taxonID	type	description	language	source
113A87FCD64AFFFAFF68FC43C355FB0D.taxon	description	(figure 1)	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64AFFFAFF68FC43C355FB0D.taxon	diagnosis	Diagnosis Dark pigmentation, with some lighter parts; normal and convex eyes; wingless; lateral furrows of pronotum narrower than other Azorean species. Integument wellpigmented with pronotum darker than elytra. Appendages less pigmented (light brown), with tibiae and tarsomeres darker than femora. First antennomere slightly brown and the remaining darker. Epipleura light brown and abdomen brown. Subnitidous with pronotum and head brighter. Microreticulation isodiametric on the head, but transverse on the pronotum and on the elytra.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64AFFFAFF68FC43C355FB0D.taxon	materials_examined	Type series HOLOTYPE: „, Açores, Terceira, Terra-Brava, 18 July – 2 August 2001 (P. A. V. Borges and F. Pereira). PARATYPES: 2 ””, same locality as holotype, 5 – 19 May 1998, 1 – 15 August 1998 (D. Silva).	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64AFFFAFF68FC43C355FB0D.taxon	discussion	Note. In addition one male and four females were collected in the same locality for molecular analysis. These are also used for measures. Additional specimens were recently collected in another locality at Terceira (Caldeira da Serra de Santa Bárbara).	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64AFFFAFF68FC43C355FB0D.taxon	description	Description Length of holotype: 3.85 mm. Length of remaining specimens: 3.36 – 3.76 mm. Medium size, convex and ovalate. Head almost 1.13 X as wide as long, 0.76 X narrower than pronotum; eyes welldeveloped, convex, diameter of eyes 2.4 X larger than temples; temples with microchaetae. Frons highly convex, the frontal furrows deeply curved and rounded, with darker pigmentation. Anterior margin of labrum very concave. Maxillary palpi long and slender. Chaetotaxy: two pairs of supraocular setae; four setae in clypeus (outer ones larger); six setae on labrum, the outer ones being the largest. Antennae 2.05 mm long (others: 1.80 – 1.98 mm), surpassing only the first quarter of elytra; the 3 rd antennomere subequal to the 2 nd; 1 st to 6 th antennomere 2 X longer than wide, 7 th antennomere 1.66 X longer than wide, 8 th to 10 th antennomere 1.3 X longer than wide, 11 th antennomere 2.36 X longer than wide. Pronotum convex, cordiform, highly transverse, 1.29 X as broad as long (others: 1.27 – 1.35 X); side margins curved till hind angle; anterior angles obtuse; hind angles obtuse, well marked and not protruding; basal border emarginated; lateral furrows narrower than in the other Azorean Trechus; basal fovea well marked. Chaetotaxy: the anterior setae is located in anterior quarter and the posterior seta located in the hind angle. Elytra ovate with rounded shoulders, 1.58 X wider than pronotum (others: 1.48 – 1.60 X) and 1.44 X as long as broad (others: 1.35 – 1.48 X); apex rounded, lateral furrows canaliculated; striae all well marked and interstriae convex. Recurrent striae well connected to the 5 th. Chaetotaxy: umbilical series typical of Trechus; preapical seta equidistant of suture and apex; the first discal seta in the 3 rd stria is located in the anterior quarter and the second discal seta approximately in the centre. Aedeagus with apex of median lobe straight and pointed (lateral view) (figure 1 a); apex rounded (dorsal view) (figure 1 c). Left and right parameres with four and five setae each, respectively (figure 1 b).	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64AFFFAFF68FC43C355FB0D.taxon	biology_ecology	Ecological notes All specimens were collected by means of pitfall traps baited with Turquin (a mixture of dark beer with preservatives) or liver, located deep inside very humid laurel forests on Terceira. More recently, several specimens were collected in leaf litter, suggesting that this is a litter species. In both Terra-Brava and Caldeira da Serra de Santa Bárbara, the terrain is basaltic with a system of cracks and deep holes and the forest floor is covered by a dense carpet of mosses and ferns with little light reaching the ground.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64AFFFAFF68FC43C355FB0D.taxon	discussion	Taxonomic notes The aedeagus of this new species clearly resembles those of the cavernicolous troglobitic species Trechus jorgensis (S. Jorge) and T. pereirai n. sp. (Pico).	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64CFFF8FE86FA99C5BCFC2A.taxon	description	(figure 2)	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64CFFF8FE86FA99C5BCFC2A.taxon	diagnosis	Diagnosis Elytra and pronotal disc slightly pigmented; elytra apical angles almost straight. Eyes reduced. Elytra and pronotal disc slightly pigmented; appendages depigmented. Microreticulation isodiametric, though not very clear. In general darker than T. picoensis, T. jorgensis and T. oromii.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64CFFF8FE86FA99C5BCFC2A.taxon	materials_examined	Type series HOLOTYPE: „, Açores, Pico, Gruta do Tio Joaquim, Ribeirinha, 29 June – 16 July 2000 (I. Amorim and F. Pereira). PARATYPE: 1 ”, Açores, Pico, Gruta das Cabras (Terra), Terra do Pão, S. Caetano, 18 – 25 August 2001 (I. Amorim and F. Pereira).	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64CFFF8FE86FA99C5BCFC2A.taxon	discussion	Note. In addition three males and one female were collected in Gruta das Cabras (Terra) for molecular analysis. These are also used for measures.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64CFFF8FE86FA99C5BCFC2A.taxon	description	Description Length of holotype: 3.20 mm. Length of other specimens: 3.02 – 3.19 mm. Small, convex and ovalate. Head normal, ovate, frontal furrows deep and curved; frons convex as in the other Azorean Trechus, but the orbitals flatter in their anterior part; eyes small and reduced, slightly convex, the diameter as long as temples. Temples curved as in T. jorgensis. Labrum arcuated. Maxillary palpi long and slender. Chaetotaxy: as in T. terrabravensis n. sp. Antennae 1.76 mm long (others: 1.68 – 1.79 mm), reaching backwards for the first quartile of elytra; 2 nd antennomere almost as long as 3 rd; 1 st to 7 th antennomere 1.8 X longer than wide, 8 th to 10 th antennomere 1.6 X longer than wide, 11 th antennomere 2.0 X longer than wide. Pronotum convex, transverse, 1.21 X as broad as long (others: 1.24 – 1.29 X); side margins curved with a prebasal small sinuation; anterior angles obtuse and slightly rounded; hind angles obtuse with small protruding corner. Median impression fine but well marked. Base emarginated with basal fovea clearly marked. Chaetotaxy: as in T. terrabravensis. Elytra ovate, convex with shoulders very rounded, 1.68 X wider than pronotum (others: 1.60 – 1.66 X) and 1.37 X as long as wide (others: 1.36 – 1.42 X). Apical angles almost sharp angled as in T. jorgensis; notch not very conspicuous. All striae well marked, with interstriae slightly convex. Chaetotaxy: as in T. terrabravensis, with setae very long and slender. Legs depigmented, similar to T. jorgensis, with protibiae not carinated; male protarsus with first two segments dilated. First metatarsomere 1.4 X as long as onychium. Aedeagus with apex of median lobe more or less straight and pointed (lateral view) (figure 2 a); apex subtruncated (dorsal view) (figure 2 c). Left and right parameres with four setae each (figure 2 b).	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64CFFF8FE86FA99C5BCFC2A.taxon	biology_ecology	Ecological notes The specimens were collected by means of liver-baited pitfall traps deep inside two small lava tubes, one located in the northeasternmost part of Pico island (Gruta do Tio Joaquim) and the other located in the central southern part of the island (Gruta das Cabras, Terra). Known from Pico island only.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64CFFF8FE86FA99C5BCFC2A.taxon	discussion	Taxonomic notes This new species is clearly similar to the troglobitic species T. jorgensis that occurs in the northwards close island of S. Jorge.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64EFFF6FEB8FBBFC527FD64.taxon	description	(figure 3)	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64EFFF6FEB8FBBFC527FD64.taxon	diagnosis	Diagnosis Characterized by the shape of the male aedeagus. Externally very similar to T. picoensis. Integument clearly depigmented, colour reddish testaceous, appendages yellowish. Microreticulation isodiametric in head and pronotum, but transverse in the elytra.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64EFFF6FEB8FBBFC527FD64.taxon	materials_examined	Type series HOLOTYPE: „, Açores, Faial, Gruta do Capelo, Parque do Capelo, 28 June – 18 July 2000 (I. Amorim and F. Pereira). PARATYPE: 1 ”, same locality as holotype, 26 August – 2 September 1999 (I. Amorim and F. Pereira).	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64EFFF6FEB8FBBFC527FD64.taxon	discussion	Note. In addition four males and six females were collected in the same cave in 2001 for molecular analysis. These are also used for measures.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64EFFF6FEB8FBBFC527FD64.taxon	description	Description Length of holotype: 4.82 mm, convex and ovalate, the males being larger than females (average length: 4.64 mm for males and 4.17 mm for females; average width: 1.91 mm for males and 1.70 mm for females). Head small and narrow as in T. picoensis, narrower than prothorax; eyes small, almost flat; temples with microchaetae and 1.10 X broader than eye length (others: 0.86 – 1.14 X). Frons convex, frontal furrows deep and curved; anterior margin of labrum concave; mandibles strong and elongate. Maxillary palpi long and slender. Chaetotaxy: as in T. terrabravensis n. sp. Antennae 3.07 mm (other specimens: 2.55 – 3.20 mm), reaching backwards for the first quartile of elytra; 3 rd antennomere 1.27 X longer than 2 nd, but quite variable (average 1.44 X in males and 1.35 X in females); 1 st to 3 rd antennomere 3.8 X longer than wide, 4 th to 7 th antennomere 3.4 X longer than wide, 8 th to 10 th antennomere 3.0 X longer than wide, 11 th antennomere 3.3 X longer than wide. Pronotum very similar to T. picoensis, convex, transverse, 1.10 X as broad as long (others: 1.15 – 1.21 X); side margins curved in front and markedly narrowed backwards in slightly curved lines up to sharp and conspicuous protruding corners; basal margin emarginated and sinuated. Median impression fine but well marked. Basal fovea clearly marked. Chaetotaxy: as in T. terrabravensis n. sp. Elytra ovate, convex with rounded shoulders, 1.90 X wider than pronotum (others: 1.70 – 1.92 X) and 1.43 X as long as wide (others: 1.42 – 1.55 X). Convex with a depressed anterior field near scutellum. Elytral apex rounded and more depigmented. Chaetotaxy: as in T. terrabravensis n. sp., with setae very long and slender. Legs depigmented, long and slender. Aedeagus apex of median lobe deeply curved upwards in lateral view (figure 3 a). Basal part of median lobe elongated and not so curved as in T. picoensis. In dorsal view the apex is narrower and longer than in T. picoensis (figure 3 c).	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64EFFF6FEB8FBBFC527FD64.taxon	biology_ecology	Ecological notes The specimens were collected by means of pitfall traps baited with cow or pig liver deep inside of a small lava tube located in Parque do Capelo (Faial). Restricted to Faial.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD64EFFF6FEB8FBBFC527FD64.taxon	discussion	Taxonomic notes This new species is clearly similar to the troglobitic species Trechus picoensis that occurs in the nearest island of Pico.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD640FFF4FE98FD02C65EFECD.taxon	description	(figure 4)	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD640FFF4FE98FD02C65EFECD.taxon	diagnosis	Diagnosis Integument depigmented; antennae with the 2 nd antennomere almost as long as 3 rd; eyes small and very flat; apical angles of elytra almost straight. Integument depigmented, testaceous; appendages and apex of elytra more yellowish; dorsal surface dull, less shining than the other Azorean species; microreticulation isodiametric but not very precise and granulated.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD640FFF4FE98FD02C65EFECD.taxon	materials_examined	Studied material One male, Açores, S. Jorge, Algar das Bocas do Fogo, 30 June – 5 August 1999 (I. Amorim and P. A. V. Borges).	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD640FFF4FE98FD02C65EFECD.taxon	description	Description Length of male: 3.20 mm (female measured 3.05 mm); width: 1.39 mm (female with 1.23 mm). Head normal, ovate, frontal furrows deep and curved; frons convex as in previous species, but the orbitals flatter in their anterior part; eyes small, clearly reduced, very flat, the diameter shorter than temples. Labrum less concave than T. terceiranus. Chaetotaxy: as in T. terrabravensis n. sp. Antennae 1.66 mm (1.61 mm in the female), reaching backwards for the first third of elytra; 3 rd antennomere almost as long as 2 nd; 1 st to 7 th antennomere 2.0 X longer than wide; 8 th to 10 th antennomere 1.6 X longer than wide, 11 th antennomere 2.0 X longer than wide. Pronotum convex, transverse, 1.27 X as broad as long (1.28 X in the female). Side margins curved, with a prebasal small sinuation; anterior angles obtuse and rounded; hind angles obtuse with small protruding corners. Base emarginated; lateral fovea not very evident; side marginal furrow narrowly canaliculated; median impression fine. Chaetotaxy: as in T. terrabravensis n. sp. Elytra ovate and convex with shoulders very rounded, as in the female 1.64 X wider than pronotum and 1.45 X (1.36 X in the female) as long as wide. Apical angles forming a right angle, notch not very conspicuous. All striae well marked. Setae long and slender with the typical chaetotaxy of the Trechus spp. Preapical seta closer to the suture than to the apex. Aedeagus with median lobe moderately arched; apex of the median lobe sharpened and slightly upward curved in lateral view (figure 4 a). In dorsal view, apical lamina subtruncated (figure 4 c). Left paramere with four setae and right paramere with five setae (figure 4 b).	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD640FFF4FE98FD02C65EFECD.taxon	biology_ecology	Ecological notes The specimen was collected by means of pitfall traps baited with cow or pig liver deep inside a volcanic pit. Restricted to S. Jorge.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
113A87FCD640FFF4FE98FD02C65EFECD.taxon	discussion	Taxonomic notes Female holotype described in Oromí and Borges (1991). This species is clearly similar to the troglobitic species Trechus pereirai n. sp. that occurs in the nearest island of Pico, and the genitalia also with T. terrabravensis n. sp. from the island of Terceira.	en	Borges, P. A. V., Serrano, A. R. M., Amorim, I. R. (2004): New species of cave-dwelling beetles (Coleoptera: Carabidae: Trechinae) from the Azores. Journal of Natural History 38 (10): 1303-1313, DOI: 10.1080/0022293031000155214, URL: http://dx.doi.org/10.1080/0022293031000155214
