identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
321A5CF67F3B50B691C9B4BFC9D1541B.text	321A5CF67F3B50B691C9B4BFC9D1541B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sellanucheza ancorata Likhitrakarn & Golovatch & Lips & Panha & Sutcharit 2026	<div><p>Sellanucheza ancorata sp. nov.</p><p>Figs 11, 12, 13</p><p>Type material.</p><p>• Holotype ♂ (SM-08), Laos, Khammouane Province, 2.5 km west-northwest of Ban Tathot, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.12381&amp;materialsCitation.latitude=17.631529" title="Search Plazi for locations around (long 105.12381/lat 17.631529)">Tham Kamouk</a>, large halled limestone cave, 200 m a. s. l., 17°37'53.5"N, 105°07'25.7"E, 06. 08. 2016, leg. P. Jäger .</p><p>Diagnosis.</p><p>The new species is distinguished from all congeners by the unique, anchor-shaped (trifid) solenophore tip. Somatically, it differs from the two other Laotian new species above by its intermediate size (ca 21 mm) and uniform brown yellowish coloration (vs small and pallid in S. laotica sp. nov.; large and dark in S. longispina sp. nov., lacking the specific sternal modifications found in either species). The gonopods (Figs 13, 14) are highly characteristic: process pb short, stout and laminiform (vs large, laminiform and bifid in S. longispina sp. nov., long and subunciform in S. jaegeri, or small and acute in S. tenebra); process pa distinctively long, slender and spiniform with an acute tip directed ventromesad (vs shorter, spiniform and suberect in S. laotica sp. nov., or large and acute in S. grandis) and the solenophore tip is split into three distinct prongs (central straight, lateral recurved), a characteristic unique to this species.</p><p>Description.</p><p>Length 20.8 mm (♂), width of midbody pro- and metazona 1.6 and 2.1 mm (♂), respectively.</p><p>Coloration of alcohol material after nine years of preservation uniform brown yellowish (Fig. 12 A – G); legs and antennae a little paler, tip of antenna dark brown.</p><p>Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae moderately long, reaching body ring 3 (♂) when stretched dorsally. In width, head &lt;ring 3 &lt;4 &lt;collum &lt;ring 2 &lt;5 &lt;6 &lt;7–17 (♂); thereafter body gently and gradually tapering.</p><p>Collum with three transverse rows of setae: 4 + 4 anterior, 2 + 2 intermediate and 3 + 3 posterior; lateral margin with a small incision in posterior half; posterior corner of paraterga very narrowly rounded, slightly declined ventrad, extending past tergal margin (Fig. 12 A, B).</p><p>Tegument smooth and shining, prozona very finely shagreened, metaterga smooth and faintly rugulose (Fig. 12 A – G); surface below paraterga finely microgranulate (Fig. 12 B, D, E). Postcollum metaterga with two transverse rows of setae: 2 + 2 in anterior (pre-sulcus) and 3 + 3 in posterior (post-sulcus) row. Tergal setae long, strong, slender, ca 1 / 3 metatergal length. Axial line faint, barely traceable on metaterga.</p><p>Paraterga well developed (Fig. 12 A, C, F), set high (at upper 1 / 3 of body), subhorizontal, lying below dorsum; anterior edge broadly rounded and narrowly bordered, fused to callus; lateral edge without incisions; posterior corner very narrowly rounded, not extending past tergal margin except for rings 17–19 where it becomes increasingly pointed (Fig. 12 E, F); posterior edge nearly straight. Paraterga 2 broad, slightly upturned posteriorly, anterior edge rounded, lateral edge with a small incision in anterior half; posterior edge concave (Fig. 12 B). Calluses on paraterga narrow, each demarcated by a sulcus only dorsally. Ozopores evident, lateral, each lying in an ovoid groove at ca 1 / 4 metatergal length in front of posterior corner (Fig. 12 B, D, E).</p><p>Transverse sulci distinct (Fig. 12 A, C, F), complete on metaterga 6–18, narrow, line-shaped, shallow, not reaching the bases of paraterga, smooth at bottom, incomplete and nearly wanting on ring 5. Stricture between pro- and metazona narrow, beaded at bottom down to base of paraterga (Fig. 12 A, C). Pleurosternal carinae complete crests on anterior rings, thereafter gradually reduced posteriad until ring 16 (♂) (Fig. 12 B, D, E).</p><p>Epiproct (Fig. 12 E – G) conical, rounded dorsoventrally, with two small and evident apical papillae; tip subtruncate; pre-apical papillae small, lying close to tip (Fig. 12 F, G). Hypoproct roundly subtriangular (Fig. 12 G), setiferous knobs at posterior edge well-separated and evident.</p><p>Sterna sparsely setose, cross-impressions shallow, without modifications; a single, linguiform, densely setose, sternal lobe between ♂ coxae 4 (Fig. 12 H, I). Legs moderately long and slender, midbody legs ca 1.1–1.3 × as long as body height; ♂ prefemora without modifications; tarsal brushes present until ♂ ring 16.</p><p>Gonopods (Figs 13, 14) simple. Coxite long, stout, subcylindrical, relatively straight, but slightly curved dorsally, with a dense group of setae distodorsally (Figs 13 A, 13 B, 13 C, 14 A, 14 B). Prefemorite (pfe) densely setose, as usual, relatively short, ca 1 / 3 length of acropodite (Figs 13 A, 13 B, 13 C, 14 A, 14 B). Femorite (fe) stout, relatively long, suberect, with a distinct. clearly visible lateral suture (Figs 13 A – D, 14 A, 14 B). Distal part of femorite supplied with two distinct processes: process pb (pb) short, stout, laminiform, elevated dorsally, with a rounded apical margin (Figs 13 A – C, 13 G, 13 F, 14 A, 14 B); process pa distinctively long, slender, spiniform, clearly curved, with an acute tip directed ventromesad (Figs 13 A, 13 B, 13 D, 13 G, 14 A). Solenophore (sph) long, slender, directed anteriad and slightly curved; tip highly characteristic, trifid; central prong straight to slightly curved, while two lateral processes recurved and pointing basally (Figs 13 A – G, 14 A, 14 B). Solenomere long, slender, almost entirely sheathed by solenophore, forming a distinct anchor shape (Figs 13 G, 14 A, 14 B).</p><p>Remarks.</p><p>It seems noteworthy that the natural structure of the gonopod is clearly shown in Fig. 14, 13 G. The structure depicted in most other views of Fig. 13 exhibits a distortion caused by the drying of the specimen during preparation. This process caused the solenomere (sl) to dislodge from the solenophore sheath and project posteriorly, an artifact that does not reflect the true morphology of the species. Geographically, Sellanucheza ancorata sp. nov. is currently known only from the type locality in Khammouane Province. This is a large-halled limestone cave (Fig. 15 D, E) which is not visited by tourists, although it is occasionally used by villagers as a shortcut between villages. The end of the cave features a ceiling breakdown with massive litter accumulation from the forest above (Fig. 15 C). In the rainy season, a river flows through the cave, and bats are present. The species appears to be narrowly endemic to its limestone karst system of the area, a pattern characteristic of many members of this genus in the Indochinese Peninsula.</p><p>Etymology.</p><p>To emphasize the characteristic anchor-shaped (trifid) tip of the solenophore; adjective.</p></div>	https://treatment.plazi.org/id/321A5CF67F3B50B691C9B4BFC9D1541B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Lips, Josiane;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Lips, Josiane, Panha, Somsak, Sutcharit, Chirasak (2026): Review of the millipede genus Sellanucheza Enghoff, Golovatch & Nguyen, 2004, with descriptions of three new species from Laos and redescription of S. variata (Attems, 1953) (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1282: 339-369, DOI: 10.3897/zookeys.1282.186746
0BED9F91B91D5AECBD9DF7BC976231AC.text	0BED9F91B91D5AECBD9DF7BC976231AC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sellanucheza Enghoff, Golovatch & Nguyen 2004	<div><p>Genus Sellanucheza Enghoff, Golovatch &amp; Nguyen, 2014</p><p>Szechuanella Hoffman 1961: 533 (D), preoccupied by Szechuanella Lu, 1959 .</p><p>Szechuanella – Hoffman 1963: 586 (M, K); Jeekel 1968: 62 (M, R); Golovatch 1984: 57 (M); Shelley et al. 2000: 132 (M); Sierwald 2009: 117 (L).</p><p>Sellanucheza Enghoff et al. 2004: 39 (M, L), nomen novum.</p><p>Cemsunguria Özdikmen 2007: 433 (M), junior objective synonym (synonymized by Nguyen 2011).</p><p>Sellanucheza – Golovatch 2013 a: 22 (M, K); 2013 b: 330 (M, K); Nguyen and Sierwald 2013: 1295 (M, L).</p><p>Diagnosis</p><p>(amended after Hoffman 1961 and Nguyen 2011). Sellanucheza is a member of the tribe Sulciferini and can be distinguished from all other genera in the tribe by the following combination of gonopodal characters: (1) telopodite provided with two or three large postfemoral processes; (2) both lamina medialis and lamina lateralis being well-developed; (3) lack of a clear demarcation between the femorite and the postfemoral region; and (4) solenophore distinctly curved ventrad to form a circular or subcircular structure together with the femorite, and bearing one or two basal processes.</p><p>General morphological characters.</p><p>A genus of medium to large-sized Paradoxosomatidae (body length ca 11–59 mm) with 20 rings and a normal pore formula. Paraterga variable, ranging from vestigial to strongly developed on midbody rings. Coloration typically dark brown to blackish but also including uniform light yellowish to pallid forms. Transverse sulcus usually distinct on metaterga. Sternal modifications in males always involving a large sternal lobe between coxae 4 on ring 5; additional modifications may include transverse lobes or paramedian cones on both rings 5 and 6, or well-developed ventroposterior tubercles near the coxae starting from ring 6 onwards. Male legs without adenostyles; tarsal brushes present, sometimes extending as far as ring 17. Epiproct long and flattened, with a pair of distinct lateral dentiform tubercles.</p><p>Gonopods relatively simple; coxite long, slender and cylindrical, with a dense group of setae distodorsally. Prefemoral part (pfe) short and densely setose, ca 1 / 3–1 / 5 length of acropodite. Femorite (fe) long and slender or relatively stout and curved, often with a distinct mesal groove. Distal part of femorite or base of solenophore (sph) supplied with two distinct processes: process pa typically long, spiniform, slender and acute, often with its tip curved ventrad, although sometimes short; process pb usually shorter, varying from a short acute spine or a long projection to a small or large lamina. Solenophore relatively long, coiled or curved to sheath the usually long and flagelliform solenomere (sl); distal margin of solenophore highly variable, ranging from truncate and subquadrate to deeply bifid or uniquely anchor-shaped (trifid).</p><p>Type species.</p><p>Szechuanella tenebra Hoffman, 1961, by original designation.</p><p>Other species included.</p><p>Sellanucheza tenebra (Hoffman, 1961), S. variata (Attems, 1953), S. grandis (Golovatch, 1984), S. hoffmani Nguyen, 2011, S. jaegeri Golovatch, 2013, S. typica Golovatch, 2013, S. laotica sp. nov., S. longispina sp. nov., S. ancorata sp. nov.</p><p>Remarks.</p><p>The genus name Sellanucheza was established by Enghoff et al. (2004) as a replacement name for the preoccupied Szechuanella Hoffman, 1961 (non Lu, 1959). Subsequently, Cemsunguria was proposed as an unnecessary replacement name by Özdikmen (2007), making it an instantaneous junior objective synonym of Sellanucheza under Article 23.1 of the ICZN (Nguyen 2011; Nguyen and Sierwald 2013). With the addition of three new species described in this study, the genus Sellanucheza now comprises nine formally described species. These new discoveries in the limestone karst systems of Khammouane and Bolikhamsay provinces, northeastern Laos (Fig. 16), represent a significant range extension for the genus, which was previously known only from southern China and northern and central Vietnam (Hoffman 1961; Golovatch 1984, 2013 a; Nguyen 2011).</p></div>	https://treatment.plazi.org/id/0BED9F91B91D5AECBD9DF7BC976231AC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Lips, Josiane;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Lips, Josiane, Panha, Somsak, Sutcharit, Chirasak (2026): Review of the millipede genus Sellanucheza Enghoff, Golovatch & Nguyen, 2004, with descriptions of three new species from Laos and redescription of S. variata (Attems, 1953) (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1282: 339-369, DOI: 10.3897/zookeys.1282.186746
85D538798BDE5EA993A715B3383699B5.text	85D538798BDE5EA993A715B3383699B5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sellanucheza grandis (Golovatch 1984)	<div><p>Sellanucheza grandis (Golovatch, 1984)</p><p>Fig. 4</p><p>Szechuanella grandis Golovatch 1983: 182 (nomen nudum).</p><p>Szechuanella grandis Golovatch 1984: 56 (D); Korsós and Golovatch 1989: 212 (R).</p><p>Sellanucheza grandis – Enghoff et al. 2004: 39 (L, R); Nguyen 2011: 59 (D, R); Golovatch 2013 a: 22 (M, K); 2013 b: 330 (M, K); Nguyen and Sierwald 2013: 1296 (L).</p><p>Material examined.</p><p>3 ♂, 4 ♀, Vietnam, Ninh binh, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.60833&amp;materialsCitation.latitude=20.31778" title="Search Plazi for locations around (long 105.60833/lat 20.31778)">Cuc Phuong National Park</a>, 300 m a. s. l., 20°19'4"N, 105°36'30"E, 26. 07. 2006, leg. S. Panha and C. Sutcharit .</p><p>Descriptive notes.</p><p>Length 54.3–56.2 (♂) or 51.8–57.3 mm (♀), width of midbody pro- and metazonae 4.8–5.2 and 5.7–6.1 mm (♂) or 5.6–6.1 and 6.7–7.0 mm (♀), respectively. Antennae moderately long, reaching body ring 5 (♂) or 4 (♀) when stretched dorsally. Sterna sparsely setose, shining, shallow cross-impressions, without modifications; a large, central, slightly bifid, setose lobe between ♂ coxae 4 (Fig. 4 D). Legs moderately long and slender, midbody legs ca 1.1–1.4 (♂) or 1.0–1.2 × (♀) as long as body height; ♂ prefemora without modifications; tarsal brushes present until ♂ ring 11.</p><p>Distribution.</p><p>Vietnam, Ninh Binh, Cuc Phuong National Park (Golovatch 1984; Korsós and Golovatch 1989; Enghoff et al. 2004; Nguyen 2011); Ha Tinh Province, Huong Son District, Son Kim commune, secondary forest; HaTinh Province, Huong Son District, Cau Treo, secondary forest; Phu Tho Province, Xuan Son National Park, forest, 500 m a. s. l.; Nghe An Province, Pu Mat National Park, Khe Thoi, closed forest, near stream, 18°58'17.4"N, 104°48'20.9"E (Nguyen 2011).</p><p>Remarks.</p><p>Originally described as Szechuanella grandis from a limestone hill in Cuc Phuong National Park, Ninh Binh Province, Vietnam (Golovatch 1984), this species was transferred to Sellanucheza by Enghoff et al. (2004). The name Szechuanella grandis was first introduced as a nomen nudum by Golovatch (1983), as it appeared in a species list without an accompanying description or diagnosis. The species was subsequently validated and formally described by Golovatch (1984). Somatically, S. grandis differs clearly from congeners, including S. tenebra and S. variata, by the considerably larger body size. In contrast to S. tenebra, where paraterga are absent from rings 3–19, S. grandis retains abbreviated paraterga demarcated by distinct dorsal sulci.</p><p>The gonopods are highly distinctive, the solenophore (sph) being conspicuously shorter, at most half as long as the strongly coiled femorite (fe) (Fig. 4 A, B). This character immediately separates S. grandis from both S. variata and S. tenebra, in which the solenophore and femorite are subequal in length (Figs 11 A, 3 A – C). The femorite is relatively straight, being only slightly curved (Fig. 4 A, B). The postfemoral part is supplied with a small, dentiform, sublateral spine (d) (Fig. 4 A, B), a large and pointed process pb (Fig. 4 A – C), as well as a large, acute, mesal process pa (Fig. 4 A, C). In addition, the solenophore is short, coiled, the tip being only faintly bifid and consisting of small, rounded, apical lobules (Fig. 4 A, C).</p><p>The new material reported above fully agrees with the original description (Golovatch 1984), showing only slight variations in standard measurements (see “ Descriptive notes ” above). In contrast to other congeners which appear to be highly localized, S. grandis seems to be relatively widespread, the maximum distance between the known localities amounting to ca 300 km. Although the distribution range has recently been extended from the type locality in Ninh Binh Province to also cover Ha Tinh, Phu Tho, and Nghe An Provinces, the species still remains endemic to northern Vietnam.</p></div>	https://treatment.plazi.org/id/85D538798BDE5EA993A715B3383699B5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Lips, Josiane;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Lips, Josiane, Panha, Somsak, Sutcharit, Chirasak (2026): Review of the millipede genus Sellanucheza Enghoff, Golovatch & Nguyen, 2004, with descriptions of three new species from Laos and redescription of S. variata (Attems, 1953) (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1282: 339-369, DOI: 10.3897/zookeys.1282.186746
4C474559951C5EA598695A1577B70CA9.text	4C474559951C5EA598695A1577B70CA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sellanucheza hoffmani Nguyen 2011	<div><p>Sellanucheza hoffmani Nguyen, 2011</p><p>Fig. 5 A – C</p><p>Sellanucheza hoffmani Nguyen 2011: 59 (D, K).</p><p>Sellanucheza hoffmani – Golovatch 2013 b: 330 (M, K); Nguyen and Sierwald 2013: 1296 (L).</p><p>Distribution.</p><p>Vietnam, Kon Tum Province, Loxo Pass, ca 80 km north of Kontum, secondary forest, 800 m a. s. l.; Quang Binh Province, Minh Hoa District, Thuong Hoa commune, Phong Nha-Ke Bang region (Nguyen 2011).</p><p>Remarks.</p><p>This species is currently known only from central Vietnam. Somatically, this species is one of the largest members of the genus (length 52–59 mm, width 5.3–6.4 mm), being comparable in size only to S. grandis . However, it is readily distinguished from all congeners by male sternum 5 showing two separate, small, setose cones between coxae 4, and by the paraterga being small, but complete, represented by keel-shaped ridges demarcated by distinct sulci both dorsally and ventrally. The gonopods are particularly distinctive, the femorite (fe) being long, marginally expanded distad, with the distal part being simultaneously twisted and curved ventrad (Fig. 5 A, B); the postfemoral region bears a long, basal, laminiform process pb (Fig. 5 A – C); and the solenophore (sph) is supplied with a very long, nearly straight, spiniform process pa at the base of the lamina medialis (Fig. 5 A, C).</p></div>	https://treatment.plazi.org/id/4C474559951C5EA598695A1577B70CA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Lips, Josiane;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Lips, Josiane, Panha, Somsak, Sutcharit, Chirasak (2026): Review of the millipede genus Sellanucheza Enghoff, Golovatch & Nguyen, 2004, with descriptions of three new species from Laos and redescription of S. variata (Attems, 1953) (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1282: 339-369, DOI: 10.3897/zookeys.1282.186746
9EC4FCDD19615BC7B68AE2565224E666.text	9EC4FCDD19615BC7B68AE2565224E666.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sellanucheza jaegeri Golovatch 2013	<div><p>Sellanucheza jaegeri Golovatch, 2013</p><p>Fig. 5 D, E</p><p>Sellanucheza jaegeri Golovatch 2013 a: 20 (D, K).</p><p>Sellanucheza jaegeri – Golovatch 2013 b: 330 (M, K); Nguyen and Sierwald 2013: 1296 (L).</p><p>Distribution.</p><p>China, Shaanxi Province, Taibai Shan, southern slope, above Houshenzi, 1,300 –1,700 m a. s. l., 33°51'N, 107°50'E (Golovatch 2013 a).</p><p>Remarks.</p><p>Sellanucheza jaegeri was described by Golovatch (2013 a) from the Taibai Shan, Shaanxi Province, China. Somatically, the species is characterized by very poorly developed paraterga which are demarcated by sulci only dorsally. The gonopods are particularly distinctive, the femorite (fe) being suberect, slender and elongate. The solenophore (sph) is relatively short, strongly coiled and deeply bifid at tip (Fig. 5 D, E). In addition, the base of the solenophore is supplied with two prominent processes, pa and pb, both being long, subunciform and directed distad (Fig. 5 D, E). The species is currently known only from the type locality in Shaanxi Province, thus being endemic to northern China.</p></div>	https://treatment.plazi.org/id/9EC4FCDD19615BC7B68AE2565224E666	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Lips, Josiane;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Lips, Josiane, Panha, Somsak, Sutcharit, Chirasak (2026): Review of the millipede genus Sellanucheza Enghoff, Golovatch & Nguyen, 2004, with descriptions of three new species from Laos and redescription of S. variata (Attems, 1953) (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1282: 339-369, DOI: 10.3897/zookeys.1282.186746
35952EB12C885DF0BC9E2701C7004591.text	35952EB12C885DF0BC9E2701C7004591.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sellanucheza laotica Likhitrakarn & Golovatch & Lips & Panha & Sutcharit 2026	<div><p>Sellanucheza laotica sp. nov.</p><p>Figs 6, 7, 8</p><p>Type material.</p><p>• Holotype ♂ (CUMZ -11647), Laos, Khammouane Province, Nong Ping, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.89361&amp;materialsCitation.latitude=17.370556" title="Search Plazi for locations around (long 105.89361/lat 17.370556)">Tham Nguen Mai</a>, 230 m a. s. l., 17°22'14"N, 105°53'37"E, 24. 02. 2016, leg. J. Lips . • Paratypes 3 ♂, 4 ♀ (CUMZ -11647), same locality, together with holotype .</p><p>Diagnosis.</p><p>The new species is distinguished from all known congeners by the following combination of characters. Somatically, it differs from the Chinese S. tenebra, S. typica and S. jaegeri by the strongly developed paraterga on midbody rings (vs paraterga virtually absent or very poorly developed). From the large Vietnamese S. grandis and S. hoffmani, it is separated by the significantly smaller body size and the uniform, light yellowish to pallid coloration. A unique apomorphy seems to be the presence of well-developed ventroposterior tubercles on sterna near coxae, starting with ring 6. The gonopods are characteristic in the relatively short solenophore showing a truncate, subquadrate distal margin and an acute ventral angle. In addition, the species differs from S. variata and S. jaegeri by process pb being short, blunt and suberect (vs long, slender and acute or subunciform), and by the short, relatively stout solenomere which is subequal in length to the solenophore (vs long and flagelliform in other congeners).</p><p>Description.</p><p>Length 11.3–13.6 (♂) or 12.1–13.2 mm (♀), width of midbody pro- and metazona 0.8–1.2 and 1.2–1.4 mm (♂) or 0.9–1.2 and 1.2–1.4 mm (♀), respectively.</p><p>Coloration of alcohol material after nine years of preservation uniform light yellowish to pallid; antennae and legs slightly paler (Fig. 6 A – I).</p><p>Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae moderately long (Fig. 6 A, B), reaching body ring 4 (♂) or 3 (♀) when stretched dorsally. In width, head &lt;ring 3 &lt;4 &lt;collum &lt;ring 5 &lt;6 &lt;2 &lt;7–17 (♂, ♀) (Fig. 6 A); thereafter body gently and gradually tapering. Collum with three transverse rows of setae: 4 + 4 anterior, 2 + 2 intermediate and 3 + 3 posterior; lateral margin with a small incision laterally in posterior half; posterior corner of paraterga very narrowly rounded, slightly declined ventrad, not extending past tergal margin (Fig. 6 A, B).</p><p>Tegument smooth and shining (Fig. 6 A – G), prozona very finely shagreened, metaterga smooth and leathery; surface below paraterga finely microgranulate (Fig. 6 B, D, E). Postcollum metaterga with two transverse rows of setae: 3 + 3 in anterior (pre-sulcus) and 3 + 3 in posterior (post-sulcus) row. Tergal setae long, strong, slender, ca 1 / 3 metatergal length. Axial line faint, barely traceable on metaterga. Paraterga strongly developed (Fig. 6 A – G), especially so in ♂, set at ca 1 / 3 midbody height, slightly upturned, but all lying below dorsum; shoulders well-developed, mostly rounded; posterior corner almost completely to very broadly rounded, all extending past tergal margin, posterior edge slightly concave, bent ventrad on rings 18 and 19 (Fig. 6 F).</p><p>Paraterga 2 broad, anterior edge angular, lateral edge with two evident incisions in anterior half; posterior edge oblique (Fig. 6 A, B). Lateral edges of paraterga 3 and 4 with two evident incisions: one in anterior 1 / 3, the other at midway (Fig. 6 A). Following paraterga with three evident incisions: largest one in anterior 1 / 3, one at midway, and another near posterior corner (Fig. 6 A, C). Paraterga thin blunt blades in lateral view, slightly thicker only on pore-bearing rings (Fig. 6 B, D, E). Calluses on paraterga demarcated by sulci only dorsally. Ozopores evident, lateral, each lying in an ovoid groove at ca 1 / 4 in front of posterior corner (Fig. 6 D, E). Transverse sulcus usually distinct, slightly incomplete on ring 4, complete and clearly visible on metaterga 5–18, usually narrow, shallow (Fig. 6 A, C, F), superficial (due to coarse texture), not reaching bases of paraterga, slightly better developed in ♀. Stricture between pro- and metazona narrow, ribbed at bottom down to base of paraterga (Fig. 6 A, C, F). Pleurosternal carinae complete crests, each usually with a sharp posterior tooth until ring 17, crests bulged anteriorly and with a small sharp posterior tooth each on rings 18 and 19 (♂, ♀) (Fig. 6 B, D, E).</p><p>Epiproct (Fig. 6 E – G) conical, rounded dorsoventrally, with two small apical papillae; tip subtruncate; pre-apical papillae evident, lying close to tip (Fig. 6 F). Hypoproct roundly subtrapeziform (Fig. 6 G), setiferous knobs at posterior edge well-separated and evident.</p><p>Sterna sparsely setose, cross-impressions shallow; starting on ring 6 with a pair of well-developed, ventroposterior tubercles near each coxa; a large, central, cordiform, densely setose lobe between ♂ coxae 4 (Fig. 6 H, I). Legs moderately long and slender, midbody legs ca 1.0–1.3 (♂) or 0.9–1.2 × (♀) as long as body height, ♂ prefemora without modifications, tarsal brushes present until ♂ legs 4.</p><p>Gonopods (Figs 7, 8) simple. Coxite long, slender, cylindrical, slightly curved posteriorly, densely setose distodorsally. Prefemorite (pfe) densely setose, as usual, ovoid, elongate, ca 1 / 3–1 / 4 length of acropodite (Figs 7 A, 7 B, 7 F, 7 G, 8 A – C). Femorite (fe) long, slender, curved (Fig. 7 B, F, G, 8 A, B); distal part supplied with a short, blunt, suberect process pb (Figs 7 B – I, 8 B, 8 C) and a short, slender, spiniform, slightly curved, mesal process pa (Figs 7 A, 7 C – G, 7 I, 8 A – D). Solenophore (sph) relatively short, supporting solenomere (sl), distal margin truncate, subquadrate, with an acute ventral angle (Figs 7 C – F, 8 C). Solenomere short, subequal in length to solenophore, relatively stout, tip acute and slightly projecting distad (Figs 7 A, 7 E – I, 8 A – D).</p><p>Remarks.</p><p>Sellanucheza laotica sp. nov. represents the first record of the genus from a limestone cave in Khammouane Province, Laos. The distinct pallid coloration strongly suggests a troglobiomorphic adaptation to its subterranean existence, a characteristic frequently observed in other cave-dwelling millipedes within the region (Likhitrakarn et al. 2017, 2021, 2022, 2024; Liu and Wynne 2019; Golovatch and Liu 2020). The new species appears to be narrowly endemic to its karst cave systems in northeastern Laos.</p><p>Etymology.</p><p>The specific epithet is a Latin adjective referring to Laos, the country of origin where the type material was collected.</p></div>	https://treatment.plazi.org/id/35952EB12C885DF0BC9E2701C7004591	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Lips, Josiane;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Lips, Josiane, Panha, Somsak, Sutcharit, Chirasak (2026): Review of the millipede genus Sellanucheza Enghoff, Golovatch & Nguyen, 2004, with descriptions of three new species from Laos and redescription of S. variata (Attems, 1953) (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1282: 339-369, DOI: 10.3897/zookeys.1282.186746
4A258ECA2C055A2D9CE2B31EF528EBEF.text	4A258ECA2C055A2D9CE2B31EF528EBEF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sellanucheza longispina Likhitrakarn & Golovatch & Lips & Panha & Sutcharit 2026	<div><p>Sellanucheza longispina sp. nov.</p><p>Figs 9, 10, 11</p><p>Type material.</p><p>• Holotype ♂ (SMF), Laos, Bolikhamsay Province, Lak Sao, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.81275&amp;materialsCitation.latitude=18.22114" title="Search Plazi for locations around (long 104.81275/lat 18.22114)">Tham Mankhone</a>, large halled tourist cave (Fig. 15 A, B), with some electric lights at the entrance, 501 m a. s. l., 18°13'16.1"N, 104°48'45.9"E, 21. 07. 2016, leg. P. Jäger .</p><p>Diagnosis.</p><p>The new species is distinguished from congeners by its large size (ca 49 mm) and the unique presence of sternal cones on both rings 5 and 6 (vs usually either ring 5 alone or ventroposterior tubercles from ring 6 on in S. laotica sp. nov.). It also differs from the pallid S. laotica sp. nov. by its dark blackish brown coloration. The gonopods are characteristic: process A (pa) long, spiniform and subequal in length to the solenophore (vs shorter, spiniform and suberect in S. laotica sp. nov., or the solenophore much shorter in S. grandis); process B (pb) large, laminiform and bifid (vs small and acute in S. variata and S. grandis); the solenophore is stout and deeply bifid, while the solenomere is flagelliform (vs stout in S. laotica sp. nov.).</p><p>Description.</p><p>Length 48.6 mm (♂), width of midbody pro- and metazona 4.8 and 5.7 mm (♂), respectively.</p><p>Coloration of alcohol material after nine years of preservation, faded to blackish brown (Fig. 9 B – H), posterior halves of metaterga, paraterga and epiproct pale light brown to yellowish (Fig. 9 A – F), head and antennae dark brown, tip of antenna dark brown, venter and legs pale brown to yellowish (Fig. 9 G – I).</p><p>Clypeolabral region densely setose (Fig. 9 B), vertex sparsely so, epicranial suture distinct. Antennae moderately long, reaching body ring 4 (♂) when stretched dorsally. In width, head &lt;ring 4 &lt;3 &lt;5 &lt;6 &lt;7 &lt;collum &lt;ring 2 &lt;8 &lt;9–17 (♂) (Fig. 9 A); thereafter body gently and gradually tapering.</p><p>Collum semilunar, setation pattern untraceable; surface rugulose, middle slightly flattened; paraterga rounded and subtriangular, devoid of lateral incisions; posterior corner very narrowly rounded, not extending past tergal margin (Fig. 9 A, B).</p><p>Tegument smooth and shining, prozona finely shagreened, metaterga smooth and leathery; surface below paraterga finely microgranulate (Fig. 9 A – G). Postcollum metaterga with two transverse rows of setae traceable at least as insertion points: 3 + 3 in anterior (pre-sulcus) and 4 + 4 in posterior (post-sulcus) row; posterior row barely traceable. Axial line visible both on pro- and metazona.</p><p>Paraterga well developed (Fig. 9 A – E, F), set high (at upper 1 / 3 of body), slightly upturned, but lying below dorsum; anterior edge broadly rounded and narrowly bordered, fused to callus; lateral edge without incisions; posterior corner very narrowly rounded, not produced past rear tergal margin except for rings 2 and 3 (Fig. 9 A, B); posterior edge nearly straight. Paraterga 2 broad, anterior edge angular and rounded, lateral edge without incisions (Fig. 9 A). Calluses on paraterga narrow, demarcated by a sulcus only dorsally. Ozopore evident, lateral, lying in an ovoid groove at ca 1 / 3 metatergal length in front of posterior corner (Fig. 9 B, D, E).</p><p>Transverse sulcus distinct (Fig. 9 A, C, F), complete on metaterga 5–18, narrow, line-shaped, deep, reaching bases of paraterga, faintly ribbed at bottom, incomplete and nearly wanting on rings 4 and 19 (Fig. 9 A, F). Stricture between pro- and metazona narrow, ribbed at bottom down to base of paraterga (Fig. 9 C, D). Pleurosternal carinae complete high crests, each with a sharp posterior tooth on rings 2–4, thereafter increasingly strongly reduced and remaining visible only as a bulge anteriorly until ring 7 (Fig. 9 B, D, E).</p><p>Epiproct (Fig. 9 E – G) conical, rounded dorsoventrally, with two small apical papillae; tip subtruncate; pre-apical papillae evident, lying close to tip (Fig. 9 G). Hypoproct nearly semi-circular (Fig. 9 G), posterior tip broadly rounded, setiferous knobs at posterior edge very small and moderately well separated.</p><p>Sterna sparsely setose, without modifications; cross-impressions shallow (Fig. 9 G). Rings 5 and 6 each with a paramedian pair of evident, basally contiguous cones located anteriorly near each coxa; cones between coxae 4 and 6 being more prominent and larger than those between coxae 5 and 7 (Fig. 9 H, I). A paramedian pair of small, but evident tubercles in front of gonopod aperture. Legs moderately long and slender, midbody legs ca 1.1–1.3 × as long as body height; ♂ prefemora without modifications; tarsal brushes present until ♂ ring 12.</p><p>Gonopods (Figs 10, 11) relatively simple, clearly coiled. Coxite long, slender, cylindrical, suberect, densely setose distodorsally (Figs 10 A, 10 D, 11 A, 11 B). Prefemorite (pfe) densely setose, as usual, ovoid, elongate, short, ca 1 / 4–1 / 5 length of acropodite (Figs 10 A – E, 11 A – C). Femorite (fe) relatively stout, long, curved dorsad (Figs 10 A, 10 B, 10 D, 10 E, 11 A, 11 B); distal part supplied with a prominent, stout, laminiform, linguiform, apically rounded process pb directed anteriad, the latter being bifid (Figs 10 A – F, 11 A – C), with ventral lobule further split into two small acute points (Figs 10 A – C, 10 F, 11 C). Process pa conspicuous, spiniform, long, slender, acute, subequal in length to solenophore, curved anteromesad (Figs 10 A – C, 11 A, 11 C). Solenophore (sph) long, stout, strongly coiled, deeply bifid distally, both lobules being blunt or rounded (Figs 10 A – E, 11 A, 11 C); solenophore almost completely sheathing solenomere. Solenomere (sl) long, slender, flagelliform, tip acute and slightly projecting distad past solenophore (Figs 10 A – E, 11 A, 11 C).</p><p>Remarks.</p><p>Sellanucheza longispina sp. nov. represents a second species of the genus to be recorded from a limestone cave environment in northeastern Laos, specifically marking the first record from Bolikhamsay Province. Compared to the pallid S. laotica sp. nov., this species retains dark pigmentation, suggesting that it may be a troglophile. Morphologically, the presence of sternal cones on both rings 5 and 6 represents a unique novelty for the genus. Geographically, the species appears to be narrowly endemic to the karst systems of Bolikhamsay Province it inhabits.</p><p>Etymology.</p><p>To emphasize the distinctively long and spiniform gonopodal process A; noun in apposition.</p></div>	https://treatment.plazi.org/id/4A258ECA2C055A2D9CE2B31EF528EBEF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Lips, Josiane;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Lips, Josiane, Panha, Somsak, Sutcharit, Chirasak (2026): Review of the millipede genus Sellanucheza Enghoff, Golovatch & Nguyen, 2004, with descriptions of three new species from Laos and redescription of S. variata (Attems, 1953) (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1282: 339-369, DOI: 10.3897/zookeys.1282.186746
3517C922DC8D55429A1867D30F606D3E.text	3517C922DC8D55429A1867D30F606D3E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sellanucheza tenebra (Hoffman 1961) Open	<div><p>Sellanucheza tenebra (Hoffman, 1961)</p><p>Fig. 1</p><p>Szechuanella tenebra Hoffman 1961: 535 (D).</p><p>Szechuanella tenebra – Hoffman 1963: 586 (M, K); Jeekel 1968: 75 (M); Wang and Mauriès 1996: 87 (L); Sierwald 2009: 125 (M).</p><p>Sellanucheza tenebra – Enghoff et al. 2004: 39 (M); Nguyen and Sierwald 2013: 1296 (L); Golovatch 2013 a: 22 (M, K); 2013 b: 330 (M, K).</p><p>Distribution.</p><p>China, Sichuan Province, Wushan, 31.07°N, 109.52°E (Hoffman 1961).</p><p>Remarks.</p><p>Sellanucheza tenebra was originally described as the type species of the genus Szechuanella Hoffman, 1961, based on a male holotype from Wushan, Sichuan Province, China (Hoffman 1961). However, as the name Szechuanella Hoffman, 1961 was found to be preoccupied by a trilobite genus (Zhang and Fan 1960), Enghoff et al. (2004) proposed Sellanucheza as a replacement name. Somatically, this species is unique among congeners due to the strongly reduced paraterga on rings 3–19, these being represented only by very faint bulges devoid of any delimiting sulci. This character makes S. tenebra clearly distinguished from all other known species, in which the paraterga, even if poorly developed, are always demarcated by at least a dorsal sulcus. In addition, male sternum 5 is characteristic in bearing a large, hemispherical lobe between coxae 4. Regarding the gonopod, S. tenebra is characterized by a long, strongly coiled solenophore (sph) which is subequal in length to a subunciform femorite (fe) (Fig. 1 A). Most notably, the gonopod bears a conspicuous, long and acute process pa directed ventrad, as well as a small, acute, basal process pb (Fig. 1 A, B). The solenophore tip is bifid, consisting of two large, rounded, subequal, hyaline, spatulate lobules (Fig. 1 A, B). These features make S. tenebra clearly distinguished from congeners such as S. variata or S. grandis, which typically show a relatively shorter or differently shaped solenophore. The species remains known only from the type locality in Sichuan Province, China.</p></div>	https://treatment.plazi.org/id/3517C922DC8D55429A1867D30F606D3E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Lips, Josiane;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Lips, Josiane, Panha, Somsak, Sutcharit, Chirasak (2026): Review of the millipede genus Sellanucheza Enghoff, Golovatch & Nguyen, 2004, with descriptions of three new species from Laos and redescription of S. variata (Attems, 1953) (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1282: 339-369, DOI: 10.3897/zookeys.1282.186746
92493876FB2059488AA0CC8ED9CF5F5B.text	92493876FB2059488AA0CC8ED9CF5F5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sellanucheza typica Golovatch 2013	<div><p>Sellanucheza typica Golovatch, 2013</p><p>Fig. 5 F, G</p><p>Sellanucheza typica Golovatch 2013 b: 329 (D, K).</p><p>Distribution.</p><p>China, Sichuan Province, Maoxian County, Southeast of Nanxinzhen, 3,155 m a. s. l., 31°34'04"N, 103°47'41"E (Golovatch 2013 b).</p><p>Remarks.</p><p>This species shows poorly developed paraterga which are nonetheless traceable as bulges demarcated by sulci at least dorsally. It differs clearly from the structurally similar S. variata by the complete absence of pleurosternal carinae (vs present until ring 5 in S. variata). The gonopods are highly diagnostic, characterized by an elongate, coiled and slender femorite (fe) (Fig. 5 F). The solenophore (sph) is subequal in length to the femorite, clearly coiled, and deeply bifurcated into an outer spiniform and an inner subspatuliform branch (Fig. 5 F). The base of the solenophore bears a long, slender, acute process pa curved anteroventrad (Fig. 5 F, G), while the distal part of the femorite is supplied with a short and spiniform process pb curved dorsad (Fig. 5 G). These gonopodal features, particularly the slenderer femorite, make S. typica readily distinguished from S. variata, the latter possessing a significantly stouter femorite. The species is currently known only from the type locality (highest elevation of 3,155 m a. s. l. in this genus) in Sichuan Province, thus being endemic to northern China.</p></div>	https://treatment.plazi.org/id/92493876FB2059488AA0CC8ED9CF5F5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Lips, Josiane;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Lips, Josiane, Panha, Somsak, Sutcharit, Chirasak (2026): Review of the millipede genus Sellanucheza Enghoff, Golovatch & Nguyen, 2004, with descriptions of three new species from Laos and redescription of S. variata (Attems, 1953) (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1282: 339-369, DOI: 10.3897/zookeys.1282.186746
0BF4A978E10055889F4A1645F4D92422.text	0BF4A978E10055889F4A1645F4D92422.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sellanucheza variata (Attems 1953) Open	<div><p>Sellanucheza variata (Attems, 1953)</p><p>Figs 2, 3</p><p>Nedyopus variatus Attems 1953: 170 (D).</p><p>Szechuanella variata – Hoffman 1963: 586 (D, K); Jeekel 1968: 62 (M); Golovatch 1983: 182 (M).</p><p>Sellanucheza variata – Enghoff et al. 2004: 39 (L); Golovatch 2013 a: 22 (M); 2013 b: 330 (M, K); Nguyen and Sierwald 2013: 1296 (L).</p><p>Type material examined.</p><p>• Syntypes 1 ♂, 1 ♀ (NHMW -3536), Vietnam, Laocai Province, Cha Pa (= Sapa), 1938–1939, leg. C. Dawydoff .</p><p>Diagnosis.</p><p>Among all known congeners, Sellanucheza variata appears to be most similar to S. typica Golovatch, 2013, but can be distinguished by the following combination of characters. Somatically, it differs from S. typica by the presence of pleurosternal carinae until ring 5 (vs absent in S. typica) and male tarsal brushes extending until ring 16 (vs ring 12 in S. typica). Regarding the gonopods, the femorite is notably stout (vs slender and elongate in S. typica). The solenophore is clearly curved and subequal in length to the femorite. Process pa is prominent, long, slender, and acute, while process pb is short, spiniform, and curved upwards.</p><p>Redescription.</p><p>Length ca 37 (♂) or 46 mm (♀), width of midbody pro- and metazona 2.8 and 3.3 mm (♂) or 4.6 and 5 mm (♀), respectively.</p><p>Coloration of alcohol material after 87 years of preservation, faded to reddish brown or black-brown (Fig. 2 A – F), posterior halves of metaterga, collum, paraterga and epiproct pale light brown to yellowish (Fig. 2 A – F), head and antennae yellow-brown, tip of antenna dark brown (Fig. 2 A), venter and legs pale brown to yellowish (Fig. 2 B – I).</p><p>Clypeolabral region and vertex sparsely setose, epicranial suture distinct. Antennae moderately long (Fig. 2 A, B), reaching body ring 3 (♂, ♀) when stretched dorsally. In width, head &lt;ring 3 = 4 &lt;2 &lt;collum &lt;ring 5 &lt;6–17 (♂); thereafter body gently and gradually tapering.</p><p>Collum semi-lunar, with an untraceable pattern of setation, surface rugulose, middle slightly flattened; paraterga rounded subtriangular, declined ventrad, devoid of lateral incisions; posterior corner very narrowly rounded, not extending past tergal margin (Fig. 2 A, B).</p><p>Tegument smooth and shining, prozona finely shagreened, metaterga finely leathery and faintly rugulose; surface below paraterga finely microgranulate (Fig. 2 A – F). Postcollum metaterga with two transverse rows of setae traceable at least as insertion points when setae broken off: 3 + 3 in anterior (pre-sulcus) and 4 + 4 in posterior (post-sulcus) row, the latter being barely traceable as insertion points (Fig. 2 A – F). Axial line visible both on pro- and metazona.</p><p>Paraterga well developed (Fig. 2 A, C, F), lying low (at 1 / 2 of body), slightly upturned, but lying below dorsum; anterior edge broadly rounded and narrowly bordered, fused to callus; lateral edge without incisions; posterior corner very narrowly rounded, not produced past rear tergal margin except for ring 2; posterior edge nearly straight. Paraterga 2 broad, anterior edge angular and rounded, lateral edge without incisions (Fig. 2 A, B). Calluses on paraterga narrow, demarcated by a sulcus only dorsally. Ozopore evident, lateral, lying in an ovoid groove at ca 1 / 3 metaterga in front of posterior corner (Fig. 2 B, D, E).</p><p>Transverse sulcus distinct (Fig. 2 A, C, F), slightly incomplete on metaterga 4, complete on metaterga 5–18, narrow, line-shaped, shallow, reaching the bases of paraterga, faintly beaded at bottom. Stricture between pro- and metazona narrow, beaded at bottom down to base of paraterga (Fig. 2 B – F).</p><p>Pleurosternal carinae complete high crests with a sharp posterior tooth on each of rings 2 and 3, thereafter remaining visible only as a bulge anteriorly until ring 5 (Fig. 2 B, D, E).</p><p>Epiproct (Fig. 2 E, F) conical, rounded dorsoventrally, with two small apical papillae; tip subtrapeziform; pre-apical papillae evident, lying close to tip. Hypoproct nearly semi-circular (Fig. 2 G), posterior tip broadly rounded, setiferous knobs at posterior edge very small and moderately well separated.</p><p>Sterna sparsely setose, without modifications (Fig. 2 G); an entire, large, rounded, linguiform, setose, sternal lobe between ♂ coxae 4 (Fig. 2 H, I). A paramedian pair of small, but evident tubercles in front of gonopod aperture. Legs moderately long and slender, midbody legs ca 1.2–1.4 (♂) or 1.1–1.3 × (♀) as long as body height, ♂ prefemora without modifications, tarsal brushes present until ♂ legs 16.</p><p>Gonopods (Fig. 3) simple. Coxite long, slender, cylindrical, slightly curved posteriorly, densely setose distodorsally. Prefemorite (pfe) densely setose, as usual, ovoid, elongate, ca 1 / 3–1 / 4 length of acropodite (Fig. 3 A – C). Femorite (fe) relatively stout, expanded basally, narrowed distad, evidently curved (Fig. 3 A – C), showing a distinct mesal groove; process pb short, spiniform, slightly curved upwards, located apicoventrally (Fig. 3 A, C). Solenophore (sph) large, broad, lamellar, strongly coiled, briefly bifid at tip, consisting of a straight, acute tooth and a rounded lobe with a faintly serrate margin (Fig. 3 C); base of solenophore with a most prominent, long, slender, curved and acute process pa projecting mesad (Fig. 3 A – C). Solenomere (sl) long, flagelliform, originating from femorite, almost fully sheathed by solenophore, with only its tip slightly exposed (Fig. 3 B, C).</p><p>Remarks.</p><p>Although Attems (1953) and Hoffman (1963) provided descriptions of this species, Sellanucheza variata has hitherto remained poorly defined, mainly due to the lack of detailed gonopod illustrations. The present study provides new and detailed drawings of the right gonopod (Fig. 3 A, B) based on the re-examination of the type material, complemented by a modified illustration of the left gonopod (Fig. 3 C) from Hoffman (1963) to provide a complete diagnostic overview. The present redescription, based on the syntypes, serves to complete the documentation and stabilize the diagnostic features of the species. In accordance with Hoffman (1963), the type locality is confirmed as being restricted to Chapa (= Sa Pa), Lao Cai Province, northern Vietnam.</p></div>	https://treatment.plazi.org/id/0BF4A978E10055889F4A1645F4D92422	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Likhitrakarn, Natdanai;Golovatch, Sergei I.;Lips, Josiane;Panha, Somsak;Sutcharit, Chirasak	Likhitrakarn, Natdanai, Golovatch, Sergei I., Lips, Josiane, Panha, Somsak, Sutcharit, Chirasak (2026): Review of the millipede genus Sellanucheza Enghoff, Golovatch & Nguyen, 2004, with descriptions of three new species from Laos and redescription of S. variata (Attems, 1953) (Diplopoda, Polydesmida, Paradoxosomatidae). ZooKeys 1282: 339-369, DOI: 10.3897/zookeys.1282.186746
