taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
BD2BFCA5A45D5702BB41006FFA58146D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540185	https://doi.org/10.3897/asp.84.e174748.figure3	Figure 3. Live specimens. a, b Caipira mineira Huber sp. nov. from Gruta do Janelão; male and female with egg-sac. c, d Caipira baiana Huber sp. nov. from W of Queimada Nova; male and female with egg-sac.	Figure 3. Live specimens. a, b Caipira mineira Huber sp. nov. from Gruta do Janelão; male and female with egg-sac. c, d Caipira baiana Huber sp. nov. from W of Queimada Nova; male and female with egg-sac.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
BD2BFCA5A45D5702BB41006FFA58146D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540187	https://doi.org/10.3897/asp.84.e174748.figure5	Figure 5. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Ar 24704. Left genital palp, prolateral (a), dorsal (b), and retrolateral (c) views. Scale bar: 0.2 mm.	Figure 5. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Ar 24704. Left genital palp, prolateral (a), dorsal (b), and retrolateral (c) views. Scale bar: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
BD2BFCA5A45D5702BB41006FFA58146D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540189	https://doi.org/10.3897/asp.84.e174748.figure7	Figure 7. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; CHNUFPI 3505 (male), ZFMK Ar 24704 (female). a, b Male chelicerae, frontal and lateral views. c, d Cleared female genitalia in ventral (external) and dorsal (internal) views. Abbreviations: ca, cheliceral apophysis; ep, epigynal pocket; pp, pore plate. Scale bars: 0.2 mm.	Figure 7. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; CHNUFPI 3505 (male), ZFMK Ar 24704 (female). a, b Male chelicerae, frontal and lateral views. c, d Cleared female genitalia in ventral (external) and dorsal (internal) views. Abbreviations: ca, cheliceral apophysis; ep, epigynal pocket; pp, pore plate. Scale bars: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
BD2BFCA5A45D5702BB41006FFA58146D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540198	https://doi.org/10.3897/asp.84.e174748.figure16	Figure 16. Caipira baiana Huber sp. nov.; male from W of Queimada Nova; ZFMK Ar 24706. a – c Left tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow: process at basis of procursus. d – f Left genital bulb, prolateral, dorsal, and retrolateral views; arrows: prolateral-dorsal ridge. Scale bars: 0.1 mm.	Figure 16. Caipira baiana Huber sp. nov.; male from W of Queimada Nova; ZFMK Ar 24706. a – c Left tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow: process at basis of procursus. d – f Left genital bulb, prolateral, dorsal, and retrolateral views; arrows: prolateral-dorsal ridge. Scale bars: 0.1 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
BD2BFCA5A45D5702BB41006FFA58146D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540197	https://doi.org/10.3897/asp.84.e174748.figure15	Figure 15. Caipira baiana Huber sp. nov.; male from W of Queimada Nova; ZFMK Ar 24706. Left genital palp, prolateral (a), dorsal (b), and retrolateral (c) views; arrow: sclerotized apophysis on femur. Scale bar: 0.2 mm.	Figure 15. Caipira baiana Huber sp. nov.; male from W of Queimada Nova; ZFMK Ar 24706. Left genital palp, prolateral (a), dorsal (b), and retrolateral (c) views; arrow: sclerotized apophysis on femur. Scale bar: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
BD2BFCA5A45D5702BB41006FFA58146D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540199	https://doi.org/10.3897/asp.84.e174748.figure17	Figure 17. Caipira baiana Huber sp. nov.; male and female from W of Queimada Nova; ZFMK Ar 24706. a Male ocular area, clypeus, and chelicerae, oblique frontal view. b, c Male chelicerae, frontal and lateral views. d, e Cleared female genitalia, ventral and dorsal views. Abbreviations: ca, cheliceral apophysis; ep, epigynal pocket; pp, pore plate. Scale bars: 0.2 mm.	Figure 17. Caipira baiana Huber sp. nov.; male and female from W of Queimada Nova; ZFMK Ar 24706. a Male ocular area, clypeus, and chelicerae, oblique frontal view. b, c Male chelicerae, frontal and lateral views. d, e Cleared female genitalia, ventral and dorsal views. Abbreviations: ca, cheliceral apophysis; ep, epigynal pocket; pp, pore plate. Scale bars: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
BD2BFCA5A45D5702BB41006FFA58146D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540200	https://doi.org/10.3897/asp.84.e174748.figure18	Figure 18. Caipira baiana Huber sp. nov.; female from W of Queimada Nova; ZFMK Ar 24706. a, b Abdomen ventral view, and epigynum at higher magnification. c, d Cleared female genitalia, ventral and dorsal views. Scale bars: 0.2 mm.	Figure 18. Caipira baiana Huber sp. nov.; female from W of Queimada Nova; ZFMK Ar 24706. a, b Abdomen ventral view, and epigynum at higher magnification. c, d Cleared female genitalia, ventral and dorsal views. Scale bars: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
BD2BFCA5A45D5702BB41006FFA58146D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540186	https://doi.org/10.3897/asp.84.e174748.figure4	Figure 4. Known geographic distribution of Caipira gen. nov., including Caipira baiana Huber sp. nov. (red square) and Caipira mineira Huber sp. nov. (green circle). Colored areas represent the main districts of the Caatinga domain sensu Moro et al. (2024): CDI – Chapada Diamantina; IRE – Irecê; PER – Peruaçu; SDS – Southern Depressão Sertaneja; and SFD – São Francisco Dunes. Other biomes are not shown. Note: Peruaçu and Irecê districts are the only limestone outcrop terrains in this region, separated by the crystalline terrains of the Southern Depressão Sertaneja and the Chapada Diamantina.	Figure 4. Known geographic distribution of Caipira gen. nov., including Caipira baiana Huber sp. nov. (red square) and Caipira mineira Huber sp. nov. (green circle). Colored areas represent the main districts of the Caatinga domain sensu Moro et al. (2024): CDI – Chapada Diamantina; IRE – Irecê; PER – Peruaçu; SDS – Southern Depressão Sertaneja; and SFD – São Francisco Dunes. Other biomes are not shown. Note: Peruaçu and Irecê districts are the only limestone outcrop terrains in this region, separated by the crystalline terrains of the Southern Depressão Sertaneja and the Chapada Diamantina.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
BD2BFCA5A45D5702BB41006FFA58146D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540201	https://doi.org/10.3897/asp.84.e174748.figure19	Figure 19. Type localities of newly described species. a Gruta do Janelão, entrance area; Caipira mineira Huber sp. nov. was found ~ 100–300 m from the cave entrance, in the aphotic zone. b Thorny woodland W of Queimada Nova; Caipira baiana Huber sp. nov. was found in the collapsed rock wall visible in the left central part of the photo. Photos BAH.	Figure 19. Type localities of newly described species. a Gruta do Janelão, entrance area; Caipira mineira Huber sp. nov. was found ~ 100–300 m from the cave entrance, in the aphotic zone. b Thorny woodland W of Queimada Nova; Caipira baiana Huber sp. nov. was found in the collapsed rock wall visible in the left central part of the photo. Photos BAH.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540185	https://doi.org/10.3897/asp.84.e174748.figure3	Figure 3. Live specimens. a, b Caipira mineira Huber sp. nov. from Gruta do Janelão; male and female with egg-sac. c, d Caipira baiana Huber sp. nov. from W of Queimada Nova; male and female with egg-sac.	Figure 3. Live specimens. a, b Caipira mineira Huber sp. nov. from Gruta do Janelão; male and female with egg-sac. c, d Caipira baiana Huber sp. nov. from W of Queimada Nova; male and female with egg-sac.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540188	https://doi.org/10.3897/asp.84.e174748.figure6	Figure 6. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; CHNUFPI 3505. a – c Left tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow: prolateral-ventral process. d – f Left genital bulb, prolateral, dorsal, and retrolateral views; arrows: prolateral-dorsal ridge. Scale bars: 0.2 mm.	Figure 6. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; CHNUFPI 3505. a – c Left tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow: prolateral-ventral process. d – f Left genital bulb, prolateral, dorsal, and retrolateral views; arrows: prolateral-dorsal ridge. Scale bars: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540189	https://doi.org/10.3897/asp.84.e174748.figure7	Figure 7. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; CHNUFPI 3505 (male), ZFMK Ar 24704 (female). a, b Male chelicerae, frontal and lateral views. c, d Cleared female genitalia in ventral (external) and dorsal (internal) views. Abbreviations: ca, cheliceral apophysis; ep, epigynal pocket; pp, pore plate. Scale bars: 0.2 mm.	Figure 7. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; CHNUFPI 3505 (male), ZFMK Ar 24704 (female). a, b Male chelicerae, frontal and lateral views. c, d Cleared female genitalia in ventral (external) and dorsal (internal) views. Abbreviations: ca, cheliceral apophysis; ep, epigynal pocket; pp, pore plate. Scale bars: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540195	https://doi.org/10.3897/asp.84.e174748.figure13	Figure 13. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Br 22-166. a Sexually dimorphic short ‘ vertical’ hair (arrow) on left tibia 1. b Putative chemoreceptor on left metatarsus 1. c Round cuticular plate (arrow) on right tarsus 4. d Tip of left tarsus 1, prolateral view. e Left tarsus 3, prolateral-distal view. f Tip of left tarsus 4, prolateral view, showing two comb hairs (arrows). Scale bars: 10 µm (a, d – f), 1 µm (b), 2 µm (c).	Figure 13. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Br 22-166. a Sexually dimorphic short ‘ vertical’ hair (arrow) on left tibia 1. b Putative chemoreceptor on left metatarsus 1. c Round cuticular plate (arrow) on right tarsus 4. d Tip of left tarsus 1, prolateral view. e Left tarsus 3, prolateral-distal view. f Tip of left tarsus 4, prolateral view, showing two comb hairs (arrows). Scale bars: 10 µm (a, d – f), 1 µm (b), 2 µm (c).	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540202	https://doi.org/10.3897/asp.84.e174748.figure20	Figure 20. a Ratio of male tibia 1 / male tibia 2 in 1662 species (x-axis) of Pholcidae, taken from the taxonomic literature and sorted in each subfamily from smallest to largest values (Art. = Arteminae; Cai. = Caipirinae subfam. nov.; Mod. = Modisiminae; Nin. = Ninetinae; Pho. = Pholcinae; Phy. = Physocyclinae subfam. nov.; Sme. = Smeringopinae). Note that the values for the two new species of Caipirinae subfam. nov. described herein are among the lowest known in Pholcidae. b Scatter plot of the ratio male tibia 1 / male tibia 2 on a proxy of male leg length (tibia 1 length). Note that the lower left corner of the plot is almost entirely filled by Ninetinae and Caipirinae subfam. nov. Erroneous and dubious measurements reported in the literature were excluded (Table S 6).	Figure 20. a Ratio of male tibia 1 / male tibia 2 in 1662 species (x-axis) of Pholcidae, taken from the taxonomic literature and sorted in each subfamily from smallest to largest values (Art. = Arteminae; Cai. = Caipirinae subfam. nov.; Mod. = Modisiminae; Nin. = Ninetinae; Pho. = Pholcinae; Phy. = Physocyclinae subfam. nov.; Sme. = Smeringopinae). Note that the values for the two new species of Caipirinae subfam. nov. described herein are among the lowest known in Pholcidae. b Scatter plot of the ratio male tibia 1 / male tibia 2 on a proxy of male leg length (tibia 1 length). Note that the lower left corner of the plot is almost entirely filled by Ninetinae and Caipirinae subfam. nov. Erroneous and dubious measurements reported in the literature were excluded (Table S 6).	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540191	https://doi.org/10.3897/asp.84.e174748.figure9	Figure 9. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Male prosoma, frontal view. b Female prosoma, frontal view. c Right male frontal cheliceral apophysis, frontal view. d Right male chelicera, frontal-lateral view. e Right female palp and chelicera, frontal-lateral view. f Male gonopore and epiandrous spigots (arrows), ventral view. Scale bars: 100 µm (a, b), 10 µm (c, d, f), 20 µm (e).	Figure 9. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Male prosoma, frontal view. b Female prosoma, frontal view. c Right male frontal cheliceral apophysis, frontal view. d Right male chelicera, frontal-lateral view. e Right female palp and chelicera, frontal-lateral view. f Male gonopore and epiandrous spigots (arrows), ventral view. Scale bars: 100 µm (a, b), 10 µm (c, d, f), 20 µm (e).	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540199	https://doi.org/10.3897/asp.84.e174748.figure17	Figure 17. Caipira baiana Huber sp. nov.; male and female from W of Queimada Nova; ZFMK Ar 24706. a Male ocular area, clypeus, and chelicerae, oblique frontal view. b, c Male chelicerae, frontal and lateral views. d, e Cleared female genitalia, ventral and dorsal views. Abbreviations: ca, cheliceral apophysis; ep, epigynal pocket; pp, pore plate. Scale bars: 0.2 mm.	Figure 17. Caipira baiana Huber sp. nov.; male and female from W of Queimada Nova; ZFMK Ar 24706. a Male ocular area, clypeus, and chelicerae, oblique frontal view. b, c Male chelicerae, frontal and lateral views. d, e Cleared female genitalia, ventral and dorsal views. Abbreviations: ca, cheliceral apophysis; ep, epigynal pocket; pp, pore plate. Scale bars: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540192	https://doi.org/10.3897/asp.84.e174748.figure10	Figure 10. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Epigynum, ventral view, showing pockets (arrows). b Male anterior lateral spinnerets (ALS) and posterior median spinnerets (PMS). c – e Female ALS and PMS. f Female PMS. Scale bars: 100 µm (a), 10 µm (b – e), 2 µm (f).	Figure 10. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Epigynum, ventral view, showing pockets (arrows). b Male anterior lateral spinnerets (ALS) and posterior median spinnerets (PMS). c – e Female ALS and PMS. f Female PMS. Scale bars: 100 µm (a), 10 µm (b – e), 2 µm (f).	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540187	https://doi.org/10.3897/asp.84.e174748.figure5	Figure 5. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Ar 24704. Left genital palp, prolateral (a), dorsal (b), and retrolateral (c) views. Scale bar: 0.2 mm.	Figure 5. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Ar 24704. Left genital palp, prolateral (a), dorsal (b), and retrolateral (c) views. Scale bar: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540197	https://doi.org/10.3897/asp.84.e174748.figure15	Figure 15. Caipira baiana Huber sp. nov.; male from W of Queimada Nova; ZFMK Ar 24706. Left genital palp, prolateral (a), dorsal (b), and retrolateral (c) views; arrow: sclerotized apophysis on femur. Scale bar: 0.2 mm.	Figure 15. Caipira baiana Huber sp. nov.; male from W of Queimada Nova; ZFMK Ar 24706. Left genital palp, prolateral (a), dorsal (b), and retrolateral (c) views; arrow: sclerotized apophysis on femur. Scale bar: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540193	https://doi.org/10.3897/asp.84.e174748.figure11	Figure 11. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Br 22-166. a Right palp, retrolateral view. b Left palp prolateral view. c Right palp, retrolateral-dorsal view. c Left palp, prolateral-dorsal view. e Left palp, dorsal view. f Tip of right procursus, retrolateral view. Arrows in a and c point at sperm duct opening. Abbreviations: b, genital bulb; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 100 µm (a – e), 20 µm (f).	Figure 11. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Br 22-166. a Right palp, retrolateral view. b Left palp prolateral view. c Right palp, retrolateral-dorsal view. c Left palp, prolateral-dorsal view. e Left palp, dorsal view. f Tip of right procursus, retrolateral view. Arrows in a and c point at sperm duct opening. Abbreviations: b, genital bulb; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 100 µm (a – e), 20 µm (f).	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540194	https://doi.org/10.3897/asp.84.e174748.figure12	Figure 12. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Right female metatarsus 4, showing similarity of trichobothrium (t) and slender metatarsal hair (arrow). b Left male metatarsus 3, prolateral view, showing three slender metatarsal hairs (arrows). c Right female metatarsus 4, bases of regular mechanoreceptor and of slender metatarsal hair (arrow). d Tip of slender metatarsal hair on left male metatarsus 3. e Female palpal tibia, showing single trichobothrium. f Prolateral trichobothrium on left male tibia 3. Scale bars: 10 µm (a, b, e, f), 3 µm (c), 2 µm (d).	Figure 12. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Right female metatarsus 4, showing similarity of trichobothrium (t) and slender metatarsal hair (arrow). b Left male metatarsus 3, prolateral view, showing three slender metatarsal hairs (arrows). c Right female metatarsus 4, bases of regular mechanoreceptor and of slender metatarsal hair (arrow). d Tip of slender metatarsal hair on left male metatarsus 3. e Female palpal tibia, showing single trichobothrium. f Prolateral trichobothrium on left male tibia 3. Scale bars: 10 µm (a, b, e, f), 3 µm (c), 2 µm (d).	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
314ECED4A18F552B8881739753D7C75F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540186	https://doi.org/10.3897/asp.84.e174748.figure4	Figure 4. Known geographic distribution of Caipira gen. nov., including Caipira baiana Huber sp. nov. (red square) and Caipira mineira Huber sp. nov. (green circle). Colored areas represent the main districts of the Caatinga domain sensu Moro et al. (2024): CDI – Chapada Diamantina; IRE – Irecê; PER – Peruaçu; SDS – Southern Depressão Sertaneja; and SFD – São Francisco Dunes. Other biomes are not shown. Note: Peruaçu and Irecê districts are the only limestone outcrop terrains in this region, separated by the crystalline terrains of the Southern Depressão Sertaneja and the Chapada Diamantina.	Figure 4. Known geographic distribution of Caipira gen. nov., including Caipira baiana Huber sp. nov. (red square) and Caipira mineira Huber sp. nov. (green circle). Colored areas represent the main districts of the Caatinga domain sensu Moro et al. (2024): CDI – Chapada Diamantina; IRE – Irecê; PER – Peruaçu; SDS – Southern Depressão Sertaneja; and SFD – São Francisco Dunes. Other biomes are not shown. Note: Peruaçu and Irecê districts are the only limestone outcrop terrains in this region, separated by the crystalline terrains of the Southern Depressão Sertaneja and the Chapada Diamantina.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540185	https://doi.org/10.3897/asp.84.e174748.figure3	Figure 3. Live specimens. a, b Caipira mineira Huber sp. nov. from Gruta do Janelão; male and female with egg-sac. c, d Caipira baiana Huber sp. nov. from W of Queimada Nova; male and female with egg-sac.	Figure 3. Live specimens. a, b Caipira mineira Huber sp. nov. from Gruta do Janelão; male and female with egg-sac. c, d Caipira baiana Huber sp. nov. from W of Queimada Nova; male and female with egg-sac.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540187	https://doi.org/10.3897/asp.84.e174748.figure5	Figure 5. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Ar 24704. Left genital palp, prolateral (a), dorsal (b), and retrolateral (c) views. Scale bar: 0.2 mm.	Figure 5. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Ar 24704. Left genital palp, prolateral (a), dorsal (b), and retrolateral (c) views. Scale bar: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540189	https://doi.org/10.3897/asp.84.e174748.figure7	Figure 7. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; CHNUFPI 3505 (male), ZFMK Ar 24704 (female). a, b Male chelicerae, frontal and lateral views. c, d Cleared female genitalia in ventral (external) and dorsal (internal) views. Abbreviations: ca, cheliceral apophysis; ep, epigynal pocket; pp, pore plate. Scale bars: 0.2 mm.	Figure 7. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; CHNUFPI 3505 (male), ZFMK Ar 24704 (female). a, b Male chelicerae, frontal and lateral views. c, d Cleared female genitalia in ventral (external) and dorsal (internal) views. Abbreviations: ca, cheliceral apophysis; ep, epigynal pocket; pp, pore plate. Scale bars: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540188	https://doi.org/10.3897/asp.84.e174748.figure6	Figure 6. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; CHNUFPI 3505. a – c Left tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow: prolateral-ventral process. d – f Left genital bulb, prolateral, dorsal, and retrolateral views; arrows: prolateral-dorsal ridge. Scale bars: 0.2 mm.	Figure 6. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; CHNUFPI 3505. a – c Left tarsus and procursus, prolateral, dorsal, and retrolateral views; arrow: prolateral-ventral process. d – f Left genital bulb, prolateral, dorsal, and retrolateral views; arrows: prolateral-dorsal ridge. Scale bars: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540191	https://doi.org/10.3897/asp.84.e174748.figure9	Figure 9. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Male prosoma, frontal view. b Female prosoma, frontal view. c Right male frontal cheliceral apophysis, frontal view. d Right male chelicera, frontal-lateral view. e Right female palp and chelicera, frontal-lateral view. f Male gonopore and epiandrous spigots (arrows), ventral view. Scale bars: 100 µm (a, b), 10 µm (c, d, f), 20 µm (e).	Figure 9. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Male prosoma, frontal view. b Female prosoma, frontal view. c Right male frontal cheliceral apophysis, frontal view. d Right male chelicera, frontal-lateral view. e Right female palp and chelicera, frontal-lateral view. f Male gonopore and epiandrous spigots (arrows), ventral view. Scale bars: 100 µm (a, b), 10 µm (c, d, f), 20 µm (e).	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540192	https://doi.org/10.3897/asp.84.e174748.figure10	Figure 10. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Epigynum, ventral view, showing pockets (arrows). b Male anterior lateral spinnerets (ALS) and posterior median spinnerets (PMS). c – e Female ALS and PMS. f Female PMS. Scale bars: 100 µm (a), 10 µm (b – e), 2 µm (f).	Figure 10. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Epigynum, ventral view, showing pockets (arrows). b Male anterior lateral spinnerets (ALS) and posterior median spinnerets (PMS). c – e Female ALS and PMS. f Female PMS. Scale bars: 100 µm (a), 10 µm (b – e), 2 µm (f).	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540196	https://doi.org/10.3897/asp.84.e174748.figure14	Figure 14. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Tarsal organ (arrow) and neighboring sensilla on right male palp. b Tip of right female palp, dorsal view, showing tarsal organ (arrow). c – f Tarsal organs on right female tarsus 1 (c), right female tarsus 2 (e), right female tarsus 3 (e), and right male tarsus 4 (f). Scale bars: 2 µm (a, c – f), 10 µm (b).	Figure 14. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Tarsal organ (arrow) and neighboring sensilla on right male palp. b Tip of right female palp, dorsal view, showing tarsal organ (arrow). c – f Tarsal organs on right female tarsus 1 (c), right female tarsus 2 (e), right female tarsus 3 (e), and right male tarsus 4 (f). Scale bars: 2 µm (a, c – f), 10 µm (b).	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540193	https://doi.org/10.3897/asp.84.e174748.figure11	Figure 11. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Br 22-166. a Right palp, retrolateral view. b Left palp prolateral view. c Right palp, retrolateral-dorsal view. c Left palp, prolateral-dorsal view. e Left palp, dorsal view. f Tip of right procursus, retrolateral view. Arrows in a and c point at sperm duct opening. Abbreviations: b, genital bulb; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 100 µm (a – e), 20 µm (f).	Figure 11. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Br 22-166. a Right palp, retrolateral view. b Left palp prolateral view. c Right palp, retrolateral-dorsal view. c Left palp, prolateral-dorsal view. e Left palp, dorsal view. f Tip of right procursus, retrolateral view. Arrows in a and c point at sperm duct opening. Abbreviations: b, genital bulb; f, femur; pa, patella; pr, procursus; ta, tarsus; ti, tibia; tr, trochanter. Scale bars: 100 µm (a – e), 20 µm (f).	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540194	https://doi.org/10.3897/asp.84.e174748.figure12	Figure 12. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Right female metatarsus 4, showing similarity of trichobothrium (t) and slender metatarsal hair (arrow). b Left male metatarsus 3, prolateral view, showing three slender metatarsal hairs (arrows). c Right female metatarsus 4, bases of regular mechanoreceptor and of slender metatarsal hair (arrow). d Tip of slender metatarsal hair on left male metatarsus 3. e Female palpal tibia, showing single trichobothrium. f Prolateral trichobothrium on left male tibia 3. Scale bars: 10 µm (a, b, e, f), 3 µm (c), 2 µm (d).	Figure 12. Caipira mineira Huber sp. nov.; male and female from Gruta do Janelão; ZFMK Br 22-166. a Right female metatarsus 4, showing similarity of trichobothrium (t) and slender metatarsal hair (arrow). b Left male metatarsus 3, prolateral view, showing three slender metatarsal hairs (arrows). c Right female metatarsus 4, bases of regular mechanoreceptor and of slender metatarsal hair (arrow). d Tip of slender metatarsal hair on left male metatarsus 3. e Female palpal tibia, showing single trichobothrium. f Prolateral trichobothrium on left male tibia 3. Scale bars: 10 µm (a, b, e, f), 3 µm (c), 2 µm (d).	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540195	https://doi.org/10.3897/asp.84.e174748.figure13	Figure 13. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Br 22-166. a Sexually dimorphic short ‘ vertical’ hair (arrow) on left tibia 1. b Putative chemoreceptor on left metatarsus 1. c Round cuticular plate (arrow) on right tarsus 4. d Tip of left tarsus 1, prolateral view. e Left tarsus 3, prolateral-distal view. f Tip of left tarsus 4, prolateral view, showing two comb hairs (arrows). Scale bars: 10 µm (a, d – f), 1 µm (b), 2 µm (c).	Figure 13. Caipira mineira Huber sp. nov.; male from Gruta do Janelão; ZFMK Br 22-166. a Sexually dimorphic short ‘ vertical’ hair (arrow) on left tibia 1. b Putative chemoreceptor on left metatarsus 1. c Round cuticular plate (arrow) on right tarsus 4. d Tip of left tarsus 1, prolateral view. e Left tarsus 3, prolateral-distal view. f Tip of left tarsus 4, prolateral view, showing two comb hairs (arrows). Scale bars: 10 µm (a, d – f), 1 µm (b), 2 µm (c).	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540190	https://doi.org/10.3897/asp.84.e174748.figure8	Figure 8. Caipira mineira Huber sp. nov.; two females from Gruta do Janelão; ZFMK Ar 24704. a, b Epigyna, ventral views. c, d Cleared female genitalia of female shown in a, ventral and dorsal views. e, f Cleared female genitalia of female shown in b, ventral and dorsal views. Scale bars: 0.2 mm.	Figure 8. Caipira mineira Huber sp. nov.; two females from Gruta do Janelão; ZFMK Ar 24704. a, b Epigyna, ventral views. c, d Cleared female genitalia of female shown in a, ventral and dorsal views. e, f Cleared female genitalia of female shown in b, ventral and dorsal views. Scale bars: 0.2 mm.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540186	https://doi.org/10.3897/asp.84.e174748.figure4	Figure 4. Known geographic distribution of Caipira gen. nov., including Caipira baiana Huber sp. nov. (red square) and Caipira mineira Huber sp. nov. (green circle). Colored areas represent the main districts of the Caatinga domain sensu Moro et al. (2024): CDI – Chapada Diamantina; IRE – Irecê; PER – Peruaçu; SDS – Southern Depressão Sertaneja; and SFD – São Francisco Dunes. Other biomes are not shown. Note: Peruaçu and Irecê districts are the only limestone outcrop terrains in this region, separated by the crystalline terrains of the Southern Depressão Sertaneja and the Chapada Diamantina.	Figure 4. Known geographic distribution of Caipira gen. nov., including Caipira baiana Huber sp. nov. (red square) and Caipira mineira Huber sp. nov. (green circle). Colored areas represent the main districts of the Caatinga domain sensu Moro et al. (2024): CDI – Chapada Diamantina; IRE – Irecê; PER – Peruaçu; SDS – Southern Depressão Sertaneja; and SFD – São Francisco Dunes. Other biomes are not shown. Note: Peruaçu and Irecê districts are the only limestone outcrop terrains in this region, separated by the crystalline terrains of the Southern Depressão Sertaneja and the Chapada Diamantina.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
1A4554885D765451AE8B227FF261DBE8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540201	https://doi.org/10.3897/asp.84.e174748.figure19	Figure 19. Type localities of newly described species. a Gruta do Janelão, entrance area; Caipira mineira Huber sp. nov. was found ~ 100–300 m from the cave entrance, in the aphotic zone. b Thorny woodland W of Queimada Nova; Caipira baiana Huber sp. nov. was found in the collapsed rock wall visible in the left central part of the photo. Photos BAH.	Figure 19. Type localities of newly described species. a Gruta do Janelão, entrance area; Caipira mineira Huber sp. nov. was found ~ 100–300 m from the cave entrance, in the aphotic zone. b Thorny woodland W of Queimada Nova; Caipira baiana Huber sp. nov. was found in the collapsed rock wall visible in the left central part of the photo. Photos BAH.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
E2C4717BF85C501793DF27CBACCE3B1B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1540183	https://doi.org/10.3897/asp.84.e174748.figure1	Figure 1. Maximum likelihood tree based on UCE (ultra-conserved elements) data. The tree was constructed with RAxML-NG and matrix 1 using a partitioned strategy and the GTR + R 4 + I + FO model. Circles on the nodes represent bootstrap support (BS) values, with green circles indicating strong support (BS: 95 % – 100 %), orange circles moderate support (BS: 70 % – 95 %), and red circles weak support (BS < 70 %). Representatives of each subfamily on the right, from top: Hoplopholcus konya Huber, 2020; Apokayana kapit (Huber, 2016); Caipira mineira Huber sp. nov.; Artema martensi Huber, 2021; Ibotyporanga walekeru Huber, 2024; Arnapa arfak Huber, 2019; Mesabolivar eberhardi Huber, 2000; photos BAH.	Figure 1. Maximum likelihood tree based on UCE (ultra-conserved elements) data. The tree was constructed with RAxML-NG and matrix 1 using a partitioned strategy and the GTR + R 4 + I + FO model. Circles on the nodes represent bootstrap support (BS) values, with green circles indicating strong support (BS: 95 % – 100 %), orange circles moderate support (BS: 70 % – 95 %), and red circles weak support (BS < 70 %). Representatives of each subfamily on the right, from top: Hoplopholcus konya Huber, 2020; Apokayana kapit (Huber, 2016); Caipira mineira Huber sp. nov.; Artema martensi Huber, 2021; Ibotyporanga walekeru Huber, 2024; Arnapa arfak Huber, 2019; Mesabolivar eberhardi Huber, 2000; photos BAH.	2026-02-16	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.		Zenodo	biologists	Meng, Guanliang;Carvalho, Leonardo S.;Podsiadlowski, Lars;Huber, Bernhard A.			
