taxonID	type	description	language	source
096987E3E443FF996D6BFD18FCA6FC80.taxon	type_taxon	Type species Chthonius terribilis With, 1906, by original designation.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E443FF936D40FCD1FC39F9D8.taxon	description	urn: lsid: zoobank. org: act: E 7931503 - 1 D 3 B- 490 F-B 3 BA- 9 E 810 FCE 1470 Figs 1 H, 2 – 6	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E443FF936D40FCD1FC39F9D8.taxon	diagnosis	Diagnosis (♂ ♀) Moderately sized troglomorphic species with elongated appendages; four eyes, anterior pair of eyes well-developed, posterior pair with flat lenses; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse; posterior margin of carapace with 2 setae; tergites I – II each with four setae. Pedipalps slender, femur 5.31 (♂), 5.14 (♀) × as long as broad, length 0.69 (♂), 0.72 (♀); chela 5.94 (♂), 5.21 (♀) × as long as broad, length 0.95 (♂), 0.99 (♀); both chelal fingers with intercalary teeth; sb closer to st than b.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E443FF936D40FCD1FC39F9D8.taxon	etymology	Etymology Named after the closest city to the type locality, Baise City. Vernacular name ÞDzƌ 伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E443FF936D40FCD1FC39F9D8.taxon	materials_examined	Type material Holotype CHINA • ♂; Guangxi, Baise City, Lingyun County, Xiaoluo Town, Dayan Cave; 24.35539 ° N, 106.708811 ° E; 735 m a. s. l.; 26 Jul. 2021; Zegang Feng, Hongru Xu, Liu Fu and Nana Zhan leg.; MHBU Ps. - MHBU-GX 2021072601. Paratypes CHINA • ♀; same data as for holotype; MHBU, Ps. - MHBU-GX 2021072602.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E443FF936D40FCD1FC39F9D8.taxon	description	Description Adult male (holotype and male paratypes; Figs 2 – 6) COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale. CEPHALOTHORAX (Figs 3 D – E, 4 A). Carapace nearly subquadrate, 1.09 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, rounded and obtuse; four eyes, anterior pair of eyes well-developed, posterior pair with flat lenses; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to the setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with 8 – 11 terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 4 D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5. CHELICERA (Figs 3 C, 4 B). Almost as long as carapace, 2.19 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13 teeth, distal one largest; movable finger with 13 contiguous small teeth; galea completely vestigial (Fig. 4 B). Serrula exterior with 20 and serrula interior with 13 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 4 C). PEDIPALP (Figs 3 A – B, F, 5 A – B). Trochanter 1.69, femur 5.31, patella 2.29, chela 5.94, hand 1.94 × as long as broad; femur 2.16 × as long as patella; movable chelal finger 2.13 × as long as hand and 0.69 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb slightly closer to st than to b; b and t situated subdistally, it slightly distal to t and t distal to b; est situated proximal to t and close to it (Figs 4 A, 5 A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 4 B, 5 B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 28 well-spaced, and pointed teeth and 25 intercalary microdenticles; movable finger with 11 well-spaced, pointed teeth, plus 11 intercalary microdenticles and 20 vestigial, rounded and contiguous basal teeth. OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 6: 5: 7: 8: 9: 10: 7: 5: T 2 T: 0. Sternal chaetotaxy IV – XII: 12: 12: 12: 12: 14: 14: 12: -: 2. Genital region: sternite II with nine setae scattered on median area, genital opening slit-like, sternite III with row of 14 setae (Fig. 3 G). LEGS (Fig. 5 C – D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.94 × as long as patella; tarsus 1.95 × as long as tibia. Leg IV: femoropatella 2.71 × as long as deep; tibia 4.67 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 2.71 × as long as deep (TS = 0.33), telotarsus 10.50 × as long as deep and 2.21 × as long as basitarsus (TS = 0.26). Setae of leg I (trochanter to tibia) 4: 10 – 11: 7: 10, setae of leg IV (trochanter to basitarsus) 2: 3: 6: 8: 7. Arolium not divided, slightly shorter than simple claws. Adult female (paratype; Figs 2 B, 3 H) Mostly same as males; tergal chaetotaxy I – XII: 4: 4: 5: 5: 6: 7: 7: 8: 9: 8: T 2 T: 0; sternal chaetotaxy IV – XII: 12: 12: 11: 15: 13: 12: 12: -: 2. Genital region: sternite II with nine setae scattered on median area, sternite III with row of 12 setae. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses) Male Body length 1.75. Pedipalps: trochanter 0.22 / 0.13 (1.69), femur 0.69 / 0.13 (5.31), patella 0.32 / 0.14 (2.29), chela 0.95 / 0.16 (5.94), hand 0.31 / 0.16 (1.94), movable chelal finger length 0.66. Chelicera 0.46 / 0.21 (2.19), movable finger length 0.26. Carapace 0.49 / 0.45 (1.09). Leg I: trochanter 0.12 / 0.09 (1.33), femur 0.35 / 0.07 (5.00), patella 0.18 / 0.07 (2.57), tibia 0.20 / 0.05 (4.00), tarsus 0.39 / 0.04 (9.75). Leg IV: trochanter 0.18 / 0.12 (1.50), femoropatella 0.57 / 0.21 (2.71), tibia 0.42 / 0.09 (4.67), basitarsus 0.19 / 0.07 (2.71), telotarsus 0.42 / 0.04 (10.50). Female Body length 1.73. Pedipalps: trochanter 0.26 / 0.14 (1.86), femur 0.72 / 0.14 (5.14), patella 0.34 / 0.16 (2.13), chela 0.99 / 0.19 (5.21), hand 0.34 / 0.19 (1.79), movable chelal finger length 0.66. Chelicera 0.52 / 0.25 (2.08), movable finger length 0.30. Carapace 0.46 / 0.58 (0.79). Leg I: trochanter 0.15 / 0.11 (1.36), femur 0.38 / 0.07 (5.43), patella 0.20 / 0.07 (2.86), tibia 0.22 / 0.05 (4.40), tarsus 0.42 / 0.05 (8.40). Leg IV: trochanter 0.21 / 0.13 (1.61), femoropatella 0.62 / 0.23 (2.70), tibia 0.42 / 0.10 (4.20), basitarsus 0.19 / 0.07 (2.71), telotarsus 0.41 / 0.04 (10.25).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E443FF936D40FCD1FC39F9D8.taxon	discussion	Remarks	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E443FF936D40FCD1FC39F9D8.taxon	biology_ecology	Ecology The specimen was collected in the completely dark zone near the cave entrance (Fig. 6).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E443FF936D40FCD1FC39F9D8.taxon	distribution	Distribution Known only from the type locality, Dayan Cave (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E449FF906EB3F919FD9FFDFD.taxon	description	Fig. 1 C	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E449FF906EB3F919FD9FFDFD.taxon	diagnosis	Diagnosis and description For details see Gao et al. (2018). Vernacular name Ṅ 形ƌ伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E449FF906EB3F919FD9FFDFD.taxon	materials_examined	Material examined Holotype CHINA • ♂; Guangxi, Guilin City, Xiufeng District, Maomaotou Cave, deep zone; 25.312811 ° N, 110.270178 ° E; 225 m a. s. l.; 15 Nov. 2016; J. J. Wynne leg.; MHBU, Ps. - MHBU-GX 16121502. Paratypes CHINA • 1 ♂, 1 ♀; same data as for holotype; MHBU, Ps. - MHBU-GX 16121503 – 04.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E449FF906EB3F919FD9FFDFD.taxon	distribution	Distribution Maomaotou Cave (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	description	urn: lsid: zoobank. org: act: 6 C 147713 - 251 A- 45 B 7 - 82 E 0 - 8 A 3 D 3 C 0 C 636 B Figs 1 L, 7 – 11	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	diagnosis	Diagnosis (♂ ♀) Moderately sized troglomorphic species with elongated appendages; four eyes, anterior pair of eyes strongly reduced, posterior pair with eyespots; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse; posterior margin of carapace with 2 setae; tergites I – II each with four setae. Pedipalps slender, femur 5.40 (♂), 4.83 – 5.50 (♀) × as long as broad, length 0.54 (♂), 0.55 – 0.58 (♀); chela 6.08 (♂), 5.13 – 5.19 (♀) × as long as broad, length 0.79 (♂), 0.82 – 0.83 (♀); both chelal fingers with intercalary teeth; sb midway between st and b.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	etymology	Etymology Named after the type locality, Fenghuang Township. Vernacular name Ṇdzƌ 伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	materials_examined	Type material Holotype CHINA • ♂; Guangxi, Hechi City, Bama Yao Autonomous County, Fenghuang Township, Unnamed Cave 3; 24.1936872 ° N, 107.390445 ° E; 467.6 m a. s. l.; 28 Jul. 2021; Zegang Feng, Hongru Xu, Liu Fu and Nana Zhan leg.; MHBU, Ps. - MHBU-GX 2021072801. Paratypes CHINA • 2 ♀♀; same data as for holotype; MHBU, Ps. - MHBU-GX 2021072802 – 03.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	description	Description Adult male (Figs 7 – 11) COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale. CEPHALOTHORAX (Figs 8 D – E, 9 A). Carapace nearly subquadrate, 1.18 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, rounded and obtuse; four eyes, anterior pair of eyes strongly reduced, posterior pair with eyespots; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with seven terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 9 D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5. CHELICERA (Figs 8 C, 9 B). Almost as long as carapace, 2.10 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13 teeth, distal one largest; movable finger with 12 contiguous small teeth; galea completely vestigial (Fig. 8 C). Serrula exterior with 19 and serrula interior with 12 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 9 C). PEDIPALP (Figs 8 A – B, F, 9 E, 10 A – B). Trochanter 1.82, femur 5.40, patella 2.00, chela 6.08, hand 1.92 × as long as broad; femur 2.25 × as long as patella; movable chelal finger 2.20 × as long as hand and 0.70 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, it slightly distal to t and t distal to est; est situated distal to b (Figs 8 A, 10 A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 8 B, 10 B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 24 well-spaced, and pointed teeth and 22 intercalary microdenticles; movable finger with 10 well-spaced, pointed teeth, plus 9 intercalary microdenticles and 13 vestigial, rounded and contiguous basal teeth. OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 5: 7: 7: 8: 8: 8: 8: 7: T 2 T: 0. Sternal chaetotaxy IV – XII: 12: 13: 12: 12: 13: 13: 11: -: 2. Genital region: sternite II with 11 setae scattered on median area, genital opening slit-like, sternite III with row of 16 setae (Fig. 8 G). LEGS (Fig. 10 C – D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.94 × as long as patella; tarsus 1.95 × as long as tibia. Leg IV: femoropatella 2.71 × as long as deep; tibia 4.67 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 2.71 × as long as deep (TS = 0.33), telotarsus 10.50 × as long as deep and 2.21 × as long as basitarsus (TS = 0.26). Setae of leg I (trochanter to tibia) 4: 10 – 11: 7: 10, setae of leg IV (trochanter to basitarsus) 2: 3: 6: 8: 7. Arolium not divided, slightly shorter than simple claws. Adult females (paratypes; Figs 7 B, 8 H) Mostly same as males; tergal chaetotaxy I – XII: 4: 4: 5: 7 – 8: 8: 8 – 9: 8 – 10: 10 – 12: 8 – 9: 5 – 7: T 2 T: 0; sternal chaetotaxy IV – XII: 11 – 12: 12 – 14: 11 – 12: 12 – 13: 13 – 16: 13: 9 – 12: -: 2. Genital region: sternite II with 10 setae scattered on median area, sternite III with row of 12 setae. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses) Male Body length 1.40. Pedipalps: trochanter 0.20 / 0.11 (1.82), femur 0.54 / 0.10 (5.40), patella 0.24 / 0.12 (2.00), chela 0.79 / 0.13 (6.08), hand 0.25 / 0.13 (1.94), movable chelal finger length 0.55. Chelicera 0.40 / 0.19 (2.10), movable finger length 0.23. Carapace 0.38 / 0.45 (0.84). Leg I: trochanter 0.12 / 0.09 (1.33), femur 0.29 / 0.05 (5.80), patella 0.14 / 0.05 (2.80), tibia 0.17 / 0.04 (4.25), tarsus 0.32 / 0.04 (8.00). Leg IV: trochanter 0.17 / 0.11 (1.54), femoropatella 0.48 / 0.19 (2.52), tibia 0.32 / 0.08 (4.00), basitarsus 0.14 / 0.06 (2.33), telotarsus 0.34 / 0.03 (11.33). Females Body length 1.42 – 1.45. Pedipalps: trochanter 0.20 – 0.21 / 0.11 (1.82 – 1.91), femur 0.55 – 0.58 / 0.10 – 0.12 (4.83 – 5.50), patella 0.27 / 0.12 (2.25), chela 0.82 – 0.83 / 0.16 (5.13 – 5.19), hand 0.26 – 0.27 / 0.16 (1.63 – 1.69), movable chelal finger length 0.57. Chelicera 0.43 – 0.45 / 0.22 (1.95 – 2.04), movable finger length 0.24 – 0.25. Carapace 0.39 – 0.48 / 0.48 – 0.49 (0.81 – 0.98). Leg I: trochanter 0.11 – 0.13 / 0.08 – 0.10 (1.30 – 1.38), femur 0.29 – 0.31 / 0.06 (4.83 – 5.17), patella 0.15 – 0.17 / 0.05 – 0.06 (2.83 – 3.00), tibia 0.16 – 0.17 / 0.04 (4.00 – 4.25), tarsus 0.34 – 0.35 / 0.04 (8.50 – 8.75). Leg IV: trochanter 0.13 – 0.18 / 0.10 – 0.11 (1.30 – 1.64), femoropatella 0.48 – 0.49 / 0.17 – 0.18 (2.72 – 2.82), tibia 0.32 – 0.35 / 0.08 (4.00 – 4.38), basitarsus 0.15 – 0.16 / 0.06 (2.50 – 2.67), telotarsus 0.34 – 0.35 / 0.03 – 0.04 (8.50 – 11.67).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	discussion	Remarks Tyrannochthonius fenghuangensis sp. nov. differs from all other hypogean species of the genus from China except Tyrannochthonius duo by the presence of four eyes, anterior pair of eyes strongly reduced, posterior pair with eyespots. It differs from T. duo by its shorter and thicker pedipalpal chela (length 0.79 mm (♂), 0.82 – 0.83 mm (♀), 6.08 (♂), 5.13 – 5.19 (♀) × as long as broad compared to length 1.11 mm (♂), 1.17 – 1.18 mm (♀), 6.94 (♂), 5.85 – 6.56 (♀) × as long as broad in T. duo), shorter and thicker pedipalpal femur (length 0.54 mm (♂), 0.55 – 0.58 mm (♀), 5.40 (♂), 4.83 – 5.50 (♀) × as long as broad compared to length 0.76 mm (♂), 0.79 – 0.81 mm (♀), 5.85 (♂), 5.79 – 6.08 (♀) × as long as broad in T. duo), the shape of epistome (epistome small, rounded and obtuse vs epistome small, triangular in T. duo) and the position of trichobothrium (sb midway between st and b compared to sb closer to b than to st in T. duo) (Hou et al. 2023 b).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	distribution	Distribution Known only from the type locality, Unnamed Cave 3 (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E451FF8C6D40FE97FC31FA81.taxon	description	urn: lsid: zoobank. org: act: D 9943 FDC-E 713 - 4225 - 8 EF 0 - 3 C 0 D 533 F 082 D Figs 1 I, 12 – 15	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E451FF8C6D40FE97FC31FA81.taxon	diagnosis	Diagnosis (♂ ♀) Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, triangular; posterior margin of carapace with 2 setae; tergites I – IV each with four setae. Pedipalps slender, femur 7.13 (♂), 6.75 (♀) × as long as broad, length 1.07 (♂), 1.08 (♀); chela 7.14 (♂), 6.58 (♀) × as long as broad, length 1.50 (♂), 1.58 (♀); chelal fingers without intercalary teeth and movable chelal finger teeth markedly retrorse and almost contiguous; sb slightly closer to st than to b.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E451FF8C6D40FE97FC31FA81.taxon	etymology	Etymology Named after the type locality, Laqiu Cave. Vernacular name ® 丘ƌ伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E451FF8C6D40FE97FC31FA81.taxon	materials_examined	Type material Holotype CHINA • ♂; Guangxi, Hechi City, Donglan County, Wuzuan Town, Laqiu Village, Laqiu Cave; 24.358433 ° N, 107.300395 ° E; 467 m a. s. l.; 11 May. 2018; Zegang Feng and Weitong Wang leg.; MHBU, Ps. - MHBU-GX 18051101. Paratype CHINA • ♀; same data as for holotype; MHBU, Ps. - MHBU-GX 18051102.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E451FF8C6D40FE97FC31FA81.taxon	description	Description Adult male (Figs 12 – 15) COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale. CEPHALOTHORAX (Figs 13 D, 14 A). Carapace nearly subquadrate, 1.02 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, triangular; without eyes or eyespots; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with nine terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 14 D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5. CHELICERA (Figs 13 C, 14 B). Almost as long as carapace, 2.44 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12 teeth, distal one largest; movable finger with 10 contiguous small teeth; galea completely vestigial (Fig. 13 C). Serrula exterior with 28 and serrula interior with 14 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 14 C). PEDIPALP (Figs 13 A – B, F, 14 E, 15 A – B). Trochanter 1.75, femur 7.13, patella 2.33, chela 7.14, hand 2.43 × as long as broad; femur 2.55 × as long as patella; movable chelal finger 1.88 × as long as hand and 0.64 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb slightly closer to st than to b; b and t situated subdistally, it slightly distal to t and t distal to est; est situated at same level as t, and distal to b (Figs 13 A, 15 A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 13 B, 15 B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 24 well-spaced, pointed and slightly retrorse macrodenticles; movable finger with 36 markedly retrorse and almost contiguous pointed teeth. OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 5: 5: 5: 4: T 2 T: 0. Sternal chaetotaxy IV – XII: 12: 8: 8: 10: 9: 9: 8: -: 2. Genital region: sternite II with nine setae scattered on median area, genital opening slit-like, sternite III with row of 14 setae (Fig. 13 F). LEGS (Fig. 15 C – D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.82 × as long as patella; tarsus 2.11 × as long as tibia. Leg IV: femoropatella 3.96 × as long as deep; tibia 6.60 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 4.00 × as long as deep (TS = 0.34), telotarsus 15.40 × as long as deep and 2.75 × as long as basitarsus (TS = 0.28). Setae of leg I (trochanter to tibia) 4: 17: 12: 13, setae of leg IV (trochanter to basitarsus) 1: 4: 7: 11: 14. Arolium not divided, slightly shorter than simple claws. Adult females (paratypes; Figs 12 B, 13 G) Mostly same as males; tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4: 5: 6: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 12: 10: 9: 9: 7: 10: 10: -: 2. Genital region: sternite II with 10 setae scattered on median area, sternite III with row of 12 setae. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses) Male Body length 1.75. Pedipalps: trochanter 0.28 / 0.16 (1.75), femur 1.07 / 0.15 (7.13), patella 0.42 / 0.18 (2.33), chela 1.50 / 0.21 (7.14), hand 0.51 / 0.21 (2.43), movable chelal finger length 0.96. Chelicera 0.61 / 0.25 (2.44), movable finger length 0.33. Carapace 0.56 / 0.55 (1.02). Leg I: trochanter 0.19 / 0.11 (1.73), femur 0.62 / 0.08 (7.75), patella 0.34 / 0.07 (4.86), tibia 0.28 / 0.05 (5.60), tarsus 0.59 / 0.05 (11.80). Leg IV: trochanter 0.24 / 0.13 (1.85), femoropatella 0.95 / 0.24 (3.96), tibia 0.66 / 0.10 (6.60), basitarsus 0.28 / 0.07 (4.00), telotarsus 0.77 / 0.05 (15.40). Female Body length 1.76. Pedipalps: trochanter 0.29 / 0.17 (1.71), femur 1.08 / 0.16 (6.75), patella 0.40 / 0.19 (2.11), chela 1.58 / 0.24 (6.58), hand 0.53 / 0.24 (2.21), movable chelal finger length 1.02. Chelicera 0.65 / 0.25 (2.48), movable finger length 0.36. Carapace 0.64 / 0.58 (1.10). Leg I: trochanter 0.20 / 0.14 (1.43), femur 0.64 / 0.08 (8.00), patella 0.37 / 0.08 (4.63), tibia 0.30 / 0.06 (5.00), tarsus 0.70 / 0.06 (11.67). Leg IV: trochanter 0.24 / 0.14 (1.71), femoropatella 0.85 / 0.19 (4.47), tibia 0.57 / 0.10 (5.70), basitarsus 0.25 / 0.07 (3.57), telotarsus 0.73 / 0.05 (14.60).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E451FF8C6D40FE97FC31FA81.taxon	discussion	Remarks Tyrannochthonius laqiuensis sp. nov. differs from all other hypogean species of the genus from China except T. tiani and T. nanxingensis by both chelal fingers lacking intercalary teeth, the presence of four setae on tergites I – III, and carapace without eyes or eyespots. It differs from T. tiani and T. nanxingensis by its shorter and thicker pedipalpal chela (length 1.50 mm (♂), 1.58 mm (♀), 7.14 (♂), 6.58 (♀) × as long as broad compared to length 1.79 mm (♀), 9.42 (♀) × as long as broad in T. tiani and length 1.92 – 1.97 mm (♂), 1.85 mm (♀), 7.04 – 7.11 (♂), 7.12 (♀) × as long as broad in T. nanxingensis) and the position of trichobothrium (sb slightly closer to st than to b compared to sb closer to b than st in T. tiani and T. nanxingensis) (Hou et al. 2023 b; Sun et al. 2025)	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E451FF8C6D40FE97FC31FA81.taxon	distribution	Distribution Known only from the type locality, Laqiu Cave (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E456FF866D74FAD2FBF3FABF.taxon	description	urn: lsid: zoobank. org: act: 4304 E 479 - 6 BBA- 47 D 9 - B 60 C- 83 FC 270 A 6091 Figs 1 N, 16 – 20	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E456FF866D74FAD2FBF3FABF.taxon	diagnosis	Diagnosis (♀) Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed, triangular; posterior margin of carapace with 2 setae; tergites I – III each with two setae, tergite IV with four setae. Pedipalps slender, femur 7.69 (♀) × as long as broad, length 1.23 (♀); chela 8.25 (♀) × as long as broad, length 1.65 (♀); both chelal fingers with intercalary teeth; sb midway between st and b.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E456FF866D74FAD2FBF3FABF.taxon	etymology	Etymology The specific name is a noun derived from the Latin words ‘ longulus ’ and ‘ membrum ’, meaning ‘ rather long’ and ‘ member’, which refers to the elongated pedipalps and legs. Vernacular name Khƌ 伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E456FF866D74FAD2FBF3FABF.taxon	materials_examined	Type material Holotype CHINA • ♀; Guangxi, Baise City, Tiandong County, Silin Town, Guangyang Village, Guangyang Cave; 23.518475 ° N, 107.393634 ° E; 363 m a. s. l.; 6 Oct. 2019; Zegang Feng and Lingchen Zhao leg.; MHBU, Ps. - MHBU-GX 2019100601.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E456FF866D74FAD2FBF3FABF.taxon	description	Description Adult female (Figs 16 – 20) COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale. CEPHALOTHORAX (Figs 17 E, 18 A). Carapace nearly subquadrate, 0.96 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome pointed, triangular; without eyes or eyespots; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with 10 terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 18 D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5. CHELICERA (Figs 17 D, 18 B). Almost as long as carapace, 2.30 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 17 teeth, distal one largest; movable finger with 15 contiguous small teeth; galea completely vestigial (Fig. 17 D). Serrula exterior with 27 and serrula interior with 11 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 18 C). PEDIPALP (Figs 17 A – B, F, 18 E, 19 A – B). Trochanter 1.65, femur 7.69, patella 2.56, chela 8.25, hand 3.05 × as long as broad; femur 2.67 × as long as patella; movable chelal finger 1.72 × as long as hand and 0.64 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, t slightly distal to it and est distal to b (Figs 17 A, 19 A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 17 B, 19 B). Both chelal fingers with row of teeth, spaced regularly along the margin, teeth smaller distally and proximally: fixed finger with 31 well-spaced, and pointed teeth and 28 intercalary microdenticles; movable finger with 20 well-spaced, pointed teeth, plus 15 intercalary microdenticles and 15 vestigial, rounded and contiguous basal teeth. OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 2: 2: 4: 4: 4: 5: 5: 5: 4: T 2 T: 0. Sternal chaetotaxy IV – XII: 12: 8: 9: 9: 10: 9: 9: -: 2. Genital region: sternite II with 10 setae scattered on median area, genital opening slit-like, sternite III with row of 12 setae (Fig. 17 C). LEGS (Fig. 19 C – D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.91 × as long as patella; tarsus 2.00 × as long as tibia. Leg IV: femoropatella 4.30 × as long as deep; tibia 6.09 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 3.33 × as long as deep (TS = 0.33), telotarsus 15.00 × as long as deep and 2.50 × as long as basitarsus (TS = 0.33). Setae of leg I (trochanter to tibia) 3: 12: 10: 10, setae of leg IV (trochanter to basitarsus) 1: 3: 7: 10: 10. Arolium not divided, slightly shorter than the simple claws. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses) Female Body length 2.04. Pedipalps: trochanter 0.33 / 0.20 (1.65), femur 1.23 / 0.16 (7.69), patella 0.46 / 0.18 (2.56), chela 1.65 / 0.20 (8.25), hand 0.61 / 0.20 (3.05), movable chelal finger length 1.05. Chelicera 0.76 / 0.33 (2.30), movable finger length 0.40. Carapace 0.66 / 0.69 (0.96). Leg I: trochanter 0.19 / 0.15 (1.27), femur 0.67 / 0.09 (7.44), patella 0.35 / 0.08 (4.38), tibia 0.32 / 0.06 (5.33), tarsus 0.64 / 0.06 (10.67). Leg IV: trochanter 0.21 / 0.15 (1.40), femoropatella 0.99 / 0.23 (4.30), tibia 0.67 / 0.11 (6.09), basitarsus 0.30 / 0.09 (3.33), telotarsus 0.75 / 0.05 (15.00).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E456FF866D74FAD2FBF3FABF.taxon	discussion	Remarks Tyrannochthonius longulimembrum sp. nov. most closely resembles T. ganshuanensis in the presence of two setae on tergites I – II, carapace without eyes or eyespots and both chelal fingers with intercalary teeth, but differs in several characters as follows: longer and thinner pedipalpal chela (length 1.65 mm (♀), 8.25 (♀) × as long as broad compared to length 1.21 – 1.44 mm (♀), 6.90 – 7.30 (♀) × as long as broad in T. ganshuanensis) and longer and thinner pedipalpal femur (length 1.23 mm (♀), 7.69 (♀) × as long as broad compared to length 0.95 – 0.97 mm (♀), 0.95 – 0.97 (♀) × as long as broad in T. ganshuanensis) (Mahnert 2009).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E456FF866D74FAD2FBF3FABF.taxon	distribution	Distribution Known only from the type locality, Guangyang Cave (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E45CFFBB6D51FAF4FC37F887.taxon	description	urn: lsid: zoobank. org: act: 77 F 8 DD 22 - F 3 A 8 - 451 C-BCA 5 - 079 F 34901 B 51 Figs 1 G, 21 – 24	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E45CFFBB6D51FAF4FC37F887.taxon	diagnosis	Diagnosis (♀) Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I – IV each with four setae. Pedipalps slender, femur 5.50 – 5.85 (♀) × as long as broad, length 0.76 – 0.77 mm (♀); chela 5.60 – 6.00 (♀) × as long as broad, length 1.12 – 1.14 mm (♀); both chelal fingers without intercalary teeth; sb slightly closer to st than to b.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E45CFFBB6D51FAF4FC37F887.taxon	etymology	Etymology The specific name is an adjective derived from a combination of the Latin words ‘ mirus ’ and ‘ dentatus ’, meaning ‘ remarkable’ and ‘ toothed’, respectively, which refers to the distinctly heteromorphic chelal teeth. Vernacular name 异 Ṅƌ 伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E45CFFBB6D51FAF4FC37F887.taxon	materials_examined	Type material Holotype CHINA • ♀; Guangxi, Hechi City, Fengshan County, Fengcheng Town, Songren Village, Xi’an Cave; 24.565644 ° N, 107.037218 ° E; 526 m a. s. l.; 5 Oct. 2019; Zegang Feng and Lingchen Zhao leg.; MHBU, Ps. - MHBU-GX 2019100501. Paratypes CHINA • 2 ♀♀; same data as for holotype; MHBU, Ps. - MHBU-GX 2019100502 – 03.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E45CFFBB6D51FAF4FC37F887.taxon	description	Description Adult females (Figs 21 – 24) COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale. CEPHALOTHORAX (Figs 22 E, 23 A). Carapace nearly subquadrate, 0.94 – 0.96 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small and pointed, triangular; without eyes or eyespots; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with seven terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 23 D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5. CHELICERA (Figs 22 D, 23 B). Almost as long as carapace, 1.92 – 2.38 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 10 – 11 teeth, distal one largest; movable finger with 12 – 14 contiguous small teeth; galea completely vestigial (Fig. 22 D). Serrula exterior with 20 – 27 and serrula interior with 12 blades. Rallum with nine blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 23 C). PEDIPALP (Figs 22 A – B, F, 23 E, 24 A – B). Trochanter 1.64 – 1.69, femur 5.50 – 5.85, patella 1.88 – 2.00, chela 5.60 – 6.00, hand 1.90 – 2.05 × as long as broad; femur 2.41 – 2.53 × as long as patella; movable chelal finger 1.87 – 2.03 × as long as hand and 0.73 – 0.77 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb slightly closer to st than to b; b and t situated subdistally, it slightly distal to t; est situated at same level as t, and distal to b (Figs 22 A, 24 A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 22 B, 24 B). Both chelal fingers with a row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 23 – 25 well-spaced, and pointed macrodenticles; movable finger with 30 – 32 contiguous remarkly retrorse and pointed teeth, plus 5 – 6 vestigial, rounded and contiguous basal teeth. OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4 – 5: 5 – 6: 5 – 7: 4: T 2 T: 0. Sternal chaetotaxy IV – XII: 12: 9 – 12: 10 – 11: 9 – 10: 10: 10 – 11: 9 – 10: -: 2. Genital region: sternite II with 10 setae scattered on median area, genital opening slit-like, sternite III with row of 10 – 12 setae (Fig. 22 C). LEGS (Fig. 24 C – D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.61 – 1.87 × as long as patella; tarsus 2.19 – 2.47 × as long as tibia. Leg IV: femoropatella 3.45 – 3.72 × as long as deep; tibia 5.50 – 5.88 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 3.00 – 4.00 × as long as deep (TS = 0.29 – 0.33), telotarsus 10.20 – 12.25 × as long as deep and 2.40 – 2.72 × as long as basitarsus (TS = 0.26 – 0.31). Setae of leg I (trochanter to tibia) 3 – 4: 10 – 11: 10: 9 – 15, setae of leg IV (trochanter to basitarsus) 2: 3 – 4: 6 – 7: 11 – 12: 9 – 10. Arolium not divided, slightly shorter than simple claws. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses) Females Body length 1.52 – 1.92. Pedipalps: trochanter 0.22 – 0.23 / 0.13 – 0.14 (1.64 – 1.69), femur 0.76 – 0.77 / 0.13 – 0.14 (5.50 – 5.85), patella 0.30 – 0.32 / 0.15 – 0.17 (1.88 – 2.00), chela 1.12 – 1.14 / 0.19 – 0.20 (5.60 – 6.00), hand 0.38 – 0.39 / 0.19 – 0.20 (1.90 – 2.05), movable chelal finger length 0.73 – 0.77. Chelicera 0.46 – 0.50 / 0.21 – 0.24 (1.92 – 2.38), movable finger length 0.26 – 0.28. Carapace 0.46 – 0.47 / 0.49 (0.94 – 0.96). Leg I: trochanter 0.15 – 0.16 / 0.09 – 0.11 (1.36 – 1.67), femur 0.42 – 0.44 / 0.07 – 0.08 (5.50 – 6.14), patella 0.23 – 0.26 / 0.06 (3.83 – 4.33), tibia 0.19 – 0.22 / 0.05 – 0.06 (3.67 – 4.20), tarsus 0.46 – 0.49 / 0.04 – 0.05 (9.2 – 11.75). Leg IV: trochanter 0.19 – 0.21 / 0.12 – 0.14 (1.50 – 1.75), femoropatella 0.65 – 0.69 / 0.18 – 0.20 (3.45 – 3.72), tibia 0.44 – 0.47 / 0.08 (5.50 – 5.88), basitarsus 0.18 – 0.20 / 0.05 – 0.06 (3.00 – 4.00), telotarsus 0.48 – 0.51 / 0.04 – 0.05 (10.20 – 12.25).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E45CFFBB6D51FAF4FC37F887.taxon	discussion	Remarks Tyrannochthonius miridentatus sp. nov. differs from all other hypogean species of the genus except T. quattuor by the movable finger of pedipalpal chela exhibits a continuous and strongly retrorse teeth, except for the few most distal ones, sb slightly closer to st than to b, the presence of four setae on tergites I – IV, epistome triangular and carapace without eyes or eyespots. It differs from T. quattuor by its shorter and thicker pedipalpal chela (length 1.12 – 1.14 mm (♀), 5.60 – 6.00 (♀) × as long as broad compared to length 1.74 – 1.86 mm (♀), 6.89 – 7.25 (♀) × as long as broad in T. quattuor) and shorter and thicker pedipalpal femur (length 0.76 – 0.77 mm (♀), 5.50 – 5.85 (♀) × as long as broad compared to compared to length 1.24 – 1.32 mm (♀), 7.33 – 7.75 (♀) × as long as broad in T. quattuor) (Hou et al. 2023 b).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E45CFFBB6D51FAF4FC37F887.taxon	distribution	Distribution Known only from the type locality, Xi’an Cave (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB86ED6FE97FD90FBCD.taxon	description	Fig. 1 B	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB86ED6FE97FD90FBCD.taxon	diagnosis	Diagnosis and description For details see Sun et al. (2025). Vernacular name 乳洞岩ƌ伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB86ED6FE97FD90FBCD.taxon	materials_examined	Material examined Holotype CHINA • ♂; Guangxi, Guilin City, Xing’an County, Rudongyan Cave; 25.574192 ° N, 110.620256 ° E; 240 m a. s. l.; 10 Oct. 2019; Zegang Feng and Lingchen Zhao leg.; MHBU, Ps. - MHBU-GX 2019101001. Paratypes CHINA • 2 ♀♀; same data as for holotype; MHBU, Ps. - MHBU-GX 2019101002 – 03; CHINA • 3 ♀♀; Guangxi, Guilin City, Xing’an County, Rudongyan Cave; 25.570469 ° N, 110.627489 ° E; 213 m a. s. l.; 4 Feb. 2023; Xiangbo Guo, Jianzhou Sun, Tao Zheng and Songtao Shi leg.; MHBU, Ps. - MHBU-GX 2023020401 – 03.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB86ED6FE97FD90FBCD.taxon	distribution	Distribution Rudongyan Cave (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB26D58FB05FC1BFD26.taxon	description	urn: lsid: zoobank. org: act: 3 F 46460 A- 87 B 2 - 4 B 81 - B 1 B 3 - F 241 F 830565 D Figs 1 F, 25 – 29	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB26D58FB05FC1BFD26.taxon	diagnosis	Diagnosis (♂ ♀) Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, triangular; posterior margin of carapace with 2 setae; tergites I – II each with two setae, tergites III – IV each with four setae. Pedipalps slender, femur 6.36 (♂), 6.00 – 6.17 (♀) × as long as broad, length 0.70 (♂), 0.66 – 0.74 (♀); chela 6.71 (♂), 6.71 – 6.80 (♀) × as long as broad, length 0.0.94 (♂), 0.94 – 1.02 (♀); both chelal fingers with intercalary teeth; sb closer to st than b.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB26D58FB05FC1BFD26.taxon	etymology	Etymology Named after the type locality, Shenlong Cave. Vernacular name DZAEƌ 伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB26D58FB05FC1BFD26.taxon	materials_examined	Type material Holotype CHINA • ♂; Guangxi, Hechi City, Huanjiang Maonan Autonomous County, Dacai Township, Shenlong Cave; 24.773162 ° N, 108.385965 ° E; 201.2 m a. s. l.; 31 Jul. 2021; Zegang Feng, Hongru Xu, Liu Fu and Nana Zhan leg.; MHBU, Ps. - MHBU-GX 2021073101. Paratypes CHINA • 2 ♀♀; same data as for holotype; MHBU, Ps. - MHBU-GX 2021073102 – 03.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB26D58FB05FC1BFD26.taxon	description	Description Adult male (Figs 25 – 29) COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale. CEPHALOTHORAX (Figs 26 D, 27 A). Carapace nearly subquadrate, 1.05 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, triangular; carapace without eyes or eyespots; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to the sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with 8 – 10 terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 27 D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5. CHELICERA (Figs 26 C, 27 B). Almost as long as carapace, 2.44 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13 teeth, distal one largest; movable finger with 13 contiguous small teeth; galea completely vestigial (Fig. 26 C). Serrula exterior with 23 and serrula interior with 13 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 27 C). PEDIPALP (Figs 26 A – B, E, 27 E, 28 A – B). Trochanter 1.64, femur 6.36, patella 2.17, chela 6.71, hand 2.50 × as long as broad; femur 2.69 × as long as patella; movable chelal finger 1.74 × as long as hand and 0.65 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb closer to st than b; b and t situated subdistally, t slightly distal to it and est slightly distal to b (Figs 26 A, 28 A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 26 B, 28 B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 32 well-spaced, slightly retrorse and pointed teeth and 19 intercalary microdenticles; movable finger with 24 almost contiguous, slightly retrorse and pointed teeth, plus nine intercalary microdenticles and six vestigial, rounded and contiguous basal teeth. OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 2: 4: 4: 4: 4: 5: 5: 5: 4: T 2 T: 0. Sternal chaetotaxy IV – XII: 12: 7: 7: 9: 8: 9: 9: -: 2. Genital region: sternite II with nine setae scattered on median area, genital opening slit-like, sternite III with row of 16 setae (Fig. 26 G). LEGS (Fig. 28 C – D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.94 × as long as patella; tarsus 2.11 × as long as tibia. Leg IV: femoropatella 2.94 × as long as deep; tibia 4.63 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 2.83 × as long as deep (TS = 0.21), telotarsus 14.00 × as long as deep and 2.47 × as long as basitarsus (TS = 0.33). Setae of leg I (trochanter to tibia) 4: 11: 11: 11, setae of leg IV (trochanter to basitarsus) 3: 3: 6: 10: 8. Arolium not divided, slightly shorter than simple claws. Adult females (paratypes; Figs 25 B, 26 G) Mostly same as males; tergal chaetotaxy I – XII: 2: 2: 4: 4: 4: 4: 4: 5: 5: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 11 – 12: 7 – 8: 7 – 8: 8: 8 – 9: 9: 9: -: 2. Genital region: sternite II with nine setae scattered on median area, sternite III with row of 10 – 12 setae. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses) Male Body length 1.40. Pedipalps: trochanter 0.20 / 0.11 (1.82), femur 0.54 / 0.10 (5.40), patella 0.24 / 0.12 (2.00), chela 0.79 / 0.13 (6.08), hand 0.25 / 0.13 (1.94), movable chelal finger length 0.55. Chelicera 0.40 / 0.19 (2.10), movable finger length 0.23. Carapace 0.38 / 0.45 (0.84). Leg I: trochanter 0.12 / 0.09 (1.33), femur 0.29 / 0.05 (5.80), patella 0.14 / 0.05 (2.80), tibia 0.17 / 0.04 (4.25), tarsus 0.32 / 0.04 (8.00). Leg IV: trochanter 0.17 / 0.11 (1.54), femoropatella 0.48 / 0.19 (2.52), tibia 0.32 / 0.08 (4.00), basitarsus 0.14 / 0.06 (2.33), telotarsus 0.34 / 0.03 (11.33). Females Body length 1.42 – 1.45. Pedipalps: trochanter 0.20 – 0.21 / 0.11 (1.82 – 1.91), femur 0.55 – 0.58 / 0.10 – 0.12 (4.83 – 5.50), patella 0.27 / 0.12 (2.25), chela 0.82 – 0.83 / 0.16 (5.13 – 5.19), hand 0.26 – 0.27 / 0.16 (1.63 – 1.69), movable chelal finger length 0.57. Chelicera 0.43 – 0.45 / 0.22 (1.95 – 2.04), movable finger length 0.24 – 0.25. Carapace 0.39 – 0.48 / 0.48 – 0.49 (0.81 – 0.98). Leg I: trochanter 0.11 – 0.13 / 0.08 – 0.10 (1.30 – 1.38), femur 0.29 – 0.31 / 0.06 (4.83 – 5.17), patella 0.15 – 0.17 / 0.05 – 0.06 (2.83 – 3.00), tibia 0.16 – 0.17 / 0.04 (4.00 – 4.25), tarsus 0.34 – 0.35 / 0.04 (8.50 – 8.75). Leg IV: trochanter 0.13 – 0.18 / 0.10 – 0.11 (1.30 – 1.64), femoropatella 0.48 – 0.49 / 0.17 – 0.18 (2.72 – 2.82), tibia 0.32 – 0.35 / 0.08 (4.00 – 4.38), basitarsus 0.15 – 0.16 / 0.06 (2.50 – 2.67), telotarsus 0.34 – 0.35 / 0.03 – 0.04 (8.50 – 11.67).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB26D58FB05FC1BFD26.taxon	discussion	Remarks Tyrannochthonius shenlongensis sp. nov. most closely resembles T. akaelus in the both chelal fingers with intercalary teeth, tergites I – II each with two setae, tergites III – IV each with four setae, tergite X with four setae; epistome small, triangular and carapace without eyes or eyespots, but differs in several characters as follows: shorter and thicker pedipalpal chela (length 0.94 – 1.02 mm (♀), 6.71 – 6.80 (♀) × as long as broad compared to length 1.28 mm (♀), 7.53 (♀) × as long as broad in T. akaelus) and shorter and thicker pedipalpal femur (length 0.66 – 0.74 mm (♀), 6.00 – 6.17 (♀) × as long as broad compared to length 0.90 mm (♀), 6.42 (♀) × as long as broad in T. akaelus) and the position of trichobothrium (sb closer to st than b compared to sb situated midway between st and b in T. akaelus) (Mahnert 2009).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB26D58FB05FC1BFD26.taxon	distribution	Distribution Known only from the type locality, Shenlong Cave (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB36E8AFE97FDB4FC20.taxon	description	Fig. 1 D	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB36E8AFE97FDB4FC20.taxon	diagnosis	Diagnosis and description For details see Sun et al. (2025). Vernacular name 田氏ƌ伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB36E8AFE97FDB4FC20.taxon	materials_examined	Material examined Holotype CHINA • ♀; Guangxi, Hechi City, Huanjiang County, The Mulun National Nature Reserve, Ganxiao Cave; 25.18312 ° N, 108.03124 ° E; 695 m a. s. l.; 8 Dec. 2023; Mingyi Tian leg.; MHBU, Ps. - MHBU-GX 2023120801.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB36E8AFE97FDB4FC20.taxon	distribution	Distribution Ganxiao Cave (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB56D55FC70FC23FDB1.taxon	description	urn: lsid: zoobank. org: act: 714 F 40 F 5 - E 81 B- 4 AC 8 - 89 BF- 8 C 6191 CC 42 DA Figs 1 K, 30 – 34	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB56D55FC70FC23FDB1.taxon	diagnosis	Diagnosis (♀) Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, triangular; posterior margin of carapace with 2 setae; tergites I – II each with four setae, tergites III – IV each with more four setae. Pedipalps slender, femur 4.89 (♀) × as long as broad, length 0.44 (♀); chela 5.25 (♀) × as long as broad, length 0.63 (♀); both chelal fingers without intercalary teeth; sb closer to st than b.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB56D55FC70FC23FDB1.taxon	etymology	Etymology Named after the type locality, Xiangu Cave. Vernacular name 仙Rƌ伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB56D55FC70FC23FDB1.taxon	materials_examined	Type material Holotype CHINA • ♀; Guangxi, Hechi City, Yizhou District, Xiayan Village, Xiangu Cave; 24.501836 ° N, 108.713608 ° E; 124 m a. s. l.; 5 Oct. 2019; Zegang Feng and Lingchen Zhao leg.; MHBU, Ps. - MHBU-GX 2019100501.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB56D55FC70FC23FDB1.taxon	description	Description Adult female (Figs 30 – 34) COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale. CEPHALOTHORAX (Figs 31 E, 32 A). Carapace nearly subquadrate, 0.94 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, triangular; without eyes or eyespots; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with a rounded anteromedial process; coxae II with six or eight terminally indented coxal spines, set as oblique and arched row, central spines slightly longer than others (Fig. 32 D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5. CHELICERA (Figs 31 D, 32 B). Almost as long as carapace, 1.83 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13 teeth, distal one largest; movable finger with nine contiguous small teeth; galea completely vestigial (Fig. 31 D). Serrula exterior with 18 and serrula interior with 12 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 32 C). PEDIPALP (Figs 31 A – B, F, 32 E, 33 A – B). Trochanter 1.75, femur 4.89, patella 2.11, chela 5.25, hand 1.67 × as long as broad; femur 2.32 × as long as patella; movable chelal finger 2.25 × as long as hand and 0.71 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb closer to st than b; b and t situated subdistally, t slightly distal to it and est distal to b (Figs 31 A, 33 A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 31 B, 33 B). Both chelal fingers with row of teeth, spaced regularly along the margin, teeth smaller distally and proximally: fixed finger with 25 well-spaced, slightly retrorse and pointed teeth; movable finger with 11 well-spaced, pointed teeth, plus 12 vestigial, rounded and contiguous basal teeth. OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 5: 6: 6: 5: 6: 5: 5: 4: T 2 T: 0. Sternal chaetotaxy IV – XII: 10: 11: 9: 9: 8: 9: 7: -: 2. Genital region: sternite II with 10 setae scattered on median area, genital opening slit-like, sternite III with row of 12 setae (Fig. 31 C). LEGS (Fig. 33 C – D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.92 × as long as patella; tarsus 2.08 × as long as tibia. Leg IV: femoropatella 2.42 × as long as deep; tibia 4.20 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 3.00 × as long as deep (TS = 0.32), telotarsus 8.00 × as long as deep and 2.00 × as long as basitarsus (TS = 0.29). Setae of leg I (trochanter to tibia) 2: 10: 9: 11, setae of leg IV (trochanter to basitarsus) 2: 3: 6: 11: 10. Arolium not divided, slightly shorter than simple claws. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses) Female Body length 1.04. Pedipalps: trochanter 0.14 / 0.08 (1.75), femur 0.44 / 0.09 (4.89), patella 0.19 / 0.09 (2.11), chela 0.63 / 0.12 (5.25), hand 0.20 / 0.12 (1.67), movable chelal finger length 0.45. Chelicera 0.33 / 0.18 (1.83), movable finger length 0.18. Carapace 0.32 / 0.34 (0.94). Leg I: trochanter 0.09 / 0.07 (1.29), femur 0.23 / 0.04 (5.75), patella 0.12 / 0.04 (3.00), tibia 0.12 / 0.03 (4.00), tarsus 0.25 / 0.03 (8.33). Leg IV: trochanter 0.11 / 0.07 (1.57), femoropatella 0.29 / 0.12 (2.42), tibia 0.29 / 0.12 (2.42), basitarsus 0.12 / 0.04 (3.00), telotarsus 0.24 / 0.03 (8.00).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB56D55FC70FC23FDB1.taxon	discussion	Remarks Tyrannochthonius xianguensis sp. nov. most closely resembles T. xinzhaiensis by the tergites I – II each with four setae, both chelal fingers without intercalary teeth, trichobothrium sb closer to st than b and carapace without eyes or eyespots, but differs in several characters as follows: shorter and thicker pedipalpal chela (length 0.63 mm (♀), 5.25 (♀) × as long as broad compared to length 1.61 – 1.64 mm (♀), 6.42 – 6.44 (♀) × as long as broad in T. xinzhaiensis) and shorter and thicker pedipalpal femur (length 0.44 mm (♀), 4.89 (♀) × as long as broad compared to length 1.14 – 1.18 mm (♀), 6.71 – 6.77 (♀) × as long as broad in T. xinzhaiensis) and tergites III – IV each with more four setae (vs tergites III – IV each with four setae in T. xinzhaiensis) (Li 2022).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB56D55FC70FC23FDB1.taxon	distribution	Distribution Known only from the type locality, Xiangu Cave (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	description	urn: lsid: zoobank. org: act: 732 B 6785 - C 1 B 6 - 4 F 71 - B 9 D 1 - 526839238 FCB Figs 1 J, 35 – 39	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	diagnosis	Diagnosis (♂ ♀) Moderately sized troglomorphic species with elongated appendages; four eyes, anterior pair of eyes reduced, posterior pair of eyespots; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse; posterior margin of carapace with 2 setae; tergites I – IV each with four setae. Pedipalps slender, femur 6.53 – 6.80 (♂), 6.75 – 7.27 (♀) × as long as broad, length 0.98 – 1.02 (♂), 1.08 – 1.09 (♀); chela 7.21 – 7.33 (♂), 6.77 – 6.91 (♀) × as long as broad, length 1.32 – 1.37 (♂), 1.45 – 1.49 (♀); both chelal fingers with intercalary teeth; sb midway between st and b.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	etymology	Etymology Named after the type locality, Xiannvyan Cave. Vernacular name 仙kƌ伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	materials_examined	Type material Holotype CHINA • ♂; Guangxi, Hechi City, Yizhou District, Qingyuan Town, Yankou Village, Xiannvyan Cave; 24.488011 ° N, 108.567428 ° E; 150 m a. s. l.; 4 Oct. 2019; Zegang Feng and Lingchen Zhao leg.; MHBU, Ps. - MHBU-GX 2019100401. Paratypes CHINA • 1 ♂, 2 ♀♀; same data as for holotype; MHBU, Ps. - MHBU-GX 2019100402 – 04.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	description	Description Adult male (Figs 35 – 39) COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale. CEPHALOTHORAX (Figs 36 D, 37 A). Carapace nearly subquadrate, 1.04 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, rounded and obtuse; four eyes, anterior pair of eyes reduced, posterior pair of eyespots; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to the setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with a rounded anteromedial process; coxae II with 12 terminally indented coxal spines on each side, set as an oblique and arched row, central spines slightly longer than others (Fig. 37 D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5. CHELICERA (Figs 36 C, 37 B). Almost as long as carapace, 1.79 – 1.86 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 9 – 12 teeth, distal one largest; movable finger with 10 – 15 contiguous small teeth; galea completely vestigial (Fig. 36 C). Serrula exterior with 20 – 22 and serrula interior with 12 – 13 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 37 C). PEDIPALP (Figs 36 A – B, F, 37 E, 38 A – B). Trochanter 1.63 – 1.81, femur 6.53 – 6.80, patella 2.29 – 2.69, chela 7.21 – 7.33, hand 2.50 – 2.68 × as long as broad; femur 2.37 – 2.51 × as long as patella; movable chelal finger 1.71 – 1.93 × as long as hand and 0.64 – 0.65 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, it slightly distal to t and t distal to est; est slightly distal to b (Figs 36 A, 38 A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 36 B, 38 B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 30 – 34 well-spaced, slightly retrorse and pointed teeth and 25 – 26 intercalary microdenticles; movable finger with 20 – 24 almost contiguous, slightly retrorse and pointed teeth, plus 9 – 14 intercalary microdenticles and 6 – 10 vestigial, rounded and contiguous basal teeth. OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4 – 5: 5 – 6: 5 – 6: 4 – 6: T 2 T: 0. Sternal chaetotaxy IV – XII: 12: 8 – 10: 6 – 8: 6 – 8: 7 – 9: 9: 9: -: 2. Genital region: sternite II with 9 – 10 setae scattered on median area, genital opening slit-like, sternite III with row of 14 – 16 setae (Fig. 36 G). LEGS (Fig. 38 C – D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.93 – 2.00 × as long as patella; tarsus 2.25 – 2.32 × as long as tibia. Leg IV: femoropatella 2.79 – 2.90 × as long as deep; tibia 5.27 – 5.36 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 2.67 – 2.78 × as long as deep (TS = 0.24 – 0.33), telotarsus 10.83 – 13.00 × as long as deep and 2.60 – 2.71 × as long as basitarsus (TS = 0.18 – 0.23). Setae of leg I (trochanter to tibia) 3: 10 – 11: 7 – 8: 8 – 10, setae of leg IV (trochanter to basitarsus) 1 – 2: 3: 6: 8 – 9: 8 – 9. Arolium not divided, slightly shorter than simple claws. Adult females (paratypes; Figs 35 B, 36 H) Mostly same as males; tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4: 5: 6: 4: T 2 T: 0; sternal chaetotaxy IV – XII: 10 – 12: 10 – 11: 6 – 8: 8 – 9: 9 – 10: 9 – 10: 9: -: 2. Genital region: sternite II with 10 setae scattered on median area, sternite III with row of 12 setae. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses) Males Body length 1.85 – 2.13. Pedipalps: trochanter 0.26 – 0.29 / 0.16 (1.63 – 1.81), femur 0.98 – 1.02 / 0.15 (6.53 – 6.80), patella 0.39 – 0.43 / 0.16 – 0.17 (2.29 – 2.69), chela 1.32 – 1.37 / 0.18 – 0.19 (7.21 – 7.33), hand 0.45 – 0.51 / 0.18 – 0.19 (2.50 – 2.68), movable chelal finger length 0.87. Chelicera 0.52 – 0.54 / 0.29 (1.79 – 1.86), movable finger length 0.30. Carapace 0.58 – 0.59 / 0.56 – 0.57 (1.04). Leg I: trochanter 0.18 – 0.21 / 0.14 – 0.15 (1.29 – 1.40), femur 0.52 – 0.56 / 0.09 (5.78 – 6.22), patella 0.26 – 0.29 / 0.07 – 0.08 (3.63 – 3.71), tibia 0.25 – 0.28 / 0.06 (4.17 – 4.67), tarsus 0.58 – 0.63 / 0.05 – 0.06 (10.50 – 11.60). Leg IV: trochanter 0.26 – 0.27 / 0.16 – 0.18 (1.44 – 1.69), femoropatella 0.81 – 0.90 / 0.29 – 0.31 (2.79 – 2.90), tibia 0.58 – 0.59 / 0.11 (5.27 – 5.36), basitarsus 0.24 – 0.25 / 0.09 (2.67 – 2.78), telotarsus 0.65 / 0.05 – 0.06 (10.83 – 13.00). Females Body length 1.90 – 2.26. Pedipalps: trochanter 0.29 – 0.32 / 0.17 (1.71 – 1.88), femur 1.08 – 1.09 / 0.15 – 0.16 (6.75 – 7.27), patella 0.43 – 0.45 / 0.18 – 0.19 (2.37 – 2.39), chela 1.45 – 1.49 / 0.21 – 0.22 (6.77 – 6.91), hand 0.53 – 0.59 / 0.21 – 0.22 (2.52 – 2.68), movable chelal finger length 0.92 – 0.94. Chelicera 0.59 – 0.61 / 0.30 – 0.32 (1.84 – 2.03), movable finger length 0.30 – 0.31. Carapace 0.63 – 0.64 / 0.60 – 0.61 (1.03 – 1.07). Leg I: trochanter 0.21 – 0.22 / 0.16 (1.32 – 1.38), femur 0.55 – 0.56 / 0.10 (5.50 – 5.60), patella 0.28 – 0.29 / 0.08 (3.50 – 3.63), tibia 0.28 – 0.29 / 0.07 (4.00 – 4.14), tarsus 0.63 – 0.65 / 0.06 (10.50 – 10.83). Leg IV: trochanter 0.27 – 0.28 / 0.17 – 0.18 (1.5 – 1.65), femoropatella 0.90 – 0.92 / 0.30 – 0.31 (2.96 – 3.00), tibia 0.53 – 0.60 / 0.10 – 0.12 (5.00 – 5.30), basitarsus 0.27 / 0.09 (3.00), telotarsus 0.67 – 0.68 / 0.05 – 0.06 (11.33 – 13.40).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	discussion	Remarks Tyrannochthonius xiannvyanensis sp. nov. most closely resembles T. fenghuangensis sp. nov. in the position of trichobothrium sb (sb midway between st and b), the shape of epistome (small, rounded and obtuse) and carapace with four eyes, anterior pair of eyes reduced, posterior pair of eyespots, but differs in several characters as follows: longer and thinner pedipalpal chela (length 1.32 – 1.37 mm (♂), 1.45 – 1.49 mm (♀), 7.21 – 7.33 (♂), 6.77 – 6.91 (♀) × as long as broad compared to length 0.79 mm (♂), 0.82 – 0.83 mm (♀), 6.08 (♂), 5.13 – 5.19 (♀) × as long as broad in T. fenghuangensis) and longer and thinner pedipalpal femur (length 0.98 – 1.02 mm (♂), 1.08 – 1.09 (♀), 6.53 – 6.80 (♂), 6.75 – 7.27 (♀) × as long as broad compared to length 0.54 mm (♂), 0.55 – 0.58 mm (♀), 5.40 (♂), 4.83 – 5.50 (♀) × as long as broad in T. fenghuangensis).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	distribution	Distribution Known only from the type locality, Xiannvyan Cave (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	description	urn: lsid: zoobank. org: act: DECB 1715 - F 328 - 41 F 5 - 971 E- 22 E 85 EF 9 BF 78 Figs 1 A, 40 – 44	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	diagnosis	Diagnosis (♂ ♀) Moderately sized troglomorphic species with elongated appendages; four eyes, anterior pair of eyes reduced, posterior pair of eyespots; anterior margin of carapace thin, finely denticulated, epistome small, triangular; posterior margin of carapace with 2 setae; tergites I – IV each with four setae. Pedipalps slender, femur 4.54 – 4.75 (♂), 4.57 – 4.92 (♀) × as long as broad, length 0.56 – 0.59 (♂), 0.64 (♀); chela 5.23 – 5.67 (♂), 5.00 – 5.28 (♀) × as long as broad, length 0.84 – 0.89 (♂), 0.95 (♀); both chelal fingers with intercalary teeth; sb closer to st than b.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	etymology	Etymology This species is named after Jiewen Xiao, who participated in field work and collected some of the specimens. Vernacular name NJ 氏ƌ伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	materials_examined	Type material Holotype CHINA • ♂; Guangxi, Guilin City, Xing’an County, cave without name (Unnamed Cave 1); 25.581534 ° N, 110.62378374 ° E; 229 m a. s. l.; 30 Nov. 2023; Jiewen Xiao leg.; MHBU, Ps. - MHBU-GX 2023113002. Paratypes CHINA • 1 ♂, 2 ♀♀; same data as for holotype; MHBU, Ps. - MHBU-GX 2023113003 – 05.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	description	Description Adult males (Figs 40 – 44) COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale. CEPHALOTHORAX (Figs 41 D, 42 A). Carapace nearly subquadrate, 0.89 – 0.90 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, triangular; four eyes, anterior pair of eyes reduced, posterior pair of eyespots; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta less than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with 9 – 10 terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 42 D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5. CHELICERA (Figs 41 C, 42 B). Almost as long as carapace, 1.75 – 1.83 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12 – 13 teeth, distal one largest; movable finger with 12 contiguous small teeth; galea completely vestigial (Fig. 41 C). Serrula exterior with 18 – 19 and serrula interior with 12 – 14 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 42 C). PEDIPALP (Figs 41 A – B, G, 42 E, 43 A – B). Trochanter 1.67 – 1.93, femur 4.54 – 4.75, patella 2.00 – 2.23, chela 5.23 – 5.67, hand 1.65 – 1.73 × as long as broad; femur 1.93 – 2.11 × as long as patella; movable chelal finger 2.21 – 2.31 × as long as hand and 0.70 – 0.71 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb closer to st than b; b and t situated subdistally, it slightly distal to t; est situated at same level as t, and distal to b (Figs 41 A, 43 A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 41 B, 43 B). Both chelal fingers with a row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 24 well-spaced, slightly retrorse and pointed teeth and 17 – 22 intercalary microdenticles; movable finger with 10 – 13 almost contiguous, slightly retrorse and pointed teeth, plus 9 – 11 intercalary microdenticles and 9 – 13 vestigial, rounded and contiguous basal teeth. OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4 – 5: 4 – 5: 4 – 6: 6: 6: 6: 5: T 2 T: 0. Sternal chaetotaxy IV – XII: 12 – 15: 9 – 10: 9 – 10: 9 – 14: 9 – 12: 9 – 13: 9: -: 2. Genital region: sternite II with 7 – 10 setae scattered on median area, genital opening slit-like, sternite III with row of 16 – 18 setae (Fig. 41 H). LEGS (Fig. 43 C – D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.56 – 2.00 × as long as patella; tarsus 1.83 – 2.00 × as long as tibia. Leg IV: femoropatella 2.35 – 3.13 × as long as deep; tibia 2.85 – 4.00 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 2.14 – 2.67 × as long as deep (TS = 0.33 – 0.38), telotarsus 8.03 – 8.75 × as long as deep and 2.06 – 2.27 × as long as basitarsus (TS = 0.23 – 0.27). Setae of leg I (trochanter to tibia) 3 – 4: 9 – 10: 7 – 10: 9 – 12, setae of leg IV (trochanter to basitarsus) 1 – 3: 3: 5 – 6: 8: 8 – 9. Arolium not divided, slightly shorter than simple claws. Adult females (paratypes; Figs 40 B, 41 F, I) Mostly same as males; tergal chaetotaxy I – XII: 4: 4: 4: 5 – 6: 6: 6: 6: 6: 6: 5: T 2 T: 0; sternal chaetotaxy IV – XII: 14: 11: 11: 13: 14: 13: 9: -: 2. Genital region: sternite II with 10 setae scattered on median area, sternite III with row of 12 setae. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses) Males Body length 1.30 – 1.66. Pedipalps: trochanter 0.20 – 0.22 / 0.11 – 0.12 (1.67 – 1.93), femur 0.56 – 0.59 / 0.12 – 0.13 (4.54 – 4.75), patella 0.28 – 0.29 / 0.13 – 0.14 (2.00 – 2.23), chela 0.84 – 0.89 / 0.15 – 0.17 (5.23 – 5.67), hand 0.26 – 0.28 / 0.15 – 0.17 (1.65 – 1.73), movable chelal finger length 0.59 – 0.62. Chelicera 0.41 – 0.44 / 0.23 – 0.24 (1.75 – 1.83), movable finger length 0.24 – 0.26. Carapace 0.41 – 0.50 / 0.46 – 0.48 (0.89 – 0.90). Leg I: trochanter 0.13 – 0.15 / 0.08 – 0.09 (1.42 – 1.87), femur 0.28 – 0.31 / 0.05 – 0.07 (4.29 – 5.43), patella 0.12 – 0.18 / 0.05 – 0.06 (2.50 – 3.60), tibia 0.13 – 0.18 / 0.04 – 0.06 (2.82 – 4.25), tarsus 0.33 – 0.35 / 0.04 (8.25 – 9.03). Leg IV: trochanter 0.15 – 0.19 / 0.11 – 0.12 (1.30 – 1.64), femoropatella 0.45 – 0.55 / 0.15 – 0.20 (2.35 – 3.13), tibia 0.25 – 0.36 / 0.09 (2.85 – 4.00), basitarsus 0.12 – 0.16 / 0.05 – 0.07 (2.14 – 2.67), telotarsus 0.31 – 0.35 / 0.04 (8.03 – 8.75). Females Body length 1.53 – 1.80. Pedipalps: trochanter 0.23 – 0.24 / 0.13 (1.77 – 1.91), femur 0.64 / 0.13 – 0.14 (4.57 – 4.92), patella 0.25 – 0.32 / 0.12 – 0.15 (2.07 – 2.13), chela 0.95 / 0.18 – 0.19 (5.00 – 5.28), hand 0.31 / 0.18 – 0.19 (1.63 – 1.72), movable chelal finger length 0.65 – 0.66. Chelicera 0.48 – 0.49 / 0.27 (1.78 – 1.81), movable finger length 0.29. Carapace 0.47 / 0.54 – 0.56 (0.84 – 0.87). Leg I: trochanter 0.14 – 0.15 / 0.10 – 0.12 (1.25 – 1.41), femur 0.33 – 0.35 / 0.06 – 0.07 (5.00 – 5.39), patella 0.15 – 0.18 / 0.06 (2.36 – 3.00), tibia 0.20 / 0.05 (3.76 – 4.00), tarsus 0.36 – 0.37 / 0.04 – 0.05 (7.40 – 9.53). Leg IV: trochanter 0.18 – 0.20 / 0.13 – 0.15 (1.33 – 1.38), femoropatella 0.53 – 0.55 / 0.12 – 0.19 (2.89 – 4.41), tibia 0.32 – 0.36 / 0.08 – 0.09 (4.00 – 4.15), basitarsus 0.15 – 0.17 / 0.06 – 0.06 (2.50 – 2.86), telotarsus 0.33 – 0.38 / 0.04 (8.25 – 9.67).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	discussion	Remarks Tyrannochthonius xiaoi sp. nov. differs from all other hypogean species of the genus from China except T. duo, T. fenghuangensis sp. nov. and T. xiannvyanensis sp. nov. by the four eyes, anterior pair of eyes reduced, posterior pair of eyespots. It differs from T. duo, T. fenghuangensis and T. xiannvyanensis by its shorter and thicker pedipalpal chela (length 0.84 – 0.89 mm (♂), 0.56 – 0.59 mm (♀), 5.23 – 5.67 (♂), 4.54 – 4.75 (♀) × as long as broad compared to length 1.11 mm (♂), 1.17 – 1.18 mm (♀), 6.94 (♂), 5.85 – 6.56 (♀) × as long as broad in T. duo, and 1.32 – 1.37 mm (♂), 1.45 – 1.49 mm (♀), 7.21 – 7.33 (♂), 6.77 – 6.91 (♀) × as long as broad in T. xiannvyanensis), the shape of epistome (epistome small, triangular compared to epistome small, rounded and obtuse in T. fenghuangensis and T. xiannvyanensis) and the position of trichobothrium (sb closer to st than b compared to sb midway between st and b in T. fenghuangensis and T. xiannvyanensis and sb closer to b than to st in T. duo) (Hou et al. 2023 b).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	biology_ecology	Ecology These specimens were collected under mud inside the cave (Fig. 44).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	distribution	Distribution Known only from the type locality, Unnamed Cave 1 (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFA66EB0FE10FDDBFBA6.taxon	description	Fig. 1 E	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFA66EB0FE10FDDBFBA6.taxon	diagnosis	Diagnosis and description For details see Sun et al. (2025). Vernacular name 岩屋ƌ伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFA66EB0FE10FDDBFBA6.taxon	materials_examined	Material examined Holotype CHINA • ♂; Guangxi, Baise City, Leye County, Yanwu Cave; 24.823900 ° N, 106.431500 ° E; 695 m a. s. l.; 28 Apr. 2024; Sunbin Huang, Mingzhi Zhao, Yan Li and Rong Chen leg.; MHBU, Ps. - MHBU-GX 2024042801.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFA66EB0FE10FDDBFBA6.taxon	distribution	Distribution Yanwu Cave (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFD96D67FBEAFC07FE48.taxon	description	urn: lsid: zoobank. org: act: D 3 D 473 B 6 - ED 3 C- 4 AA 5 - 8 FFE- 5 F 3 D 0773 CDFB Figs 45 – 49	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFD96D67FBEAFC07FE48.taxon	diagnosis	Diagnosis (♂ ♀) Moderately sized troglomorphic species with elongated appendages; four eyes, anterior pair of eyes reduced, posterior pair of eyespots; anterior margin of carapace thin, finely denticulated, epistome small, triangular; posterior margin of carapace with 2 setae; tergites I – IV each with four setae. Pedipalps slender, femur 4.54 – 4.75 (♂), 4.57 – 4.92 (♀) × as long as broad, length 0.56 – 0.59 (♂), 0.64 (♀); chela 5.23 – 5.67 (♂), 5.00 – 5.28 (♀) × as long as broad, length 0.84 – 0.89 (♂), 0.95 (♀); both chelal fingers with intercalary teeth; sb closer to st than b.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFD96D67FBEAFC07FE48.taxon	etymology	Etymology Named after the type locality, Yilingyan Cave. Vernacular name 伊岭岩ƌ伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFD96D67FBEAFC07FE48.taxon	materials_examined	Type material Holotype CHINA • ♂; Guangxi, Nanning City, Wuming District, Yilingyan Cave; 23.041315 ° N, 108.294759 ° E; 96 m a. s. l.; 8 Oct. 2019; Zegang Feng and Lingchen Zhao leg.; MHBU, Ps. - MHBU-GX 2019100801. Paratypes CHINA • 1 ♂, 2 ♀♀; same data as for holotype; MHBU, Ps. - MHBU-GX 2019100802 – 04.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFD96D67FBEAFC07FE48.taxon	description	Description Adult males (Figs 45 – 49) COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale. CEPHALOTHORAX (Figs 46 D, 47 A). Carapace nearly subquadrate, 0.89 – 0.90 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, triangular; four eyes, anterior pair of eyes reduced, posterior pair of eyespots; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with 7 – 8 terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 47 D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5. CHELICERA (Figs 46 C, 47 B). Almost as long as carapace, 2.04 – 2.14 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13 – 14 teeth, distal one largest; movable finger with 13 – 15 contiguous small teeth; galea completely vestigial (Fig. 46 C). Serrula exterior with 16 – 21 and serrula interior with 10 – 12 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 47 C). PEDIPALP (Figs 46 A – B, F, 47 E, 48 A – B). Trochanter 1.83 – 1.92, femur 5.15 – 5.67, patella 1.93 – 2.21, chela 6.47 – 6.80, hand 2.20 – 2.33 × as long as broad; femur 2.19 – 2.31 × as long as patella; movable chelal finger 1.94 – 1.97 × as long as hand and 0.67 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb closer to st than b; b and t situated subdistally, it situated at same level as t; est slightly distal to b (Figs 46 A, 48 A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 46 B, 48 B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 28 – 33 well-spaced, slightly retrorse and pointed teeth and 26 – 27 intercalary microdenticles; movable finger with 15 – 20 almost contiguous, slightly retrorse and pointed teeth, plus 15 – 16 intercalary microdenticles and 14 – 20 vestigial, rounded and contiguous basal teeth. OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 4: 4: 4: 4: 4: 4: 4: 5: 5: 4: T 2 T: 0. Sternal chaetotaxy IV – XII: 12: 10: 9: 9: 9: 9: 9: -: 2. Genital region: sternite II with nine setae scattered on median area, genital opening slit-like, sternite III with row of 14 setae (Fig. 46 G). LEGS (Fig. 48 C – D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 2.05 – 2.17 × as long as patella; tarsus 2.00 – 2.21 × as long as tibia. Leg IV: femoropatella 2.60 – 2.86 × as long as deep; tibia 4.67 – 5.00 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 3.00 – 3.33 × as long as deep (TS = 0.24), telotarsus 10.83 – 13.00 × as long as deep and 2.28 – 2.35 × as long as basitarsus (TS = 0.32 – 0.33). Setae of leg I (trochanter to tibia) 3 – 4: 11 – 12: 8: 9 – 12, setae of leg IV (trochanter to basitarsus) 2 – 3: 3: 5 – 8: 9 – 10: 8 – 9. Arolium not divided, slightly shorter than simple claws. Adult females (paratypes; Figs 45 B, 46 H) Mostly same as males; tergal chaetotaxy I – XII: 4: 4: 4: 5 – 6: 6: 6: 6: 6: 6: 5: T 2 T: 0; sternal chaetotaxy IV – XII: 14: 11: 11: 13: 14: 13: 9: -: 2. Genital region: sternite II with 10 setae scattered on median area, sternite III with row of 12 setae. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses) Males Body length 1.39 – 1.45. Pedipalps: trochanter 0.22 – 0.23 / 0.12 (1.83 – 1.92), femur 0.67 – 0.68 / 0.12 – 0.13 (5.15 – 5.67), patella 0.29 – 0.31 / 0.14 – 0.15 (1.93 – 2.21), chela 0.97 – 1.02 / 0.15 (6.47 – 6.80), hand 0.33 – 0.35 / 0.15 (2.20 – 2.33), movable chelal finger length 0.65 – 0.68. Chelicera 0.47 – 0.49 / 0.22 – 0.24 (2.04 – 2.14), movable finger length 0.24 – 0.25. Carapace 0.44 – 0.45 / 0.50 – 0.51 (0.88). Leg I: trochanter 0.14 – 0.15 / 0.09 (1.56 – 1.67), femur 0.39 – 0.41 / 0.07 (5.57 – 5.86), patella 0.18 – 0.20 / 0.06 – 0.07 (2.57 – 3.33), tibia 0.19 – 0.21 / 0.04 – 0.05 (3.80 – 5.25), tarsus 0.42 / 0.04 (10.50). Leg IV: trochanter 0.18 – 0.20 / 0.11 (1.64 – 1.82), femoropatella 0.63 – 0.65 / 0.22 – 0.25 (2.60 – 2.86), tibia 0.42 – 0.45 / 0.09 (4.67 – 5.00), basitarsus 0.20 – 0.21 / 0.06 – 0.07 (3.00 – 3.33), telotarsus 0.47 – 0.48 / 0.04 (11.75 – 12.00). Females Body length 1.39 – 1.53. Pedipalps: trochanter 0.25 / 0.13 – 0.14 (1.78 – 1.92), femur 0.72 – 0.76 / 0.14 – 0.15 (5.07 – 5.14), patella 0.32 – 0.33 / 0.16 – 0.17 (1.94 – 2.00), chela 1.06 – 1.09 / 0.18 – 0.19 (5.58 – 6.05), hand 0.36 – 0.38 / 0.18 – 0.19 (1.89 – 2.11), movable chelal finger length 0.71 – 0.73. Chelicera 0.52 – 0.55 / 0.25 – 0.27 (2.04 – 2.08), movable finger length 0.27 – 0.29. Carapace 0.47 – 0.54 / 0.54 – 0.56 (0.84 – 1.00). Leg I: trochanter 0.14 – 0.17 / 0.10 – 0.12 (1.40 – 1.42), femur 0.40 – 0.43 / 0.07 – 0.08 (5.38 – 5.71), patella 0.21 – 0.22 / 0.07 (3.00 – 3.14), tibia 0.22 / 0.05 – 0.06 (3.67 – 4.40), tarsus 0.44 – 0.45 / 0.04 – 0.05 (9.00 – 11.00). Leg IV: trochanter 0.20 – 0.21 / 0.13 – 0.14 (1.50 – 1.54), femoropatella 0.66 – 0.68 / 0.23 – 0.24 (2.83 – 2.87), tibia 0.46 / 0.08 – 0.10 (4.60 – 5.75), basitarsus 0.21 / 0.07 – 0.08 (2.63 – 3.00), telotarsus 0.49 / 0.04 – 0.05 (9.80 – 12.25).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFD96D67FBEAFC07FE48.taxon	discussion	Remarks Tyrannochthonius yilingyanensis sp. nov. most closely resembles T. xiaoi sp. nov. in the both chelal fingers with intercalary teeth, trichobothrium sb closer to st than b, epistome small, triangular, tergites I – IV each with four setae and four eyes, anterior pair of eyes reduced, posterior pair of eyespots, but differs in several characters as follows: longer and thinner pedipalpal chela (length 0.97 – 1.02 mm (♂), 1.06 – 1.09 mm (♀), 6.47 – 6.80 (♂), 5.58 – 6.05 (♀) × as long as broad compared to length 0.84 – 0.89 mm (♂), 0.95 mm (♀), 5.23 – 5.67 (♂), 5.00 – 5.28 (♀) × as long as broad in T. xiaoi) and longer and thinner pedipalpal femur (length 0.67 – 0.68 mm (♂), 0.72 – 0.76 (♀), 5.15 – 5.67 (♂), 5.07 – 5.14 (♀) × as long as broad compared to length 0.56 – 0.59 mm (♂), 0.64 mm (♀), 4.54 – 4.75 (♂), 4.57 – 4.92 (♀) × as long as broad in T. xiaoi).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFD96D67FBEAFC07FE48.taxon	distribution	Distribution Known only from the type locality, Yilingyan Cave (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	description	urn: lsid: zoobank. org: act: 219044 D 3 - 5956 - 477 A-B 9 C 0 - CF 290452 B 267 Figs 1 M, 50 – 54	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	diagnosis	Diagnosis (♂) Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed, triangular; posterior margin of carapace with 2 setae; tergites I – II each with two setae, tergite III – IV each with four setae. Pedipalps slender, femur 5.00 (♂) × as long as broad, length 0.40 (♂); chela 6.44 (♂) × as long as broad, length 0.58 (♂); both chelal fingers without intercalary teeth; sb closer to st than b.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	etymology	Etymology This species is named after Kun Yu, who collected this specimen. Vernacular name Ḕ 氏ƌ伪 $.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	materials_examined	Type material Holotype CHINA • ♂; Guangxi, Liuzhou City, Yufeng District, Liyong Town, Unnamed Cave 2; 24.20260909 ° N, 109.49899591 ° E; 94 m a. s. l.; 9 Aug. 2023; Kun Yu leg.; MHBU, Ps. - MHBU-GX 2023080901.	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	description	Description Adult male (Figs 50 – 54) COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale. CEPHALOTHORAX (Figs 51 E, 52 A). Carapace nearly subquadrate, 0.94 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, triangular; without eyes or eyespots; with 18 setae arranged s 4 s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I rounded anteromedial process; coxae II with seven or eight terminally indented coxal spines, set as oblique and arched row, central spines slightly longer than others (Fig. 52 D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5. CHELICERA (Figs 51 D, 52 B). Almost as long as carapace, 1.93 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with nine teeth, distal one largest; movable finger with 10 contiguous small teeth; galea completely vestigial (Fig. 51 D). Serrula exterior with 17 and serrula interior with 11 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 52 C). PEDIPALP (Figs 51 A – B, F, 52 E, 53 A – B). Trochanter 1.75, femur 5.00, patella 2.00, chela 6.44, hand 2.11 × as long as broad; femur 2.22 × as long as patella; movable chelal finger 2.05 × as long as hand and 0.67 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb closer to st than b; b and t situated subdistally, it slightly distal to t; est situated at same level as t and distal to b (Figs 51 A, 53 A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 51 B, 53 B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 25 well-spaced, slightly retrorse and pointed teeth and 12 intercalary microdenticles; movable finger with 10 almost contiguous, slightly retrorse and pointed teeth, plus nine intercalary microdenticles and nine vestigial, rounded and contiguous basal teeth. OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I – XII: 2: 4: 4: 4: 4: 5: 5: 5: 5: 4: T 2 T: 0. Sternal chaetotaxy IV – XII: 12: 8: 7: 7: 6: 7: 7: -: 2. Genital region: sternite II with 10 setae scattered on median area, genital opening slit-like, sternite III with row of 14 setae (Fig. 51 C). LEGS (Fig. 53 C – D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 2.09 × as long as patella; tarsus 2.00 × as long as tibia. Leg IV: femoropatella 2.57 × as long as deep; tibia 5.00 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 3.00 × as long as deep (TS = 0.29), telotarsus 8.33 × as long as deep and 2.08 × as long as basitarsus (TS = 0.32). Setae of leg I (trochanter to tibia) 4: 10: 8: 9, setae of leg IV (trochanter to basitarsus) 2: 3: 6: 9: 10. Arolium not divided, slightly shorter than simple claws. Dimensions (length / breadth or, in the case of the legs, length / depth in mm; ratios in parentheses) Male Body length 0.98. Pedipalps: trochanter 0.14 / 0.08 (1.75), femur 0.40 / 0.08 (5.00), patella 0.40 / 0.08 (5.00), chela 0.58 / 0.09 (6.44), hand 0.19 / 0.09 (2.11), movable chelal finger length 0.39. Chelicera 0.29 / 0.15 (1.93), movable finger length 0.17. Carapace 0.31 / 0.33 (0.94). Leg I: trochanter 0.08 / 0.04 (2.00), femur 0.23 / 0.04 (5.75), patella 0.11 / 0.04 (2.75), tibia 0.12 / 0.03 (4.00), tarsus 0.24 / 0.03 (8.00). Leg IV: trochanter 0.11 / 0.08 (1.38), femoropatella 0.36 / 0.14 (2.57), tibia 0.25 / 0.05 (5.00), basitarsus 0.12 / 0.04 (3.00), telotarsus 0.25 / 0.03 (8.33).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	discussion	Remarks Tyrannochthonius yui sp. nov. most closely resembles T. yanwuensis by the tergite I with two setae, tergite III – IV and X each with four setae, both chelal fingers with intercalary teeth and carapace without eyes or eyespots, but differs in several characters as follows: tergite II with four setae (vs tergite II with two setae in T. yanwuensis), shorter pedipalpal chela (length 0.58 mm (♂) compared to length 1.31 mm (♂) in T. yanwuensis) and shorter pedipalpal femur (length 0.40 mm (♂) compared to length 0.94 mm (♂) in T. yanwuensis) (Sun et al. 2025).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	biology_ecology	Ecology The specimen was collected from bat guano inside the cave (Fig. 54).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	distribution	Distribution Known only from the type locality, Unnamed Cave 2 (Guangxi, China).	en	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
