identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5123327E9E3C5C1B99DFEF94880853E9.text	5123327E9E3C5C1B99DFEF94880853E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liodessus cerrodelamuerte Balke & Bilton & Springer & Villastrigo & Neven & Hendrich 2026	<div><p>Liodessus cerrodelamuerte sp. nov.</p><p>Figs 1 A – E, 3, 4 A – C</p><p>Type locality.</p><p>Costa Rica, San José, Cerro de la Muerte.</p><p>Holotype, male: • Costa Rica, Cerro de la Muerte, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.75411&amp;materialsCitation.latitude=9.57041" title="Search Plazi for locations around (long -83.75411/lat 9.57041)">Calle de Torres</a>, 3,300 m, 27. ii. 2024, 9.57041°N, 83.75411°W, Springer &amp; Balke (CR_MB_2024_03) (MZUCR) .</p><p>Paratypes (N = 157). 115 exx. Same label data as holotype (MZUCR, ZSM); • 1 ex. Costa Rica, Cerro de la Muerte, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.75469&amp;materialsCitation.latitude=9.56658" title="Search Plazi for locations around (long -83.75469/lat 9.56658)">Calle de Torres</a>, 3,350 m, 27. ii. 2024, 9.56658°N, 83.75469°W, Springer &amp; Balke (CR_MB_2024_04) (MZUCR, ZSM) ; • 4 males, 2 females, 32 exx., Costa Rica, Cerro de la Muerte, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.75411&amp;materialsCitation.latitude=9.57041" title="Search Plazi for locations around (long -83.75411/lat 9.57041)">Calle de Torres</a>, 3,300 m, 17. vii. 2024, 9.57041°N, 83.75411°W, Bilton (CBP, ZSM) ; • 1 male, 1 female, Costa Rica, San José, Chirripó summit region, 18. iii. 1971, Hohmann (loc. 2) (NMB) (voucher MB 9030) .</p><p>Note.</p><p>We saw one additional specimen from the Páramo de Chirripó in the Costa Rican National Museum insect collection which we could, however, not borrow.</p><p>Description.</p><p>Habitus with slight but obvious discontinuity between pronotum and elytra (Fig. 1 A, B), pronotum widest before base and more parallel sided laterally. Total length (TL) of beetle: 2.0 mm; length without head (TL - h): 1.8 mm; maximum width (TW): 1 mm.</p><p>Colour. Dark brown to blackish dorsally and ventrally, basal antennomeres, base of meso- and metatibia contrastingly of lighter colour (as in Fig. 1 B).</p><p>Surface sculpture. Head with few larger and smaller setiferous punctures; with distinct microreticulation except on middle of head between the eyes (as in Fig. 1 B). Pronotum and elytra shiny; with moderately dense and coarse setiferous punctation (as in Fig. 1 B).</p><p>Structures. Antenna filiform. Head without occipital line; with rounded clypeus. Pronotum with distinct lateral bead; with distinct long and deep basal striae (Fig. 1 A). Elytra with distinct basal striae; without sutural line; without basal epipleural transverse carina. Metathoracic wings not examined in holotype (in paratype: short, about half the length of elytra, Fig. 1 B).</p><p>Genitalia. Median lobe of aedeagus slender with simple curvature in basal 1 / 3, then broadening and forming a convexity on dorsal margin (in lateral view), then significantly narrowing towards tip (Fig. 1 C); gradually narrowing towards a narrow tip in ventral view (Fig. 1 D); parameres of simple “ Bidessini ” type, two-segmented (Fig. 1 E).</p><p>Female. Pronotum and elytra can show a varying extent of microreticulation. This ranges from having a few meshes to being very obviously microreticulate on the disc of the pronotum and the elytra being only faintly microreticulate apically, to showing visible microreticulation on entire apical 1 / 3 to ½. Very few specimens, including the single female from Páramo de Chirripó, have a completely dull dorsal surface.</p><p>Variation. Size variation of the paratypes is TL 2.0– 2.2 mm; TL - h 1.7–1.9 mm; TW 1.0– 1.1 mm (specimen from Páramo de Chirripó slightly larger, TL 2.3 mm; TL - h 2.0 mm; TW 1.1 mm). Dorsal colour can be lighter brown to mostly dark orange-brown (see Fig. 1 A). Usually without occipital line, few specimens with faint line, more like a few serial punctures connected by a very shallow impression.</p><p>Etymology.</p><p>Named after the type locality, the mountain Cerro de la Muerte.</p><p>Differential diagnosis.</p><p>Distinguished from the other species of Liodessus by the diagnostic combination of the following features: species from high altitude of Costa Rica; with slight but obvious discontinuity between pronotum and elytra; head without (or with really faint) occipital line; pronotal and elytral striae present; wings reduced considerably; shape of median lobe.</p><p>Distribution.</p><p>Known from high altitudes of Costa Rica&gt; 3,000 m (Fig. 3).</p><p>Habitat.</p><p>Collected from small, shallow to very shallow puddles beside a gravel road, likely with running seepage water for much of the year (Fig. 4 A, B) as well as water at a small well nearby (Fig. 4 C).</p></div>	https://treatment.plazi.org/id/5123327E9E3C5C1B99DFEF94880853E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Bilton, David;Springer, Monika;Villastrigo, Adrián;Neven, Katja;Hendrich, Lars	Balke, Michael, Bilton, David, Springer, Monika, Villastrigo, Adrián, Neven, Katja, Hendrich, Lars (2026): Two new species of Liodessus diving beetles from Costa Rica (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 10: 147-154, DOI: 10.3897/alpento.10.193709
6C0B8183C2D754A68B1613BFB6DA3D53.text	6C0B8183C2D754A68B1613BFB6DA3D53.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liodessus quetzalicus Balke & Bilton & Springer & Villastrigo & Neven & Hendrich 2026	<div><p>Liodessus quetzalicus sp. nov.</p><p>Figs 2 A – D, 3, 4 D – F</p><p>Type locality.</p><p>Costa Rica, Cartago, Paraíso Quetzal.</p><p>Holotype, male: • Costa Rica, Páramo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.8452&amp;materialsCitation.latitude=9.6478" title="Search Plazi for locations around (long -83.8452/lat 9.6478)">Paraíso Quetzal</a>, 2,760 m, 10. - 11. iii. 2022, 9.6478°N, 83.8452°W, Springer &amp; Balke (CR_MB_2022_01) (MZUCR) .</p><p>Paratypes (N = 224). 130 exx., same data as holotype (MZUCR, ZSM); • 49 exx, Costa Rica, Páramo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.84528&amp;materialsCitation.latitude=9.64788" title="Search Plazi for locations around (long -83.84528/lat 9.64788)">Paraíso Quetzal</a>, 2,760 m, 27. ii. 2024, 9.64788°N, 83.84528°W, Springer &amp; Balke (CR_MB_2024_05) (MZUCR, ZSM) ; • 3 males, 3 females, 13 exx, Costa Rica, Páramo, Paraíso Quetzal, 2,760 m, Bilton (CBP, ZSM) ; • 6 exx., Costa Rica: Cartago, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.968&amp;materialsCitation.latitude=9.86012" title="Search Plazi for locations around (long -83.968/lat 9.86012)">Coris</a>, roadside ditch, 1,380 m, 28. ii. 2024, 9.86012°N, 83.96800°W, Springer &amp; Balke (CR_MB_2024_06) (MZUCR, ZSM) (incl. vouchers MB 10101 – MB 10108, MB 10278, MB 10279) ; • 6 exx., Costa Rica: Cartago, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-83.9882&amp;materialsCitation.latitude=9.86271" title="Search Plazi for locations around (long -83.9882/lat 9.86271)">Rio Coris</a>, creek, 1,390 m, 28. ii. 2024, 9.86271°N, 83.98820°W, Springer &amp; Balke (CR_MB_2024_07) (MZUCR, ZSM) ; • 14 exx., Costa Rica: Cartago, Alto de Ochomogo, 1,500 m, 20. xi. 1955, Malkin (CAS, ZSM) (incl. vouchers MB 9303, MB 9304) .</p><p>Description.</p><p>Habitus only with slight discontinuity between pronotum and elytra (as in Fig. 2 A, D), pronotum widest before base and more rounded laterally. TL: 2.2 mm; TL - h: 1.9 mm; TW: 1.0 mm.</p><p>Colour. Very dark brown to blackish dorsally and ventrally, pronotum of lighter colour towards lateral sides, lateral margin yellowish; elytra with lighter colour pattern in the form of longitudinal patches as in Fig. 2 D. Basal antennomeres and legs contrastingly of lighter colour (Fig. 2 D).</p><p>Surface sculpture. Head with few larger and smaller setiferous punctures (as in Fig. 2 A, D); with distinct microreticulation except on middle of head between the eyes. Pronotum and elytra shiny; with moderately dense and coarse setiferous punctation.</p><p>Structures. Antenna filiform. Head with occipital line; with rounded clypeus. Pronotum with distinct lateral bead; with distinct long and deep basal striae (as in Fig. 2 A, D). Elytra with distinct basal striae; without sutural line; without basal epipleural transverse carina. Metathoracic wings not examined in holotype (in paratype: fully developed).</p><p>Genitalia. Median lobe of aedeagus simply curved and relatively broad, narrowing towards apex; tip broadly rounded in ventral view, of characteristic spoon- or duck beaked shape; parameres of simple “ Bidessini ” type, two segmented (Fig. 2 C).</p><p>Female. Elytra with visible microreticulation on apical 1 / 3 to ½.</p><p>Variation. Size variation of the paratypes is TL 2.2–2.3 mm; TL - h 1.9–2.0 mm; TW 1.1–1.2 mm. The elytra may show contrasting pattern of yellowish bands in freshly hatched or generally very pale specimens. The beetles from Alto de Ochomogo (CAS) appear dark orange, although a darker colour pattern is still obvious. This appearance might be an artefact of longer storage and / or method of preservation. Our freshly collected specimens from the nearby Coris area are darker.</p><p>Etymology.</p><p>Named after the type locality, which is 650 m north-east from the Paraíso Quetzal Lodge.</p><p>Differential diagnosis.</p><p>This species can be distinguished from other species of Liodessus by the following combination of diagnostic characters: Occurrence in Costa Rica; shape of median lobe (tip in ventral view of characteristic spoon- or duck beaked shape).</p><p>Distribution.</p><p>Known from the central valley near San José and higher altitudes of Costa Rica nearby, up to around 2,800 m (Fig. 3).</p><p>Habitat.</p><p>Collected from small, shallow to very shallow puddles as well as edge of lagoon (Fig. 4 D) in Páramo, as well as densely vegetated roadside ditch (Fig. 4 E) and densely vegetated margin of creek (Rio Coris) (Fig. 4 F).</p></div>	https://treatment.plazi.org/id/6C0B8183C2D754A68B1613BFB6DA3D53	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Balke, Michael;Bilton, David;Springer, Monika;Villastrigo, Adrián;Neven, Katja;Hendrich, Lars	Balke, Michael, Bilton, David, Springer, Monika, Villastrigo, Adrián, Neven, Katja, Hendrich, Lars (2026): Two new species of Liodessus diving beetles from Costa Rica (Coleoptera, Dytiscidae, Bidessini). Alpine Entomology 10: 147-154, DOI: 10.3897/alpento.10.193709
