identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
35E889A30718542E95AE1918E9FEC1C0.text	35E889A30718542E95AE1918E9FEC1C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calamaria similis Qi & Nguyen & Yang & Xu & David & Shi & Liu & Rong & Korolev & Poyarkov & Wang 2026	<div><p>Calamaria similis sp. nov.</p><p>Tables 4, 5; Figs 4, 5, 6, 7, 8, 9</p><p>Chresonymy.</p><p>Calamaria pavimentata – Yang and Zheng (2018, in part); Lee (2021, in part); Yeung et al. (2022; in part); Cai et al. (2023: in part); Poyarkov et al. (2023: in part); Liang et al. (2024, in part); Zhang et al. (2025 a, 2025 b; in part); Lalremsanga et al. (2026, in part).</p><p>Calamaria pavimentata pavimentata – David et al. (2023: in part).</p><p>Type material.</p><p>Holotype: China • 1 ♂; Guangdong Province, Yangjiang City, Yangchun City, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=111.41305&amp;materialsCitation.latitude=22.070618" title="Search Plazi for locations around (long 111.41305/lat 22.070618)">Shuangjiao Town</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=111.41305&amp;materialsCitation.latitude=22.070618" title="Search Plazi for locations around (long 111.41305/lat 22.070618)">Qifendong Village [now Dahe Village]</a>; 22.07061848°N, 111.41304597°E; altitude 44 m a. sl.; collected by JJL on 12 August 2017; SYS r 001816 . Paratypes: China • 1 subadult ♀; Guangxi ZAR, Chongzuo City, Longzhou County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.97141&amp;materialsCitation.latitude=22.461645" title="Search Plazi for locations around (long 106.97141/lat 22.461645)">Nonggang National Nature Reserve</a>; 22.46164593°N, 106.97141398°E; altitude 350 m a. s. l.; collected by CZR on 24 May 2017; SYS r 001725 • 1 subadult ♂; Guangxi ZAR, Chongzuo City, Longzhou County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.97141&amp;materialsCitation.latitude=22.461645" title="Search Plazi for locations around (long 106.97141/lat 22.461645)">Nonggang National Nature Reserve</a>; 22.46164593°N, 106.97141398°E; altitude 350 m a. s. l.; collected by CZR on 24 May 2017; SYS r 001726 • 1 subadult ♂; Guangxi ZAR, Chongzuo City, Longzhou County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.02262&amp;materialsCitation.latitude=22.317844" title="Search Plazi for locations around (long 107.02262/lat 22.317844)">Shuikou Town</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.02262&amp;materialsCitation.latitude=22.317844" title="Search Plazi for locations around (long 107.02262/lat 22.317844)">Shangjin Village</a>; 22.317845°N, 107.022622°E; altitude ca. 300 m a. s. l.; collected by JHY in 2013; KFBG 14507 .</p><p>Diagnosis.</p><p>Calamaria similis sp. nov. is distinguished from all other congeners by the combination of the following morphological characters: Maxillary teeth eight, unmodified; rostral higher than wide; prefrontal shorter than frontal and contacting the first two supralabials; mental not in contact with anterior chin shields; dorsal scales in 13–13 – 13 rows, smooth throughout; a single preocular and postocular; four supralabials, with the 2 nd and 3 rd contacting the eye; five infralabials; six scales surrounding the paraparietal; 145–168 ventrals (145–155 in males, 168 in female); 15–23 paired subcaudals (20–23 in males, 15 in female); a relatively short tail (6.4–9.3 % of total length in males, 6.1 % in the female), thick and nearly cylindrical, gradually tapering to an obtuse point. Dorsal coloration dark brown to blackish brown, the body marked with five longitudinal rows of dark-edged scales, forming indistinct, discontinuous stripes of variable clarity along the body; two nuchal collars are present, the first is a larger, dark brown collar, and the second is smaller and without pattern; two pairs of light spots are present on the dorsal tail, variably developed and sometimes indistinct. The ventral surface yellowish white, immaculate; underside of tail lacking a broad median black stripe extending posteriorly to the tail tip.</p><p>Description of the holotype, SYS r 001816, adult male (see Figs 4, 5 A – C).</p><p>Specimen in good condition, slightly discolored due to preservation after approximately nine years in preservative fluid (Fig. 4). Body slender, cylindrical (SVL 170.0 mm); tail short, thinner than body, thick and nearly cylindrical, gradually tapering to an obtuse point (TaL 17.4 mm, TaL / TL 9.3 %). Head small, elliptical in dorsal view (head length 6.1 mm; head width 3.5 mm; head height 2.4 mm). Eye small, round (eye diameter 0.8 mm), smaller than eye-nostril distance 1.1 mm but larger than eye-mouth distance 0.5 mm.</p><p>Head scalation. Rostral broader than high (rostral width 1.2 mm, rostral height 0.9 mm); dorsal portion visible from above approximately half the length of the prefrontal suture. Prefrontal shorter than frontal, not entering orbit, in contact with the first and second supralabials. Frontal hexagonal, longer than wide (frontal length 1.8 mm, frontal width 1.4 mm). Temporals absent. Paraparietal surrounded by six scales. One preocular on each side, higher than wide, slightly taller than postocular but shorter than eye diameter. One postocular on each side, also higher than wide. Nasals divided, small, bordered by rostral, prefrontal, and 1 st supralabial. Four supralabials on each side, the 2 nd and 3 rd entering the orbit; the 4 th largest in height (relative size: 4&gt; 2&gt; 3&gt; 1), but the 2 nd one, rather low, is as long as the 4 th. Five infralabials on each side, the first three pairs contacting anterior chin shields; the 1 st pair meeting at the midline; the 4 th largest. Anterior chin shields much longer than wide, slightly pentagonal, meet completely along their longest sides. Posterior chin shields are shorter, in contact anteriorly, and separated by the preventral at approximately one-third of their length.</p><p>Body scalation. Dorsal scale rows 13–13 – 13 throughout body, all smooth. Ventrals 150 (+ 1 preventral); subcaudals 21, all paired; dorsocaudal reduction from 9 to 8 occurring posterior to subcaudal 3, from 8 to 6 occurring posterior to subcaudal 4, and from 6 to 5 occurring posterior to subcaudal 15; Cloacal plate undivided.</p><p>Coloration of the holotype. In life (Fig. 5 A – C), the dorsal surface of head is reddish brown, with blackish pigment forming a cap-like marking dorsally; the lateral surfaces of the head are slightly paler than the dorsum, with blackish pigment forming a wavy longitudinal mask extending from the rostral region to the nape. The ventral surface of the head is pale yellow, with irregular dark brown markings restricted to the mental, the infralabials, and anterior chin-shields. The dorsal surface of the body is reddish brown. Two nuchal collars are present, the first is a distinct dark brown collar, slightly shorter than the head length and extending laterally onto the sides of the neck; the second is smaller, without pattern, and approximately one to two dorsal scales in length. Five longitudinal series of interrupted dark brown stripes are visible along the body, extending from behind the posteriormost collar to the tip of the tail. The margins of the dorsal scales are heavily suffused with black pigment, forming an almost reticulate pattern. The ventrals are pale yellow, with dark brown blotches along the outermost lateral margins, forming a dashed line parallel to the longitudinal stripes. The tail dorsum is slightly darker than the body dorsum, and the longitudinal stripes fade posteriorly. Two pairs of small, faint yellowish-brown spots are present on the dorsal tail, one pair at the base and one posteriorly. Subcaudal surface pale yellow, generally immaculate.</p><p>In preservative (Fig. 4), the dorsal surface of the head, body, and tail fade to brown. Two nuchal collars remain in strong contrast. The dark brown longitudinal stripes become much more diffuse, whereas the reticulate pattern formed by the heavily pigmented margins of the dorsal scales become more conspicuous. The pale spots on the tail turn light yellowish brown. The ventral surfaces of the head, body, and tail are uniformly pale yellowish white.</p><p>Skull morphology of holotype. (Fig. 6) The skull is subcylindrical in general shape. In dorsal view, the anterior margin of the premaxilla is flat, the nasals and septomaxillae are relatively strong and laterally expanded, the conchal processes of the septomaxillae are broad, flattened, and well-developed. The frontals are approximately right-triangle-shaped with their anterior tip rounded. The parietal is scutiform, longer than wide, while the supraoccipital is subpentagonal. The prootic and exoccipitals are both well developed and similar in size, forming the widest part of the braincase.</p><p>In ventral view, the vomers are well-developed with their anterior tips nearly reach the vomerine processes of premaxilla. The maxillary processes of palatines are triangular and laterally expanded while the choanal processes degenerated, almost invisible. The pterygoid is slender and lanceolate in shape, twice the length of the palatine. The basisphenoid and basioccipital lack a ventral process. The parasphenoid rostrum of the basisphenoid is sharp and triangular shaped. The basioccipital is relatively flat and laterally expanded.</p><p>In lateral view, the ascending process of premaxilla is slender and greatly reduced. The conchal process of the septomaxilla is anteroposteriorly expanded, reaching the lateral edge of nasals. The prefrontal and postorbital are similar in height, the bottom tip of the postorbital reaches the top margin of the maxilla. The ectopterygoid is biforked, the labial furcula of ectopterygoid is oval, and distinct from the lingual process. The lingual process projects anterodorsally, overlaps the dorsal margin ectopterygoid process of the maxilla, visible in lateral view but not visible in ventral view.</p><p>The mandible is slender and moderately curved. The prearticular crest of the compound bone is prominent while the surangular crest is absent.</p><p>Dentition of the holotype. (Fig. 6) The maxillary teeth are conical, laterally compressed, and posteriorly recurved at the tip, consistent with the description of unmodified maxillary teeth in Inger and Marx (1965). The morphology of palatine teeth, pterygoid teeth, and dentary teeth show no significant specialization relative to the maxillary teeth. Maxillary teeth 8, palatine teeth 5, pterygoid teeth 11, dentary teeth 7 on each side.</p><p>Variation.</p><p>(Figs 7, 8, 9) Variation in measurements and scalation of the type series is summarized in Table 4. Sexual dimorphism is evident in several characters. The female possesses significantly more ventral scales than males (VEN 168 vs 145–155), but fewer subcaudal scales (SC 15 in female vs 21–23 in males). Tail proportions also differ markedly between sexes: males have relatively longer tails (TaL / TL ratio 6.4–9.3 %), whereas the female exhibits a shorter relative tail length (TaL / TL ratio 6.1 %). Coloration shows moderate intraspecific variation. The dorsal ground color varies from reddish brown to dark brown, with five dark longitudinal stripes running along the body. Supralabials and infralabials are cream to yellowish white, upper portions of supralabials occasionally with scattered dark pigmentation. Ventral coloration is uniform, ranging from yellowish white to pale yellow, without markings. The subcaudal region is similar in color to the ventrals or slightly darker and lacks a dark longitudinal stripe on the underside of the tail. The tail tip varies in pigmentation, being uniformly dark, distinctly darker than the remainder of the tail, or only weakly pigmented.</p><p>Sources: 1 = Duméril et al. (1854); 2 = Fan (1931); 3 = this study.</p><p>Etymology.</p><p>The specific epithet similis is a Latin adjective meaning similar, referring to the strong morphological resemblance of this species to Calamaria pavimentata . This similarity has resulted in the species being previously misidentified and treated as C. pavimentata in numerous earlier studies (e. g., Yang and Zheng 2018). We propose the following common names for the new species: “ Similar Reed Snake ” (English), “ 拟尖尾两头蛇 ” (nĬ jiān wěi liăng tóu shé, Chinese), “ R ắn mai g ầm tương đ ồng ” (Vietnamese), and “ Сходная карликовая змея ” (Skhodnaya karlikovaya zmeya, Russian).</p><p>Comparison.</p><p>Comparative morphological data for the new species and currently recognized members of the genus Calamaria from mainland China and the Indo-Burma Region are presented in Table 5. Calamaria similis sp. nov. differs from all congeners, particularly those referred to the Calamaria pavimentata species complex, by a unique combination of meristic characters, body proportions, and coloration, as detailed below:</p><p>Calamaria similis sp. nov. is readily distinguished from C. lumbricoidea (distributed in Thailand, Malaysia, Singapore, Indonesia, Brunei, and the Philippines) and C. schlegeli (Thailand, Malaysia, Indonesia, and Singapore) by having four supralabials, with the second and third in contact with the eye (vs five supralabials, with the third and fourth in contact). Furthermore, both species are restricted to areas south of the Isthmus of Kra in Peninsular Malaysia and have not been recorded from mainland Indochina.</p><p>Calamaria similis sp. nov. also differs from C. lovii ingermarxorum Darevsky &amp; Orlov, 1992 (Gia Lai Province, Vietnam), C. nebulosa (Phongsaly Province, Laos), C. thanhi Ziegler &amp; Le, 2005 (Quang Tri Province, Vietnam), and C. yunnanensis (Yunnan Province, China; possibly Dien Bien Province, Vietnam) by presence of a preocular scale (vs absent). It further differs from C. buchi Marx &amp; Inger, 1955 (Lam Dong Province, Vietnam), C. concolor Orlov, Nguyen, Nguyen, Ananjeva &amp; Ho, 2010 (Hue and Da Nang cities, Vietnam), C. gialaiensis Ziegler, Nguyen &amp; Nguyen, 2008 (Gia Lai Province, Vietnam), C. lovii ingermarxorum, C. lumbricoidea, and C. sangi Nguyen, Koch &amp; Ziegler, 2009 (Quang Ngai Province, Vietnam) in having the mental not contacting the chin shields (vs contacting).</p><p>Calamaria similis sp. nov. differs from C. abramovi Orlov, 2009 (Quang Ngai Province, Vietnam) by having slightly fewer ventral scales in females (VEN 168 vs 174), fewer subcaudal scales in both sexes (SC 20–23 in males and 15 in females vs 26 in males and 20 in females), a slightly shorter relative tail length in males (TaL / TL 6.4–9.3 % vs 13.3 %), a blackish-brown dorsal coloration with narrow dark longitudinal stripes (vs uniformly black), and presence of a nuchal collar without a pattern (vs absent).</p><p>Compared with Calamaria andersoni (Yunnan Province, China), Calamaria similis sp. nov. has lower ventral scale counts in both sexes (VEN 145–155 in males and 168 in females vs 164–172 in males and 186 in females), a smaller total length in both sexes (maximum TL 187 mm in males and 154 mm in females vs 351 mm in males and 312 mm in females).</p><p>It is distinct from Calamaria dominici Ziegler, Tran &amp; Nguyen, 2019 (Lam Dong Province, Vietnam) in having lower ventral scale counts in females (168 vs 174), slightly fewer subcaudal scales in females (15 vs 17–18), a smaller total length in females (maximum TL 154 mm vs 421 mm), a blackish-brown dorsal coloration with narrow dark longitudinal stripes (vs dark with irregular yellow blotches), and presence of patternless nuchal collar (vs absent).</p><p>It differs from Calamaria septentrionalis (southern China and northern Vietnam) in having more subcaudal scales in both sexes (SC 22–23 in males and 15 in females vs 15–19 in males and 6–11 in females), a smaller total length in both sexes (maximum TL 187 mm in males and 154 mm in females vs 344 mm in males and 384 mm in females), a longer relative tail length in females (TaL / TL 6.1 % vs 2.6–4.3 %), and an obtusely pointed tail tip (vs broadly rounded).</p><p>Compared with Calamaria strigiventris Poyarkov, Nguyen, Orlov &amp; Vogel, 2019 (Lam Dong and Khanh Hoa provinces, Vietnam), Calamaria similis sp. nov. differs by having lower ventral scale counts in females (VEN 168 vs 176–183), fewer subcaudal scales in both sexes (SC 20–23 in males and 15 in females vs 29–31 in males and 20–22 in females), a smaller total length in both sexes (maximum TL 187 mm in males and 154 mm in females vs 362 mm in males and 367 mm in females), a shorter relative tail length in both sexes (TaL / TL 6.4–9.3 % in males and 6.1 % in females vs 11.2–17.9 % in males and 8.4–8.6 % in females), presence of patternless nuchal collar (vs absent), presence of two pairs of light spots on the dorsal tail (vs absent).</p><p>From members of the Calamaria pavimentata species complex: Calamaria similis sp. nov. differs from C. annamensis (Quang Tri Province, Vietnam) by having lower ventral scale counts in both sexes (VEN 145–155 in males and 168 in females vs 182 in males and 196–204 in females), a smaller total length in both sexes (maximum TL 187 mm in males and 154 mm in females vs 304 mm in males and 552 mm in females), a slightly longer relative tail length in females (TaL / TL 6.1 % vs 3.9–4.7 %), yellowish-white, immaculate ventrals (vs light orange to salmon with small black spots).</p><p>It is distinguished from Calamaria arcana (Guangdong, Fujian, and Hunan provinces, possibly occurring in Zhejiang Province, China) by having lower ventral scale counts in both sexes (VEN 145–155 in males and 168 in females vs 170–176 in males and 192 in females), slightly higher subcaudal counts in females (SC 15 vs 12), a smaller total length in males (maximum TL 187 mm vs 304 mm), a slightly longer relative tail length in females (TaL / TL 6.1 % vs 4.7 %), fewer maxillary teeth (MT 8 vs 10).</p><p>Calamaria similis sp. nov. further differs from C. berezowskii (Sichuan Province, China) by having a smaller total length in both sexes (maximum TL 187 mm in males and 154 mm in females vs 290 mm in males and 245 mm in females); a broad and distinct nuchal collar (vs narrow and indistinct); and presence of two pairs of light spots on the dorsal tail (vs absent).</p><p>It can be distinguished from Calamaria jinggangensis (Jiangxi, Guizhou, and possibly Hunan provinces, China) by having slightly lower ventral scale counts in both sexes (VEN 145–155 in males and 168 in females vs 157–158 in males and 179 in females), slightly higher subcaudal counts in females (SC 15 vs 12), a shorter relative tail length in males (TaL / TL 6.4–9.3 % vs 15 %) but a longer relative tail length in females (TaL / TL 6.1 % vs 3.6 %).</p><p>Calamaria similis sp. nov. differs from C. mizoramensis (Mizoram State, India and Bangladesh) by having a smaller total length in both sexes (maximum TL 187 mm in males and 154 mm in females vs 251 mm in males and 258 mm in females), a shorter relative tail length in males (TaL / TL 6.4–9.3 % vs 10.3–13.1 %), and presence of two pairs of light spots on the dorsal tail (vs absent).</p><p>Calamaria similis sp. nov. differs from C. synergis (Yunnan Province, China) by having six scales surrounding the paraparietal (vs five), lower ventral scale counts in males (VEN 145–155 vs 161–166), a slightly smaller maximum total length in males (maximum TL 187 mm vs 284 mm), and presence of two pairs of light spots on the dorsal tail (vs absent).</p><p>Calamaria similis sp. nov. differs from C. pfefferi Stejneger, 1901 (Okinawa Island, Japan) by having higher ventral scale counts in females (VEN 168 vs 158–165), fewer subcaudal scales in males (SC 20–23 vs 24–26), presence of a nuchal collar without pattern (vs absent), and presence of two pairs of light spots on the dorsal tail (vs absent) (Stejneger 1907; Maki 1931; Yoshigo et al. 2003; Nakamura et al. 2025).</p><p>Calamaria similis sp. nov. differs from C. pavimentata sensu stricto (Java Island, Indonesia; data based on the combined morphology of the holotypes of C. pavimentata and its junior synonym C. quadrimaculata following Inger and Marx 1965) by having higher ventral scale counts in females (VEN 168 vs 136), fewer subcaudal scales in males (SC 20–23 vs 27) but slightly higher subcaudal counts in females (SC 15 vs 13), a smaller total length in males (maximum TL 187 mm vs 206 mm), and a shorter relative tail length in both sexes (TaL / TL 6.4–9.3 % in males and 6.1 % in females vs 11.6 % in males and 8.0 % in females). Historical illustrations of the holotypes of C. pavimentata and its junior synonym C. quadrimaculata (Jan and Sordelli 1860–1866) indicate presence of a midventral longitudinal stripe on the tail in C. pavimentata and subcaudal blotches in C. quadrimaculata (Appendix 1); however, such markings appear to be variable within members of the C. pavimentata species complex and may also occur in the new species, and therefore were not used as diagnostic characters here.</p><p>Lastly, Calamaria similis sp. nov. differs from all five other junior synonyms and subspecies of C. pavimentata . Among these, it can be distinguished from Calamaria siamensis Günther, 1864 by having a lower maximum ventral scale count (VEN 168 vs 190) and by presence of nuchal collar with no pattern (vs absent) (Günther, 1864). It further differs from Calamaria pavimentata var. uniformis Smith, 1921 by having lower subcaudal scale counts (SC 20–23 in males and 15 in females vs 30–34 in males and 18–19 in females), and by presence of two pairs of light spots on the dorsal tail (vs absent) (Smith 1921). From Calamaria pavimentata formosana Maki, 1931, it differs by having slightly lower ventral scale counts in males (VEN 145–155 vs 153–178 a smaller maximum total length (max TL 187 mm in males and 154 mm in females vs 280 mm in males and 310 mm in females), and presence of two pairs of light spots on the dorsal tail (vs absent) (Stejneger 1907; Maki 1931). From Calamaria pavimentata banaensis Bourret, 1934, it differs by having lower ventral scale counts in both sexes (VEN 145–155 in males and 168 in female vs 157–160 in males and 178–179 in females), lacking a dark midventral stripe on the belly (vs present), and by presence of two pairs of light spots on the dorsal tail (Bourret 1934). Finally, Calamaria similis sp. nov. differs from Calamaria pavimentata miyarai Takara, 1962 by having lower ventral scale counts in males (VEN 145–155 vs 157–174), fewer subcaudal scales in females (SC 15 vs 17–19), a smaller maximum total length (187 mm in males and 154 mm in females vs 290 mm in males and 368 mm in females), a shorter relative tail length in females (TaL / TL 6.1 % vs 7.4–7.6 %), and presence of two pairs of light spots on the dorsal tail (Takara 1962; Ota 1982; Ota and Hokama 1996).</p><p>Distribution, natural history notes, and conservation status.</p><p>Calamaria similis sp. nov. had long been misidentified as Calamaria pavimentata . However, due to the lack of available molecular material, its confirmed distribution is currently restricted to Yangchun City, Yangjiang City, Guangdong Province, and Longzhou County, Chongzuo City, Guangxi ZAR, China. All specimens were collected during the daytime. The holotype (SYS r 001816) was found beneath a thick leaf-litter layer in a hilly landscape, whereas two paratypes (SYS r 001725 and SYS r 001726) were encountered along a roadside in limestone karst forest. Given its semi-fossorial habits, the species is currently known from only a few collected specimens from Guangdong Province and Guangxi ZAR, China, and a small number of photographic records. Notably, the straight-line distance between the confirmed localities in Guangdong Province and Guangxi ZAR exceeds 450 km, encompassing extensive areas that remain unsurveyed. Virtually nothing is known about its population size, ecology, or precise distributional limits. Given this paucity of information, we recommend that Calamaria similis sp. nov. be provisionally assessed as Data Deficient (DD) under the IUCN Red List criteria (IUCN Standards and Petitions Subcommittee 2025).</p></div>	https://treatment.plazi.org/id/35E889A30718542E95AE1918E9FEC1C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Qi, Shuo;Nguyen, Tan Van;Yang, Jian-Huan;Xu, Yu-Hao;David, Patrick;Shi, Jing-Song;Liu, Jing-Jian;Rong, Can-Zhong;Korolev, Alexey M.;Poyarkov, Nikolay A.;Wang, Ying-Yong	Qi, Shuo, Nguyen, Tan Van, Yang, Jian-Huan, Xu, Yu-Hao, David, Patrick, Shi, Jing-Song, Liu, Jing-Jian, Rong, Can-Zhong, Korolev, Alexey M., Poyarkov, Nikolay A., Wang, Ying-Yong (2026): A new species of Calamaria (Squamata, Calamariidae) from southern China, previously confused with Calamaria pavimentata Duméril, Bibron & Duméril, 1854. ZooKeys 1277: 245-280, DOI: 10.3897/zookeys.1277.187107
