taxonID	type	description	language	source
2764BD61A91BFFEBFE8EF8F0FCD1F92F.taxon	type_taxon	TYPE SPECIES. — Similiclypeina conradi Bucur, 1993, designated by original description.	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A91BFFEBFE8EF8F0FCD1F92F.taxon	discussion	REMARKS The genus diagnosis by Bucur (1993: 105) is as follows: “ Thallus made up of overlapping whorls of primary ramifications. Ramifications slightly tilted with respect to the axial cavity, welded at their proximal part and, usually, loose at their distal end. Overlapping whorls touching one another, partly welded into a more or less compact calcareous sheath ”. Some species that were doubtfully attributed to the genus Clypeina Michelin, namely Clypeina alrawii Radoičić, 1978, and Clypeina somalica Conrad, Peybernès & Masse, 1983 have been transferred to the genus Similyclypeina, along with the creation of the new species, Similiclypeina conradi (see Bucur 1993). For discussion on the taxonomy of Similiclypeina Bucur, 1993 and the close (synonymous?) genus Piriferella Sokač, 1996, see Bucur (2000), Conrad et al. (2009), Bucur et al. (2000, 2021), and Granier (2013). This controversial taxonomic issue however is not influencing the present generic assignment of the Tibetan alga as, if both genera are identical, Similiclypeina would then have priority.	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A918FFE6FF24F970FE5FFACC.taxon	description	(Figs 5 A, B; 6; 7)	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A918FFE6FF24F970FE5FFACC.taxon	materials_examined	TYPE MATERIAL. — Tibet • Northern Lhasa Terrane, Roma Chambo section. The section crops out along the eastern side of the Roma Chambo river, about 15 km SW of the Lake Dong Co in the Gerze county, Tibet (China). The coordinates of the section base are c. 32 ° 04 ’ 42 ” N, 84 ° 31 ’ 34 ” E, and encompasses a part of the Langshan Formation with a thickness of 80 m that can be assigned to the Aptian. Holotype: longitudinal thallus section illustrated in Figure 7 E; thin-section 21 LMC- 66 F 1 stored at the School of Earth Sciences and Resources, China University of Geosciences in Nanjing.	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A918FFE6FF24F970FE5FFACC.taxon	etymology	ETYMOLOGY. — The species name refers to the Lower-mid Cretaceous Langshan Formation cropping out in the Lhasa Terrane of Tibet.	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A918FFE6FF24F970FE5FFACC.taxon	materials_examined	TYPE LOCALITY AND AGE. — Tibet, northern Lhasa Terrane, Roma Chambo section (Figs 1; 2), late early (Praeorbitolina cormyi biozone of Schroeder et al. 2010) to early late Aptian. OCCURRENCE AND STRATIGRAPHY. — The species is widely distributed in the Roma Chambo section spanning an interval from the late early Aptian (Praeorbitolina biozone) and parts of the late Aptian.	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A918FFE6FF24F970FE5FFACC.taxon	diagnosis	DIAGNOSIS. — Species of Similiclypeina with relatively thick calcareous skeleton exhibiting spinose outer appearance, rather low ratio inner-outer diameter and tilted uncompressed tubular branches.	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A918FFE6FF24F970FE5FFACC.taxon	description	DIMENSIONS. — See Table 1. DESCRIPTION Thallus medium-sized, and cylindrical with verticils of primary laterals. Laterals tilted towards the main axis and arranged in quincunxes. The calcification consists of sparry calcite, is variable, and often displays a thick envelope around the main axis resulting in a low d / D ratio (e. g. Fig. 6 B, H). Mostly the elongated tubular (uncompressed) laterals (phloiophorous type), 6 to 10 per vertical, are straight and directed away from the main axis, sometimes (top part of thallus?) bending upwards (Fig. 7 F). Along their length, the laterals only slightly widen. The verticils are rather close-set; the individual calcareous coating of the laterals results in a spinose appearance at the distal endings (Fig. 6 A). In these cases, the calcified wall shows a slight concave depression at the thallus surface between the laterals. Transverse sections of the laterals as shown in tangential thallus sections are always rounded. The inner surface (axial cavity) is smooth and well delimited. COMPARISONS The main feature that in our opinion justifies the separation of Similiclypeina langshanensis sp. nov. from allied taxa are the shape of the laterals. In Similiclypeina somalica (Conrad, Peybernès & Masse), the laterals are directed almost perpendicular to the main axis and egg-shaped (Fig. 8 A, B). The laterals of the type-species S. conradi Conrad typically are flattened vertically at their proximal parts and continuously decrease in diameter outwards (Fig. 8 E, F). In addition, the diameter of the laterals is larger in S. conradi (Table 2). In Piriferella spinosa Sokač, the laterals are similar to S. somalica (Fig. 8 C, D). The laterals of S. langshanensis sp. nov. attain a length almost double than that reported from S. verrucosa Sokač and also the vertical spacing is distinctly lower (Fig. 8 G, H; Table 2). For other differences in biometric parameters see Table 2.	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A915FFE6FECEFA51FD97F9FD.taxon	type_taxon	TYPE SPECIES. — Bakalovaella elitzae (Bakalova) Bucur, 1993 [basionym: Cylindroporella elitzae Bakalova, 1971].	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A915FFE1FF60F930FD61F849.taxon	description	(Fig. 9 A-E, H)	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A915FFE1FF60F930FD61F849.taxon	distribution	STRATIGRAPHY. — Mu (1986) reported Heteroporella xizangensis from the northern slope of Langshan Mountain in the Duoba District, Baingoin County. The type-level is indicated as belonging to the lower part of the Langshan Formation containing Morelletpora turgida (Radoičić) and Palorbitolina lenticularis (Blumenbach) indicating an early Aptian age.	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A915FFE1FF60F930FD61F849.taxon	description	DESCRIPTION The alga has a cylindrical thallus with a main axis that corresponds to about 1 / 3 of the total diameter (Fig. 9 H). Sterile laterals and fertile whorls are arranged perpendicularly towards the axis in an alternating manner. In the last third (towards the algal surface), the fertile ampullae are touching each other (vertically and horizontally) appearing as a contiguous undulating micritic bands between the levels of sterile laterals (Fig. 9 C). The fertile ampullae are inflated with a flattened distal end thereby occupying the space between the sterile laterals almost completely .. The inner surface of the main axis is smooth.	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A915FFE1FF60F930FD61F849.taxon	discussion	REMARKS Heteroporella xizangensis Mu was originally described as belonging to Heteroporella Praturlon, 1967. Mu (1986) designated one holotype (PB 10994, pl. 4 [wrongly indicated as pl. VI = 6], fig. 1) and two paratypes (PB 10995, pl. 4, fig. 4 and PB 10696, pl. 4, fig. 5). Although validly described, it has neither been included in the critical literature inventory of species (Granier & Deloffre 1993; Bucur 1999) nor the taxonomic revision of the genus Heteroporella and its “ species ” (Granier et al. 1994). This led to the reduction of the inventory of the genus Heteroporella to its type-species H. lepina Praturlon, while all other species previously described as belonging to it were assigned to other genera. Heteroporella xizangensis Mu is herewith assigned to the genus Bakalovaella Bucur, 1993. This genus is characterized by thalli bearing alternating sterile and fertile whorls that are connected to the former that exhibit a distal widening to form a cortex (see also Granier & Bucur 2019). Bakalovaella xizangensis has been observed only in the Mendang section (upper Aptian-middle Albian). COMPARISONS Bakalovaella elitzae (Bakalova) (Barremian-Aptian of Bulgaria), another species of the genus that may be associated to B. xizangensis, is much smaller and displays almost spherical fertile ampullae (Fig. 9 I, J). Bakalovaella benizarensis (Fourcade et al. ex Jaffrezo in Bassoullet et al. 1978 (Barremian of Spain) represents a small-sized species (mean D: 0.60 mm to 0.85 mm) with six subspherical ampullae per vertical and a reduced vertical spacing (h: 0.13 - 0.18). Bakalovaella deloffrei Granier & Bucur, 2019 (Hauterivian of western France) represents another small representative of the genus (mean D: c. 0.70 mm) displaying a subcylindrical main axis bearing deeply imbricated verticils of 5 to 8 laterals each. Bakalovaella filipovici (Radoičić, 2006) (Cenomanian-Turonian of Kosovo) represents a species with a comparably narrow main axis (d: 0.075 - 0.177 mm) and subspherical fertile ampullae.	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A912FFE3FC7AFA51FCDEFE93.taxon	description	(Fig. 9 I, J)	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A912FFE3FC7AFA51FCDEFE93.taxon	discussion	REMARKS Bakalovaella elitzae is relatively rare in the Langshan Formation and has been observed in both sections studied. In the Mendang section it may co-occur with B. xizangensis in the same samples. For a detailed description of the species see Bucur (1993), Taherpour Khalil Abad et al. (2010), and Bucur et al. (2012) which includes a tentative thallus reconstruction.	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A910FFE3FF65FBF6FE12F84E.taxon	description	(Figs 5 E, F; 9 F, K)	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A910FFE3FF65FBF6FE12F84E.taxon	distribution	OCCURRENCES. — Roma Chambo section and Mendang section; Aptian and Albian.	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A910FFE3FF65FBF6FE12F84E.taxon	discussion	REMARKS Morelletpora turgida represents a rather common species in the upper Aptian-Albian parts of the Langshan Formation, often dispersed with individual thallus fragments in orbitolinid-rich facies (Fig. 5 D). Here the typical barrel-shaped isolated segments prevail. Packstones with abundant thallus fragments and are orbitolinids characterized by more elongated, almost cylindrical segments (Fig. 5 F).	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
2764BD61A910FFE3FF39FED5FCDEFE46.taxon	type_taxon	TYPE SPECIES. — Salpingoporella muehlbergii (Lorenz, 1902) Pia in Trauth (1918) [basionym: Diplopora muehlbergii Lorenz, 1902].	en	Sun, Qi, Schlagintweit, Felix, Li, Xianghui (2025): New records of Dasycladales (green algae) from the Aptian-Albian Langshan Formation of Tibet. Comptes Rendus Palevol 24 (20): 381-395, DOI: 10.5852/cr-palevol2025v24a20, URL: https://doi.org/10.5852/cr-palevol2025v24a20
