin of eye at about or slightly forward of midpoint (Figs 5 – 9); bothrium (Fig. 5) in deep, rounded trichobothrial depression; trichobothrial depression without setae (Figs 5 – 9). Epigenal ridge extending dorsoanteriorly from trichobothrial depression to anterior margin of antennal hump; ridge knife-edged for entire length (Figs 5 – 9); groove present and contiguous with full length of dorsoposterior side of ridge; groove moderately wide to narrow and continuous with trichobothrial depression (Figs 5 – 9). Eye with posterior margin broadly rounded (Fig. 6) to slightly flattened (Figs 7, 8); corneal sensilla absent. Gular sutures moderately widely separated anteriorly, widely separated posteriorly (Fig. 18). Gula depressed; surface microreticulate anteriorly and polished posteriorly; extent of microreticulation variable, restricted to small anterior region to extending to about posterior third and remainder polished; some species with matt surface behind microreticulation. Mandibles with three large denticles on left and four on right (Fig. 16); both mandibles with minute, basal denticule or point present and distinct or hardly evident or absent (Note: Minute basal denticule or point not included for count of denticles). Labrum quadridentate (Figs 10 – 14), or rarely hexadentate with additional, small, apicolateral denticle (Fig. 15); denticles tapered to acute apex, for some species apparent sublateral denticle evident as pointed (Fig. 10) or rounded (Figs 13, 14) lobe; submedial denticle larger and longer than sublateral denticle; submedial and sublateral denticles rarely of same length (Fig. 15); sublateral denticle moderately large to tiny. Antennal scape stout, large and parallel-sided to near base, then slightly narrowed in some species (Figs 1 – 4, 18). Prothorax strongly hexagonal in dorsal view (Figs 1 – 4, 43 – 60); length slightly to moderately longer than wide (length / width: 1.01 – 1.13, Ø 1.04); dorsal surface with or without narrow, midlongitudinal, impunctate strip and without midlongitudinal groove on most of length; dorsal surface broadly and shallowly convex; surface with dense, very fine to moderately coarse, setate punctation; punctures discrete and separated (Fig. 2 a), or surface with jumbled mix of short, contiguous rows of punctures and irregularly arranged punctures (Figs 1 b, 3 b), or with long, more regular, longitudinal rows, and surface striate and appearing wrinkled (Fig. 4 b); macrosetae absent. Prohypomeron with postprocoxal lobe separated from remainder of hypomeron by prohypomeronal transverse ridge (Figs 19, 20). Mesoventrite with deep, oval, median depression (e. g. Fig. 25); surface of ventrite with (Fig. 22) or without (Fig. 21) reticulate sculpturing; extent of surface reticulation variable (Figs 23 – 25); median depression and posterior median surface often without reticulation, but present in some species; surface without sculpturing polished; surface with few ridges; anterior mesanapleural ridge present (e. g. Fig. 22), often strongly reduced; posterior mesanapleural ridge present (e. g. Fig. 24) or absent (e. g. Fig. 25). Mesofurcasternum (e. g. Fig. 22) without internal furcasternal apophysis on posteromedial margin (cf. Herman 2023: Figs 82, 241, 250). Mesofemur without plectral ridges (Herman 2023: Figs 85, 106, 121, 185, 203). Elytral surface with dense, moderately large to tiny, setate granules (Figs 1 c – 4 c); seta arising from top or posterior side of granule; surface with three rows of punctures (rows most effectively viewed using diffused light); punctures few and within each row well separated; rows extending from near anterior elytral margin to about one to two thirds of elytral length; sutural row deeply impressed, most strongly developed, and always present; median and lateral rows moderately impressed to not impressed and slightly to poorly developed to imperceptible, but row of punctures usually discernable. Wings fully developed (and likely permitting flight) to reduced to short pads of about three-fifths to about equal to elytral length (and likely flightless); some species with macropterous and micropterus individuals in same population. Metaventrite without stridular file (cf. Herman 2023: Figs 105, 125, 154, 155, 189). Metakatepisternal process present as small, apically rounded bump or knob on posterior, metaventral margin (Fig. 26; cf. Herman 2023: Figs 26, 89, 97, 241, 270, 335). Metatibia with large inner and small outer apical comb (Figs 30 – 33); inner comb with many long tines; outer comb with significantly fewer and shorter tines than inner comb; outer comb with about four to eight tines. Sternite II with median point of posterior margin present (Fig. 27). Sternite III with long, intercoxal carina (Fig. 27). Sternite IV with (apparent) glandular lobe on anterior margin (Fig. 28). Male: Sternite VII with or without modification of median surface and posterior margin; median surface often with broad, shallow depression (Figs 206, 223) and medioposterior cluster of dark macrosetae (Figs 217, 219, 221, 223); posterior margin of most species truncate (Fig. 206) or some species with shallow (Figs 217, 225) to moderately deep (Fig. 233) median emargination. Sternite VIII with emarginate posterior margin; emargination with variable depth, width, and form (e. g. Figs 199, 200, 227, 232, 234). Aedeagus with dorsal surface of median lobe midlongitudinally membranous from base to ostium (Figs 36 – 38, 42) or with elongate membranous oval and separated from ostium by sclerotization (Figs 39 – 41). Female: Sternite VII with truncate posterior margin. Sternite VIII with rounded posterior margin, perhaps rarely with shallow emargination (Fig. 212). Abdominal segment IX with one, moderately wide to very wide gonocoxal plate (Fig 29); posterior margin emarginate; lateral margin sinuate with base slightly rounded and about as wide as apex (Figs 249 – 265) to greatly enlarged and significantly wider than apex (Figs 266 – 269). Spermatheca hyaline, vesicular (Fig. 239), connected to sclerotized sperm pump (Figs 237 – 247) consisting of chamber segment (Fig. 239) and accessory terminal segment (Fig. 239); chamber segment without process of Scopaeus (cf. Frisch et al. 2002: 28); bursal duct right proximal of sperm pump straight (Figs 237 – 244) or curved (Figs 245 – 247), gradually more weakly sclerotized proximally.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFFA1417FF5EE1DD055358A0.taxon	distribution	Distribution: Frischianus is known from the Indian Subcontinent (India: Kerala, Tamil Nadu) and Southeast Asia (Thailand: Chiang Mai; Malaysia: Pahang, Penang, Perak, Sabah, Sarawak, Selangor; Indonesia: Aceh, Bali, Banten, Jambi, Jawa Barat, Jawa Tengah, Sulawesi Tengah, Sumatera Barat, Sumatera Utara; the Philippines: Mindanao). It is likely species will be found in southern China, Myanmar, Laos, Vietnam, and Brunei.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFFA1417FF5EE1DD055358A0.taxon	biology_ecology	Habitat: Label data and personal collection experience of Frisch suggest a riparian habitat for Frischianus similar to the preference of Scopaeus (Frisch et al. 2002: 28). In Java, Frisch collected series of F. laticollis, F. similis sp. nov., F. communis sp. nov., and the holotype of F. setifer sp. nov. in humid, slightly vegetated places along banks of creeks and rivers, below thin layers of phytodebris, under stones, and in the upper layer of sandy, gravelly soil. Frisch observed that the abundance of specimens and diversity of species decreased significantly as he moved toward shadier and dryer places away from the water. The sparse habitat information on the labels of specimens borrowed from collections is imprecise and generalized (secondary forest, logged forest, oilpalm plantation, rainforest, primary forest), but often indicates humid and riparian habitats (paddy field, riparian forest strip, near stream, Poring Hot Springs, sifted from phytodebris accumulated by flood). Frischianus uhligi sp. nov. was sifted from forest litter, although it remains unclear whether the collection site was near water. Species of Frischianus were collected from lowlands to 2,600 meters elevation (F. pedator sp. nov.; Mt. Singgalang, Sumatra) and 3,000 meters (F. subalpinus sp. nov.; Mt. Kerinci, Sumatra).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFFA1417FF5EE1DD055358A0.taxon	discussion	Remarks: The presence in Frischianus of the paraocular trichobothrium (Figs 5 – 9), tripartite ligular lobe (Fig. 17), long and tapered profurcasternum (in ventral view, Fig. 19), trilobed anterior margin of sternite II (Fig. 27), and absence of the pronotal marginal ridge (Fig. 20) support placing the genus in the Scopaeina. Four species are transferred herein from Scopaeus as new combinations in Frischianus. Frischianus rufulus (Kraatz 1859: 140), from Myanmar, was described in Lithocharis, transferred to Medon in the subgenus Lithocharis by Bernhauer and Schubert (1912: 243), then to Scopaeus by Cameron (1931: 177). Frischianus laticollis (Cameron 1925: 183), from Indonesia, was described in Scopaeus. Frischianus miscellus and Frischianus coriaceus, both from Malaysia, were described in Medon by Cameron (1932: 129 and 136, respectively), who (Cameron 1950: 15) later moved them to Scopaeus. Along with these four, Frisch describes 30 new species in the present work. By the present study, we now know that paederines with the cephalic and cervical characters of Scopaeus, but for which the pronotal length is only slightly greater than the width, is a signal that card mounted specimens must be examined for the presence of the epigenal ridge and / or removed from the card to ascertain the existence of a stridulum and condition of the metakatepisternal processes. In a key to the genera of the Scopaeina (Herman 2023: 61), Frischianus runs to Hyperscopaeus in couplet 4. Characters separating the two genera are listed above in the second paragraph of the Diagnosis for Frischianus.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFFA1417FF5EE1DD055358A0.taxon	etymology	Etymology: By choosing the name Frischianus, Herman honors his long-time friend and colleague, Johannes Frisch, of the Museum für Naturkunde Berlin, for his insightful work on Scopaeus that has resulted in numerous beautifully detailed studies and illustrations of the widespread, speciose genus Scopaeus and who was the first to identify both this cluster of species as differing from Scopaeus along with the discovery that Hyperscopaeus was also a genus separate from Scopaeus. His studies of Scopaeus are among the best ever published for the genus and will be a model and guide for work on the genus far into the future. The gender of the name is masculine.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFF71417FCFCE09D030F5942.taxon	diagnosis	Diagnosis: Distinguished from remaining species lineages of F. laticollis subgroup (chapter 3.2.2.1.) by ventral endophallic sclerite apically extended in deeply emarginate ventral lobe and two dorsolateral spines with crossed ends strongly protruding from ostium of aedeagus (Figs 61 – 66), and by gonocoxal plate of ♀ with sclerotization triangularly extended across membranous, basolateral dilation (Figs 248, 249). Separate from F. schuriani and F. setifer species lineages furthermore by presence of dorsal endophallic spine (Figs 61 – 66).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFF71417FCFCE79D03295B21.taxon	diagnosis	Diagnosis: Separated from F. rufulus subgroup (chapter 3.2.2.2.) as follows: Submedial labral denticles pointed, usually longer than pointed, rarely obtuse, sublateral labral denticles (Figs 10 – 12, 14). Abdominal sternite VIII of ♂ with medioposterior incision narrow (e. g. Figs 199 – 205). Aedeagus (Figs 61 – 156) with apical lobe present and ventral process absent; apical lobe more or less, often strongly, shifted sinistrad and curved dextrad; endophallic sclerite variable in size and shape, small to enlarged with various modifications, with foramen indiscernible or moderately developed. Gonocoxal plate of ♀ (unknown in eight of 22 species included) with posterior margin widely, but moderately deeply emarginate (Figs 248 – 256, 258 – 262).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFF71417FF5EE23D02D75C21.taxon	diagnosis	Diagnosis: Separated from F. strigaticollis species group (chapter 3.2.3.) as follows: Pronotal punctation type 1 – 3 (Figs 1 b – 3 b). Aedeagus (Figs 61 – 180) asymmetrical; dorsal lobe absent; parameres vestigial, longitudinally contiguous with circoforamen, in ventral and dorsal view not laterally protruding from contours of phallobase (e. g. Figs 64 – 66); dorsal, midlongitudinal split of aedeagus ending proximally of and not opened into ostium (types A, B, C; Figs 36 – 41). Gonocoxal plate of ♀ (unknown in 11 of 29 species included) subrectangular without hypertrophic, anterior enlargement (Figs 248 – 265). Bursal duct at transition to sperm pump straight or slightly curved (Figs 237 – 244). Distributed in Southeast Asia [Myanmar, Thailand (Chiang Mai), southern Philippines (Mindanao), Malay Peninsula, Borneo (Sabah, Sarawak), Sumatra, Sulawesi, Java, Bali].	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFF71408FCFCE27C07E35981.taxon	description	(Figs 1, 43, 61 – 66, 199, 237, 248, 249)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFF71408FCFCE27C07E35981.taxon	materials_examined	Type specimen: Lectotype ♂, Malaysia, Selangor: The Gap, leg. Cameron (NHML); labelled “ Type ” (round, printed, red edged label), “ The Gap, / Selangor, F. M. S. / Dr. Cameron. ” (printed), “ M. Cameron / Bequest / B. M. 1955 - 147. ” (printed), “ Bank / of Stream ” (printed), “ Debris ” (printed), “ M. / coriaceus / TYPE Cam. ” (handwritten, “ TYPE ” in red), “ Lectotype / Medon coriaceus / Cameron, 1932 / des. Frisch & Herman, 2025 ”; here designated. The original description of Medon coriaceus (Cameron 1932: 136) contains no information regarding the number of underlying specimens nor does it specify a holotype by original designation (ICZN 1999: Article 73.1.1.). However, as male characters are described, at least one male must have been available to the author. To stabilize the name Medon coriaceus according to ICZN 1999, Article 74.1., we designate as the lectotype the only available specimen, a male from the Cameron collection at NHML, which is labelled “ Type ”, originates from the type locality The Gap and bears a handwritten identification label which agrees with the example of Cameron’s handwriting in Horn et al. (1990: 477). New records: Malaysia, Pahang: Ringlet, 1250 m, 26. III. 1993, leg. Calame & Löbl (MHNG); Bukit Fraser, Tiong-Layang trail, 1300 m, 18. III. 1993, leg. Calame & Löbl (MHNG); Bukit Fraser, 1280 m, 17. IX. 1972, leg. Jaccoud (MFNB); Tanah Rata (Cameron Highlands), 19. – 23.3.2008, leg. Hammond (NHML).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFF71408FCFCE27C07E35981.taxon	description	Description: Habitus (Figs 1, 43). Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color unicolored medium brown or elytra with darker brown lateroposterior angles in variable expansion; tip of abdomen, antennae, palps, and legs yellow brown. Labral denticulation type 1 (Fig. 10). Forebody surface microsculpture type 1 (Fig. 1). Elytral rows of punctures present; sutural row of punctures faintly impressed anteriorly, occupying about anterior 0.7 of elytral sutural length. Total body length 3.2 – 3.6 mm; forebody length 1.8 – 2.0 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with triangular, medioposterior incision with concave lateral margins, occupying roughly 0.3 of sternite length (Fig. 199). Aedeagus (Figs 61 – 66) with short apical lobe; apical lobe in lateral view slender with apical end minutely pointed ventrally (Figs 61, 64), in ventral view widely triangular with very narrow, short apical end shifted sinistrad and strongly bent dextrad (Figs 62, 65). Endophallus in ventral and dorsal view situated medially in aedeagus (Figs 62, 63, 65, 66). Endophallic sclerite with robust, broad base, continued distally in large ventral lobe and two dorsolateral spines (Figs 61 – 66); ventral lobe and dorsolateral spines of endophallic sclerite strongly protruding from ostium; ventral lobe of endophallic sclerite by deep, wide, ventral emargination divided in two ventrolateral lobes with round apical end (Figs 61 – 66); dorsolateral spines of endophallic sclerite slightly curved ventrad (Figs 61, 64), in dorsal view aligned toward each other with ends crossing (Figs 63, 66). Dorsal endophallic spine thin, reaching apices of dorsolateral spines of endophallic sclerite, faintly curved ventrad (Figs 61, 64), in dorsal view straight, pointing longitudinally, and with apical end running ventral of crossed ends of dorsolateral endophallic spines (Figs 63, 66). Dorsal, midlongitudinal split of phallobase typeA (Fig. 36). Aedeagus with ventral margin between postforamen and apical lobe straight (Figs 61, 64), in dorsal view oval with slightly curved lateral margins (Figs 63, 66). Length of aedeagus 0.43 – 0.46 mm. Female: Gonocoxal plate (Figs 248, 249) with basolateral region membranous and gradually widened laterally; sclerotized, setose portion of gonocoxal plate widely emarginate posteriorly, triangularly extended across membranous enlargement anteriad, and ending in variable tip; sclerotized, triangular extension of gonocoxal plate without basolateral ridge, not hard set off from adjacent membrans.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFF71408FCFCE27C07E35981.taxon	distribution	Distribution: Frischianus coriaceus is widely distributed in the Malay Peninsula and recorded from the Cameron Highlands (Ringlet) in the north as far south as The Gap and Fraser’s Hill.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE81408FCFCE09D03D65921.taxon	diagnosis	Diagnosis: Separated from remaining lineages of F. laticollis subgroup (chapter 3.2.2.1.) by small (Figs 67 – 105) to moderately enlarged (Figs 106 – 117) or long spiniform (Figs 118 – 135) endophallic sclerite. Different from F. schuriani and F. setifer lineages also by presence of dorsal endophallic spine (Figs 67 – 135).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE81408FCFCE09D03D65921.taxon	discussion	Remarks: With 14 species included, the Frischianus laticollis lineage of the F. laticollis subgroup is the most speciose and most widespread terminal lineage of Frischianus as far as known presently. It is recorded from the Malay Peninsula, Sumatra, Borneo, Java and Bali.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE8140DFCFCE29C05925981.taxon	description	(Figs 2, 44, 67 – 69, 200, 238, 250)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE8140DFCFCE29C05925981.taxon	materials_examined	Type specimens examined: Lectotype ♀, Indonesia, Jawa Barat, Cibodas, 400 m, VIII. 1921, leg. Dammerman; labelled “ Tjibodas / 1400 M. / VIII. 1921 ” (printed except for date), “ Scopaeus / laticollis Cam. / TYPE ” (handwritten), “ Type ” (red, printed), “ Lectotype / Scopaeus laticollis / Cameron, 1925 / des. Frisch & Herman, 2025 ” (NHML); here designated. Paralectotypes (2 specimens): 1 ♀, labelled “ Tjibodas / 1400 M. / VIII. 1921 ” (printed except for date), “ Cameron det. ” (handwritten), “ Syntypus ” (red, printed), “ Scopaeus / laticollis Cam ” (handwritten), “ coll. DEI / Eberswalde ” (printed), “ Paralectotype / Scopaeus laticollis / Cameron, 1925 / des. Frisch & Herman, 2025 ” (SDEI); 1 ♀, labelled “ Tjibodas / 1400 M. / VIII. 1921 ” (printed except for date), “ Scopaeus laticollis Cam. / Cotype ”, “ M. Cameron / Bequest / B. M. 1955 - 147. ” (printed), “ Paralectotype / Scopaeus laticollis / Cameron, 1925 / des. Frisch & Herman, 2025 ” (NHML). Cameron (1925: 183, 184) described Scopaeus laticollis based on “ 3 examples ” from Cibodas [Tjibodas], which we examined. Even though Cameron labeled one of them as “ Type ”, these specimens are syntypes, because the author did not publish a holotype by original designation (ICZN 1999: Article 73.1.1.). Thus, a lectotype designation is required to stabilize the name S. laticollis Cameron, 1925 according to ICZN 1999, Article 74.1. As no male syntype is available, we select as the lectotype a female from the Cameron collection at NHML, which was collected at the type locality Cibodas and labeled “ TYPE ” by Cameron himself as can be seen from the example of his handwriting in Horn et al. (1990: 477). Frischianus laticollis is one of three closely related Javanese species that can accurately be distinguished only by the aedeagus. The other two species are described below as F. similis sp. nov. and F. communis sp. nov.. Even though no male types are available, F. laticollis can be interpreted quite reliably. Both the female lectotype and the two female paralectotypes differ from the females of F. similis and F. communis by the slightly longer gonocoxal plate (Fig. 250) and agree in this feature with a series that was later collected at the type locality Cibodas of F. laticollis and also contains males. We consider it a pragmatic solution to refer the species group name laticollis Cameron, 1925 to this species. New records: Indonesia: Jawa Barat: Bogor, NOslope of Mt Salak (06 ° 39 ' 55 " S, 106 ° 45 ' 36 " E), 640 m, 13. IX. 2015, leg. Frisch (MFNB, ZMB); Bogor: Lake Warna, VIII. 1984, leg. Robert (MHNG); Bogor, Cibodas, 1400 m, 3. – 6. IX. 1989, leg. Agosti, Burckhardt & Löbl (MHNG); Sukabumi, NNW Cidahu: Cirasamala River (S-slope of Mt Salak), (06 ° 44 ' 18 " S, 106 ° 42 ' 52 " E), 1210 m, 29. IX. 2015, leg. Frisch (MFNB); Sukabumi, Cikaniki: Cikaniki River (Mt Halimun, Halimun – Salak National Park) (06 ° 44 ' 46 " S, 106 ° 32 ' 25 " E), 1020 m, 5. X. 2015, leg. Frisch (ZMB); Sukabumi, Sirnarasa: Cimaja River (S-slope of Mt Halimun), (06 ° 51 ' 32 " S, 106 ° 31 ' 06 " E), 670 m, 21. IX. 2015, leg. Frisch (MFNB, ZMB); Cianjur, Mt Gede, 1400 – 1500 m, 24. – 28. V. 1997, leg. Kurbatov (MHNG). Jawa Tengah: Pekalongan, Petungkriono (Nslope Dieng Plateau) (07 ° 06.42 ' S, 109 ° 44.51 '), 1115 m, 22. VIII. 2006, leg. Riedel (SMNK).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE8140DFCFCE29C05925981.taxon	description	Description: Habitus (Figs 2, 44). Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color medium brown to dark brown with lighter brown humeral angles; light brown humeral color often extended across anterior third of elytra and along suture more or less reducing dark color to lateroposterior angles of elytra; tip of abdomen, antennae, palps, and legs light brown. Labral denticulation type 1 (Fig. 10). Forebody surface microsculpture type 2 (Fig. 2). Elytral rows of punctures distinct; sutural row of punctures impressed, occupying anterior half of elytral sutural length. Total body length 3.0 – 3.6 mm; forebody length 1.8 – 2.0 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with deep, narrow, medioposterior incision with feebly concave lateral margins, occupying about 0.4 of sternite length (Fig. 200). Aedeagus (Figs 67 – 69) with apical lobe narrow, arising from sinistral half of truncate apical margin of aedeagus (Figs 68, 69); apical lobe with straight dorsal and convex ventral margin, gradually narrowed toward small, triangular, ventrad pointing apical tooth (Fig. 67). Position of endophallus in aedeagus unknown, because evaginated in all male type specimens. Endophallic sclerite elongate, curved dorsally, with strongly sclerotized, subcircular median portion (Fig. 67), in evaginated state broad in dorsal and ventral view (Fig. 68, 69). Dorsal endophallic spine, in lateral view, curved ventrad and narrowed toward thin, acute apical end (Fig. 67), not protruding from ostium when invaginated (cf. Figs 70, 97), in ventral and dorsal view almost straight with parallel lateral margins and acute apical end (Figs 68, 69). Dorsal, midlongitudinal split of phallobase type B (Fig. 39). Aedeagus with ventral margin between postforamen and apical lobe slightly concave (Fig. 67), in dorsal view long oval, slightly asymmetrical with dextral lateral margin more strongly curved than almost straight, sinistal lateral margin, and somewhat tapered apicad (Fig. 69). Length of aedeagus 0.44 – 0.45 mm. Female: Gonocoxal plate 1.7 times as long as wide, without basal and subbasal ridge, with subparallel, slightly concave lateral margins, shallowly emarginate posterior margin, and biconvex basal margin (Fig. 250).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE8140DFCFCE29C05925981.taxon	distribution	Distribution: Frischianus laticollis is spread in western and Central Java, Indonesia, and recorded from 06 ° 51 ' 32 " S, 106 ° 31 ' 06 " E at Mount Halimun, Jawa Barat, as far east as 07 ° 06.42 ' S, 109 ° 44.51 ', Dieng Plateau in Jawa Tengah.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFEF1403FF5EE03C07155FA1.taxon	description	(Figs 45, 70 – 75, 239, 251)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFEF1403FF5EE03C07155FA1.taxon	materials_examined	Type specimens: Indonesia: Holotype ♂, Banten, Lebak: Citorek Kidul (06 ° 44 ' 51 " S, 106 ° 19 ' 13 " E), 870 m, 23. V. 2016, leg. Frisch (ZMB). Paratypes (16 specimens): 5 ♂, 9 ♀, same data as holotype (MFNB, ZMB); 2 ♂, Jawa Barat, Sukabumi: Ciptarasa (Halimun – Salak National Park) (06 ° 49 ' 52 " S, 106 ° 30 ' 05 " E), 19. IX. 2005, leg. Riedel (SMNK).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFEF1403FF5EE03C07155FA1.taxon	description	Description: Habitus (Fig. 45). Body size, exoskeletal characters, coloring, and shape of abdominal sternite VIII as in Frischianus laticollis (cf. Figs 2, 44, 200), but different according to shape of aedeagus and female gonocoxal plate as follows. Total body length 3.1 – 3.6 mm; forebody length 1.8 – 1.9 mm. Male: Aedeagus (Figs 70 – 72) with apical lobe widened dextrad at base (Figs 71, 72); apical lobe with slightly concave dorsal margin, strongly triangular ventral margin, and round apical end without apicoventral tooth (Fig. 70). Length of aedeagus 0.44 – 0.45 mm. Female: Gonocoxal plate shorter, only about 1.6 times as long as wide, with lateral margins somewhat more deeply concave (Fig. 251).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFEF1403FF5EE03C07155FA1.taxon	distribution	Distribution: Frischianus similis is known only from western Java, Indonesia, where it was found in two close localities in the east of Banten and the very west of Jawa Barat.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFEF1403FF5EE03C07155FA1.taxon	etymology	Etymology: The epithet similis (adjective, Latin: similar) emphasizes the similarity of the aedeagus of this new species to the aedeagus of Scopaeus laticollis and the close relationship of these species, respectively.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFEF1403FF5EE03C07155FA1.taxon	discussion	Comparative note: Frischianus laticollis and F. similis resemble each other in their external morphology (Figs 44, 45). Males can be distinguished according to the shape of the aedeagus. In F. laticollis the aedeagus exhibits a ventral apical tooth of the apical lobe (Fig. 67), while this tooth is absent in F. similis (Fig. 70). A single specimen with an apical tooth of the apical lobe (Figs 73 – 75), that was found within a F. similis sample, perhaps belongs to F. similis according to the subtriangular, angled ventral margin of the apical lobe and might reflect a possible variability of the aedeagus. Its apical tooth is furthermore broad and obtuse (Fig. 73), while in F. laticollis this tooth is narrower and acute (Fig. 67). This specimen is not included in the type series. Females can seemingly be distinguished according to the slightly different proportions of the gonocoxal plate, which in examined specimens of F. laticollis including the lectotype is longer (Fig. 250) than in F. similis (Fig. 251). Examination of further females is, however, necessary to confirm the reliability of the gonocoxal plate proportions for species identification. As far as known, the distributional pattern of F. similis follows that of the closely related F. laticollis immediately to the west, which is why both taxa might be allopatric vicariants at the species or, perhaps, at the subspecies level.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE31403FF5EE71C05B85B01.taxon	description	(Figs 76 – 99, 240, 252)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE31403FF5EE71C05B85B01.taxon	materials_examined	Type specimens: Holotype ♂, Indonesia, Jawa Barat, Bogor, SW Pamijahan (Cimuara Herang River), (06 ° 42 ' 41 " S, 106 ° 41 ' 03 " E), 990 m, 3. X. 2015, leg. Frisch (ZMB). Paratypes (60 specimens): Indonesia: Aceh: Ketambe (Mt Leuser National Park): 1 ♂, 450 m, 26. II. – 1. III. 1991, leg. Bocák & Bocáková (NHMB), 4 ♂, 7 ♀, 300 – 500 m, 23. – 30. XI. 1989, leg. Agosti, Burckhardt & Löbl (MHNG). Jawa Barat: 1 ♀, same data as holotype (MFNB); 1 ♂, 1 ♀, Sukabumi, Cisalimas: Cisalimas River (E-slope of Mt Halimun) (06 ° 45 ' 21 " S, 106 ° 33 ' 38 " E), 870 m, 19. IX. 2015, leg. Frisch (MFNB, ZMB); 1 ♂, 5 ♀, Sukabumi, Sirnarasa: Cimaja River (S-slope of Mt Halimun) (06 ° 51 ' 32 " S, 106 ° 31 ' 06 " E), 670 m, 21. IX. 2015, leg. Frisch (MFNB, ZMB); 1 ♂, 1 ♀, Sukabumi, Simaresmi: Cisareno River (06 ° 49 ' 02 " S, 106 ° 30 ' 09 " E), 1000 m, 22. IX. 2015, leg. Frisch (MFNB, ZMB); 1 ♂, Sukabumi, NNW Cidahu: Cirasamala River (S-slope of Mt Salak) (06 ° 44 ' 18 " S, 106 ° 42 ' 52 " E), 1210 m, 29. IX. 2015, leg. Frisch (ZMB); 2 ♂, Sukabumi, Cikaniki: Cikaniki River (Mt Halimun, Halimun – Salak National Park) (06 ° 44 ' 46 " S, 106 ° 32 ' 25 " E), 1020 m, 5. X. 2015, leg. Frisch (MFNB, ZMB). Jawa Tengah: 1 ♂, Pekalongan, E Petungkriono: mountain N Tinalum (07 ° 06.418 ' S, 109 ° 44.514 E), 1120 m, 22. VIII. 2006, leg. Riedel (SMNK). Bali: 1 ♂, 1 ♀, Mt Agung, 19. VII. 1982, leg. Rougemont (HECO); 3 ♂, 4 ♀, Mt Batukaru [Mt Batukai, Mt Baturau], 500 – 700 m, 28. – 29. X. 1991, leg. Löbl (MHNG, MFNB); 2 ♂, Tabanan: Batukaru, 700 m, 12. VI. 1994, leg. Wunderle (PWCM); 1 ♂, Tabanan: Bedugul, 1200 m, 13. VI. 1994, leg. Wunderle (MFNB). Malaysia: 1 ♂, Pahang, Awana (Genting Highlands), 1150 m, 3. IV. 1993, leg. Löbl & Calame (MHNG); 2 ♂, 2 ♀, Sabah, Koto Marudu, Sorinsim – Tokou [Topou] (6 ° 127.278 N, 116 ° 42.417 E), 22. – 8. III. 1997, leg. Floren (NHMW); Sabah, Ranau, Poring Hot Spring: 1 ♂, 9. IV. 1990, leg. Rougemont (HECO), 8 ♂, 3 ♀, 7. – 12. V. 1987, leg. Burckhardt & Löbl (MHNG); 2 ♂, 2 ♀, Sarawak, Mt Mulu National Park, 50 – 100 m, V. – VIII. 1978, leg. Hammond & Marshall (NHML).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE31403FF5EE71C05B85B01.taxon	description	Description: Body size, exoskeletal characters, coloring (cf. Figs 2, 44), and shape of medioposterior incision of abdominal sternite VIII (cf. Fig. 200) as in Frischianus laticollis, but different according to shape of aedeagus and female gonocoxal plate as follows. Total body length 2.9 – 3.6 mm; forebody length 1.7 – 1.9 mm. Male: Aedeagus (Figs 76 – 99) with apical lobe short, strongly curved ventrad, round to right-angled widened ventrally, variable in width (Figs 76, 79, 82, 85, 88, 91, 94, 97). Dorsal endophallic spine variable, in lateral view slender, weakly to strongly curved ventrad, and with apical end thin and acute, in dorsal view wider, usually curved sinistrad with longitudinally or slightly dextrad directed, more or less dilated apical end (Figs 78, 81, 84, 87, 90, 93, 96). Length of aedeagus 0.41 – 0.44 mm. Female: Gonocoxal plate shorter, only about 1.6 times as long as wide; lateral margins somewhat more deeply concave (Fig. 252).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE31403FF5EE71C05B85B01.taxon	distribution	Distribution: Frischianus communis is widespread in Indonesia and Malaysia. It is recorded from the north of Sumatra (Aceh), the Malay Peninsula (Pahang), and northern Borneo (Sabah, Sarawak) as far south as Java and Bali. Though no records are known from southern parts of Sumatra and Borneo, the species is certainly present there.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE31403FF5EE71C05B85B01.taxon	etymology	Etymology: The epithet communis (adjective, Latin: common, frequent) refers to the widespread distribution of Frischianus communis in Southeast Asia compared to the presumably much smaller distribution areas of the other species of this genus.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE31403FF5EE71C05B85B01.taxon	discussion	Comparative note: Frischianus laticollis, F. similis, and F. communis cannot be identified based on their external appearance alone. The distinguishing characters of F. laticollis and F. similis are discussed above in the species chapter for F. similis. Males of F. communis can easily be distinguished according to the short, strongly ventrad curved apical lobe of the adeagus (Figs 76, 79, 82, 85, 88, 91, 94, 97), while the apical lobe is straight in F. laticollis and F. similis (Figs 67, 70, 73). In F. laticollis, the apical lobe moreover exhibits a ventral apical tooth (Fig. 67). While females of F. laticollis seem to differ from females of F. similis and F. communis according to the longer gonocoxal plate (Fig. 250), females of the latter two species, which are sympatric in Java, agree in the shape of the gonocoxal plate (Figs 251, 251) and cannot be distinguished.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE31405FCFCE0BC04D95B01.taxon	description	(Figs 100 – 102, 201, 253)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE31405FCFCE0BC04D95B01.taxon	materials_examined	Type specimens: Holotype ♂, Malaysia, Perak: Cascade Sungei Simei (Cameron Highlands), III. 1977, leg. Jaccoud (MHNG). Paratypes: 2 ♀, same data as holotype (MFNB, MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE31405FCFCE0BC04D95B01.taxon	description	Description: Habitus cf. Figs 2, 44, 45. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color medium brown with darker brown lateroposterior angles; antennae, palps, and legs light brown. Labral denticulation type 1 (Fig. 10). Forebody surface microsculpture type 2 (Fig. 2). Elytral rows of punctures present; sutural row of punctures impressed, extending over anterior half of elytral sutural length. Total body length 3.3 – 3.4 mm; forebody length 1.8 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with narrow triangular, medioposterior incision with straight lateral margins, occupying about 0.3 of sternite length (Fig. 201). Aedeagus (Figs 100 – 102) with apical lobe arising from aedeagus sinistrally; apical lobe narrower than half of distal width of aedeagus (Fig. 101), strongly curved ventrad, and slightly widened toward round end with proximad pointing tooth (Fig. 100). Position of endophallus in aedeagus unknown, because evaginated in male type specimen. Endophallic sclerite in lateral view elongate with strongly sclerotized apical portion convex dorsally (Fig. 100), in dorsal view with strongly sclerotized, laterally straight and apically pointed, sinistral section sharply set off against less sclerotized, laterally curved, dextral portion (Fig. 102). Dorsal endophallic spine in lateral view curved ventrad, slightly widened toward apex (Fig. 100), when invaginated probably not protruding from ostium (cf. Figs 70, 97), in dorsal view strongly curved sinistrad with longitudinally directed apical end (Figs 102). Dorsal, midlongitudinal split of phallobase type B (cf. Fig. 39). Aedeagus with ventral margin strongly concave between postforamen and apical lobe (Fig. 100), in dorsal view narrow and symmetrical with almost straight lateral margins feebly, gradually narrowed apicad (Fig. 102). Length of aedeagus 0.43 mm. Female: Gonocoxal plate about 1.7 times as long as wide, without basal and subbasal ridge, with subparallel, slightly concave lateral margins, shallowly emarginate posterior margin, and faintly biconvex anterior margin (Fig. 253).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE31405FCFCE0BC04D95B01.taxon	distribution	Distribution: Frischianus mariae is known only from the type locality Cascade Sungei Simei in the Cameron Highlands of Perak, Malaysia.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE31405FCFCE0BC04D95B01.taxon	etymology	Etymology: With choosing the epithet mariae (Latinized noun, derived from the first name Maria, genitive, singular) Johannes Frisch dedicates this new species to his dear wife Maria Frisch in gratitude for her love that survived all the worries that come with being in a relationship with a passionate entomologist.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE51405FF5EE0BC03075B41.taxon	description	(Figs 103 – 105, 202)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE51405FF5EE0BC03075B41.taxon	materials_examined	Type specimens: Malaysia: Holotype ♂, Sabah, Sandakan: S Lokan, III. 1997, leg. Chung (NHML). Paratypes: 1 ♂, 2 ♀, Ranau, Poring Hot Springs (6 ° 03.547 ' N, 116 ° 42.181 ' E), 9. VIII. 2009, leg. Floren (MFNB, NHMW).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE51405FF5EE0BC03075B41.taxon	description	Description: Habitus cf. Figs 2, 44, 45. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color unicolored reddish medium brown; tip of abdomen, antennae, palps, and legs yellow brown. Labral denticulation type 1 (Fig. 10). Forebody surface microsculpture type 2, but pronotal and elytral punctation somewhat coarser than in Figs 2 b, 2 c and elytral punctation stronger granular than in Fig. 2 c. Sutural elytral row of punctures impressed, occupying anterior half of elytral sutural length; medial row of punctures distinct, lateral row of punctures indistinct. Total body length 3.0 mm; forebody length 1.7 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with narrow triangular, medioposterior incision with straight lateral margins, occupying about 0.3 of sternite length (Fig. 202). Aedeagus (Figs 103 – 105) with apical lobe arising from sinistral half of distal width of aedeagus and being evenly curved dextrad (Figs 104, 105); apical lobe in lateral view short, wide and parallel, strongly curved ventrad, and evenly rounded apically toward proximally pointing, obtuse angle (Fig. 103). Endophallus in dorsal view inserted dextrally in aedeagus (Fig. 105). Endophallic sclerite short, in ventral view with two round, subapical, lateral sclerotizations (Fig. 104). Dorsal endophallic spine strong, quite far protruding from ostium, in lateral view evenly, weakly curved distoventrad with minutely serrate apical end (Fig. 103), in dorsal view from narrow, furcate basal end curved sinistrad toward sinistrally widened, longitudinally directed apical end (Fig. 105). Dextral half of ventral, ostial margin of aedeagus convex (Figs 104, 105). Dorsal, midlongitudinal split of phallobase type B (cf. Fig. 39). Aedeagus with ventral margin between postforamen and apical lobe slightly concave (Fig. 103), in dorsal view long oval, gradually narrowed apicad with dextral lateral margin more strongly curved than almost straight sinistral lateral margin (Fig. 105). Length of aedeagus 0.4 mm. Female: Gonocoxal plate as in F. communis (Fig. 252), about 1.6 times as long as wide.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE51405FF5EE0BC03075B41.taxon	distribution	Distribution: Frischianus brenneri was discovered in northern Borneo, where it was collected south of Lokan, Sabah Province, Malaysia.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE51405FF5EE0BC03075B41.taxon	etymology	Etymology: With the epithet brenneri (Latinized noun, derived from the surname Brenner, genitive, singular) Johannes Frisch honours his friend Uli Brenner, Schlüchtern-Breitenbach, Germany, coleopterist experienced in faunistics and ecology, in memory of joint collecting trips and the good times spent together.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE51407FCFCE07C07825B61.taxon	description	(Figs 106 – 108, 203, 254)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE51407FCFCE07C07825B61.taxon	materials_examined	Type specimens: Holotype ♂, Indonesia, Jambi, Kerinci: W Mt Tujuh Lake, 1400 m, 14. XI. 1989, leg. Agosti, Burckhardt & Löbl (MHNG). Paratypes: 3 ♀, same data as holotype (MFNB, MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE51407FCFCE07C07825B61.taxon	description	Description: Habitus cf. Figs 2, 44, 45. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color reddish medium brown with somewhat darker brown lateroposterior angles of elytra and abdomen; tip of abdomen, antennae, palps, and legs yellow brown. Labral denticulation type 1 (Fig. 10). Forebody surface punctation type 2, but pronotal punctation notably coarser than in Fig. 2 b. Sutural elytral row of punctures impressed, occupying anterior half of elytral sutural length; medial and lateral elytral row of punctures indistinct. Total body length 3.6 – 4.0 mm; forebody length 1.9 – 2.1 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with narrow triangular, medioposterior incision with slightly convex lateral margins, occupying about 0.3 of sternite length; anterior end of incision shortly narrowed and subparallel (Fig. 203). Aedeagus (Figs 106 – 108) with apical lobe arising subsinistrally from asymmetrically subtriangular, ventrodistal end of aedeagus (Figs 107, 108); apical lobe in lateral view elongate, subparallel, and hooked bent ventrad with apical end acute and somewhat curved proximad (Fig. 106). Endophallus in ventral and dorsal view situated submedially in aedeagus (Figs 107, 108). Endophallic sclerite large, in lateral view elongate with pointed proximal and round apical end (Fig. 106), in ventral view concave laterally, forcipate apically, and with tapered, diverging proximal ends (Fig. 107). Dorsal endophallic spine thin with feebly serrate apical end somewhat protruding from ostium, in lateral view straight with basal portion somewhat curved ventrad (Fig. 106), in dorsal view from notably widened, triangular, furcate basal end slightly curved sinistrad (Fig 108). Type of dorsal, midlongitudinal split of phallobase unclear. Aedeagus with ventral margin between postforamen and apical lobe strongly concave (Fig. 106), in dorsal view with very narrow, parallel distal half and slightly narrowed, somewhat dextrad bent proximal end (Figs 108). Length of aedeagus 0.51 mm. Female: Gonocoxal plate about 1.6 times as long as wide, without basal and subbasal ridge, with bisinuate anterior margin, subparallel, slightly concave lateral margins, and slightly emarginate posterior margin (Fig. 254).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE51407FCFCE07C07825B61.taxon	distribution	Distribution: Frischianus hamatus was found only at the type locality near Mount Kerinci in the Indonesian province of Jambi, Sumatra.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE51407FCFCE07C07825B61.taxon	etymology	Etymology: The epithet hamatus (adjective, Latin: hook-like, uncinate) refers to the hook-like apex of the aedeagus of this new species.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE71407FF5EE05C02A55861.taxon	description	(Figs 109 – 111, 204, 255)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE71407FF5EE05C02A55861.taxon	materials_examined	Type specimens: Holotype ♂, Indonesia, Sumatera Utara, Karo, 5 km W Brastagi: Tongkoh, 1450 m, 3. XII. 1989, leg. Agosti, Burckhardt & Löbl (MHNG). Paratypes: 1 ♂, 2 ♀, same data as holotype (MFNB, MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE71407FF5EE05C02A55861.taxon	description	Description: Habitus cf. Figs 2, 44, 45. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color unicolored reddish medium brown, or darker reddish brown with abdomen and lateroposterior angles of elytra somewhat darkened; tip of abdomen, antennae, palps, and legs light brown. Labral denticulation type 1 (Fig. 10). Forebody surface microsculpture type 2, but pronotal punctation compared to Fig. 2 b notably coarser and mediolongitudinally almost compacted. Sutural, elytral row of punctures impressed, occupying anterior half of elytral sutural length; medial and lateral row of punctures indistinct. Total body length 3.4 – 3.6 mm; forebody length 1.8 – 2.0 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with narrow triangular, medioposterior incision with lateral margins slightly convex, occupying about 0.3 of sternite length; anterior end of incision narrow, subparallel (Fig. 204). Aedeagus (Figs 109 – 111) with apical lobe arising from sinistral lateral margin of aedeagus; apical lobe in dorsal view strongly curved dextrad (Fig. 111), in lateral view elongate, evenly curved ventrad, and tapered toward acute, proximad curved apical end [Fig. 109 (outermost tip of apical lobe broken off)]. Endophallus in ventral and dorsal view situated medially in aedeagus (Figs 110, 111). Endophallic sclerite large, in lateral view triangular with right-angled tip pointing dorsad and seemingly inserted into right-angled curved, dorsal endophallic spine ventrally (Fig. 109), in ventral view subrectangular with strongly sclerotized, lateral thirds proximally connected by basally extended, biangulate, sclerotized band (Fig. 110). Dorsal endophallic spine strong, in lateral view at about midlength right-angled bent with ventroproximad pointing proximal portion and longer, narrow, straight distal portion with round, feebly serrate apical end far protruding from ostium (Fig. 109); dorsal endophallic spine in dorsal view sagittate, aligned midlongitudinally, from broadly furcate base evenly tapered toward acute apical end (Fig. 111). Ventral margin of ostium in dorsal view widely biconvex (Fig. 111). Type of dorsal, midlongitudinal split of phallobase unclear. Aedeagus with ventral margin distal of postforamen shortly concave, afterwards running straight toward apical lobe (Fig. 109), in dorsal view slightly, gradually widened apicad (Fig. 111). Length of aedeagus 0.46 mm. Female: Gonocoxal plate 1.6 times as long as wide, without basal and subbasal ridge, with bisinuate anterior margin, subparallel, slightly concave lateral margins, and emarginate posterior margin (Fig. 255).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE71407FF5EE05C02A55861.taxon	distribution	Distribution: Frischianus loebli is currently known only from the type locality Tongkoh in the Indonesian province of Sumatera Utara.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE71407FF5EE05C02A55861.taxon	etymology	Etymology: With choosing the epithet loebli (Latinized noun, derived from the surname Löbl, genitive, singular) Johannes Frisch dedicates this new species to his dear friend Ivan Löbl, specialist of Scaphidiinae at Muséum d’histoire naturelle, Geneva, Switzerland, in gratitude for his friendly support over the years and in appreciation of his important life’s work.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE71439FCFCE35C04B05CA1.taxon	description	(Figs 46, 112 – 114, 205, 256)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE71439FCFCE35C04B05CA1.taxon	materials_examined	Type specimens: Holotype ♂, Indonesia, Jambi, Kerinci: Mt Kerinci, 3000 m, 12. XI. 1989, leg. Agosti, Burckhardt & Löbl (MHNG). Paratypes: 2 ♀, same data as holotype (MFNB, MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE71439FCFCE35C04B05CA1.taxon	description	Description: Habitus (Fig. 46). Microphthalmous, micropterous without palisade fringe of abdominal tergite VII. Body color unicolored light, reddish medium brown; tip of abdomen, antennae, palps, and legs yellow brown. Labral denticulation type 2 (cf. Figs 11, 12). Forebody surface microsculpture type 2; compared to Figs 2 a, 2 b cephalic and pronotal punctation finer and more widely spaced, and elytra notably more coarsely punctate with somewhat uneven, almost rugulose surface. Eyes strongly reduced, 0.2 times as long as temples, in longitudinal diameter comprising 4 – 5 ommatidia. Temples notably widened posterior of eyes. Elytral sutural length 0.7 times as long as pronotal length. Elytra strongly narrowed across rounded humeri and with shallow, transverse depression in anterior half. Metathoracic wings strongly reduced in length and width to very narrow, elongate strip 0.6 times as long as elytral sutural length and 4.7 times longer than wide. Elytral sutural row of punctures not impressed, comprising only about three large punctures in anterior third of elytral sutural length; medial and lateral elytral row of punctures imperceptible in coarse elytral sculpturing. Total body length 4.3 – 4.9 mm; forebody length 2.1 – 2.3 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with very narrow, deep, medioposterior incision with lateral margins convex, occupying about 0.4 of sternite length (Fig. 205). Aedeagus (Figs 112 – 114) with apical lobe situated dextrally at strongly asymmetrical, subtriangular distal end of aedeagus (Figs 113, 114); apical lobe in lateral view elongate, strongly curved ventrad, ventrally extended in strong, proximad pointing tooth, and with short, blunt, somewhat distad curved tip (Fig. 112). Endophallus in ventral and dorsal view situated submedially in aedeagus (Figs 113, 114). Endophallic sclerite relatively small, elongate, in lateral view dentiform with sharp apical end (Fig. 112), in ventral view about half as wide as aedeagus, oval, with convex lateral margins evenly narrowed toward round apical end (Fig. 113). Dorsal endophallic spine slender with narrow base, feebly curved dextrad, slightly narrowed medially and with acute end (Figs 112 – 114), in dorsal view strongly protruding from ostium (Fig. 114), but only slightly projecting dorsally in lateral view (Fig. 112). Dorsal, midlongitudinal split of phallobase type B (cf. Fig. 39). Aedeagus with ventral margin between postforamen and apical lobe feebly inverted S-shaped (Fig. 112), in dorsal view long oval with subparallel lateral margins and slightly curved dextrad at midlength (Fig. 114). Length of aedeagus 0.69 mm. Female: Gonocoxal plate about 1.4 times as long as wide (measured at greatest width across feebly concave, transverse basal ridge excluding acute, lateroanterior ends), without subbasal ridge, with subparallel, concave lateral margins, and almost semicircularly emarginate posterior margin (Fig. 256).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE71439FCFCE35C04B05CA1.taxon	distribution	Distribution: Frischianus subalpinus is known only from the type locality in the subalpine zone of Mt Kerinci, Indonesia’s highest mountain on Sumatra, and in view of its flightlessness probably endemic there.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFE71439FCFCE35C04B05CA1.taxon	etymology	Etymology: The epithet subalpinus (adjective, Latin: subalpine) refers to the subalpine altitude where Frischianus subalpinus was found on Mount Kerinci.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD91439FF5EE61C05CA5FE1.taxon	description	(Figs 47, 257)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD91439FF5EE61C05CA5FE1.taxon	materials_examined	Type specimens: Holotype ♀, Indonesia, Jambi, Kerinci: Mt Kerinci, 1900 m, 13. XI. 1989, leg. Agosti, Burckhardt & Löbl (MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD91439FF5EE61C05CA5FE1.taxon	description	Description: Habitus (Fig. 47). Similar to Frischianus subalpinus from which it differs as follows: Body color dark medium brown with posterior half of elytra blackish brown; tip of abdomen, antennae, palps, and legs light brown. Head and pronotum with denser, coarser punctation; elytral surface not rugulose with finer punctation notably coarser than punctation of head and pronotum. Eyes somewhat larger, 0.4 times as long as temples; longitudinal diameter comprising 6 – 7 ommatidia. Temples parallel, not widened posterior of eyes. Elytra slightly longer, sutural length 0.7 times as long as pronotal length, less narrowed across rounded humeri, and without clear depression in anterior half. Somewhat smaller; total body length 4.0 mm, forebody length 2.0 mm. Male unknown. Female: Gonocoxal plate slender, about 2.1 times as long as wide, with roundly subtriangularly extended basal margin without basal and subbasal ridge, subparallel, slightly concave lateral margins, and deeply, narrowly emarginate posterior margin (Fig. 257).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD91439FF5EE61C05CA5FE1.taxon	distribution	Distribution: Frischianus curtipennis is native to the montane zone of Mt Kerinci, Indonesia’s highest mountain on Sumatra, where it is probably endemic due to its flightlessness.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD91439FF5EE61C05CA5FE1.taxon	etymology	Etymology: The epithet curtipennis [adjective, Latin, composed of the adjective curtus (short) and the noun penna (wing)] refers to the reduced elytra and metathoracic wings of this new species.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD91439FF5EE61C05CA5FE1.taxon	discussion	Remarks: The metathoracic wings were not examined to avoid damage of the single type specimen. They are expected to be reduced judging by the absence of the palisade fringe of abdominal tergite VII. Frischianus curtipennis is treated in the chapter of the F. laticollis lineage of the F. laticollis species group for purely practical reasons due to its similarity with F. subalpinus, to which it is compared. The long gonocoxal plate with a deep, narrow posterior emargination (Fig. 257) is more suggestive of the F. rufulus subgroup of the F. laticollis species group (cf. Figs 263 – 265). Frischianus curtipennis is not assigned to the F. laticollis lineage, but a species incertae sedis as long as male characters are unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD91438FCFCE4DC06FD5641.taxon	description	(Figs 48, 115 – 117, 206, 207, 258)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD91438FCFCE4DC06FD5641.taxon	materials_examined	Type specimens: Holotype ♂, Indonesia, Sumatera Barat, Bukittinggi: Mt Singgalang, 2100 – 2600 m, 16. X. 1990, leg. Riedel (SMNS). Paratypes: 2 ♂, 1 ♀, same data as holotype (MFNB, NHMW, SMNS).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD91438FCFCE4DC06FD5641.taxon	description	Description: Habitus (Fig. 48). Microphthalmous, micropterous without palisade fringe of abdominal tergite VII. Body color unicolored dark brown except for medium brown tip of abdomen; antennae, palps, and legs light brown to medium brown. Labral denticulation type 2 (Figs 11, 12). Forebody surface microsculpture type 2 (Fig. 2), but cephalic punctation finer and elytral punctation coarser. Eyes strongly reduced, 0.2 times as long as temples, in longitudinal diameter comprising 4 – 5 ommatidia. Temples notably widened posterior of eyes. Elytral sutural length 0.6 times as long as pronotal length. Elytra strongly narrowed across rounded humeri and with shallow, transverse depression in anterior half. Metathoracic wings strongly reduced in length, about as long as humeral length of elytra. Sutural elytral row of punctures short, impressed, occupying anterior third of elytral sutural length; medial and lateral elytral row of punctures imperceptible in coarse elytral sculpturing. Total body length 3.6 – 4.0 mm; forebody length 2.0 – 2.1 mm. Male: Abdominal sternite VII with longitudinal depression in posterior two-thirds of median third of sternite width covered with widely spaced, laterally more densely arranged, black macrosetae (Fig. 206). Abdominal sternite VIII with narrow, triangular, medioposterior incision with concave lateral margins occupying about 0.3 of sternite length (Fig. 207). Aedeagus (Figs 115 – 117) with long, slender apical lobe situated medially at slightly asymmetrical, subtriangular, distal end of aedeagus (Figs 116, 117); apical lobe evenly curved ventrad, with subapical, ventral tooth and minute, sharp, dorsad bent, apical tip (Fig. 115). Ostium considerably retracted laterally, uncovering distal half of dorsal endophallic spine and distal fourth of endophallic sclerite (Figs 115, 117). Endophallus in ventral and dorsal view situated medially in aedeagus (Figs 116, 117). Endophallic sclerite large, in lateral view elongate (Fig. 115), in ventral view widely oval, almost occupying aedeagal width, and with convex lateral margins evenly tapered toward round apical end (Fig. 116). Dorsal endophallic spine long, thin and straight, with short, broad, basal dilation (Figs 115, 117). Dorsal, midlongitudinal split of phallobase type A (cf. Figs 36 – 38). Aedeagus with ventral margin between postforamen and apical lobe weakly concave (Fig. 115), in dorsal view long oval with dextral lateral margin slightly curved and sinistral lateral margin straight, and distally of dorsal margin of ostium gradually, convexly tapered toward apical lobe (Fig. 117). Length of aedeagus 0.66 mm. Female: Gonocoxal plate about 1.4 times as long as wide, widest across medially divided basal ridge with posteriorly curved median ends, without subbasal ridge, somewhat concave laterally and somewhat narrowed toward emarginate posterior margin (Fig. 258).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD91438FCFCE4DC06FD5641.taxon	distribution	Distribution: Frischianus pedator was discovered at high altitude on Mount Singgalang, Sumatera Barat, and is expected to be endemic to this volcano judging from its flightlessness.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD91438FCFCE4DC06FD5641.taxon	etymology	Etymology: The epithet pedator (noun, Latin, nominative: pedestrian) refers to the fact that this new species can only travel by walking due to its inability to fly.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD8143AFCFCE0FC07615821.taxon	description	(Figs 49, 118 – 120, 208)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD8143AFCFCE0FC07615821.taxon	materials_examined	Type specimen: Holotype ♂, Malaysia, Sabah (4 ° 63 ' 50 " – 4 ° 77 ' 16 " N, 117 ° 43 ' 83 – 117 ° 70 ' 31 " E), 300 m, XI. – XII. 2011, leg. Ewers et al. (SAFE Project) (NHML).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD8143AFCFCE0FC07615821.taxon	description	Description: Habitus (Fig. 49). Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color reddish medium brown with head and elytra except for humera darker brown; tip of abdomen, antennae, palps, and legs light brown. Labral denticulation type 2 (cf. Figs 11, 12). Forebody surface microsculpture type 1 (Fig. 1); elytral punctation much denser compared to Fig. 1 c. Sutural elytral row of punctures not much impressed, extending over anterior 0.7 of elytral sutural length; medial and lateral row of punctures indistinct, reduced to few, widely spaced punctures. Total body length 3.1 mm; forebody length 1.7 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with narrow triangular, medioposterior incision with slightly concave lateral margins, occupying about 0.3 of sternite length (Fig. 208). Aedeagus (Figs 118 – 120) with apical lobe situated in sinistral nine-tenth of distal width of aedeagus (Figs 119, 120); apical lobe long, occupying almost half of aedeagal length, in lateral view directed distoventrad, parallel in proximal half, somewhat curved ventrad at about midlength, and afterwards narrowed toward shortly distad curved tip (Fig. 118); in dorsal view, apical lobe broad with short, obtuse, dextrobasal dilatation, faintly concave dextral margin, convex sinistral margin, and gradually tapered toward narrow, acute, somewhat dextrad bent apical end (Fig. 120). Endophallus in dorsal view with base inserted sinistrally in aedeagus (Fig. 120). Endophallic sclerite from wide base sinistrally extended in long, narrow spine almost reaching apical end of dorsal endophallic spine and far protruding from ostium, in dorsal view with sinistral curvature running sinistrally of dorsal endophallic spine (Fig. 120). Dorsal endophallic spine long and slender, reaching about half of length of apical lobe, slightly curved ventrad, and considerably protruding from ostium (Figs 118, 120), with broad (Fig. 120), ventrad curved (Fig. 118) basal end and narrow, somewhat dorsosinistrad curved apical end (Fig. 118) with short, longitudinally directed tip (Fig. 120). Dorsal, midlongitudinal split of phallobase type A (cf. Figs 36 – 38). Aedeagus with ventral margin between postforamen and apical lobe weakly convex (Fig. 118), in dorsal view oval with lateral margins weakly, equally convex (Fig. 120). Length of aedeagus 0.49 mm. Female unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD8143AFCFCE0FC07615821.taxon	distribution	Distribution: Frischianus uhligi is native to northeastern Borneo. The exact type locality in the Malaysian province of Sabah is unknown, because the holotype label lacks a locality name, and the given coordinates comprise a large area.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD8143AFCFCE0FC07615821.taxon	etymology	Etymology: With naming this new species uhligi (Latinized noun, derived from the surname Uhlig, genitive, singular) Johannes Frisch thankfully honours his dear colleague Manfred Uhlig, specialist of Erichsonius at the Museum für Naturkunde Berlin, to whom he owes a lot in his life.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDA143DFF5EE39C04055F01.taxon	description	(Figs 50, 121 – 123, 209, 259)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDA143DFF5EE39C04055F01.taxon	materials_examined	Type specimens: Malaysia, Sabah: Holotype ♂, Ranau: Poring Hot Springs, 500 m, 8. V. 1987, leg. Burckhardt & Löbl (MHNG). Paratypes (6 specimens): 2 ♂, 1 ♀, same data as holotype (MFNB, MHNG); 2 ♀, same data as holotype, but 7. V. 1987 and 11. V. 1987 (MHNG); 1 ♂, Ranau, Poring Hot Springs: Langanan Falls, 900 – 950 m, 12. V. 1987, leg. Burckhardt & Löbl (MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDA143DFF5EE39C04055F01.taxon	description	Description: Habitus (Fig. 50). Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color unicolored light reddish brown to medium brown; tip of abdomen, antennae, palps, and legs yellow brown. Labral denticulation types 1 or 2 (Figs 10 – 12); submedial denticles more or less pointed, longer than or about as long as pointed or obtuse, sublateral denticles. Forebody surface microsculpture type 2 (Fig. 2), but pronotal and elytral punctation coarser than in Figs 2 b, 2 c. Elytral rows of punctures occupying about anterior 0.7 of elytral sutural length; sutural row of punctures impressed anteriorly; medial and lateral row of punctures often reduced to few, widely spaced punctures. Total body length 3.1 – 3.4 mm; forebody length 1.6 – 1.8 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with narrow triangular, medioposterior incision with slightly concave lateral margins, occupying roughly 0.3 of sternite length (Fig. 209). Aedeagus (Figs 121 – 123) with apical lobe situated in about sinistral half of distal aedeagal width (Figs 122, 123); apical lobe long and slender, at base strongly and obtuse-angled bent distoventrad, approximately at midlength with slight, dorsally directed curvature, after that gradually tapered toward acute, slightly distad curved apical end (Fig. 121), in ventral view close to base somewhatcurvedsinistrally, afterthatslightlyundulateand gradually tapered toward thin, acute apical end (Fig. 123). Endophallus in ventral and dorsal view with base inserted submedially in aedeagus (Figs 122, 123). Endophallic sclerite in lateral view slender, notably projecting from ostium but significantly shorter than dorsal endophallic spine, curved distoventrad, and gradually tapered toward subacute end (Fig. 121); endophallic sclerite in dorsal view broadly suboval with almost straight dextral margin and convex sinistral margin, gradually tapered toward acute apical end, and clearly protruding dextrally over dorsal endophallic spine and apical lobe (Fig. 123). Dorsal endophallic spine long and slender, reaching about half of length of apical lobe, straight in proximal two-thirds situated within aedeagus, but in distal third considerably projecting from ostium, strongly curved distoventrad in line with apical lobe, and with shortly dorsad curved apical end (Fig. 121); dorsal endophallic spine in dorsal view straight, aligned longitudinally, with moderately dilated base and acute, sinistrad curved apical end (Fig. 123). Dorsal, midlongitudinal split of phallobase type A (cf. Figs 36 – 38). Aedeagus with ventral margin between postforamen and apical lobe slightly convex (Fig. 121), in dorsal view narrow with straight lateral margins and somewhat tapered proximad (phallobase collapsed?) (Fig. 123). Length of aedeagus 0.46 – 0.5 mm. Female: Gonocoxal plate long, about 2.5 times as long as wide, triangular anteriorly; basal ridge angulate in middle and medially curved at ends; subbasal ridge triangular; lateral margins subparallel; medioposterior emargination short, truncate, and laterally delimited by subacute ends (Fig. 259).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDA143DFF5EE39C04055F01.taxon	distribution	Distribution: Frischianus burckhardti occurs in the Malaysian province of Sabah, northeastern Borneo, where it was discovered at Poring Hot Springs east of Mount Kinabalu.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDA143DFF5EE39C04055F01.taxon	etymology	Etymology: With choosing the epithet burckhardti (Latinized noun, derived from the surname Burckhardt, genitive, singular) Johannes Frisch thanks Daniel Burckhardt, phylogeographer and specialist of Psylloidea at Naturhistorisches Museum Basel, Switzerland, who co-collected the type specimens, for friendly advice at an early stage of his work on the Scopaeina.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDD143DFF5EE4BC030E5FE2.taxon	description	(Figs 124 – 126, 210)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDD143DFF5EE4BC030E5FE2.taxon	materials_examined	Type specimen: Holotype ♂, Malaysia, Sabah, Sandakan: Sepilok, X. 1996, leg. Chung (NHML).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDD143DFF5EE4BC030E5FE2.taxon	description	Description: Habitus (Fig. 50). Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color unicolored light reddish brown; tip of abdomen, antennae, palps, and legs yellow brown. Labral denticulation type 4 (cf. Fig. 15), but in comparison with closely related species probably non-species-specific, individual variation. Forebody surface microsculpture type 1 (Fig. 1). Elytral rows of punctures long and distinct, extending over anterior 0.7 of elytral sutural length; sutural row of punctures not much impressed. Total body length 3.1 mm; forebody length 1.9 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with narrow triangular, medioposterior incision with slightly concave lateral margins, occupying about 0.3 of sternite length (Fig. 210). Aedeagus (Figs 124 – 126) with apical lobe situated in sinistral six-seventh of distal aedeagal width (Figs 125, 126); apical lobe long, occupying half of aedeagal length, in lateral view slender, somewhat curved ventrad, in about apical fourth evenly tapered toward distad curved, acute end (Fig. 124), in dorsal view broad, evenly curved dextrad, and gradually tapered toward abruptly narrowed, slim, acute apical end (Fig. 126). Endophallus in dorsal view with base inserted subsinistrally in aedeagus, with endophallic sclerite and dorsal endophallic spine considerably projecting from ostium (Fig. 126). Endophallic sclerite extended in long, narrow spine almost reaching apical end of dorsal endophallic spine, in lateral view straight and gradually narrowed toward acute apical end (Fig. 124); endophallic sclerite in dorsal view from sinistral, weakly sclerotized base aligned dextrodistad, running dextrally of dorsal endophallic spine, and with tip somewhat curved sinistrad (Fig. 126). Dorsal endophallic spine with ventrad curved, furcate base (Figs 124, 126), in distal half densely studded with laterodistad directed spinules / denticles (echinate, spinulate) except for short, acute, inechinate apical end, in lateral view thin and straight (Fig. 124), in dorsal view feebly inverted Sshaped with echinate portion somewhat widened (Fig. 125). Dorsal, midlongitudinal split of phallobase type A (cf. Figs 36 – 38). Aedeagus with ventral margin between postforamen and apical lobe straight (Fig. 124), in dorsal view long oval, subparallel with lateral margins weakly, equally curved (Fig. 126). Length of aedeagus 0.43 mm. Female unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDD143DFF5EE4BC030E5FE2.taxon	distribution	Distribution: Frischianus echinatospinatus is known only from northern Borneo, where it was discovered near Sandakan in the northeast of Sabah, Malaysia.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDD143DFF5EE4BC030E5FE2.taxon	etymology	Etymology: The epithet echinatospinatus [adjective, Latin, composed of the adjectives echinatus (spiny, thorny, prickly, spinulate) and spinatus (indicating a feature of the species related to the spine)] refers to the echinate dorsal endophallic spine of the aedeagus (Figs 124 – 126).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDD143FFCFCE4DC070B5DC2.taxon	description	(Figs 51, 127 – 129, 211, 212, 260)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDD143FFCFCE4DC070B5DC2.taxon	materials_examined	Type specimens: Holotype ♂, Malaysia, Sabah, 60 km road Tambunan – Mt Kinabalu (Crocker Range), 1270 m, 17. V. 1987, leg. Burckhardt & Löbl (MHNG). Paratype: 1 ♀, same data as holotype (MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDD143FFCFCE4DC070B5DC2.taxon	description	Description: Habitus (Fig. 51). Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color unicolored reddish medium brown to dark brown; tip of abdomen, antennae, palps, and legs yellow brown. Labral denticulation type 1 (Fig. 10). Forebody surface microsculpture type 2 (Fig. 2), but pronotal and elytral punctation coarser than in Figs 2 b, 2 c. Sutural elytral row of punctures impressed anteriorly, occupying approximately anterior 0.7 of elytral sutural length; medial and lateral elytral row of punctures reduced to few, widely spaced punctures. Total body length 3.1 mm; forebody length 1.6 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with short, triangular, medioposterior incision with slightly concave lateral margins, occupying roughly 0.2 of sternite length (Fig. 211). Aedeagus (Figs 127 – 129) with apical lobe situated in about sinistral half of distal aedeagal width (Figs 128, 129); apical lobe long, in lateral view from wide base curved ventrad, and gradually tapered toward distad curved, acute apical end (Fig. 127), in ventral view from wide base gradually tapered toward thin distal half with slightly sinistrad bent apical end (Fig. 128). Endophallus in ventral and dorsal view with base inserted medially in aedeagus; dorsal endophallic spine and endophallic sclerite equally long and considerably projecting from ostium (Figs 128, 129). Endophallic sclerite distally extended in long, evenly ventrad curved, spiniform process with two large, subapical, ventrodextral teeth (Fig. 127); endophallic sclerite in dorsal view wide basally, slightly aligned dextrad, with convex sinistral margin and slightly concave, bidentate dextral margin (Fig. 128), somewhat protruding over dextral margin of apical lobe, and gradually tapered toward short, narrow, longitudinad curved apical end (Fig. 129). Dorsal endophallic spine long, evenly curved and gradually tapered ventrad, and extending beyond midlength of apical lobe; ventral margin of dorsal endophallic spine strongly serrate in distal half except apically (Fig. 127); dorsal endophallic spine in dorsal view from broad, excised proximal end somewhat directed distodextrad, and gradually tapered toward sinistrad curved end strongly protruding apical lobe dextrally (Fig. 129). Dorsal, midlongitudinal split of phallobase type A (cf. Figs 36 – 38). Aedeagus with ventral margin between postforamen and apical lobe straight (Fig. 127), in dorsal view long oval, with almost straight lateral margins gradually tapered proximad (phallobase collapsed?) (Fig. 129). Length of aedeagus 0.46 mm. Female: Abdominal sternite VIII with very short, concave, posterior emargination (Fig. 212). Gonocoxal plate long, about 2.4 times as long as wide, in anterior half gradually, subtriangularly narrowed with truncate anterior end; basal and subbasal ridge triangular; lateral margins shortly concave in midlength of sternite, but somewhat convex in posterior half of sternite length; posterior margin with short emargination delimited laterally by subacute ends (Fig. 260).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDD143FFCFCE4DC070B5DC2.taxon	distribution	Distribution: Frischianus puthzi is known only from the type locality in the Crocker Range in northeastern Borneo, Sabah, Malaysia.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDD143FFCFCE4DC070B5DC2.taxon	etymology	Etymology: Johannes Frisch dedicates this new species to Volker Puthz, Schlitz, Germany, to honour his great life’s work on the taxonomy of the Steninae and Euaesthetinae, and to thank Volker for introducing him to the Staphylinidae back when he was a teenager (epithet puthzi: Latinized noun, derived from the surname Puthz, genitive, singular).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDF143FFCFCE39D021C5641.taxon	diagnosis	Diagnosis: Separated from remaining lineages of F. laticollis subgroup (chapter 3.2.2.1.) by straight, not ventrad curved apical lobe of aedeagus (Figs 136 – 138) and enlarged endophallic sclerite deeply divided in two lateral halves parallel to each other (Figs 136 – 138). Different from F. coriaceus and F. laticollis lineages furthermore by absence of dorsal endophallic spine.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDF143FFCFCE39D021C5641.taxon	discussion	Remark: The Frischianus schuriani lineage is named after F. schuriani sp. nov. from the Malay Peninsula, the only representative so far.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDF143FFF5EE6FC03135821.taxon	description	(Figs 130 – 132, 213)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDF143FFF5EE6FC03135821.taxon	materials_examined	Type specimen: Holotype ♂, Malaysia, Sabah, Lahad Datu: Ulu Segama (Danum Valley Forest Reserve), (04 ° 57 ' N, 117 ° 48 ' E), 200 m, 3. – 5. XI. 2005, leg. Mann, Slade & Villanueva (HECO).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDF143FFF5EE6FC03135821.taxon	description	Description: Habitus cf. Fig. 51. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color reddish medium brown with head and posterior half of elytra slightly darker brown; tip of abdomen, antennae, palps, andlegsyellowbrown. Labraldenticulation type 1 (Fig. 10). Forebody surface microsculpture type 2 (Fig. 2), but pronotal and elytral punctation somewhat coarser than in Figs 2 b, 2 c. Sutural elytral row of punctures impressed anteriorly, extending over anterior 0.7 of elytral sutural length; medial and lateral elytral row of punctures reduced to few, widely spaced punctures. Total body length 3.1 mm; forebody length 1.6 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with narrow triangular, medioposterior incision with slightly concave lateral margins, occupying roughly 0.3 of sternite length (Fig. 213). Aedeagus (Figs 130 – 132) with apical lobe strongly narrowed basally to about sinistral fourth of distal aedeagal width (Fig. 131); apical lobe long and slender, occupying almost half of aedeagal length, in lateral view evenly curved ventrad and gradually tapered toward distad curved tip (Fig. 130); in ventral view, apical lobe narrow, projecting sinistrally over distal process of endophallic sclerite, in distal half somewhat dextracurved and gradually tapered toward thin apical end with tiny, subapical, dextrolateral tooth (Fig. 131). Endophallus in ventral and dorsal view with base inserted medially in aedeagus (Figs 131, 132); endophallic sclerite and dorsal endophallic spine strongly lengthened and considerably projecting from ostium (Figs 130 – 132). Endophallic sclerite with distal process slightly curved ventrad (Fig. 130); distal process of endophallic sclerite, in dorsal view, long subrectangular, in proximal portion somewhat dextracurved, then narrowed toward sinistracurved distal portion bearing distolaterad pointing sinistroapical tooth, distad pointing dextroapical tooth, and ventrad pointing subdextroapical tooth (Figs 130 – 132). Dorsal endophallic spine long, evenly curved ventrad, with strongly serrate ventral margin and more finely serrate dorsal margin gradually tapered toward very slender, acute, more strongly ventrad curved apical end (Fig. 130), in dorsal view evenly, moderately curved sinistrad (Fig. 132). Dorsal, midlongitudinal split of phallobase type A (cf. Figs 36 – 38). Aedeagus with ventral margin between postforamen and apical lobe straight (Fig. 130), in dorsal view long oval with straight lateral margins (Fig. 132). Length of aedeagus 0.53 mm. Female unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDF143FFF5EE6FC03135821.taxon	distribution	Distribution: Frischianus serratispinatus is native to northeastern Borneo and was collected in the Danum Valley in eastern Sabah, Malaysia.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDF143FFF5EE6FC03135821.taxon	etymology	Etymology: The epithet serratispinatus [adjective, Latin, composed of the adjectives serratus (serrate) and spinatus (indicating a feature of the species related to the spine)] refers to the serrate dorsal endophallic spine of the aedeagus (Fig. 130).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFDF143FFF5EE6FC03135821.taxon	discussion	Remark: A ♂ and a ♀ of Frischianus from the Tawau Hills [Malaysia, Sabah, Tawau (4 ° 24.009 ' N, 117 ° 53.412 ' E), 5. IX. 2009, leg. Floren (NHMW)], which is situated only about 60 km south of the type locality of S. serratispinatus, might be conspecific according to the basically comparable structure of the aedeagus (Figs 133 – 135). Nevertheless, the aedeagus of the ♂ from Tawau differs by the straight, not dextracurved distal portion of the apical lobe, the almost edentate dorsal endophallic spine, which is only weakly dentate basally, and the distolaterad pointing dextroapical tooth of the endophallic sclerite. More material is necessary to answer the question of whether the specimens from the Tawau Hills and the Danum Valley Forest Research represent different species or just intraspecific variations.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD11431FF5EE5BC023F5D02.taxon	description	(Figs 136 – 138, 214)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD11431FF5EE5BC023F5D02.taxon	materials_examined	Type specimen: Holotype ♂, Malaysia, Pahang, 4 mls NE Cameron Highlands, 23. – 25. IV. 1977, leg. Watrous (FMNH).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD11431FF5EE5BC023F5D02.taxon	description	Description: Habitus cf. Fig. 50. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color reddish medium brown with slightly darker brown head; tip of abdomen, antennae, palps, and legs light brown. Labral denticulation type 2 (cf. Figs 11, 12). Forebody surface microsculpture type 1 (Fig. 1). Sutural elytral row of punctures weakly impressed, in about anterior fourth of elytral sutural length densely punctate, afterwards with widely spaced punctures continued posteriorly; medial and lateral elytral row of punctures indistinct, comprising few, widely spaced punctures. Total body length 3.9 mm; forebody length 2.1 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with deep, narrow, medioposterior incision with convex lateral margins occupying about 0.4 of sternite length (Fig. 214). Aedeagus (Figs 136 – 138) in lateral view with slender, straight apical lobe at base somewhat bent ventrad (Fig. 136); apical lobe in ventral view occupying median third of aedeagal apical width, lobiform and subparallel laterally with small, dextral indentation and round apical end (Fig. 137). Endophallic sclerite considerably protruding from wide ostium (Figs 136, 138), deeply divided into two lateral halves each remarkably enlarged dorsally in proximal portion, then abruptly curved ventrad and ending in distal process with short dorsal and longer ventral apical tooth (Fig. 136); base of endophallic sclerite in sinistral position within aedeagus and, both lateral halves taken together, gradually widened toward broad, subparallel median portion abruptly narrowed toward slender, gradually tapered, pointed distal portion (Figs 137, 138). Dorsal, midlongitudinal split of phallobase type B (Fig. 39). Aedeagus with ventral margin between postforamen and apical lobe moderately inverted Sshaped (Fig. 136), in dorsal view oval with dextral lateral margin more strongly curved than sinistral lateral margin (Fig. 138). Length of aedeagus 0.43 mm. Female unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD11431FF5EE5BC023F5D02.taxon	distribution	Distribution: Frischianus schuriani is native to the Cameron Highlands, Malay Peninsula.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD11431FF5EE5BC023F5D02.taxon	etymology	Etymology: With the dedication of this new species, Johannes Frisch thanks his dear colleague Bernhard Schurian, photographer at the Museum für Naturkunde Berlin, for the perfect high-resolution habitus images published in this article (Figs 1 – 9, 47, 57) (epithet schuriani: Latinized noun, derived from the surname Schurian, genitive, singular).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD11430FCFCE6BD04675CE1.taxon	diagnosis	Diagnosis: Separated from remaining lineages of F. laticollis subgroup (chapter 3.2.2.1.) by elongate, nonspiniform, non-lobiform, moderately dilated endophallic sclerite with round apical end and usually well visible, subapical endophallic foramen (Figs 139 – 156). Different from F. coriaceus and F. laticollis lineages also by absence of dorsal endophallic spine. Separated from F. schuriani lineage by ventrad curved apical lobe of aedeagus and different shape of endophallic sclerite (cf. Figs 136 – 138). Separated from F. coriaceus, F. laticollis, and F. schuriani lineages also by dorsal, midlongitudinal split of phallobase short oval and widely separate from ostium (type C; Fig. 40).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD11430FCFCE6BD04675CE1.taxon	discussion	Remarks: The Frischianus setifer lineage of the F. laticollis subgroup, named after F. setifer, whose characters represent this phylogenetic clade well, currently comprises six species, all of which are new to science. They are described below. The Frischianus setifer lineage is recorded from the Malay Peninsula, northern Borneo (Sabah, Sarawak), Java, and Sulawesi.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01430FCFCE7DC02B05941.taxon	description	(Figs 142 – 144, 216)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01430FCFCE7DC02B05941.taxon	materials_examined	Type specimen: Holotype ♂, Malaysia, Pahang, 19 mls NE Kuala Lumpur, 28. IV. 1977, leg. Watrous (FMNH).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01430FCFCE7DC02B05941.taxon	description	Description: Similar to Frischianus latilobatus (cf. Fig. 50), but labral denticulation type 1 (Fig. 10). Elytra moderately shiny with fine and dense, somewhat granular punctation with interspaces smaller than puncture diameters. Sutural elytral row of punctures faintly impressed anteriorly, extending over anterior 0.7 of elytral sutural length; medial and lateral row of punctures indistinct. Smaller than F. latilobatus: total body length 3.0 mm, forebody length 1.7 mm, length of head and pronotum 1.1 mm. Male: Aedeagus (Figs 142 – 144) similar to that of F. latilobatus, but different as follows: Apical lobe, in lateral view, narrower with acute apical end; ventral tooth less pointed and setulose proximoventral margin straight (Fig. 142). Endophallic sclerite with apical end not uncinate (Fig. 142), in ventral and dorsal view narrower medially and with base inserted medially in aedeagus (Figs 143, 144); endophallic foramen without straight proximal margin (Fig. 144). Aedeagus with ventral margin between postforamen and base of apical lobe less concave (Fig. 142). Length of aedeagus 0.4 mm. Female unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01430FCFCE7DC02B05941.taxon	distribution	Distribution: Frischianus latilobatoides is native to Pahang in the Malay Peninsula.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01430FCFCE7DC02B05941.taxon	etymology	Etymology: The epithet latilobatoides [adjective, Latin, composed of the adjectives latus (broad), lobatus (indicating a characteristic / feature of the lobe) and the Greek, Latinized suffix “ ides ” (referring to a similarity)] refers to the resemblance and close relationship of this new species to Frischianus latilobatus.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01430FF5EE7DC02975CE1.taxon	description	(Figs 139 – 141, 215)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01430FF5EE7DC02975CE1.taxon	materials_examined	Type specimen: Holotype ♂, Malaysia, Sabah (4 ° 63 ' 50 " – 4 ° 77 ' 16 " N, 117 ° 43 ' 83 " – 117 ° 70 ' 31 " E), 300 m, XI. – XII. 2011, leg. Ewers et al. (NHML).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01430FF5EE7DC02975CE1.taxon	description	Description: Habitus cf. Fig. 50. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color medium brown; tip of abdomen, antennae, palps, and legs light brown. Labral denticulation type 2 (Figs 11, 12). Forebody surface microsculpture type 1 (Fig. 1); elytra of holotype lost. Total body length 3.4 mm; length of head and pronotum (elytra missing) 1.3 mm. Male: Abdominal sternite VII without diagnostic characters. Abdominal sternite VIII with narrow triangular, medioposterior incision with slightly concave lateral margins, occupying about 0.25 of sternite length (Fig. 215). Aedeagus (Figs 139 – 141) with large apical lobe arising from sinistral half of distal aedeagal width (Figs 140, 141); apical lobe in lateral view as broad as aedeagus distally, evenly curved ventrad with more strongly ventrad curved, right-angled apical end; ventral margin of apical lobe concave between convex proximal portion studded with few setulae and widely triangular distal portion with small, acute, median tooth (Fig. 139); apical lobe in ventral view from dextrally situated proximal end in moderate, sinistral curve running toward narrow, shortly dextracurved, rounded apical end (Fig. 140). Endophallic sclerite, even though strongly projecting from ostium in dorsal view (Fig. 141), not protruding over aedeagus dorsally (Fig. 139); endophallic sclerite in lateral view moderately curved ventrad, from narrow base gradually thickened toward midlength, after that gradually tapered toward shortly ventrad bent, uncinate apical end (Fig. 139), in dorsal view wide and subparallel, from sinistrally inserted base directed distodextrad, and in distal fifth evenly tapered toward triangular, subacute tip slightly surpassing aedeagus dextrally (Fig. 141); apical portion of endophallic sclerite with clear, wide endophallic foramen with straight proximal margin (Fig. 141). Aedeagus with ventral margin between postforamen and base of apical lobe deeply concave (Fig. 139), in dorsal view long oval and gradually tapered distad with dextral lateral margin slightly more strongly curved than sinistral lateral margin (Fig. 141). Length of aedeagus 0.46 mm. Female unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01430FF5EE7DC02975CE1.taxon	distribution	Distribution: Frischianus latilobatus was collected in Sabah, Northeast Borneo, where it was discovered north of Tawau. The exact type locality is unknown, because the coordinates of the location label of the holotype comprise a large area.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01430FF5EE7DC02975CE1.taxon	etymology	Etymology: The epithet latilobatus [adjective, Latin, composed of the adjectives latus (broad) and lobatus (indicating a characteristic / feature of the lobe)] refers to the apical lobe of the aedeagus of this new species, which is broad in lateral view compared to the other species of Frischianus known so far (Fig. 139).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01435FCFCE27C04CA58C1.taxon	description	(Figs 52, 145 – 147, 217, 218, 241, 261)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01435FCFCE27C04CA58C1.taxon	materials_examined	Type specimens: Malaysia: Holotype ♂, Sabah (04 ° 63 ' 05 " – 04 ° 77 ' 16 " N, 117 ° 43 ' 83 " – 117 ° 70 ' 31 " E), 300 m, II. – III. 2011, leg. Ewers et al. (SAFE-Projekt) (HECO). Paratypes (29 specimens): 5 ♂, 14 ♀, same data as holotype (HECO, MFNB, NHML); 3 ♂, 5 ♀, Sabah: Tawau (04 ° 66 ' N, 117 ° 06 ' E), 100 m, 11. – 17. X. 2012, leg. Gray (HECO, MFNB); 1 ♂, 1 ♀, Sarawak: Mt Mulu National Park, 50 – 100 m, V. – VIII. 1978, leg. Hammond & Marshall (NHML).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01435FCFCE27C04CA58C1.taxon	description	Description: Habitus (Fig. 52). Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color medium brown to dark brown with elytra unicolored dark brown or lighter brown anteriorly in variable extent; tip of abdomen, antennae, palps, and legs light brown. Labral denticulation type 1 or 2 (cf. Figs 10 – 12). Forebody surface microsculpture type 1 (Fig. 1). Sutural elytral row of punctures impressed, extending over anterior half of elytral sutural length; medial and lateral elytral row of punctures reduced to few, widely spaced punctures. Total body length 2.7 – 3.4 mm; forebody length 1.6 – 1.8 mm. Male: Abdominal sternite VII with midlongitudinal deplanation from about median third of sternite width slightly narrowed toward faintly emarginate posterior sternite margin; posterior third of midlongitudinal deplanation with irregularly limited, indistinct cluster of black macrosetae (Fig. 217). Abdominal sternite VIII with narrow triangular, medioposterior incision with slightly concave lateral margins, occupying about posterior 0.3 of sternite length (Fig. 218). Aedeagus (Figs 145 – 147) with long, narrow apical lobe situated in sinistral half of distal aedeagal width (Figs 146, 147). Apical lobe in lateral view strongly curved ventrad and evenly tapered toward more strongly bent apical end (Fig. 145), in dorsal view narrow, somewhat thickened at midlength, and with subacute end (Fig. 147). Endophallic sclerite situated subsinistrally in aedeagus (Figs 146, 147), in lateral view slender basally, somewhat widened toward ventrad curved, acute apical end, not protruding from aedeagus dorsally (Fig. 145); endophallic sclerite in ventral and dorsal view subparallel with narrow, furcate base, at about midlength slightly narrowed, and with wide, truncate apical end (Figs 146, 147). Ostium with dextral margin slightly extended into short, round, dextrodistal lobe (Figs 146, 147). Aedeagus with ventral margin between postforamen and base of apical lobe evenly, moderately concave (Fig. 145), in dorsal view long oval with lateral margins bent almost equally, gradually tapered distad (Fig. 147). Length of aedeagus 0.37 – 0.4 mm. Female: Gonocoxal plate short, about 1.5 times as long as wide, without basal and subbasal ridge, with anterior margin triangularly emarginate, lateral margins slightly concave, and posterior margin almost semicircularly emarginate with subacute lateroposterior ends (Fig. 261).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01435FCFCE27C04CA58C1.taxon	distribution	Distribution: Frischianus telnovi is distributed in the Malaysian provinces of Sabah and Sarawak, northern Borneo.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01435FCFCE27C04CA58C1.taxon	etymology	Etymology: Johannes Frisch dedicates this new species to his dear colleague Dmitry Telnov, Natural History Museum London, United Kingdom, in gratitudefor his generous support during a recent visit to the Coleoptera collection there (epithet telnovi: Latinized noun, derived from the surname Telnov, genitive, singular).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD01435FCFCE27C04CA58C1.taxon	discussion	Comment: A female of Frischianus from the Philippines (Mindanao, E-slope Mt McKinley, 28. VIII. 1946, leg. Werner, FMNH) shares with F. telnovi the shape of the female gonocoxal plate with a triangularly emarginate basal margin (cf. Fig. 248). It furthermore resembles F. telnovi in general appearance and the sculpturing of the body surface, but it differs by somewhat smaller eyes and unicolored medium brown body coloring. It may represent an undescribed species, but males are unknown and in the Scopaeina both coloring and eye size often vary infraspecifically and geographically.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD51435FF5EE3FC05975821.taxon	description	(Figs 148 – 150, 219, 220)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD51435FF5EE3FC05975821.taxon	materials_examined	Type specimen: Holotype ♂, Indonesia, Banten, Lebak: Citorek Kidul (06 ° 44 ' 51 " S, 106 ° 19 ' 13 " E), 870 m, 23. V. 2016, leg. Frisch (ZMB).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD51435FF5EE3FC05975821.taxon	description	Description: Habitus cf. Fig. 52. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Head medium brown; pronotum, elytra, and abdomen light brown; antennae, palps, and legs yellow brown. Labral denticulation type 2 to 3 (cf. Figs 11 – 14). Forebody surface microsculpture type 1 (Fig. 1). Sutural elytral row of punctures distinct, but weakly impressed, occupying anterior 0.7 of elytral sutural length; medial and lateral row of punctures reduced to few, widely spaced punctures. Total body length 3.1 mm; forebody length 1.7 mm. Male: Abdominal sternite VII in basal half with shallow, longitudinal, ovoidal impression occupying median third of sternite width followed posteriorly by two adjacent, submedian clusters of black macrosetae pointing medioposteriorly toward each other (Fig. 219). Abdominal sternite VIII with narrow, medioposterior incision with concave lateral margins, occupying about 0.4 of sternite length (Fig. 220). Aedeagus (Figs 148 – 150) with apical lobe situated in sinistral half of distal aedeagal width (Fig. 149). Apical lobe in lateral view about as long as wide and triangular with acute, moderately ventrad curved apical end (Fig. 148), in ventral view with straight dextral margin and evenly dextrad curved sinistral margin gradually tapered toward rounded apical end (Fig. 149). Endophallic sclerite situated submedially in aedeagus (Fig. 150), in lateral view evenly, weakly dilated distad with subacute, moderately ventrad curved apical end slighly protruding from aedeagus dorsally (Fig. 148), in dorsal view from narrow, furcate base weakly curved dextrad with wide, round apical end almost reaching apex of apical lobe; in ventral and dorsal view, endophallic sclerite notably surpassing dextral margin of apical lobe (Figs 149, 150). Ostium with dextral margin strongly extended into long, longitudinally directed, dextrodistal spine of half length of apical lobe (Figs 149, 150). Aedeagus with ventral margin between postforamen and base of apical lobe concave (Fig. 148), in dorsal view long oval with both lateral margins bent equally, gradually tapered distad (Fig. 150). Length of aedeagus 0.36 mm. Female unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD51435FF5EE3FC05975821.taxon	distribution	Distribution: Frischianus setifer is native to western Java and was discovered in the east of Banten Province.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD51435FF5EE3FC05975821.taxon	etymology	Etymology: The epithet setifer [adjective, Latin, composed of the noun seta (bristle) and the verb ferre (to carry)] refers to the remarkable pair of posterior clusters of bristles of abdominal sternite VII of this new species, the arrangement of which is unique within Frischianus so far (Fig. 219).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD51434FCFCE39C07EF5841.taxon	description	(Figs 151 – 153, 221, 222)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD51434FCFCE39C07EF5841.taxon	materials_examined	Type specimens: Holotype ♂, Malaysia, Penang: Penang Island, X. 1913, leg. Bryant (NHML). Paratype: 1 ♂, Perak, leg. Doherty (NHML).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD51434FCFCE39C07EF5841.taxon	description	Description: Habitus cf. Fig. 52. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color reddish medium brown with light brown antennae, palps, and legs. Labral denticulation type 2 (cf. Figs 11, 12). Forebody surface microsculpture type 1 (Fig. 1). Sutural elytral row of punctures distinct, but weakly impressed, occupying about anterior half of elytral sutural length; medial and lateral elytral row of punctures weakly developed. Total body length 3.1 mm; forebody length 1.7 mm. Male: Abdominal sternite VII with shallow, midlongitudinal deplanation from about median third of basal sternite width slightly narrowed toward truncate posterior sternite margin; midlongitudinal deplanation in about posterior fifth with two adjacent, not clearly separated, submedial clusters of short, black macrosetae each slightly aligned medioposteriad (Fig. 221). Abdominal sternite VIII with narrow triangular, medioposterior incision with concave lateral margins, occupying about 0.3 of sternite length (Fig. 222). Aedeagus (Figs 151 – 153) with long, slender apical lobe arising from sinistral half of distal aedeagal width (Figs 152, 153). Apical lobe in lateral view slightly curved ventrad, parallel basally, in distal half moderately, convexly dilated ventrally, after that evenly tapered toward acute, slightly ventracurved apical end (Fig. 151); in dorsal view, apical lobe narrow, slightly bent dextrad in basal half, at about midlength somewhat dilated, afterwards feebly curved longitudinally with thin, blunt end (Fig. 153). Endophallic sclerite in aedeagus situated dextrally (Figs 152, 153), relatively slender, in lateral view with ventrad curved, notably dilated apex (Fig. 151), in dorsal view narrow, subparallel, with deeply furcate base and wide, sinistracurved, subacute apical end slightly protruding from ostium (Fig. 153). Ostium with dextral margin extended into longitudinally aligned, dextrodistal tooth (Figs 152, 153). Aedeagus with ventral margin between postforamen and apical lobe convex proximally and concave distally (Fig. 151), in dorsal view long oval with lateral margins curved equally, and gradually tapered distad (Fig. 153). Length of aedeagus 0.43 mm. Female unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD51434FCFCE39C07EF5841.taxon	distribution	Distribution: Frischianus barclayi is distributed in the Malay Peninsula and known from the neighbouring provinces of Penang and Perak.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD51434FCFCE39C07EF5841.taxon	etymology	Etymology: With choosing the epithet barclayi (Latinized noun, derived from the surname Barclay, genitive, singular) Johannes Frisch warmly thanks Max Barclay, Natural History Museum, London, United Kingdom, for his hospitality during a recent visit to the Coleoptera collection there.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD41434FF5EE37C02B65981.taxon	description	(Figs 154 – 156, 223, 224, 242, 262)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD41434FF5EE37C02B65981.taxon	materials_examined	Type specimens: Holotype ♂, Indonesia, Sulawesi Tengah, Banggai: N Kayutanyo (Matanyo Forest), 120 m, 2. – 7. XI. 1989, leg. Krikken & van der Blom (NBCL). Paratype: 1 ♀, same data as holotype (NBCL).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD41434FF5EE37C02B65981.taxon	description	Description: Habitus cf. Fig. 52. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color unicolored reddish medium brown with light brown antennae, palps, and legs. Labral denticulation type 2 (cf. Figs 11, 12). Forebody surface microsculpture resembling type 1 (Fig. 1), but very dense pronotal punctation not umbilicate. Sutural elytral row of punctures distinct, but weakly impressed, occupying anterior half of elytral sutural length; medial elytral row of punctures reduced to few, widely spaced punctures; lateral elytral row of punctures indistinct. Total body length 3.6 mm; forebody length 1.8 mm. Male: Abdominal sternite VII with shallow, midlongitudinal deplanation gradually narrowed from median third of basal sternite width to slightly concave median sixth of posterior sternite margin; midlongitudinal deplanation with sparse primary setation, on either side delimited with irregular row of black macrosetae increasingly more densely arranged posteriad (Fig. 223). Abdominal sternite VIII with narrow, medioposterior incision with concave lateral margins, occupying roughly 0.3 of sternite length (Fig. 224). Aedeagus (Figs 154 – 156) with dextrad curved distal end and apical lobe notably shifted dextrally (Figs 155, 156); apical lobe in lateral view long, curved ventrad at base, with straight dorsal and convex ventral margin at about midlength, then gradually tapered toward strongly ventrad curved, uncinate apical end (Fig. 154); apical lobe in ventral and dorsal view with strongly convex dextral margin and moderately convex sinistral margin strongly tapered toward subacute tip; dextral margin of apical lobe shortly concave apically (Figs 155, 156). Endophallic sclerite situated subdextrally in aedeagus (Figs 155, 156), not much protruding from ostium (Fig. 156), in lateral view slender with strongly ventrad curved apical end (Fig. 154), in ventral and dorsal view broad, in about distal half somewhat convexly widened, ending in strongly dextrad curved, minutely pointed apical end (Figs 155, 156). Ostium with dorsodextral margin convexly enlarged (Fig. 154); dorsodextral margin in dorsal view from sinistal side of aedeagus running obliquely distodextrad (Fig. 156). Aedeagus with ventral margin between postforamen and base of apical lobe feebly convex proximally but strongly convex distally (Fig. 154), in dorsal view long oval with dextral lateral margin somewhat more strongly convex than almost straight, sinistral lateral margin, and gradually tapered distad (Fig. 156). Length of aedeagus 0.46 mm. Female: Gonocoxal plate short, about 1.3 times as long as wide, widest across line-like thin, concave basal ridge, without subbasal ridge; lateral margins straight, gradually narrowed toward more strongly narrowed posterior seventh; posterior emargination narrow, almost semicircular, delimited laterally by subacute ends (Fig. 262).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD41434FF5EE37C02B65981.taxon	distribution	Distribution: Frischianus huijbregtsi is native to Sulawesi, where it was found in the Matanyo Forest, Sulawesi Tengah, Indonesia.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD41434FF5EE37C02B65981.taxon	etymology	Etymology: The epithet huijbregtsi (Latinized noun, derived from the surname Huijbregts, genitive, singular) honours Hans Huijbregts, Naturalis Biodiversity Center, Leiden, the Netherlands, who kindly hosted Johannes Frisch during a visit to the Coleoptera collection there where this new species was discovered.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD61436FF5EE19C02CA58C2.taxon	description	(Figs 157 – 159, 225, 226)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD61436FF5EE19C02CA58C2.taxon	materials_examined	Type specimens: Holotype ♂, Thailand, Chiang Mai: Doi Chiang Dao, 880 m, 21. II. 1986, leg. Schwendiger (MHNG). Paratype: 1 ♂, same data as holotype, but 760 m, 16.10.1986 (MFNB).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD61436FF5EE19C02CA58C2.taxon	description	Description: Habitus cf. Fig. 54. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color unicolored medium brown with light brown tip of abdomen, palps, and legs; antennae from medium brown scapus gradually lighter to light brown distal segments. Labral denticulation type 4 (cf. Fig. 15). Forebody surface microsculpture not assignable to types 1 – 4 (Figs 1 – 4); cephalic punctation as in Fig. 3 a; pronotum polished with very dense, non-umbilicate punctation; elytral punctation slightly coarser and less dense than cephalic punctation, not clearly granular. Sutural elytral row of punctures notably impressed anteriorly, occupying about anterior 0.7 of elytral sutural length; medial and lateral elytral rows of punctures indistinct. Total body length 4.2 mm; forebody length 2.1 mm. Male: Abdominal sternite VII with very short, wide emargination of posterior margin and shallow, midlongitudinal depression gradually widened posteriorly to middle third of sternite width and here covered with two adjacent, submedial clusters of few, short, black macrosetae (Fig. 225). Abdominal sternite VIII in posterior 0.3 of length with widely triangular emargination continued anteriorly into narrow, longitudinally aligned end (Fig. 226). Aedeagus (Figs 157 – 159) without apical lobe. Long ventral process arising from aedeagus shifted far proximodextrally with apical end reaching apical end of endophallic sclerite; ventral process in lateral view thin with incrassate, dorsally denticulate basal third directed distoventrally and sublongitudinally aligned distal two-thirds (Fig. 157), in ventral view with narrow base situated close to dextral margin of aedeagus, followed by strongly sinistrad curved midlength portion sinistrally strongly convexly dilated, after that evenly dextracurved and gradually tapered toward narrow, blunt apical end (Fig. 158). Endophallic sclerite in ventral and dorsal view situated medially in aedeagus (Figs 158, 159), in lateral view from narrow, dorsad curved base remarkably widened distad and extended in massive apical portion convexly dilated dorsally, projecting from ostium, and directly at margin of ostium strongly curved ventrad (Fig. 157); apical portion of endophallic sclerite with thin, apical spine pointing proximad (Fig. 157), dextrodorsad pointing ventral tooth (Figs 157, 158), and short, sinistral tooth pointing proximolaterad (Figs 158, 159). Endophallic foramen remarkably triangularly enlarged dextrally with denticulate distal margin (Figs 158, 159). Dorsal endophallic spine present, long and slender with acute apex, in lateral view adjacent to endophallic sclerite, curved ventrad, and with slightly distad curved end faintly protruding over apical portion of endophallic sclerite distally, in dorsal view with distal half somewhat curved dextrad (Fig. 159). Ostium oblique, with sinistrally convex, dextrally concave dorsal margin, extended dextrodistad and beveled sinistroproximad (Figs 157 – 159). Dorsal, midlongitudinal split of phallobase type C (cf. Figs 40, 41). Aedeagus with ventral margin distal of postforamen straight (Fig. 157), in dorsal view short oval with convex lateral margins (Fig. 159). Length of aedeagus 0.51 mm. Female unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD61436FF5EE19C02CA58C2.taxon	distribution	Distribution: Frischianus crassiphallatus occurs in the very north of Thailand in the province of Chiang Mai and is the northernmost known representative of the genus.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD61436FF5EE19C02CA58C2.taxon	etymology	Etymology: The epithet crassiphallatus [adjective, Latin, composed of the adjectives crassus (thick) and phallatus (derived from the noun phallus with suffix - atus, expressing a quality / attribute of the species related to the phallus) refers to the thick, robust endophallic sclerite of this new species (Figs 157 – 159).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD61436FF5EE5BD06AD5A21.taxon	diagnosis	Diagnosis: Separated from F. laticollis subgroup (chapter 3.2.1.1.) as follows: Labral denticulation variable, but tending to type 4 (Fig. 15); labrum usually short with submedial denticles reduced to triangular tooth or obtuse angle not protruding over anterior labral margin; submedial denticles usually not longer or even shorter than pointed or obtuse sublateral denticles. Abdominal sternite VIII of ♂ with wide, triangular, medioposterior emargination with abruptly narrowed anterior end aligned longitudinally or somewhat dextrad (Figs 226 – 230). Aedeagus with long, prominent, distoventrally oriented ventral process (Figs 157 – 180); apical lobe absent (Figs 157 – 162) or present (Figs 163 – 180), neither shifted sinistrad nor curved dextrad (homology with apical lobe of F. laticollis subgroup questionable); endophallic sclerite hypertrophic with well developed foramen often enlarged laterally. Gonocoxal plate of ♀ (unknown in three of six species included) with posterior margin narrowly, triangularly incised (Figs 263 – 265).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD61436FF5EE5BD06AD5A21.taxon	discussion	Remarks: The Frischianus rufulus subgroup comprises F. rufulus comb. nov., the first described species of this phylogenetic clade, F. miscellus comb. nov., and four species new to science, that are described or redescribed below. The F. rufulus subgroup is recorded from Myanmar, Thailand (Chiang Mai), and the Malay Peninsula.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD61428FCFCE3FC05C55D21.taxon	description	(Figs 53, 160 – 162, 227)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD61428FCFCE3FC05C55D21.taxon	materials_examined	Type specimen examined: Lectotype ♂, Myanmar; labelled “ 76. ” (handwritten), “ India / orient. ” (handwritten), “ Holotypus ” (red, printed), “ Lithochar. / rufulus Kr. ” (handwritten), “ Coll. Kraatz ” (printed), “ Coll. DEI / Eberswalde ” (printed), “ Lectotype / Lithocharis rufulus / Kraatz, 1859 / des. Frisch & Herman, 2025 ” (SDEI); here designated. The number of specimens from which Kraatz (1859: 140, 141) described Lithocharis rufulus is not stated in the original description, but as male characters are described, at least one male must have been available to the author. Kraatz also did not designate a “ type ” [holotype by original designation (ICZN 1999: Article 73.1.1.)]. Thus, to stabilize the name Lithocharis rufulus according to ICZN 1999, Article 74.1., the only available specimen is designated the lectotype, a male from the Kraatz collection at SDEI, which is labelled “ Holotype ”, whose locality label is in accordance with the original description, and which bears handwritten labels which agree with the example of Kraatz’s handwriting in Horn et al. (1990: 482). New record: Thailand, Ranong, Kapoe: Khlong Nakha Wildlife Sanctuary (Phuket Mountain Range), 30 m, 29. I. 1991, leg. Schwendinger (MFNB, MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD61428FCFCE3FC05C55D21.taxon	description	Redescription: Habitus (Fig. 53). Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color unicolored medium brown; tip of abdomen, antennae, palps, and legs yellow brown. Labral denticulation type 4 (cf. Fig. 15); in lectotype sublateral labral denticles asymmetrically developed, obtuse dextrally and short-pointed sinistrally. Forebody surface microsculpture type 3 (Fig. 3). Sutural elytral row of punctures weakly impressed anteriorly, extending over approximately anterior 0.7 of elytral sutural length; medial and lateral elytral rows of punctures indistinct. Total body length 3.6 mm; forebody length 1.9 mm. Male: Abdominal sternite VII with very shallow, triangular emargination in median third of posterior margin. Abdominal sternite VIII in roughly posterior 0.2 of length with widely triangular incision continued anteriorly into very narrow, longitudinally aligned end (Fig. 227). Aedeagus (Figs 160 – 162) without apical lobe. Ventral process arising from aedeagus subapically and far dextrally and exceeding apical end of endophallic sclerite considerably in length; ventral process in lateral view thin, directed distoventrad, seemingly somewhat twisted in proximal half followed by somewhat wider distal half with slightly ventrad curved, subacute apical end (Fig. 160), in ventral view with narrow base close to dextral aedeagal margin, after that strongly curved distosinistrad toward sinistrally and convexly widened midlength portion, and gradually tapered toward narrow, acute, somewhat dextrad curved apical end (Fig. 162). Endophallic sclerite in dorsal and ventral view with elongate base almost as wide as width of aedeagus and in aedeagus situated medially (Figs 161, 162); endophallic sclerite mainly sinistrally expanded into huge, lobiform apical portion evenly curved ventrad, about heart-shaped in dorsal view, and considerably projecting from ostium (Figs 160 – 162); apical portion of endophallic sclerite at broad, emarginate apex ventrally extended in round dextral lobe (Figs 160, 161), shorter, narrow median denticle with proximad pointing tip (Figs 160, 161), and longer, wider sinistral tooth with ventroproximad pointing tip (Figs 160, 161). Endophallic foramen with smooth proximal margin, situated in median third of subapical width of apical end of endophallic sclerite (Fig. 162). Dorsal endophallic spine absent. Ostium with convex ventral margin and dorsal margin somewhat emarginate dextrally (Figs 161, 162). Shape of dorsal, midlongitudinal split of phallobase between types B and C (cf. Figs 39 – 41). Aedeagus in dorsal view narrow with parallel lateral margins (Fig. 162). Length of aedeagus 0.42 mm. Female unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFD61428FCFCE3FC05C55D21.taxon	distribution	Distribution: Frischianus rufulus was most probably described from southern Myanmar. The species that Kraatz (1859) described from “ India orient ” were collected by Helfer 1836 – 1839 in the very south of Myanmar between Dawei [Tavoy] and the Tenasserim Range (cf. Kraatz 1859: 3, Schmidt-Göbel 1849: III). 130 years later, the species was eventually rediscovered in the south of Thailand.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC8142AFCFCE69C07ED59E1.taxon	description	(Figs 54, 163 – 165, 228, 243, 263)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC8142AFCFCE69C07ED59E1.taxon	materials_examined	Type specimens: Lectotype ♂, Malaysia, Penang, leg. Cameron; labelled “ Penang, / Dr. Cameron. ” (printed), “ Paddy (handwritten) / Debris ” (printed), “ M. / miscellus / TYPE Cam. ” (handwritten, “ TYPE ” in red), “ M. Cameron / Bequest / B. M. 1955 - 147. ” (printed), “ Lectotype / Medon miscellus / Cameron, 1932 / des. Frisch & Herman, 2025 ” (NHML); here designated. Paralectotypes (1 ♂, 2 ♀), same locality label as lectotype, paralectotype labels by Frisch & Herman, 2025 (NHML). In the original description, Cameron (1932: 128 – 130) neither stated the number of specimens from which he described Medon miscellus nor did he designate a holotype by original designation (ICZN 1999: Article 73.1.1.). Thus, a lectotype designation is required to stabilize the name Medon miscellus according to ICZN 1999, Article 74.1. From among the four syntypes from Penang, the male which Cameron had labelled as “ TYPE ”, judging from the example of Cameron’s handwriting in Horn et al. (1990: 477), is selected as the lectotype. New record: Malaysia, Penang, Penang Island: Penang Hill, 700 m, 9. XI. 1999, leg. Cuccodoro & Löbl (MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC8142AFCFCE69C07ED59E1.taxon	description	Redescription: Habitus (Fig. 54). Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color unicolored medium brown, or head slightly darker brown and elytra with darker brown to blackish brown lateroposterior angles in variable extent up to blackish brown lateral thirds leaving only wide suture strip and humera median brown; tip of abdomen, antennae, palps, and legs yellow brown. Labral denticulation variable; submedial denticles pointed and longer than pointed sublateral denticles (type 1, cf. Fig. 10), or submedial denticles very short, triangular, and sublateral denticles reduced (type 4, Fig. 15). Forebody microsculpture type 3 (Fig. 3), but elytral punctation less granular than in Fig. 3 c. Sutural elytral row of punctures weakly impressed anteriorly, occupying roughly anterior 0.7 of elytral sutural length; medial and lateral rows of punctures indistinct. Total body length 3.3 – 3.5 mm; forebody length 1.8 – 1.9 mm. Male: Abdominal sternite VII with very short, triangular emargination in median third of posterior margin. Abdominal sternite VIII in roughly posterior 0.3 of length with widely triangular incision continued anteriorly into very narrow, slightly dextrally aligned end (Fig. 228). Aedeagus (Figs 163 – 165) with thin apical lobe running longitudinally between ventral process and process of endophallic sclerite (Fig. 163). Ventral process long, exceeding apical end of endophallic sclerite considerably in length, in lateral view thin with longitudinally curved, somewhat incrassate median third followed by first shortly narrowed, then moderately widened apical third with subacute tip (Fig. 163); in ventral view, ventral process arising from aedeagus medially, from base strongly widened toward median portion with concave dextral and convex sinistral margin, afterwards strongly, evenly tapered toward thin, acute, longitudinal apical end (Fig. 164). Endophallic sclerite spiniform and considerably projecting from ostium, with base inserted in aedeagus sinistrally (Fig. 165); endophallic sclerite in lateral view uncinate and from wide median portion gradually tapered toward thin, acute, evenly ventroproximad curved apical tip (Fig. 163), in dorsal view relatively broad, somewhat directed dextrad, gradually, weakly widened and protruding over apical lobe dextrally, afterwards abruptly narrowed into thin, sinistral, sinistroproximad curved apical spine (Fig. 165). Endophallic foramen with extremely finely denticulate proximal margin, situated in about subapical midwidth of wide portion of endophallic sclerite (Fig. 165). Dorsal endophallic spine absent. Ostium ventrally obliquely projected distodextrad (Figs 163 – 165), dorsally strongly concave proximad (Fig. 165). Dorsal, midlongitudinal split of aedeagus type B (cf. Fig. 39). Aedeagus in dorsal view oval with subparallel lateral margins (Fig. 165). Length of aedeagus 0.43 – 0.44 mm. Female: Gonocoxal plate stout, about 1.6 times as long as wide, with trapezoidal basal ridge, without subbasal ridge, from enlarged lateroposterior ends of basal ridge concavely narrowed toward deep posterior emargination laterally delimited by acute lateroposterior ends (Fig. 263).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC8142AFCFCE69C07ED59E1.taxon	distribution	Distribution: Frischianus miscellus was described from Penang, Malaysia, but the exact type locality was not published by Cameron (1932: 129, 130). It was later collected on Penang Island.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCA142DFCFCE5BC07B45C61.taxon	description	(Figs 166 – 168, 229)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCA142DFCFCE5BC07B45C61.taxon	materials_examined	Type specimens: Holotype ♂, Malaysia, Selangor: Sungai Buloh, 27. VII. 1972, leg. Jaccoud (MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCA142DFCFCE5BC07B45C61.taxon	description	Description: Habitus cf. Fig. 55. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color medium brown with head and lateroposterior angles of elytra slightly darker brown; tip of abdomen, antennae, palps, and legs yellow brown. Labral denticulation type 4 (cf. Fig. 15), with submedial labral denticles strongly reduced to obtuse angle, not protruding over anterior labral margin, and not longer than obtuse sublateral labral denticles. Forebody surface microsculpture not assignable to types 1 – 4 (Figs 1 – 4); cephalic punctation as in Fig. 3 a; pronotum polished with very dense, non-umbilicate punctation; elytral punctation slightly coarser and less dense than cephalic punctation. Sutural elytral row of punctures weakly impressed anteriorly, occupying about anterior 0.7 of elytral sutural length; medial and lateral rows of punctures indistinct. Total body length 3.3 mm; forebody length 1.9 mm. Male: Abdominal sternite VII with extremely short, inconspicuous, median emargination of posterior margin. Abdominal sternite VIII in roughly posterior 0.3 of length with widely triangular incision continued anteriorly into very narrow, somewhat dextrally aligned end (Fig. 229). Aedeagus (Figs 166 – 168) with large apical lobe somewhat displaced dorsodextrad (Fig. 166); apical lobe in lateral view broad with narrow, abruptly tapered apex, running distoventrad between ventral lobe and spiniform endophallic sclerite, shorter than ventral lobe, and only half as long as from ostium protruding portion of endophallic sclerite (Fig. 166); apical lobe in ventral view lobiform with broadly convex end, somewhat directed dextrad (Fig. 167). Ventral process long, arising from aedeagus subapically (Fig. 166) and subdextrally (Fig. 167), in lateral view thin and directed distoventrad with slightly distad curved apical end (Fig. 166), in ventral view close to base notably dilated sinistrally, somewhat aligned sinistrad, and over almost entire length gradually tapered toward strongly dextrad bent, acute apical end (Fig. 167). Endophallic sclerite spiniform, significantly projecting from ostium, and situated sinistrally in aedeagus (Fig. 168); basal portion of endophallic sclerite straight with parallel margins, in lateral view aligned longitudinally (Fig. 166), in dorsal view aligned somewhat dextrad (Fig. 168); endophallic sclerite in lateral view notably incrassate at about midlength, after that strongly curved ventrad toward gradually tapered, distad curved apical portion (Fig. 166); in dorsal view, apical portion of endophallic sclerite evenly curved sinistrad and gradually tapered toward acute apical end (Fig. 168). Endophallic foramen (Figs 166, 168) narrow, slightly protruding beyond dextral margin of endophallic sclerite (Fig. 168), with straight, slightly denticulate proximal margin. Dorsal endophallic spine absent. Ostium with dorsal margin convexly projected distosinistrad (Fig. 168). Dorsal, midlongitudinal split of phallobase type B (cf. Fig. 39). Aedeagus stout, in lateral view from broad distal end (without distal lobes) strongly narrowed proximad (Fig. 166), in dorsal view short oval and with subparallel lateral margins (Fig. 168). Length of aedeagus 0.53 mm. Female unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCA142DFCFCE5BC07B45C61.taxon	distribution	Distribution: Frischianus tridens is known only from Sungai Buloh near Kuala Lumpur in the province of Selangor, Malaysia.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCA142DFCFCE5BC07B45C61.taxon	etymology	Etymology: The epithet tridens (noun, Latin, nominativ: trident) was selected to reflect the shape of the aedeagus (Fig. 166), which is reminiscent of a trident, a forklike instrument or weapon.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCD142DFF5EE75C033A5B61.taxon	description	(Figs 3, 55, 169 – 174, 230, 244, 264)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCD142DFF5EE75C033A5B61.taxon	materials_examined	Type specimens: Malaysia, Pahang: Holotype ♂, Ringlet, 1250 m, 26. III. 1993, leg. Calame & Löbl (MHNG). Paratypes (4 specimens): 2 ♀, same data as holotype (MHNG); 1 ♂, 1 ♀, Tanah Rata (Cameron Highlands), 19. – 23. III. 2008, leg. Hammond (MFNB, NHML).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCD142DFF5EE75C033A5B61.taxon	description	Description: Habitus (Figs 3, 55). Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Specimens from Ringlet medium brown or reddish brown with head slightly darker brown; specimens from Tanah Rata darker colored with dark brown head, contrasted reddish brown pronotum, dark brown elytra except for reddish brown humera, and medium brown abdomen gradually darker posteriorly; tip of abdomen, antennae, palps, and legs yellow brown to light brown. Labral denticulation type 4 (cf. Fig. 15), variable; submedial denticles pointed, somewhat longer than triangular sublateral denticles, or submedial denticles reduced to obtuse angle not protruding over anterior labral margin and not longer than reduced sublateral denticles. Forebody surface microsculpture type 3 (Fig. 3). Sutural elytral row of punctures weakly impressed anteriorly, occupying about anterior 0.7 of elytral sutural length; medial and lateral rows of punctures indistinct. Total body length 3.3 – 3.6 mm; forebody length 1.8 – 2.0 mm. Male: Abdominal sternite VII without modification. Abdominal sternite VIII in about posterior 0.3 of length with widely triangular incision continued anteriorly into very narrow, somewhat dextrally aligned end (Fig. 230). Aedeagus (Figs 169 – 174) with apical lobe strongly displaced dorsodextrad, thus located just ventral to midwidth of dextral ostial margin (Figs 169, 172); apical lobe spiniform, pointing distally, shorter than midlength of ventral process, not reaching apex of endophallic sclerite (Figs 169, 172), and not projecting over ventral process laterally (Figs 170, 173). Ventral process arising from aedeagus ventroapically (Figs 169, 172) and subdextrally (Figs 170, 173) and aligned distoventrally (Figs 169, 172); ventral process in lateral view with very narrow base, somewhat incrassate at midlength, and slightly narrowed toward subacute apical end (Figs 169, 172), in ventral view aligned longitudinally and from slightly sinistrally widened base evenly tapered toward shortly dextracurved, acute apical end (Figs 170, 173). Endophallic sclerite spiniform, significantly projecting from ostium, with base situated sinistrally in aedeagus (Figs 171, 174); endophallic sclerite in lateral view with broad, ventrally convexly dilated, longitudinally aligned basal portion (Figs 169, 172), at endophallic foramen abruptly narrowed, and continuing in distoventrad bent, thin distal portion with slight, dorsally convex curvature gradually tapered toward thin, acute apical end; in dorsal view, endophallic sclerite clearly directed distodextrad, with straight, parallel basal portion, expanded median portion with convex sinistral margin and strongly, triangularly dextrad protruding endophallic foramen with concave, denticulate proximal margin, and strongly tapered, somewhat sinistrad curved apical portion (Figs 171, 174). Dorsal endophallic spine absent. Ostium with dorsal margin convexly projected distosinistrad (Figs 171, 174). Dorsal, midlongitudinal split of aedeagus type C (cf. Figs 40, 41), indistinct in dorsal view. Aedeagus stout, with dextrad shifted median foramen (Figs 170, 173), in lateral view from broad distal end strongly narrowed posteriorly (Figs 169, 172), in dorsal view short oval with convex lateral margins (Figs 171, 174). Length of aedeagus 0.54 mm. Female: Gonocoxal plate long, about 2.0 times as long as wide, with narrow, transverse basal ridge, without subbasal ridge, in posterior half gradually narrowed toward narrow, triangular posterior emargination laterally delimited by subacute ends (Fig. 264).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCD142DFF5EE75C033A5B61.taxon	distribution	Distribution: Frischianus rubiginicollis was discovered near the towns of Ringlet and Tanah Rata in the Cameron Highlands, Malay Peninsula.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCD142DFF5EE75C033A5B61.taxon	etymology	Etymology: The epithet rubiginicollis [adjective, Latin, composed of the adjective rubiginosus (reddish brown) and the noun collum (neck)] is derived from the reddish brown pronotum of the specimens from Tanah Rata, that is contrasted with darker head and elytra.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCD142EFCFCE05C07E65A62.taxon	description	(Figs 56, 175 – 180, 265)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCD142EFCFCE05C07E65A62.taxon	materials_examined	Type specimens: Malaysia: Holotype ♂, Selangor, The Gap, 900 m, 14. III. 1993, leg. Calame & Löbl (MHNG). Paratypes (4 specimens): 1 ♂, 1 ♀, Selangor, Gombak Valley, 460 m, 13. VII. 1968, leg. Taylor (MFNB, MHNG); 1 ♂, Pahang, 19 km NE Kuala Lumpur, 28. IV. 1977, leg. Watrous (FMNH); 1 ♀, Pahang, 15 km N Kuala Lumpur, 29. IV. 1977, leg. Watrous (FMNH).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCD142EFCFCE05C07E65A62.taxon	description	Description: Habitus (Fig. 56), exoskeletal characters (cf. Fig. 3), and shape of abdominal sternite VIII of ♂ (cf. Fig. 230) as in Frischianus rubiginicollis, but different as follows. Body color unicolored light reddish brown (teneral?) to reddish medium brown with slightly darker brown head and dark brown lateroposterior angles of elytra. Total body length 3.4 – 3.6 mm; forebody length 1.8 – 1.9 mm. Male: Aedeagus (Figs 175 – 180) with apical lobe aligned distad (Fig. 175) or feebly bent dorsad (Fig. 178). Endophallic sclerite in lateral view with moderate to slight, convex, dorsal dilatation in basal portion and thinner, dorsally concave apical portion (Figs 175, 178); in dorsal view, endophallic sclerite with notably thinner, longitudinally aligned basal portion clearly set off from sinistrally dilated, dextrad pointing median portion followed by spiniform, moderately sinistrad curved apical portion (Figs 177, 180). Endophallic foramen apparently more strongly projecting dextrally due to narrow base of apical spine of endophallic sclerite, and with convex proximal margin (Figs 177, 180). Dorsal margin of ostium convexly projected distomediad (Fig. 180). Length of aedeagus 0.5 – 0.53 mm. Female: Gonocoxal plate subrhomboid, about 2.0 times as long as wide, from widest diameter at midlength equally narrowed both anteriorly and posteriorly, with less narrow triangular posterior emargination laterally delimited by less pointed ends (Fig. 265).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCD142EFCFCE05C07E65A62.taxon	distribution	Distribution: Frischianus karneri was collected in the mountainous border region of the neighboring Malay provinces of Pahang and Selangor northeast of Kuala Lumpur.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCD142EFCFCE05C07E65A62.taxon	etymology	Etymology: Johannes Frisch dedicates this new species to his dear friend Michael Karner, Frankfurt am Main, specialist of Psammoecus (Silvanidae), in memory of joint entomological expeditions and in gratitude for inspiring discussions about entomology (e. g. clear taxa delimitations versus rice pudding) and life in general (epithet karneri: Latinized noun, derived from the surname Karner, genitive, singular).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCE142EFF5EE15D03CA5FA1.taxon	diagnosis	Diagnosis: Separated from F. laticollis species group (chapter 3.2.2.) as follows: Pronotal punctures tend to fuse longitudinally, forming somewhat undulate, longitudinal grooves continuous or joined in places and separated by alternating ridges formed by longitudinally confluent puncture interspaces (pronotal microsculpture type 4); alternating pronotal furrows and ridges clear (Fig. 4 b) or only partially developed with visibility depending on lighting conditions (Figs 57, 58). Aedeagus symmetrical (Figs 187 – 198) or slightly asymmetrical (Figs 181 – 186); apical lobe absent, dorsal lobe present; parameres placed transversely, on their narrow side shortly contiguous with circoforamen and extended dorsad, therefore in ventral and dorsal view protruding laterally from contours of phallobase (e. g. Figs 181 – 183), possibly absent in one species; dorsal, midlongitudinal split of aedeagus narrow, but widely opened into ostium, with lateral margins curved toward each other or overlapping at midlength (type D, Fig. 42). Gonocoxal plate of ♀ (unknown in one of five species included) strongly enlarged anteriorly, hypertrophic (Figs 266 – 269). Bursal duct at transition to sperm pump strongly curved or undulate (Figs 245 – 247). Distribution in India (Kerala, Tamil Nadu).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCE142EFF5EE15D03CA5FA1.taxon	discussion	Remarks: In Figs 181 – 198, the transverse parameres of the F. strigaticollis species group are little contrasted, often indistinct and thus in the print version of this publication probably hardly recognizable in the lateral view of the aedeagus, but certainly well visible in a high-resolution pdf. The laterad protruding parameres are well discernible in the ventral and dorsal view of the aedeagus. The Frischianus strigaticollis species group currently comprises five species new to science, which are described below. Due to their considerable differences in male and female sexual characters, it can be assumed that they represent different clades, but the introduction of subgroups is not yet appropriate given the presumably large number of undiscovered species throughout India.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCE1420FCFCE71C040E5CE1.taxon	description	(Figs 57, 181 – 186, 231, 245, 266)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCE1420FCFCE71C040E5CE1.taxon	materials_examined	Type specimens: India, Tamil Nadu: Holotype ♂, 16 km E Kodaikanal (Palni Hills), 1400 m, 15. XI. 1972, leg. Besuchet, Löbl & Mussard (MHNG). Paratypes (44 specimens): 17 ♂, 19 ♀, same data as holotype (MFNB, MHNG); 6 ♂, 2 ♀, 23 km E Kodaikanal (Palni Hills), 1200 m, 16. XI. 1972, leg. Besuchet, Löbl & Mussard (MFNB, MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCE1420FCFCE71C040E5CE1.taxon	description	Description: Habitus (Fig. 57). Presumably pterodimorphous with small eyes and palisade fringe of abdominal tergite VII. Type specimens micropterous with short, narrow elytra; elytral width equal to pronotal width; elytral sutural length about 0.2 times shorter than pronotum; elytra with shallow, transverse, subbasal elytral depression; metathoracic wings strongly reduced in length, usually 1.1 – 1.2 times as long as elytral sutural length; macropterous specimens expected to appear at least temporarily due to presence of palisade fringe of abdominal tergite VII in micropterous specimens. Body color unicolored reddish light brown to medium brown, elytra rarely darker brown lateroposteriorly; tip of abdomen, antennae, palps, and legs light brown. Labral denticulation type 1 or 2 (cf. Figs 10 – 12). Forebody surface microsculpture not assignable to types 1 – 4; head little shiny with fine, compacted, setose punctation; pronotal punctation similar type 3 (Fig. 3 b), in places puncture interstices confluent to very narrow, somewhat undulate, indistinct, longitudinal ridges; elytra shiny with punctation slightly coarser and less dense than cephalic punctation, vaguely granular. Sutural elytral row of punctures shallow, usually not exceeding anterior half of elytral sutural length; medial and lateral elytral row of punctures reduced to few, indistinct punctures obscured by elytral sculpturing. Total body length 2.7 – 3.0 mm; forebody length 1.5 – 1.6 mm. Male: Abdominal sternite VII without modification. Abdominal sternite VIII with narrow, medioposterior incision with straight lateral margins, occupying about 0.2 of sternite length (Fig. 231). Aedeagus (Figs 181 – 186) asymmetrical with apical portion slightly aligned dextrad (Figs 182, 183). Dorsal lobe movably attached to aedeagus, somewhat longer than endophallic sclerite, in lateral view straight and thin (Figs 181, 184), in dorsal view triangular and somewhat aligned dextrad (Figs 183, 186). Ventral process close to ventrally projected postforamen, in lateral view slender, straight, and pointing distad (Figs 181, 184), in ventral view about as wide as third of aedeagal width and gradually widened toward round, slightly dextrad curved end (Figs 182, 185). Endophallic sclerite elongate, strongly protruding beyond ostium, in ventral and dorsal view lobate, gradually narrowed distad, and extended into longer, dextral apical tooth and shorter, sinistral apical tooth (Figs 182, 183, 185, 186) both curved ventrad (Fig. 181); apical teeth of endophallic sclerite often reduced (Figs 184 – 185); endophallic sclerite finely setulose dorsoproximally with setulae visible in lateral view (Figs 181, 184). Aedeagus in lateral view slender with ventral margin distal of ventral process almost straight with convex, subapical dilatation (Figs 181, 184), in dorsal view narrow with slightly concave lateral margins (Figs 183, 186). Length of aedeagus 0.23 – 0.4 mm. Female: Gonocoxal plate from enlarged anterior portion moderately narrowed toward setose posterior fourth; posterior margin of gonocoxal plate weakly, concavely emarginate; enlarged anterior portion of gonocoxal plate membranous and hyaline with strongly sclerotized, thin, sublateral bands densely lined with medioposteriorly pointing spinules in few, close, irregular rows (Fig. 266).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCE1420FCFCE71C040E5CE1.taxon	distribution	Distribution: Frischianus parentium is native to the Palni Hills in Tamil Nadu in the south of India.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFCE1420FCFCE71C040E5CE1.taxon	etymology	Etymology: With choosing the epithet parentium (noun, Latin, genitive: of the parents) Johannes Frisch dedicates this new species to his beloved parents Gisela (†) and Walther Frisch (†), Fulda, Germany, who supported his passion for entomology from the start.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC01420FF5EE7DC02705B02.taxon	description	(Figs 58, 187 – 189, 232, 267)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC01420FF5EE7DC02705B02.taxon	materials_examined	Type specimens: India: Holotype ♂, Kerala, Pambanar-Peermade, 950 m, 5. – 9. XI. 1972, leg. Besuchet, Löbl & Mussard (MHNG). Paratypes (67 specimens): Kerala: 7 ♂, 9 ♀, same data as holotype (MFNB, MHNG); 5 ♂, 1 ♀, 5 km E Kumily (Cardamom Hills), 1000 m, 6. XI. 1972, leg. Besuchet, Löbl & Mussard (MFNB, MHNG); 1 ♀, Nelliampathi Hills, 900 m, 30. XI. 1972, leg. Besuchet, Löbl & Mussard (MHNG); 2 ♂, Kumily, Periyar Wildlife Sanctuary, 2. IX. 1989, leg. Riedel (SMNS); 3 ♂, 10 km SW Kumily: Vallakadavu (Cardamom Hills) (09 ° 31 ' N, 77 ° 7 ' E), 1000 m, 24. XII. 1993, leg. Boukal & Kejval (NHMW). Tamil Nadu: 22 ♂, 9 ♀, 18 km N Valparai (Anaimalai Hills), 1250 m, 18. XI. 1972, leg. Besuchet, Löbl & Mussard (MFNB, MHNG); 4 ♂, 4 ♀, Valparai (Anaimalai Hills), 1100 m, 20. XI. 1972, leg. Besuchet, Löbl & Mussard (MFNB, MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC01420FF5EE7DC02705B02.taxon	description	Description: Habitus (Fig. 58). Pterodimorphous with small eyes and palisade fringe of abdominal tergite VII in both macropterous and micropterous specimens. Macropterous specimens: elytra somewhat wider than pronotal width; elytral sutural length subequal to pronotal length; metathoracic wings fully developed. Micropterous specimens: elytral width equal to pronotal width; elytral sutural length subequal to or up to 0.2 times shorter than pronotum; elytra with shallow, transverse, subbasal depression; metathoracic wings reduced in length to varying degrees, in examined specimens longer than elytral sutural length. Body color unicolored medium brown to blackish brown, or medium brown with elytra darker brown posteriorly in variable extent; tip of abdomen, antennae, palps, and legs light brown to medium brown. Labral denticulation variable, types 1 to 3; submedial denticles usually pointed, longer than pointed (cf. Fig. 10) to widely obtuse (Fig. 14) sublateral denticles. Forebody surface microsculpture type 4 (Fig. 4), but cephalic punctation finer than in Fig. 4 a and pronotal furrows and ridges much finer, denser, and less distinct than in Fig. 4 b. Sutural elytral row of punctures shallow, short, usually not exceeding anterior half of elytral sutural length; medial and lateral elytral row of punctures reduced to few, indistinct punctures. Total body length 3.0 – 3.3 mm; forebody length 1.5 – 1.7 mm. Male: Abdominal sternite VII without modification. Abdominal sternite VIII in about posterior 0.15 of length with narrow incision (Fig. 232). Aedeagus (Figs 187 – 189) symmetrical. Dorsal lobe movably attached, in lateral view thin and straight (Fig. 187), in dorsal view triangular and gradually tapered toward somewhat dextrad curved, thin, subacute apical end (Fig. 189). Ventral process situated close to postforamen, in lateral view slender, slightly curved distad, with strong, dorsal subapical tooth and moderately ventrad curved apical end (Fig. 187), in ventral view about half as wide as aedeagus, parallel with wide, round apical end (Fig. 188). Endophallic sclerite triangular, protruding beyond ostium distoventrally (Fig. 187), in ventral view with transverse, rectangular apical end (Fig. 188). Aedeagus in lateral view with ventral margin distal of ventral process almost straight (Fig. 187), in dorsal view very narrow with slightly concave lateral margins (Fig. 189). Ostium of aedeagus with convex lateral margins (Fig. 187). Length of aedeagus 0.46 – 0.5 mm. Female: Gonocoxal plate from enlarged anterior portion strongly narrowed toward subparallel, setose posterior fourth with concavely emarginate posterior margin; enlarged anterior portion of gonocoxal plate membranous and hyaline with strongly sclerotized, sublateral bands densely lined with medioposteriorly pointing spinules in few, loose, irregular rows (Fig. 267).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC01420FF5EE7DC02705B02.taxon	distribution	Distribution: Frischianus strigaticollis is distributed across the mountains of Southwest India and recorded from the Nelliampathi Hills in Tamil Nadu as far south as the Cardamom Hills, Kerala.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC01420FF5EE7DC02705B02.taxon	etymology	Etymology: The epithet strigaticollis [adjective, Latin, composed of the adjective strigatus (striped) and the noun collum (neck)] refers to the longitudinally lineate, strigate sculpture of the pronotal surface of this new species (Fig. 58).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC01422FCFCE0BC044C58C1.taxon	description	(Figs 59, 190 – 192, 233, 234, 246, 268)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC01422FCFCE0BC044C58C1.taxon	materials_examined	Type specimens: India, Kerala: Holotype ♂, Nelliampathi Hills, 900 m, 30. XI. 1972, leg. Besuchet, Löbl & Mussard (MHNG). Paratypes (94 specimens): 11 ♂, 78 ♀, same data as holotype (MFNB, MHNG); 1 ♂, 2 ♀, 10 km SW Kumily: Vallakadavu (Cardamom Hills) (09 ° 31 ' N, 77 ° 7 ' E), 1000 m, 24. XII. 1993, leg. Boukal & Kejval (NHMW); 1 ♂, 1 ♀, Periyar, 800 m, 13. I. 1972, leg. Mussard (MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC01422FCFCE0BC044C58C1.taxon	description	Description: Habitus (Fig. 59). Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color reddish medium brown with darker brown head and lateroposterior angles of elytra up to dark brown body with black head and blackish brown elytra; tip of abdomen, antennae, palps, and legs light brown to medium brown. Labral denticulation type 1 or 2 (cf. Figs 10 – 12). Forebody surface microsculpture type 4 (Fig. 4), but cephalic punctation finer than in Fig. 4 a and pronotal furrows and ridges finer and denser than in Fig. 4 b, indistinct. Sutural elytral row of punctures shallow, occupying about anterior half of sutural length with densely arranged punctures; medial and lateral row of punctures reduced to few punctures. Total body length 2.4 – 2.7 mm; forebody length 1.4 – 1.5 mm. Male: Abdominal sternite VII with median strip of ventral surface flattened and somewhat widened toward shortly emarginate median third of posterior margin delimited by strongly sclerotized, lateroposterior angles (Fig. 233). Abdominal sternite VIII in about posterior 0.15 of length with wide, triangular emargination with straight, strongly sclerotized lateral margins and obtuse-angled anterior end (Fig. 234). Aedeagus (Figs 190 – 192) symmetrical. Dorsal lobe short, not protruding apex of ventral process distally, in lateral view thin with moderately dilated, round apical end (Fig. 190), in dorsal view about as wide as aedeagus, but in apical portion on both sides concavely narrowed toward broad, transversely rectangular apical end (Fig. 192). Ventral process very close to postforamen, directed distoventrad, and reaching apex of dorsal lobe; in lateral view, distal third of ventral process somewhat curved dorsad, dorsally convexly widened, and gradually tapered toward acute apical end (Fig. 190); ventral process in ventral view almost as wide as aedeagus, subparallel, and in apical third gradually tapered toward subacute apical end (Fig. 191). Endophallic sclerite in lateral view oblong with tapered, subacute distal and proximal ends (Fig. 190), made up of three lateral, paired, elongate sclerites (Fig. 192) extended into processes directed toward each other visible in ventral view (Fig. 191): outer paired sclerites both extended into thin spine aligned distomedially; distal, inner paired sclerites short, lobiform, and aligned longitudinally; proximal, inner paired sclerites both extended into stout, apically round process aligned distomedially. Aedeagus strongly convex dorsally and with ventral margin distal of ventral process almost obtuse-angled curved (Fig. 190), in dorsal view narrow with subparallel lateral margins (Fig. 192). Length of aedeagus 0.34 – 0.37 mm. Female: Gonocoxal plate enlarged basally, but without huge, oval, hyaline, laterally spinulose anterior portion of other species of F. strigatocollis group; gonocoxal plate from wide, deeply emarginate proximal portion gradually tapered toward subparallel, setose posterior third with emarginate posterior margin; basal ridge delimiting deep, anterior emargination of gonocoxal plate, strongly sclerotized only at proximolateral angles and shortly laterad of middle of sclerite; anterior emargination of gonocoxal plate filled membranously and exhibits two lateral, medioposteriorly curved straps followed proximally by lateral, strongly sclerotized, shortly ribbed strips (Fig. 268).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC01422FCFCE0BC044C58C1.taxon	distribution	Distribution: Frischianus keralensis is native to southwestern India and recorded from the Nelliampathi Hills in the north as far south as the Cardamom Hills.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC01422FCFCE0BC044C58C1.taxon	etymology	Etymology: The epithet keralensis (adjective, Latin: from Kerala) refers to the distribution of this new species in the Indian federal state of Kerala.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC21422FF5EE3FC05B75821.taxon	description	(Figs 193 – 195, 235)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC21422FF5EE3FC05B75821.taxon	materials_examined	Type specimen: Holotype ♂, India, Kerala, 50 km SO Munnar: Valara Waterfalls, 450 – 500 m, 25. XI. 1972, leg. Besuchet, Löbl & Mussard (MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC21422FF5EE3FC05B75821.taxon	description	Description: Habitus cf. Fig. 60. Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color reddish brown with head slightly darker reddish brown and elytra dark brown in about posterior two-thirds; tip of abdomen, antennae, palps, and legs light brown. Labral denticulation type 3 (cf. Figs 13, 14). Forebody surface microsculpture type 4 (Fig. 4), but cephalic punctation and alternating furrows and ridges of elyra finer than in Figs 4 a, 4 b. Sutural elytral row of punctures shallow, not exceeding anterior half of elytral sutural length; medial and lateral row of punctures reduced to few, indistinct punctures obscured by elytral punctation. Total body length 2.9 mm; forebody length 1.6 mm. Male: Abdominal sternite VII without modification. Abdominal sternite VIII with medioposterior incision with convex lateral margins and narrow anterior end, occupying about 0.4 of sternite length (Fig. 235). Aedeagus (Figs 193 – 195) symmetrical with testaceous dorsal lobe with wide, shallowly emarginate apex (Figs 194, 195) and ventrad curved lateral margins enclosing endophallic sclerite dorsally (Fig. 193). Ventral process situated distally of postforamen, in lateral view thin, slightly curved distad and with dorsal apical tooth (Fig. 193), in ventral view long triangular and gradually tapered toward narrow, round end (Fig. 194). Endophallic sclerite in lateral view long subrectangular with tapered basal end and deeply emarginate distal end extended in wider dorsoapical lobe with round end and narrower ventroapical lobe with dilated, slightly undulate, truncate end protruding beyond dorsal lobe ventrally (Fig. 193); in ventral view, endophallic sclerite nearly as broad as aedeagus, gradually, convexly narrowed toward shortly emarginate apical ends of dorsoapical and ventroapical lobes (Fig. 194); endophallic sclerite in dorsal view in wide, oval median portion with lateral rows of mediodistad pointing setulae (Fig. 195) slightly projecting beyond dorsal margin of endophallic sclerite in lateral view (Fig. 193). Ostium strongly, concavely retracted both ventrally and dorsally (Figs 194, 195), with lateral margins strongly projected at right angles distally (Figs 193, 195). Aedeagus in dorsal view narrow with subparallel lateral margins (Figs 195). Length of aedeagus 0.44 mm. Female unknown.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC21422FF5EE3FC05B75821.taxon	distribution	Distribution: Frischianus lineolatus is known only from the type locality Valara Waterfalls in Kerala in the southwest of India.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC21422FF5EE3FC05B75821.taxon	etymology	Etymology: The epithet lineolatus (adjective, Latin: weakly lineate) refers to the finely lineate pronotum of this new species.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC2145DFCFCE39C06A15B42.taxon	description	(Figs 60, 196 – 198, 236, 247, 269)	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC2145DFCFCE39C06A15B42.taxon	materials_examined	Type specimens: India, Kerala: Holotype ♂, 10 km SW Kumily: Vallakadavu (Cardamom Hills) (09 ° 31 ' N, 77 ° 7 ' E), 1000 m, 24. XII. 1993, leg. Boukal & Kejval (NHMW). Paratypes (9 specimens): 1 ♂, 2 ♀, same data as holotype (MFNB, NHMW); 1 ♂, Nelliampathi Hills, 300 m, 30. XI. 1972, leg. Besuchet, Löbl & Mussard (MFNB); 3 ♂, Kumily: Periyon Wildlife Sanctuary (Cardamon Hills), 900 m, 7. XI. 1972, leg. Besuchet, Löbl & Mussard (MHNG); 1 ♂, Wayanad: Kottiyoor, 650 m, 7. IV. 1969, leg. Soans & Brown (MHNG).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC2145DFCFCE39C06A15B42.taxon	description	Description: Habitus (Figs 4, 60). Macrophthalmous, macropterous with palisade fringe of abdominal tergite VII. Body color light reddish brown to reddish medium brown; head slightly darker reddish brown, elytra dark brown in about posterior two-thirds; tip of abdomen, antennae, palps, and legs light brown. Labral denticulation type 3 (cf. Figs 13, 14), sublateral labral denticles obtusely angled or convex. Forebody surface microsculpture type 4 (Fig. 4). Sutural elytral row of punctures impressed with punctures densely arranged, usually not exceeding anterior half of elytral sutural length; medial and lateral row of punctures reduced to few punctures. Total body length 2.9 – 3.2 mm; forebody length 1.6 – 1.8 mm. Male: Abdominal sternite VII with very short, wide, posterior emargination feebly convex in median fourth. Abdominal sternite VIII with medioposterior incision with convex lateral margins and narrow anterior end, occupying about posterior 0.3 of length (Fig. 236). Aedeagus (Figs 196 – 198) symmetrical. Dorsal lobe testaceous and subhyaline with wide, emarginate apex, ventrally strongly expanded with convex lateral margins enclosing endophallic sclerite dorsally (Figs 196 – 198). Ventral process situated distally of postforamen, but connected with postforamen by thick sclerotization, in lateral view stocky, somewhat curved ventrad, bearing large, right-angled dorsal tooth, and gradually tapered toward subacute apical end armed with minute, dorsoapical denticle (Fig. 196); in ventral view, ventral process long subtriangular with convex lateral margins, gradually tapered toward acute apical end (Fig. 197). Endophallic sclerite stout, convex dorsally, and triangularly extended distally (Fig. 196), in ventral view nearly as wide as aedeagus, and in distal third gradually narrowed toward wide, truncate end (Fig. 197), in dorsal view proximally with V-shaped dorsal margin and distally subcircular (Fig. 198). Ostium strongly, concavely retracted both ventrally and dorsally (Figs 197, 198) with rounded lateral margins strongly projected distally (Figs 196, 198). Aedeagus with ventral margin distal of ventral process straight and finely serrate (Fig. 196), in dorsal view narrow with subparallel lateral margins (Figs 198). Length of aedeagus 0.37 – 0.4 mm. Female: Gonocoxal plate with enlarged, hyaline, anterior portion with strongly sclerotized, thin, sublateral bands densely lined with medially directed spinules in few, loose, irregular rows; anterior portion of gonocoxal plate strongly narrowed toward subparallel, setose, posterior fourth with concave posterior margin (Fig. 269).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC2145DFCFCE39C06A15B42.taxon	distribution	Distribution: Frischianus lineatocollis is widely distributed across Southwest India, where it is recorded from Kottiyoor southward to the Cardamom Hills. A female from Goa (NHMW), which is not included into the type series, is probably conspecific.	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
2B718793FFC2145DFCFCE39C06A15B42.taxon	etymology	Etymology: The epithet lineatocollis [adjective, Latin, composed of the adjective lineatus (lineate) and the noun collum (neck)] describes the lineate, strigate, longitudinal sculpture of the pronotum of this new species (Fig. 4 b).	en	Frisch, Johannes, Herman, Lee (2025): Frischianus gen. nov., a new genus of the Scopaeina Mulsant & Rey, 1878 from the Oriental Region (Coleoptera: Staphylinidae: Paederinae: Lathrobiini). Megataxa 17 (2): 217-290, DOI: 10.11646/megataxa.17.2.1
