taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
713901F9CEE75D2C93224A253ADDBAB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790698	https://doi.org/10.3897/zse.102.206462.figure2	Figure 2. The map of collection sites along the Black Sea coastline (above) and a COI - based maximum likelihood time-calibrated phylogenetic tree (reconstruction) of the relationships between Niphargus almensis sp. nov. and Niphargus uchansi sp. nov., and their congeners from the “ tarkhankuticus ” ingroup (below). The results from different species delimitation methods (1 – ABGD; 2 – ASAP; 3 – PTP / bPTP; 4 – morphology) are depicted with colored bars highlighting congruence across methods. Additional sequences of N. longicaudatus (A. Costa in Hope, 1851) (KY 706921 and KY 706922), used as an outgroup, were taken from the NCBI (GenBank) database. The estimated divergence times is indicated after Guy-Haim et al. (2018) and Copilaş-Ciocianu and Petrusek (2015).	Figure 2. The map of collection sites along the Black Sea coastline (above) and a COI - based maximum likelihood time-calibrated phylogenetic tree (reconstruction) of the relationships between Niphargus almensis sp. nov. and Niphargus uchansi sp. nov., and their congeners from the “ tarkhankuticus ” ingroup (below). The results from different species delimitation methods (1 – ABGD; 2 – ASAP; 3 – PTP / bPTP; 4 – morphology) are depicted with colored bars highlighting congruence across methods. Additional sequences of N. longicaudatus (A. Costa in Hope, 1851) (KY 706921 and KY 706922), used as an outgroup, were taken from the NCBI (GenBank) database. The estimated divergence times is indicated after Guy-Haim et al. (2018) and Copilaş-Ciocianu and Petrusek (2015).	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
713901F9CEE75D2C93224A253ADDBAB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790699	https://doi.org/10.3897/zse.102.206462.figure3	Figure 3. A general view of the alive specimens (a) and the characteristic biotopes where Niphargus almensis sp. nov. was discovered on the Alma River (b, c) and its tributary, the Bodrak River (d). The yellow arrows indicate the exact sampling locations in the riverine hyporheic habitats.	Figure 3. A general view of the alive specimens (a) and the characteristic biotopes where Niphargus almensis sp. nov. was discovered on the Alma River (b, c) and its tributary, the Bodrak River (d). The yellow arrows indicate the exact sampling locations in the riverine hyporheic habitats.	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
713901F9CEE75D2C93224A253ADDBAB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790704	https://doi.org/10.3897/zse.102.206462.figure7	Figure 7. Niphargus almensis sp. nov., holotype ♂ (ZMMU Mb- 1317) (a – k, m, o) and paratype ♀ (ZMMU Mb- 1318) (l, n, p): a – c. Epimeral plates I – III; g, h. Telson; i. Pleopod III; j. Retinacula of pleopod III; k, l. Uropod I; m, n. Uropod II; o, p. Uropod III.	Figure 7. Niphargus almensis sp. nov., holotype ♂ (ZMMU Mb- 1317) (a – k, m, o) and paratype ♀ (ZMMU Mb- 1318) (l, n, p): a – c. Epimeral plates I – III; g, h. Telson; i. Pleopod III; j. Retinacula of pleopod III; k, l. Uropod I; m, n. Uropod II; o, p. Uropod III.	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
713901F9CEE75D2C93224A253ADDBAB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790700	https://doi.org/10.3897/zse.102.206462.figure4	Figure 4. Niphargus almensis sp. nov., holotype ♂ (ZMMU Mb- 1317): a. Antenna I; b. Accessory flagellum of antenna I; c. Antenna II; d. Gnathopod I; e. Distoventral corner of propodus (palm) of GnI; f. Gnathopod II; g. Distoventral palmar margin of propodus (palm) of GnII.	Figure 4. Niphargus almensis sp. nov., holotype ♂ (ZMMU Mb- 1317): a. Antenna I; b. Accessory flagellum of antenna I; c. Antenna II; d. Gnathopod I; e. Distoventral corner of propodus (palm) of GnI; f. Gnathopod II; g. Distoventral palmar margin of propodus (palm) of GnII.	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
713901F9CEE75D2C93224A253ADDBAB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790703	https://doi.org/10.3897/zse.102.206462.figure6	Figure 6. Niphargus almensis sp. nov., holotype ♂ (ZMMU Mb- 1317): a. Pereopod III; b. Dactylus of PIII; c. Pereopod IV; d. Dactylus of PIV; e. Pereopod V; f. Dactylus of PV; g. Pereopod VI; h. Dactylus of PVI; i. Pereopod VII; j. Dactylus of PVII.	Figure 6. Niphargus almensis sp. nov., holotype ♂ (ZMMU Mb- 1317): a. Pereopod III; b. Dactylus of PIII; c. Pereopod IV; d. Dactylus of PIV; e. Pereopod V; f. Dactylus of PV; g. Pereopod VI; h. Dactylus of PVI; i. Pereopod VII; j. Dactylus of PVII.	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
713901F9CEE75D2C93224A253ADDBAB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790701	https://doi.org/10.3897/zse.102.206462.figure5	Figure 5. Niphargus almensis sp. nov., holotype ♂ (ZMMU Mb- 1317): a. Labrum (upper lip); b. Labium (lower lip); c, e. Mandible; d, f. Incisor process and pars incisiva of mandibles; g. Maxilla I; h. Same, distal margin of outer lobe; i. Maxilla II; j. Maxilliped.	Figure 5. Niphargus almensis sp. nov., holotype ♂ (ZMMU Mb- 1317): a. Labrum (upper lip); b. Labium (lower lip); c, e. Mandible; d, f. Incisor process and pars incisiva of mandibles; g. Maxilla I; h. Same, distal margin of outer lobe; i. Maxilla II; j. Maxilliped.	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
713901F9CEE75D2C93224A253ADDBAB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790697	https://doi.org/10.3897/zse.102.206462.figure1	Figure 1. A map of the main rivers on the Crimean Peninsula, along with the locations of our study sites and records of hyporheic crustaceans, including Niphargus spp. and Litorogammarus spp. Two ingroups of the genus Niphargus (Amphipoda: Niphargidae), “ tarkhankuticus ” and “ tauricus ”, have been specifically identified as having adapted to and inhabiting riverine hyporheic habitats. For further information, please see the text. Green circles with black stars indicate the localities where the species described in the article were discovered (see below).	Figure 1. A map of the main rivers on the Crimean Peninsula, along with the locations of our study sites and records of hyporheic crustaceans, including Niphargus spp. and Litorogammarus spp. Two ingroups of the genus Niphargus (Amphipoda: Niphargidae), “ tarkhankuticus ” and “ tauricus ”, have been specifically identified as having adapted to and inhabiting riverine hyporheic habitats. For further information, please see the text. Green circles with black stars indicate the localities where the species described in the article were discovered (see below).	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
713901F9CEE75D2C93224A253ADDBAB9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790708	https://doi.org/10.3897/zse.102.206462.figure11	Figure 11. Niphargus uchansi sp. nov., holotype ♂ (ZMMU Mb- 1319): a. Pereopod III; b. Dactylus of PIII; c. Pereopod IV; d. Pereopod V; e. Dactylus of PV; f. Pereopod VI; g. Dactylus of PVI; h. Pereopod VII; i. Dactylus of PVII.	Figure 11. Niphargus uchansi sp. nov., holotype ♂ (ZMMU Mb- 1319): a. Pereopod III; b. Dactylus of PIII; c. Pereopod IV; d. Pereopod V; e. Dactylus of PV; f. Pereopod VI; g. Dactylus of PVI; h. Pereopod VII; i. Dactylus of PVII.	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
F629596244FF5BF7BB5D5C70D3C5BF10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790706	https://doi.org/10.3897/zse.102.206462.figure9	Figure 9. Niphargus uchansi sp. nov., holotype ♂ (ZMMU Mb- 1319) (a, c, e, f, h) and paratype ♀ (ZMMU Mb- 1320) (b, d, g): a, b. Gnathopod I; c, d. Gnathopod II; e. Armature of urosomites I and II; f, g. Telson; h. Hooks in retinacula of pleopod III.	Figure 9. Niphargus uchansi sp. nov., holotype ♂ (ZMMU Mb- 1319) (a, c, e, f, h) and paratype ♀ (ZMMU Mb- 1320) (b, d, g): a, b. Gnathopod I; c, d. Gnathopod II; e. Armature of urosomites I and II; f, g. Telson; h. Hooks in retinacula of pleopod III.	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
F629596244FF5BF7BB5D5C70D3C5BF10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790707	https://doi.org/10.3897/zse.102.206462.figure10	Figure 10. Niphargus uchansi sp. nov., holotype ♂ (ZMMU Mb- 1319): a. Labrum (upper lip); b. Labium (lower lip); c, e. Mandible; d, f. Incisor process and pars incisiva of mandibles; g. Maxilla I; h. Same, distal margin of outer lobe; i. Maxilla II; j. Maxilliped.	Figure 10. Niphargus uchansi sp. nov., holotype ♂ (ZMMU Mb- 1319): a. Labrum (upper lip); b. Labium (lower lip); c, e. Mandible; d, f. Incisor process and pars incisiva of mandibles; g. Maxilla I; h. Same, distal margin of outer lobe; i. Maxilla II; j. Maxilliped.	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
F629596244FF5BF7BB5D5C70D3C5BF10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790708	https://doi.org/10.3897/zse.102.206462.figure11	Figure 11. Niphargus uchansi sp. nov., holotype ♂ (ZMMU Mb- 1319): a. Pereopod III; b. Dactylus of PIII; c. Pereopod IV; d. Pereopod V; e. Dactylus of PV; f. Pereopod VI; g. Dactylus of PVI; h. Pereopod VII; i. Dactylus of PVII.	Figure 11. Niphargus uchansi sp. nov., holotype ♂ (ZMMU Mb- 1319): a. Pereopod III; b. Dactylus of PIII; c. Pereopod IV; d. Pereopod V; e. Dactylus of PV; f. Pereopod VI; g. Dactylus of PVI; h. Pereopod VII; i. Dactylus of PVII.	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
F629596244FF5BF7BB5D5C70D3C5BF10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790709	https://doi.org/10.3897/zse.102.206462.figure12	Figure 12. Niphargus uchansi sp. nov., holotype ♂ (ZMMU Mb- 1319) (a, c, f, g, i, k) and paratype ♀ (ZMMU Mb- 1320) (b, d, f, h, j, l): a, b. Epimeral plate I; c, d. Epimeral plate II; e, f. Epimeral plate III; g, h. Uropod I; i, j. Uropod II; k, l. Uropod III.	Figure 12. Niphargus uchansi sp. nov., holotype ♂ (ZMMU Mb- 1319) (a, c, f, g, i, k) and paratype ♀ (ZMMU Mb- 1320) (b, d, f, h, j, l): a, b. Epimeral plate I; c, d. Epimeral plate II; e, f. Epimeral plate III; g, h. Uropod I; i, j. Uropod II; k, l. Uropod III.	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
F629596244FF5BF7BB5D5C70D3C5BF10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1790698	https://doi.org/10.3897/zse.102.206462.figure2	Figure 2. The map of collection sites along the Black Sea coastline (above) and a COI - based maximum likelihood time-calibrated phylogenetic tree (reconstruction) of the relationships between Niphargus almensis sp. nov. and Niphargus uchansi sp. nov., and their congeners from the “ tarkhankuticus ” ingroup (below). The results from different species delimitation methods (1 – ABGD; 2 – ASAP; 3 – PTP / bPTP; 4 – morphology) are depicted with colored bars highlighting congruence across methods. Additional sequences of N. longicaudatus (A. Costa in Hope, 1851) (KY 706921 and KY 706922), used as an outgroup, were taken from the NCBI (GenBank) database. The estimated divergence times is indicated after Guy-Haim et al. (2018) and Copilaş-Ciocianu and Petrusek (2015).	Figure 2. The map of collection sites along the Black Sea coastline (above) and a COI - based maximum likelihood time-calibrated phylogenetic tree (reconstruction) of the relationships between Niphargus almensis sp. nov. and Niphargus uchansi sp. nov., and their congeners from the “ tarkhankuticus ” ingroup (below). The results from different species delimitation methods (1 – ABGD; 2 – ASAP; 3 – PTP / bPTP; 4 – morphology) are depicted with colored bars highlighting congruence across methods. Additional sequences of N. longicaudatus (A. Costa in Hope, 1851) (KY 706921 and KY 706922), used as an outgroup, were taken from the NCBI (GenBank) database. The estimated divergence times is indicated after Guy-Haim et al. (2018) and Copilaş-Ciocianu and Petrusek (2015).	2026-09-25	Marin, Ivan;Palatov, Dmitry		Zenodo	biologists	Marin, Ivan;Palatov, Dmitry			
