identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
12E40BB3BBB45C9588F5E67A7763F921.text	12E40BB3BBB45C9588F5E67A7763F921.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riukiaria Attems 1938	<div><p>Genus Riukiaria Attems, 1938</p><p>Type species.</p><p>Riukiaria pugionifera Verhoeff, 1936 .</p><p>Generic diagnosis.</p><p>Body with 20 rings, compact and relatively small, displaying an often vivid colour pattern. Posterolateral corners of paraterga blunt on rings 1–4, becoming acutely produced on rings 5–18. Gonopods simple, biramous, and forceps-like; solenomere consistently longer and stouter than prefemoral process (when present). Gonopod coxite with 0–2 setae (Korsós et al. 2011; Golovatch 2014).</p><p>Included species or subspecies.</p><p>Riukiaria anachoreta Tanabe, 1988, R. belousovi Golovatch, 2014, R. bifida (Takakuwa, 1942), R. capaca Wang &amp; Zhang, 1993, R. chelifera (Takakuwa, 1941), R. chinensis Tanabe, Ishii &amp; Yin, 1996, R. cohaesiva (Wang, 1957), R. contigua (Wang, 1957), R. cornuta (Haga, 1968), R. cucfuongensis (Nguyen, 2016), R. datei (Miyosi, 1957), R. davidiani Golovatch, 2014, R. diacantha (Miyosi, 1952), R. falcifera Verhoeff, R. geniculata (Takakuwa, 1941), R. holstii (Pocock, 1895), R. jamila Tanabe, 1990, R. kabaki Golovatch, 2014, R. korolevi Golovatch, 2014, R. maculata Korsós, Nakamura &amp; Tanabe, 2011, R. marinae (Golovatch, 1978), R. martensi Golovatch, 2014, R. montana (Haga, 1968), R. mundyi Korsós, Nakamura &amp; Tanabe, 2011, R. ochracea (Gressitt, 1941), R. puella Tanabe, 1988, R. pugionifera Verhoeff, 1936, R. rosulans (Tömösváry, 1885), R. scutata (Takakuwa, 1942), R. semicircularis semicircularis (Takakuwa, 1941), R. semicircularis hosidei (Miyosi, 1952), R. spatuliformis Golovatch, 2015, R. spina Chen, Zheng &amp; Jiang, 2023, R. spiralipes (Takakuwa, 1942), R. taiwana (Takakuwa, 1942), R. tianmu (Tanabe, Ishii &amp; Yin, 1996), R. uncata (Haga, 1968), R. uraensis (Wang, 1956) .</p></div>	https://treatment.plazi.org/id/12E40BB3BBB45C9588F5E67A7763F921	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Han, Zewu;Liu, Xin;Wang, Jiajia;Dong, Yan	Han, Zewu, Liu, Xin, Wang, Jiajia, Dong, Yan (2026): Description of a new species of the genus Riukiaria (Diplopoda, Polydesmida, Xystodesmidae) from eastern China, with the characterization of its complete mitochondrial genome. ZooKeys 1272: 315-335, DOI: 10.3897/zookeys.1272.182977
4DC1038EEAA95AD4A30A68F74816DFE8.text	4DC1038EEAA95AD4A30A68F74816DFE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Riukiaria langyaensis Han & Wang 2026	<div><p>Riukiaria langyaensis Han &amp; Wang sp. nov.</p><p>Figs 1, 2, 3, 4, 5</p><p>Material examined.</p><p>Holotype: China • ♂, Anhui, Chuzhou City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=118.46917&amp;materialsCitation.latitude=32.433056" title="Search Plazi for locations around (long 118.46917/lat 32.433056)">Langya Mountain Forest Farm</a>, 32°25'59"N, 118°28'09"E, alt. 100 m, 7 May 2025, Z. W. Han &amp; J. J Wang leg, NMCHZU DH 28. GenBank: PX 436095 . Paratypes: • 1 ♀, same locality and date as holotype, NMCHZU DH 29 .</p><p>Other material.</p><p>China • 3 ♂ 2 ♀, Anhui, Chuzhou County, Niutou Mountain, 32°27'47"N, 118°29'97"E, alt. 87 m, 25 May 2025, Z. W. Han &amp; J. J Wang leg, NMCHZU DH 30 -34 .</p><p>Diagnosis.</p><p>This species closely resembles Riukiaria mundyi Korsós, Nakamura &amp; Tanabe, 2011 in tergal features and body coloration. However, Riukiaria langyaensis sp. nov. differs by having a lighter body color than R. mundyi, with antennae and walking legs exhibiting a wheat-colored hue and the paratergal margins appearing translucent. In terms of gonopod structure, the prefemorite of Riukiaria langyaensis sp. nov. is similar to that of Riukiaria contigua (Wang, 1957), but its acropodite is more inflated and nearly spherical, with an indistinctly forked tip, and the prefemoral processes are completely missing.</p><p>Description.</p><p>Length of holotype 23.6 mm, midbody paranotal width 4.8 mm, metatergal width 2.9 mm. Female body length 25.1 mm, midbody paranotal width 5.1 mm, metatergal width 3.4 mm.</p><p>Coloration in life nearly uniform dark orange, with posterior 2 / 5 of body occasionally pale orange in some individuals. Epicranium concolorous with body; antennae and legs wheat-coloured. No sexual dimorphism observed. Under UV light, entire body exhibiting strong sky-blue fluorescence. In alcohol, vivid coloration fading rapidly, most metaterga becoming light marbled orange.</p><p>Body with 20 rings. Pro- and metazonae smooth, devoid of tubercles, punctation or wrinkles, except for corrugated folds on ventral side of metaterga. Head smooth, epicranial suture distinct; forehead raised, surface uneven. Mandibular stipes ventrally depressed. Several setae scattered above clypeus, with two dense transverse rows at its margin and on labrum. Antennae straight, reaching behind midway of body ring 3 when stretched dorsally; first antennomere globose, second slightly clavate, subequal with antennomeres 3–5, sixth longest, c. 1.2 times as long as the fifth, seventh small, slender, slightly longer than wide.</p><p>In width, head &lt;collum &lt;ring 2 &lt;3 &lt;4 = 11, thereafter body gradually tapering towards telson. Collum convex, smooth, shiny; lateral and posterior margin with weak ridge, lateral corners triangular, slightly directed caudad. All paraterga clearly delimited by a dorsal sulcus, lateral calluses broad and distinct; margins translucent. Caudolateral corner of paraterga invariably rounded, in collum acute, in rings 2–4 subrectangular, in following rings growing increasingly acute and drawn well caudad behind rear tergal margin, nearly beak-shaped in posterior rings (Fig. 2 A – C). Pore formula normal (5, 7, 9, 10, 12, 13, 15–18), ozopores lying in median excavation of paraterga (in lateral view). Epiproct short, in dorsal view subtriangular, in lateral view protruding, parallel-sided, slightly curved ventrad, with 7 + 7 setae, 2 + 2 of them on knobs. Paraprocts strongly marginate with 2 + 2 setae; hypoproct with 1 + 1 setae on knobs (Fig. 2 F).</p><p>Sterna without modifications, mostly smooth; very narrow between legs 1 and 2, clearly broader until ring 7, especially broad behind gonopods. Legs generally densely setose, robust, c. 1.1–1.2 times as long as midbody height, last pair considerably shorter than previous ones (Fig. 4 B, C). Coxae 3–7 especially densely setose ventrally. Prefemora with a moderately developed ventral spine, growing increasingly stronger towards body end; femur straight, c. 2.2 times as long as prefemur; postfemur stout; tibia straight, subequal to postfemur; tarsus slender, c. 1.8 times as long as tibia; claw curved, c. 0.25 mm long. Coxa 2 with a strong, median projection about half as long as coxa proper apically with a membranous tubula surrounded by strong setae; prefemoral joint with 1 + 1 macrosetae (Fig. 4 A).</p><p>Gonopod aperture transversely oblong, c. 1.75 times as broad as long, with elevated lateral and caudal edges. Gonopod coxa short, stout, subcylindrical, with two coxal macrosetae, and a normal, prominent, mesal cannula. Prefemoral process entirely absent. Prefemorite with dense setae on left side. Acropodite inflated, oval, distinctly curved, with a prostatic groove. Solenomere apical, bifurcated (Fig. 4 D, E).</p><p>Distribution.</p><p>Known only from the type locality in Chuzhou, Anhui Province, China.</p><p>Etymology.</p><p>The species name is derived from the geographic name of the location where the species was first discovered.</p><p>Remarks.</p><p>The genus Riukiaria comprises 38 species distributed across China, Japan, and Vietnam (Marek et al. 2014; Golovatch 2015; Chen et al. 2023; Korsós and Nakamura 2025). In China, 18 species have been reported from Shaanxi (1 species), Sichuan (5 species), Zhejiang (2 species), Fujian (1 species), Chongqing (1 species), and Taiwan (8 species). Riukiaria langyaensis sp. nov. is the first species of Riukiaria reported from Anhui Province.</p></div>	https://treatment.plazi.org/id/4DC1038EEAA95AD4A30A68F74816DFE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Han, Zewu;Liu, Xin;Wang, Jiajia;Dong, Yan	Han, Zewu, Liu, Xin, Wang, Jiajia, Dong, Yan (2026): Description of a new species of the genus Riukiaria (Diplopoda, Polydesmida, Xystodesmidae) from eastern China, with the characterization of its complete mitochondrial genome. ZooKeys 1272: 315-335, DOI: 10.3897/zookeys.1272.182977
