identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
7E9E563864345A1587851F6D0D763C51.text	7E9E563864345A1587851F6D0D763C51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax cristatus (Mollendorff 1886)	<div><p>Dicharax cristatus (Möllendorff, 1886)</p><p>Figs 5 A, 6</p><p>Alycaeus cristatus Möllendorff, 1886: 168, pl. 5, fig. 6. Type locality: Hunan, China.</p><p>Dicharax cristatus – Páll-Gergely et al. 2017: 34–43, figs 12 c, d, 13 c, 24–27, 28 a – d, 29 c, d, 30. Páll-Gergely et al. 2020 d: 57.</p><p>Material examined.</p><p>Laos • 139 shells; Vientiane Province, Feuang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.038445&amp;materialsCitation.latitude=18.674778" title="Search Plazi for locations around (long 102.038445/lat 18.674778)">unnamed limestone hill (loc. MF 1)</a>; 18°40'29.2"N, 102°02'18.4"E; 320 m a. s. l.; 7 Oct. 2024; leg. CU - NUOL expedition team; CUMZ 14480 (Figs 5 A, 6) • 34 shells; Vientiane Province, Feuang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.11748&amp;materialsCitation.latitude=18.739" title="Search Plazi for locations around (long 102.11748/lat 18.739)">Pha Reung (loc. MF 14)</a>; 18°44'20.4"N, 102°07'02.9"E; 360 m a. s. l.; 10 Oct. 2024; leg. CU - NUOL expedition team; CUMZ 14481 .</p><p>Diagnosis.</p><p>Shell depressed-conical. Spire ~ 1 / 5 of shell height. R 1 with regular ribs; R 2 as long as or slightly shorter than R 3. R 3 with sharp or moderately sharp swelling, in the middle of R 3 or close to the aperture. Inner peristome sometimes with conspicuous upper incision. Outer peristome expanded, not reflected.</p><p>Differential diagnosis.</p><p>See Páll-Gergely et al. (2017: 43).</p><p>Distribution.</p><p>Gansu, Shaanxi, Sichuan, Yunnan, Hunan, Hubei, Guangxi provinces, China; northern Vietnam; Luang Prabang Province (Páll-Gergely et al. 2017), and Feuang District, Vientiane Province, Laos.</p><p>Remarks.</p><p>The current species boundary of D. cristatus based on morphological characters is wide, varying in shell size, spire height (flat to somewhat elevated), peristome (strongly crenulated to not crenulated), R 1 sculpture (smooth to regularly ribbed), and the absence or presence or the position of the swelling on R 3. Therefore, there are no unique set of characters unequivocally defining D. cristatus (Páll-Gergely et al. 2017) . It is possible that future molecular studies may show that this nominal taxon contains multiple lineages. See earlier taxonomic treatment of D. cristatus in Páll-Gergely et al. (2017, 2020 d).</p></div>	https://treatment.plazi.org/id/7E9E563864345A1587851F6D0D763C51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jirapatrasilp, Parin;Inkhavilay, Khamla;Somvongsa, Chanvilay;Malavong, Banchai;Janjai, Teerangkul;Jeratthitikul, Ekgachai;Likhitrakarn, Natdanai;Srisonchai, Ruttapon;Sutcharit, Chirasak	Jirapatrasilp, Parin, Inkhavilay, Khamla, Somvongsa, Chanvilay, Malavong, Banchai, Janjai, Teerangkul, Jeratthitikul, Ekgachai, Likhitrakarn, Natdanai, Srisonchai, Ruttapon, Sutcharit, Chirasak (2026): The operculate micro land snails from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Laos, with description of new species. ZooKeys 1290: 257-291, DOI: 10.3897/zookeys.1290.198096
7843E635DF5B551892790FADDA8BB80B.text	7843E635DF5B551892790FADDA8BB80B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax feuangensis Jirapatrasilp & Inkhavilay 2026	<div><p>Dicharax feuangensis Jirapatrasilp &amp; Inkhavilay sp. nov.</p><p>Fig. 8</p><p>Type material.</p><p>Holotype. Laos • 1 shell. SH = 1.0 mm, SW = 1.9 mm; Vientiane Province, Feuang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.038445&amp;materialsCitation.latitude=18.674778" title="Search Plazi for locations around (long 102.038445/lat 18.674778)">unnamed limestone hill (loc. MF 1)</a>; 18°40'29.2"N, 102°02'18.4"E; 320 m a. s. l.; 7 Oct. 2024; leg. CU - NUOL expedition team; CUMZ 14483 (Fig. 8 A) . Paratypes. Laos • 1 shell; same data as for holotype; CUMZ 14484 (Fig. 8 B) .</p><p>Diagnosis.</p><p>Shell lenticular, depressed-conical. Spire ~ 1 / 5 of shell height. R 1 ~ 1 ½ whorls with regular, sharp radial ribs. R 2 very short; with ~ 10 ribs; ribs consistently thickened throughout. R 3 ~ 1 / 5 whorl with ~ 21 ribs. Apertural tube coiling parallel to horizontal axis of the shell, slightly expanding near the end of the aperture. Inner peristome slim, very slightly protruding. Outer peristome thickened, slightly expanded at the columellar corner, not reflected.</p><p>Differential diagnosis.</p><p>Among the tiny Dicharax species (SH and SW &lt;2 mm), D. feuangensis sp. nov. is most similar to D. parvulus (Möllendorff, 1887) (Páll-Gergely et al. 2021: fig. 31 b) in its low spire, short R 2, and a nearly consistent radial rib density along the teleoconch, rendering the boundaries between R 1 and R 2, and R 2 and R 3 indistinct. However, the new species has a longer R 3 with more ribs (21 vs 14), a less widening R 3, and an outer peristome with only a slight expansion at the columellar corner. This new species is also similar to D. abdoui Páll-Gergely, 2017 (Páll-Gergely et al. 2017: fig. 6) in R 2 length, but D. feuangensis sp. nov. has distinct higher elevated radial ribs throughout R 1 – R 3, and a smooth protoconch, while D. abdoui has nearly smooth R 1 and R 3, and a protoconch with extremely fine pits arranged in spiral rows.</p><p>Description.</p><p>Shell lenticular, depressed-conical, solid, translucent, SH 1.0 mm, SW 1.9 mm (n = 2). Shell outline oval in apical view, spire ~ 1 / 5 of shell height. Whorls ~ 3 ½; spiral striation lacking on the entire shell. Protoconch low, ~ 1 ¾ whorls, smooth. Radial ribs strong, regular, starting from R 1, becoming sharper, thicker, and less close-set toward the end of aperture. R 1 ~ 1 ½ whorls with ~ 33 ribs in ¼ whorl adjoining R 2. Boundary between R 1 and R 2 indistinct as R 2 contains nearly the same rib density and thickness as in R 1. R 2 very short; with ~ 10 ribs; ribs consistently thickened throughout. Boundary between R 2 and R 3 indistinct as R 3 contains nearly the same rib density and thickness as in R 2. R 3 ~ 1 / 5 whorl with ~ 21 ribs; apertural tube coiling parallel to horizontal axis of the shell, slightly expanding near the end of the aperture. Aperture round. Peristome double with regular outer peristome. Inner peristome slim, slightly protruding. Outer peristome thickened, multilayered (visible in lateral view), slightly expanded at the columellar corner, not reflected. Umbilicus elliptical, open, approximately 1 / 3 of shell width. Operculum thin, translucent; multispiral ridges not elevated.</p><p>Etymology.</p><p>The species name is derived from the type locality, Feuang District in Vientiane Province, Laos.</p><p>Distribution.</p><p>Known only from the type locality in Feuang District, Vientiane Province, Laos.</p><p>Remarks.</p><p>The length and the less widening of R 3, and a smooth protoconch are consistent across all specimens of this new species, suggesting that these are important diagnostic characters.</p></div>	https://treatment.plazi.org/id/7843E635DF5B551892790FADDA8BB80B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jirapatrasilp, Parin;Inkhavilay, Khamla;Somvongsa, Chanvilay;Malavong, Banchai;Janjai, Teerangkul;Jeratthitikul, Ekgachai;Likhitrakarn, Natdanai;Srisonchai, Ruttapon;Sutcharit, Chirasak	Jirapatrasilp, Parin, Inkhavilay, Khamla, Somvongsa, Chanvilay, Malavong, Banchai, Janjai, Teerangkul, Jeratthitikul, Ekgachai, Likhitrakarn, Natdanai, Srisonchai, Ruttapon, Sutcharit, Chirasak (2026): The operculate micro land snails from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Laos, with description of new species. ZooKeys 1290: 257-291, DOI: 10.3897/zookeys.1290.198096
2AD2885A2C1E50E69FDD189701DAB90B.text	2AD2885A2C1E50E69FDD189701DAB90B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax Kobelt & Mollendorff 1900	<div><p>Dicharax Kobelt &amp; Möllendorff, 1900</p><p>Alycaeus (Dicharax) Kobelt &amp; Möllendorff, 1900: 186 (replacement name for Charax Benson, 1859, non Charax Scopoli, 1777 [Pisces]).</p><p>Dicharax – Jirapatrasilp et al. 2021: 4. Páll-Gergely et al. 2021: 21. Gittenberger et al. 2024: 199. Matsuda and Yano 2024: 3. Páll-Gergely et al. 2025: 50.</p><p>Type species.</p><p>Alycaeus hebes Benson, 1857, by subsequent designation (Gude 1921: 236).</p><p>Remarks.</p><p>This genus is diagnosed by a low protoconch, and the entire shell lacking spiral striation. This genus comprises nearly 200 species and is widely distributed from the Western Ghats of India and in the southwestern and southeastern Himalayan region to Japan, through the Malay Peninsula to the southern arc of the Malay Archipelago up to Sumatra and Java (Páll-Gergely et al. 2020 d). See earlier taxonomic treatment, extended diagnosis, and other taxonomic remarks of Dicharax in Jirapatrasilp et al. (2021).</p></div>	https://treatment.plazi.org/id/2AD2885A2C1E50E69FDD189701DAB90B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jirapatrasilp, Parin;Inkhavilay, Khamla;Somvongsa, Chanvilay;Malavong, Banchai;Janjai, Teerangkul;Jeratthitikul, Ekgachai;Likhitrakarn, Natdanai;Srisonchai, Ruttapon;Sutcharit, Chirasak	Jirapatrasilp, Parin, Inkhavilay, Khamla, Somvongsa, Chanvilay, Malavong, Banchai, Janjai, Teerangkul, Jeratthitikul, Ekgachai, Likhitrakarn, Natdanai, Srisonchai, Ruttapon, Sutcharit, Chirasak (2026): The operculate micro land snails from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Laos, with description of new species. ZooKeys 1290: 257-291, DOI: 10.3897/zookeys.1290.198096
9FDE9B761CE95B2B9ACCD60A5FBFCA72.text	9FDE9B761CE95B2B9ACCD60A5FBFCA72.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicharax pallgergelyi Jirapatrasilp & Inkhavilay 2026	<div><p>Dicharax pallgergelyi Jirapatrasilp &amp; Inkhavilay sp. nov.</p><p>Figs 5 B, 7</p><p>Type material.</p><p>Holotype. Laos • 1 shell, SH = 1.5 mm, SW = 2.3 mm; Vientiane Province, Feuang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.038445&amp;materialsCitation.latitude=18.674778" title="Search Plazi for locations around (long 102.038445/lat 18.674778)">unnamed limestone hill (loc. MF 1)</a>; 18°40'29.2"N, 102°02'18.4"E; 320 m a. s. l.; 7 Oct. 2024; leg. CU - NUOL expedition team; CUMZ 14482 (Fig. 5 B) . Paratypes. Laos • 5 shells; same data as for holotype; CUMZ 14482 A (Fig. 7) .</p><p>Diagnosis.</p><p>Shell depressed-conical. Spire ~ 1 / 5 of shell height. R 1 ~ 1 ¾ whorls with regular, very low ribs. R 2 ~ 1 / 5 whorl and nearly as long as or slightly shorter than R 3; with ~ 25 ribs; ribs becoming thicker at the end of R 2. R 3 ~ 1 / 5 whorl with very fine, close-set ribs. Apertural tube coiling slightly downward from the horizontal axis of the shell, expanding near the end of the aperture. Inner peristome blunt, thick, protruding. Outer peristome thickened, multilayered, expanded, not reflected.</p><p>Differential diagnosis.</p><p>Dicharax pallgergelyi sp. nov. is similar to D. notus (Godwin-Austen, 1914) (Páll-Gergely et al. 2025: fig. 71 a – e) in shell shape and size, but the new species possesses a lesser number of R 2 ribs (25–28) with lower rib height, compared to those in D. notus (48–50, more elevated R 2 ribs with T-shaped cross section). In addition, D. pallgergelyi sp. nov. also differs by a distinct protruding inner peristome which is blunt and thick.</p><p>Description.</p><p>Shell depressed-conical, solid, translucent, SH 1.4–1.5 mm, SW 2.1–2.4 mm (n = 6). Shell outline oval in apical view, spire ~ 1 / 5 of shell height. Whorls ~ 3 ¾; spiral striation lacking on the entire shell. Protoconch low, ~ 1 ¾ whorls, smooth. Radial ribs regular and very low, indistinct starting from R 1, becoming more distinct toward the end of R 2, and finer, more close-set, and less distinct in R 3. R 1 ~ 1 ¾ whorls with ~ 38 ribs in ¼ whorl adjoining R 2. Boundary between R 1 and R 2 distinct as R 2 contains thicker and more distinct ribs. R 2 ~ 1 / 5 whorl and nearly as long as or slightly shorter than R 3; with ~ 25 ribs; ribs becoming thicker at the end of R 2. Boundary between R 2 and R 3 distinct as a slight constriction. R 3 ~ 1 / 5 whorl with very fine, close-set ribs; apertural tube coiling slightly downward from the horizontal axis of the shell, expanding near the end of the aperture. Aperture round. Peristome double with prominent outer peristome. Inner peristome blunt, thick, protruding. Outer peristome thickened, multilayered (visible in lateral view), expanded, not reflected. Umbilicus elliptical, open, approximately one third of shell width. Operculum unknown.</p><p>Etymology.</p><p>The species name is in honour of Dr. Barna Páll-Gergely, a prominent Hungarian malacologist, who extensively studied the taxonomy of the Alycaeinae and first recognised this species as new to science.</p><p>Distribution.</p><p>Known only from the type locality in Feuang District, Vientiane Province, Laos.</p><p>Remarks.</p><p>The number range of R 2 ribs and the occurrence of the protruding inner peristome are consistent across all specimens of this new species, suggesting that these are important diagnostic characters.</p></div>	https://treatment.plazi.org/id/9FDE9B761CE95B2B9ACCD60A5FBFCA72	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jirapatrasilp, Parin;Inkhavilay, Khamla;Somvongsa, Chanvilay;Malavong, Banchai;Janjai, Teerangkul;Jeratthitikul, Ekgachai;Likhitrakarn, Natdanai;Srisonchai, Ruttapon;Sutcharit, Chirasak	Jirapatrasilp, Parin, Inkhavilay, Khamla, Somvongsa, Chanvilay, Malavong, Banchai, Janjai, Teerangkul, Jeratthitikul, Ekgachai, Likhitrakarn, Natdanai, Srisonchai, Ruttapon, Sutcharit, Chirasak (2026): The operculate micro land snails from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Laos, with description of new species. ZooKeys 1290: 257-291, DOI: 10.3897/zookeys.1290.198096
A9F0F327D8D85449BB8304EF204C367D.text	A9F0F327D8D85449BB8304EF204C367D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dioryx Benson 1859	<div><p>Dioryx Benson, 1859</p><p>Alycæus (Dioryx) Benson, 1859: 177 .</p><p>Dioryx – Páll-Gergely et al. 2020 d: 121–122.</p><p>Type species.</p><p>Alycaeus amphora Benson, 1856, by subsequent designation (Gude 1921: 198).</p><p>Remarks.</p><p>This genus is diagnosed by a globular or high-spired shell, sometimes the body whorl is angled or keeled, a smooth protoconch, a reduced sculpture in teleoconch, and an absence of R 3. At present, there are 27 species of Dioryx known from the southeastern Himalayan region to Taiwan in the east, and down to the northern part of the Malay Peninsula and southern Vietnam in the south (Páll-Gergely et al. 2020 d). See earlier taxonomic treatment, extended diagnosis, and other taxonomic remarks of Dioryx in Páll-Gergely et al. (2020 d).</p></div>	https://treatment.plazi.org/id/A9F0F327D8D85449BB8304EF204C367D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jirapatrasilp, Parin;Inkhavilay, Khamla;Somvongsa, Chanvilay;Malavong, Banchai;Janjai, Teerangkul;Jeratthitikul, Ekgachai;Likhitrakarn, Natdanai;Srisonchai, Ruttapon;Sutcharit, Chirasak	Jirapatrasilp, Parin, Inkhavilay, Khamla, Somvongsa, Chanvilay, Malavong, Banchai, Janjai, Teerangkul, Jeratthitikul, Ekgachai, Likhitrakarn, Natdanai, Srisonchai, Ruttapon, Sutcharit, Chirasak (2026): The operculate micro land snails from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Laos, with description of new species. ZooKeys 1290: 257-291, DOI: 10.3897/zookeys.1290.198096
92E5B2325A76571C864360157360FD8F.text	92E5B2325A76571C864360157360FD8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dioryx dongiensis Varga 1972	<div><p>Dioryx dongiensis Varga, 1972</p><p>Fig. 2 B</p><p>Dioryx dongiensis Varga, 1972: 135, 137, figs 21–23. Type locality: Vietnam, Tonkin, Cuc Phuong bei Dong [Cuc Phuong, Ninh Binh]. Páll-Gergely et al. 2020 d: 125. von Oheimb et al. 2025: tables 1, 3, 4, fig. 8 h.</p><p>Material examined.</p><p>Laos • 1 shell; Vientiane Province, Feuang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.97522&amp;materialsCitation.latitude=18.550138" title="Search Plazi for locations around (long 101.97522/lat 18.550138)">limestone hill behind Xinxayyaram Temple (loc. MF 10)</a>; 18°33'00.5"N, 101°58'30.8"E; 265 m a. s. l.; 9 Oct. 2024; leg. CU - NUOL expedition team; CUMZ 14476 (Fig. 2 B) .</p><p>Diagnosis.</p><p>Shell conical, globular. Spire ~ 30 % of shell height. Protoconch smooth. R 1 ~ 3 ½ whorls, sculptured with indistinct spiral striation. R 2 ~ ¼ whorl, with thin ribs; the end of R 2 slightly narrower. Peristome double with expanding outer peristome; totally adnate to the last whorl. Umbilicus slightly open.</p><p>Differential diagnosis.</p><p>Dioryx dongiensis is different from D. kobeltianus (Möllendorff, 1874) in having a shallower umbilicus with a less pronounced edge of the last whorl’s lower periphery. In addition, the peristome of D. kobeltianus is not totally adnate to the last whorl (Páll-Gergely et al. 2017).</p><p>Distribution.</p><p>The type locality (von Oheimb et al. 2025), and Feuang District, Vientiane Province, Laos.</p><p>Remarks.</p><p>This is the first record of Dioryx dongiensis in Laos, apart from the type locality in Vietnam (von Oheimb et al. 2025). Although the locality in Laos is more than 400 km away from the type locality, this shell is herein identified to this species based on its shell shape and the peristome totally adnate to the last whorl. Moreover, several Alycaeinae species have a wide distribution from Vietnam to Laos (e. g., Dicharax depressus; Páll-Gergely et al. 2017). As the shell of this species is very small, soil sieving and floatation methods (Liew et al. 2008) are preferred to retrieve more shells, and this species might be collected from more localities located between Vietnam and Laos. See earlier taxonomic treatment of Dioryx dongiensis in Páll-Gergely et al. (2020 d).</p></div>	https://treatment.plazi.org/id/92E5B2325A76571C864360157360FD8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jirapatrasilp, Parin;Inkhavilay, Khamla;Somvongsa, Chanvilay;Malavong, Banchai;Janjai, Teerangkul;Jeratthitikul, Ekgachai;Likhitrakarn, Natdanai;Srisonchai, Ruttapon;Sutcharit, Chirasak	Jirapatrasilp, Parin, Inkhavilay, Khamla, Somvongsa, Chanvilay, Malavong, Banchai, Janjai, Teerangkul, Jeratthitikul, Ekgachai, Likhitrakarn, Natdanai, Srisonchai, Ruttapon, Sutcharit, Chirasak (2026): The operculate micro land snails from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Laos, with description of new species. ZooKeys 1290: 257-291, DOI: 10.3897/zookeys.1290.198096
30AF2FFEC95D56EA86C3787C7ED13CA5.text	30AF2FFEC95D56EA86C3787C7ED13CA5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dioryx feuangensis Jirapatrasilp & Inkhavilay 2026	<div><p>Dioryx feuangensis Jirapatrasilp &amp; Inkhavilay sp. nov.</p><p>Fig. 4</p><p>Type material.</p><p>Holotype. Laos • 1 shell, SH = 9.3 mm, SW = 8.3 mm; Vientiane Province, Feuang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.131805&amp;materialsCitation.latitude=18.702389" title="Search Plazi for locations around (long 102.131805/lat 18.702389)">Pha Thor (loc. MF 13)</a>; 18°42'08.6"N, 102°07'54.5"E; 225 m a. s. l.; 10 Oct. 2024; leg. CU - NUOL expedition team; CUMZ 14477 (Fig. 4 A) . Paratypes. Laos • 33 shells; same data as for holotype; CUMZ 14478 • 3 shells; same data as for holotype; NHMUK 20260369 • 3 shells; same data as for holotype; SMF .</p><p>Other material.</p><p>Laos • 224 shells; Vientiane Province, Feuang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.11748&amp;materialsCitation.latitude=18.739" title="Search Plazi for locations around (long 102.11748/lat 18.739)">Pha Reung (loc. MF 14)</a>; 18°44'20.4"N, 102°07'02.9"E; 360 m a. s. l.; 10 Oct. 2024; leg. CU - NUOL expedition team; CUMZ 14479 (Fig. 4 B) .</p><p>Diagnosis.</p><p>Shell conical, globular. Spire ~ ¼ of shell height. Spiral striation lacking on the entire shell. R 1 ~ 2 ½ whorls; R 2 ~ ½ whorl, with indistinct ribs; the end of R 2 narrower, appearing as a broad constriction between R 2 and apertural tube. Peristome double with prominent, expanding outer peristome with irregular perimeter. Umbilicus open.</p><p>Differential diagnosis.</p><p>Dioryx feuangensis sp. nov. is distinct in having a prominent, expanding outer peristome with irregular perimeter. Apart from this character, this new species is similar to D. compactus (Bavay &amp; Dautzenberg, 1900) (Páll-Gergely et al. 2017: fig. 4 c) in its globular shell shape with a SW: SH ratio approaching 1, but the new species differs by a larger shell (SH 8.5–9.5 mm, SW 8.0– 8.5 mm vs SH 4.7 mm, SW 4.2 mm) with matt surface. This new species is also similar to D. rosea (Bavay &amp; Dautzenberg, 1900) in its shell colour and shape, but the new species differs by a lack of spiral striation on the entire shell, and a less protruding inner peristome. The new species’ matt shell surface matches more to D. globuloides Zilch, 1957 (Páll-Gergely et al. 2017: fig. 4 d), but the latter species is smaller, with a shorter sutural tube at R 2, and the outer peristome is expanded to a lesser extent. Dioryx feuangensis sp. nov. is similar to D. requiescens (Mabille, 1887) in its R 2 length, but D. feuangensis sp. nov. is more globular in shell shape, and has an outer peristome with irregular perimeter, and an apertural tube opening parallel to the horizontal axis of the shell. Dioryx requiescens is less globular (SW: SH ratio &lt;1), and has a regular outer peristome, and an apertural tube opening obliquely downward to the horizontal axis of the shell.</p><p>Description.</p><p>Shell conical, globular, solid, matt, yellowish to pale crimson, SH 8.5–9.5 mm, SW 8.0– 8.5 mm (n = 10). Shell outline round in apical view, spire ~ ¼ of shell height. Whorls ~ 4 ½, last whorl large, round; spiral striation lacking on the entire shell. Protoconch low, ~ 1 ½ whorls, glossy, smooth. R 1 ~ 2 ½ whorls; R 2 ~ ½ whorl, with indistinct ribs; the end of R 2 narrower, appearing as a broad constriction between R 2 and apertural tube. R 3 absent; apertural tube expanding from the end of R 2, opening parallel to horizontal axis of the shell. Aperture round; peristome double with prominent outer peristome with irregular perimeter. Inner peristome blunt, slightly protruding. Outer peristome not thickened, expanded, not reflected. Umbilicus open. Operculum unknown.</p><p>Etymology.</p><p>The species name is derived from the type locality, Feuang District in Vientiane Province, Laos.</p><p>Distribution.</p><p>Known only from the limestone areas in Feuang District, Vientiane Province, Laos.</p><p>Remarks.</p><p>The lack of spiral striation on the entire shell and a prominent, expanding outer peristome with irregular perimeter are consistent across all specimens of this new species, suggesting that these are important diagnostic characters.</p></div>	https://treatment.plazi.org/id/30AF2FFEC95D56EA86C3787C7ED13CA5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jirapatrasilp, Parin;Inkhavilay, Khamla;Somvongsa, Chanvilay;Malavong, Banchai;Janjai, Teerangkul;Jeratthitikul, Ekgachai;Likhitrakarn, Natdanai;Srisonchai, Ruttapon;Sutcharit, Chirasak	Jirapatrasilp, Parin, Inkhavilay, Khamla, Somvongsa, Chanvilay, Malavong, Banchai, Janjai, Teerangkul, Jeratthitikul, Ekgachai, Likhitrakarn, Natdanai, Srisonchai, Ruttapon, Sutcharit, Chirasak (2026): The operculate micro land snails from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Laos, with description of new species. ZooKeys 1290: 257-291, DOI: 10.3897/zookeys.1290.198096
521FD8D6E33D53B69A7571C45778F2EE.text	521FD8D6E33D53B69A7571C45778F2EE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metalycaeus heudei (Bavay & Dautzenberg 1900)	<div><p>Metalycaeus heudei (Bavay &amp; Dautzenberg, 1900)</p><p>Figs 2 A, 3</p><p>Alycaeus (Charax) heudei Bavay &amp; Dautzenberg, 1900: 121–122 . Type locality: Haut-Tonkin.</p><p>Metalycaeus heudei – Páll-Gergely et al. 2020 d: 143 –144. Páll-Gergely et al. 2021: 53, fig. 41.</p><p>Material examined.</p><p>Laos • 2 shells; Houaphanh Province, Viengxay District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.145386&amp;materialsCitation.latitude=20.457806" title="Search Plazi for locations around (long 104.145386/lat 20.457806)">Ban Nathan village</a>; 20°27'28.1"N, 104°08'43.4"E; 965 m a. s. l.; 6 Oct. 2022; leg. K. Inkhavilay; CUMZ 14473 –14474 (Figs 2 A, 3) • 17 shells; Houaphanh Province, Viengxay District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.17461&amp;materialsCitation.latitude=20.46075" title="Search Plazi for locations around (long 104.17461/lat 20.46075)">Ban Hang Long village</a>; 20°27'38.7"N, 104°10'28.6"E; 840 m a. s. l.; 6 Oct. 2022; leg. K. Inkhavilay; CUMZ 14475 .</p><p>Diagnosis.</p><p>Shell depressed to conical. Protoconch scaly; spiral striation starting from R 1, continuing to the end of R 2. R 1 ~ 1 ¾ whorls with regular, strong radial ribs. R 2 ~ ¼ whorl and nearly twice as long as R 3; with ~ 32 ribs. R 3 with less distinct ribs, strongly coiling downwards being highly oblique to horizontal axis of the shell, slightly expanding near the end of the aperture. Inner peristome blunt, slightly protruding. Outer peristome thickened, multilayered, expanded in various degrees, not reflected. Apertural lip finely granulated, mostly with 4–6 wavy ridges inside the apertural tube at the position when the tube starting to coil downward. Umbilicus round, close.</p><p>Differential diagnosis.</p><p>See Páll-Gergely et al. (2017: 83).</p><p>Distribution.</p><p>Yunnan, Hunan and Hubei provinces, China; Shan and Kayah States, Myanmar; northern Vietnam; Luang Prabang (Páll-Gergely et al. 2017, 2021) and Houaphanh provinces, Laos.</p><p>Remarks.</p><p>Metalycaeus heudei is one of the most widely distributed alycaein species known, with high variation in shell size, shell shape from depressed to conical, R 2 length and shape (bulging or not), and the strength of the swelling on R 3. The shells from Houaphanh Province, Laos in this study correspond to the conical end of the shell shape spectrum (paviei form). These shells are also very similar to the type material of Alycaeus paviei var. minor Bavay &amp; Dautzenberg, 1912 in having a close umbilicus and ridges inside the apertural tube at the position when the tube starting to coil downward. In addition, a lack of spiral striation on the protoconch was detected in these shells, aligning to the previous report that not all shells from the same Metalycaeus species possess this character (Páll-Gergely et al. 2017, 2021). See earlier taxonomic treatment of M. heudei in Páll-Gergely et al. (2021).</p></div>	https://treatment.plazi.org/id/521FD8D6E33D53B69A7571C45778F2EE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jirapatrasilp, Parin;Inkhavilay, Khamla;Somvongsa, Chanvilay;Malavong, Banchai;Janjai, Teerangkul;Jeratthitikul, Ekgachai;Likhitrakarn, Natdanai;Srisonchai, Ruttapon;Sutcharit, Chirasak	Jirapatrasilp, Parin, Inkhavilay, Khamla, Somvongsa, Chanvilay, Malavong, Banchai, Janjai, Teerangkul, Jeratthitikul, Ekgachai, Likhitrakarn, Natdanai, Srisonchai, Ruttapon, Sutcharit, Chirasak (2026): The operculate micro land snails from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Laos, with description of new species. ZooKeys 1290: 257-291, DOI: 10.3897/zookeys.1290.198096
8F03BD7842805C2A8AD4625CA6266A97.text	8F03BD7842805C2A8AD4625CA6266A97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Metalycaeus Pilsbry 1900	<div><p>Metalycaeus Pilsbry, 1900</p><p>Alycæus (Metalycæus) Pilsbry, 1900: 382 .</p><p>Metalycaeus – Páll-Gergely et al. 2020 d: 137 –138. Páll-Gergely et al. 2021: 45. Gittenberger et al. 2024: 190.</p><p>Type species.</p><p>Alycaeus (Metalycaeus) melanopoma Pilsbry, 1900 (synonym of Alycaeus nipponensis Reinhardt, 1877, see Minato 1988), by subsequent designation (Thiele 1929).</p><p>Remarks.</p><p>This genus is diagnosed by the presence of spiral striation on both the protoconch and teleoconch. However, there is at least one species, M. pygmachos Páll-Gergely &amp; Hunyadi, 2021 from Myanmar, lacking spiral striation on the entire shell, but this species is assigned to Metalycaeus based on its large size and the depressed swelling on R 3. Therefore, the lack of spiral striation on the protoconch is deemed a secondary reduction, and it is necessary to consider other shell characters in order to classify the specimens into Metalycaeus . At present, there are 70 species of Metalycaeus known from southeastern Himalaya to the south of Honshu Island, Japan, the northern islands of the Philippines, northern Laos and Vietnam, and central Thailand (Páll-Gergely et al. 2020 d, 2021; Gittenberger et al. 2024). See earlier taxonomic treatment, extended diagnosis, and other taxonomic remarks of Metalycaeus in Páll-Gergely et al. (2020 d) and Gittenberger et al. (2024).</p></div>	https://treatment.plazi.org/id/8F03BD7842805C2A8AD4625CA6266A97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jirapatrasilp, Parin;Inkhavilay, Khamla;Somvongsa, Chanvilay;Malavong, Banchai;Janjai, Teerangkul;Jeratthitikul, Ekgachai;Likhitrakarn, Natdanai;Srisonchai, Ruttapon;Sutcharit, Chirasak	Jirapatrasilp, Parin, Inkhavilay, Khamla, Somvongsa, Chanvilay, Malavong, Banchai, Janjai, Teerangkul, Jeratthitikul, Ekgachai, Likhitrakarn, Natdanai, Srisonchai, Ruttapon, Sutcharit, Chirasak (2026): The operculate micro land snails from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Laos, with description of new species. ZooKeys 1290: 257-291, DOI: 10.3897/zookeys.1290.198096
C104AF4B16EB53F69294EA48BECAD50B.text	C104AF4B16EB53F69294EA48BECAD50B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pincerna panhai Jirapatrasilp & Inkhavilay 2026	<div><p>Pincerna panhai Jirapatrasilp &amp; Inkhavilay sp. nov.</p><p>Figs 10, 11</p><p>Type material.</p><p>Holotype. Laos • 1 shell, SH = 3.9 mm, SW = 4.2 mm; Vientiane Province, Feuang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.038445&amp;materialsCitation.latitude=18.674778" title="Search Plazi for locations around (long 102.038445/lat 18.674778)">unnamed limestone hill (loc. MF 1)</a>; 18°40'29.2"N, 102°02'18.4"E; 320 m a. s. l.; 7 Oct. 2024; leg. CU - NUOL expedition team; CUMZ 14486 (Fig. 10) . Paratypes. Laos • 5 shells; same data as for holotype; CUMZ 14487 (Fig. 11 A) • 2 shells; same data as for holotype; NHMUK 20260370 • 2 shells; same data as for holotype; SMF .</p><p>Other material.</p><p>Laos • 5 shells; Vientiane Province, Feuang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.97972&amp;materialsCitation.latitude=18.544945" title="Search Plazi for locations around (long 101.97972/lat 18.544945)">limestone hill opposite to Xinxayyaram Temple (loc. MF 8)</a>; 18°32'41.8"N, 101°58'47.0"E; 280 m a. s. l.; 8 Oct. 2024; leg. CU - NUOL expedition team; CUMZ 14488 (Fig. 11 B) • 5 shells; Vientiane Province, Feuang District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.11748&amp;materialsCitation.latitude=18.739" title="Search Plazi for locations around (long 102.11748/lat 18.739)">Pha Reung (loc. MF 14)</a>; 18°44'20.4"N, 102°07'02.9"E; 360 m a. s. l.; 10 Oct. 2024; leg. CU - NUOL expedition team; CUMZ 14489 .</p><p>Diagnosis.</p><p>R 1 ~ 1 ¾ whorls with regular, sharp radial ribs. R 2 ~ ¼ whorl and nearly as long as R 3; with ~ 25 thicker, blunter, and more close-set ribs. R 3 ~ 3 ribs; apertural tube coiling more or less parallel to horizontal axis of the shell, expanding near the end of the aperture. Inner peristome blunt, slightly protruding. Outer peristome thickened, multilayered, expanded, not reflected. Umbilicus closed by the expanded columellar extension of outer peristome.</p><p>Differential diagnosis.</p><p>Among the ribbed species of Pincerna, P. panhai sp. nov. is similar to P. costulosus (Bavay &amp; Dautzenberg, 1912) (Páll-Gergely 2023 b: figs 8, 9) in its sharp, spaced radial ribs, and an umbilicus closed by the expanded columellar extension of outer peristome. However, the new species has a wider shell shape, a longer R 2 (~ 25 vs 6–14 ribs), and R 3 with sharper, more distinct ribs. The sharp, distinct R 3 ribs of the new species match more with P. acroptychia Páll-Gergely, 2023 (Páll-Gergely 2023 b: fig. 5 a – e), but the new species has a wider shell shape, less close-set radial ribs, and a longer R 2 (~ 25 vs 16 ribs). Pincerna panhai sp. nov. is also similar to P. viginticostatus Páll-Gergely, 2023 (Páll-Gergely 2023 b: fig. 14 a – f) in shell shape and R 2 length, and a close umbilicus. However, the new species has sharper and less close-set radial ribs, and less R 3 ribs (~ 3 vs 8 ribs).</p><p>Description.</p><p>Shell conical, globose, solid, translucent, whitish to pale crimson, SH 3.8–4.0 mm, SW 4.1–4.3 mm (n = 10). Shell outline round in apical view, spire ~ ¼ of shell height. Whorls ~ 4 ¼, last whorl large, round; spiral striation lacking on the entire shell. Protoconch high, ~ 1 ¾ whorls, glossy, smooth. Radial ribs strong, regular, starting from R 1, continuing to the end of aperture. R 1 ~ 1 ¾ whorls; with ~ 9 sharp ribs in ¼ whorl adjoining R 2. Boundary between R 1 and R 2 distinct as R 2 contains more close-set and thicker, blunter ribs than R 1. R 2 ~ ¼ whorl and nearly as long as R 3; with ~ 25 ribs; ribs consistently thickened throughout. Boundary between R 2 and R 3 distinct as R 3 abruptly expanded, and contains much less close-set and sharper ribs than R 1 and R 2. R 3 with fine growth lines and ~ 3 ribs; coiling more or less parallel to horizontal axis of the shell, expanding near the end of the aperture. Aperture round; peristome double with prominent outer peristome. Inner peristome blunt, slightly protruding. Outer peristome thickened, multilayered (visible in lateral view), expanded, not reflected. Umbilicus closed by the expanded columellar extension of outer peristome. Operculum thin, translucent; multispiral ridges not elevated.</p><p>Etymology.</p><p>The species name is in honour of Prof. Somsak Panha, a prominent Thai malacologist and our beloved professor, who supervised the taxonomic and systematic study of land snails and other terrestrial invertebrates in Laos, as well as other countries in mainland Southeast Asia.</p><p>Distribution.</p><p>Known only from the limestone areas in Feuang District, Vientiane Province, Laos.</p><p>Remarks.</p><p>The occurrence of regular, sharp radial ribs, the number range of R 2 and R 3 ribs, and a close umbilicus are consistent across all specimens of this new species, suggesting that these are important diagnostic characters.</p></div>	https://treatment.plazi.org/id/C104AF4B16EB53F69294EA48BECAD50B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jirapatrasilp, Parin;Inkhavilay, Khamla;Somvongsa, Chanvilay;Malavong, Banchai;Janjai, Teerangkul;Jeratthitikul, Ekgachai;Likhitrakarn, Natdanai;Srisonchai, Ruttapon;Sutcharit, Chirasak	Jirapatrasilp, Parin, Inkhavilay, Khamla, Somvongsa, Chanvilay, Malavong, Banchai, Janjai, Teerangkul, Jeratthitikul, Ekgachai, Likhitrakarn, Natdanai, Srisonchai, Ruttapon, Sutcharit, Chirasak (2026): The operculate micro land snails from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Laos, with description of new species. ZooKeys 1290: 257-291, DOI: 10.3897/zookeys.1290.198096
4E19A43A89F0597881FF4A9DE04FD1E0.text	4E19A43A89F0597881FF4A9DE04FD1E0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pincerna Preston 1907	<div><p>Pincerna Preston, 1907</p><p>Alycæus (Pincerna) Preston, 1907: 206 .</p><p>Pincerna – Do and Nguyen 2022: 365. Páll-Gergely 2023 b: 259.</p><p>Type species.</p><p>Alycæus (Pincerna) liratula Preston, 1907, by original designation.</p><p>Remarks.</p><p>This genus is diagnosed by a high spired, globose triangular shell, a smooth protoconch, a very short to short R 2, and a closed umbilicus. At present, there is a total of 13 species of Pincerna known from northern Vietnam and northern Laos, and Sumatra, the southern part of the Malay Peninsula, and northern Borneo (Páll-Gergely et al. 2020 d; Gittenberger et al. 2022). See earlier taxonomic treatment, extended diagnosis and other taxonomic remarks of Pincerna in Do and Nguyen (2022) and Páll-Gergely et al. (2024).</p></div>	https://treatment.plazi.org/id/4E19A43A89F0597881FF4A9DE04FD1E0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jirapatrasilp, Parin;Inkhavilay, Khamla;Somvongsa, Chanvilay;Malavong, Banchai;Janjai, Teerangkul;Jeratthitikul, Ekgachai;Likhitrakarn, Natdanai;Srisonchai, Ruttapon;Sutcharit, Chirasak	Jirapatrasilp, Parin, Inkhavilay, Khamla, Somvongsa, Chanvilay, Malavong, Banchai, Janjai, Teerangkul, Jeratthitikul, Ekgachai, Likhitrakarn, Natdanai, Srisonchai, Ruttapon, Sutcharit, Chirasak (2026): The operculate micro land snails from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Laos, with description of new species. ZooKeys 1290: 257-291, DOI: 10.3897/zookeys.1290.198096
54BDA81375B65412AFB6A3B52425C691.text	54BDA81375B65412AFB6A3B52425C691.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pincerna vanbuensis (Bavay & Dautzenberg 1900)	<div><p>Pincerna vanbuensis (Bavay &amp; Dautzenberg, 1900)</p><p>Fig. 9</p><p>Alycaeus (Dyorix [sic]) vanbuensis Bavay &amp; Dautzenberg, 1900: 120 . Type locality: Van Bu [Son La Province, Vietnam].</p><p>Alycaeus vanbuensis – Inkhavilay et al. 2019: 13, fig. 4 c – e.</p><p>Pincerna vanbuensis – Do and Nguyen 2022: 369, 372, figs 3 a – e, 6 e.</p><p>Pincerna (?) vanbuesis – Páll-Gergely 2023 b: 269–272, figs 3 h, i, 12, 14 g.</p><p>Material examined.</p><p>Laos • 5 shells; Houaphanh Province, Viengxay District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.17461&amp;materialsCitation.latitude=20.46075" title="Search Plazi for locations around (long 104.17461/lat 20.46075)">Ban Hang Long village</a>; 20°27'38.7"N, 104°10'28.6"E; 840 m a. s. l.; 6 Oct. 2022; leg. K. Inkhavilay; CUMZ 14485 (Fig. 9) .</p><p>Diagnosis.</p><p>R 1 ~ 3 ¾ whorls with regular, fine radial ribs. R 2 ~ ¼ whorl and nearly as long as R 3; R 2 with very fine and close-set ribs. R 3 with less distinct ribs; apertural tube coiling slightly oblique to horizontal axis of the shell, expanding near the end of the aperture. Inner peristome blunt, slightly protruding. Outer peristome thickened, multilayered, expanded, slightly reflected. Umbilicus mostly closed by the expanded columellar extension of outer peristome.</p><p>Differential diagnosis.</p><p>See Páll-Gergely (2023 b: 270).</p><p>Distribution.</p><p>Southern China, northern Vietnam; Luang Prabang, Luang Namtha, Khammouane (Páll-Gergely 2023 b), and Houaphanh provinces, Laos.</p><p>Remarks.</p><p>Samples from different localities are variable in shell size, density, and strength of R 1 ribs, and the extent of close umbilicus in shells with narrower body whorl (Páll-Gergely 2023 b). See earlier taxonomic treatment of P. vanbuensis in Do and Nguyen (2022) and Páll-Gergely (2023 b).</p></div>	https://treatment.plazi.org/id/54BDA81375B65412AFB6A3B52425C691	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Jirapatrasilp, Parin;Inkhavilay, Khamla;Somvongsa, Chanvilay;Malavong, Banchai;Janjai, Teerangkul;Jeratthitikul, Ekgachai;Likhitrakarn, Natdanai;Srisonchai, Ruttapon;Sutcharit, Chirasak	Jirapatrasilp, Parin, Inkhavilay, Khamla, Somvongsa, Chanvilay, Malavong, Banchai, Janjai, Teerangkul, Jeratthitikul, Ekgachai, Likhitrakarn, Natdanai, Srisonchai, Ruttapon, Sutcharit, Chirasak (2026): The operculate micro land snails from the Alycaeinae (Caenogastropoda, Cyclophoridae) in Laos, with description of new species. ZooKeys 1290: 257-291, DOI: 10.3897/zookeys.1290.198096
