taxonID	type	description	language	source
265D8C4EFFB1FF89FF23DE5DFD47FBE8.taxon	description	(Figs. 1 – 3)	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFB1FF89FF23DE5DFD47FBE8.taxon	materials_examined	Voucher: 2021 - FLP-IMM- 0217 (GenBank voucher: ON 259521).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFB1FF89FF23DE5DFD47FBE8.taxon	description	Egg. Not documented. First instar (Figs. 1 a, b; 2 a). Head capsule width: 0.9 mm (n = 1). Head capsule appearing black, epicranial notch insignificant and vertex flattened, with two rectangular scoli, each with two flattened, rather fanned setae with broad apex (scolus length 0.2 mm (n = 1 )); three chalazae visible in frontal view, M 1, M 2, and M 3, with seta on each of M 1 – 3; six setae visible on one side of labrum; six stemmata visible, with stemma 1 and 6 somewhat semi-transparent and thus insignificant, stemma 3 largest, closer to stemma 2 than 4. Body smooth, integument semi-transparent and creamy-white, but also receiving influence from green intestinal content; pair of white mid-dorsal lines visible, two-toned sub-dorsal line present, brownish above and whitish below, another two-toned lateral line present in supraspiracular region, whitish above and brownish below, additional whitish lateral subspiracular line present below; spiracles appearing light brown, tracheal system not discernable; chaetotaxy not fully discernible based on photographs but apical shape of primary setae strongly bulbed (sensu Murray 2001 b: Fig. 3.13 D), XD 1 and XD 2 visible on T 1 (apparently equal in length), D 1, D 2, SD 1 and L 1 apparently present from T 1 to A 10 (D 1 about three times longer than D 2 on A 8 and A 9), SV 1, SV 2, SV 3, L 2 and PP 1 noticeable on A 10; whitish bifid caudal filaments, appearing similar or shorter than A 8 in dorsal view. Body length: 5.4 mm (n = 1). Duration: Unknown, molted 4 days after sampling (n = 1). Second instar (Figs. 1 c, d; 2 b). Head capsule width: 1.1 mm (n = 1). Head capsule appearing dark brown with minute sculpting, epicranial notch and vertex as in previous instar, with numerous secondary setae and two developed horn-like scoli, inner margin appearing darker (darker area extends to vertex), with seta on each tip (scolus length 0.4 mm (n = 1 )); seven whitish chalazae visible on lateral side, M 1 – 3, P 1 – 4 with primary seta on each; six setae on one side of labrum; six stemmata visible, arrangement similar to previous instar. Body overall appearing dark flesh and granular; pair of dorsal lines changing [from first instar] to creamy-whitish, brown lines in the subdorsal and supra-spiracular regions changing [from first instar] to pinkish, both accompanied by two creamy-whitish lines below and above, respectively, line below spiracles insignificant; pair of brownish smudge-like markings appearing on posterior side of sub-dorsal area on A 1, A 2, and A 3, prominent on A 3; setae thread-like and shorter; spiracles brownish, tracheal system not discernable; bifid caudal filaments, appearing longer than A 8 in dorsal view, innermargin pinkish. Body length: 8.3 mm (n = 1). Duration: 8 days (n = 1). Third instar (Figs. 1 e, f; 2 c). Head capsule width: 1.4 mm (n = 1). Head capsule similar to previous instar (scolus length 0.5 mm (n = 1 )); six setae on one side of labrum. Body similar to previous instar except for brownish smudge-like markings appearing more significant, noticeable on A 4, A 5, A 6, and A 7; spiracles appearing darker. Body length: 13.1 mm (n = 1). Duration: 7 days (n = 1). Fourth instar (Figs. 1 g, h; 2 d). Head capsule width: 1.7 mm (n = 1). Head capsule similar to previous instar except for epicranial notch somewhat noticeable, overall appearing more elongate and contrast between dark area and paler area more prominent (scolus length 0.6 mm (n = 1 )); six setae on one side of labrum. Body similar to previous instar except for overall appearing paler and less pinkish, terminal abdominal segments appearing darker dorsally (due to darker markings along mid-dorsal lines); mid-dorsal to lateral creamy-white lines delineated with pinkish border, line below spiracles more defined; brownish smudge-like markings appearing somewhat reduced, breaking into four small spots on A 3. Body length: 23.5 mm (n = 1). Duration: 9 days (n = 1). Fifth (ultimate) instar (Figs. 1 i, j; 2 e). Head capsule width: 2.6 mm (n = 1). Head capsule similar to previous instar except for darker area more prominent and labrum appearing reduced (scolus length 0.9 mm (n = 1 )); reduced labrum, six setae on one side labrum. Body similar to previous instar except for overall appearing ocher; dark brownish shading present dorsally, “ X-shaped ” and prominent from T 2 to A 2, fading but still visible from A 3 onwards and noticeable again on terminal few abdominal segments; dark brown brownish markings visible below sub-dorsal line, appearing as band in dorsal view. Body length: 31.8 mm (n = 1). Duration: 18 days (n = 1). Pupa (Figs. 1 k – m). Short and relatively smooth, overall appearing light brownish and somewhat mottled with dark brown, ventrally creamy white scattered with dark brown, including antennae, wing case, and legs, mesothorax dorsally marked with prominent darker patch, middle part of thorax and posterior margin of wing case appears carinate with small keel, two longitudinal rows of protuberances on abdomen marked partially dark brown, as well as cremaster [details not discernible due to apparent subsequent loss]. Body length: 12.4 mm (n = 1). Duration: 15 days (n = 1).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFB1FF89FF23DE5DFD47FBE8.taxon	biology_ecology	Host plant of Caeruleuptychia urania Taquara micrantha (Kunth) I. L. C. Oliveira & R. P. Oliveira (Poaceae: Bambusoideae: Olyreae) (Fig. 3). This plant species associated with C. urania is a species of herbaceous bamboo that grows at the edge, along forest trails, and in light gaps in the forest, as well as grouped with other individuals of the same species. For 2021 - FLP-IMM- 0217, the host plant associated with the larva was found in a light gap near the edge of the forest, in an area adjacent to a reforestation plot. On the date of collection (March 9, 2021) the host plant was 1.5 m in height with the larva located 1.1 m above the ground and situated on the abaxial surface, in the middle part of the leaf blade, between the midrib and the leaf margin (Table 2).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFBCFF8EFF23DFCAFC0CFE6C.taxon	description	(Figs. 4 – 7)	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFBCFF8EFF23DFCAFC0CFE6C.taxon	materials_examined	Vouchers: 2021 - FLP-IMM- 0277, 0526 (GenBank voucher: ON 259527).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFBCFF8EFF23DFCAFC0CFE6C.taxon	description	Egg. Not documented. First instar (Figs. 4 a, b; 5 a). Head capsule width: 0.75 mm (n = 1). Head capsule appearing brown to black, epicranial notch insignificant and vertex rather flattened, with two rectangular scoli, each accompanied with two thread-like setae equal in length (scolus length 0.2 mm (n = 1 )); three chalazae visible in frontal view, M 1, M 2, M 3, with seta on each of M 1 – 3; six setae visible on one side of labrum; six stemmata visible, with stemma 1 and 6 paler and thus somewhat insignificant, stemma 3 largest, closer to stemma 2 than 4. Body integument creamy whitish and semi-transparent (small white stippling noticeable throughout), but also receiving influence from green intestinal content; pair of faint white lines in subdorsal area extending to A 10, faint reddish dorsal and lateral lines noticeable (prominent posteriorly); chaetotaxy not fully discernible based on photographs but primary setae weakly bulbed at apex (sensu Murray 2001 b: Fig 3.13 C), XD 1 and XD 2 visible on T 1, similar in length; D 1, D 2, SD 1 and L 1 apparently present from T 1 to A 10, D 1 becoming progressively longer towards posterior abdominal segments, D 1 about three times longer than D 2 on A 8 and A 9; ventral prolegs present on A 3 to A 6, caudal prolegs present on A 10; caudal filament short (shorter than A 8 in dorsal view). Body length: 4.3 mm (n = 1). Duration: Unknown, molted the day following capture. Second instar (Figs. 4 c, d; 5 b). Head capsule width: 0.9 mm (n = 1). Head appearing black with numerous secondary setae; vertex and epicranial notch as in previous instar; “ horn-like ” developed scolus, without setae, appearing concolorous except for somewhat insignificant whitish stripe on posterior side extending beyond base (scolus length 0.4 mm (n = 1 )); chalazae appearing reduced on lateral side, with primary seta on each of these chalazae discernible; six stemmata visible, with stemma 1 and 6 somewhat semi-transparent and thus insignificant, stemma 3 largest, closer to stemma 2 than 4; six setae visible on one side of labrum. Body greenish-yellow to chartreuse, purple band extending along mid-dorsal area, well-defined between T 1 to T 3 and A 8 to A 10, somewhat faint in between; creamy white insignificant band traversing dorso-lateral area, with jagged dark purplish band below, well defined along couple of terminal abdominal segments; spiracles visible as brownish spots, prominent on T 1 and A 8, tracheal system not visible; ventral prolegs present on A 3 to A 6, caudal prolegs present on A 10; bifid caudal filaments similar in length to A 8 in dorsal view, appearing purplish dorsally. Body length: 6.9 mm (n = 1). Duration: 7 days (n = 1). Third instar (Figs. 4 e, f; 5 c). Head capsule width: 1.0 mm (n = 1). Head mostly black but partially appearing russet with numerous secondary setae; “ horn-like ” scolus appearing concolorous except for anteriorly appearing reddish (with brownish tip) and whitish stripe clearly visible and well defined on posterior side extending beyond base (scolus length 0.40 mm (n = 1 )); three semi-transparent chalazae visible on lateral side (M 1 – 3), with primary seta on each of these three chalazae; six stemmata visible, with stemma 1 and 6 somewhat semi-transparent and thus insignificant, stemma 3 largest, closer to stemma 2 than 4; six setae visible on one side of labrum. Body overall grayish-green but T 1 – 3 and terminal abdominal segments somewhat paler, chartreuse to rather grayish-green, dark band extending along mid-dorsal area, well-defined between T 1 to T 3 and A 5 to A 8, somewhat insignificant from A 1 to A 4; creamy white insignificant band traversing dorso-lateral area; pair of brownish markings present on posterior end of A 3; laterally scattered with freckle-like brownish spots; spiracles visible as brownish spots, prominent on T 1 and A 8; ventral prolegs present on A 3 to A 6, caudal prolegs present on A 10; bifid caudal filaments elongate (twice as long as A 8 in dorsal view), appearing creamy white laterally, inner side pink. Body length: 7.7 mm (n = 1). Duration: 9 days (n = 1). Fourth instar (Figs. 4 g, h; 5 d). Head capsule width: 1.2 mm (n = 1). Head appearing light olive to brown with numerous secondary setae; “ horn-like ” scolus appearing orangish distally, light stripe at posterior side of scolus delineating dark area on vertex extending towards T 1, as well as lateral side of head (scolus length 0.5 mm (n = 1 )); three semi-transparent chalazae visible on lateral side (M 1 – 3), with primary seta on each of these three chalazae; stemmata arrangement similar to previous instar; six setae visible on one side of labrum. Body overall beige to grayish-brown (becoming whitish day prior to molt); general pattern similar to previous instar except creamy-white dorso-lateral band jagged and more defined; spiracles darker; caudal filaments similar in length to A 8 in dorsal view. Body length: 12.4 mm (n = 1). Duration: 7 days (n = 1). Fifth (penultimate) instar (Figs. 4 i, j; 5 e). Head capsule width: 1.7 mm (n = 1). Head appearing light olive with numerous secondary setae; “ horn-like ” scoli appearing orangish distally, dark stripe on lateral side extending beyond base, as well as inner side extending to cover vertex (scolus length 0.8 mm (n = 1 )); three semi-transparent chalazae visible on lateral side (M 1 – 3), with primary seta on each of these three chalazae; stemmata arrangement similar to previous instar; six setae visible on one side of labrum. Body overall grayish to ocher-rust, rather orangish in 2021 - FLP-IMM- 0277; general pattern similar to previous instar except for mid-dorsal band appearing to become defined in A 3 and A 4, dark jagged bands appearing in dorso-lateral and lateral area from A 2 to A 5, inner side of caudal filaments dark. Body length: 18.3 mm (n = 1). Duration: 8 days (n = 1). Sixth (ultimate) instar (Figs. 4 k, l; 5 f). Head capsule width: 2.5 mm (n = 1). Head similar to previous instar except for base of scolus broadening and labrum reduced (scolus length 1.0 mm (n = 1 )). Body ocher to grayishocher, general pattern similar to previous instar. Body length: 31.7 mm (n = 1). Duration: 16 days (n = 1). Pupa (Figs. 4 m-o). Body creamy-white and overall mottled with light brown; wing case following same pattern; body short and round; small cream and dark brown protuberances present in two rows along dorsal area of abdomen except for pair on A 1 lacking dark brown coloration; spiracles pink; dark brown short and broad cremaster, appearing rectangular in lateral view. Body length: 12.2 mm (n = 1). Duration: 12 days (n = 1).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFBCFF8EFF23DFCAFC0CFE6C.taxon	discussion	Note. A possible case of individual variation in larval instar numbers (five or six) has been observed for D. ocypete. We interpreted that 2021 - FLP-IMM- 0277 was collected in its second larval instar on March 24, 2021, based on its size and coloration, subsequently molting three times.	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFBCFF8EFF23DFCAFC0CFE6C.taxon	biology_ecology	Host plants of Deltaya ocypete Pariana lunata Nees (Poaceae: Bambusoideae: Olyreae) (Fig. 6). This plant species associated with D. ocypete is an herbaceous bamboo that grows at the edge of the forest, along forest trails, in light gaps, and also sometimes at the edge of the palm swamp. This plant species potentially exhibits an ephemeral lifestyle in response to the intense seasonal pressures present at FLP during the dry season, based on observations of severe mortality. For 2021 - FLP-IMM- 0277, the associated host plant was located at the edge of a forest trail. On the date of the collection (March 24, 2021), the plant was 1.3 m tall with the larva located 0.3 m above the ground and positioned on the abaxial surface of the leaf, near the base of the blade, on the midrib of the leaf blade (Table 2). Pariana vulgaris Tutin (Poaceae: Bambusoideae: Olyreae) (Fig. 7). This plant species, also associated with D. ocypete, is an herbaceous bamboo that grows at the edge of the forest adjacent to reforestation plots, usually near the edge of the palm swamp, along forest trails and light gaps. During the dry season, while the plants in the forest were dry, this plant species associated with 2021 - FLP-IMM- 0526 was vigorous, as they were located at the edge of the palm swamp. On the day of the collection (September 1, 2021) the plant was 0.35 m tall with the larva located 0.1 m above the ground and positioned on the abaxial surface of the leaf, near the base of the blade, on the midrib of the leaf blade (Table 2).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFBBFF92FF23DAB7FEFDFF3E.taxon	description	(Figs. 3, 8 – 10)	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFBBFF92FF23DAB7FEFDFF3E.taxon	materials_examined	Vouchers: 2021 - FLP-IMM- 0240, 0304, 0311, 0374, 0406, 0564, 0565; 2022 - FLP-IMM- 0138, 0139, 0162 (GenBank vouchers: ON 259522 – ON 259526).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFBBFF92FF23DAB7FEFDFF3E.taxon	description	Egg (Figs. 8 a, b). Spherical, slightly elongated vertically, semi-transparent and pearl-like and whitish in color, with hexagonal markings covering the surface. Diameter: 1.2 mm (n = 1). Duration: Unknown, hatched 2 – 6 days (n = 3) after collection. First instar (Figs. 8 c, d; 9 a). Head capsule width: 0.8 mm (n = 1). Head capsule appearing dark brown, epicranial notch insignificant and vertex flattened, with two rectangular scoli, each with two spatula-like setae equal in length and appearing dark, with narrow tips (scolus length 0.2 mm (n = 1 )); three chalazae visible in frontal view, M 1, M 2, and M 3, with spatula-like seta on each of M 1 – 3 (posterior chalazae reduced); six setae visible on one side of labrum; six stemmata visible, with stemma 1 and 6 somewhat semi-transparent and thus insignificant, stemma 3 largest, closer to stemma 2 than 4. Body integument semi-transparent and creamy-white, but also receiving influence from dark green to light green intestinal content; tracheal system noticeable as white line at spiracles; chaetotaxy not fully discernible based on photographs but primary setae appearing transparent and strongly bulbous at apex (sensu Murray 2001 b: Fig 3.13 D), XD 1 and XD 2 visible on T 1, D 1, D 2, SD 1 and L 1 apparently present from T 1 to A 10, D 1 appearing longer than D 2 on A 8 and A 9, but weakly so; short, bifid caudal filaments (shorter than A 8 in dorsal view). Body length: 4.9 – 6.2 mm (n = 2). Duration: 7 – 9 days (n = 3). Second instar (Figs. 8 e, f; 9 b). Head capsule width: 1.1 mm (n = 1). Head capsule appearing brown with numerous secondary setae and pair of “ antler-like ” bifurcating scoli, inner short fork positioned perpendicular to horizontal plane, appearing darker distally, with seta on both tips (scolus length 0.6 mm (n = 1 )); seven whitish chalazae visible on lateral side, M 1 – 3 clearly noticeable with primary seta on each, P 1 - 4 insignificant; six setae on one side of labrum; six stemmata visible, with stemma 1 and 6 somewhat semi-transparent and thus insignificant, stemma 3 largest, close to stemma 2 than 4. Body integument creamy-white, with whitish spots scattered across body, intestinal content influencing body color as in previous instar; setae thread-like and shorter; tracheal system noticeable as white line at spiracles; bifid caudal filaments, length similar to A 8 in dorsal view, dorsally pinkish. Body length: 7.5 – 8.8 mm (n = 2). Duration: 6 – 10 days (n = 3). Third instar (Figs. 8 g, h; 9 c). Head capsule width: 1.5 mm (n = 1). Head capsule appearing brown with numerous secondary setae and pair of “ antler-like ” bifurcating scoli, inner short fork positioned perpendicular to horizontal plane, appearing darker distally (narrow white stripe visible posteriorly), with seta on both tips (scolus length 0.6 mm (n = 1 )); seven whitish chalazae visible on lateral side, M 1 – 3 with primary seta on each, P 1 – 4 more prominent compared to preceding instar; six setae on one side of labrum; six stemmata visible, with stemma 1 and 6 somewhat semi-transparent and thus insignificant, stemma 3 largest, close to stemma 2 than 4. Body integument creamy-white, intestinal content influencing body color as in previous instar but body turning whitish day prior to molt and gut content not visible; setae thread-like and shorter; pair of whitish jagged stripes present dorsally, concolorous stripe in sub-dorsal area extending from scolus to caudal filament, area between these stripes appearing whitish roughly from A 2 to A 6, two rather insignificant lateral stripes present; tracheal system noticeable as white line at spiracles; bifid caudal filaments, length longer than A 8 in dorsal view, dorsally pinkish. Body length: 14.2 – 18.0 mm (n = 2). Duration: 6 – 10 days (n = 3). Fourth (penultimate) instar (Figs. 8 i, j; 9 d). See Nakahara et al. (2022). Fifth (ultimate) instar (Figs. 8 k, l; 9 e). See Nakahara et al. (2022). Pupa (Fig. 4: m – o). Short and relatively smooth, ventrally overall appearing somewhat mottled, creamy white scattered with dark brown, including antennae, wing case, and legs, dorsally creamy white scattered with dark brown except for rather pinkish two longitudinal rows of protuberances on abdomen and small areas of creamy-white elsewhere, cremaster brown with whitish pitted sculpture, broadens horizontally in middle with brownish rectangular plate (in posterior view), otherwise narrowing towards distal end. Body length: 11.2 mm (n = 1). Duration: 11 days (n = 1). Note. An individual variation in larval instar numbers (four or five) has been observed for C. philippa. Since passing through five larval instars was more common, the description above is based on that instar number.	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFBBFF92FF23DAB7FEFDFF3E.taxon	biology_ecology	Host plants of Cisandina philippa Taquara micrantha (Kunth) I. L. C. Oliveira & R. P. Oliveira (Poaceae: Bambusoideae: Olyreae) (Fig. 3). Following Nakahara et al. (2022 b), we also report Taquara micrantha as a host plant associated with C. philippa. See section above for further details on the habitat of this plant species. For 2021 - FLP-IMM- 0565, the associated host plant was located near the edge of a palm swamp. Ot the date of collection (November 8, 2021), the plant was 0.6 m tall with the larva located 0.6 m above the ground and positioned on the abaxial surface of the leaf, between the base and apex of the leaf blade, near the leaf margin (Table 2). Olyra latifolia L. (Poaceae: Bambusoideae: Olyreae) (Fig. 10). This plant species associated with C. philippa is an herbaceous bamboo that grows inside the forest, alongside dense understory vegetation and along forest trails as well. For 2021 - FLP-IMM- 0374, the associated host plant was located at the edge of a forest trail. On the date of collection (May 12, 2021), the plant was 1.5 m tall with the larva located 1.5 m above the ground and positioned on the abaxial surface of the leaf, in the middle part, between the midrib and the margin of the leaf blade (Table 2). Olyra caudata Trin. (Poaceae: Bambusoideae: Olyreae). This plant species associated with C. philippa is a rare herbaceous bamboo that grows inside the forest, alongside dense understory vegetation at FLP. For 2021 - FLP-IMM- 0304, the associated host plant was located at the edge of a forest trail. On the date of collection (April 3, 2021), the plant was 0.9 m tall with the larva located 0.5 m above the ground and positioned on the abaxial surface of the leaf, near to the base, between the midrib and margin of the leaf blade (Table 2).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA6FF95FF23DBDCFCC8FC79.taxon	description	(Figs. 10 – 13)	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA6FF95FF23DBDCFCC8FC79.taxon	materials_examined	Vouchers: 2021 - FLP-IMM- 0376; 2022 - FLP-IMM- 0094, 0095; 2023 - FLP-IMM- 0122, 0145 *, 0210, 0274 *, 0275, 0342, 0343 (GenBank voucher: ON 514278).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA6FF95FF23DBDCFCC8FC79.taxon	description	* denotes individuals used for body length measurements. Egg (Figs. 11 a, b). Spherical, similar to C. philippa, except for more rounded and appearing more yellowish. Diameter: 1.1 mm (n = 1). Duration: Unknown, hatched 5 days after collection (n = 1). First instar (Figs. 11 c, d; 12 a). Head capsule width: 0.88 mm (n = 1). Head capsule overall similar to C. philippa, except for appearing paler (scolus length 0.2 mm (n = 1 )). Body overall similar to C. philippa but slightly paler. Body length: 5.1 – 5.6 mm (n = 2). Duration: 6 – 9 days (n = 2). Second instar (Figs. 11 e, f; 12 b). Head capsule width: 1.1 mm (n = 1). Head capsule appearing light brown with numerous secondary setae, with pair of “ antler-like ” scoli with several “ wart-like ” projections, appearing darker distally, with seta on all projections (scolus length 0.4 mm (n = 1 )); seven whitish chalazae visible on lateral side, M 1 – 3 and P 1 – 4 (these may appear reduced), with primary setae on each; setae on labrum not examined; six stemmata visible, with stemma 1 and 6 somewhat semi-transparent and thus insignificant, stemma 3 largest, close to stemma 2 than 4. Body similar to immediately preceding species, greenish and tracheal system visible. Body length: 8.2 – 8.7 mm (n = 2). Duration: 8 – 10 days (n = 2). Third instar (Figs. 11 g, h; 12 c). Head and body not described herein, see Nakahara et al. (2022 c). Body length: 11.4 – 12.1 mm (n = 2). Duration: 7 – 10 days (n = 2). Fourth (penultimate or ultimate) instar (Figs. 11 i, j; 12 d). Head and body not described herein, see Nakahara et al. (2022 c). Body length: 19.1 – 19.4 mm (n = 2). Duration: 8 – 11 days (n = 2). Fifth (ultimate) instar (Figs. 11 k, l, 12 e). Head and body not described herein, see Nakahara et al. (2022 c). Body length: 25.7 – 27.6 mm (n = 2). Duration: 13 days (n = 1). Pupa (Figs. 11 m – o). Not described herein, see Nakahara et al. (2022 c). Duration: 12 days (n = 1).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA6FF95FF23DBDCFCC8FC79.taxon	discussion	Note. An individual variation in larval instar numbers (four or five) has been observed for C. castanya. The mature larva descriptions can be supplemented by Nakahara et al. (2022 c) but it must be noted that the present study suggests the difference in larval instar number occurs at a similar frequency.	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA6FF95FF23DBDCFCC8FC79.taxon	biology_ecology	Host plants of Cisandina castanya Olyra latifolia L. (Poaceae: Bambusoideae: Olyreae) (Fig. 10). Following Nakahara et al. (2022 b), we also report Olyra latifolia as a host plant associated with C. castanya. See the section above for further details on the habitat of this plant species. For 2021 - FLP-IMM- 0376, the associated host plant was located at the edge of a forest trail. On the date of collection (May 12, 2021), the plant was 1.5 m tall with the larva located 1 m above the ground and positioned on the abaxial surface of the leaf, in the middle part of the leaf blade, between the midrib and the margin of the leaf blade (Table 2). Piresia macrophylla Soderstr. (Poaceae: Bambusoideae: Olyreae) (Fig. 13). This plant species associated with C. castanya is an herbaceous bamboo that grows inside the forest, alongside dense understory vegetation, and along forest trails. For 2022 - FLP-IMM- 0095, the associated host plant was located at the edge of a forest trail. On the date of collection (February 23, 2022), the plant was 0.5 m tall with the larva located 0.5 m above the ground and positioned on the abaxial surface of the leaf, in the middle part of the leaf blade, between the midrib and the margin of the leaf blade (Table 2).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA0FF99FF23DF5BFCE5FEA2.taxon	description	(Figs. 14 – 16)	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA0FF99FF23DF5BFCE5FEA2.taxon	materials_examined	Vouchers: 2022 - FLP-IMM- 0136; 2025 - FLP-IMM- 0028 (GenBank voucher: PX 102130).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA0FF99FF23DF5BFCE5FEA2.taxon	description	Egg (Figs. 14 a, b). Spherical, yellow and semi-translucent, with apparently somewhat irregular polygonal markings covering the surface. Head capsule visible day prior to hatching. Diameter: 1.2 mm (n = 1). Duration: Unknown, hatched 4 days (n = 1) after collection. First instar (Figs. 14 c, d). Head capsule width: 1.0 mm (n = 1). Head capsule appearing light olive, epicranial notch insignificant and vertex rather flattened, with two scoli, appearing with rectangular basal half bifurcating into triangular distal half, inner triangular projection appearing darker, each triangular projection with seemingly thread-like setae equal in length (scolus length 0.2 mm (n = 1 )) (presence / absence of additional pore / setae not discernible based on photos); three chalazae visible on one side in frontal view, M 1, M 2, M 3, with seta on each of M 1 – 3 (M 1 / 2 darker); clypeus and mandibles brown; four setae visible on side of labrum; six stemmata present mostly located on dark patch, stemma 3 largest, slightly closer to stemma 2 than 4. Body integument creamy whitish and semi-transparent, but also receiving influence from green intestinal content and revealing pair of faint white lines in subdorsal area extending to A 10; tracheal system visible along spiracles as white line; chaetotaxy not fully discernible based on photographs but primary setae weakly bulbed at apex (sensu Murray 2001 b: Fig 3.13 C), XD 1 and XD 2 apparently present on T 1; D 1, D 2, SD 1 and L 1 apparently present from T 1 to A 10, D 1 becoming longer towards posterior abdominal segments (appearing darker and slightly thicker on A 10), D 1 and D 2 similar in length on A 8 and A 9; ventral prolegs present on A 3 to A 6, caudal prolegs present on A 10; caudal filament pink and short (shorter than A 8 in dorsal view). Body length: 6.9 mm (n = 1). Duration: 8 days (n = 1). Second instar (Figs. 14 e, f; 15 a). Head capsule width: 1.4 mm (n = 1). Head colour and general appearance similar to preceding instar, rough texture with subcircular protuberances regularly distributed, notable differences include growth of secondary setae, scolus and chalazae developed and “ horn-like ”, darkening as they grow (scolus length 0.3 mm (n = 1 )), as well as P 1 – 4, and six other minor chalazae accompanied with seta located posteriorly (additional chalazae visible behind head scoli and dark patch reduced compared to previous instar); Body greenish-yellow to chartreuse (becoming darker later in this instar), white band extending along mid-dorsal area, well defined from A 1, becoming more defined later in this instar; few creamy white lines traversing dorso-lateral and lateral area; spiracles visible as whitish spots, prominent on T 1 and A 8, tracheal system visible as in preceding instar; ventral prolegs present on A 3 to A 6, caudal prolegs present on A 10; bifid caudal filaments similar in length to A 8 in dorsal view, appearing pink, with base of setae appearing black. Body length: 12.0 mm (n = 1). Duration: 12 days (n = 1). Third instar (Figs. 14 g, h; 15 b). Head capsule width: 2.1 mm (n = 1). Head (scolus length 0.5 mm (n = 1 )) and body similar to immediately preceding instar except for head colour turning into olive green, dark apical area of chalazae and scoli reduced, and body overall appearing mint green and irregular whitish markings visible dorsally. Body length: 19.1 – 22.9 mm (n = 2). Duration: 7 – 10 days (n = 2). Fourth (ultimate) instar (Figs. 14 i – l; 15 c). Head capsule width: 3.5 mm (n = 1). Head capsule similar to previous instar except for overall appearing brown and dull, surface rather rough, chalazae appearing stubby, darker area prominent and labrum appearing reduced (scolus length 0.9 mm (n = 1 )); six setae on one side of labrum. Body similar to previous instar except for overall appearing light ocher to beige, poorly defined light ocher bands with pink borders in subdorsal region, with some dark brownish shading noticeable but becoming reduced later in fourth instar; turning into purple with orange markings on last day of fourth (ultimate) instar and appearing concolorous during prepupa; bifid caudal filaments dark gray and stubby. Body length: 38.3 – 38.6 mm (n = 2). Duration: 11 – 13 days (n = 2). Pupa (Figs. 14 m – o). Body chestnut brown to dark chestnut brown, dorsally darker, somewhat mottled dorsally and grayish patches visible; squared wing case concolourous but overall appearing darker; body overall appearing angular in lateral view, dorsal margin of thorax angular in lateral view; rather developed two-toned (black and ocher) protuberances present in two rows along dorsal area of abdomen forming bumpy dorsal margin in lateral view; spiracles dark brown; cremaster dark brown and elongate, broad at base and gradually narrowing distally in dorsal view appearing trapezoidal, rather rectangular and evenly broad in lateral view. Body length: 17.7 mm (n = 1). Duration: 13 days (n = 1).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA0FF99FF23DF5BFCE5FEA2.taxon	biology_ecology	Host plant of Harjesia blanda Guadua weberbaueri Pilg. (Poaceae: Bambusoideae: Bambuseae) (Fig. 16). This plant species associated with H. blanda is a bamboo species that grows grouped with other individuals of the same species at the edge of land areas of palm swamp, and also in some deforested areas. For both vouchers, the associated host plant was located at the edge of a trail next to palm swamps, adjacent to agricultural areas. On the date of collection (March 15, 2022), for voucher 2022 - FLP-IMM- 0136 the host plant was 4 m tall with the egg located 1.3 m above the ground and situated on the abaxial surface of the leaf, near the base of the blade, between the midrib and the margin of the leaf blade, whereas for voucher 2025 - FLP-IMM- 0028 the host plant was 3 m tall with the larva located 1 m above the ground and situated on the abaxial surface of the leaf, near to the base of the leaf, on the margin of the leaf blade (Table 2).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFACFF9BFF23DAFDFAA7F853.taxon	description	(Figs. 17 – 19)	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFACFF9BFF23DAFDFAA7F853.taxon	materials_examined	Voucher: 2021 - FLP-IMM- 0511 (GenBank voucher: PX 102131).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFACFF9BFF23DAFDFAA7F853.taxon	description	Egg. Not documented. First instar (Figs. 17 a, b; 18 a). Head capsule width: 0.9 mm (n = 1). Head capsule appearing dark brown, epicranial notch insignificant and vertex flattened, with two trapezoidal shaped scoli, with pair of thread-like setae dilated at apex (scolus length 0.1 mm (n = 1 )); three chalazae visible in frontal view, M 1, M 2, M 3, with seta on each of M 1 – 3, additional seta noticeable above stemma 1 (labeled as A 4 in Murray (2001 b), but see also below); six stemmata visible, with stemma 1 and 6 somewhat semi-transparent and thus insignificant, stemma 3 largest, closer to stemma 2 than 4. Body pink and semi-transparent, receiving influence from dark green (T 1 to A 5) to lighter (A 6 to A 8) gut content; pair of white stripes in subdorsal area; three white lateral stripes present, parallel to previous stripe; chaetotaxy not fully discernible based on photographs but primary setae weakly bulbed at apex (sensu Murray 2001 b: Fig 3.13 C), XD 1 and XD 2 present on T 1, long and similar in length; D 1, D 2, SD 1 and L 1 present from T 1 to A 10, D 1 relatively lengthy throughout all body segments, D 1 longer than D 2 on A 8 and A 9 but weakly so; L 2 present from A 3 to A 10 (SD 2 not discernible); tracheal system barely noticeable at spiracles; bifid caudal filaments short (shorter than A 8 in dorsal view), with D 1, D 2 and SD 1 present (SD 2 not discernible); ventral prolegs present on A 3 to A 6, caudal prolegs present on A 10. Body length: 5.3 mm (n = 1). Duration: 8 days (n = 1). Second instar (Figs. 17 c, d; 18 b). Head capsule width: 1.5 mm (n = 1). Head appearing light russet with numerous secondary setae; pair of short and stubby “ horn-like ” scoli appearing darker (scolus length 0.3 mm (n = 1 )); three chalazae (M 1 – 3) noticeable; six stemmata visible, arrangement similar to previous instar. Body pink, pair of creamy-white lines extending along mid-dorsal area from T 1 to A 10, three dorso-lateral to lateral lines from T 1 to A 10 (lateral line below spiracles terminating at A 8); creamy white insignificant band traversing dorso-lateral area; spiracles visible as light brownish spots, prominent on T 1 and A 8; ventral prolegs present on A 3 to A 6, caudal prolegs present on A 10; bifid caudal filaments short (similar or shorter than A 8 in dorsal view), appearing creamy white dorsally, inner side pink. Body length: 6.4 mm (n = 1). Duration: 8 days (n = 1). Third instar (Figs. 17 e, f; 18 c). Head capsule width: 2.1 mm (n = 1). Head (scolus length 0.4 mm (n = 1 )) and body similar to immediately preceding instar except for light area posterior of scoli more prominent; dorso-lateral line jagged from A 2 to A 7. Body length: 14.3 mm (n = 1). Duration: 12 days (n = 1). Fourth instar (Figs. 17 g, h). Head capsule width: not measured. Head (scolus length not measured) and body similar to immediately preceding instar except for body overall darker, subdorsal white line appearing more sinuous, and spiracles appearing dark brownish. Body length: 18 mm (n = 1). Duration: 14 days (n = 1). Fifth (ultimate) instar (Figs. 17 i, j; 18 d, e). Head capsule width: 3.0 mm (n = 1). Head (scolus length 0.4 mm (n = 1 )) and body similar to immediately preceding instar except for scolus appearing stubby, setae base more developed and prominent, and labrum reduced; body overall darker, sinuous subdorsal line less prominent, and caudal filaments shorter. Body length: 32.0 mm (n = 1). Duration: 18 days (n = 1). Pupa (Fig. 9: k – m). Body chestnut brown, dorsally darker; creamy-white stripe present on dorsal surface of thorax; wing case delineated with creamy-white border; body overall appearing short and round in lateral view; small creamy-white protuberances present in two rows along dorsal area of abdomen somewhat forming stripes from A 3 onwards; spiracles white; chestnut brown cremaster, appearing rectangular in lateral view but somewhat elongate. Body length: 11.8 mm (n = 1). Duration: 14 days (n = 1). Note. Determination of the molt date from second instar to third instar involves guesswork as we used recording of two consecutive days of no feeding to narrow down this date (August 24, 2021); although the molt is an estimation by monitoring, the head capsule was collected for this particular molt. Additionally, we determined a molt from fourth to fifth instar between September 13 and 29 by examining larva images of both dates, although the fourth instar head capsule was not collected after this molt.	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFACFF9BFF23DAFDFAA7F853.taxon	biology_ecology	Host plant of Yphthimoides renata Lasiacis ligulata Hitchc. & Chase (Poaceae: Panicoideae: Paniceae) (Fig. 19). This plant species associated with Y. renata is a species of herbaceous bamboo that grows generally at the edge of the forest, and along trails within the forest. The phenology observed consists of flowering and fruiting between June and July in the study area. The associated individual with Y. renata was located at the edge of the forest. On the date of collection (August 9, 2021), the plant was 2.5 m tall. The larva was collected 1.1 m above the ground and positioned on the abaxial surface of the leaf, near the base of the leaf blade, at the leaf margin (Table 2).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA9FF9CFF23DECCFE86F81B.taxon	description	(Fig. 20)	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA9FF9CFF23DECCFE86F81B.taxon	materials_examined	Voucher: 2021 - FLP-IMM- 0312.	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA9FF9CFF23DECCFE86F81B.taxon	description	Egg to Pupa. Not documented and discussed herein, see Janzen & Hallwachs (2018).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA9FF9CFF23DECCFE86F81B.taxon	biology_ecology	Host plant of Magneuptychia libye Pharus virescens Döll (Poaceae: Pharoideae: Phareae) (Fig. 20). This plant species associated with M. libye is an herbaceous bamboo that grows in light gaps and along trails within the forest at FLP. This annual species is present during the rainy season and appears less vigorous in the dry season in this study area. For 2021 - FLP-IMM- 0312, the host plant from which the egg was collected was located in a light gap. On the date of collection (April 8, 2021) the plant was 1 m tall with the larva located 0.5 m above the ground and situated on the abaxial surface of the leaf, near the base of the blade, between the midrib and the leaf margin (Table 2).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA8FF9EFF23DC04FDB0F83D.taxon	description	(Fig. 21)	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA8FF9EFF23DC04FDB0F83D.taxon	materials_examined	Vouchers: 2023 - FLP-IMM- 0316, 0327 (GenBank vouchers: PX 102128, PX 102129).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA8FF9EFF23DC04FDB0F83D.taxon	description	Egg to Pupa. Not documented and discussed herein, see Nakahara et al. (2020).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
265D8C4EFFA8FF9EFF23DC04FDB0F83D.taxon	biology_ecology	Host plant of Modestia harpyia Euterpe precatoria Martius (Arecaceae: Arecoideae: Euterpeae) (Fig. 21). This species associated with M. harpyia is a palm species that grows within the forest with abundance being prominent in the canopy, and it should also be noted that there is an abundance of seedlings during the rainy season. For 2023 - FLP-IMM- 0316, the host plant was a seedling located inside the forest. On the date of collection (March 25, 2023), the seedling was 0.2 m tall with the egg located 0.1 m above the ground, situated on one of the leaflets on the abaxial side in the middle part of the leaf, on midrib of the leaf blade. For 2023 - FLP-IMM- 0327, the host plant was a seedling located inside the forest. At the collection date (March 26, 2023), the seedling was 0.1 m tall with the larva located 0.1 above the ground and situated on the adaxial surface of the leaf blade, in the distal part between the midrib and the leaf margin (Table 2).	en	Nakahara, Shinichi, Vega, Yeison, Corahua-Espinoza, Thalia, Kabir, Jamal, Maclure, Calum, Duerr, Nathan, Gallice, Geoffrey, Cervantes-Martinez, Joselin, Arteaga, Zunilda Escalante, Tejeira, Rafael, Ccahuana, Rodrigo, Baine, Quinlyn (2026): Immature stages and new host plant records for eight euptychiine butterflies in southeastern Peru (Lepidoptera: Nymphalidae: Satyrinae: Satyrini). Zootaxa 5748 (2): 151-192, DOI: 10.11646/zootaxa.5748.2.1, URL: https://doi.org/10.11646/zootaxa.5748.2.1
