identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
23ECA58C637A5DF383EE28BDD503B971.text	23ECA58C637A5DF383EE28BDD503B971.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Habeastrum decussatum Hausdorf & Tapia 2026	<div><p>Habeastrum decussatum Hausdorf &amp; Tapia sp. nov.</p><p>Fig. 1 M – R, Tables 1, 2</p><p>Type material.</p><p>Holotype: Ecuador, Sucumbios province: • Lumbaqui, via la Troncal km 12, sector Dumbique, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.2997&amp;materialsCitation.latitude=-0.0364" title="Search Plazi for locations around (long -77.2997/lat -0.0364)">Dashino River</a>, in leaf litter at base of non-calcareous rocky wall in a small river cliff, 0.0364°S, 77.2997°W, 500 m a. s. l., 22. 07. 2025, E. Tapia &amp; A. Guasti leg., QCAZI 282661 . Paratypes: • same data as holotype: QCAZI 282662 / 1; ZMH -MOL-165491 / 2 .</p><p>Diagnosis.</p><p>Shell minute (1.3 mm high), corneous-brown, wide, elongated conical, with 4.5 strongly convex whorls, protoconch (2 whorls) with dense, irregular indentations, teleoconch with fine ribs (24–25 ribs / 0.5 mm) crossed by spiral lines, and a narrow umbilicus taking 8–10 % of the shell diameter.</p><p>Description.</p><p>Shell elongated conical; 4.5 strongly convex whorls; protoconch (c. 2 whorls) with dense, irregular indentations; teleoconch finely ribbed (24–25 ribs / 0.5 mm on ventral side of body whorl) crossed by distinct spiral lines; corneous-brown; aperture circular, prosocline; peristome continuous, attached to body whorl, not expanded, not reflexed and hardly thickened on the inside; umbilicus open, narrow, taking 8–10 % of shell diameter. Measurements see Table 2.</p><p>Etymology.</p><p>Named after the decussate sculpture consisting of fine radial ribs crossed by spiral lines (Latin decussatus, marked with lines intersecting cross-wise, used as an adjective).</p><p>Distribution.</p><p>Habeastrum decussatum sp. nov. is only known from piedmont evergreen forest in the north of the Eastern Cordillera in Sucumbios province in Ecuador at 500 m a. s. l.</p><p>Remarks.</p><p>Habeastrum decussatum sp. nov. differs from all other Habeastrum species in the wide conical shell with fine radial ribs crossed by distinct spiral lines on the teleoconch, and distinct irregular indentations on the protoconch (Table 2).</p></div>	https://treatment.plazi.org/id/23ECA58C637A5DF383EE28BDD503B971	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Hausdorf, Bernhard;Roosen, Marijn T.;Tapia, Elicio	Hausdorf, Bernhard, Roosen, Marijn T., Tapia, Elicio (2026): First records of Habeastrum species from Ecuador (Gastropoda, Diplommatinidae). Evolutionary Systematics 10 (2): 295-301, DOI: 10.3897/evolsyst.10.199967
E4C832D7C15B573E8237846CAD9DE4A2.text	E4C832D7C15B573E8237846CAD9DE4A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Habeastrum Simone 2019	<div><p>Habeastrum Simone, 2019</p><p>Habeastrum Simone, 2019: 287 . Type species (by original designation): Habeastrum parafusum Simone, 2019 .</p><p>Diagnosis.</p><p>Habeastrum is characterised by minute dextral, elongated conical to cylindrical shells with regularly increasing whorls, an open umbilicus, and a non-expanded peristome.</p></div>	https://treatment.plazi.org/id/E4C832D7C15B573E8237846CAD9DE4A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Hausdorf, Bernhard;Roosen, Marijn T.;Tapia, Elicio	Hausdorf, Bernhard, Roosen, Marijn T., Tapia, Elicio (2026): First records of Habeastrum species from Ecuador (Gastropoda, Diplommatinidae). Evolutionary Systematics 10 (2): 295-301, DOI: 10.3897/evolsyst.10.199967
2EF7F0E7A77057AC89C8F11EBF81EDCF.text	2EF7F0E7A77057AC89C8F11EBF81EDCF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Habeastrum striolatum Hausdorf & Tapia 2026	<div><p>Habeastrum striolatum Hausdorf &amp; Tapia sp. nov.</p><p>Fig. 1 A – F, Tables 1, 2</p><p>Type material.</p><p>Holotype: Ecuador, Pastaza province: • Puyo, Chuwitayo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.7884&amp;materialsCitation.latitude=-1.9511" title="Search Plazi for locations around (long -77.7884/lat -1.9511)">Cueva de los Tayos</a>, inside the cave on the bottom of a 100 m vertical sinkhole, 1.9511°S, 77.7884°W, 740 m a. s. l., 24. 11. 2025, E. Tapia leg., QCAZI 282654 . Paratypes: • same data as holotype: QCAZI 282655 / 1; ZMH -MOL-165097 / 2 .</p><p>D, large diameter of shell; H, height of shell; n, number of measured individuals; std. dev., standard deviation; U, width of umbilicus; W, number of whorls.</p><p>D, large diameter of shell; H, height of shell; da, diameter of aperture; ha, height of aperture; R, ribs / 0.5 mm on body whorl; U, width of umbilicus; W, number of whorls.</p><p>Diagnosis.</p><p>Shell minute (1.3 mm high), white, elongated conical, with 5.25–5.5 strongly convex whorls, protoconch with indistinct, radial wrinkles, teleoconch with dense riblets (26–38 ribs / 0.5 mm), and a narrow umbilicus taking 9–11 % of the shell diameter.</p><p>Description.</p><p>Shell elongated conical; 5.25–5.5 strongly convex whorls; protoconch (c. 2 whorls) with indistinct, radial wrinkles; teleoconch with dense riblets (26–38 riblets / 0.5 mm on ventral side of body whorl); white; aperture almost circular, prosocline; peristome continuous, attached to body whorl, not expanded, not reflexed and hardly thickened on the inside; umbilicus open, narrow, taking 9–11 % of shell diameter. Measurements see Table 2.</p><p>Etymology.</p><p>Named after the striolate sculpture with fine linear ridges (Latin striolatus, striated, used as an adjective).</p><p>Distribution.</p><p>Habeastrum striolatum sp. nov. is known only from the inside of the Cueva de los Tayos in piedmont evergreen forest on limestone outcrops of the Amazonian Cordilleras in Pastaza province in Ecuador at 740 m a. s. l.</p><p>Remarks.</p><p>Habeastrum striolatum sp. nov. resembles the sympatric Habeastrum tayos sp. nov. in shell form and size, but differs from it in the finer and denser ribbing (Table 2). Habeastrum striolatum sp. nov. is smaller than the Brazilian Habeastrum species. Concerning shell form, Habeastrum striolatum sp. nov. is intermediate between the shorter conical Habeastrum omphalium Simone, 2019, and longer conical Habeastrum strangei Simone, Cavallari &amp; Salvador, 2020. Habeastrum parafusum Simone, 2019 has a more cylindrical-conical shell. The sculpture of Habeastrum striolatum sp. nov. (26–38 riblets / 0.5 mm on ventral side of body whorl) is most similar to that of H. omphalium (ca. 22 riblets / 0.5 mm), but this species has faint spiral lines between the ribs, which are lacking in H. striolatum sp. nov. The ribbing is more dense in H. parafusum (ca. 40 riblets / 0.5 mm), and less dense in H. strangei (ca. 9 riblets / 0.5 mm).</p></div>	https://treatment.plazi.org/id/2EF7F0E7A77057AC89C8F11EBF81EDCF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Hausdorf, Bernhard;Roosen, Marijn T.;Tapia, Elicio	Hausdorf, Bernhard, Roosen, Marijn T., Tapia, Elicio (2026): First records of Habeastrum species from Ecuador (Gastropoda, Diplommatinidae). Evolutionary Systematics 10 (2): 295-301, DOI: 10.3897/evolsyst.10.199967
A7B0546D5AE15FF99E8AA9208BB2B53D.text	A7B0546D5AE15FF99E8AA9208BB2B53D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Habeastrum tayos Hausdorf, Roosen & Tapia 2026	<div><p>Habeastrum tayos Hausdorf, Roosen &amp; Tapia sp. nov.</p><p>Fig. 1 G – L, Tables 1, 2</p><p>Type material.</p><p>Holotype: Ecuador, Pastaza province: • Puyo, Chiwitayu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.7875&amp;materialsCitation.latitude=-1.9478" title="Search Plazi for locations around (long -77.7875/lat -1.9478)">Cueva de los Tayos area</a>, 1.9478°S, 77.7875°W, 740 m a. s. l., 07. 11. 2025, E. Tapia &amp; A. Guasti leg., QCAZI 282656 . Paratypes: Ecuador, Pastaza province: • same data as holotype. QCAZI 282657 / 1; ZMH -MOL-165103 / 5 . • Puyo, Chiwitayu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.7877&amp;materialsCitation.latitude=-1.9477" title="Search Plazi for locations around (long -77.7877/lat -1.9477)">Cueva de los Tayos area</a>, 1.9477°S, 77.7877°W, 750 m a. s. l., 07. 11. 2025, E. Tapia &amp; A. Guasti leg., ZMH -MOL-165577 / 1 . • Puyo, Chiwitayu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.7879&amp;materialsCitation.latitude=-1.9475" title="Search Plazi for locations around (long -77.7879/lat -1.9475)">Cueva de los Tayos area</a>, 1.9475°S, 77.7879°W, 760 m a. s. l., 08. 11. 2025, E. Tapia &amp; A. Guasti leg., ZMH -MOL-165578 / 2 . • Puyo, Chiwitayu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.7875&amp;materialsCitation.latitude=-1.9473" title="Search Plazi for locations around (long -77.7875/lat -1.9473)">Cueva de los Tayos area</a>, 1.9473°S, 77.7875°W, 750 m a. s. l., 08. 11. 2025, E. Tapia &amp; A. Guasti leg., INABIO -MCL-1255 / 1, ZMH -MOL-165579 / 1 . • Puyo, Chiwitayu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.7873&amp;materialsCitation.latitude=-1.9479" title="Search Plazi for locations around (long -77.7873/lat -1.9479)">Cueva de los Tayos area</a>, 1.9479°S, 77.7873°W, 720 m a. s. l., 08. 11. 2025, E. Tapia &amp; A. Guasti leg., QCAZI 282658 / 3, ZMH -MOL-165580 / 3 . • Puyo, Chiwitayu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.7869&amp;materialsCitation.latitude=-1.9481" title="Search Plazi for locations around (long -77.7869/lat -1.9481)">Cueva de los Tayos area</a>, 1.9481°S, 77.7869°W, 730 m a. s. l., 09. 11. 2025, E. Tapia leg., QCAZI 282659 / 2, ZMH -MOL-165581 / 2 . • Puyo, Chiwitayu, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.7867&amp;materialsCitation.latitude=-1.9482" title="Search Plazi for locations around (long -77.7867/lat -1.9482)">Cueva de los Tayos area</a>, 1.9482°S, 77.7867°W, 730 m a. s. l., 09. 11. 2025, E. Tapia leg., QCAZI 282660 / 1, ZMH -MOL-165582 / 2 . – Ecuador, Morona-Santiago province: • Sunka Biological Station, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.75&amp;materialsCitation.latitude=-2.0667" title="Search Plazi for locations around (long -77.75/lat -2.0667)">sampling station SU 7</a>, 2.0667°S, 77.7500°W, 1032 m a. s. l., 01. 03. 2026, M. T. Roosen &amp; L. Tucupi Catan leg., INABIO -MCL-1253 / 1, INABIO -MCL-1254 / 1 .</p><p>Other material.</p><p>Ecuador, Napo province: • Tena, Cotundo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.04959&amp;materialsCitation.latitude=-0.22109" title="Search Plazi for locations around (long -78.04959/lat -0.22109)">Santa Helena de los Guacamayos towards Cosanga</a>, 0.6609°S, 77.7930°W, 1700 m a. s. l., 07. 11. 2025, D. Kammin, K. Kopp &amp; E. Tapia leg., ZMH -MOL-165576 / 1 . Chaco, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.04959&amp;materialsCitation.latitude=-0.22109" title="Search Plazi for locations around (long -78.04959/lat -0.22109)">Oyacachi</a>, 0.22109°S, 78.04959°W, 3090 m a. s. l., 16. 11. 2025, D. Kammin, K. Kopp &amp; E. Tapia leg., ZMH -MOL-165583 / 1 .</p><p>Diagnosis.</p><p>Shell minute (1.2–1.6 mm high), white, elongated conical, with 4.5–6 strongly convex whorls, coarse ribs (8–12 ribs / 0.5 mm), and a narrow umbilicus taking 7 % of the shell diameter.</p><p>Description.</p><p>Shell elongated conical; 4.5–6 strongly convex whorls; protoconch (c. 2 whorls) with a dense pattern of wavy wrinkles; teleoconch coarsely ribbed (8–12 ribs / 0.5 mm on ventral side of body whorl); white; aperture circular, prosocline; peristome continuous, attached to body whorl, not expanded, not reflexed and hardly thickened on the inside; umbilicus open, narrow, taking up to 7 % of shell diameter. Measurements see Table 2.</p><p>Etymology.</p><p>The specific epithet refers to the oilbird ( Steatornis caripensis Humboldt, 1817), also known as the ‘ tayo’ in Spanish (used as a noun in apposition). This species breeds in caves, many of which are named after it, including caves at the type locality and around Sunka.</p><p>Distribution.</p><p>Habeastrum tayos sp. nov. is known from piedmont evergreen forest on limestone outcrops in the Amazonian Cordilleras in Pastaza and Morona-Santiago provinces at 720–1032 m a. s. l. It was found in litter between calcareous rocks. Similar shells were found at higher altitudes in low montane evergreen forest of the northern Eastern Andes (1700 m a. s. l.) and in high montane evergreen forest of the northern Eastern Andes (3090 m a. s. l.) in Napo province. There they occurred in a very acidic habitat without calcareous rocks. Their specific identity has to be proven, given the different habitats. There were no statistical differences between the eleven measured specimens from Pastaza and the two specimens from the Eastern Andes in the shell dimensions (Student’s t-tests, two-tailed: diameter of shell, p = 0.37; height of shell, p = 0.27).</p><p>Remarks.</p><p>Habeastrum tayos sp. nov. differs from all the other known Habeastrum species in the much coarser ribbing (Table 2).</p></div>	https://treatment.plazi.org/id/A7B0546D5AE15FF99E8AA9208BB2B53D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Hausdorf, Bernhard;Roosen, Marijn T.;Tapia, Elicio	Hausdorf, Bernhard, Roosen, Marijn T., Tapia, Elicio (2026): First records of Habeastrum species from Ecuador (Gastropoda, Diplommatinidae). Evolutionary Systematics 10 (2): 295-301, DOI: 10.3897/evolsyst.10.199967
