taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700566/files/figure.png	https://doi.org/10.5281/zenodo.18700566	FIG. 2. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MNHN.F.PPI282 (holotype) in dorsal view (A) with explanatory line drawing and detail of the vertex in dorsal view (B). Grey shading for hardened sediment. Scale bars: A, 100 mm; B, 20 mm. Photos: Olivier Lambert.	FIG. 2. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MNHN.F.PPI282 (holotype) in dorsal view (A) with explanatory line drawing and detail of the vertex in dorsal view (B). Grey shading for hardened sediment. Scale bars: A, 100 mm; B, 20 mm. Photos: Olivier Lambert.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700568/files/figure.png	https://doi.org/10.5281/zenodo.18700568	FIG. 3. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MNHN.F.PPI283 (paratype) in dorsal view with explanatory line drawing. Grey shading for hardened sediment; hatching for main break surfaces; dashed lines for more tentative interpretations of sutures and edges. Scale bar: 100 mm.	FIG. 3. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MNHN.F.PPI283 (paratype) in dorsal view with explanatory line drawing. Grey shading for hardened sediment; hatching for main break surfaces; dashed lines for more tentative interpretations of sutures and edges. Scale bar: 100 mm.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700570/files/figure.png	https://doi.org/10.5281/zenodo.18700570	FIG. 4. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MNHN.F.PPI282 (holotype) in right lateral view with explanatory line drawing. Grey shading for hardened sediment;hatching for main break surfaces;dashed lines for more tentative interpretations of sutures.Scale bar:100 mm. Photos:Olivier Lambert.	FIG. 4. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MNHN.F.PPI282 (holotype) in right lateral view with explanatory line drawing. Grey shading for hardened sediment;hatching for main break surfaces;dashed lines for more tentative interpretations of sutures.Scale bar:100 mm. Photos:Olivier Lambert.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700572/files/figure.png	https://doi.org/10.5281/zenodo.18700572	FIG. 5. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MNHN.F.PPI283 (paratype) in right (A) and left (B, without the anterior part of the rostrum) lateral view. Dashed lines for main sutures and bone edges. Scale bars: 100 mm. Photos: Olivier Lambert.	FIG. 5. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MNHN.F.PPI283 (paratype) in right (A) and left (B, without the anterior part of the rostrum) lateral view. Dashed lines for main sutures and bone edges. Scale bars: 100 mm. Photos: Olivier Lambert.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700574/files/figure.png	https://doi.org/10.5281/zenodo.18700574	FIG. 6. — Cranium of Mesokentriodon protohumboldti n.gen.,n. sp.MNHN.F.PPI282 (holotype) in ventral view (A) with explanatory line drawing and detail of right side of basicranium and palate in ventral view (B). Grey shading for hardened sediment;hatching for main break surfaces.Scale bars:100 mm.Photos:Olivier Lambert.	FIG. 6. — Cranium of Mesokentriodon protohumboldti n.gen.,n. sp.MNHN.F.PPI282 (holotype) in ventral view (A) with explanatory line drawing and detail of right side of basicranium and palate in ventral view (B). Grey shading for hardened sediment;hatching for main break surfaces.Scale bars:100 mm.Photos:Olivier Lambert.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700576/files/figure.png	https://doi.org/10.5281/zenodo.18700576	FIG. 7. — Cranium of Mesokentriodon protohumboldti n. gen.,n. sp.MNHN.F.PPI283 (paratype) in ventral view (A) and detail of left side of basicranium and palate in anteroventral and slightly lateral view (B). Dashed lines for main sutures,bone edges,and other anatomical features.Scale bars:100 mm.Photos:Olivier Lambert.	FIG. 7. — Cranium of Mesokentriodon protohumboldti n. gen.,n. sp.MNHN.F.PPI283 (paratype) in ventral view (A) and detail of left side of basicranium and palate in anteroventral and slightly lateral view (B). Dashed lines for main sutures,bone edges,and other anatomical features.Scale bars:100 mm.Photos:Olivier Lambert.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700578/files/figure.png	https://doi.org/10.5281/zenodo.18700578	FIG. 8. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MNHN.F.PPI282 (holotype) in anterior (A, lacking most of the rostrum), posterior (B), and right anterodorsolateral (C) views. Dashed lines for sutures. Scale bars: 100 mm. Photos: Olivier Lambert.	FIG. 8. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MNHN.F.PPI282 (holotype) in anterior (A, lacking most of the rostrum), posterior (B), and right anterodorsolateral (C) views. Dashed lines for sutures. Scale bars: 100 mm. Photos: Olivier Lambert.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700582/files/figure.png	https://doi.org/10.5281/zenodo.18700582	FIG. 9. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MNHN.F.PPI283 (paratype) in anterodorsal (A, lacking part of the rostrum) and left anterodorsolateral (B) views. Scale bars: 100 mm. Photos: Olivier Lambert.	FIG. 9. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MNHN.F.PPI283 (paratype) in anterodorsal (A, lacking part of the rostrum) and left anterodorsolateral (B) views. Scale bars: 100 mm. Photos: Olivier Lambert.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700584/files/figure.png	https://doi.org/10.5281/zenodo.18700584	FIG. 10. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MUSM4693 in dorsal (A), left lateral (B), and posterior (C) views. Scale bar: 100 mm. Photos: Olivier Lambert.	FIG. 10. — Cranium of Mesokentriodon protohumboldti n. gen., n. sp. MUSM4693 in dorsal (A), left lateral (B), and posterior (C) views. Scale bar: 100 mm. Photos: Olivier Lambert.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700586/files/figure.png	https://doi.org/10.5281/zenodo.18700586	FIG. 11. — Left periotic of Mesokentriodon protohumboldti n. gen., n. sp. MUSM4693 in ventral (A, with explanatory line drawing), medial (B), lateral (C), dorsal (D, with explanatory line drawing), posterior (E), anterior (F), and dorsomedial (G) views.Hatching for break surfaces. Scale bar: 10 mm. Photos: Olivier Lambert.	FIG. 11. — Left periotic of Mesokentriodon protohumboldti n. gen., n. sp. MUSM4693 in ventral (A, with explanatory line drawing), medial (B), lateral (C), dorsal (D, with explanatory line drawing), posterior (E), anterior (F), and dorsomedial (G) views.Hatching for break surfaces. Scale bar: 10 mm. Photos: Olivier Lambert.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700588/files/figure.png	https://doi.org/10.5281/zenodo.18700588	FIG. 12. — Partial left tympanic bulla of Mesokentriodon protohumboldti n. gen., n. sp. MUSM4693 in medial (A), dorsal (B), and ventral (C) view. Scale bar: 10 mm. Photos: Olivier Lambert.	FIG. 12. — Partial left tympanic bulla of Mesokentriodon protohumboldti n. gen., n. sp. MUSM4693 in medial (A), dorsal (B), and ventral (C) view. Scale bar: 10 mm. Photos: Olivier Lambert.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700590/files/figure.png	https://doi.org/10.5281/zenodo.18700590	FIG. 13. — Detached posterior lower right tooth of Mesokentriodon protohumboldti n. gen., n. sp. MUSM4693 in mesial (A), lingual (B), and distal (C) view. Scale bar: 10 mm. Photos: Olivier Lambert.	FIG. 13. — Detached posterior lower right tooth of Mesokentriodon protohumboldti n. gen., n. sp. MUSM4693 in mesial (A), lingual (B), and distal (C) view. Scale bar: 10 mm. Photos: Olivier Lambert.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700564/files/figure.png	https://doi.org/10.5281/zenodo.18700564	FIG. 1. — Schematic map of the East Pisco Basin, modified from Marx et al. (2017), showing the localities (red stars) where specimens of Mesokentriodon protohumboldti n. gen., n. sp. were collected:Mal Paso for the referred specimen MUSM4693 and Santa Rosa for both the holotype MNHN.F.PPI282 and the paratype MNHN.F.PPI283. Inset: schematic map of Peru with the position of the East Pisco Basin.	FIG. 1. — Schematic map of the East Pisco Basin, modified from Marx et al. (2017), showing the localities (red stars) where specimens of Mesokentriodon protohumboldti n. gen., n. sp. were collected:Mal Paso for the referred specimen MUSM4693 and Santa Rosa for both the holotype MNHN.F.PPI282 and the paratype MNHN.F.PPI283. Inset: schematic map of Peru with the position of the East Pisco Basin.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700592/files/figure.png	https://doi.org/10.5281/zenodo.18700592	FIG. 14. — Periotic, malleus, incus, tympanic bulla, and teeth of Incacetus broggii AMNH32656 (holotype): right periotic in ventral (A), medial (B), posterior (C), lateral (D), dorsal (E), and dorsomedial (F) views; right malleus in posteromedial view (G); right incus in medial view (H); two detached posterior teeth in mesial or distal view (I); right tympanic in medial (J), ventral (K), and lateral (L) views. Scale bars: A-F, I-L, 10 mm; G, H, 5 mm. Photos: Olivier Lambert.	FIG. 14. — Periotic, malleus, incus, tympanic bulla, and teeth of Incacetus broggii AMNH32656 (holotype): right periotic in ventral (A), medial (B), posterior (C), lateral (D), dorsal (E), and dorsomedial (F) views; right malleus in posteromedial view (G); right incus in medial view (H); two detached posterior teeth in mesial or distal view (I); right tympanic in medial (J), ventral (K), and lateral (L) views. Scale bars: A-F, I-L, 10 mm; G, H, 5 mm. Photos: Olivier Lambert.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
1A4AA5001C2BFD51D3ADFA2725AFF97D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/18700594/files/figure.png	https://doi.org/10.5281/zenodo.18700594	FIG. 15. — Phylogenetic relationships of Mesokentriodon protohumboldti n. gen., n. sp. with other early delphinidans: A, single most parsimonious tree resulting from the heuristic search based on the morphological matrix with the deletion of 45 taxa for which most of the periotic scores are lacking, with implied weights (K = 3) and with a molecular constraint as a backbone. Tree length = 4294; CI = 0.17;RI = 0.48.Mysticeti and early stem Odontoceti are omitted and Delphinoidea is collapsed for clarity (see complete tree in Appendix 4). The positions of Arimidelphis sorbinii (sister group to Lipotes vexillifer) and Parapontoporia sternbergi (sister group to Platanista gangetica) are extremely unlikely, as the former is a delphinid (Bianucci 2005) and the latter a lipotid (Muizon 1988b), as confirmed in many other analyses (including some in Appendix 4). B, early delphinidan part of the single most parsimonious tree resulting from the heuristic search based on the morphological matrix with the addition of five ‘kentriodontid’ s.l. species for which most of the periotic scores are lacking, with implied weights (K = 12) and with a molecular constraint as a backbone (see complete tree in Appendix 4). Tree length = 4353; CI = 0.17; RI = 0.48. Bootstrap and jackknife support values higher than 50 are shown respectively above and below the corresponding nodes.	FIG. 15. — Phylogenetic relationships of Mesokentriodon protohumboldti n. gen., n. sp. with other early delphinidans: A, single most parsimonious tree resulting from the heuristic search based on the morphological matrix with the deletion of 45 taxa for which most of the periotic scores are lacking, with implied weights (K = 3) and with a molecular constraint as a backbone. Tree length = 4294; CI = 0.17;RI = 0.48.Mysticeti and early stem Odontoceti are omitted and Delphinoidea is collapsed for clarity (see complete tree in Appendix 4). The positions of Arimidelphis sorbinii (sister group to Lipotes vexillifer) and Parapontoporia sternbergi (sister group to Platanista gangetica) are extremely unlikely, as the former is a delphinid (Bianucci 2005) and the latter a lipotid (Muizon 1988b), as confirmed in many other analyses (including some in Appendix 4). B, early delphinidan part of the single most parsimonious tree resulting from the heuristic search based on the morphological matrix with the addition of five ‘kentriodontid’ s.l. species for which most of the periotic scores are lacking, with implied weights (K = 12) and with a molecular constraint as a backbone (see complete tree in Appendix 4). Tree length = 4353; CI = 0.17; RI = 0.48. Bootstrap and jackknife support values higher than 50 are shown respectively above and below the corresponding nodes.	2026-02-19	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni		Zenodo	biologists	Lambert, Olivier;Oberlin, Maëlle;Muizon, Christian de;Collareta, Alberto;Urbina, Mario;Bianucci, Giovanni			
