identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
6C3F6F09B0DE5CD4BC5F6661B93E89F4.text	6C3F6F09B0DE5CD4BC5F6661B93E89F4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche (Atopsaura) callosa (Navas 1924)	<div><p>Atopsyche (Atopsaura) callosa (Navás, 1924)</p><p>Atopsyche (Atopsaura) callosa Navás, 1924: 78 [Type locality: Costa Rica; MNHNP; ♂; in Ventrarma]. Flint 1975: 566 [synonymy]. McElravy et al. 1981: 152 [distribution]. Holzenthal 1988: 53 [distribution]. Sykora 1991: 250 [distribution]. Flint 1991: 18 [♂; distribution]. Aguila 1992: 533 [distribution]. Flint 1996: 379 [distribution]. Angrisano 1999: 26 [checklist]. Muñoz-Quesada 2000: 275 [checklist]. Holzenthal and Cressa 2002: 134 [distribution]. Rueda Martín 2006: 58 [larva; pupa; distribution]. Isa Miranda and Rueda Martín 2014: 199 [distribution]. Armitage et al. 2015: 8 [checklist]. Armitage and Cornejo 2015: 197 [checklist].</p><p>Atopsyche (Atopsaura) alconura Ross, 1953: 288 [Type locality: Peru, Dpto. Cuzco, Pcia. Paucartambo, Cosnipata Valley; INHS; ♂ ]. Flint 1975: 566 [to synonymy].</p><p>Atopsyche (Atopsaura) schmidi Denning, 1965: 267 [Type locality: Costa Rica, San Jose; CAS; ♂]. Flint 1975: 566 [to synonymy].</p><p>Material examined.</p><p>Peru • 2 ♂ 24 ♀; Pasco, small creek in Yanachaga-Chemillén NP buffer zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.4732&amp;materialsCitation.latitude=-10.4278" title="Search Plazi for locations around (long -75.4732/lat -10.4278)">San Daniel sector</a>; 10.4278°S, 75.4732°W, 2134 m a. s. l.; 16 Nov. 2010; C. Carranza, J. Peralta leg.; light trap; MUSM . • 1 ♂ 6 ♀; same data as preceding, but 17 Nov. 2010; MUSM .</p><p>Distribution.</p><p>Argentina, Bolivia, Colombia, Costa Rica, Ecuador, Panama, Peru (Cuzco Department, Pasco Department), Venezuela.</p><p>Remarks.</p><p>Atopsyche callosa is one of the most widely distributed species in the genus, recorded from Costa Rica to Argentina (Holzenthal and Calor 2017). In Peru, it has previously been reported from Cuzco (Flint 1996). In this study, we provide the first record of A. callosa from Pasco.</p></div>	https://treatment.plazi.org/id/6C3F6F09B0DE5CD4BC5F6661B93E89F4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
D61A64C1835D50368D2CD5CD7F96F81A.text	D61A64C1835D50368D2CD5CD7F96F81A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche (Atopsaura) lobosa Ross & King 1952	<div><p>Atopsyche (Atopsaura) lobosa Ross &amp; King, 1952</p><p>Atopsyche (Atopsaura) lobosa Ross &amp; King, 1952: 200 [Type locality: Bolivia, Incachaca; CMNH; ♂ ]. Flint 1996: 380 [distribution]. Angrisano 1999: 26 [checklist]. Ríos-Touma et al. 2017: 27 [distribution]. Mey and Ospina-Torres 2018: 24 [distribution]. Rázuri-Gonzales et al. 2025: 27 [distribution].</p><p>Material examined.</p><p>Peru • 1 ♂; Pasco, small creek in Yanachaga-Chemillén NP buffer zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.4732&amp;materialsCitation.latitude=-10.4278" title="Search Plazi for locations around (long -75.4732/lat -10.4278)">San Daniel sector</a>; 10.4278°S, 75.4732°W, 2134 m a. s. l.; 16 Jun. 2010; E. Rázuri, C. Carranza leg.; light trap; MUSM . • 1 ♀; same data as preceding, but 17 Jun. 2010; MUSM . • 1 ♂ 4 ♀; Pasco, Yanachaga-Chemillén NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.3578&amp;materialsCitation.latitude=-10.5452" title="Search Plazi for locations around (long -75.3578/lat -10.5452)">Quebrada San Alberto at Refugio El Cedro</a>; 10.5452°S, 75.3578°W, 2421 m a. s. l.; 27 Aug 2015; E. Rázuri, L. Figueroa, B. Portuguez leg.; light trap; MUSM .</p><p>Distribution.</p><p>Bolivia, Ecuador, Colombia, Peru (Cuzco, Pasco).</p><p>Remarks.</p><p>Atopsyche lobosa was originally described from Bolivia, and subsequently also found in Peru (Cuzco Department). Later, Ríos-Touma et al. (2017) recorded it from Ecuador for the first time. Mey and Ospina-Torres (2018) have also recorded it from Colombia. Rázuri-Gonzales et al. (2025) recorded it from the Zamora Chinchipe Province (Ecuador). Here we also record it from the Pasco Department.</p></div>	https://treatment.plazi.org/id/D61A64C1835D50368D2CD5CD7F96F81A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
9AA73A42ECA856E18ECFD27F442218BD.text	9AA73A42ECA856E18ECFD27F442218BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche (Atopsaura) puharcocha Schmid 1989	<div><p>Atopsyche (Atopsaura) puharcocha Schmid, 1989</p><p>Atopsyche (Atopsaura) puharcocha Schmid, 1989: 129 [Type locality: Bolivia, Prov. Cochabamba, Rio Ronquito, rd. to Villa Tunari, Chapare; NMNH; ♂ ]. Sykora 1991: 150 [distribution]. Flint 1996: 380 [distribution]. Angrisano 1999: 27 [checklist].</p><p>Material examined.</p><p>Peru • 1 ♂ 2 ♀; Pasco, small creek in Yanachaga-Chemillén NP buffer zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.4732&amp;materialsCitation.latitude=-10.4278" title="Search Plazi for locations around (long -75.4732/lat -10.4278)">San Daniel sector</a>; 10.4278°S, 75.4732°W, 2134 m a. s. l.; 03 May 2011; C. Carranza, M. Alvarado, L. Figueroa leg.; light trap; MUSM . • 2 ♀; same data as preceding, but 04 May 2011; MUSM .</p><p>Distribution.</p><p>Bolivia, Ecuador, Peru (Cuzco Department, Pasco Department).</p><p>Remarks.</p><p>This species was first described by Schmid (1989) from Bolivia, and later reported by Sykora (1991) from Ecuador. Flint (1996) recorded it from Puente San Pedro in the Cosñipata Valley (Cuzco Department). In this study, we report it in the San Daniel Sector of the Yanachaga-Chemillén NP (Pasco Department).</p></div>	https://treatment.plazi.org/id/9AA73A42ECA856E18ECFD27F442218BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
44E2937ABB3453FF8316FCC99EF6305A.text	44E2937ABB3453FF8316FCC99EF6305A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche (Atopsaura) tincuracu Schmid 1989	<div><p>Atopsyche (Atopsaura) tincuracu Schmid, 1989</p><p>Atopsyche (Atopsaura) tincuracu Schmid, 1989: 130 [Type locality: Bolivia, Prov. La Paz, Unduavi / Coroico; NMNH; ♂ ]. Angrisano 1999: 27 [checklist].</p><p>Material examined.</p><p>Peru • 1 ♂ 1 ♀; Pasco, Yanachaga-Chemillén NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.3578&amp;materialsCitation.latitude=-10.5452" title="Search Plazi for locations around (long -75.3578/lat -10.5452)">Quebrada San Alberto at Refugio El Cedro</a>; 10.5452°S, 75.3578°W, 2421 m a. s. l.; 27 Aug 2015; E. Rázuri, L. Figueroa, B. Portuguez leg.; light trap; MUSM . • 1 ♂; same data as preceding; UMSP .</p><p>Distribution.</p><p>Bolivia, Peru (Pasco Department).</p><p>Remarks.</p><p>Atopsyche tincuracu was originally described from Bolivia. In this study, we record its occurrence in Peru (Pasco Department), approximately 1000 km northwest of the type locality.</p></div>	https://treatment.plazi.org/id/44E2937ABB3453FF8316FCC99EF6305A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
9B567C8953885D22B255D364D7934DC9.text	9B567C8953885D22B255D364D7934DC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche (Atopsyche) explanata Ross 1953	<div><p>Atopsyche (Atopsyche) explanata Ross, 1953</p><p>Atopsyche (Atopsyche) explanata Ross, 1953: 288 [Type locality: Peru, Dpto. Cuzco, Pcia. Paucartambo, Cosnipata Valley; INHS; ♂ ].</p><p>Material examined.</p><p>Peru • 5 ♂; Pasco, small creek in Yanachaga-Chemillén NP buffer zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.4732&amp;materialsCitation.latitude=-10.4278" title="Search Plazi for locations around (long -75.4732/lat -10.4278)">San Daniel sector</a>; 10.4278°S, 75.4732°W, 2134 m a. s. l.; 16 Jun. 2010; E. Rázuri, C. Carranza leg.; light trap; MUSM . • 4 ♂; same data as preceding, but 17 Jun. 2010; MUSM . • 4 ♂; same data as preceding, but 03 May 2011; C. Carranza, M. Alvarado, L. Figueroa leg.; MUSM . • 3 ♂; same data as preceding, but 04 May 2011; MUSM . • 1 ♂; same data as preceding, but 24 Aug. 2015; E. Rázuri, L. Figueroa, B. Portuguez leg.; UMSP .</p><p>Distribution.</p><p>Peru (Cuzco Department, Pasco Department).</p><p>Remarks.</p><p>Ross (1953) described Atopsyche explanata from an unspecified locality in the Cosñipata Valley in the Cuzco Department (Peru). It has also been recorded from Puente Unión in the Cosñipata Valley (UMSP; GBIF. org 2025). We record it for the first time from the Pasco Department, approximately 520 km northwest of Puente Unión.</p></div>	https://treatment.plazi.org/id/9B567C8953885D22B255D364D7934DC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
3EDD2084E8AE51A0A9F78F85756A3D1B.text	3EDD2084E8AE51A0A9F78F85756A3D1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche (Atopsyche) ikonnikovi Martynov 1912	<div><p>Atopsyche (Atopsyche) ikonnikovi Martynov, 1912</p><p>Atopsyche (Atopsyche) ikonnikovi Martynov, 1912: 34 [Type locality: Peru, La Merced; type depository unknown; ♂ ]. Ulmer 1913: 384 [distribution]. Ross and King 1952: 195 [♂]. Angrisano 1999: 27 [checklist].</p><p>Material examined.</p><p>Peru: 1 ♂; Huánuco, Monson [Monzón] Valley, Tingo María; 23 Sep. 1954; E. I. Schlinger, E. S. Ross leg.; CAS .</p><p>Distribution.</p><p>Argentina, Bolivia, Peru (Huánuco Department, Junín Department).</p><p>Remarks.</p><p>This species was originally described from La Merced in central Peru (Martynov 1912). Ulmer (1913) later recorded it from Salta, Argentina, and Angrisano (1999) noted its presence in Bolivia. We now present a new record from Tingo María, located approximately 200 km north of La Merced.</p></div>	https://treatment.plazi.org/id/3EDD2084E8AE51A0A9F78F85756A3D1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
886EFE6676D35290A85F830C4F71B2D8.text	886EFE6676D35290A85F830C4F71B2D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche (Atopsyche) vatucra Ross 1953	<div><p>Atopsyche (Atopsyche) vatucra Ross, 1953</p><p>Atopsyche (Atopsyche) vatucra Ross, 1953: 290 [Type locality: Peru, Dpto. Cuzco, Pcia. Paucartambo, Cosnipata Valley; INHS; ♂ ]. Flint 1996: 379 [distribution].</p><p>Material examined.</p><p>Peru • 2 ♂ 1 ♀; Pasco, small creek in Yanachaga-Chemillén NP buffer zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.4732&amp;materialsCitation.latitude=-10.4278" title="Search Plazi for locations around (long -75.4732/lat -10.4278)">San Daniel sector</a>; 10.4278°S, 75.4732°W, 2134 m a. s. l.; 17 Jun. 2010; E. Rázuri, C. Carranza leg.; light trap; MUSM . • 1 ♂; same data as preceding, but 16 Nov. 2010; C. Carranza, J. Peralta leg.; MUSM . • 2 ♂; same data as preceding, but 17 Nov. 2010; MUSM . • 5 ♂ 1 ♀; same data as preceding, but 03 May 2011; C. Carranza, M. Alvarado, L. Figueroa leg.; MUSM . • 5 ♂ 2 ♀; same data as preceding, but 04 May 2011; MUSM .</p><p>Distribution.</p><p>Bolivia, Ecuador, Peru (Cuzco Department, Pasco Department), Venezuela.</p><p>Remarks.</p><p>Atopsyche vatucra was originally described from an unspecified locality in the Cosñipata Valley, Cuzco, Peru (Ross 1953), and subsequently reported from additional sites within the valley (Flint 1996). This species has also been documented in Bolivia, Ecuador, and Venezuela (UMSP; GBIF. org 2025). In this study, we report this species from the San Daniel sector in the Yanachaga-Chemillén National Park buffer zone, located approximately 500 km northwest of previously known localities for this species in Peru.</p></div>	https://treatment.plazi.org/id/886EFE6676D35290A85F830C4F71B2D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
DE8EF66EA2475A6E8462D6A0D9743618.text	DE8EF66EA2475A6E8462D6A0D9743618.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche (Dolochorema) bispinosa Schmid 1989	<div><p>Atopsyche (Dolochorema) bispinosa Schmid, 1989</p><p>Fig. 16 (wing venation), Fig. 17 (male genitalia)</p><p>Atopsyche (Dolochorema) bispinosa Schmid, 1989: 120 [Type locality: Bolivia, Dpto. La Paz, Unduavi-Coroico; NMNH; ♂ ]. Angrisano 1999: 27 [checklist].</p><p>Material examined.</p><p>Paratype. Bolivia • ♂; La Paz, Unduavi / Coroico; 2500 m a. s. l.; 19–25 Nov. 1984; L. E. Peña G. leg.; USNM .</p><p>Distribution.</p><p>Bolivia.</p></div>	https://treatment.plazi.org/id/DE8EF66EA2475A6E8462D6A0D9743618	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
6E4E3262C7865B35AE57A51959387962.text	6E4E3262C7865B35AE57A51959387962.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche (Dolochorema) huascarani Rázuri-Gonzales & Holzenthal 2025	<div><p>Atopsyche (Dolochorema) huascarani sp. nov.</p><p>Figs 8, 9</p><p>Type material.</p><p>Holotype. Peru • 1 ♂; Ancash, Huascarán NP, Puesto de control Llaca, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.4481&amp;materialsCitation.latitude=-9.4375" title="Search Plazi for locations around (long -77.4481/lat -9.4375)">Laguna Llaca</a>; 9.4375°S, 77.4481°W, 4421 m a. s. l.; 09 May 2016; P. Ancajima and R. Pradel leg.; light trap; MUSM -ENT-320969 (MUSM) . Paratypes. Peru • 1 ♂; same data as the holotype (UMSP) • 1 ♀; same data as the holotype (MUSM) .</p><p>Diagnosis.</p><p>Atopsyche yanachaga belongs to a group of species traditionally included in the subgenus Dolochorema . These species are characterized by the insertion of the second segment of the inferior appendage, which is positioned on the mesal surface of the first segment and is largely hidden in lateral view. Among these species, A. yanachaga is most similar to A. irregularis (Banks, 1913) (Peru) and A. bispinosa Schmid, 1989 (Bolivia) based on the shape of the parapods in lateral and dorsal view and the inferior appendages in ventral view. All three species have an elongate, narrow parapod, but both A. yanachaga and A. irregularis possess a strongly bent parapod at the base. In contrast, in dorsal view, A. yanachaga has an inflated, rounded parapod apically, while A. irregularis has a straight, narrow one. The first segment of the inferior appendages in these species is mitten-shaped in ventral view, but they can be distinguished by the shape of the second segment. In A. yanachaga, this segment extends beyond the posterior margin of the first segment, while in the other two species, they are small. Notably, A. yanachaga is the only species in this group that lacks the ventrolateral branch of the phallic apparatus.</p><p>Description.</p><p>Adult. Forewing length: male (12.5 mm, n = 2), female (13.5 mm, n = 1). Body brown (in alcohol), wings pale brown (in alcohol). Wing venation as in Fig. 8. Sterna III and IV without glands; sternum V with a pair of long, membranous glands; processes on sterna VI and VII short, triangular in ventral view, setose.</p><p>Male genitalia (Fig. 9). Segment IX, in lateral view, pentagonal, slightly higher than long, with setae on ventral and posterolateral surface (Fig. 9 A). Parapod, in lateral view, elongate, narrow, broader and strongly bent basally, apex rounded, dorsal margin and lateral surface with a few setae mesally (Fig. 9 A); in dorsal view, elongate, lateral margin sinuous, mesal margin convex mesally, setae on dorsal surface at mid-length, apex inflated and rounded (Fig. 9 B). Filipod digitate, as long as parapods, setose, apex rounded (Fig. 9 A). Preanal appendage short, rounded, setose (Fig. 9 A). First segment of inferior appendage, in lateral view, rectangular, ventral margin straight, slightly expanded basally and apically (less produced in the paratype), dorsal margin convex at mid-length, produced mesad into a rugose, triangular projection, bearing single spine-like setae apically (less slender in one of the sides of the paratype), with setae along margins and lateral surface (Fig. 9 A); in ventral view, mitten-shape, setose, lateral margin convex, mesal margin concave (Fig. 9 C); second segment of inferior appendage, in lateral view, elongate, bilobed, dorsal lobe subacute and directed dorsad, ventral lobe triangular, inserted on the mesal surface of the first segment of the inferior appendage (Fig. 9 A); in ventral view, roughly quadrangular, apicolateral corner produced into a rounded lobe, apex truncate (Fig. 9 C). Proctiger, in lateral view, narrow basally, slightly wider apically, with a short carina basolaterally, ventral margin membranous, densely covered with very fine setae, apex truncate (Fig. 9 A). Phallic apparatus simple; phallotheca broadly rounded basally, phallic apodeme indiscernible; with narrow ventral process articulating with inferior appendages, slightly broader at mid-length; ventrolateral branches of phallotheca absent; dorsal process of phallotheca absent; posterior section of phallotheca, in ventral view, broad basally, tapering towards apex, directed posterad, apex subacute (Fig. 9 D); in dorsal view, with a shallow notch mesally (Fig. 9 E); phallic spine, stout, spine-like structure, with a strong convex curvature near base, then slightly sinuous (Fig. 9 D); in dorsal view, apex acute (Fig. 9 E).</p><p>Distribution.</p><p>Peru: Ancash Department.</p><p>Etymology.</p><p>The specific epithet refers to the Huascarán National Park, from where the type specimen was collected.</p></div>	https://treatment.plazi.org/id/6E4E3262C7865B35AE57A51959387962	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
F4AE598EC4F351AC9F2A0940629F4ED8.text	F4AE598EC4F351AC9F2A0940629F4ED8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche (Dolochorema) irregularis (Banks 1913)	<div><p>Atopsyche (Dolochorema) irregularis (Banks, 1913)</p><p>Fig. 18 (wing venation), Fig. 19 (male genitalia)</p><p>Atopsyche (Dolochorema) irregularis Banks, 1913: 240 [Type locality: Peru, S. E., Cuzco; MCZ; ♂; in Dolochorema]. Ross 1956: 125 [♂]. Schmid 1989: 160 [to Atopsyche].</p><p>Material examined.</p><p>Holotype. Peru • ♂; Cuzco, S. E. Peru; 2300 m a. s. l.; Anton Heinrich Hermann Fassl leg.; Type 11785; MCZ.</p><p>Distribution.</p><p>Peru.</p><p>Remarks.</p><p>Atopsyche irregularis was the first species described in the genus Dolochorema (Banks, 1913), and later synonymized with Atopsyche (Schmid 1989) . It has only been recorded from the Cuzco Department in Peru.</p></div>	https://treatment.plazi.org/id/F4AE598EC4F351AC9F2A0940629F4ED8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
A8045900F681565FAC8A2F27B2F63BFF.text	A8045900F681565FAC8A2F27B2F63BFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche (Dolochorema) major Schmid 1989	<div><p>Atopsyche (Dolochorema) major Schmid, 1989</p><p>Fig. 20 (male genitalia)</p><p>Atopsyche (Dolochorema) major Schmid, 1989: 126 [Type locality: Bolivia, Prov. La Paz, Rio Zongo; NMNH; ♂ ]. Angrisano 1999: 27 [checklist].</p><p>Material examined.</p><p>Holotype. Bolivia • ♂; La Paz, Tarata, Río Zongo; 3200 m a. s. l.; 24–30 Oct. 1984; L. E. Peña G. leg.; USNM.</p><p>Distribution.</p><p>Bolivia.</p></div>	https://treatment.plazi.org/id/A8045900F681565FAC8A2F27B2F63BFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
6747C89A47C9592EAA05F961AD12D250.text	6747C89A47C9592EAA05F961AD12D250.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche cedroi Rázuri-Gonzales & Holzenthal 2025	<div><p>Atopsyche cedroi sp. nov.</p><p>Figs 2, 3</p><p>Type material.</p><p>Holotype. Peru • 1 ♂; Pasco, Yanachaga-Chemillén NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.3578&amp;materialsCitation.latitude=-10.5452" title="Search Plazi for locations around (long -75.3578/lat -10.5452)">Quebrada San Alberto at Refugio El Cedro</a>; 10.5452°S, 75.3578°W, 2421 m a. s. l.; 27 Aug. 2015; E. Rázuri, L. Figueroa and B. Portuguez leg.; light trap; UMSP 000220106 (MUSM) . Paratype. Peru • 1 ♀; same data as the holotype (MUSM) .</p><p>Diagnosis.</p><p>Atopsyche cedroi belongs to the bicolorata species group of Schmid (1989), characterized by having short inferior appendages and a broad notch apically on the first segment of these appendages. The second segment is reduced and inserted at the posterodorsal corner of or in the notch of the first segment. Among the species in this group, A. cedroi most closely resembles A. unicolorata Schmid, 1989 (Bolivia). In both species, the first segment of the inferior appendage is apically notched, with the second segment inserted at the posterodorsal corner of the first segment. However, A. cedroi differs in several key features. The apical notch is broad, and the posteroventral corner is rectangular, while in A. unicolorata, the apical notch is narrower, and the posteroventral corner is digitate. Additionally, the second segment of the inferior appendage in A. cedroi is elongated and downturned, whereas in A. unicolorata, it is subtriangular and directed posterad. Although both species have a simple phallic apparatus, the base of the phallotheca is broad in A. cedroi and narrowly rounded in A. unicolorata . Finally, while the parapods are similarly shaped in both species, in A. cedroi, they are broader mesally and feature a pair of prominent spine-like setae on their lateral surface, which are absent in A. unicolorata .</p><p>Description.</p><p>Adult. Forewing length: male (9.5 mm, n = 1), female (13.5 mm, n = 1). Body and wings mostly pale brown, head and thorax dark brown. Forewing with scattered brown setae and longer, erect straw-colored setae along major longitudinal veins; with irregular patch of dark brown setae delimited by the posterior margin, the stem of Cu 1, the apical stem of 1 A + 2 A + 3 A, and the base of the wing (corresponds with darker wing membrane) and adjacent much smaller patch of black setae on fork V; pterostigma slightly coriaceous. Wing venation as in Fig. 2. Sterna III and IV without glands; sternum V with a pair of long, membranous glands; process on sternum VI slightly shorter than its segment, digitate, bearing short setae on basal third and spine-like setae on apical two-thirds; process on sternum VII very short, digitate, bare.</p><p>Male genitalia (Fig. 3). Segment IX, in lateral view, subtriangular, slightly longer than high (Fig. 3 A). Parapod, in lateral view, shorter than inferior appendage, broader mesally, tapering towards apex, directed slightly posteroventrad, with setae on apical half and a pair of peg-like setae on mesoventral surface, apex rounded (Fig. 3 A); in dorsal view, digitate, lateral margin slightly sinuous, mesal margin straight, with setae apically, apex rounded (Fig. 3 B). Filipod digitate, shorter than parapods, setose, apex acute (Fig. 3 A). Preanal appendage short, rounded, setose (Fig. 3 A). First segment of inferior appendage, in lateral view, roughly quadrangular, ventral margin slightly sinuous, dorsal margin slightly convex apically, posteroventral corner produced into quadrate process, posterodorsal corner produced posterad, broadly notched apically, with long setae on ventral margin, short setae on lateral surface, and pair of peg-like setae subapically on mesal surface (three peg-like setae on opposite side) (Fig. 3 A); in ventral view, mitten-shaped, setose, lateral margin convex, mesal margin slightly sinuous with a quadrate process medially (Fig. 3 C); second segment of inferior appendage, in lateral view, digitate, with a few very short setae basally, slightly curved posteroventrad (Fig. 3 A); in ventral view, digitate, slightly curved mesad, apex narrowly rounded (Fig. 3 C). Proctiger, in lateral view, broadly widened apically, with truncate apical margin, covered with small setae (visible at 100 ×) (Fig. 3 A); in dorsal view, tapering apically. Phallic apparatus simple; phallotheca broadly rounded basally, phallic apodeme indiscernible; with ventral process articulating with inferior appendages, broad, constricted mesally; ventrolateral branches of phallotheca absent; dorsal process of phallotheca absent; posterior section of the phallotheca, in lateral view, broad basally, tapering towards apex, directed posterad, apex acute, membranous apically (Fig. 3 D); in dorsal view, with a notch mesally, apex directed posterad (Fig. 3 E); phallic spine elongate, stout, with a slight curvature near the base (Fig. 3 D); in dorsal view, with four erect setae on small, lateral bulges, apex acute (Fig. 3 E).</p><p>Distribution.</p><p>Peru: Pasco Department.</p><p>Etymology.</p><p>This new species is named after the locality where the type was collected, Refugio El Cedro (Yanachaga-Chemillén NP, Pasco Department, Peru).</p></div>	https://treatment.plazi.org/id/6747C89A47C9592EAA05F961AD12D250	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
76787BDA787E5434A9A97C78AAAE9CB4.text	76787BDA787E5434A9A97C78AAAE9CB4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche chemillen Rázuri-Gonzales & Holzenthal 2025	<div><p>Atopsyche chemillen sp. nov.</p><p>Figs 4, 5</p><p>Type material.</p><p>Holotype. Peru • 1 ♂; Pasco, small creek in Yanachaga-Chemillén NP buffer zone, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.4732&amp;materialsCitation.latitude=-10.4278" title="Search Plazi for locations around (long -75.4732/lat -10.4278)">San Daniel sector</a>; 10.4278°S, 75.4732°W, 2134 m a. s. l.; 03 May 2011; C. Carranza, M. Alvarado, L. Figueroa leg.; light trap; MUSM -ENT-320759 (MUSM) . Paratypes. Peru • 3 ♂; same data as the holotype, but 04 May 2011 • 1 ♂; same data as the holotype, but 16 Jun. 2010; E. Rázuri, C. Carranza leg. (MUSM) • 1 ♂; same data as the holotype, but 16 Nov. 2010; C. Carranza, J. Peralta leg. (MUSM) • 1 ♂ 1 ♀; same data as the holotype, but 24 Aug. 2015; E. Rázuri, L. Figueroa, B. Portuguez leg. (UMSP) • 2 ♀; same data as the holotype, but 24 Aug. 2015; E. Rázuri, L. Figueroa, B. Portuguez leg. (MUSM) .</p><p>Diagnosis.</p><p>Atopsyche chemillen is most similar to members of the falina species group of Ross and King (1952). This species group is characterized by the slightly produced posteroventral corner of the first segment of the inferior appendage. From all the members included therein, the new species is most similar to A. falina (Navás, 1930) (Argentina), A. mayucapac Schmid, 1989 (Venezuela), A. neolobosa Flint, 1963 (Ecuador), and A. yunguensis Rueda Martín, 2006 (Argentina, Bolivia) based on the presence of a spine-bearing dorsal process on the phallotheca. Atopsyche chemillen is most similar to A. mayucapac based on the straight inferior appendage and the shape of the phallotheca, but differs in the shape of the parapods in lateral view (the dorsal margin of this structure is biconcave in the new species but almost straight in A. mayucapac) and the shape of segment IX (quadrate in the new species but the dorsal margin is obliterated in A. mayucapac).</p><p>Description.</p><p>Adult. Forewing length: male (9 mm, n = 1), female (11 mm, n = 1). Body pale brown, wings brown. Forewing with scattered dark brown setae and longer straw-colored and brown setae along major longitudinal veins and wing margin. Wing venation as in Fig. 4. Tergum IV with elongate, flattened gland, opening on posterolateral margin, lined internally with minute spines; sternum V with a pair of tiny protuberances; process on sternum VI as long as its segment, curved, bearing spine-like setae along its length (the last one is peg-like), process on sternum VII shorter than process on previous segment, digitate, bearing very fine, short setae (only visible at 200 ×).</p><p>Male genitalia (Fig. 5). Segment IX, in lateral view, quadrangular, slightly higher than long (Fig. 5 A). Parapod, in lateral view, shorter than inferior appendage, same width along its length, with a spine-like lobe subapically on ventral margin, directed posterodorsad, with setae apically, apex concave (Fig. 5 A); in dorsal view, short, lateral margin with a spine-like projection subapically, mesal margin straight, with setae apically, apex upturned, concave (Fig. 5 B). Filipod digitate, longer than parapods, setose, apex slightly capitate (Fig. 5 A). Preanal appendage short, rounded, setose (Fig. 5 A). First segment of inferior appendage, in lateral view, long, straight, same width along its length, posteroventral corner slightly produced posterad, with setae on lateral and ventral surfaces (Fig. 5 A); in ventral view, slightly C-shaped, setose, lateral margin almost straight, mesal margin slightly concave, posterior margin truncate (Fig. 5 C); second segment of inferior appendage, in lateral view, subtriangular, very short, setose, dorsal and ventral margins straight, apex rounded (Fig. 5 A); in ventral view, falcate, apex acute (Fig. 5 C). Proctiger, in lateral view, broadly widened apically, with truncate apical margin, with a few long setae on its basal half; in dorsal view, tapering apically (Fig. 5 A). Phallic apparatus complex; phallotheca broadly rounded basally, phallic apodeme indiscernible; with ventral process articulating with inferior appendages, narrow, tapering towards apex; ventrolateral branches of phallotheca absent; dorsal process of phallotheca present, divided into pair of recurved, rounded lobes, bearing short spines on apical half; posterior section of phallotheca, in lateral view, broad basally, tapering towards apex, directed posterad, apical half covered with short setae, lateral surface with a serrated ridge, apex narrowly rounded (Fig. 5 D); in dorsal view, with a deep notch mesally, apex directed laterad (Fig. 5 E); phallic spine elongate, stout, with a strong curvature near the base (Fig. 5 D); in dorsal view, apex acute (Fig. 5 E).</p><p>Distribution.</p><p>Peru: Pasco Department.</p><p>Etymology.</p><p>Atopsyche chemillen is named after the Yanachaga-Chemillén National Park, where the type locality is situated. Chemillen means black or burned in Yanesha, referring to the dark appearance of these mountains during sunsets on sunny days.</p></div>	https://treatment.plazi.org/id/76787BDA787E5434A9A97C78AAAE9CB4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
F5F70BF4D0015CDD8CF81F7B000625A8.text	F5F70BF4D0015CDD8CF81F7B000625A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche corcuerai Rázuri-Gonzales & Holzenthal 2025	<div><p>Atopsyche corcuerai sp. nov.</p><p>Figs 6, 7</p><p>Type material.</p><p>Holotype. Peru • 1 ♂; Cajamarca; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.7809&amp;materialsCitation.latitude=-7.39689" title="Search Plazi for locations around (long -78.7809/lat -7.39689)">ACP Bosque Cachil</a>; 7.39689°S, 78.7809°W, 2522 m a. s. l.; 21–22 Apr. 2015; J. Peralta, P. Sánchez, M. Rodríguez, E. Sánchez, D. Silva, J. Grados leg.; light trap; MUSM -ENT-320967 (MUSM) . Paratypes. Peru • 9 ♂ 1 ♀; same data as the holotype (MUSM) • 2 ♂; same data as the holotype (UMSP) • 2 ♂; same data as the holotype (NMNH) • 2 ♂; same data as the holotype (SMF) .</p><p>Diagnosis.</p><p>This new species closely resembles Atopsyche cajas Harper &amp; Turcotte, 1985 (Ecuador), particularly in the position of the second segment of the inferior appendage, which is inserted on the mesal surface of the first segment. However, the two species can easily be separated based on differences in the shape of the parapod, inferior appendage, and phallic apparatus. In the new species, the parapod is narrow and tapering in lateral view, while in A. cajas, the parapod is broader and roughly uniform in width. In dorsal view, the parapod of the A. corcuerai is narrow and truncate apically, whereas in A. cajas, it is wider mesally and rounded apically. The apex of the first segment of the inferior appendages in A. corcuerai is falcate, while in A. cajas, it is broadly rounded. In ventral view, the second segment of the inferior appendages in A. corcuerai is roughly one-third the length of the first segment, while in A. cajas, it is about half the length of the first segment. Additionally, the mesal surface of the first segment of the inferior appendage is produced in A. corcuerai, whereas it is rounded in A. cajas . Finally, the posterior processes of the phallotheca in the new species are longer and narrower than those in A. cajas, and the phallic spine is straight in the new species, but slightly curved in A. cajas .</p><p>Description.</p><p>Adult. Forewing length: male (9 mm, n = 16), female (10 mm, n = 1). Body brown (in ethanol), wings pale brown (in ethanol). Wing venation as in Fig. 6. Sternum III – IV without glands; sternum V with a pair of long, membranous glands; process on sternum VI slightly longer than its segment, slightly curved, bearing fine setae on its basal half and spine-like setae on its apical half, process on sternum VII very short, subtriangular, bearing very fine setae.</p><p>Male genitalia (Fig. 7). Segment IX, in lateral view, pentagonal, shorter than high, with setae on posterolateral and posteroventral surface (Fig. 7 A). Parapod, in lateral view, elongate, sausage-shaped, broad mesally, tapering towards apex, directed slightly ventrad, dorsal and mesal surface with a few setae subapically, apex narrowly rounded (Fig. 7 A); in dorsal view, elongate, lateral margin slightly sinuous, mesal margin almost straight, setae on dorsal surface on apical half, apex truncate (Fig. 7 B). Filipod digitate, as long as parapods, setose, apex rounded (Fig. 7 A). Preanal appendage short, rounded, setose (Fig. 7 A). First segment of inferior appendage, in lateral view, oval, ventral margin convex, dorsal margin slightly convex subapically, posterior margin produced into short, subacute process, with setae along on ventral margin and dorsal surface subapically (Fig. 7 A); in ventral view, reniform, setose, lateral margin convex, mesal margin concave and slightly membranous with a mesal projection directed posteromesad (this projection is pointed on the right side and presents a slight cleft on the left side) (Fig. 7 C); second segment of inferior appendage, in lateral view, less than half the length of the first segment, ovate, dorsal and ventral margins slightly constricted mesally, apex subacute (Fig. 7 A); in ventral view, subtriangular, posterior margin slightly projected posterad, apex narrowly rounded (Fig. 7 C). Proctiger, in lateral view, narrow basally, wider apically, with a short carina laterodorsally on basal half, covered with minute setae, apex slightly rounded (Fig. 7 A); in dorsal view, tapering apically, with a slight mesal notch. Phallic apparatus complex; phallotheca broadly rounded basally, phallic apodeme indiscernible; with ventral process articulating with inferior appendages, broad, slightly broader basally; ventrolateral branches of phallotheca present, roughly digitate, approximately 0.66 times as long as posterior half of phallotheca, blunt apically; dorsal process of phallotheca absent; posterior section of phallotheca, in lateral view, broad basally, tapering towards apex, directed posterad, apex narrowly rounded (Fig. 7 D); in dorsal view, with a deep notch mesally (Fig. 7 E); phallic spine elongate, stout, spine-like structure, without basal curvature, straight (Fig. 7 D); in dorsal view, slightly pointed apically (Fig. 7 E).</p><p>Distribution.</p><p>Peru: Cajamarca Department.</p><p>Etymology.</p><p>This new species is named in honor of the Peruvian poet Marco Antonio Corcuera Díaz, who heralded the protection of Bosques de Cachil, where the type series was collected.</p></div>	https://treatment.plazi.org/id/F5F70BF4D0015CDD8CF81F7B000625A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
0EDB7B8C72615FB99824A246DB506B00.text	0EDB7B8C72615FB99824A246DB506B00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche refulioae Rázuri-Gonzales & Holzenthal 2025	<div><p>Atopsyche refulioae sp. nov.</p><p>Figs 10, 11</p><p>Type material.</p><p>Holotype. Peru • 1 ♂; Pasco, Yanachaga-Chemillén NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.3578&amp;materialsCitation.latitude=-10.5452" title="Search Plazi for locations around (long -75.3578/lat -10.5452)">Quebrada San Alberto at Refugio El Cedro</a>; 10.5452°S, 75.3578°W, 2421 m a. s. l.; 27 Aug. 2015; E. Rázuri, L. Figueroa and B. Portuguez leg.; light trap; UMSP 000220104 (MUSM) . Paratypes. Peru • 1 ♂ 2 ♀; same data as the holotype (MUSM) • 1 ♂ 2 ♀; same data as the holotype (UMSP) .</p><p>Diagnosis.</p><p>Atopsyche refulioae is related to species included in the A. batesi species group due to the unpaired dorsobasal process of the phallic apparatus. Among these species, A. refulioae is most similar to A. callosa Navás, 1924 (Colombia, Costa Rica, Ecuador, Panama, Peru, Venezuela), A. majada Ross, 1947 (Belize, Costa Rica, Guatemala, Honduras, Nicaragua, Mexico, Panama), A. minimajada Blahnik &amp; Gottschalk, 1997 (Costa Rica), and A. puharcocha Schmid, 1989 (Bolivia, Ecuador, Peru) based on the shape of segment IX in lateral view and the presence of dorsal process on the phallic apparatus. Atopsyche refulioae can easily be distinguished from these species based on the shape of the first segment of the inferior appendage with the posteroventral corner produced into a point, which is produced into a digitate process of varying length in the other species. Also, the claw-like shape of the parapod in dorsal view (elongate and sinuous in lateral view) sets the new species apart from the other species, which have shorter and wider parapods.</p><p>Description.</p><p>Adult. Forewing length: male (7.25 mm, n = 2). Body pale brown, wings dark brown in the area delimited by the costal vein, the bifurcation of the R 1, the base of the M vein, and the vein that closes the discal cell, and along the apical margin, otherwise pale brown. Forewing with erect setae on veins forming irregular pattern of alternate dark brown and yellow setae, with dark brown setae along costal margin; apex of wing with fringe of brown and yellow setae. Wing venation as in Fig. 10 A, B. Terga III and IV with oval glands, lined internally with spines (glands on tergum IV are smaller) (Fig. 10 C); process on sternum VI longer than its segment, slightly curved, bearing fine setae on its basal half increasing in thickness towards its apex, last few setae very prominent and spine-like; process on sternum VII short, less than half the length of its segment, straight, bare.</p><p>Male genitalia (Fig. 11). Segment IX, in lateral view, quadrangular, much higher than long, dorsal margin very short, with setae on posterolateral and ventral surfaces (Fig. 11 A). Parapod in lateral view, broad basally, narrow apically, sinuous, mesally with a spinose acute process and a dorsal, rounded projection bearing long setae, apex acute and upturned, bearing three long setae (Fig. 11 A); in dorsal view, broad basally with transverse carina, mesal margin concave and lateral margin biconcave, mesal rounded projection, spinose acute process, and acute apex directed laterad (Fig. 11 B). Parapod, in lateral view, elongate, sinuous, broader basally, tapering into an acute, upturned apex, mesally with a spinose subacute process and a dorsal rounded projection bearing long setae, with short setae apically (Fig. 11 A); in dorsal view, elongate, sinuous, mesal processes directed laterad, most basal process rounded and bearing long setae, process at mid-length acute and bearing spine-like setae, acute apically. Filipod digitate, longer than parapods, with a patch of setae apically (Fig. 11 A). Preanal appendage short, rounded, setose (Fig. 11 A). First segment of inferior appendage, in lateral view, trapezoidal, ventral margin slightly concave, dorsal margin slightly convex subapically, posteroventral corner produced into a slightly upturned point, with setae on lateral surface and along ventral margin (Fig. 11 A); in ventral view, C-shaped, setose, lateral margin convex, mesal margin sinuous (this margin differs between the left and right sides) (Fig. 11 C); second segment of inferior appendage, in lateral view, finger-like, narrow at mid-length, slightly inflated apically, setose, dorsal margin convex, ventral margin concave, apex truncate (Fig. 11 A); in ventral view, digitate, slightly curved mesad, apex rounded (Fig. 11 C). Proctiger, in lateral view, narrow basally, wider apically, with a carina laterodorsally, ventral margin membranous basally, with long setae on apicodorsal corner and along carina, apex truncate (Fig. 11 A). Phallic apparatus complex; phallotheca broadly rounded basally, phallic apodeme indiscernible; with ventral process of the phallotheca articulating with inferior appendages; ventrolateral branches of the phallotheca absent, dorsal process of the phallotheca present, single, elongate, bearing short spines apically, apex acute, with ventrolateral flanges fused to ventral process of the phallotheca; posterior section of the phallotheca, in lateral view, same width throughout its length, directed slightly posteroventrad, apex covered with short setae, lateral surface produced into a large flap, produced laterad, apex rounded (Fig. 11 D); in dorsal view, with a narrow notch mesally, apex directed posterad (Fig. 11 E); phallic spine elongate, stout, with a slight ventral curvature near base, then straight (Fig. 11 D); in dorsal view, apex acute (Fig. 11 E).</p><p>Distribution.</p><p>Peru: Pasco Department.</p><p>Etymology.</p><p>The first author would like to dedicate this new species to Sonia Refulio for her support throughout the years.</p></div>	https://treatment.plazi.org/id/0EDB7B8C72615FB99824A246DB506B00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
50F5A396B209535080E56DEA8BCE81BB.text	50F5A396B209535080E56DEA8BCE81BB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche sofiae Rázuri-Gonzales & Holzenthal 2025	<div><p>Atopsyche sofiae sp. nov.</p><p>Figs 12, 13</p><p>Type material.</p><p>Holotype. Peru • 1 ♂; La Libertad, Provincia Bolivar, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.5376&amp;materialsCitation.latitude=-7.6049" title="Search Plazi for locations around (long -77.5376/lat -7.6049)">Laguna Quishuar</a>; 7.6049°S, 77.5376°W, 3482 m a. s. l.; 31 Mar. 2011; C. Carranza leg.; light trap; MUSM -ENT-320966 (MUSM) . Paratype. Peru • 1 ♂; same data as the holotype (UMSP) .</p><p>Diagnosis.</p><p>Atopsyche sofiae also belongs to the bicolorata species group of Schmid (1989), characterized by having short inferior appendages and a broad notch apically on the first segment of these appendages. The second segment is reduced and inserted at the posterodorsal corner or in the notch of the first segment. Among the species in this group, the new species most closely resembles A. yupanqui Schmid, 1989 (Venezuela). In both species, the first segment of the inferior appendage is rectangular, with the posterodorsal and posteroventral corners slightly projecting mesad rather than posterad, and the second segment is inserted in the notch formed by these corners. In A. sofiae, however, these corners are more pronounced, forming a concavity where the strongly reduced second segment is inserted (not visible in lateral view). Additionally, the posterodorsal corner is densely covered with peg-like setae and the posteroventral corner bears thick, spine-like setae (features absent in A. yupanqui). Though both species have a quite simple phallic apparatus, the ventrolateral branch of the phallotheca in A. sofiae curves ventrad, while in A. yupanqui it is directed posterad. Finally, the parapods are slender in both species, but in A. sofiae, the parapods taper to an acute apex and bear a pair a peg-like setae basally. In contrast, the parapods in A. yupanqui do not taper, have a notched apex, and bear no peg-like setae.</p><p>Description.</p><p>Adult. Forewing length: male (13.5 mm, n = 2). Body and wings pale brown. Forewings with erect setae on veins without distinct pattern, with dark brown setae along costal margin, wing apex with fringe of brown setae. Wing venation as in Fig. 12. Sternum III and IV without glands; sternum V with a pair of long, membranous glands; processes on sternum VI and VII strongly reduced; process on sternum VI rounded in ventral view, with four spine-like setae, process on sternum VI triangular in ventral view, with densely clumped group of 8 spine-like setae with blunt apices (this process is absent in the paratype).</p><p>Male genitalia (Fig. 13). Segment IX, in lateral view, quadrangular, almost as high as long, posteroventral margin produced and blunt (Fig. 13 A). Parapod, in lateral view, elongate, digitate, slightly broader basally, tapering towards apex, directed slightly ventrad, with a group of setae apically and pair of peg-like setae basolaterally, apex narrowly rounded (Fig. 13 A); in dorsal view, flattened, lateral margin slightly sinuous, mesal margin almost straight, with a group of setae apically and peg-like setae on lateral flange, apex acute (Fig. 13 B). Filipod digitate, longer than parapods, setose (Fig. 13 A). Preanal appendage short, rounded, setose (Fig. 13 A). First segment of inferior appendage, in lateral view, rectangular, ventral margin almost straight, slightly concave basally (left side on paratype much more convex), dorsal margin slightly convex mesally, posteroventral and posterodorsal corner produced mesad forming a socket where the second segment is inserted (left posteroventral corner on paratype directed more ventromesad), posterodorsal corner densely covered with short, peg-like setae, posteroventral corner with long, spine-like setae, with setae on dorsal, ventral, and apical margins and lateral surface (Fig. 13 A); in ventral view, reniform, setose, lateral margin slightly convex, mesal margin convex with apical truncate projection (Fig. 13 C); second segment of inferior appendage, in lateral view, reduced, rounded (Fig. 13 A); in ventral view, reniform, slightly curved mesad, apex truncate (Fig. 13 C). Proctiger, in lateral view, narrow basally, slightly wider apically, with a long diagonal carina, slightly membranous, apex truncate (Fig. 13 A). Phallic apparatus complex; phallotheca broadly rounded basally, phallic apodeme indiscernible; with ventral process articulating with inferior appendages, narrow, same width throughout its length; ventrolateral branches of phallotheca present, hooked, ~ 0.25 × as long as posterior half of phallotheca, curved ventrad, acute apically; dorsal process of phallotheca absent; posterior section of phallotheca, in lateral view, broad basally, tapering towards apex, with a faint ridge on lateral margin, directed slightly posteroventrad, apex narrowly rounded (Fig. 13 D); in dorsal view, with a deep notch mesally (Fig. 13 E); phallic spine elongate, stout, spine-like structure, with slight curvature near base, then slightly sinuous (Fig. 13 D); in dorsal view, apex acute (Fig. 13 E).</p><p>Distribution.</p><p>Peru: La Libertad Department.</p><p>Etymology.</p><p>This species is named after Sofia Carranza Pereyra, daughter of the collector of the type specimens.</p></div>	https://treatment.plazi.org/id/50F5A396B209535080E56DEA8BCE81BB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
E0838A9086245782A93D0A7AF5A316F7.text	E0838A9086245782A93D0A7AF5A316F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopsyche yanachaga Rázuri-Gonzales & Holzenthal 2025	<div><p>Atopsyche yanachaga sp. nov.</p><p>Figs 14, 15</p><p>Type material.</p><p>Holotype. Peru • 1 ♂; Pasco, Yanachaga-Chemillén NP, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.3578&amp;materialsCitation.latitude=-10.5452" title="Search Plazi for locations around (long -75.3578/lat -10.5452)">Quebrada San Alberto at Refugio El Cedro</a>; 10.5452°S, 75.3578°W, 2421 m a. s. l.; 07 May 2011; C. Carranza, M. Alvarado, L. Figueroa leg.; light trap; MUSM -ENT-320971 (MUSM) . Paratypes. Peru • 1 ♂; same data as the holotype, but 12 Nov. 2010; C. Carranza, J. Peralta leg. (MUSM) • 1 ♂; same data as the holotype, but 13 Jun. 2010; E. Rázuri, C. Carranza leg. (UMSP) .</p><p>Diagnosis.</p><p>Atopsyche yanachaga is the second species related to the species traditionally included in the subgenus Dolochorema . It is most similar to A. major Schmid, 1989 (Bolivia). In lateral view, both species have an inferior appendage slightly longer than high, a low and quadrate segment IX, and relatively narrow, apically rounded parapods. However, certain details differ in these structures. The posteroventral margin of the first segment of the inferior appendage is more pronounced in the new species, whereas in A. major, the posterodorsal margin is more pronounced. Additionally, the second segment of the inferior appendage is longer. Furthermore, in the new species, the anterior margin of segment IX is shorter than the posterior margin, whereas both margins are of equal size in A. major . Finally, the parapods of the new species is slightly bent downwards at the base and have a sinuous dorsal margin, whereas those of A. major are straight, with a sharp point and a rounded indentation on the apical half of the dorsal margin.</p><p>Description.</p><p>Adult. Forewing length: male (9.5 mm, n = 3). Body and wings brown. Forewing with scattered brown and straw-colored setae along major longitudinal veins; wing membrane with brown and straw-colored setae. Wing venation as in Fig. 14. Sterna III and IV without glands; sternum V with a pair of long, membranous glands; process on sternum VI slightly longer than its segment, straight, bearing fine setae on its basal half and spine-like setae on its apical half, process on sternum VII very short, subtriangular, bearing spine-like setae.</p><p>Male genitalia. Segment IX, in lateral view, quadrangular, posterior margin almost twice the length of the anterior margin, both margins almost straight (Fig. 15 A). Parapod, in lateral view, elongate, sausage-shaped, slightly broader mesally and apically, slightly bent ventrad, dorsal surface with a few setae and 5 peg-like setae mesally, apex rounded (Fig. 15 A); in dorsal view, elongate, slightly flattened, lateral margin sinuous with acute projections basally and subapically, mesal margin straight, setae on dorsal surface and peg-like setae mesally, apex subacute (Fig. 15 B). Filipod long, slender, with setae of different size along length, apex rounded (Fig. 15 A). Preanal appendage short, rounded, setose (Fig. 15 A). First segment of inferior appendage, in lateral view, quadrangular, ventral margin slightly convex mesally, dorsal margin convex subapically (forming a broad, bilobed process in ventral view), posterior margin rounded, with setae on margins and lateral surface, mesal surface with a group of ~ 8 long, thick spine-like setae basally and three short, spine-like setae mesally (only 2 spine-like setae on paratype) (Fig. 15 A); in ventral view, subtriangular, setose, lateral margin convex, mesal margin sinuous (Fig. 15 C); second segment of inferior appendage, in lateral view, triangular, with a few minute setae apically, dorsal margin straight, ventral margin convex, apex narrowly rounded (Fig. 15 A); in ventral view, thumb-shaped, apex rounded (Fig. 15 C). Proctiger, in lateral view, narrow basally, slightly wider apically, membranous, apex truncate (Fig. 15 A). Phallic apparatus complex; phallotheca broadly rounded basally, phallic apodeme indiscernible; with ventral process articulating with inferior appendages, narrow, same width throughout its length; ventrolateral branches of phallotheca present, leaf-shaped, as long as posterior half of phallotheca, acute apically; dorsal process of phallotheca absent; posterior section of phallotheca, in lateral view, broad basally, tapering towards apex, directed slightly posteroventrad, apex rounded, capitate (Fig. 15 D); in dorsal view, with a shallow notch mesally (Fig. 15 E); phallic spine elongate, stout, spine-like structure, with a strong convex curvature near base, then slightly sinuous (Fig. 15 D); in dorsal view, apex acute (Fig. 15 E).</p><p>Distribution.</p><p>Peru: Pasco Department.</p><p>Etymology.</p><p>Atopsyche yanachaga is named after the Yanachaga-Chemillén National Park, where the type locality is situated. Yanachaga means black bump in Quechua, referring to the dark appearance of these mountains during sunsets on sunny days.</p></div>	https://treatment.plazi.org/id/E0838A9086245782A93D0A7AF5A316F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Rázuri-Gonzales, Ernesto;Holzenthal, Ralph W.	Rázuri-Gonzales, Ernesto, Holzenthal, Ralph W. (2025): The genus Atopsyche (Trichoptera, Hydrobiosidae) in Peru, with the description of seven new species. ZooKeys 1263: 441-478, DOI: 10.3897/zookeys.1263.150396
