taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
6019203B28275126925C7B3160E21525.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462894	https://doi.org/10.3897/zookeys.1259.168521.figure1	Figure 1. Baetis procul sp. nov., larva, habitus. a. Male, dorsal view; b. Male, lateral view; c. Female, dorsal view; d. Female, ventral view. Scale bars: 1 mm.	Figure 1. Baetis procul sp. nov., larva, habitus. a. Male, dorsal view; b. Male, lateral view; c. Female, dorsal view; d. Female, ventral view. Scale bars: 1 mm.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
6019203B28275126925C7B3160E21525.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462895	https://doi.org/10.3897/zookeys.1259.168521.figure2	Figure 2. Baetis procul sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Head; i. Hind protopteron (arrows: costal margin). Scale bars: 100 µm (h); 20 µm (a, c – g); 10 µm (b, i).	Figure 2. Baetis procul sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Head; i. Hind protopteron (arrows: costal margin). Scale bars: 100 µm (h); 20 µm (a, c – g); 10 µm (b, i).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
6019203B28275126925C7B3160E21525.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462896	https://doi.org/10.3897/zookeys.1259.168521.figure3	Figure 3. Baetis procul sp. nov., larva. a. Hind leg; b. Femur, outer margin; c. Femur, inner margin; d. Tibia; e. Tarsus, claw; f. Femoral patch, fore leg; g. Femoral patch, middle leg; h. Femoral patch, hind leg; i. Abdomen; j. Abdominal segments; k, l. Tergalius IV; m. Paraproct. Scale bars: 100 µm (i); 50 µm (a); 20 µm (k); 10 µm (b – h, j, l, m).	Figure 3. Baetis procul sp. nov., larva. a. Hind leg; b. Femur, outer margin; c. Femur, inner margin; d. Tibia; e. Tarsus, claw; f. Femoral patch, fore leg; g. Femoral patch, middle leg; h. Femoral patch, hind leg; i. Abdomen; j. Abdominal segments; k, l. Tergalius IV; m. Paraproct. Scale bars: 100 µm (i); 50 µm (a); 20 µm (k); 10 µm (b – h, j, l, m).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
6019203B28275126925C7B3160E21525.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462897	https://doi.org/10.3897/zookeys.1259.168521.figure4	Figure 4. Baetis procul sp. nov., female imago. a. Habitus, lateral view (arrow: hind wing); b. Habitus, dorsal view; c. Fore leg; d. Wings; e. Hind wing; f. Apex of fore wing. Scale bars: 1 mm (a, b); 200 µm (d); 100 µm (c); 20 µm (e); 10 µm (f).	Figure 4. Baetis procul sp. nov., female imago. a. Habitus, lateral view (arrow: hind wing); b. Habitus, dorsal view; c. Fore leg; d. Wings; e. Hind wing; f. Apex of fore wing. Scale bars: 1 mm (a, b); 200 µm (d); 100 µm (c); 20 µm (e); 10 µm (f).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
6019203B28275126925C7B3160E21525.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462929	https://doi.org/10.3897/zookeys.1259.168521.figure36	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
6019203B28275126925C7B3160E21525.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462930	https://doi.org/10.3897/zookeys.1259.168521.figure37	Figure 37. Habitats. a. Creek near Outback Hotel (location D 1); b. Sabeto River, near Sabeto (location I; lowland river); c. Biausevu River, near Biausevu (location G); d. Waisila creek, Colo-I-Suva Forest Park (location H). Characteristics of locations see Table 3.	Figure 37. Habitats. a. Creek near Outback Hotel (location D 1); b. Sabeto River, near Sabeto (location I; lowland river); c. Biausevu River, near Biausevu (location G); d. Waisila creek, Colo-I-Suva Forest Park (location H). Characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
6019203B28275126925C7B3160E21525.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462932	https://doi.org/10.3897/zookeys.1259.168521.figure39	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
FA2598CA45AE547CA1C42BCE52C22212.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462899	https://doi.org/10.3897/zookeys.1259.168521.figure6	Figure 6. Papuanatula (Fijifiliola) aji sp. nov., larva, habitus. a. dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 6. Papuanatula (Fijifiliola) aji sp. nov., larva, habitus. a. dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
FA2598CA45AE547CA1C42BCE52C22212.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462900	https://doi.org/10.3897/zookeys.1259.168521.figure7	Figure 7. Papuanatula (Fijifiliola) aji sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Paracercus; i. Cercus; j. Male mature larva, developing subimaginal gonostylus; k. Fore protoptera, protuberance. Scale bars: 20 µm (a, c – h). 10 µm (b, i – k).	Figure 7. Papuanatula (Fijifiliola) aji sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Paracercus; i. Cercus; j. Male mature larva, developing subimaginal gonostylus; k. Fore protoptera, protuberance. Scale bars: 20 µm (a, c – h). 10 µm (b, i – k).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
FA2598CA45AE547CA1C42BCE52C22212.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462901	https://doi.org/10.3897/zookeys.1259.168521.figure8	Figure 8. Papuanatula (Fijifiliola) aji sp. nov., larva. a. Hind leg; b. Femur, outer margin; c. Tibia; d. Tarsus, outer margin; e. Tarsus, claw; f. Abdomen; g. Abdominal terga; h. Paraproct; i. Tergalius IV; j. Head. Scale bars: 100 µm (f, j). 50 µm (a); 10 µm (b – e, g – i).	Figure 8. Papuanatula (Fijifiliola) aji sp. nov., larva. a. Hind leg; b. Femur, outer margin; c. Tibia; d. Tarsus, outer margin; e. Tarsus, claw; f. Abdomen; g. Abdominal terga; h. Paraproct; i. Tergalius IV; j. Head. Scale bars: 100 µm (f, j). 50 µm (a); 10 µm (b – e, g – i).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
FA2598CA45AE547CA1C42BCE52C22212.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462929	https://doi.org/10.3897/zookeys.1259.168521.figure36	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
FA2598CA45AE547CA1C42BCE52C22212.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462930	https://doi.org/10.3897/zookeys.1259.168521.figure37	Figure 37. Habitats. a. Creek near Outback Hotel (location D 1); b. Sabeto River, near Sabeto (location I; lowland river); c. Biausevu River, near Biausevu (location G); d. Waisila creek, Colo-I-Suva Forest Park (location H). Characteristics of locations see Table 3.	Figure 37. Habitats. a. Creek near Outback Hotel (location D 1); b. Sabeto River, near Sabeto (location I; lowland river); c. Biausevu River, near Biausevu (location G); d. Waisila creek, Colo-I-Suva Forest Park (location H). Characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
FA2598CA45AE547CA1C42BCE52C22212.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462931	https://doi.org/10.3897/zookeys.1259.168521.figure38	Figure 38. Habitats. a. Tributary to Sekawa River, near Nakawaga (location E; clean lowland river); b. Sekawa River near Nakawaga (location E 1; lowland river, disturbed by human activities; many filamentous green algae); c. Seaqaqa River, bridge near Saivou (location F). Characteristics of locations see Table 3.	Figure 38. Habitats. a. Tributary to Sekawa River, near Nakawaga (location E; clean lowland river); b. Sekawa River near Nakawaga (location E 1; lowland river, disturbed by human activities; many filamentous green algae); c. Seaqaqa River, bridge near Saivou (location F). Characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
FA2598CA45AE547CA1C42BCE52C22212.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462932	https://doi.org/10.3897/zookeys.1259.168521.figure39	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
197B2BA0998D5793B6655DEF9025D817.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462902	https://doi.org/10.3897/zookeys.1259.168521.figure9	Figure 9. Papuanatula (Fijifiliola) aurantica sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 9. Papuanatula (Fijifiliola) aurantica sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
197B2BA0998D5793B6655DEF9025D817.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462903	https://doi.org/10.3897/zookeys.1259.168521.figure10	Figure 10. Papuanatula (Fijifiliola) aurantica sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Head; i. Paracercus; j, k. Cerci. Scale bars: 100 µm (h); 20 µm (a, c – g, i); 10 µm (b, j, k).	Figure 10. Papuanatula (Fijifiliola) aurantica sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Head; i. Paracercus; j, k. Cerci. Scale bars: 100 µm (h); 20 µm (a, c – g, i); 10 µm (b, j, k).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
197B2BA0998D5793B6655DEF9025D817.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462904	https://doi.org/10.3897/zookeys.1259.168521.figure11	Figure 11. Papuanatula (Fijifiliola) aurantica sp. nov., larva. a. Hind leg; b. Femur, outer margin; c. Femur, inner margin; d. Tibia; e. Tarsus, outer margin; f. Tarsus, claw; g. Abdominal terga; h. Tergalius III; i. Paraproct. Scale bars: 50 µm (a); 10 µm (b – i).	Figure 11. Papuanatula (Fijifiliola) aurantica sp. nov., larva. a. Hind leg; b. Femur, outer margin; c. Femur, inner margin; d. Tibia; e. Tarsus, outer margin; f. Tarsus, claw; g. Abdominal terga; h. Tergalius III; i. Paraproct. Scale bars: 50 µm (a); 10 µm (b – i).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
197B2BA0998D5793B6655DEF9025D817.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462930	https://doi.org/10.3897/zookeys.1259.168521.figure37	Figure 37. Habitats. a. Creek near Outback Hotel (location D 1); b. Sabeto River, near Sabeto (location I; lowland river); c. Biausevu River, near Biausevu (location G); d. Waisila creek, Colo-I-Suva Forest Park (location H). Characteristics of locations see Table 3.	Figure 37. Habitats. a. Creek near Outback Hotel (location D 1); b. Sabeto River, near Sabeto (location I; lowland river); c. Biausevu River, near Biausevu (location G); d. Waisila creek, Colo-I-Suva Forest Park (location H). Characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
197B2BA0998D5793B6655DEF9025D817.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462932	https://doi.org/10.3897/zookeys.1259.168521.figure39	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
E884A3E29D2351F983039F86830F8025.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462905	https://doi.org/10.3897/zookeys.1259.168521.figure12	Figure 12. Papuanatula (Fijifiliola) bula sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 12. Papuanatula (Fijifiliola) bula sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
E884A3E29D2351F983039F86830F8025.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462906	https://doi.org/10.3897/zookeys.1259.168521.figure13	Figure 13. Papuanatula (Fijifiliola) bula sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Maxillary palp; i. Paracercus; j, k. Cerci; l. Head. Scale bars: 100 µm (l); 20 µm (a, c – g, i); 10 µm (b, h, j, k).	Figure 13. Papuanatula (Fijifiliola) bula sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Maxillary palp; i. Paracercus; j, k. Cerci; l. Head. Scale bars: 100 µm (l); 20 µm (a, c – g, i); 10 µm (b, h, j, k).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
E884A3E29D2351F983039F86830F8025.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462907	https://doi.org/10.3897/zookeys.1259.168521.figure14	Figure 14. Papuanatula (Fijifiliola) bula sp. nov., larva. a. Fore leg; b. Femur, outer margin; c. Femur, inner margin; d. Tibia; e. Tarsus, outer margin; f. Tarsus, claw; g. Abdomen; h. Abdominal terga; i. Tergalius IV; j. Paraproct. Scale bars: 100 µm (g); 50 µm (a); 10 µm (b – f, h – j).	Figure 14. Papuanatula (Fijifiliola) bula sp. nov., larva. a. Fore leg; b. Femur, outer margin; c. Femur, inner margin; d. Tibia; e. Tarsus, outer margin; f. Tarsus, claw; g. Abdomen; h. Abdominal terga; i. Tergalius IV; j. Paraproct. Scale bars: 100 µm (g); 50 µm (a); 10 µm (b – f, h – j).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
E884A3E29D2351F983039F86830F8025.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462930	https://doi.org/10.3897/zookeys.1259.168521.figure37	Figure 37. Habitats. a. Creek near Outback Hotel (location D 1); b. Sabeto River, near Sabeto (location I; lowland river); c. Biausevu River, near Biausevu (location G); d. Waisila creek, Colo-I-Suva Forest Park (location H). Characteristics of locations see Table 3.	Figure 37. Habitats. a. Creek near Outback Hotel (location D 1); b. Sabeto River, near Sabeto (location I; lowland river); c. Biausevu River, near Biausevu (location G); d. Waisila creek, Colo-I-Suva Forest Park (location H). Characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
E884A3E29D2351F983039F86830F8025.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462932	https://doi.org/10.3897/zookeys.1259.168521.figure39	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
F7FA939D52CA596F901BF29529A4F755.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462908	https://doi.org/10.3897/zookeys.1259.168521.figure15	Figure 15. Papuanatula (Fijifiliola) claudia sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 15. Papuanatula (Fijifiliola) claudia sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
F7FA939D52CA596F901BF29529A4F755.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462909	https://doi.org/10.3897/zookeys.1259.168521.figure16	Figure 16. Papuanatula (Fijifiliola) claudia sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Paracercus; i. Cercus; j. Head; k. Mature female larva, abdominal segments III and IV, eggs; l. Eggs in mature female larva. Scale bars: 100 µm (j); 20 µm (a, c – h); 10 µm (b, i, k, l).	Figure 16. Papuanatula (Fijifiliola) claudia sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Paracercus; i. Cercus; j. Head; k. Mature female larva, abdominal segments III and IV, eggs; l. Eggs in mature female larva. Scale bars: 100 µm (j); 20 µm (a, c – h); 10 µm (b, i, k, l).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
F7FA939D52CA596F901BF29529A4F755.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462910	https://doi.org/10.3897/zookeys.1259.168521.figure17	Figure 17. Papuanatula (Fijifiliola) claudia sp. nov., larva. a. Fore leg; b. Femur, outer margin; c. Femur, inner margin; d. Tibia; e. Tarsus, outer margin; f. Tarsus, claw; g. Abdomen; h. Abdominal terga; i, j. Tergalius IV; k. Paraproct. Scale bars: 100 µm (g); 50 µm (a); 20 µm (i); 10 µm (b – f, h, j, k).	Figure 17. Papuanatula (Fijifiliola) claudia sp. nov., larva. a. Fore leg; b. Femur, outer margin; c. Femur, inner margin; d. Tibia; e. Tarsus, outer margin; f. Tarsus, claw; g. Abdomen; h. Abdominal terga; i, j. Tergalius IV; k. Paraproct. Scale bars: 100 µm (g); 50 µm (a); 20 µm (i); 10 µm (b – f, h, j, k).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
F7FA939D52CA596F901BF29529A4F755.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462929	https://doi.org/10.3897/zookeys.1259.168521.figure36	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
F7FA939D52CA596F901BF29529A4F755.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462932	https://doi.org/10.3897/zookeys.1259.168521.figure39	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
648B9C5EC7EB586EA0285FE183F0080A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462911	https://doi.org/10.3897/zookeys.1259.168521.figure18	Figure 18. Papuanatula (Fijifiliola) crussetae sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 18. Papuanatula (Fijifiliola) crussetae sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
648B9C5EC7EB586EA0285FE183F0080A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462912	https://doi.org/10.3897/zookeys.1259.168521.figure19	Figure 19. Papuanatula (Fijifiliola) crussetae sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Head; i. Paracercus; j. Cercus. Scale bars: 100 µm (h); 20 µm (a, c – g, i); 10 µm (b, j).	Figure 19. Papuanatula (Fijifiliola) crussetae sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Head; i. Paracercus; j. Cercus. Scale bars: 100 µm (h); 20 µm (a, c – g, i); 10 µm (b, j).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
648B9C5EC7EB586EA0285FE183F0080A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462913	https://doi.org/10.3897/zookeys.1259.168521.figure20	Figure 20. Papuanatula (Fijifiliola) crussetae sp. nov., larva. a. Middle leg; b. Femur, outer margin; c. Femur, inner margin; d. Tibia, outer margin; e. Tibia, inner margin; f. Tarsus, outer margin; g. Tarsus, inner margin; h. Claw (arrow: posterior seta); i. Abdomen; j. Abdominal terga; k. Paraproct; l, m. Tergalius VI. Scale bars: 100 µm (i); 50 µm (a, l); 10 µm (b – h, j, k, m).	Figure 20. Papuanatula (Fijifiliola) crussetae sp. nov., larva. a. Middle leg; b. Femur, outer margin; c. Femur, inner margin; d. Tibia, outer margin; e. Tibia, inner margin; f. Tarsus, outer margin; g. Tarsus, inner margin; h. Claw (arrow: posterior seta); i. Abdomen; j. Abdominal terga; k. Paraproct; l, m. Tergalius VI. Scale bars: 100 µm (i); 50 µm (a, l); 10 µm (b – h, j, k, m).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
648B9C5EC7EB586EA0285FE183F0080A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462929	https://doi.org/10.3897/zookeys.1259.168521.figure36	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
648B9C5EC7EB586EA0285FE183F0080A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462932	https://doi.org/10.3897/zookeys.1259.168521.figure39	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
2035864D449056DCB9FF4955E3DE41C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462914	https://doi.org/10.3897/zookeys.1259.168521.figure21	Figure 21. Papuanatula (Fijifiliola) flowersi sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 21. Papuanatula (Fijifiliola) flowersi sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
2035864D449056DCB9FF4955E3DE41C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462915	https://doi.org/10.3897/zookeys.1259.168521.figure22	Figure 22. Papuanatula (Fijifiliola) flowersi sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible (detail: subtriangular process); f. Right mandible; g. Maxilla; h. Head; i. Paracercus; j. Cercus. Scale bars: 100 µm (h); 20 µm (a, c – g, i); 10 µm (b, j).	Figure 22. Papuanatula (Fijifiliola) flowersi sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible (detail: subtriangular process); f. Right mandible; g. Maxilla; h. Head; i. Paracercus; j. Cercus. Scale bars: 100 µm (h); 20 µm (a, c – g, i); 10 µm (b, j).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
2035864D449056DCB9FF4955E3DE41C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462916	https://doi.org/10.3897/zookeys.1259.168521.figure23	Figure 23. Papuanatula (Fijifiliola) flowersi sp. nov., larva. a. Fore leg; b. Femur, outer margin; c. Tibia; d. Tarsus, outer margin; e. Tarsus, claw; f. Abdomen; g. Abdominal terga; h. Paraproct; i, j. Tergalius IV. Scale bars: 100 µm (f); 50 µm (a); 20 µm (i); 10 µm (b – e, g, h, j).	Figure 23. Papuanatula (Fijifiliola) flowersi sp. nov., larva. a. Fore leg; b. Femur, outer margin; c. Tibia; d. Tarsus, outer margin; e. Tarsus, claw; f. Abdomen; g. Abdominal terga; h. Paraproct; i, j. Tergalius IV. Scale bars: 100 µm (f); 50 µm (a); 20 µm (i); 10 µm (b – e, g, h, j).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
2035864D449056DCB9FF4955E3DE41C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462929	https://doi.org/10.3897/zookeys.1259.168521.figure36	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
2035864D449056DCB9FF4955E3DE41C1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462932	https://doi.org/10.3897/zookeys.1259.168521.figure39	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	Figure 39. Distribution of described species. a. Baetis procul sp. nov. (locations B, I); b. Papuanatula (Fijifiliola) aji sp. nov. (locations A, B, C, C 1, D, E, E 1, F, I); c. Papuanatula (Fijifiliola) aurantica sp. nov. (location G); d. Papuanatula (Fijifiliola) bula sp. nov. (locations D, D 1, G, H); e. Papuanatula (Fijifiliola) claudia sp. nov., Papuanatula (Fijifiliola) flowersi sp. nov. (locations C, C 1); f. Papuanatula (Fijifiliola) crussetae sp. nov. (location C).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
C992ED638E9253DA8E2146D5C18F23E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462917	https://doi.org/10.3897/zookeys.1259.168521.figure24	Figure 24. Papuanatula (Fijifiliola) gattolliati sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 24. Papuanatula (Fijifiliola) gattolliati sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
C992ED638E9253DA8E2146D5C18F23E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462918	https://doi.org/10.3897/zookeys.1259.168521.figure25	Figure 25. Papuanatula (Fijifiliola) gattolliati sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Head; i. Paracercus; j. Cercus. Scale bars: 100 µm (h); 20 µm (a, c – g, i); 10 µm (b, j).	Figure 25. Papuanatula (Fijifiliola) gattolliati sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Head; i. Paracercus; j. Cercus. Scale bars: 100 µm (h); 20 µm (a, c – g, i); 10 µm (b, j).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
C992ED638E9253DA8E2146D5C18F23E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462919	https://doi.org/10.3897/zookeys.1259.168521.figure26	Figure 26. Papuanatula (Fijifiliola) gattolliati sp. nov., larva. a. Fore leg; b. Femur, outer margin; c. Tibia; d. Tarsus, outer margin; e. Tarsus, claw; f. Abdomen; g. Abdominal terga; h. Paraproct; i, j. Tergalius IV. Scale bars: 100 µm (f); 50 µm (a); 20 µm (i); 10 µm (b – e, g, h, j).	Figure 26. Papuanatula (Fijifiliola) gattolliati sp. nov., larva. a. Fore leg; b. Femur, outer margin; c. Tibia; d. Tarsus, outer margin; e. Tarsus, claw; f. Abdomen; g. Abdominal terga; h. Paraproct; i, j. Tergalius IV. Scale bars: 100 µm (f); 50 µm (a); 20 µm (i); 10 µm (b – e, g, h, j).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
C992ED638E9253DA8E2146D5C18F23E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462931	https://doi.org/10.3897/zookeys.1259.168521.figure38	Figure 38. Habitats. a. Tributary to Sekawa River, near Nakawaga (location E; clean lowland river); b. Sekawa River near Nakawaga (location E 1; lowland river, disturbed by human activities; many filamentous green algae); c. Seaqaqa River, bridge near Saivou (location F). Characteristics of locations see Table 3.	Figure 38. Habitats. a. Tributary to Sekawa River, near Nakawaga (location E; clean lowland river); b. Sekawa River near Nakawaga (location E 1; lowland river, disturbed by human activities; many filamentous green algae); c. Seaqaqa River, bridge near Saivou (location F). Characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
C992ED638E9253DA8E2146D5C18F23E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462933	https://doi.org/10.3897/zookeys.1259.168521.figure40	Figure 40. Distribution of described species. a. Papuanatula (Fijifiliola) gattolliati sp. nov. (locations E, F); b. Papuanatula (Fijifiliola) grisea sp. nov. (locations A, B, I); c. Papuanatula (Fijifiliola) sekawa sp. nov. (location E); d. Papuanatula (Fijifiliola) tenebris sp. nov. (location C 1).	Figure 40. Distribution of described species. a. Papuanatula (Fijifiliola) gattolliati sp. nov. (locations E, F); b. Papuanatula (Fijifiliola) grisea sp. nov. (locations A, B, I); c. Papuanatula (Fijifiliola) sekawa sp. nov. (location E); d. Papuanatula (Fijifiliola) tenebris sp. nov. (location C 1).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
D358252C08B15275A4E5A747C26DE594.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462920	https://doi.org/10.3897/zookeys.1259.168521.figure27	Figure 27. Papuanatula (Fijifiliola) grisea sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 27. Papuanatula (Fijifiliola) grisea sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
D358252C08B15275A4E5A747C26DE594.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462921	https://doi.org/10.3897/zookeys.1259.168521.figure28	Figure 28. Papuanatula (Fijifiliola) grisea sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Paracercus; i. Cercus; j. Head; k. Male mature larva, developing subimaginal gonostylus. Scale bars: 100 µm (j); 50 µm (c); 20 µm (e – h); 10 µm (b, d, i, k).	Figure 28. Papuanatula (Fijifiliola) grisea sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Paracercus; i. Cercus; j. Head; k. Male mature larva, developing subimaginal gonostylus. Scale bars: 100 µm (j); 50 µm (c); 20 µm (e – h); 10 µm (b, d, i, k).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
D358252C08B15275A4E5A747C26DE594.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462922	https://doi.org/10.3897/zookeys.1259.168521.figure29	Figure 29. Papuanatula (Fijifiliola) grisea sp. nov., larva. a. Fore leg; b. Femur, outer margin; c. Tibia; d. Tarsus; e. Claw; f. Abdomen; g. Abdominal terga; h. Paraproct; i. Tergalius IV. Scale bars: 100 µm (f); 50 µm (a); 10 µm (b – e, g – i).	Figure 29. Papuanatula (Fijifiliola) grisea sp. nov., larva. a. Fore leg; b. Femur, outer margin; c. Tibia; d. Tarsus; e. Claw; f. Abdomen; g. Abdominal terga; h. Paraproct; i. Tergalius IV. Scale bars: 100 µm (f); 50 µm (a); 10 µm (b – e, g – i).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
D358252C08B15275A4E5A747C26DE594.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462929	https://doi.org/10.3897/zookeys.1259.168521.figure36	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
D358252C08B15275A4E5A747C26DE594.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462930	https://doi.org/10.3897/zookeys.1259.168521.figure37	Figure 37. Habitats. a. Creek near Outback Hotel (location D 1); b. Sabeto River, near Sabeto (location I; lowland river); c. Biausevu River, near Biausevu (location G); d. Waisila creek, Colo-I-Suva Forest Park (location H). Characteristics of locations see Table 3.	Figure 37. Habitats. a. Creek near Outback Hotel (location D 1); b. Sabeto River, near Sabeto (location I; lowland river); c. Biausevu River, near Biausevu (location G); d. Waisila creek, Colo-I-Suva Forest Park (location H). Characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
D358252C08B15275A4E5A747C26DE594.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462933	https://doi.org/10.3897/zookeys.1259.168521.figure40	Figure 40. Distribution of described species. a. Papuanatula (Fijifiliola) gattolliati sp. nov. (locations E, F); b. Papuanatula (Fijifiliola) grisea sp. nov. (locations A, B, I); c. Papuanatula (Fijifiliola) sekawa sp. nov. (location E); d. Papuanatula (Fijifiliola) tenebris sp. nov. (location C 1).	Figure 40. Distribution of described species. a. Papuanatula (Fijifiliola) gattolliati sp. nov. (locations E, F); b. Papuanatula (Fijifiliola) grisea sp. nov. (locations A, B, I); c. Papuanatula (Fijifiliola) sekawa sp. nov. (location E); d. Papuanatula (Fijifiliola) tenebris sp. nov. (location C 1).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
894E9BAB13B6544F9EA6721A6D829125.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462898	https://doi.org/10.3897/zookeys.1259.168521.figure5	Figure 5. Papuanatula, comparison of main characters between subgenera: left column: Papuanatula s. str.; middle column: Papuafiliola; right column: Fijifiliola subgen. nov.; row one (top): labrum, shape, and dorsal submarginal arc of setae; row two: labial palp segment II / III; row three: right mandible; row four: femur, outer margin row of setae; row five (bottom): tibia, anterior surface, row of setae.	Figure 5. Papuanatula, comparison of main characters between subgenera: left column: Papuanatula s. str.; middle column: Papuafiliola; right column: Fijifiliola subgen. nov.; row one (top): labrum, shape, and dorsal submarginal arc of setae; row two: labial palp segment II / III; row three: right mandible; row four: femur, outer margin row of setae; row five (bottom): tibia, anterior surface, row of setae.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
894E9BAB13B6544F9EA6721A6D829125.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462901	https://doi.org/10.3897/zookeys.1259.168521.figure8	Figure 8. Papuanatula (Fijifiliola) aji sp. nov., larva. a. Hind leg; b. Femur, outer margin; c. Tibia; d. Tarsus, outer margin; e. Tarsus, claw; f. Abdomen; g. Abdominal terga; h. Paraproct; i. Tergalius IV; j. Head. Scale bars: 100 µm (f, j). 50 µm (a); 10 µm (b – e, g – i).	Figure 8. Papuanatula (Fijifiliola) aji sp. nov., larva. a. Hind leg; b. Femur, outer margin; c. Tibia; d. Tarsus, outer margin; e. Tarsus, claw; f. Abdomen; g. Abdominal terga; h. Paraproct; i. Tergalius IV; j. Head. Scale bars: 100 µm (f, j). 50 µm (a); 10 µm (b – e, g – i).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
894E9BAB13B6544F9EA6721A6D829125.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462900	https://doi.org/10.3897/zookeys.1259.168521.figure7	Figure 7. Papuanatula (Fijifiliola) aji sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Paracercus; i. Cercus; j. Male mature larva, developing subimaginal gonostylus; k. Fore protoptera, protuberance. Scale bars: 20 µm (a, c – h). 10 µm (b, i – k).	Figure 7. Papuanatula (Fijifiliola) aji sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Paracercus; i. Cercus; j. Male mature larva, developing subimaginal gonostylus; k. Fore protoptera, protuberance. Scale bars: 20 µm (a, c – h). 10 µm (b, i – k).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
E0640DB98ED65F88851AC3932204375F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462923	https://doi.org/10.3897/zookeys.1259.168521.figure30	Figure 30. Papuanatula (Fijifiliola) sekawa sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 30. Papuanatula (Fijifiliola) sekawa sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
E0640DB98ED65F88851AC3932204375F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462924	https://doi.org/10.3897/zookeys.1259.168521.figure31	Figure 31. Papuanatula (Fijifiliola) sekawa sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Head; i. Paracercus; j. Cercus. Scale bars: 100 µm (h); 20 µm (a, c – g, i); 10 µm (b, j).	Figure 31. Papuanatula (Fijifiliola) sekawa sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Head; i. Paracercus; j. Cercus. Scale bars: 100 µm (h); 20 µm (a, c – g, i); 10 µm (b, j).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
E0640DB98ED65F88851AC3932204375F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462925	https://doi.org/10.3897/zookeys.1259.168521.figure32	Figure 32. Papuanatula (Fijifiliola) sekawa sp. nov., larva. a. Hind leg; b. Femur, outer margin; c. Tibia; d. Tarsus, outer margin; e. Tarsus, claw; f. Abdomen; g. Abdominal terga; h. Tergalius III; i. Paraproct. Scale bars: 100 µm (f); 50 µm (a); 10 µm (b – e, g – i).	Figure 32. Papuanatula (Fijifiliola) sekawa sp. nov., larva. a. Hind leg; b. Femur, outer margin; c. Tibia; d. Tarsus, outer margin; e. Tarsus, claw; f. Abdomen; g. Abdominal terga; h. Tergalius III; i. Paraproct. Scale bars: 100 µm (f); 50 µm (a); 10 µm (b – e, g – i).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
E0640DB98ED65F88851AC3932204375F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462931	https://doi.org/10.3897/zookeys.1259.168521.figure38	Figure 38. Habitats. a. Tributary to Sekawa River, near Nakawaga (location E; clean lowland river); b. Sekawa River near Nakawaga (location E 1; lowland river, disturbed by human activities; many filamentous green algae); c. Seaqaqa River, bridge near Saivou (location F). Characteristics of locations see Table 3.	Figure 38. Habitats. a. Tributary to Sekawa River, near Nakawaga (location E; clean lowland river); b. Sekawa River near Nakawaga (location E 1; lowland river, disturbed by human activities; many filamentous green algae); c. Seaqaqa River, bridge near Saivou (location F). Characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
E0640DB98ED65F88851AC3932204375F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462933	https://doi.org/10.3897/zookeys.1259.168521.figure40	Figure 40. Distribution of described species. a. Papuanatula (Fijifiliola) gattolliati sp. nov. (locations E, F); b. Papuanatula (Fijifiliola) grisea sp. nov. (locations A, B, I); c. Papuanatula (Fijifiliola) sekawa sp. nov. (location E); d. Papuanatula (Fijifiliola) tenebris sp. nov. (location C 1).	Figure 40. Distribution of described species. a. Papuanatula (Fijifiliola) gattolliati sp. nov. (locations E, F); b. Papuanatula (Fijifiliola) grisea sp. nov. (locations A, B, I); c. Papuanatula (Fijifiliola) sekawa sp. nov. (location E); d. Papuanatula (Fijifiliola) tenebris sp. nov. (location C 1).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
95828F6EA5F655A896610F2B1DE454F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462926	https://doi.org/10.3897/zookeys.1259.168521.figure33	Figure 33. Papuanatula (Fijifiliola) tenebris sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	Figure 33. Papuanatula (Fijifiliola) tenebris sp. nov., larva, habitus. a. Dorsal view; b. Lateral view; c. Ventral view. Scale bars: 1 mm.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
95828F6EA5F655A896610F2B1DE454F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462927	https://doi.org/10.3897/zookeys.1259.168521.figure34	Figure 34. Papuanatula (Fijifiliola) tenebris sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Head; i. Paracercus; j. Cercus. Scale bars: 100 µm (h); 50 µm (c, e – g); 20 µm (a, d); 10 µm (b, i, j).	Figure 34. Papuanatula (Fijifiliola) tenebris sp. nov., larva. a, b. Labrum; c. Labium; d. Hypopharynx and superlinguae; e. Left mandible; f. Right mandible; g. Maxilla; h. Head; i. Paracercus; j. Cercus. Scale bars: 100 µm (h); 50 µm (c, e – g); 20 µm (a, d); 10 µm (b, i, j).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
95828F6EA5F655A896610F2B1DE454F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462928	https://doi.org/10.3897/zookeys.1259.168521.figure35	Figure 35. Papuanatula (Fijifiliola) tenebris sp. nov., larva. a. Middle leg; b. Femur, outer margin; c. Femur, inner margin; d. Tibia; e. Tibia, inner margin; f. Tarsus, outer margin; g. Tarsus, claw; h. Abdomen; i. Abdominal terga; j, k. Tergalius IV; l. Paraproct. Scale bars: 100 µm (h); 50 µm (a, j); 10 µm (b – g, i, k, l).	Figure 35. Papuanatula (Fijifiliola) tenebris sp. nov., larva. a. Middle leg; b. Femur, outer margin; c. Femur, inner margin; d. Tibia; e. Tibia, inner margin; f. Tarsus, outer margin; g. Tarsus, claw; h. Abdomen; i. Abdominal terga; j, k. Tergalius IV; l. Paraproct. Scale bars: 100 µm (h); 50 µm (a, j); 10 µm (b – g, i, k, l).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
95828F6EA5F655A896610F2B1DE454F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462929	https://doi.org/10.3897/zookeys.1259.168521.figure36	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	Figure 36. Habitats. a. Overview map; b, c. Ba River near Outback Hotel (locations A, B; lowland river); d. Savuione River, close to beyond the Savuione waterfall, near Abaca village (location C; mountain river); e. Savuione River, downstream of the waterfall, near Abaca village (location C 1; mountain river). For characteristics of locations see Table 3.	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
95828F6EA5F655A896610F2B1DE454F2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1462933	https://doi.org/10.3897/zookeys.1259.168521.figure40	Figure 40. Distribution of described species. a. Papuanatula (Fijifiliola) gattolliati sp. nov. (locations E, F); b. Papuanatula (Fijifiliola) grisea sp. nov. (locations A, B, I); c. Papuanatula (Fijifiliola) sekawa sp. nov. (location E); d. Papuanatula (Fijifiliola) tenebris sp. nov. (location C 1).	Figure 40. Distribution of described species. a. Papuanatula (Fijifiliola) gattolliati sp. nov. (locations E, F); b. Papuanatula (Fijifiliola) grisea sp. nov. (locations A, B, I); c. Papuanatula (Fijifiliola) sekawa sp. nov. (location E); d. Papuanatula (Fijifiliola) tenebris sp. nov. (location C 1).	2025-11-11	Kaltenbach, Thomas		Zenodo	biologists	Kaltenbach, Thomas			
