taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
14DB66CA0EDF58FD95099C200E13C085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518371	https://doi.org/10.3897/zse.102.165834.figure3	Figure 3. Crassignatha hekou Wu & Lin, sp. nov. A. Male habitus, dorsal view; B. Ditto, ventral view; C. Ditto, lateral view; D. Female habitus, dorsal view; E. Ditto, ventral view; F. Ditto, lateral view. Scale bars: 0.50 mm (A – F).	Figure 3. Crassignatha hekou Wu & Lin, sp. nov. A. Male habitus, dorsal view; B. Ditto, ventral view; C. Ditto, lateral view; D. Female habitus, dorsal view; E. Ditto, ventral view; F. Ditto, lateral view. Scale bars: 0.50 mm (A – F).	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
14DB66CA0EDF58FD95099C200E13C085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518372	https://doi.org/10.3897/zse.102.165834.figure4	Figure 4. Crassignatha hekou Wu & Lin, sp. nov. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, dorsal view; G. Ditto, ventral view. Scale bars: 0.10 mm (A – G).	Figure 4. Crassignatha hekou Wu & Lin, sp. nov. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, dorsal view; G. Ditto, ventral view. Scale bars: 0.10 mm (A – G).	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
14DB66CA0EDF58FD95099C200E13C085.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518381	https://doi.org/10.3897/zse.102.165834.figure13	Figure 13. Distribution records of Crassignatha species in southern China and Southeast Asia. 1 C. rostriformis; 2 C. bangbie; 3 C. thamphra; 4 C. gucheng; 5 C. baihua; 6 C. quadriventris; 7 C. mengla; 8 C. dongnai; 9 C. liangdu sp. nov.; 10 C. si; 11 C. yamu; 12 C. panlong sp. nov.; 13 C. hekou sp. nov.; 14 C. ertou; 15 C. yinzhi; 16 C. pianma; 17 C. qingxu sp. nov.; 18 C. nasalis; 19 C. shunani; 20 C. nantou; 21 C. shiluensis; 22 C. danaugirangensis; 23 C. haeneli; 24 C. seedam; 25 C. seeliam; 26 C. bicorniventris; 27 C. quanqu; 28 C. gudu; 29 C. changyan; 30 C. xichou. Red, purple, blue, and yellow symbols correspond to branches in Figs 1, 2, whereas green symbols represent species not included in the phylogenetic analyses.	Figure 13. Distribution records of Crassignatha species in southern China and Southeast Asia. 1 C. rostriformis; 2 C. bangbie; 3 C. thamphra; 4 C. gucheng; 5 C. baihua; 6 C. quadriventris; 7 C. mengla; 8 C. dongnai; 9 C. liangdu sp. nov.; 10 C. si; 11 C. yamu; 12 C. panlong sp. nov.; 13 C. hekou sp. nov.; 14 C. ertou; 15 C. yinzhi; 16 C. pianma; 17 C. qingxu sp. nov.; 18 C. nasalis; 19 C. shunani; 20 C. nantou; 21 C. shiluensis; 22 C. danaugirangensis; 23 C. haeneli; 24 C. seedam; 25 C. seeliam; 26 C. bicorniventris; 27 C. quanqu; 28 C. gudu; 29 C. changyan; 30 C. xichou. Red, purple, blue, and yellow symbols correspond to branches in Figs 1, 2, whereas green symbols represent species not included in the phylogenetic analyses.	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
CDECE949A228597EA167D26BDAE9C7EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518373	https://doi.org/10.3897/zse.102.165834.figure5	Figure 5. Crassignatha liangdu Wu & Lin, sp. nov. A. Male habitus, dorsal view; B. Ditto, ventral view; C. Ditto, lateral view; D. Female habitus, dorsal view; E. Ditto, ventral view; F. Ditto, lateral view. Scale bars: 0.50 mm (A – F).	Figure 5. Crassignatha liangdu Wu & Lin, sp. nov. A. Male habitus, dorsal view; B. Ditto, ventral view; C. Ditto, lateral view; D. Female habitus, dorsal view; E. Ditto, ventral view; F. Ditto, lateral view. Scale bars: 0.50 mm (A – F).	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
CDECE949A228597EA167D26BDAE9C7EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518374	https://doi.org/10.3897/zse.102.165834.figure6	Figure 6. Crassignatha liangdu Wu & Lin, sp. nov. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, dorsal view; G. Ditto, ventral view. Scale bars: 0.10 mm (A – G).	Figure 6. Crassignatha liangdu Wu & Lin, sp. nov. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, dorsal view; G. Ditto, ventral view. Scale bars: 0.10 mm (A – G).	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
CDECE949A228597EA167D26BDAE9C7EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518381	https://doi.org/10.3897/zse.102.165834.figure13	Figure 13. Distribution records of Crassignatha species in southern China and Southeast Asia. 1 C. rostriformis; 2 C. bangbie; 3 C. thamphra; 4 C. gucheng; 5 C. baihua; 6 C. quadriventris; 7 C. mengla; 8 C. dongnai; 9 C. liangdu sp. nov.; 10 C. si; 11 C. yamu; 12 C. panlong sp. nov.; 13 C. hekou sp. nov.; 14 C. ertou; 15 C. yinzhi; 16 C. pianma; 17 C. qingxu sp. nov.; 18 C. nasalis; 19 C. shunani; 20 C. nantou; 21 C. shiluensis; 22 C. danaugirangensis; 23 C. haeneli; 24 C. seedam; 25 C. seeliam; 26 C. bicorniventris; 27 C. quanqu; 28 C. gudu; 29 C. changyan; 30 C. xichou. Red, purple, blue, and yellow symbols correspond to branches in Figs 1, 2, whereas green symbols represent species not included in the phylogenetic analyses.	Figure 13. Distribution records of Crassignatha species in southern China and Southeast Asia. 1 C. rostriformis; 2 C. bangbie; 3 C. thamphra; 4 C. gucheng; 5 C. baihua; 6 C. quadriventris; 7 C. mengla; 8 C. dongnai; 9 C. liangdu sp. nov.; 10 C. si; 11 C. yamu; 12 C. panlong sp. nov.; 13 C. hekou sp. nov.; 14 C. ertou; 15 C. yinzhi; 16 C. pianma; 17 C. qingxu sp. nov.; 18 C. nasalis; 19 C. shunani; 20 C. nantou; 21 C. shiluensis; 22 C. danaugirangensis; 23 C. haeneli; 24 C. seedam; 25 C. seeliam; 26 C. bicorniventris; 27 C. quanqu; 28 C. gudu; 29 C. changyan; 30 C. xichou. Red, purple, blue, and yellow symbols correspond to branches in Figs 1, 2, whereas green symbols represent species not included in the phylogenetic analyses.	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
CDECE949A228597EA167D26BDAE9C7EA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518376	https://doi.org/10.3897/zse.102.165834.figure8	Figure 8. Crassignatha panlong Wu & Lin, sp. nov. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, dorsal view; G. Ditto, ventral view. Scale bars: 0.10 mm (A – G).	Figure 8. Crassignatha panlong Wu & Lin, sp. nov. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, dorsal view; G. Ditto, ventral view. Scale bars: 0.10 mm (A – G).	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
387158897D935B5AA5634EC01ADF0DB2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518375	https://doi.org/10.3897/zse.102.165834.figure7	Figure 7. Crassignatha panlong Wu & Lin, sp. nov. A. Male habitus, dorsal view; B. Ditto, ventral view; C. Ditto, lateral view; D. Female habitus, dorsal view; E. Ditto, ventral view; F. Ditto, lateral view. Scale bars: 0.50 mm (A – F).	Figure 7. Crassignatha panlong Wu & Lin, sp. nov. A. Male habitus, dorsal view; B. Ditto, ventral view; C. Ditto, lateral view; D. Female habitus, dorsal view; E. Ditto, ventral view; F. Ditto, lateral view. Scale bars: 0.50 mm (A – F).	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
387158897D935B5AA5634EC01ADF0DB2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518376	https://doi.org/10.3897/zse.102.165834.figure8	Figure 8. Crassignatha panlong Wu & Lin, sp. nov. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, dorsal view; G. Ditto, ventral view. Scale bars: 0.10 mm (A – G).	Figure 8. Crassignatha panlong Wu & Lin, sp. nov. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, dorsal view; G. Ditto, ventral view. Scale bars: 0.10 mm (A – G).	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
387158897D935B5AA5634EC01ADF0DB2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518381	https://doi.org/10.3897/zse.102.165834.figure13	Figure 13. Distribution records of Crassignatha species in southern China and Southeast Asia. 1 C. rostriformis; 2 C. bangbie; 3 C. thamphra; 4 C. gucheng; 5 C. baihua; 6 C. quadriventris; 7 C. mengla; 8 C. dongnai; 9 C. liangdu sp. nov.; 10 C. si; 11 C. yamu; 12 C. panlong sp. nov.; 13 C. hekou sp. nov.; 14 C. ertou; 15 C. yinzhi; 16 C. pianma; 17 C. qingxu sp. nov.; 18 C. nasalis; 19 C. shunani; 20 C. nantou; 21 C. shiluensis; 22 C. danaugirangensis; 23 C. haeneli; 24 C. seedam; 25 C. seeliam; 26 C. bicorniventris; 27 C. quanqu; 28 C. gudu; 29 C. changyan; 30 C. xichou. Red, purple, blue, and yellow symbols correspond to branches in Figs 1, 2, whereas green symbols represent species not included in the phylogenetic analyses.	Figure 13. Distribution records of Crassignatha species in southern China and Southeast Asia. 1 C. rostriformis; 2 C. bangbie; 3 C. thamphra; 4 C. gucheng; 5 C. baihua; 6 C. quadriventris; 7 C. mengla; 8 C. dongnai; 9 C. liangdu sp. nov.; 10 C. si; 11 C. yamu; 12 C. panlong sp. nov.; 13 C. hekou sp. nov.; 14 C. ertou; 15 C. yinzhi; 16 C. pianma; 17 C. qingxu sp. nov.; 18 C. nasalis; 19 C. shunani; 20 C. nantou; 21 C. shiluensis; 22 C. danaugirangensis; 23 C. haeneli; 24 C. seedam; 25 C. seeliam; 26 C. bicorniventris; 27 C. quanqu; 28 C. gudu; 29 C. changyan; 30 C. xichou. Red, purple, blue, and yellow symbols correspond to branches in Figs 1, 2, whereas green symbols represent species not included in the phylogenetic analyses.	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
387158897D935B5AA5634EC01ADF0DB2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518374	https://doi.org/10.3897/zse.102.165834.figure6	Figure 6. Crassignatha liangdu Wu & Lin, sp. nov. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, dorsal view; G. Ditto, ventral view. Scale bars: 0.10 mm (A – G).	Figure 6. Crassignatha liangdu Wu & Lin, sp. nov. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, dorsal view; G. Ditto, ventral view. Scale bars: 0.10 mm (A – G).	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
F645963986955B8E9CC8CB84AA5B337B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518377	https://doi.org/10.3897/zse.102.165834.figure9	Figure 9. Crassignatha qingxu Wu & Lin, sp. nov. A. Male habitus, dorsal view; B. Ditto, ventral view; C. Ditto, lateral view; D. Female habitus, dorsal view; E. Ditto, ventral view; F. Ditto, lateral view. Scale bars: 0.50 mm (A – F).	Figure 9. Crassignatha qingxu Wu & Lin, sp. nov. A. Male habitus, dorsal view; B. Ditto, ventral view; C. Ditto, lateral view; D. Female habitus, dorsal view; E. Ditto, ventral view; F. Ditto, lateral view. Scale bars: 0.50 mm (A – F).	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
F645963986955B8E9CC8CB84AA5B337B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518378	https://doi.org/10.3897/zse.102.165834.figure10	Figure 10. Crassignatha qingxu Wu & Lin, sp. nov. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, ventral view; G. Ditto, dorsal view. Scale bars: 0.10 mm (A – G).	Figure 10. Crassignatha qingxu Wu & Lin, sp. nov. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, ventral view; G. Ditto, dorsal view. Scale bars: 0.10 mm (A – G).	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
F645963986955B8E9CC8CB84AA5B337B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518381	https://doi.org/10.3897/zse.102.165834.figure13	Figure 13. Distribution records of Crassignatha species in southern China and Southeast Asia. 1 C. rostriformis; 2 C. bangbie; 3 C. thamphra; 4 C. gucheng; 5 C. baihua; 6 C. quadriventris; 7 C. mengla; 8 C. dongnai; 9 C. liangdu sp. nov.; 10 C. si; 11 C. yamu; 12 C. panlong sp. nov.; 13 C. hekou sp. nov.; 14 C. ertou; 15 C. yinzhi; 16 C. pianma; 17 C. qingxu sp. nov.; 18 C. nasalis; 19 C. shunani; 20 C. nantou; 21 C. shiluensis; 22 C. danaugirangensis; 23 C. haeneli; 24 C. seedam; 25 C. seeliam; 26 C. bicorniventris; 27 C. quanqu; 28 C. gudu; 29 C. changyan; 30 C. xichou. Red, purple, blue, and yellow symbols correspond to branches in Figs 1, 2, whereas green symbols represent species not included in the phylogenetic analyses.	Figure 13. Distribution records of Crassignatha species in southern China and Southeast Asia. 1 C. rostriformis; 2 C. bangbie; 3 C. thamphra; 4 C. gucheng; 5 C. baihua; 6 C. quadriventris; 7 C. mengla; 8 C. dongnai; 9 C. liangdu sp. nov.; 10 C. si; 11 C. yamu; 12 C. panlong sp. nov.; 13 C. hekou sp. nov.; 14 C. ertou; 15 C. yinzhi; 16 C. pianma; 17 C. qingxu sp. nov.; 18 C. nasalis; 19 C. shunani; 20 C. nantou; 21 C. shiluensis; 22 C. danaugirangensis; 23 C. haeneli; 24 C. seedam; 25 C. seeliam; 26 C. bicorniventris; 27 C. quanqu; 28 C. gudu; 29 C. changyan; 30 C. xichou. Red, purple, blue, and yellow symbols correspond to branches in Figs 1, 2, whereas green symbols represent species not included in the phylogenetic analyses.	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
CEDF2935C9CC50F89824A3C349D71AEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518379	https://doi.org/10.3897/zse.102.165834.figure11	Figure 11. Crassignatha shunani Lin & S. Q. Li, 2020. A. Male habitus, dorsal view; B. Ditto, ventral view; C. Ditto, lateral view; D. Female habitus, dorsal view; E. Ditto, ventral view; F. Ditto, lateral view. Scale bars: 0.50 mm (A – F).	Figure 11. Crassignatha shunani Lin & S. Q. Li, 2020. A. Male habitus, dorsal view; B. Ditto, ventral view; C. Ditto, lateral view; D. Female habitus, dorsal view; E. Ditto, ventral view; F. Ditto, lateral view. Scale bars: 0.50 mm (A – F).	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
CEDF2935C9CC50F89824A3C349D71AEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518380	https://doi.org/10.3897/zse.102.165834.figure12	Figure 12. Crassignatha shunani Lin & S. Q. Li, 2020. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, ventral view; G. Ditto, dorsal view. Scale bars: 0.10 mm (A – G).	Figure 12. Crassignatha shunani Lin & S. Q. Li, 2020. A. Male palp, prolateral view; B. Ditto, ventral view; C. Ditto, retrolateral view; D. Epigyne, ventral view; E. Ditto, lateral view; F. Vulva, ventral view; G. Ditto, dorsal view. Scale bars: 0.10 mm (A – G).	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
CEDF2935C9CC50F89824A3C349D71AEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518381	https://doi.org/10.3897/zse.102.165834.figure13	Figure 13. Distribution records of Crassignatha species in southern China and Southeast Asia. 1 C. rostriformis; 2 C. bangbie; 3 C. thamphra; 4 C. gucheng; 5 C. baihua; 6 C. quadriventris; 7 C. mengla; 8 C. dongnai; 9 C. liangdu sp. nov.; 10 C. si; 11 C. yamu; 12 C. panlong sp. nov.; 13 C. hekou sp. nov.; 14 C. ertou; 15 C. yinzhi; 16 C. pianma; 17 C. qingxu sp. nov.; 18 C. nasalis; 19 C. shunani; 20 C. nantou; 21 C. shiluensis; 22 C. danaugirangensis; 23 C. haeneli; 24 C. seedam; 25 C. seeliam; 26 C. bicorniventris; 27 C. quanqu; 28 C. gudu; 29 C. changyan; 30 C. xichou. Red, purple, blue, and yellow symbols correspond to branches in Figs 1, 2, whereas green symbols represent species not included in the phylogenetic analyses.	Figure 13. Distribution records of Crassignatha species in southern China and Southeast Asia. 1 C. rostriformis; 2 C. bangbie; 3 C. thamphra; 4 C. gucheng; 5 C. baihua; 6 C. quadriventris; 7 C. mengla; 8 C. dongnai; 9 C. liangdu sp. nov.; 10 C. si; 11 C. yamu; 12 C. panlong sp. nov.; 13 C. hekou sp. nov.; 14 C. ertou; 15 C. yinzhi; 16 C. pianma; 17 C. qingxu sp. nov.; 18 C. nasalis; 19 C. shunani; 20 C. nantou; 21 C. shiluensis; 22 C. danaugirangensis; 23 C. haeneli; 24 C. seedam; 25 C. seeliam; 26 C. bicorniventris; 27 C. quanqu; 28 C. gudu; 29 C. changyan; 30 C. xichou. Red, purple, blue, and yellow symbols correspond to branches in Figs 1, 2, whereas green symbols represent species not included in the phylogenetic analyses.	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
CEDF2935C9CC50F89824A3C349D71AEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518934	https://doi.org/10.3897/zse.102.165834.figure1	Figure 1. Tree topology obtained using maximum likelihood analysis in IQ-TREE v 2.0. Numbers at nodes indicate bootstrap values. Twenty-one species representing the genus Crassignatha cluster together to form a monophyletic clade with high support (yellow, red, blue, and purple boxes). Numbering and branch colors correspond to those used in Fig. 13. Species within the gray box represent the outgroup. Red text indicates newly described species.	Figure 1. Tree topology obtained using maximum likelihood analysis in IQ-TREE v 2.0. Numbers at nodes indicate bootstrap values. Twenty-one species representing the genus Crassignatha cluster together to form a monophyletic clade with high support (yellow, red, blue, and purple boxes). Numbering and branch colors correspond to those used in Fig. 13. Species within the gray box represent the outgroup. Red text indicates newly described species.	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
CEDF2935C9CC50F89824A3C349D71AEF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1518935	https://doi.org/10.3897/zse.102.165834.figure2	Figure 2. Tree topology obtained from Bayesian inference analysis in MrBayes v 3.2. 7. Numbers at nodes indicate posterior probabilities. Other settings and annotations are the same as in Fig. 1. The phylogenetic tree constructed using the Bayesian method supports the monophyly of the four newly described species.	Figure 2. Tree topology obtained from Bayesian inference analysis in MrBayes v 3.2. 7. Numbers at nodes indicate posterior probabilities. Other settings and annotations are the same as in Fig. 1. The phylogenetic tree constructed using the Bayesian method supports the monophyly of the four newly described species.	2026-01-23	Wu, Haihong;Lin, Yucheng		Zenodo	biologists	Wu, Haihong;Lin, Yucheng			
