taxonID	type	description	language	source
121B6D3BFF982740FF04D58DFE1A129A.taxon	type_taxon	Type species. Piratula hygrophila (Thorell, 1872) from England (Blackwall 1861), by subsequent designation (Roewer 1955).	en	Sudhin, Puthoor Pattammal, Sankaran, Pradeep M., Sen, Souvik (2025): First record of the genus Piratula Roewer, 1960 from India, with the description of a new species (Araneae: Lycosidae: Zoicinae). Zootaxa 5659 (4): 593-598, DOI: 10.11646/zootaxa.5659.4.9, URL: https://doi.org/10.11646/zootaxa.5659.4.9
121B6D3BFF982740FF04D58DFE1A129A.taxon	discussion	Remarks. We considered both Pirata and Piratula for the inclusion of this new species, however, it best fits the generic characters of the genus Piratula as defined by Omelko et al. (2011): large, longer than wide and C-shaped tegular apophysis, underdeveloped subterminal apophysis, sperm duct with an almost transverse course, and having a prolateral tibial spine on leg I in female.	en	Sudhin, Puthoor Pattammal, Sankaran, Pradeep M., Sen, Souvik (2025): First record of the genus Piratula Roewer, 1960 from India, with the description of a new species (Araneae: Lycosidae: Zoicinae). Zootaxa 5659 (4): 593-598, DOI: 10.11646/zootaxa.5659.4.9, URL: https://doi.org/10.11646/zootaxa.5659.4.9
121B6D3BFF992741FF04D3BBFEDD120B.taxon	description	Figs 1 – 3	en	Sudhin, Puthoor Pattammal, Sankaran, Pradeep M., Sen, Souvik (2025): First record of the genus Piratula Roewer, 1960 from India, with the description of a new species (Araneae: Lycosidae: Zoicinae). Zootaxa 5659 (4): 593-598, DOI: 10.11646/zootaxa.5659.4.9, URL: https://doi.org/10.11646/zootaxa.5659.4.9
121B6D3BFF992741FF04D3BBFEDD120B.taxon	etymology	Etymology. The specific epithet is an adjective and refers to the acute basal arm of the tegular apophysis of the new species (Figs 1 I, 2 A). The Latin term ‘ acuminatus’ means sharp.	en	Sudhin, Puthoor Pattammal, Sankaran, Pradeep M., Sen, Souvik (2025): First record of the genus Piratula Roewer, 1960 from India, with the description of a new species (Araneae: Lycosidae: Zoicinae). Zootaxa 5659 (4): 593-598, DOI: 10.11646/zootaxa.5659.4.9, URL: https://doi.org/10.11646/zootaxa.5659.4.9
121B6D3BFF992741FF04D3BBFEDD120B.taxon	materials_examined	Type material. Holotype ♂ (NZC-ZSI- 9647 / 18) and paratypes 1 ♂, 1 ♀ (NZC-ZSI- 9648 / 18), INDIA: West Bengal: South 24 Parganas district, Kastala (21 ° 52 ’ N 88 ° 08 ’ E; 16 m a. s. l.), 03 November 2019, leg. K. Valarmathi.	en	Sudhin, Puthoor Pattammal, Sankaran, Pradeep M., Sen, Souvik (2025): First record of the genus Piratula Roewer, 1960 from India, with the description of a new species (Araneae: Lycosidae: Zoicinae). Zootaxa 5659 (4): 593-598, DOI: 10.11646/zootaxa.5659.4.9, URL: https://doi.org/10.11646/zootaxa.5659.4.9
121B6D3BFF992741FF04D3BBFEDD120B.taxon	diagnosis	Diagnosis. Males of P. acuminata sp. nov. are most similar to the males of Piratula piratoides (Bösenberg & Strand, 1906) as both share a ridge-like subterminal apophysis, conductor which is absent, and basolaterally located subtegulum. It can be distinguished from the latter species by the apical arm of tegular apophysis with comparatively narrow proximal part (vs. broad in P. piratoides), and tegular apophysis with single tooth (vs. three in P. piratoides; cf. Figs 1 G, I – K, 2 A, C and Omelko et al. 2011: figs 109 – 110, 113). Females of the new species are similar to those of Piratula yaginumai (Tanaka, 1974) as both share a triangular epigynal plate with concave posterior margin, and short and thick copulatory ducts, but can be separated from the latter by ovate spermathecae (vs. elongate in P. yaginumai; cf. Figs 1 M – N, 2 E – F and Omelko et al. 2011: fig. 156 a – b).	en	Sudhin, Puthoor Pattammal, Sankaran, Pradeep M., Sen, Souvik (2025): First record of the genus Piratula Roewer, 1960 from India, with the description of a new species (Araneae: Lycosidae: Zoicinae). Zootaxa 5659 (4): 593-598, DOI: 10.11646/zootaxa.5659.4.9, URL: https://doi.org/10.11646/zootaxa.5659.4.9
121B6D3BFF992741FF04D3BBFEDD120B.taxon	description	Description. Male in ethanol (holotype, Fig. 1 A, C, E). Color overall pale creamy white with brown shades (Fig. 1 A; the yellowish colouration of Fig. 1 C – F, I — L is due to increased light exposure); opisthosoma covered with chalk white spots, dorsum posterolaterally provided with a pair of light brown stripes. Cheliceral pro- and retromargins with three teeth each (Fig. 1 E). Opisthosoma nearly ovate, hirsute (Fig. 1 A). Body length 8.81. Carapace 5.08 long, 3.82 wide. Opisthosoma 3.73 long, 2.28 wide. Eye sizes and interdistances: ALE 0.13, AME 0.17, PLE 0.18, PME 0.28; AME – ALE 0.05, AME – AME 0.11, AME – PME 0.12, PME – PLE 0.16, PME – PME 0.22. Clypeus height at AMEs 0.09, at ALEs 0.15. Chelicerae 1.53 long. Length of palp and legs: palp 4.89 [1.85, 0.86, 0.97, 1.21], I 12.48 [3.67, 1.40, 3.04, 3.13, 1.24], II 11.86 [3.55, 1.25, 2.64, 3.20, 1.22], III 10.81 [3.23, 1.22, 2.25, 3.04, 1.07], IV 13.90 [4.02, 1.48, 3.08, 3.97, 1.35]. Leg formula: 4123. Spination of palp: femur pl 2 do 3 rl 1; legs: femur I pl 1 do 2, II pl 1 do 3 rl 1, III pl 2 do 2 rl 2, IV pl 2 do 3 rl 1; patellae I – II pl 1, III pl 1 do 1 rl 1, IV pl 1 rl 1; tibia I plv 2 rlv 1 v 1, II pl 2 plv 2 rlv 2, III – IV pl 2 plv 3 do 2 rl 2 rlv 3; metatarsus I plv 3 rlv 3, II pl 3 plv 2 rlv 3 v 1, III – IV pl 3 plv 3 rl 3 rlv 3 v 1; tarsi I – IV spineless. Palp (Figs 1 G – L, 2 A – D). Subtegulum small, basolaterally located (Figs 1 G, 2 A). Tegular apophysis with long, curved apical arm having smoothly rounded tip and short, triangular basal arm with acute tip, with broadly triangular basal tooth (Figs 1 G, I arrow 1, 2 A). Terminal apophysis broad at base, with blunt tip, dorsally having a longitudinal groove to accommodate embolus (Figs 1 K – L, 2 C – D). Subterminal apophysis ridge-like, lying very close to terminal apophysis (Figs 1 K arrow 2, 2 C). Conductor absent. Embolus short, apically curved, with acute tip (Figs 1 L arrow 3, 2 D). Female in ethanol (paratype, Fig. 1 B, D, F). Habitus and details like male except for the following: opisthosoma without prominent patches / markings (Fig. 1 B). Body length 10.57. Carapace 4.77 long, 3.52 wide. Opisthosoma 5.80 long, 4.44 wide. Eye sizes and interdistances: ALE 0.12, AME 0.20, PLE 0.22, PME 0.34; AME – ALE 0.09, AME – AME 0.11, AME – PME 0.11, PME – PLE 0.22, PME – PME 0.23. Clypeus height at AMEs 0.14, at ALEs 0.17. Chelicerae 2.12 long. Length of palp and legs: palp 4.53 [1.66, 0.65, 1.09, 1.13], I 10.74 [3.24, 1.39, 2.59, 2.45, 1.07], II 10.05 [3.10, 1.50, 2.17, 2.23, 1.05], III 9.79 [2.96, 1.09, 2.12, 2.57, 1.05], IV 13.14 [3.84, 1.39, 3.06, 3.61, 1.24]. Spination of palp: femur pl 1 do 3; legs: femur III pl 2 do 3 rl 2; patella IV pl 1 do 1 rl 1; tibia I plv 2 rlv 1, II pl 2 rlv 1; metatarsus II pl 2 plv 3 rlv 3 v 1. Genitalia (Figs 1 M – N, 2 E – F). Epigynal plate broadly triangular, with concave posterior margin (Figs 1 M, 2 E). Copulatory openings indistinct. Copulatory ducts short, thick, sclerotised, converging (Figs 1 N, 2 F). Spermathecae small, ovate, widely separated, slightly converging (Figs 1 N, 2 F). Fertilization ducts narrow, curved medially, diverging (Figs 1 N, 2 F). Variation. Male (n = 2): body length 6.93 – 8.81. The carapace length of the new species described here is nearly five (female) to six (male) millimetres, which is higher than in other Piratula species (<3 mm) examined by Omelko et al. (2011).	en	Sudhin, Puthoor Pattammal, Sankaran, Pradeep M., Sen, Souvik (2025): First record of the genus Piratula Roewer, 1960 from India, with the description of a new species (Araneae: Lycosidae: Zoicinae). Zootaxa 5659 (4): 593-598, DOI: 10.11646/zootaxa.5659.4.9, URL: https://doi.org/10.11646/zootaxa.5659.4.9
121B6D3BFF992741FF04D3BBFEDD120B.taxon	distribution	Distribution. Known only from the type locality in West Bengal, India (Fig. 3). This is the first record of the genus from India.	en	Sudhin, Puthoor Pattammal, Sankaran, Pradeep M., Sen, Souvik (2025): First record of the genus Piratula Roewer, 1960 from India, with the description of a new species (Araneae: Lycosidae: Zoicinae). Zootaxa 5659 (4): 593-598, DOI: 10.11646/zootaxa.5659.4.9, URL: https://doi.org/10.11646/zootaxa.5659.4.9
