taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883170/files/figure.png	https://doi.org/10.5281/zenodo.17883170	FIGURE 1. A, Head, anterolateral view; B, frons, dorsal view; C, frons, anterior; D, head, ventrolateral. A, B, C, Phymatocera peregrinator comb. nov., female (B, C, lectotype). D, P. nipponica, female. B, C, Photographed by H. Vårdal.	FIGURE 1. A, Head, anterolateral view; B, frons, dorsal view; C, frons, anterior; D, head, ventrolateral. A, B, C, Phymatocera peregrinator comb. nov., female (B, C, lectotype). D, P. nipponica, female. B, C, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883195/files/figure.png	https://doi.org/10.5281/zenodo.17883195	FIGURE 5. Dorsal tentorial pit and its neighboring areas, lateroanterior view. A, Phymatocera satoi sp. nov., female. B, Nesotomostethus religiosa, female. C, Phymatoceropsis japonica, female. D, Pareophora gracilis, female. E, Ardis pallipes, female. F, Periclista nigrifrons, female.	FIGURE 5. Dorsal tentorial pit and its neighboring areas, lateroanterior view. A, Phymatocera satoi sp. nov., female. B, Nesotomostethus religiosa, female. C, Phymatoceropsis japonica, female. D, Pareophora gracilis, female. E, Ardis pallipes, female. F, Periclista nigrifrons, female.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883217/files/figure.png	https://doi.org/10.5281/zenodo.17883217	FIGURE 13. Phymatocera membra.A, Female, dorsal view; B, C, male, dorsal and lateroventral views; D, tarsal claw; E, apex of female abdomen, lateral view; F, ovipositor, lateral view; G, lancet; H, ventral parts of first to third annuli of lancet; I, apical part of lancet (arrows indicating some marginal sensilla); J, K, male genitalia, dorsal and ventral views; L, penis valves, lateral view (left dorsal).	FIGURE 13. Phymatocera membra.A, Female, dorsal view; B, C, male, dorsal and lateroventral views; D, tarsal claw; E, apex of female abdomen, lateral view; F, ovipositor, lateral view; G, lancet; H, ventral parts of first to third annuli of lancet; I, apical part of lancet (arrows indicating some marginal sensilla); J, K, male genitalia, dorsal and ventral views; L, penis valves, lateral view (left dorsal).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883219/files/figure.png	https://doi.org/10.5281/zenodo.17883219	FIGURE 14. Phymatocera nipponica. A, B, Female, dorsal and lateral views; C, male, dorsal view; D, tarsal claw, female; E, apex of female abdomen, lateral view; F, ovipositor; G, ventral parts of first to fourth annuli of lancet; H–J, apical part of lancet; K, male genitalia, dorsal view (penis valves removed); L, ditto (penis valves not removed), ventral view; M, penis valve, lateral view (left dorsal). A, B, Photographed just after killing.	FIGURE 14. Phymatocera nipponica. A, B, Female, dorsal and lateral views; C, male, dorsal view; D, tarsal claw, female; E, apex of female abdomen, lateral view; F, ovipositor; G, ventral parts of first to fourth annuli of lancet; H–J, apical part of lancet; K, male genitalia, dorsal view (penis valves removed); L, ditto (penis valves not removed), ventral view; M, penis valve, lateral view (left dorsal). A, B, Photographed just after killing.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883229/files/figure.png	https://doi.org/10.5281/zenodo.17883229	FIGURE 16. Phymatocera satoi sp. nov. A, B, Female, dorsal and lateral views; C, male, dorsal view; D, tarsal claw, female; E, apex of female abdomen, lateral view; F, ovipositor, lateral view; G, ventral parts of first to third annuli; H, apical part of lancet; I, K, male genitalia, dorsal and ventral views; L, penis valve, lateral view (left dorsal). A, B, D–F, H, Holotype (F, H, reversed).	FIGURE 16. Phymatocera satoi sp. nov. A, B, Female, dorsal and lateral views; C, male, dorsal view; D, tarsal claw, female; E, apex of female abdomen, lateral view; F, ovipositor, lateral view; G, ventral parts of first to third annuli; H, apical part of lancet; I, K, male genitalia, dorsal and ventral views; L, penis valve, lateral view (left dorsal). A, B, D–F, H, Holotype (F, H, reversed).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883233/files/figure.png	https://doi.org/10.5281/zenodo.17883233	FIGURE 17. Phymatocera fuscata comb. nov. A, B, Female, laterodorsal and lateroventral views (head, dorsal view); C, D, male, laterodorsal and lateroventral views; E, posterior part of thorax and abdominal tergum 1, female, dorsal view; F, tarsal claw, female; G, ditto, male; H, posteroproximal part of fore wing, female; I, ditto, male; J, apex of abdomen, female, lateral views; K, male genitalia, ventral view (penis valves removed). A, B, E, F, H, J, Holotype.	FIGURE 17. Phymatocera fuscata comb. nov. A, B, Female, laterodorsal and lateroventral views (head, dorsal view); C, D, male, laterodorsal and lateroventral views; E, posterior part of thorax and abdominal tergum 1, female, dorsal view; F, tarsal claw, female; G, ditto, male; H, posteroproximal part of fore wing, female; I, ditto, male; J, apex of abdomen, female, lateral views; K, male genitalia, ventral view (penis valves removed). A, B, E, F, H, J, Holotype.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883235/files/figure.png	https://doi.org/10.5281/zenodo.17883235	FIGURE 18. Phymatocera peregrinator comb. nov. A, B, C, Female, laterodorsal, lateral and laterodorsal views; D, male, laterodorsal view; E, tarsal claw, female; F, ditto, male; G, metathorax and abdominal tergum 1, female, dorsal view; H, apex of abdomen, female, lateral view; I, ovipositor; J, middle of lancet; K, ventral parts of basal annuli; L, apex of lancet; M, N, male genitalia, dorsal and ventral views (N, penis valves removed); O, penis valve (left dorsal).A, B, G, Lectotype of Neotomostethus peregrinator, photographed by H. Vårdal. C, Holotype of Phymatocera hokkaidonis.	FIGURE 18. Phymatocera peregrinator comb. nov. A, B, C, Female, laterodorsal, lateral and laterodorsal views; D, male, laterodorsal view; E, tarsal claw, female; F, ditto, male; G, metathorax and abdominal tergum 1, female, dorsal view; H, apex of abdomen, female, lateral view; I, ovipositor; J, middle of lancet; K, ventral parts of basal annuli; L, apex of lancet; M, N, male genitalia, dorsal and ventral views (N, penis valves removed); O, penis valve (left dorsal).A, B, G, Lectotype of Neotomostethus peregrinator, photographed by H. Vårdal. C, Holotype of Phymatocera hokkaidonis.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883211/files/figure.png	https://doi.org/10.5281/zenodo.17883211	FIGURE 10. A, B, Frons, dorsoanterior view; C, D, anterodorsal part of mesepisternum, anteroventral view; E–H, antenna, lateral or medial view. A, C, E, F, Phymatocera fuscata comb. nov.: A, C, E, female (holotype); F, male. B, D, G, H, P. peregrinator comb. nov.: B, D, G, female (B, lectotype); H, male. B, Photographed by H. Vårdal.	FIGURE 10. A, B, Frons, dorsoanterior view; C, D, anterodorsal part of mesepisternum, anteroventral view; E–H, antenna, lateral or medial view. A, C, E, F, Phymatocera fuscata comb. nov.: A, C, E, female (holotype); F, male. B, D, G, H, P. peregrinator comb. nov.: B, D, G, female (B, lectotype); H, male. B, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883201/files/figure.png	https://doi.org/10.5281/zenodo.17883201	FIGURE 6.A–D, Mesoscutellum, laterodorsal view.E–H, Mesonotum, laterodorsal view (arrow indicating middle of notaulus). I, Head anterolateral view.A, E, Phymatoera satoi sp. nov., female. B, F, Paracharactus leucopodus, female. C, G, Nesotomostethus religiosa, female. D, H, Masaakia longivaginata, female. I, Phymatoera nipponica, female (antenna removed).	FIGURE 6.A–D, Mesoscutellum, laterodorsal view.E–H, Mesonotum, laterodorsal view (arrow indicating middle of notaulus). I, Head anterolateral view.A, E, Phymatoera satoi sp. nov., female. B, F, Paracharactus leucopodus, female. C, G, Nesotomostethus religiosa, female. D, H, Masaakia longivaginata, female. I, Phymatoera nipponica, female (antenna removed).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883179/files/figure.png	https://doi.org/10.5281/zenodo.17883179	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883191/files/figure.png	https://doi.org/10.5281/zenodo.17883191	FIGURE 4. A–E, Fore wing (c, cell; cv, crossvein; v, vein). G–I, Anterior part of thorax, anteroventral view. A, Nipponocampa esakii, female. B, I, Tomostethus nigritus, female. C, Apareophora japonica, female. D, G, Phymatoera nipponica, male. E, Phymatoceriola nipponensis, female. F, H, Hoplocampoides longiserrus, female (holotype).	FIGURE 4. A–E, Fore wing (c, cell; cv, crossvein; v, vein). G–I, Anterior part of thorax, anteroventral view. A, Nipponocampa esakii, female. B, I, Tomostethus nigritus, female. C, Apareophora japonica, female. D, G, Phymatoera nipponica, male. E, Phymatoceriola nipponensis, female. F, H, Hoplocampoides longiserrus, female (holotype).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17884946/files/figure.png	https://doi.org/10.5281/zenodo.17884946	FIGURE 11. A, B, Fore wing (c = cell, v = vein); C–E, male genitalia, lateroventral view; F–H, ventral parts of fifth and sixth annuli of lancet. A, C, F, Phymatocera nipponica. B, E, H, P. satoi sp. nov. D, G, P. membra.	FIGURE 11. A, B, Fore wing (c = cell, v = vein); C–E, male genitalia, lateroventral view; F–H, ventral parts of fifth and sixth annuli of lancet. A, C, F, Phymatocera nipponica. B, E, H, P. satoi sp. nov. D, G, P. membra.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFEFFA3FF7B9E8CFE26EE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883223/files/figure.png	https://doi.org/10.5281/zenodo.17883223	FIGURE 15. Phymatocera nipponica. A, Middle instar larva, 6 June 2019; B, last feeding instar larva (penultimate instar larva), 7 June 2019; C, mature larva (ultimate instar larva), 18 June 2019. Photographed by S. Ibuki.	FIGURE 15. Phymatocera nipponica. A, Middle instar larva, 6 June 2019; B, last feeding instar larva (penultimate instar larva), 7 June 2019; C, mature larva (ultimate instar larva), 18 June 2019. Photographed by S. Ibuki.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFAFFA1FF7B9DEDFA05E8E9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883186/files/figure.png	https://doi.org/10.5281/zenodo.17883186	FIGURE 3. A, C, D, Hind tarsus. B, Middle tarsus.A, Tomostethus nigritus, female. B, C, Onychostethomostus gilvipes, female (holotype) (C, reversed). D, Esehabachia luteipes, female (holotype).	FIGURE 3. A, C, D, Hind tarsus. B, Middle tarsus.A, Tomostethus nigritus, female. B, C, Onychostethomostus gilvipes, female (holotype) (C, reversed). D, Esehabachia luteipes, female (holotype).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFAFFA1FF7B9DEDFA05E8E9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883191/files/figure.png	https://doi.org/10.5281/zenodo.17883191	FIGURE 4. A–E, Fore wing (c, cell; cv, crossvein; v, vein). G–I, Anterior part of thorax, anteroventral view. A, Nipponocampa esakii, female. B, I, Tomostethus nigritus, female. C, Apareophora japonica, female. D, G, Phymatoera nipponica, male. E, Phymatoceriola nipponensis, female. F, H, Hoplocampoides longiserrus, female (holotype).	FIGURE 4. A–E, Fore wing (c, cell; cv, crossvein; v, vein). G–I, Anterior part of thorax, anteroventral view. A, Nipponocampa esakii, female. B, I, Tomostethus nigritus, female. C, Apareophora japonica, female. D, G, Phymatoera nipponica, male. E, Phymatoceriola nipponensis, female. F, H, Hoplocampoides longiserrus, female (holotype).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFFAFFA1FF7B9DEDFA05E8E9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883195/files/figure.png	https://doi.org/10.5281/zenodo.17883195	FIGURE 5. Dorsal tentorial pit and its neighboring areas, lateroanterior view. A, Phymatocera satoi sp. nov., female. B, Nesotomostethus religiosa, female. C, Phymatoceropsis japonica, female. D, Pareophora gracilis, female. E, Ardis pallipes, female. F, Periclista nigrifrons, female.	FIGURE 5. Dorsal tentorial pit and its neighboring areas, lateroanterior view. A, Phymatocera satoi sp. nov., female. B, Nesotomostethus religiosa, female. C, Phymatoceropsis japonica, female. D, Pareophora gracilis, female. E, Ardis pallipes, female. F, Periclista nigrifrons, female.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF8FFAEFF7B9B91FA05EC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883207/files/figure.png	https://doi.org/10.5281/zenodo.17883207	FIGURE 9. A, B, Head, lateral view; C, D, ventral part of head, ventrolateral view; E–H, flagellomere 1 of left antenna, dorsal view, female; I–L, ditto, male; M, N, male genitalia, ventral view (penis valves removed). A, C, F, J, M, Phymatocera peregrinator comb. nov.; A, C, F, female (F, holotype of P. hokkaidonis); J, M, male. B, H, K, P. nipponica: B, H, female; K, male. D, G, P. satoi, female (G, holotype). E, I, N, P. fuscata: E, female (holotype); I, N, male. L, P. sp.-1, male.	FIGURE 9. A, B, Head, lateral view; C, D, ventral part of head, ventrolateral view; E–H, flagellomere 1 of left antenna, dorsal view, female; I–L, ditto, male; M, N, male genitalia, ventral view (penis valves removed). A, C, F, J, M, Phymatocera peregrinator comb. nov.; A, C, F, female (F, holotype of P. hokkaidonis); J, M, male. B, H, K, P. nipponica: B, H, female; K, male. D, G, P. satoi, female (G, holotype). E, I, N, P. fuscata: E, female (holotype); I, N, male. L, P. sp.-1, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF8FFAEFF7B9B91FA05EC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883179/files/figure.png	https://doi.org/10.5281/zenodo.17883179	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF8FFAEFF7B9B91FA05EC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17884946/files/figure.png	https://doi.org/10.5281/zenodo.17884946	FIGURE 11. A, B, Fore wing (c = cell, v = vein); C–E, male genitalia, lateroventral view; F–H, ventral parts of fifth and sixth annuli of lancet. A, C, F, Phymatocera nipponica. B, E, H, P. satoi sp. nov. D, G, P. membra.	FIGURE 11. A, B, Fore wing (c = cell, v = vein); C–E, male genitalia, lateroventral view; F–H, ventral parts of fifth and sixth annuli of lancet. A, C, F, Phymatocera nipponica. B, E, H, P. satoi sp. nov. D, G, P. membra.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF8FFAEFF7B9B91FA05EC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883211/files/figure.png	https://doi.org/10.5281/zenodo.17883211	FIGURE 10. A, B, Frons, dorsoanterior view; C, D, anterodorsal part of mesepisternum, anteroventral view; E–H, antenna, lateral or medial view. A, C, E, F, Phymatocera fuscata comb. nov.: A, C, E, female (holotype); F, male. B, D, G, H, P. peregrinator comb. nov.: B, D, G, female (B, lectotype); H, male. B, Photographed by H. Vårdal.	FIGURE 10. A, B, Frons, dorsoanterior view; C, D, anterodorsal part of mesepisternum, anteroventral view; E–H, antenna, lateral or medial view. A, C, E, F, Phymatocera fuscata comb. nov.: A, C, E, female (holotype); F, male. B, D, G, H, P. peregrinator comb. nov.: B, D, G, female (B, lectotype); H, male. B, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF8FFAEFF7B9B91FA05EC6F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883215/files/figure.png	https://doi.org/10.5281/zenodo.17883215	FIGURE 12. A, B, Ocellar and frontal areas, anterodorsal view; C–F, abdominal terga 1–3, laterodorsal view. A, C, D, Phymatocera membra: A, C, female; D, male. B, E, F, P. nipponica: B, E, female; F, male.	FIGURE 12. A, B, Ocellar and frontal areas, anterodorsal view; C–F, abdominal terga 1–3, laterodorsal view. A, C, D, Phymatocera membra: A, C, female; D, male. B, E, F, P. nipponica: B, E, female; F, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF5FFABFF7B997DFEC8EA2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883205/files/figure.png	https://doi.org/10.5281/zenodo.17883205	FIGURE 8. A, B, Female abdomen, ventrolateral view. C–E, Head, lateral view. F, G, Basal part of male antenna, dorsal view (below lateral). A, D, Phymatoera nipponica, female. B, G, Phymatoceriola suigenensis: B, female (lectotype); G, male. C, Phymatocera fuscata, female (holotype). E, Phymatoceriola ussuriensis, male. F, Phymatoera membra, male.	FIGURE 8. A, B, Female abdomen, ventrolateral view. C–E, Head, lateral view. F, G, Basal part of male antenna, dorsal view (below lateral). A, D, Phymatoera nipponica, female. B, G, Phymatoceriola suigenensis: B, female (lectotype); G, male. C, Phymatocera fuscata, female (holotype). E, Phymatoceriola ussuriensis, male. F, Phymatoera membra, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF5FFABFF7B997DFEC8EA2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17884946/files/figure.png	https://doi.org/10.5281/zenodo.17884946	FIGURE 11. A, B, Fore wing (c = cell, v = vein); C–E, male genitalia, lateroventral view; F–H, ventral parts of fifth and sixth annuli of lancet. A, C, F, Phymatocera nipponica. B, E, H, P. satoi sp. nov. D, G, P. membra.	FIGURE 11. A, B, Fore wing (c = cell, v = vein); C–E, male genitalia, lateroventral view; F–H, ventral parts of fifth and sixth annuli of lancet. A, C, F, Phymatocera nipponica. B, E, H, P. satoi sp. nov. D, G, P. membra.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF5FFABFF7B997DFEC8EA2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883215/files/figure.png	https://doi.org/10.5281/zenodo.17883215	FIGURE 12. A, B, Ocellar and frontal areas, anterodorsal view; C–F, abdominal terga 1–3, laterodorsal view. A, C, D, Phymatocera membra: A, C, female; D, male. B, E, F, P. nipponica: B, E, female; F, male.	FIGURE 12. A, B, Ocellar and frontal areas, anterodorsal view; C–F, abdominal terga 1–3, laterodorsal view. A, C, D, Phymatocera membra: A, C, female; D, male. B, E, F, P. nipponica: B, E, female; F, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF5FFABFF7B997DFEC8EA2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883217/files/figure.png	https://doi.org/10.5281/zenodo.17883217	FIGURE 13. Phymatocera membra.A, Female, dorsal view; B, C, male, dorsal and lateroventral views; D, tarsal claw; E, apex of female abdomen, lateral view; F, ovipositor, lateral view; G, lancet; H, ventral parts of first to third annuli of lancet; I, apical part of lancet (arrows indicating some marginal sensilla); J, K, male genitalia, dorsal and ventral views; L, penis valves, lateral view (left dorsal).	FIGURE 13. Phymatocera membra.A, Female, dorsal view; B, C, male, dorsal and lateroventral views; D, tarsal claw; E, apex of female abdomen, lateral view; F, ovipositor, lateral view; G, lancet; H, ventral parts of first to third annuli of lancet; I, apical part of lancet (arrows indicating some marginal sensilla); J, K, male genitalia, dorsal and ventral views; L, penis valves, lateral view (left dorsal).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF5FFABFF7B997DFEC8EA2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883179/files/figure.png	https://doi.org/10.5281/zenodo.17883179	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883170/files/figure.png	https://doi.org/10.5281/zenodo.17883170	FIGURE 1. A, Head, anterolateral view; B, frons, dorsal view; C, frons, anterior; D, head, ventrolateral. A, B, C, Phymatocera peregrinator comb. nov., female (B, C, lectotype). D, P. nipponica, female. B, C, Photographed by H. Vårdal.	FIGURE 1. A, Head, anterolateral view; B, frons, dorsal view; C, frons, anterior; D, head, ventrolateral. A, B, C, Phymatocera peregrinator comb. nov., female (B, C, lectotype). D, P. nipponica, female. B, C, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883179/files/figure.png	https://doi.org/10.5281/zenodo.17883179	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883191/files/figure.png	https://doi.org/10.5281/zenodo.17883191	FIGURE 4. A–E, Fore wing (c, cell; cv, crossvein; v, vein). G–I, Anterior part of thorax, anteroventral view. A, Nipponocampa esakii, female. B, I, Tomostethus nigritus, female. C, Apareophora japonica, female. D, G, Phymatoera nipponica, male. E, Phymatoceriola nipponensis, female. F, H, Hoplocampoides longiserrus, female (holotype).	FIGURE 4. A–E, Fore wing (c, cell; cv, crossvein; v, vein). G–I, Anterior part of thorax, anteroventral view. A, Nipponocampa esakii, female. B, I, Tomostethus nigritus, female. C, Apareophora japonica, female. D, G, Phymatoera nipponica, male. E, Phymatoceriola nipponensis, female. F, H, Hoplocampoides longiserrus, female (holotype).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883201/files/figure.png	https://doi.org/10.5281/zenodo.17883201	FIGURE 6.A–D, Mesoscutellum, laterodorsal view.E–H, Mesonotum, laterodorsal view (arrow indicating middle of notaulus). I, Head anterolateral view.A, E, Phymatoera satoi sp. nov., female. B, F, Paracharactus leucopodus, female. C, G, Nesotomostethus religiosa, female. D, H, Masaakia longivaginata, female. I, Phymatoera nipponica, female (antenna removed).	FIGURE 6.A–D, Mesoscutellum, laterodorsal view.E–H, Mesonotum, laterodorsal view (arrow indicating middle of notaulus). I, Head anterolateral view.A, E, Phymatoera satoi sp. nov., female. B, F, Paracharactus leucopodus, female. C, G, Nesotomostethus religiosa, female. D, H, Masaakia longivaginata, female. I, Phymatoera nipponica, female (antenna removed).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883203/files/figure.png	https://doi.org/10.5281/zenodo.17883203	FIGURE 7.A–D, Tarsal claw.E, F, Head, ventrolateral view.A, Phymatoceriola nipponensis, female (holotype).B, Phymatocera peregrinator comb. nov., female. C, E, Phymatoera nipponica, female. D, F, Paracharactus leucopodus, female.	FIGURE 7.A–D, Tarsal claw.E, F, Head, ventrolateral view.A, Phymatoceriola nipponensis, female (holotype).B, Phymatocera peregrinator comb. nov., female. C, E, Phymatoera nipponica, female. D, F, Paracharactus leucopodus, female.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883205/files/figure.png	https://doi.org/10.5281/zenodo.17883205	FIGURE 8. A, B, Female abdomen, ventrolateral view. C–E, Head, lateral view. F, G, Basal part of male antenna, dorsal view (below lateral). A, D, Phymatoera nipponica, female. B, G, Phymatoceriola suigenensis: B, female (lectotype); G, male. C, Phymatocera fuscata, female (holotype). E, Phymatoceriola ussuriensis, male. F, Phymatoera membra, male.	FIGURE 8. A, B, Female abdomen, ventrolateral view. C–E, Head, lateral view. F, G, Basal part of male antenna, dorsal view (below lateral). A, D, Phymatoera nipponica, female. B, G, Phymatoceriola suigenensis: B, female (lectotype); G, male. C, Phymatocera fuscata, female (holotype). E, Phymatoceriola ussuriensis, male. F, Phymatoera membra, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883207/files/figure.png	https://doi.org/10.5281/zenodo.17883207	FIGURE 9. A, B, Head, lateral view; C, D, ventral part of head, ventrolateral view; E–H, flagellomere 1 of left antenna, dorsal view, female; I–L, ditto, male; M, N, male genitalia, ventral view (penis valves removed). A, C, F, J, M, Phymatocera peregrinator comb. nov.; A, C, F, female (F, holotype of P. hokkaidonis); J, M, male. B, H, K, P. nipponica: B, H, female; K, male. D, G, P. satoi, female (G, holotype). E, I, N, P. fuscata: E, female (holotype); I, N, male. L, P. sp.-1, male.	FIGURE 9. A, B, Head, lateral view; C, D, ventral part of head, ventrolateral view; E–H, flagellomere 1 of left antenna, dorsal view, female; I–L, ditto, male; M, N, male genitalia, ventral view (penis valves removed). A, C, F, J, M, Phymatocera peregrinator comb. nov.; A, C, F, female (F, holotype of P. hokkaidonis); J, M, male. B, H, K, P. nipponica: B, H, female; K, male. D, G, P. satoi, female (G, holotype). E, I, N, P. fuscata: E, female (holotype); I, N, male. L, P. sp.-1, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17884946/files/figure.png	https://doi.org/10.5281/zenodo.17884946	FIGURE 11. A, B, Fore wing (c = cell, v = vein); C–E, male genitalia, lateroventral view; F–H, ventral parts of fifth and sixth annuli of lancet. A, C, F, Phymatocera nipponica. B, E, H, P. satoi sp. nov. D, G, P. membra.	FIGURE 11. A, B, Fore wing (c = cell, v = vein); C–E, male genitalia, lateroventral view; F–H, ventral parts of fifth and sixth annuli of lancet. A, C, F, Phymatocera nipponica. B, E, H, P. satoi sp. nov. D, G, P. membra.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883215/files/figure.png	https://doi.org/10.5281/zenodo.17883215	FIGURE 12. A, B, Ocellar and frontal areas, anterodorsal view; C–F, abdominal terga 1–3, laterodorsal view. A, C, D, Phymatocera membra: A, C, female; D, male. B, E, F, P. nipponica: B, E, female; F, male.	FIGURE 12. A, B, Ocellar and frontal areas, anterodorsal view; C–F, abdominal terga 1–3, laterodorsal view. A, C, D, Phymatocera membra: A, C, female; D, male. B, E, F, P. nipponica: B, E, female; F, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883219/files/figure.png	https://doi.org/10.5281/zenodo.17883219	FIGURE 14. Phymatocera nipponica. A, B, Female, dorsal and lateral views; C, male, dorsal view; D, tarsal claw, female; E, apex of female abdomen, lateral view; F, ovipositor; G, ventral parts of first to fourth annuli of lancet; H–J, apical part of lancet; K, male genitalia, dorsal view (penis valves removed); L, ditto (penis valves not removed), ventral view; M, penis valve, lateral view (left dorsal). A, B, Photographed just after killing.	FIGURE 14. Phymatocera nipponica. A, B, Female, dorsal and lateral views; C, male, dorsal view; D, tarsal claw, female; E, apex of female abdomen, lateral view; F, ovipositor; G, ventral parts of first to fourth annuli of lancet; H–J, apical part of lancet; K, male genitalia, dorsal view (penis valves removed); L, ditto (penis valves not removed), ventral view; M, penis valve, lateral view (left dorsal). A, B, Photographed just after killing.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883223/files/figure.png	https://doi.org/10.5281/zenodo.17883223	FIGURE 15. Phymatocera nipponica. A, Middle instar larva, 6 June 2019; B, last feeding instar larva (penultimate instar larva), 7 June 2019; C, mature larva (ultimate instar larva), 18 June 2019. Photographed by S. Ibuki.	FIGURE 15. Phymatocera nipponica. A, Middle instar larva, 6 June 2019; B, last feeding instar larva (penultimate instar larva), 7 June 2019; C, mature larva (ultimate instar larva), 18 June 2019. Photographed by S. Ibuki.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883229/files/figure.png	https://doi.org/10.5281/zenodo.17883229	FIGURE 16. Phymatocera satoi sp. nov. A, B, Female, dorsal and lateral views; C, male, dorsal view; D, tarsal claw, female; E, apex of female abdomen, lateral view; F, ovipositor, lateral view; G, ventral parts of first to third annuli; H, apical part of lancet; I, K, male genitalia, dorsal and ventral views; L, penis valve, lateral view (left dorsal). A, B, D–F, H, Holotype (F, H, reversed).	FIGURE 16. Phymatocera satoi sp. nov. A, B, Female, dorsal and lateral views; C, male, dorsal view; D, tarsal claw, female; E, apex of female abdomen, lateral view; F, ovipositor, lateral view; G, ventral parts of first to third annuli; H, apical part of lancet; I, K, male genitalia, dorsal and ventral views; L, penis valve, lateral view (left dorsal). A, B, D–F, H, Holotype (F, H, reversed).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFF3FFB6FF7B9EB0FEBEE81C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883217/files/figure.png	https://doi.org/10.5281/zenodo.17883217	FIGURE 13. Phymatocera membra.A, Female, dorsal view; B, C, male, dorsal and lateroventral views; D, tarsal claw; E, apex of female abdomen, lateral view; F, ovipositor, lateral view; G, lancet; H, ventral parts of first to third annuli of lancet; I, apical part of lancet (arrows indicating some marginal sensilla); J, K, male genitalia, dorsal and ventral views; L, penis valves, lateral view (left dorsal).	FIGURE 13. Phymatocera membra.A, Female, dorsal view; B, C, male, dorsal and lateroventral views; D, tarsal claw; E, apex of female abdomen, lateral view; F, ovipositor, lateral view; G, lancet; H, ventral parts of first to third annuli of lancet; I, apical part of lancet (arrows indicating some marginal sensilla); J, K, male genitalia, dorsal and ventral views; L, penis valves, lateral view (left dorsal).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEEFFB4FF7B9CE8FBC7EC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883179/files/figure.png	https://doi.org/10.5281/zenodo.17883179	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEEFFB4FF7B9CE8FBC7EC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883195/files/figure.png	https://doi.org/10.5281/zenodo.17883195	FIGURE 5. Dorsal tentorial pit and its neighboring areas, lateroanterior view. A, Phymatocera satoi sp. nov., female. B, Nesotomostethus religiosa, female. C, Phymatoceropsis japonica, female. D, Pareophora gracilis, female. E, Ardis pallipes, female. F, Periclista nigrifrons, female.	FIGURE 5. Dorsal tentorial pit and its neighboring areas, lateroanterior view. A, Phymatocera satoi sp. nov., female. B, Nesotomostethus religiosa, female. C, Phymatoceropsis japonica, female. D, Pareophora gracilis, female. E, Ardis pallipes, female. F, Periclista nigrifrons, female.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEEFFB4FF7B9CE8FBC7EC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883201/files/figure.png	https://doi.org/10.5281/zenodo.17883201	FIGURE 6.A–D, Mesoscutellum, laterodorsal view.E–H, Mesonotum, laterodorsal view (arrow indicating middle of notaulus). I, Head anterolateral view.A, E, Phymatoera satoi sp. nov., female. B, F, Paracharactus leucopodus, female. C, G, Nesotomostethus religiosa, female. D, H, Masaakia longivaginata, female. I, Phymatoera nipponica, female (antenna removed).	FIGURE 6.A–D, Mesoscutellum, laterodorsal view.E–H, Mesonotum, laterodorsal view (arrow indicating middle of notaulus). I, Head anterolateral view.A, E, Phymatoera satoi sp. nov., female. B, F, Paracharactus leucopodus, female. C, G, Nesotomostethus religiosa, female. D, H, Masaakia longivaginata, female. I, Phymatoera nipponica, female (antenna removed).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEEFFB4FF7B9CE8FBC7EC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883207/files/figure.png	https://doi.org/10.5281/zenodo.17883207	FIGURE 9. A, B, Head, lateral view; C, D, ventral part of head, ventrolateral view; E–H, flagellomere 1 of left antenna, dorsal view, female; I–L, ditto, male; M, N, male genitalia, ventral view (penis valves removed). A, C, F, J, M, Phymatocera peregrinator comb. nov.; A, C, F, female (F, holotype of P. hokkaidonis); J, M, male. B, H, K, P. nipponica: B, H, female; K, male. D, G, P. satoi, female (G, holotype). E, I, N, P. fuscata: E, female (holotype); I, N, male. L, P. sp.-1, male.	FIGURE 9. A, B, Head, lateral view; C, D, ventral part of head, ventrolateral view; E–H, flagellomere 1 of left antenna, dorsal view, female; I–L, ditto, male; M, N, male genitalia, ventral view (penis valves removed). A, C, F, J, M, Phymatocera peregrinator comb. nov.; A, C, F, female (F, holotype of P. hokkaidonis); J, M, male. B, H, K, P. nipponica: B, H, female; K, male. D, G, P. satoi, female (G, holotype). E, I, N, P. fuscata: E, female (holotype); I, N, male. L, P. sp.-1, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEEFFB4FF7B9CE8FBC7EC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17884946/files/figure.png	https://doi.org/10.5281/zenodo.17884946	FIGURE 11. A, B, Fore wing (c = cell, v = vein); C–E, male genitalia, lateroventral view; F–H, ventral parts of fifth and sixth annuli of lancet. A, C, F, Phymatocera nipponica. B, E, H, P. satoi sp. nov. D, G, P. membra.	FIGURE 11. A, B, Fore wing (c = cell, v = vein); C–E, male genitalia, lateroventral view; F–H, ventral parts of fifth and sixth annuli of lancet. A, C, F, Phymatocera nipponica. B, E, H, P. satoi sp. nov. D, G, P. membra.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEEFFB4FF7B9CE8FBC7EC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883229/files/figure.png	https://doi.org/10.5281/zenodo.17883229	FIGURE 16. Phymatocera satoi sp. nov. A, B, Female, dorsal and lateral views; C, male, dorsal view; D, tarsal claw, female; E, apex of female abdomen, lateral view; F, ovipositor, lateral view; G, ventral parts of first to third annuli; H, apical part of lancet; I, K, male genitalia, dorsal and ventral views; L, penis valve, lateral view (left dorsal). A, B, D–F, H, Holotype (F, H, reversed).	FIGURE 16. Phymatocera satoi sp. nov. A, B, Female, dorsal and lateral views; C, male, dorsal view; D, tarsal claw, female; E, apex of female abdomen, lateral view; F, ovipositor, lateral view; G, ventral parts of first to third annuli; H, apical part of lancet; I, K, male genitalia, dorsal and ventral views; L, penis valve, lateral view (left dorsal). A, B, D–F, H, Holotype (F, H, reversed).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEEFFB4FF7B9CE8FBC7EC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883205/files/figure.png	https://doi.org/10.5281/zenodo.17883205	FIGURE 8. A, B, Female abdomen, ventrolateral view. C–E, Head, lateral view. F, G, Basal part of male antenna, dorsal view (below lateral). A, D, Phymatoera nipponica, female. B, G, Phymatoceriola suigenensis: B, female (lectotype); G, male. C, Phymatocera fuscata, female (holotype). E, Phymatoceriola ussuriensis, male. F, Phymatoera membra, male.	FIGURE 8. A, B, Female abdomen, ventrolateral view. C–E, Head, lateral view. F, G, Basal part of male antenna, dorsal view (below lateral). A, D, Phymatoera nipponica, female. B, G, Phymatoceriola suigenensis: B, female (lectotype); G, male. C, Phymatocera fuscata, female (holotype). E, Phymatoceriola ussuriensis, male. F, Phymatoera membra, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEEFFB4FF7B9CE8FBC7EC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883233/files/figure.png	https://doi.org/10.5281/zenodo.17883233	FIGURE 17. Phymatocera fuscata comb. nov. A, B, Female, laterodorsal and lateroventral views (head, dorsal view); C, D, male, laterodorsal and lateroventral views; E, posterior part of thorax and abdominal tergum 1, female, dorsal view; F, tarsal claw, female; G, ditto, male; H, posteroproximal part of fore wing, female; I, ditto, male; J, apex of abdomen, female, lateral views; K, male genitalia, ventral view (penis valves removed). A, B, E, F, H, J, Holotype.	FIGURE 17. Phymatocera fuscata comb. nov. A, B, Female, laterodorsal and lateroventral views (head, dorsal view); C, D, male, laterodorsal and lateroventral views; E, posterior part of thorax and abdominal tergum 1, female, dorsal view; F, tarsal claw, female; G, ditto, male; H, posteroproximal part of fore wing, female; I, ditto, male; J, apex of abdomen, female, lateral views; K, male genitalia, ventral view (penis valves removed). A, B, E, F, H, J, Holotype.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEEFFB4FF7B9CE8FBC7EC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883235/files/figure.png	https://doi.org/10.5281/zenodo.17883235	FIGURE 18. Phymatocera peregrinator comb. nov. A, B, C, Female, laterodorsal, lateral and laterodorsal views; D, male, laterodorsal view; E, tarsal claw, female; F, ditto, male; G, metathorax and abdominal tergum 1, female, dorsal view; H, apex of abdomen, female, lateral view; I, ovipositor; J, middle of lancet; K, ventral parts of basal annuli; L, apex of lancet; M, N, male genitalia, dorsal and ventral views (N, penis valves removed); O, penis valve (left dorsal).A, B, G, Lectotype of Neotomostethus peregrinator, photographed by H. Vårdal. C, Holotype of Phymatocera hokkaidonis.	FIGURE 18. Phymatocera peregrinator comb. nov. A, B, C, Female, laterodorsal, lateral and laterodorsal views; D, male, laterodorsal view; E, tarsal claw, female; F, ditto, male; G, metathorax and abdominal tergum 1, female, dorsal view; H, apex of abdomen, female, lateral view; I, ovipositor; J, middle of lancet; K, ventral parts of basal annuli; L, apex of lancet; M, N, male genitalia, dorsal and ventral views (N, penis valves removed); O, penis valve (left dorsal).A, B, G, Lectotype of Neotomostethus peregrinator, photographed by H. Vårdal. C, Holotype of Phymatocera hokkaidonis.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEEFFB4FF7B9CE8FBC7EC24.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883211/files/figure.png	https://doi.org/10.5281/zenodo.17883211	FIGURE 10. A, B, Frons, dorsoanterior view; C, D, anterodorsal part of mesepisternum, anteroventral view; E–H, antenna, lateral or medial view. A, C, E, F, Phymatocera fuscata comb. nov.: A, C, E, female (holotype); F, male. B, D, G, H, P. peregrinator comb. nov.: B, D, G, female (B, lectotype); H, male. B, Photographed by H. Vårdal.	FIGURE 10. A, B, Frons, dorsoanterior view; C, D, anterodorsal part of mesepisternum, anteroventral view; E–H, antenna, lateral or medial view. A, C, E, F, Phymatocera fuscata comb. nov.: A, C, E, female (holotype); F, male. B, D, G, H, P. peregrinator comb. nov.: B, D, G, female (B, lectotype); H, male. B, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFECFFB2FF7B98B0FE60EAF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883205/files/figure.png	https://doi.org/10.5281/zenodo.17883205	FIGURE 8. A, B, Female abdomen, ventrolateral view. C–E, Head, lateral view. F, G, Basal part of male antenna, dorsal view (below lateral). A, D, Phymatoera nipponica, female. B, G, Phymatoceriola suigenensis: B, female (lectotype); G, male. C, Phymatocera fuscata, female (holotype). E, Phymatoceriola ussuriensis, male. F, Phymatoera membra, male.	FIGURE 8. A, B, Female abdomen, ventrolateral view. C–E, Head, lateral view. F, G, Basal part of male antenna, dorsal view (below lateral). A, D, Phymatoera nipponica, female. B, G, Phymatoceriola suigenensis: B, female (lectotype); G, male. C, Phymatocera fuscata, female (holotype). E, Phymatoceriola ussuriensis, male. F, Phymatoera membra, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFECFFB2FF7B98B0FE60EAF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883207/files/figure.png	https://doi.org/10.5281/zenodo.17883207	FIGURE 9. A, B, Head, lateral view; C, D, ventral part of head, ventrolateral view; E–H, flagellomere 1 of left antenna, dorsal view, female; I–L, ditto, male; M, N, male genitalia, ventral view (penis valves removed). A, C, F, J, M, Phymatocera peregrinator comb. nov.; A, C, F, female (F, holotype of P. hokkaidonis); J, M, male. B, H, K, P. nipponica: B, H, female; K, male. D, G, P. satoi, female (G, holotype). E, I, N, P. fuscata: E, female (holotype); I, N, male. L, P. sp.-1, male.	FIGURE 9. A, B, Head, lateral view; C, D, ventral part of head, ventrolateral view; E–H, flagellomere 1 of left antenna, dorsal view, female; I–L, ditto, male; M, N, male genitalia, ventral view (penis valves removed). A, C, F, J, M, Phymatocera peregrinator comb. nov.; A, C, F, female (F, holotype of P. hokkaidonis); J, M, male. B, H, K, P. nipponica: B, H, female; K, male. D, G, P. satoi, female (G, holotype). E, I, N, P. fuscata: E, female (holotype); I, N, male. L, P. sp.-1, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFECFFB2FF7B98B0FE60EAF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883211/files/figure.png	https://doi.org/10.5281/zenodo.17883211	FIGURE 10. A, B, Frons, dorsoanterior view; C, D, anterodorsal part of mesepisternum, anteroventral view; E–H, antenna, lateral or medial view. A, C, E, F, Phymatocera fuscata comb. nov.: A, C, E, female (holotype); F, male. B, D, G, H, P. peregrinator comb. nov.: B, D, G, female (B, lectotype); H, male. B, Photographed by H. Vårdal.	FIGURE 10. A, B, Frons, dorsoanterior view; C, D, anterodorsal part of mesepisternum, anteroventral view; E–H, antenna, lateral or medial view. A, C, E, F, Phymatocera fuscata comb. nov.: A, C, E, female (holotype); F, male. B, D, G, H, P. peregrinator comb. nov.: B, D, G, female (B, lectotype); H, male. B, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFECFFB2FF7B98B0FE60EAF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883233/files/figure.png	https://doi.org/10.5281/zenodo.17883233	FIGURE 17. Phymatocera fuscata comb. nov. A, B, Female, laterodorsal and lateroventral views (head, dorsal view); C, D, male, laterodorsal and lateroventral views; E, posterior part of thorax and abdominal tergum 1, female, dorsal view; F, tarsal claw, female; G, ditto, male; H, posteroproximal part of fore wing, female; I, ditto, male; J, apex of abdomen, female, lateral views; K, male genitalia, ventral view (penis valves removed). A, B, E, F, H, J, Holotype.	FIGURE 17. Phymatocera fuscata comb. nov. A, B, Female, laterodorsal and lateroventral views (head, dorsal view); C, D, male, laterodorsal and lateroventral views; E, posterior part of thorax and abdominal tergum 1, female, dorsal view; F, tarsal claw, female; G, ditto, male; H, posteroproximal part of fore wing, female; I, ditto, male; J, apex of abdomen, female, lateral views; K, male genitalia, ventral view (penis valves removed). A, B, E, F, H, J, Holotype.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEAFFBFFF7B9E8CFC77ED4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883170/files/figure.png	https://doi.org/10.5281/zenodo.17883170	FIGURE 1. A, Head, anterolateral view; B, frons, dorsal view; C, frons, anterior; D, head, ventrolateral. A, B, C, Phymatocera peregrinator comb. nov., female (B, C, lectotype). D, P. nipponica, female. B, C, Photographed by H. Vårdal.	FIGURE 1. A, Head, anterolateral view; B, frons, dorsal view; C, frons, anterior; D, head, ventrolateral. A, B, C, Phymatocera peregrinator comb. nov., female (B, C, lectotype). D, P. nipponica, female. B, C, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEAFFBFFF7B9E8CFC77ED4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883179/files/figure.png	https://doi.org/10.5281/zenodo.17883179	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEAFFBFFF7B9E8CFC77ED4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883203/files/figure.png	https://doi.org/10.5281/zenodo.17883203	FIGURE 7.A–D, Tarsal claw.E, F, Head, ventrolateral view.A, Phymatoceriola nipponensis, female (holotype).B, Phymatocera peregrinator comb. nov., female. C, E, Phymatoera nipponica, female. D, F, Paracharactus leucopodus, female.	FIGURE 7.A–D, Tarsal claw.E, F, Head, ventrolateral view.A, Phymatoceriola nipponensis, female (holotype).B, Phymatocera peregrinator comb. nov., female. C, E, Phymatoera nipponica, female. D, F, Paracharactus leucopodus, female.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEAFFBFFF7B9E8CFC77ED4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883207/files/figure.png	https://doi.org/10.5281/zenodo.17883207	FIGURE 9. A, B, Head, lateral view; C, D, ventral part of head, ventrolateral view; E–H, flagellomere 1 of left antenna, dorsal view, female; I–L, ditto, male; M, N, male genitalia, ventral view (penis valves removed). A, C, F, J, M, Phymatocera peregrinator comb. nov.; A, C, F, female (F, holotype of P. hokkaidonis); J, M, male. B, H, K, P. nipponica: B, H, female; K, male. D, G, P. satoi, female (G, holotype). E, I, N, P. fuscata: E, female (holotype); I, N, male. L, P. sp.-1, male.	FIGURE 9. A, B, Head, lateral view; C, D, ventral part of head, ventrolateral view; E–H, flagellomere 1 of left antenna, dorsal view, female; I–L, ditto, male; M, N, male genitalia, ventral view (penis valves removed). A, C, F, J, M, Phymatocera peregrinator comb. nov.; A, C, F, female (F, holotype of P. hokkaidonis); J, M, male. B, H, K, P. nipponica: B, H, female; K, male. D, G, P. satoi, female (G, holotype). E, I, N, P. fuscata: E, female (holotype); I, N, male. L, P. sp.-1, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEAFFBFFF7B9E8CFC77ED4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883211/files/figure.png	https://doi.org/10.5281/zenodo.17883211	FIGURE 10. A, B, Frons, dorsoanterior view; C, D, anterodorsal part of mesepisternum, anteroventral view; E–H, antenna, lateral or medial view. A, C, E, F, Phymatocera fuscata comb. nov.: A, C, E, female (holotype); F, male. B, D, G, H, P. peregrinator comb. nov.: B, D, G, female (B, lectotype); H, male. B, Photographed by H. Vårdal.	FIGURE 10. A, B, Frons, dorsoanterior view; C, D, anterodorsal part of mesepisternum, anteroventral view; E–H, antenna, lateral or medial view. A, C, E, F, Phymatocera fuscata comb. nov.: A, C, E, female (holotype); F, male. B, D, G, H, P. peregrinator comb. nov.: B, D, G, female (B, lectotype); H, male. B, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFEAFFBFFF7B9E8CFC77ED4C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883235/files/figure.png	https://doi.org/10.5281/zenodo.17883235	FIGURE 18. Phymatocera peregrinator comb. nov. A, B, C, Female, laterodorsal, lateral and laterodorsal views; D, male, laterodorsal view; E, tarsal claw, female; F, ditto, male; G, metathorax and abdominal tergum 1, female, dorsal view; H, apex of abdomen, female, lateral view; I, ovipositor; J, middle of lancet; K, ventral parts of basal annuli; L, apex of lancet; M, N, male genitalia, dorsal and ventral views (N, penis valves removed); O, penis valve (left dorsal).A, B, G, Lectotype of Neotomostethus peregrinator, photographed by H. Vårdal. C, Holotype of Phymatocera hokkaidonis.	FIGURE 18. Phymatocera peregrinator comb. nov. A, B, C, Female, laterodorsal, lateral and laterodorsal views; D, male, laterodorsal view; E, tarsal claw, female; F, ditto, male; G, metathorax and abdominal tergum 1, female, dorsal view; H, apex of abdomen, female, lateral view; I, ovipositor; J, middle of lancet; K, ventral parts of basal annuli; L, apex of lancet; M, N, male genitalia, dorsal and ventral views (N, penis valves removed); O, penis valve (left dorsal).A, B, G, Lectotype of Neotomostethus peregrinator, photographed by H. Vårdal. C, Holotype of Phymatocera hokkaidonis.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFE3FFBBFF7B9961FC3CEE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883191/files/figure.png	https://doi.org/10.5281/zenodo.17883191	FIGURE 4. A–E, Fore wing (c, cell; cv, crossvein; v, vein). G–I, Anterior part of thorax, anteroventral view. A, Nipponocampa esakii, female. B, I, Tomostethus nigritus, female. C, Apareophora japonica, female. D, G, Phymatoera nipponica, male. E, Phymatoceriola nipponensis, female. F, H, Hoplocampoides longiserrus, female (holotype).	FIGURE 4. A–E, Fore wing (c, cell; cv, crossvein; v, vein). G–I, Anterior part of thorax, anteroventral view. A, Nipponocampa esakii, female. B, I, Tomostethus nigritus, female. C, Apareophora japonica, female. D, G, Phymatoera nipponica, male. E, Phymatoceriola nipponensis, female. F, H, Hoplocampoides longiserrus, female (holotype).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFE3FFBBFF7B9961FC3CEE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883195/files/figure.png	https://doi.org/10.5281/zenodo.17883195	FIGURE 5. Dorsal tentorial pit and its neighboring areas, lateroanterior view. A, Phymatocera satoi sp. nov., female. B, Nesotomostethus religiosa, female. C, Phymatoceropsis japonica, female. D, Pareophora gracilis, female. E, Ardis pallipes, female. F, Periclista nigrifrons, female.	FIGURE 5. Dorsal tentorial pit and its neighboring areas, lateroanterior view. A, Phymatocera satoi sp. nov., female. B, Nesotomostethus religiosa, female. C, Phymatoceropsis japonica, female. D, Pareophora gracilis, female. E, Ardis pallipes, female. F, Periclista nigrifrons, female.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFE3FFBBFF7B9961FC3CEE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883186/files/figure.png	https://doi.org/10.5281/zenodo.17883186	FIGURE 3. A, C, D, Hind tarsus. B, Middle tarsus.A, Tomostethus nigritus, female. B, C, Onychostethomostus gilvipes, female (holotype) (C, reversed). D, Esehabachia luteipes, female (holotype).	FIGURE 3. A, C, D, Hind tarsus. B, Middle tarsus.A, Tomostethus nigritus, female. B, C, Onychostethomostus gilvipes, female (holotype) (C, reversed). D, Esehabachia luteipes, female (holotype).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFE3FFBBFF7B9961FC3CEE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883201/files/figure.png	https://doi.org/10.5281/zenodo.17883201	FIGURE 6.A–D, Mesoscutellum, laterodorsal view.E–H, Mesonotum, laterodorsal view (arrow indicating middle of notaulus). I, Head anterolateral view.A, E, Phymatoera satoi sp. nov., female. B, F, Paracharactus leucopodus, female. C, G, Nesotomostethus religiosa, female. D, H, Masaakia longivaginata, female. I, Phymatoera nipponica, female (antenna removed).	FIGURE 6.A–D, Mesoscutellum, laterodorsal view.E–H, Mesonotum, laterodorsal view (arrow indicating middle of notaulus). I, Head anterolateral view.A, E, Phymatoera satoi sp. nov., female. B, F, Paracharactus leucopodus, female. C, G, Nesotomostethus religiosa, female. D, H, Masaakia longivaginata, female. I, Phymatoera nipponica, female (antenna removed).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFE3FFBBFF7B9961FC3CEE80.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883203/files/figure.png	https://doi.org/10.5281/zenodo.17883203	FIGURE 7.A–D, Tarsal claw.E, F, Head, ventrolateral view.A, Phymatoceriola nipponensis, female (holotype).B, Phymatocera peregrinator comb. nov., female. C, E, Phymatoera nipponica, female. D, F, Paracharactus leucopodus, female.	FIGURE 7.A–D, Tarsal claw.E, F, Head, ventrolateral view.A, Phymatoceriola nipponensis, female (holotype).B, Phymatocera peregrinator comb. nov., female. C, E, Phymatoera nipponica, female. D, F, Paracharactus leucopodus, female.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFE3FFBBFF7B9DA5FB24EADC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883179/files/figure.png	https://doi.org/10.5281/zenodo.17883179	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	FIGURE 2. A, Thorax, laterodorsal view; B, anterior part of mesepisternum, lateroventral view; C, metathorax, lateral view; D, anterior apical spur of fore tibia, anterior view; E, hind tarsus, anterior view; F, tarsal claw; G, fore wing (c = cell, v = vein); H, I, posteroproximal part of fore wing; J, female hind wing (cv = crossvein); K, female abdomen, ventrolateral view; L, second to fourth annuli of lancet; M, N, male genitalia, dorsal and ventral views. A, C, G, J, Phymatocera peregrinator comb. nov., female (A, lectotype). B, E, H, I, K, L, P. nipponica, female. D, P. sp.-1, male. F, P. satoi sp. nov. female. M, N, P. aterrima, male. A, Photographed by H. Vårdal.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFE2FFBAFF7B9DECFC08EC6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883195/files/figure.png	https://doi.org/10.5281/zenodo.17883195	FIGURE 5. Dorsal tentorial pit and its neighboring areas, lateroanterior view. A, Phymatocera satoi sp. nov., female. B, Nesotomostethus religiosa, female. C, Phymatoceropsis japonica, female. D, Pareophora gracilis, female. E, Ardis pallipes, female. F, Periclista nigrifrons, female.	FIGURE 5. Dorsal tentorial pit and its neighboring areas, lateroanterior view. A, Phymatocera satoi sp. nov., female. B, Nesotomostethus religiosa, female. C, Phymatoceropsis japonica, female. D, Pareophora gracilis, female. E, Ardis pallipes, female. F, Periclista nigrifrons, female.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFE2FFBAFF7B9DECFC08EC6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883191/files/figure.png	https://doi.org/10.5281/zenodo.17883191	FIGURE 4. A–E, Fore wing (c, cell; cv, crossvein; v, vein). G–I, Anterior part of thorax, anteroventral view. A, Nipponocampa esakii, female. B, I, Tomostethus nigritus, female. C, Apareophora japonica, female. D, G, Phymatoera nipponica, male. E, Phymatoceriola nipponensis, female. F, H, Hoplocampoides longiserrus, female (holotype).	FIGURE 4. A–E, Fore wing (c, cell; cv, crossvein; v, vein). G–I, Anterior part of thorax, anteroventral view. A, Nipponocampa esakii, female. B, I, Tomostethus nigritus, female. C, Apareophora japonica, female. D, G, Phymatoera nipponica, male. E, Phymatoceriola nipponensis, female. F, H, Hoplocampoides longiserrus, female (holotype).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFE2FFBAFF7B9DECFC08EC6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883203/files/figure.png	https://doi.org/10.5281/zenodo.17883203	FIGURE 7.A–D, Tarsal claw.E, F, Head, ventrolateral view.A, Phymatoceriola nipponensis, female (holotype).B, Phymatocera peregrinator comb. nov., female. C, E, Phymatoera nipponica, female. D, F, Paracharactus leucopodus, female.	FIGURE 7.A–D, Tarsal claw.E, F, Head, ventrolateral view.A, Phymatoceriola nipponensis, female (holotype).B, Phymatocera peregrinator comb. nov., female. C, E, Phymatoera nipponica, female. D, F, Paracharactus leucopodus, female.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFE2FFBAFF7B9DECFC08EC6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883205/files/figure.png	https://doi.org/10.5281/zenodo.17883205	FIGURE 8. A, B, Female abdomen, ventrolateral view. C–E, Head, lateral view. F, G, Basal part of male antenna, dorsal view (below lateral). A, D, Phymatoera nipponica, female. B, G, Phymatoceriola suigenensis: B, female (lectotype); G, male. C, Phymatocera fuscata, female (holotype). E, Phymatoceriola ussuriensis, male. F, Phymatoera membra, male.	FIGURE 8. A, B, Female abdomen, ventrolateral view. C–E, Head, lateral view. F, G, Basal part of male antenna, dorsal view (below lateral). A, D, Phymatoera nipponica, female. B, G, Phymatoceriola suigenensis: B, female (lectotype); G, male. C, Phymatocera fuscata, female (holotype). E, Phymatoceriola ussuriensis, male. F, Phymatoera membra, male.	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
26267A73FFE2FFBAFF7B9DECFC08EC6D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17883186/files/figure.png	https://doi.org/10.5281/zenodo.17883186	FIGURE 3. A, C, D, Hind tarsus. B, Middle tarsus.A, Tomostethus nigritus, female. B, C, Onychostethomostus gilvipes, female (holotype) (C, reversed). D, Esehabachia luteipes, female (holotype).	FIGURE 3. A, C, D, Hind tarsus. B, Middle tarsus.A, Tomostethus nigritus, female. B, C, Onychostethomostus gilvipes, female (holotype) (C, reversed). D, Esehabachia luteipes, female (holotype).	2025-10-17	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege		Zenodo	biologists	Hara, Hideho;Ibuki, Shinichi;Vårdal, Hege			
