identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
50AE4B6884D65937A4845E1B2DDE2230.text	50AE4B6884D65937A4845E1B2DDE2230.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrobiomorpha bachmanni Urcola & Rodriguez & Michat & Torres 2026	<div><p>Hydrobiomorpha bachmanni sp. nov.</p><p>Figs 1 A, 2 A, 3 A, 4 A, 4 B, 5 A – D, 6 A – L, 7 A, 7 G, 8</p><p>Type locality.</p><p>Argentina: Corrientes Province, Parque Nacional Iberá, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.195835&amp;materialsCitation.latitude=-28.54611" title="Search Plazi for locations around (long -57.195835/lat -28.54611)">Portal Laguna Iberá</a>, 28°32'46"S, 57°11'45"W, elevation 67 m a. s. l.</p><p>Type material.</p><p>Holotype • ♂, deposited in the entomological collection of the Museo Argentino de Ciencias Naturales “ Bernardino Rivadavia ” (MACN), Buenos Aires, Argentina, labelled: “ ARGENTINA. Corrientes / Province. PN Iberá. Portal / Laguna Iberá 8.xi.2019 / JI Urcola and G Rodriguez leg. ” [typed, white label], “ HOLOTYPE / Hydrobiomorpha bachmanni sp. nov. / Urcola, Rodriguez / Michat, Torres, 2026 " [typed, red label], “ MACN_En / 46706 " [typed, white label] . Paratypes: • 10 ♂♂ and 10 ♀♀ from type locality (MACN_En 46707 to 46726). All paratypes with the following labels: “ ARGENTINA. Corrientes / Province. PN Iberá. Portal / Laguna Iberá 14.xi.2019 / JI Urcola and G Rodriguez leg. ” [typed, white label], “ PARATYPE / Hydrobiomorpha bachmanni sp. nov. / Urcola, Rodriguez / Michat, Torres, 2026 " [typed, yellow label] .</p><p>Other material examined.</p><p>Paraguay: • Presidente Hayes Department, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.3225&amp;materialsCitation.latitude=-24.385277" title="Search Plazi for locations around (long -57.3225/lat -24.385277)">Estancia Campo Grande</a>, 24°23'07"S, 57°19'21"W, 5–8.xii.2002, OR Di Iorio leg., light trap, 6 ♂♂ and 5 ♀♀ (LEBA) .</p><p>Diagnosis.</p><p>L: 15.5–19.0 mm (males), 16.5–20.5 mm (females). W: 7.5–8.5 mm (males), 8.0– 9.4 mm (females). Body shape (Figs 1 A, 2 A, 3 A): elliptical, sides subparallel, rather convex; elytral margins scarcely expanded outward on the anterior half. Prosternal keel (Fig. 3 A) slightly concave; posterior spine acute, moderately long, reaching approximately mid-length of procoxae. Metaventral keel (Fig. 3 A) almost straight in lateral view, with small anterior notch and midline depression; posterior portion short, weakly arched ventrally, apex obtusely pointed, not reaching first abdominal ventrite. Male: mesotibial setigerous furrow present, extending for ~ 1 / 3 of tibial length (Table 1; Fig. 7 A, G); protarsal claws robust, anterior claw with a pointed proximal tooth; meso- and metatarsal claws with a conspicuous blunter proximal tooth; fifth abdominal ventrite with glabrous area occupying ~ 2 / 5 of its total length (Fig. 4 A); genitalia: median lobe broad, apex deeply bifurcate into two long, narrow, subparallel projections; accessory appendages of median lobe broad, sclerotized, abruptly narrowed distally and terminating in short, apical hooks directed outward; parameres moderately broad in lateral view, apex acute and strongly sclerotized, curved toward midline (Fig. 5 A – D). Female: fifth abdominal ventrite with glabrous area ~ 2 / 5 of its total length, though narrower than in males (Fig. 4 B).</p><p>Differential diagnosis.</p><p>Hydrobiomorpha bachmanni sp. nov. can be readily distinguished from the other Argentine Hydrobiomorpha species by the unique combination of a glabrous area present in both sexes and occupying ~ 2 / 5 of the length of the last abdominal ventrite (Fig. 4 A, B), together with a male mesotibial setigerous furrow extending along about the distal 1 / 3 of the tibia (Fig. 7 A, G). In H. iguazu, the glabrous area of the last abdominal ventrite is much larger in both sexes (Fig. 4 E, F) and the male mesotibial setigerous furrow extending along about the distal 1 / 4 of the tibia; in H. spinosa, the male mesotibial setigerous furrow extending along about the distal 1 / 2 of the tibia (Fig. 7 F, J) and the glabrous area of the last abdominal ventrite is minute in male and absent in female (Fig. 4 M, N); in H. irina irina, the glabrous area of last abdominal ventrite is much smaller in both sexes (Fig. 4 G, H) and males lack the mesotibial furrow. The species also differs from H. corumbaensis and H. longa, both of which lack a glabrous area on the last abdominal ventrite in females (Fig. 4 D, J) and a mesotibial furrow in males (Fig. 7 B, D, H). The male genitalia of H. bachmanni (Fig. 5 A – D) is readily distinguished from those of all other Argentine species by the median lobe with apex deeply bifurcate into two long, narrow, subparallel projections, and by the parameres with acute, strongly sclerotized apices and broad accessory appendages ending in short outwardly directed hooks. The most similar male genitalia are found in H. utiariti (Fig. 5 AC – AF), but that species differs in having the median lobe apically truncate, with two shorter, slightly divergent projections, and parameres that are more strongly curved and denticulate subdistally.</p><p>Description of holotype.</p><p>Size. L: 16.8 mm; W: 8.0 mm.</p><p>Habitus. Body elliptical, sides subparallel, rather convex.</p><p>Color. Integument very dark brown to black; venter slightly paler. Legs dark brown to black, with tarsi paler. Antennae and palpi light reddish.</p><p>Head (see Fig. 8 A, D – F, paratype). Ground punctation double, with one type of punctures very fine, regular, and dense, the other fine, somewhat sparser and irregular; both well impressed, over very fine, regular microreticulation with a hexagonal pattern (Fig. 8 D – F). Clypeus with anterior margin broadly emarginate, exposing a narrow membranous area; surface shining, without evident microreticulation. Four groups of coarse, deep punctures bearing trichobothria (systematic punctures, sensu Hansen 1991) on each side of clypeal midline and along inner margin of eyes (Fig. 8 D, E). Labrum strongly transverse, anterolaterally rounded, with broad and shallow anteromedian emargination; surface shiny, without microreticulation, with fine double punctation. Two distinct pits, each bearing a tuft of trichobothria, present on either side of midline. Mentum with coarse, dense punctation. Submentum with sparse punctures restricted to lateral margins; central area smooth and shiny.</p><p>Pronotum (see Fig. 8 B, G – I, paratype). Transverse, arched, broadest at posterior margin, gradually narrowing toward anterior angles. Anterior margin arched; posterior margin very weakly bisinuate medially. Lateral margins narrowly reflexed. Anterior and posterior angles obtusely rounded. Surface shining, with very weakly impressed microreticulation of hexagonal pattern (Fig. 8 I). Punctation double, less impressed than on head (Fig. 8 G). Additionally, shallow coarse punctures bearing trichobothria present, on anterior angles, on lateral portions of disc (Fig. 8 H), and along anterior third on each side of midline (Fig. 8 G).</p><p>Elytra (see Fig. 8 C, J, K, paratype). Elongate-oval, broadest near mid-length. Subparallel in anterior 2 / 3, then uniformly rounded toward apex; weakly emarginate at suture. Anterior and lateral margins weakly reflexed, forming a narrow lateral bead (broadest at humerus and in anterior two-thirds, gradually narrowing toward apex), followed mesally by irregular field of moderate, rather dense punctures, extending from humerus to apex. Surface shining, with fine, shallow, isodiametric microreticulation of hexagonal pattern (Fig. 8 J, K). Ground punctation composed of fine to very fine punctures, variable in size and density, not forming a clearly double punctation. Systematic punctures arranged in six rows, outer row intermingled with lateral puncture field.</p><p>Venter (see Figs 2 A, 4 A). Prosternum with glabrous, slightly concave keel; posterior spine acute, moderately long, extending to mid-length of procoxae. Meso-metaventral keel with profile almost straight in lateral view, bearing small notch on anterior margin and depression along midline of metaventrite, area in which it appears slightly widened. Metaventral keel with posterior portion short, weakly arched ventrally, apex obtusely pointed, not reaching first abdominal ventrite. Fifth abdominal ventrite with glabrous area subtriangular, very broad, ~ 2 / 5 of its total length (Fig. 4 A).</p><p>Legs (see Figs 2 A, 6 A – F, paratype, 7 A, G, paratype). Mesofemora with dense punctation, more impressed toward dorsal (anterior) margin; metafemora with very sparse and shallow punctation. Mesotibiae with setigerous furrow extending approximately along distal 1 / 3, gradually widening toward distal end and provided with dense brush of reddish setae (Fig. 7 A, G). Protarsal claws robust, anterior (inner) claw bearing robust proximal tooth with pointed apex; meso- and metatarsal claws with conspicuous proximal tooth, though blunter (Fig. 6 A – F).</p><p>Genitalia (see Fig. 5 A – D, paratype). Median lobe broad, apex deeply bifurcate into two long, narrow, subparallel projections. Dorsal sclerite markedly widened on distal third, gradually tapering proximally. Ventral membranous portion very narrow, distally bounded by more sclerotized bridge connecting both distal ends. Ventral sclerite long and narrow, clearly visible. Accessory appendages of median lobe broad, sclerotized, abruptly narrowed distally and terminating in short, apical hooks directed outward. Parameres moderately broad in lateral view, apex acute and strongly sclerotized, curved toward midline.</p><p>Female. Similar to male but slightly larger; differing in absence of mesotibial setigerous furrow. Glabrous area of fifth abdominal ventrite same length as in male but notably narrower (Fig. 4 B). Claws falciform, relatively slender, with short distal portion; less robust and less strongly angulate than in male (Fig. 6 G – L).</p><p>Etymology.</p><p>The species is named after Dr. Axel O. Bachmann (1927–2017), in recognition of his contributions to the taxonomy of South American Hydrophilidae and his lasting impact on the study of Hydrobiomorpha, including the description and revision of several Argentine species, as well as his role in building and curating reference collections that continue to support systematic research in the region.</p><p>Distribution.</p><p>Paraguay, Argentina: Corrientes Province.</p><p>Biology.</p><p>Unknown. This species was collected using light traps.</p></div>	https://treatment.plazi.org/id/50AE4B6884D65937A4845E1B2DDE2230	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Urcola, Matías R.;Rodriguez, Georgina;Michat, Mariano C.;Torres, Patricia L. M.	Urcola, Matías R., Rodriguez, Georgina, Michat, Mariano C., Torres, Patricia L. M. (2026): Revision of the genus Hydrobiomorpha Blackburn, 1888 (Coleoptera, Hydrophilidae) from Argentina, with the description of a new species and an updated identification key. ZooKeys 1289: 109-139, DOI: 10.3897/zookeys.1289.202453
E6B43B050A8B59D5AADB92B7BE4C0892.text	E6B43B050A8B59D5AADB92B7BE4C0892.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrobiomorpha Blackburn 1888	<div><p>Genus Hydrobiomorpha Blackburn, 1888</p><p>Hydrobiomorpha Blackburn, 1888: 814–815. (Type species: Hydrobiomorpha bovilli Blackburn, 1889, by subsequent designation of Knisch (1924: 234).</p><p>Neohydrophilus d’Orchymont, 1911: 59 . (Type species: Neohydrophilus deplanatus, d’Orchymont, 1919, by original designation). Originally described as a subgenus of Hydrophilus; raised to generic rank by d’Orchymont 1919: 160–161; synonymized with Hydrobiomorpha by Mouchamps 1959: 296.</p><p>Taxonomic history.</p><p>Blackburn (1888) established the genus Hydrobiomorpha for two Australian species, H. bovilli and H. tepperi, but no type species was designated. Later, Knisch (1924) designated H. bovilli Blackburn as the type species. Hydrobiomorpha bovilli is a poorly known species and its type material has not been examined by recent authors. Bachmann (1988) examined specimens of H. tepperi, considered closely similar to H. bovilli by Blackburn (1888) and Mouchamps (1959), and concluded that the placement of H. bovilli in Hydrobiomorpha is justified. d’Orchymont (1911) established the subgenus Hydrophilus (Neohydrophilus) for several species, without designating a type species. In 1919, he raised Neohydrophilus to generic rank and designated N. deplanatus as the type species. Mouchamps (1959) revised the group, synonymized Neohydrophilus with Hydrobiomorpha, and added several new species, four from the Americas. Bachmann (1988) revised the American members of the group and added 19 new species. The genus currently comprises 61 species, 56 of which are extant and have a pantropical distribution (Hansen 1999; Short and Hebauer 2006; Short 2010; Short and Fikáček 2011; Newton 2022).</p><p>Diagnosis.</p><p>Medium-sized to large beetles, length 13.7–28.0 mm (most species 17.0–22.0 mm); body moderately to strongly convex in lateral view; color uniformly black, or more frequently dark brown, without metallic sheen or paler areas; legs slightly paler, rarely yellowish brown (only some H. rudesculpta); clypeus excavate anteriorly, separated from labrum by membranous stripe; labrum with two coarse setigerous punctures; genae near gula partly glabrous; several antennomeres (including club) moderately expanded, divided, with long setae; prosternum, forming a median keel, ventral border generally straight and subparallel to body axis, sometimes rounded, oblique or inflexed, and most frequently with posterior sharp point, rarely very short; meso-metasternal keel straight or slightly concave in lateral view, with small anterior notch; posterior portion short, rarely surpassing posterior margin of first abdominal ventrite, ending bluntly and laterally compressed, sometimes subconical and sharply pointed, or slightly hooked; furcasternum pubescent; mesotibiae of males sometimes with setigerous furrow along distal half of ventral (inner) face, of variable length, covered with dense brush of golden to reddish setae; claws falciform, with proximal denticle, claws of same leg frequently differing slightly in size or shape; male claws strongly angulate; elytra without sutural stria; marginal series of setigerous punctures very regular, sometimes irregular, rarely indistinct; abdominal ventrites totally covered by hydrophobic pubescence, except for fifth ventrite, which shows semicircular, subrectangular or subtriangular posterior glabrous area, typically larger in males, rarely minute, or absent in females; male genitalia complex, median lobe with dorsal sclerite large, variously shaped, laterally compressed, and frequently with raised dorsal lamina; gonopore located proximally or at mid-length, with conspicuous ventral sclerite (“ ventral strut ” of Leech in Bachmann 1988); accessory appendages of median lobe variable in shape, ranging from broad to elongate and from straight to curved or sigmoid, with apices acute, truncate, rounded, or bearing short outwardly directed hooks; parameres bent, with accessory appendages robust, sclerotized, rarely at right angle, or largely bifid.</p><p>Distribution.</p><p>Pantropical, mainly Neotropical. Contartese and Bachmann (1987) presented maps with the distribution of the species of Argentina.</p><p>Biology.</p><p>Adults and larvae inhabit vegetated permanent ponds but are infrequently collected; to date, no records from rain pools are known. Adults are mostly scavengers or periphyton eaters, and larvae are predaceous (Archangelsky et al. 2004). Adults are good swimmers, occasionally flying at night and attracted to artificial lights.</p></div>	https://treatment.plazi.org/id/E6B43B050A8B59D5AADB92B7BE4C0892	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Urcola, Matías R.;Rodriguez, Georgina;Michat, Mariano C.;Torres, Patricia L. M.	Urcola, Matías R., Rodriguez, Georgina, Michat, Mariano C., Torres, Patricia L. M. (2026): Revision of the genus Hydrobiomorpha Blackburn, 1888 (Coleoptera, Hydrophilidae) from Argentina, with the description of a new species and an updated identification key. ZooKeys 1289: 109-139, DOI: 10.3897/zookeys.1289.202453
1B2260200A1A585D9E649CFE39BDF1F9.text	1B2260200A1A585D9E649CFE39BDF1F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrobiomorpha corumbaensis Mouchamps 1959	<div><p>Hydrobiomorpha corumbaensis Mouchamps, 1959</p><p>Figs 1 B, 2 B, 3 B, 4 C, 4 D, 5 E – H, 6 M, 7 B</p><p>Hydrobiomorpha [(Hydrobiomorpha)] longa subsp. corumbaensis Mouchamps, 1959: 331 (original description).</p><p>Hydrobiomorpha corumbaensis: Contartese and Bachmann 1987: fig. 4 (distribution only).</p><p>Hydrobiomorpha corumbaensis: Bachmann 1988: 10, 21–22, fig. 8 A – K (specific rank; redescription; key to species of the Americas).</p><p>Type locality.</p><p>Brazil: Mato Grosso, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.65278&amp;materialsCitation.latitude=-19.008888" title="Search Plazi for locations around (long -57.65278/lat -19.008888)">Corumbá</a> [19°00'32"S, 57°39'10"W].</p><p>Material examined.</p><p>Argentina: • Corrientes Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.833332&amp;materialsCitation.latitude=-27.816668" title="Search Plazi for locations around (long -56.833332/lat -27.816668)">Cambyretá</a>, 27°49'00"S, 56°50'00"W, 16.xi.2018, JI Urcola, G Rodriguez and SA Mazzucconi leg., light trap, 2 ♂♂ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.295555&amp;materialsCitation.latitude=-29.357224" title="Search Plazi for locations around (long -59.295555/lat -29.357224)">Manantiales</a> [29°21'26"S, 59°17'44"W], 9.x.1962, 1 ♂ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.5125&amp;materialsCitation.latitude=-27.370277" title="Search Plazi for locations around (long -58.5125/lat -27.370277)">San Cosme</a> [27°22'13"S, 58°30'45"W], 1937, J Wurth leg., 1 ♀ (MACN -En) and 1 ♂ (LEBA) . • Entre Ríos Province: El Palmar National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.317776&amp;materialsCitation.latitude=-31.865278" title="Search Plazi for locations around (long -58.317776/lat -31.865278)">small lagoon near Capilla Stream</a>, 31°51'55"S, 58°19'04"W, 28.ii.2004, PLM Torres leg., 1 ♂ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.644447&amp;materialsCitation.latitude=-30.740557" title="Search Plazi for locations around (long -59.644447/lat -30.740557)">La Paz</a> [30°44'26"S, 59°38'40"W], xii.1972, 1 ♂ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.166668&amp;materialsCitation.latitude=-32.616665" title="Search Plazi for locations around (long -60.166668/lat -32.616665)">Victoria</a> [32°37'00"S, 60°10'00"W], 8–12.ii.1982, AO Bachmann leg., light trap, 1 ♂ (MACN -En) .</p><p>Diagnosis.</p><p>L: 15.5–17.5 mm (males), 15.5–18.6 mm (females). W: 6.6–7.6 mm (males), 6.6–8.1 mm (females). Body shape (Figs 1 B, 2 B, 3 B): navicular, tapered posteriorly and less so anteriorly, slightly convex; elytral margins not reflexed. Prosternal keel (Fig. 3 B): ventral margin nearly straight; posterior spine acute, moderately long. Metaventral keel (Fig. 3 B): posteriorly blunt and ventrally directed, not reaching first abdominal ventrite. Male: mesotibial setigerous furrow absent (Table 1; Fig. 7 B); fifth abdominal ventrite with a short, poorly delimited glabrous area ~ 1 / 4 (in some cases ≤ 1 / 3) of its total length (Fig. 4 C); protarsal claws dissimilar, with rounded proximal internal process; mesotarsal claws robust, with rounded proximal tooth; metatarsal claws very dissimilar, posterior claw with blunt, rounded preapical expansion (Fig. 6 M); genitalia: median lobe ending in two very short projections; accessory appendages of median lobe broad in proximal half, becoming narrower distally and diverging outward, with rounded distal ends; parameres broad, abruptly narrowed distally (Fig. 5 E – H). Female: glabrous area of fifth abdominal ventrite absent (Fig. 4 D).</p><p>Distribution.</p><p>Brazil (Mouchamps 1959; Bachmann 1988), Paraguay (Bachmann 1988), Argentina (provinces): Corrientes (Bachmann 1988; Gómez Lutz et al. 2012; this study), Entre Ríos (Torres et al. 2007; this study).</p><p>Biology.</p><p>This species inhabits small permanent ponds (2,500–5,000 m 2 in surface area) fully exposed to sunlight, with vegetated margins and centers, semi-transparent water, muddy substrate, and depth of &lt;2 m (Torres et al. 2007; Gómez Lutz et al. 2012).</p><p>Remarks.</p><p>It can be distinguished from other similar species by the unusual posterior metatarsal claw of males, which bears a blunt, rounded preapical expansion (Fig. 6 M); in H. longa this expansion is angulate, rather than rounded (Fig. 6 N). Male genitalia of H. corumbaensis (Fig. 5 E – H) are similar to those of H. longa (Fig. 5 Q – T) but differ in the median lobe, which ends in two short, blunt projections rather than long and sharp as in H. longa .</p></div>	https://treatment.plazi.org/id/1B2260200A1A585D9E649CFE39BDF1F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Urcola, Matías R.;Rodriguez, Georgina;Michat, Mariano C.;Torres, Patricia L. M.	Urcola, Matías R., Rodriguez, Georgina, Michat, Mariano C., Torres, Patricia L. M. (2026): Revision of the genus Hydrobiomorpha Blackburn, 1888 (Coleoptera, Hydrophilidae) from Argentina, with the description of a new species and an updated identification key. ZooKeys 1289: 109-139, DOI: 10.3897/zookeys.1289.202453
67E4541AC6BA5F2FBB444EB104463DE8.text	67E4541AC6BA5F2FBB444EB104463DE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrobiomorpha iguazu Bachmann 1988	<div><p>Hydrobiomorpha iguazu Bachmann, 1988</p><p>Figs 1 C, 2 C, 3 C, 4 E, 4 F, 5 I – L</p><p>Hydrobiomorpha iguazu: Contartese and Bachmann 1987: fig. 3 (nomen nudum, distribution only).</p><p>Hydrobiomorpha iguazu Bachmann, 1988: 9, 15–16, fig. 31 A – H (original description; key to species of the Americas).</p><p>Hydrobiomorpha iguazu: Bachmann et al. 2002: 141 (female description).</p><p>Type locality.</p><p>Argentina: Misiones Province, Parque Nacional Iguazú [25°31'05"S, 54°08'00"W].</p><p>Material examined.</p><p>Argentina: • Misiones Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-54.133335&amp;materialsCitation.latitude=-25.518055" title="Search Plazi for locations around (long -54.133335/lat -25.518055)">Iguazú National Park</a> [25°31'05"S, 54°08'00"W], x.1980, 1 ♂ (holotype, MACN -En 2213) ; • same locality iv.2001, PLM Torres and MC Michat leg., 1 ♀ (MACN -En) .</p><p>Diagnosis.</p><p>L: 22.6 mm (male holotype), 21.0 mm (female). W: 10.6 mm (both sexes). Body shape (Figs 1 C, 2 C, 3 C): subelliptical, slightly convex; elytral margin moderately reflexed on the posterior 2 / 3. Prosternal keel (Fig. 3 C): moderately long, ventral margin nearly straight; posterior spine very short. Metaventral keel (Fig. 3 C): slightly broadened, ending posteriorly in a short, laterally compressed, blunt tip, reaching approximately mid-length of first abdominal ventrite. Male: mesotibial setigerous furrow present, short, extending for ~ 1 / 4 of tibial length, with a short reddish setal brush (Table 1); fifth abdominal ventrite with a very large, subquadrate glabrous area, ~ 3 / 4 of its total length (Fig. 4 E); protarsal claws unknown, not preserved in holotype (only known male); mesotarsal claws robust, with conspicuous proximal tooth; metatarsal claws robust, with conspicuous proximal tooth; genitalia: strongly sclerotized, median lobe subparallel, abruptly narrowed distally, apex bifurcate into two short sharp projections; accessory appendages of median lobe elongate, strongly sclerotized, with acute distal ends; parameres shorter than median lobe, broad in lateral view, narrowed distally and ending in projections, slightly curved towards midline (Fig. 5 I – L). Female: fifth abdominal ventrite with a very large subtriangular glabrous area, ~ 2 / 3 of its total length (Fig. 4 F).</p><p>Distribution.</p><p>Argentina: Misiones Province (Bachmann 1988; Bachmann et al. 2002).</p><p>Biology.</p><p>The female was collected from a shallow pond of turbid water (~ 20 cm deep), near the entrance to the Macuco Trail in Iguazú National Park. No habitat data are available for the holotype.</p><p>Remarks.</p><p>The large size, 21.0– 22.6 mm and dark dorsal coloration resemble those of H. utinga Bachmann, 1988, H. mucajai Bachmann, 1988, H. solitaria Bachmann, 1988, and H. praesumptapolita Bachmann, 1988 . The median lobe of the male genitalia resembles that of H. praesumptapolita (Bachmann, 1988) . The large bare area on the last abdominal ventrite, and the short setigerous furrow, of 1 / 4 the length of the mesotibiae of males, are distinctive; the above mentioned species have the furrow longer, half as long as the tibiae, and in H. mucajai it is absent.</p></div>	https://treatment.plazi.org/id/67E4541AC6BA5F2FBB444EB104463DE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Urcola, Matías R.;Rodriguez, Georgina;Michat, Mariano C.;Torres, Patricia L. M.	Urcola, Matías R., Rodriguez, Georgina, Michat, Mariano C., Torres, Patricia L. M. (2026): Revision of the genus Hydrobiomorpha Blackburn, 1888 (Coleoptera, Hydrophilidae) from Argentina, with the description of a new species and an updated identification key. ZooKeys 1289: 109-139, DOI: 10.3897/zookeys.1289.202453
141480FD9D835135990A4D0FAA1A3EA4.text	141480FD9D835135990A4D0FAA1A3EA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrobiomorpha irina subsp. irina (Brulle 1837)	<div><p>Hydrobiomorpha irina irina (Brullé, 1837)</p><p>Figs 1 D, 2 D, 3 D, 4 G – H, 5 M – P, 7 C</p><p>Hydrophilus irinus Brullé, 1837: 55 (original description, see Bousquet (2016)).</p><p>Neohydrophilus irinus: Knisch 1924: 235.</p><p>Neohydrophilus medius: d'Orchymont 1939 b: 263 –264 (misidentification).</p><p>Hydrobiomorpha [(Hydrobiomorpha)] irina: Mouchamps 1959: 322, 326–327.</p><p>Hydrobiomorpha (Hydrobiomorpha) irina: Bachmann 1963: 30 –31, fig. 1 (key to Argentine species).</p><p>Hydrobiomorpha irina irina: Contartese and Bachmann 1987: fig. 1 (distribution only).</p><p>Hydrobiomorpha irina irina: Bachmann 1988: 8, 26–28, fig. 12 A – I (subspecific rank; redescription; key to species of the Americas).</p><p>Type locality.</p><p>[Argentina]: Rivière du Parana, “ entre son embouchure et Corrientes ”.</p><p>Material examined.</p><p>Argentina: • Buenos Aires Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.333332&amp;materialsCitation.latitude=-35.383335" title="Search Plazi for locations around (long -57.333332/lat -35.383335)">Verónica</a> [35°23'00"S, 57°20'00"W], x.1937, JB Daguerre leg., 1 ♂ (MACN -En) . • Chaco Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.616665&amp;materialsCitation.latitude=-25.95" title="Search Plazi for locations around (long -60.616665/lat -25.95)">Juan José Castelli</a> [25°57'00"S, 60°37'00"W], 30.iii.1978, GJ Williner leg., 1 ♀ (MACN -En) . • Corrientes Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.833332&amp;materialsCitation.latitude=-27.816668" title="Search Plazi for locations around (long -56.833332/lat -27.816668)">Cambyretá</a>, 27°49'00"S, 56°50'00"W, 16.xi.2018, JI Urcola, G Rodriguez and SA Mazzucconi leg., light trap, 2 ♂♂ and 1 ♀ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.195835&amp;materialsCitation.latitude=-28.54611" title="Search Plazi for locations around (long -57.195835/lat -28.54611)">Iberá Lagoon</a>, 28°32'46"S, 57°11'45"W, 8.xi.2019, JI Urcola and G Rodriguez leg., light trap, 1 ♂ (LEBA) ; • Santo Tomé [28°33'00"S, 56°03'00"W] [no date], G Pellerano leg., 1 ♂ and 1 ♀ (MACN -En) . • Entre Ríos Province: El Palmar National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.317776&amp;materialsCitation.latitude=-31.865278" title="Search Plazi for locations around (long -58.317776/lat -31.865278)">small lagoon near the mouth of Capilla Stream</a>, 31°51'55"S, 58°19'04"W, 28.ii.2004, PLM Torres leg., 2 ♂♂ and 1 ♀ (LEBA) . • Santa Fe Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.10472&amp;materialsCitation.latitude=-30.018055" title="Search Plazi for locations around (long -60.10472/lat -30.018055)">El Cristal Natural Reserve</a>, 30°01'05"S, 60°06'17"W, xii.2010, MC Michat leg., 1 ♀ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.639446&amp;materialsCitation.latitude=-32.9575" title="Search Plazi for locations around (long -60.639446/lat -32.9575)">Rosario</a> [32°57'27"S, 60°38'22"W], ii.1930, A Stevenin leg., 1 ♂ (MACN -En) .</p><p>Diagnosis.</p><p>L: 14.2–16.0 mm (males), 14.5–17.2 mm (females). W: 6.4–7.3 mm (males), 6.5–8.0 mm (females). Body shape (Figs 1 D, 2 D, 3 D): navicular, more or less attenuated anteriorly and posteriorly, slightly convex; elytral margin weakly reflexed or not reflexed. Prosternal keel (Fig. 3 D): ventral margin nearly straight; posterior spine acute, moderately long. Metaventral keel (Fig. 3 D): long and sharp, reaching or slightly surpassing posterior margin of first abdominal ventrite. Male: mesotibial setigerous furrow absent (Table 1; Fig. 7 C); fifth abdominal ventrite with small glabrous area, &lt;1 / 4 of its total length (Fig. 4 G); protarsal claws nearly straight, with long, slender proximal tooth; mesotarsal claws similar, with long, slender, slightly hooked proximal tooth; metatarsal claws slightly dissimilar, anterior claw with shorter, more strongly hooked proximal tooth; genitalia: median lobe broad proximally, narrowed at mid-length, and expanded distally, apex rounded, with two lateral projections directed proximally; accessory appendages of median lobe elongate, excurved, with acute distal ends; parameres shorter than median lobe, narrow and weakly curved ventrally (Fig. 5 M – P). Female: fifth abdominal ventrite with a small glabrous area, ~ 1 / 5 or less of its total length (Fig. 4 H).</p><p>Distribution.</p><p>Brazil (Mouchamps 1959; Bachmann 1963), Paraguay (Mouchamps 1959), Uruguay (Bachmann 1988), Argentina (provinces): Buenos Aires (Bachmann 1963, 1988; this study), Chaco (Mouchamps 1959; Bachmann 1963, 1988; Libonatti et al. 2013; this study), Corrientes (Bachmann 1963, 1988; Gómez Lutz et al. 2012; this study), Entre Ríos (Bachmann 1963, 1988; Torres et al. 2007), Santa Fe (Bachmann 1963, 1988; Macchia et al. 2015; this study), Tucumán (Bachmann 1988).</p><p>Biology.</p><p>The species was collected from a variety of lentic habitats, including temporary and semi-permanent ponds, shallow pools, and flooded areas. Habitats ranged from small, sun-exposed ponds (&lt;20 cm deep and ≤ 100 m 2) with turbid water and muddy substrate (Libonatti et al. 2013; Torres et al. 2007) to larger, permanent water bodies (≤ 2 m deep and ≤ 2500 m 2) (Torres et al. 2007; Gómez Lutz et al. 2012). Sites had variable turbidity (from semi-transparent to highly turbid water rich in organic matter), muddy substrates, and marginal and emergent aquatic vegetation including floating macrophytes. Some localities were temporary, with surface area and depth fluctuating due to desiccation, while others were more permanent. All collection sites were exposed to full sunlight and located in areas with low anthropogenic impact. Adults were also collected using light traps (Libonatti et al. 2013; Macchia et al. 2015; this study).</p><p>Remarks.</p><p>It can be distinguished from other similar species by the small glabrous area of the fifth abdominal ventrite (~ 1 / 4 of its total length in males (Fig. 4 G), 1 / 5 in females (Fig. 4 H)) and the absence of the male mesotibial setigerous furrow (Fig. 7 C). Hydrobiomorpha spinosa is the most similar species, sharing a long metaventral keel that reaches or occasionally surpasses the posterior margin of the first abdominal ventrite; however, H. spinosa has a very long male mesotibial furrow (~ 2 / 3 of tibial length (Fig. 7 F, J)) and a minute or absent glabrous area on the fifth abdominal ventrite (Fig. 4 M). Hydrobiomorpha irina sesquirina, H. irina varginha, and H. irinoides (all Bachmann 1988) are larger; in H. irina sesquirina the fifth abdominal ventrite is slightly tectiform, with the glabrous area extending anteriorly as a narrow strip.</p></div>	https://treatment.plazi.org/id/141480FD9D835135990A4D0FAA1A3EA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Urcola, Matías R.;Rodriguez, Georgina;Michat, Mariano C.;Torres, Patricia L. M.	Urcola, Matías R., Rodriguez, Georgina, Michat, Mariano C., Torres, Patricia L. M. (2026): Revision of the genus Hydrobiomorpha Blackburn, 1888 (Coleoptera, Hydrophilidae) from Argentina, with the description of a new species and an updated identification key. ZooKeys 1289: 109-139, DOI: 10.3897/zookeys.1289.202453
B590A4CC7FEB57D9A39A4F93E1CD0496.text	B590A4CC7FEB57D9A39A4F93E1CD0496.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrobiomorpha longa (Bruch 1915)	<div><p>Hydrobiomorpha longa (Bruch, 1915)</p><p>Figs 1 E, 2 E, 3 E, 4 I – J, 5 Q – T, 6 N, 7 D, 7 H</p><p>Hydrophilus longus Bruch, 1915: 448–449, fig. 1 (original description).</p><p>Neohydrophilus longus: d’Orchymont 1919: 161 –162.</p><p>Hydrobiomorpha [(Hydrobiomorpha)] longa: Mouchamps 1959: 327, 330–331.</p><p>Hydrobiomorpha (Hydrobiomorpha) longa: Bachmann 1963: 30 –32, fig. 2 (key to Argentine species).</p><p>Hydrobiomorpha longa: Contartese and Bachmann 1987: fig. 5 (distribution only).</p><p>Hydrobiomorpha longa: Bachmann 1988: 10, 19–21, fig. 7 A – H (redescription; key to species of the Americas).</p><p>Type locality.</p><p>Argentina: Santa Fe Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.383335&amp;materialsCitation.latitude=-29.566668" title="Search Plazi for locations around (long -60.383335/lat -29.566668)">La Gallareta</a> [29°34'00"S, 60°23'00"W].</p><p>Material examined.</p><p>Argentina: • Buenos Aires Province: Zárate, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.93667&amp;materialsCitation.latitude=-33.909725" title="Search Plazi for locations around (long -58.93667/lat -33.909725)">camping Isla Talavera</a>, 33°54'35"S, 58°56'12"W, 15.xi.2025, G Rodriguez leg., light trap, 1 ♂ (LEBA) . • Chaco Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.45&amp;materialsCitation.latitude=-26.783333" title="Search Plazi for locations around (long -60.45/lat -26.783333)">Presidencia Roque Sáenz Peña</a> [26°47'00"S, 60°27'00"W], 30.xi.1980, 2 ♂♂ (MACN -EN) . • Entre Ríos Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.017223&amp;materialsCitation.latitude=-32.397224" title="Search Plazi for locations around (long -58.017223/lat -32.397224)">Concordia</a> [32°23'50"S, 58°01'02"W] [no date], JB Daguerre leg., 1 ♂ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.644447&amp;materialsCitation.latitude=-30.740557" title="Search Plazi for locations around (long -59.644447/lat -30.740557)">La Paz</a> [30°44'26"S, 59°38'40"W], xii.1974, 1 ♂ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.433334&amp;materialsCitation.latitude=-32.35" title="Search Plazi for locations around (long -58.433334/lat -32.35)">Pronunciamiento</a> [32°21'00"S, 58°26'00"W], ii.1958, L Gontero leg., 1 ♀ (MACN -En) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.191113&amp;materialsCitation.latitude=-32.153336" title="Search Plazi for locations around (long -58.191113/lat -32.153336)">Liebig</a> [32°09'12"S, 58°11'28"W], x.1988, MR Zelich leg., light trap, 2 ♂♂ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.166668&amp;materialsCitation.latitude=-32.616665" title="Search Plazi for locations around (long -60.166668/lat -32.616665)">Victoria</a> [32°37'00"S, 60°10'00"W], 21–26.iii.1981, AO Bachmann leg., 2 ♂♂ and 5 ♀♀ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.40056&amp;materialsCitation.latitude=-32.163055" title="Search Plazi for locations around (long -58.40056/lat -32.163055)">Villa Elisa</a> [32°09'47"S, 58°24'02"W], xi.1973, L Gontero leg., 2 ♂♂ (LEBA) . • Santa Fe Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.639446&amp;materialsCitation.latitude=-32.9575" title="Search Plazi for locations around (long -60.639446/lat -32.9575)">Rosario</a> [32°57'27"S, 60°38'22"W], ii.1930, A Stevenin leg., 1 ♂ (MACN -En) .</p><p>Diagnosis.</p><p>L: 14.0– 15.3 mm (males), 14.8–16.6 mm (females). W: 6.2–6.8 mm (males), 6.9–7.5 mm (females). Body shape (Figs 1 E, 2 E, 3 E): narrow, sides subparallel at mid-length, slightly tapered posteriorly and anteriorly, slightly convex; elytral margins not reflexed. Prosternal keel (Fig. 3 E): moderately long, ventral margin nearly straight; posterior spine short, acute. Metaventral keel (Fig. 3 E): moderately developed, posterior tip short and sharp, not reaching first abdominal ventrite. Male: mesotibial setigerous furrow absent (Table 1; Fig. 7 D, H); ventral inner face of mesotibia with only a small distal group of spines; fifth abdominal ventrite with glabrous area ~ 1 / 3 of its total length (Fig. 4 I); protarsal claws with robust proximal tooth; mesotarsal claws with rounded, broad proximal tooth; metatarsal claws with posterior claw bearing conspicuous angular preapical expansion (Fig. 6 N); genitalia: median lobe slender, gradually tapering distally; apex bifurcate with two long sharp projections directed distally; accessory appendages of median lobe elongate, sclerotized, divergent distally, with truncate distal ends; parameres shorter than median lobe, narrow in lateral view, gradually narrowed distally and ending in rather blunt projections directed dorsally (Fig. 5 Q – T). Female: fifth abdominal ventrite without glabrous area (Fig. 4 J).</p><p>Distribution.</p><p>Paraguay (Mouchamps 1959; Bachmann 1963, 1988), Argentina (provinces): Buenos Aires (Bachmann 1963, 1988; this study), Chaco (Bachmann 1988; Libonatti et al. 2013), Corrientes (Bachmann 1963, 1988), Entre Ríos (Bachmann 1963, 1988), Formosa (Bachmann 1963, 1988), Salta (Bachmann 1963), Santa Fe (Bruch 1915; d’Orchymont 1928; Mouchamps 1959; Bachmann 1963, 1988), Tucumán (Bachmann 1988).</p><p>Biology.</p><p>Adults inhabit moderately vegetated permanent ponds (this study) and have been recorded from a stream with unshaded, muddy substrate, relatively clear water, and floating vegetation (Libonatti et al. 2013), representing the only lotic habitat record for the genus in Argentina. Adults occasionally fly at night and are attracted to artificial lights (this study).</p><p>Remarks.</p><p>It can be distinguished from other similar species by the posterior metatarsal claw of males, which bears a conspicuous angular preapical expansion (Fig. 6 N), and by being the smallest and narrowest species in the genus (14.0– 16.6 mm). The largest specimens of H. longa resemble externally the smallest ones of H. irina irina; however, H. irina irina has a longer and sharper metaventral keel and a denser, more regular marginal series of elytral punctures. H. corumbaensis, originally described as a subspecies of H. longa, is larger and has a blunt metaventral keel and the posterior metatarsal claw of males bears a rounded rather than angular preapical expansion (Fig. 6 M). The median lobe of the male genitalia ends in two long sharp projections in H. longa (Fig. 5 Q, R) whereas in H. corumbaensis the projections are short and blunt (Fig. 5 E, F).</p></div>	https://treatment.plazi.org/id/B590A4CC7FEB57D9A39A4F93E1CD0496	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Urcola, Matías R.;Rodriguez, Georgina;Michat, Mariano C.;Torres, Patricia L. M.	Urcola, Matías R., Rodriguez, Georgina, Michat, Mariano C., Torres, Patricia L. M. (2026): Revision of the genus Hydrobiomorpha Blackburn, 1888 (Coleoptera, Hydrophilidae) from Argentina, with the description of a new species and an updated identification key. ZooKeys 1289: 109-139, DOI: 10.3897/zookeys.1289.202453
7B5F145CEA0E57C197F87608D3CA7724.text	7B5F145CEA0E57C197F87608D3CA7724.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrobiomorpha rudesculpta (d'Orchymont 1939)	<div><p>Hydrobiomorpha rudesculpta (d’Orchymont, 1939)</p><p>Figs 1 F, 2 F, 3 F, 4 K, 4 L, 5 U – X, 6 O, 7 E, 7 I</p><p>Neohydrophilus rudesculptus d’Orchymont, 1939 a: 376–378 (original description).</p><p>Hydrobiomorpha rudesculpta: Mouchamps 1959: 324, 333.</p><p>Hydrobiomorpha (Hydrobiomorpha) rudesculpta: Bachmann 1963: 29, 32, fig. 3 (key to Argentine species).</p><p>[Neohydrophilus] rudesculptus: Döbler 1977: 386.</p><p>Hydrobiomorpha rudesculpta: Contartese and Bachmann 1987: fig. 2 (distribution only).</p><p>Hydrobiomorpha rudesculpta: Bachmann 1988: 8, 45–46, fig. 30 A – H (redescription; key to species of the Americas).</p><p>Type locality.</p><p>Paraguay: Río Pilcomayo.</p><p>Material examined.</p><p>Argentina: • Chaco Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.986668&amp;materialsCitation.latitude=-27.45139" title="Search Plazi for locations around (long -58.986668/lat -27.45139)">Resistencia</a> [27°27'05"S, 58°59'12"W] [no date], JB Daguerre leg., 1 ♂ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.708336&amp;materialsCitation.latitude=-27.291666" title="Search Plazi for locations around (long -60.708336/lat -27.291666)">San Bernardo</a> [27°17'30"S, 60°42'30"W], ii.1980, OR Di Iorio leg., light trap, 1 ♂ and 2 ♀♀ (LEBA) . • Corrientes Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.195835&amp;materialsCitation.latitude=-28.54611" title="Search Plazi for locations around (long -57.195835/lat -28.54611)">Iberá lagoon</a>, 28°32'46"S, 57°11'45"W, 7.xi.2019, JI Urcola and G Rodriguez leg., light trap, 1 ♀ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.295555&amp;materialsCitation.latitude=-29.357224" title="Search Plazi for locations around (long -59.295555/lat -29.357224)">Manantiales</a> [29°21'26"S, 59°17'44"W], ix.1960, 1 ♀ (LEBA) . • Salta Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-64.46889&amp;materialsCitation.latitude=-24.754723" title="Search Plazi for locations around (long -64.46889/lat -24.754723)">Piquete</a> [24°45'17"S, 64°28'08"W], 18.i.1928, A Bridarolli leg., 1 ♀ (MACN -En) . • Santa Fe Province: Chaco [likely refers to the biogeographic province] [no date], C Bruch leg., 1 ♂ (MACN -En) . • No locality data: [no date], A Breyer leg., 1 ♀ (MACN -En) .</p><p>Diagnosis.</p><p>L: 16.3–18.8 mm (males), 17.2–20.3 mm (females). W: 8.8–9.9 mm (males), 9.4–11.6 mm (females). Body shape (Figs 1 F, 2 F, 3 F): broadly oval, widest slightly posterior to mid-length of elytra, strongly convex (subglobose); elytral margins not reflexed. Prosternal keel (Fig. 3 F): strongly oblique, posterior spine very long. Metaventral keel (Fig. 3 F): very short, posterior tip blunt, not reaching mid-length of first abdominal ventrite. Male: mesotibial setigerous furrow absent (Table 1; Fig. 7 E, I); ventral inner face of mesotibia with a very dense distal patch of spinules; claws highly distinctive; protarsal claws with enormous spatulate proximal tooth on inner claw; mesotarsal claws with very large, rounded, oblique proximal tooth; metatarsal claws long, with large, rounded proximal tooth (Fig. 6 O); fifth abdominal ventrite with small, wide, poorly delimited glabrous area (Fig. 4 K); genitalia: median lobe very broad, largely membranous, apex with two large rounded lobes; accessory appendages of median lobe narrow in dorsal view, broad in lateral view, strongly sclerotized, distally bent at a right angle toward the midline, bearing an acute outwardly directed process at the point of inflection; parameres slightly shorter than median lobe, with a subdistal dorsal projection directed proximally and a complex basal lobe (Fig. 5 U – X). Female: fifth abdominal ventrite without glabrous area (Fig. 4 L).</p><p>Distribution.</p><p>Paraguay (d’Orchymont 1939 a; Mouchamps 1959; Bachmann 1963, 1988), Argentina (provinces): Chaco (Bachmann 1963, 1988; this study), Corrientes (Bachmann 1963, 1988; Torres et al. 2012; this study), Entre Ríos (Bachmann 1963, 1988), Formosa (Bachmann 1963, 1988), Salta (Bachmann 1963, 1988), Santa Fe (Bachmann 1963).</p><p>Biology.</p><p>To date, the biology of this species remains unknown; however, it is frequently collected using light traps (Torres et al. 2012; this study).</p><p>Remarks.</p><p>Hydrobiomorpha rudesculpta is the most anatomically distinctive Argentine species of the genus, recognizable by its subglobose body shape, the spatulate proximal tooth of the anterior protarsal claw in males (Fig. 6 O), and the distinctive structure of the male genitalia (Fig. 5 U – X; see Diagnosis for details). No other species combines these characters.</p></div>	https://treatment.plazi.org/id/7B5F145CEA0E57C197F87608D3CA7724	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Urcola, Matías R.;Rodriguez, Georgina;Michat, Mariano C.;Torres, Patricia L. M.	Urcola, Matías R., Rodriguez, Georgina, Michat, Mariano C., Torres, Patricia L. M. (2026): Revision of the genus Hydrobiomorpha Blackburn, 1888 (Coleoptera, Hydrophilidae) from Argentina, with the description of a new species and an updated identification key. ZooKeys 1289: 109-139, DOI: 10.3897/zookeys.1289.202453
AFB0D82B5B985A6284CA5A2BF54D87B6.text	AFB0D82B5B985A6284CA5A2BF54D87B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrobiomorpha spinosa (d'Orchymont 1928)	<div><p>Hydrobiomorpha spinosa (d’Orchymont, 1928)</p><p>Figs 1 G, 2 G, 3 G, 4 M, 4 N, 5 Y – AB, 7 F, 7 J</p><p>Hydrophilus medius: Brullé, 1837: 54–55 (partim, only the record from Rio Negro, Patagonia).</p><p>Hydrochares medius: Berg 1884: C [= 100] (fide Bachmann 1988).</p><p>Neohydrophilus medius var. spinosus d’Orchymont, 1928: 164 (original description).</p><p>Hydrobiomorpha [(Hydrobiomorpha)] spinosa: Mouchamps 1959: 322, 327, 328.</p><p>Hydrobiomorpha spinosa: Bachmann 1963: 30, 32, 33, fig. 4 (key to Argentine species).</p><p>Hydrobiomorpha spinosa: Contartese and Bachmann 1987: fig. 4 (distribution only).</p><p>Hydrobiomorpha spinosa: Bachmann 1988: 8, 37–39, fig. 23 A – I (redescription; key to species of the Americas).</p><p>Hydrobiomorpha spinosa: Archangelsky et al. 2004: 253 –264 (description of immature stages).</p><p>Type localities.</p><p>Argentina: Santiago del Estero Province, bords du Rio Salado, environs d’Icaño, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-62.883892&amp;materialsCitation.latitude=-28.678055" title="Search Plazi for locations around (long -62.883892/lat -28.678055)">Mistol Paso</a> [28°40'41"S, 62°53'02"W], and Chaco de Santiago del Estero, Rio Salado; Santa Fe Province, Chaco de Santa Fe, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.50028&amp;materialsCitation.latitude=-28.846111" title="Search Plazi for locations around (long -59.50028/lat -28.846111)">Las Garzas</a> [28°50'46"S, 59°30'01"W], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.50028&amp;materialsCitation.latitude=-28.846111" title="Search Plazi for locations around (long -59.50028/lat -28.846111)">bords du Rio Las Garzas, 20 km à l’Ouest d’Ocampo</a>.</p><p>Material examined.</p><p>Argentina: • Buenos Aires Province: Olavarría, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.330555&amp;materialsCitation.latitude=-36.893333" title="Search Plazi for locations around (long -60.330555/lat -36.893333)">Tapalqué stream</a>, 36°53'36"S, 60°19'50"W, 23.i.2026, TA Favier Dubois leg., 3 ♀♀ (LEBA) . • Chaco Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.606667&amp;materialsCitation.latitude=-26.80861" title="Search Plazi for locations around (long -59.606667/lat -26.80861)">Chaco National Park</a> [26°48'31"S, 59°36'24"W], 18.i.2011, MC Michat and SA Mazzucconi leg, 2 ♂♂ and 2 ♀♀ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.45&amp;materialsCitation.latitude=-26.783333" title="Search Plazi for locations around (long -60.45/lat -26.783333)">Presidencia Roque Sáenz Peña</a> [26°47'00"S, 60°27'00"W], 22.xii.1991, light trap, 1 ♂ (LEBA) . • Entre Ríos Province: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.2275&amp;materialsCitation.latitude=-31.869444" title="Search Plazi for locations around (long -58.2275/lat -31.869444)">El Palmar National Park</a>, 31°52'10"S, 58°13'39"W, 11.xi.2023, G Rodriguez leg., 1 ♀ (LEBA) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.166668&amp;materialsCitation.latitude=-32.616665" title="Search Plazi for locations around (long -60.166668/lat -32.616665)">Victoria</a> [32°37'00"S, 60°10'00"W], 23–26.iii.1981, AO Bachmann leg., 1 ♂ and 1 ♀ (LEBA) . • Salta Province: Rosario de la Frontera, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-64.943054&amp;materialsCitation.latitude=-25.834167" title="Search Plazi for locations around (long -64.943054/lat -25.834167)">Camping ACA</a> [25°50'03"S, 64°56'35"W], 27.xi.2003, light trap, PLM Torres and MC Michat leg., 1 ♂ and 4 ♀♀ (LEBA) .</p><p>Diagnosis.</p><p>L: 13.7–17.9 mm (males), 14.0– 20.5 mm (females). W: 6.6–8.3 mm (males), 6.7–10.3 mm (females). Body shape (Figs 1 G, 2 G, 3 G): variable, ranging from elliptical to navicular, anterior and posterior ends tapered to different degrees, moderately convex; elytral margins not reflexed. Prosternal keel (Fig. 3 G): variable, generally long; posterior spine either very long and robust or short and acute. Metaventral keel (Fig. 3 G): very long, sharp, often slightly hooked, reaching or surpassing posterior margin of first abdominal ventrite. Male: mesotibial setigerous furrow present, very long, ~ 1 / 2–2 / 3 of tibial length, with dense reddish setal brush (Table 1; Fig. 7 F, J); fifth abdominal ventrite with minute triangular glabrous area, commonly &lt;1 / 5 of its total length (Fig. 4 M); protarsal claws slightly dissimilar, proximal tooth long, slender, somewhat flattened; mesotarsal claws robust, proximal tooth long, blunt; metatarsal claws slightly dissimilar, proximal tooth long, blunt; genitalia: median lobe narrow, subparallel; apex truncate, with two short rounded lateral projections directed proximally, visible in ventral view; dorsal sclerite as a prominent lamina projecting perpendicularly to the longitudinal axis of the median lobe; accessory appendages of median lobe elongate, excurved, slightly sigmoid, with acute distal ends; parameres shorter than median lobe, moderately narrow in lateral view, strongly curved ventrally (Fig. 5 Y – AB). Female: fifth abdominal ventrite without glabrous area (Fig. 4 N).</p><p>Distribution.</p><p>Brazil (d’Orchymont 1928), Paraguay (Mouchamps 1959; Bachmann 1963), Uruguay (Bachmann 1963; Benetti and Garrido 2004), Argentina (provinces): Buenos Aires (d’Orchymont 1928; Mouchamps 1959; Bachmann 1963, 1988; von Ellenrieder and Fernández 2000; Archangelsky et al. 2004; Fontanarrosa et al. 2004; Fernández et al. 2010; Macchia 2022; Macchia and Cicchino 2023; this study), Chaco (Bachmann 1963, 1988; Libonatti et al. 2013; this study), Chubut (Bachmann 1963), Córdoba (Bachmann 1963, 1988), Corrientes (Bachmann 1963, 1988; Torres et al. 2012), Entre Ríos (Bachmann 1963, 1988; Torres et al. 2007; this study), Formosa (Bachmann 1963, 1988), La Pampa (Bachmann 1988), Mendoza (Bachmann 1963, 1988), Misiones (Bachmann 1963, 1988), Río Negro (Bachmann 1988), Salta (Bachmann 1963, 1988; this study), San Juan (Bachmann 1988), San Luis (Bachmann 1988), Santa Fe (d’Orchymont 1928; Mouchamps 1959; Bachmann 1963, 1988), Santiago del Estero (Mouchamps 1959; Bachmann 1963, 1988), Tucumán (Bachmann 1963, 1988).</p><p>Biology.</p><p>Hydrobiomorpha spinosa is the most widely distributed American species of the genus. Adults are scavengers or periphyton eaters, inhabiting vegetated ponds, characterized by abundant organic matter, turbid water, and a muddy substrate. Adults occasionally fly at night and are attracted to artificial lights (Torres et al. 2007, 2012; Libonatti et al. 2013; Macchia and Cicchino 2023). Larvae are active predators but are rarely collected. Archangelsky et al. (2004) provided the first description of the egg-case for the genus and described all larval instars and the pupa.</p><p>Remarks.</p><p>It can be distinguished from other similar species by the very long male mesotibial setigerous furrow (1 / 2–2 / 3 of tibial length) and the minute or absent glabrous area of the fifth abdominal ventrite. Small specimens resemble H. irina irina; however, H. irina irina has a shorter metaventral keel, lacks the male mesotibial furrow, and has a well-developed glabrous area in both sexes (~ 1 / 4 of its total length in males, 1 / 5 in females). In H. spinosa, the glabrous area typically occupies about 1 / 5 of the total length of the fifth abdominal ventrite but may be reduced to ~ 1 / 10 (Bachmann 1988). The median lobe of the male genitalia of H. irina irina is broader, with a rounded apex bearing lateral projections (Fig. 5 M, N), whereas in H. spinosa the apex is subtruncate with rounded lateral expansions (Fig. 5 Y, Z).</p></div>	https://treatment.plazi.org/id/AFB0D82B5B985A6284CA5A2BF54D87B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Urcola, Matías R.;Rodriguez, Georgina;Michat, Mariano C.;Torres, Patricia L. M.	Urcola, Matías R., Rodriguez, Georgina, Michat, Mariano C., Torres, Patricia L. M. (2026): Revision of the genus Hydrobiomorpha Blackburn, 1888 (Coleoptera, Hydrophilidae) from Argentina, with the description of a new species and an updated identification key. ZooKeys 1289: 109-139, DOI: 10.3897/zookeys.1289.202453
A34ABB4E3A365C0CA802A7F2C61C2073.text	A34ABB4E3A365C0CA802A7F2C61C2073.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydrobiomorpha utiariti Bachmann 1988	<div><p>Hydrobiomorpha utiariti Bachmann, 1988</p><p>Figs 5 AC – AF, 9</p><p>Hydrobiomorpha utiariti Bachmann, 1988: 8, 18, 19, fig. 6 A – K (original description).</p><p>Type locality.</p><p>Brazil: Mato Grosso Norte, Utiariti, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.2825&amp;materialsCitation.latitude=-13.027222" title="Search Plazi for locations around (long -58.2825/lat -13.027222)">Río Papagaio</a> [13°01'38"S, 58°16'57"W] .</p><p>Material examined.</p><p>Brazil: • Mato Grosso, Utiariti, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-58.2825&amp;materialsCitation.latitude=-13.027222" title="Search Plazi for locations around (long -58.2825/lat -13.027222)">Río Papagaio</a> [13°01'38"S, 58°16'57"W], xi.1966, JM Viana leg., 1 ♂ (holotype, MACN -En 2746) and 1 ♀ (allotype of Bachmann 1988, MACN -En 2747) . Paraguay: • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-57.95139&amp;materialsCitation.latitude=-22.1575" title="Search Plazi for locations around (long -57.95139/lat -22.1575)">Puerto Vallemí</a> [22°09'27"S, 57°57'05"W], 11.vi.1952, AO Bachmann leg., 1 ♀ (paratype, MACN -En 2748) .</p><p>Diagnosis.</p><p>L: 22.0 mm (male holotype), 21.0– 22.5 mm (females). W: 10.0 mm (male holotype), 9.4–10.0 mm (females). Body shape (Fig. 9 A, B): navicular, sides subparallel, slightly expanded at mid-length, moderately convex; elytral margins slightly reflexed, especially posteriorly, more distinctly in males than in females. Prosternal keel (Fig. 9 C): ventral margin straight, posterior spine very short. Metaventral keel (Fig. 9 C): slightly broadened, with a fine longitudinal groove; posterior end rounded, reaching middle of first abdominal ventrite. Male: mesotibial setigerous furrow present, ~ 1 / 2 of tibial length, narrow, with a dense brush of reddish setae (Table 1); fifth abdominal ventrite with a subtriangular, very large glabrous area, ~ 2 / 3 of its total length (Fig. 9 E); claws robust; protarsal claws with robust proximal tooth on inner claw; mesotarsal claws with somewhat spatulate proximal tooth; metatarsal claws unknown, not preserved in the holotype (only known male); genitalia: median lobe broad, apex truncate, bearing two slightly divergent projections arising centrally from the margin; accessory appendages of median lobe elongate, strongly sclerotized, abruptly narrowed distally, terminating in short outwardly directed apical hooks; parameres as long as the median lobe, broad in lateral view, strongly curved, with subdistal denticulations and apices directed toward the midline (Fig. 5 AC – AF). Female: fifth abdominal ventrite with subtriangular glabrous area, &lt;1 / 2 of its total length (Fig. 9 F).</p><p>Distribution.</p><p>Brazil (Bachmann 1988), Paraguay (Bachmann 1988).</p><p>Biology.</p><p>Unknown.</p><p>Remarks.</p><p>Hydrobiomorpha utiariti is included herein because it overlaps in distribution with H. bachmanni sp. nov. in Paraguay and presents similar anatomical features, particularly in body size, convexity, and development of male mesotibial setigerous furrow and glabrous area of fifth abdominal ventrite, which may lead to confusion between the two species. Hydrobiomorpha utiariti differs from H. bachmanni sp. nov. by its larger size, darker coloration, longer male mesotibial setigerous furrow (~ 1 / 2 of tibial length, whereas in H. bachmanni sp. nov. it extends ~ 1 / 3), and larger glabrous area in both sexes (~ 2 / 3 of its total length in males (Fig. 9 E) and &lt;1 / 2 in females (Fig. 9 F), whereas ~ 2 / 5 in both sexes in H. bachmanni sp. nov. (Fig. 4 A, B). It also differs in the prosternal keel, which is shorter in H. utiariti and bears a short posterior spine, and in the shape of the male genitalia: the median lobe has two slightly divergent apical projections in H. utiariti (Fig. 5 AC – AF), whereas in H. bachmanni sp. nov. these are subparallel (Fig. 5 A – D); and the parameres of H. utiariti bear conspicuous subdistal internal denticulations that are absent in H. bachmanni sp. nov.</p></div>	https://treatment.plazi.org/id/A34ABB4E3A365C0CA802A7F2C61C2073	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Urcola, Matías R.;Rodriguez, Georgina;Michat, Mariano C.;Torres, Patricia L. M.	Urcola, Matías R., Rodriguez, Georgina, Michat, Mariano C., Torres, Patricia L. M. (2026): Revision of the genus Hydrobiomorpha Blackburn, 1888 (Coleoptera, Hydrophilidae) from Argentina, with the description of a new species and an updated identification key. ZooKeys 1289: 109-139, DOI: 10.3897/zookeys.1289.202453
