identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
096987E3E443FF996D6BFD18FCA6FC80.text	096987E3E443FF996D6BFD18FCA6FC80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius Chamberlin 1929	<div><p>Genus Tyrannochthonius Chamberlin, 1929</p><p>Type species</p><p>Chthonius terribilis With, 1906, by original designation.</p></div>	https://treatment.plazi.org/id/096987E3E443FF996D6BFD18FCA6FC80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E443FF936D40FCD1FC39F9D8.text	096987E3E443FF936D40FCD1FC39F9D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius baiseensis Sun & Guo & Zhang 2026	<div><p>Tyrannochthonius baiseensis sp. nov.</p><p>urn:lsid:zoobank.org:act: E7931503-1D3B-490F-B3BA-9E810FCE1470</p><p>Figs 1H, 2–6</p><p>Diagnosis</p><p>(♂ ♀) Moderately sized troglomorphic species with elongated appendages; four eyes, anterior pair of eyes well-developed, posterior pair with flat lenses; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse; posterior margin of carapace with 2 setae; tergites I–II each with four setae. Pedipalps slender, femur 5.31 (♂), 5.14 (♀) × as long as broad, length 0.69 (♂), 0.72 (♀); chela 5.94 (♂), 5.21 (♀) × as long as broad, length 0.95 (♂), 0.99 (♀); both chelal fingers with intercalary teeth; sb closer to st than b.</p><p>Etymology</p><p>Named after the closest city to the type locality, Baise City.</p><p>Vernacular name</p><p>ÞDzƌ伪$.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♂; Guangxi, Baise City, Lingyun County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.70881&amp;materialsCitation.latitude=24.35539" title="Search Plazi for locations around (long 106.70881/lat 24.35539)">Xiaoluo Town</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.70881&amp;materialsCitation.latitude=24.35539" title="Search Plazi for locations around (long 106.70881/lat 24.35539)">Dayan Cave</a>; 24.35539° N, 106.708811° E; 735 m a.s.l.; 26 Jul. 2021; Zegang Feng, Hongru Xu, Liu Fu and Nana Zhan leg.; MHBU Ps.- MHBU-GX2021072601.</p><p>Paratypes</p><p>CHINA • ♀; same data as for holotype; MHBU, Ps.- MHBU-GX2021072602 .</p><p>Description</p><p>Adult male (holotype and male paratypes; Figs 2–6)</p><p>COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale.</p><p>CEPHALOTHORAX (Figs 3D–E, 4A). Carapace nearly subquadrate, 1.09 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, rounded and obtuse; four eyes, anterior pair of eyes well-developed, posterior pair with flat lenses; with 18 setae arranged s4s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to the setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with 8–11 terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 4D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5.</p><p>CHELICERA (Figs 3C, 4B). Almost as long as carapace, 2.19 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13 teeth, distal one largest; movable finger with 13 contiguous small teeth; galea completely vestigial (Fig. 4B). Serrula exterior with 20 and serrula interior with 13 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 4C).</p><p>PEDIPALP (Figs 3A–B, F, 5A–B). Trochanter 1.69, femur 5.31, patella 2.29, chela 5.94, hand 1.94 × as long as broad; femur 2.16 × as long as patella; movable chelal finger 2.13 × as long as hand and 0.69 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb slightly closer to st than to b; b and t situated subdistally, it slightly distal to t and t distal to b; est situated proximal to t and close to it (Figs 4A, 5A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 4B, 5B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 28 well-spaced, and pointed teeth and 25 intercalary microdenticles; movable finger with 11 well-spaced, pointed teeth, plus 11 intercalary microdenticles and 20 vestigial, rounded and contiguous basal teeth.</p><p>OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 6: 5: 7: 8: 9: 10: 7: 5: T2T: 0. Sternal chaetotaxy IV–XII: 12: 12: 12: 12: 14:14: 12: -: 2. Genital region: sternite II with nine setae scattered on median area, genital opening slit-like, sternite III with row of 14 setae (Fig. 3G).</p><p>LEGS (Fig. 5C–D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.94 × as long as patella; tarsus 1.95 × as long as tibia. Leg IV: femoropatella 2.71 × as long as deep; tibia 4.67 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 2.71 × as long as deep (TS = 0.33), telotarsus 10.50 × as long as deep and 2.21 × as long as basitarsus (TS = 0.26). Setae of leg I (trochanter to tibia) 4: 10–11: 7: 10, setae of leg IV (trochanter to basitarsus) 2: 3: 6: 8:7. Arolium not divided, slightly shorter than simple claws.</p><p>Adult female (paratype; Figs 2B, 3H)</p><p>Mostly same as males; tergal chaetotaxy I–XII: 4: 4: 5: 5: 6: 7: 7: 8: 9: 8: T2T: 0; sternal chaetotaxy IV–XII: 12: 12:11: 15: 13: 12: 12: -: 2. Genital region: sternite II with nine setae scattered on median area, sternite III with row of 12 setae.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses)</p><p>Male</p><p>Body length 1.75. Pedipalps: trochanter 0.22/0.13 (1.69), femur 0.69/0.13 (5.31), patella 0.32/0.14 (2.29), chela 0.95/0.16 (5.94), hand 0.31/0.16 (1.94), movable chelal finger length 0.66. Chelicera 0.46/0.21 (2.19), movable finger length 0.26. Carapace 0.49/0.45 (1.09). Leg I: trochanter 0.12/0.09 (1.33), femur 0.35/0.07 (5.00), patella 0.18/0.07 (2.57), tibia 0.20/0.05 (4.00), tarsus 0.39/0.04 (9.75). Leg IV: trochanter 0.18/0.12 (1.50), femoropatella 0.57/0.21 (2.71), tibia 0.42/0.09 (4.67), basitarsus 0.19/0.07 (2.71), telotarsus 0.42/0.04 (10.50).</p><p>Female</p><p>Body length 1.73. Pedipalps: trochanter 0.26/0.14 (1.86), femur 0.72/0.14 (5.14), patella 0.34/0.16 (2.13), chela 0.99/0.19 (5.21), hand 0.34/0.19 (1.79), movable chelal finger length 0.66. Chelicera 0.52/0.25 (2.08), movable finger length 0.30. Carapace 0.46/0.58 (0.79). Leg I: trochanter 0.15/0.11 (1.36), femur 0.38/0.07 (5.43), patella 0.20/0.07 (2.86), tibia 0.22/0.05 (4.40), tarsus 0.42/0.05 (8.40). Leg IV: trochanter 0.21/0.13 (1.61), femoropatella 0.62/0.23 (2.70), tibia 0.42/0.10 (4.20), basitarsus 0.19/0.07 (2.71), telotarsus 0.41/0.04 (10.25).</p><p>Remarks</p><p>Tyrannochthonius baiseensis sp. nov. most closely resembles T. robustus Beier, 1951 in the position of the trichobothrium (sb closer to st than b), the chaetotaxy of tergites (I–II each with four setae), and the number of eyes, but differs in several characters as follows: longer pedipalpal chela (0.99 mm (♀) compared to 0.79 mm (♀) in T. robustus Beier, 1951), longer pedipalpal femur (0.72 mm (♀) compared to 0.47 mm (♀) in T. robustus Beier, 1951), and the shape of epistome (small, rounded and obtuse compared to pointed, triangular in T. robustus Beier, 1951) (Beier 1951).</p><p>Ecology</p><p>The specimen was collected in the completely dark zone near the cave entrance (Fig. 6).</p><p>Distribution</p><p>Known only from the type locality, Dayan Cave (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E443FF936D40FCD1FC39F9D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E449FF906EB3F919FD9FFDFD.text	096987E3E449FF906EB3F919FD9FFDFD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius chixingi Gao, Wynne & Zhang 2018	<div><p>Tyrannochthonius chixingi Gao, Wynne &amp; Zhang, 2018</p><p>Fig. 1C</p><p>Tyrannochthonius chixingi Gao, Wynne &amp; Zhang, 2018: 173–195, figs 1, 6a–b.</p><p>Diagnosis and description</p><p>For details see Gao et al. (2018).</p><p>Vernacular name</p><p>Ṅ形ƌ伪$.</p><p>Material examined</p><p>Holotype</p><p>CHINA • ♂; Guangxi, Guilin City, Xiufeng District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.27018&amp;materialsCitation.latitude=25.31281" title="Search Plazi for locations around (long 110.27018/lat 25.31281)">Maomaotou Cave</a>, deep zone; 25.312811° N, 110.270178° E; 225 m a.s.l.; 15 Nov. 2016; J.J. Wynne leg.; MHBU, Ps.- MHBU-GX16121502.</p><p>Paratypes</p><p>CHINA • 1 ♂, 1 ♀; same data as for holotype; MHBU, Ps.- MHBU-GX 16121503 –04 .</p><p>Distribution</p><p>Maomaotou Cave (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E449FF906EB3F919FD9FFDFD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E44AFF8A6D6DFD35FBFCFEC0.text	096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius fenghuangensis Sun & Guo & Zhang 2026	<div><p>Tyrannochthonius fenghuangensis sp. nov.</p><p>urn:lsid:zoobank.org:act: 6C147713-251A-45B7-82E0-8A3D3C0C636B</p><p>Figs 1L, 7–11</p><p>Diagnosis</p><p>(♂ ♀) Moderately sized troglomorphic species with elongated appendages; four eyes, anterior pair of eyes strongly reduced, posterior pair with eyespots; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse; posterior margin of carapace with 2 setae; tergites I–II each with four setae. Pedipalps slender, femur 5.40 (♂), 4.83–5.50 (♀) × as long as broad, length 0.54 (♂), 0.55– 0.58 (♀); chela 6.08 (♂), 5.13–5.19 (♀) × as long as broad, length 0.79 (♂), 0.82–0.83 (♀); both chelal fingers with intercalary teeth; sb midway between st and b.</p><p>Etymology</p><p>Named after the type locality, Fenghuang Township.</p><p>Vernacular name</p><p>Ṇdzƌ伪$.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♂; Guangxi, Hechi City, Bama Yao Autonomous County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.39044&amp;materialsCitation.latitude=24.193687" title="Search Plazi for locations around (long 107.39044/lat 24.193687)">Fenghuang Township</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.39044&amp;materialsCitation.latitude=24.193687" title="Search Plazi for locations around (long 107.39044/lat 24.193687)">Unnamed Cave</a> 3; 24.1936872° N, 107.390445° E; 467.6 m a.s.l.; 28 Jul. 2021; Zegang Feng, Hongru Xu, Liu Fu and Nana Zhan leg.; MHBU, Ps.- MHBU-GX2021072801.</p><p>Paratypes</p><p>CHINA • 2 ♀♀; same data as for holotype; MHBU, Ps.- MHBU-GX2021072802–03 .</p><p>Description</p><p>Adult male (Figs 7–11)</p><p>COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale.</p><p>CEPHALOTHORAX (Figs 8D–E, 9A). Carapace nearly subquadrate, 1.18 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, rounded and obtuse; four eyes, anterior pair of eyes strongly reduced, posterior pair with eyespots; with 18 setae arranged s4s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with seven terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 9D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5.</p><p>CHELICERA (Figs 8C, 9B). Almost as long as carapace, 2.10 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13 teeth, distal one largest; movable finger with 12 contiguous small teeth; galea completely vestigial (Fig. 8C). Serrula exterior with 19 and serrula interior with 12 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 9C).</p><p>PEDIPALP (Figs 8A–B, F, 9E, 10A–B). Trochanter 1.82, femur 5.40, patella 2.00, chela 6.08, hand 1.92 × as long as broad; femur 2.25 × as long as patella; movable chelal finger 2.20 × as long as hand and 0.70 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, it slightly distal to t and t distal to est; est situated distal to b (Figs 8A, 10A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 8B, 10B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 24 well-spaced, and pointed teeth and 22 intercalary microdenticles; movable finger with 10 well-spaced, pointed teeth, plus 9 intercalary microdenticles and 13 vestigial, rounded and contiguous basal teeth.</p><p>OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 5: 7: 7: 8: 8: 8: 8: 7: T2T: 0. Sternal chaetotaxy IV–XII: 12: 13:12: 12: 13: 13: 11:-: 2. Genital region: sternite II with 11 setae scattered on median area, genital opening slit-like, sternite III with row of 16 setae (Fig. 8G).</p><p>LEGS (Fig. 10C–D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.94 × as long as patella; tarsus 1.95 × as long as tibia. Leg IV: femoropatella 2.71 × as long as deep; tibia 4.67 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 2.71 × as long as deep (TS = 0.33), telotarsus 10.50 × as long as deep and 2.21 × as long as basitarsus (TS = 0.26). Setae of leg I (trochanter to tibia) 4: 10–11: 7: 10, setae of leg IV (trochanter to basitarsus) 2: 3: 6: 8:7. Arolium not divided, slightly shorter than simple claws.</p><p>Adult females (paratypes; Figs 7B, 8H)</p><p>Mostly same as males; tergal chaetotaxy I–XII: 4: 4: 5: 7–8: 8: 8–9: 8–10: 10–12: 8–9: 5–7: T2T: 0; sternal chaetotaxy IV–XII: 11–12: 12–14: 11–12:12–13: 13–16: 13: 9–12: -: 2. Genital region: sternite II with 10 setae scattered on median area, sternite III with row of 12 setae.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses)</p><p>Male</p><p>Body length 1.40. Pedipalps: trochanter 0.20/0.11 (1.82), femur 0.54/0.10 (5.40), patella 0.24/0.12 (2.00), chela 0.79/0.13 (6.08), hand 0.25/0.13 (1.94), movable chelal finger length 0.55. Chelicera 0.40/0.19 (2.10), movable finger length 0.23. Carapace 0.38/0.45 (0.84). Leg I: trochanter 0.12/0.09 (1.33), femur 0.29/0.05 (5.80), patella 0.14/0.05 (2.80), tibia 0.17/0.04 (4.25), tarsus 0.32/0.04 (8.00). Leg IV: trochanter 0.17/0.11 (1.54), femoropatella 0.48/0.19 (2.52), tibia 0.32/0.08 (4.00), basitarsus 0.14/0.06 (2.33), telotarsus 0.34/0.03 (11.33).</p><p>Females</p><p>Body length 1.42–1.45. Pedipalps: trochanter 0.20–0.21/0.11 (1.82–1.91), femur 0.55–0.58/0.10–0.12 (4.83–5.50), patella 0.27/0.12 (2.25), chela 0.82–0.83/0.16 (5.13–5.19), hand 0.26–0.27/0.16 (1.63– 1.69), movable chelal finger length 0.57. Chelicera 0.43–0.45/0.22 (1.95–2.04), movable finger length 0.24–0.25. Carapace 0.39–0.48/0.48–0.49 (0.81–0.98). Leg I: trochanter 0.11–0.13/0.08–0.10 (1.30– 1.38), femur 0.29–0.31/0.06 (4.83–5.17), patella 0.15–0.17/0.05–0.06 (2.83–3.00), tibia 0.16–0.17/0.04 (4.00–4.25), tarsus 0.34–0.35/0.04 (8.50–8.75). Leg IV: trochanter 0.13–0.18/0.10–0.11 (1.30–1.64), femoropatella 0.48–0.49/0.17–0.18 (2.72–2.82), tibia 0.32–0.35/0.08 (4.00–4.38), basitarsus 0.15– 0.16/0.06 (2.50–2.67), telotarsus 0.34–0.35/0.03–0.04 (8.50–11.67).</p><p>Remarks</p><p>Tyrannochthonius fenghuangensis sp. nov. differs from all other hypogean species of the genus from China except Tyrannochthonius duo by the presence of four eyes, anterior pair of eyes strongly reduced, posterior pair with eyespots. It differs from T. duo by its shorter and thicker pedipalpal chela (length 0.79 mm (♂), 0.82–0.83 mm (♀), 6.08 (♂), 5.13–5.19 (♀) × as long as broad compared to length 1.11 mm (♂), 1.17–1.18 mm (♀), 6.94 (♂), 5.85–6.56 (♀) × as long as broad in T. duo), shorter and thicker pedipalpal femur (length 0.54 mm (♂), 0.55–0.58 mm (♀), 5.40 (♂), 4.83–5.50 (♀) × as long as broad compared to length 0.76 mm (♂), 0.79–0.81 mm (♀), 5.85 (♂), 5.79–6.08 (♀) × as long as broad in T. duo), the shape of epistome (epistome small, rounded and obtuse vs epistome small, triangular in T. duo) and the position of trichobothrium (sb midway between st and b compared to sb closer to b than to st in T. duo) (Hou et al. 2023b).</p><p>Distribution</p><p>Known only from the type locality, Unnamed Cave 3 (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E44AFF8A6D6DFD35FBFCFEC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E451FF8C6D40FE97FC31FA81.text	096987E3E451FF8C6D40FE97FC31FA81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius laqiuensis Sun & Guo & Zhang 2026	<div><p>Tyrannochthonius laqiuensis sp. nov.</p><p>urn:lsid:zoobank.org:act: D9943FDC-E713-4225-8EF0-3C0D533F082D</p><p>Figs 1I, 12–15</p><p>Diagnosis</p><p>(♂ ♀) Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, triangular; posterior margin of carapace with 2 setae; tergites I–IV each with four setae. Pedipalps slender, femur 7.13 (♂), 6.75 (♀) × as long as broad, length 1.07 (♂), 1.08 (♀); chela 7.14 (♂), 6.58 (♀) × as long as broad, length 1.50 (♂), 1.58 (♀); chelal fingers without intercalary teeth and movable chelal finger teeth markedly retrorse and almost contiguous; sb slightly closer to st than to b.</p><p>Etymology</p><p>Named after the type locality, Laqiu Cave.</p><p>Vernacular name</p><p>®丘ƌ伪$.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♂; Guangxi, Hechi City, Donglan County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.30039&amp;materialsCitation.latitude=24.358433" title="Search Plazi for locations around (long 107.30039/lat 24.358433)">Wuzuan Town</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.30039&amp;materialsCitation.latitude=24.358433" title="Search Plazi for locations around (long 107.30039/lat 24.358433)">Laqiu Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.30039&amp;materialsCitation.latitude=24.358433" title="Search Plazi for locations around (long 107.30039/lat 24.358433)">Laqiu Cave</a>; 24.358433° N, 107.300395° E; 467 m a.s.l.; 11 May. 2018; Zegang Feng and Weitong Wang leg.; MHBU, Ps.- MHBU-GX18051101.</p><p>Paratype</p><p>CHINA • ♀; same data as for holotype; MHBU, Ps.- MHBU-GX 18051102 .</p><p>Description</p><p>Adult male (Figs 12–15)</p><p>COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale.</p><p>CEPHALOTHORAX (Figs 13D, 14A). Carapace nearly subquadrate, 1.02 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, triangular; without eyes or eyespots; with 18 setae arranged s4s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with nine terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 14D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5.</p><p>CHELICERA (Figs 13C, 14B). Almost as long as carapace, 2.44 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12 teeth, distal one largest; movable finger with 10 contiguous small teeth; galea completely vestigial (Fig. 13C). Serrula exterior with 28 and serrula interior with 14 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 14C).</p><p>PEDIPALP (Figs 13A–B, F, 14E, 15A–B). Trochanter 1.75, femur 7.13, patella 2.33, chela 7.14, hand 2.43 × as long as broad; femur 2.55 × as long as patella; movable chelal finger 1.88 × as long as hand and 0.64 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb slightly closer to st than to b; b and t situated subdistally, it slightly distal to t and t distal to est; est situated at same level as t, and distal to b (Figs 13A, 15A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 13B, 15B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 24 well-spaced, pointed and slightly retrorse macrodenticles; movable finger with 36 markedly retrorse and almost contiguous pointed teeth.</p><p>OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 5: 5: 5: 4: T2T: 0. Sternal chaetotaxy IV–XII: 12: 8: 8: 10: 9: 9: 8: -: 2. Genital region: sternite II with nine setae scattered on median area, genital opening slit-like, sternite III with row of 14 setae (Fig. 13F).</p><p>LEGS (Fig. 15C–D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.82 × as long as patella; tarsus 2.11 × as long as tibia. Leg IV: femoropatella 3.96 × as long as deep; tibia 6.60 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 4.00 × as long as deep (TS = 0.34), telotarsus 15.40 × as long as deep and 2.75 × as long as basitarsus (TS = 0.28). Setae of leg I (trochanter to tibia) 4: 17: 12: 13, setae of leg IV (trochanter to basitarsus) 1: 4: 7: 11: 14. Arolium not divided, slightly shorter than simple claws.</p><p>Adult females (paratypes; Figs 12B, 13G)</p><p>Mostly same as males; tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4: 5: 6: 4: T2T: 0; sternal chaetotaxy IV–XII: 12: 10:9: 9: 7: 10: 10: -: 2. Genital region: sternite II with 10 setae scattered on median area, sternite III with row of 12 setae.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses)</p><p>Male</p><p>Body length 1.75. Pedipalps: trochanter 0.28/0.16 (1.75), femur 1.07/0.15 (7.13), patella 0.42/0.18 (2.33), chela 1.50/0.21 (7.14), hand 0.51/0.21 (2.43), movable chelal finger length 0.96. Chelicera 0.61/0.25 (2.44), movable finger length 0.33. Carapace 0.56/0.55 (1.02). Leg I: trochanter 0.19/0.11 (1.73), femur 0.62/0.08 (7.75), patella 0.34/0.07 (4.86), tibia 0.28/0.05 (5.60), tarsus 0.59/0.05 (11.80). Leg IV: trochanter 0.24/0.13 (1.85), femoropatella 0.95/0.24 (3.96), tibia 0.66/0.10 (6.60), basitarsus 0.28/0.07 (4.00), telotarsus 0.77/0.05 (15.40).</p><p>Female</p><p>Body length 1.76. Pedipalps: trochanter 0.29/0.17 (1.71), femur 1.08/0.16 (6.75), patella 0.40/0.19 (2.11), chela 1.58/0.24 (6.58), hand 0.53/0.24 (2.21), movable chelal finger length 1.02. Chelicera 0.65/0.25 (2.48), movable finger length 0.36. Carapace 0.64/0.58 (1.10). Leg I: trochanter 0.20/0.14 (1.43), femur 0.64/0.08 (8.00), patella 0.37/0.08 (4.63), tibia 0.30/0.06 (5.00), tarsus 0.70/0.06 (11.67). Leg IV: trochanter 0.24/0.14 (1.71), femoropatella 0.85/0.19 (4.47), tibia 0.57/0.10 (5.70), basitarsus 0.25/0.07 (3.57), telotarsus 0.73/0.05 (14.60).</p><p>Remarks</p><p>Tyrannochthonius laqiuensis sp. nov. differs from all other hypogean species of the genus from China except T. tiani and T. nanxingensis by both chelal fingers lacking intercalary teeth, the presence of four setae on tergites I–III, and carapace without eyes or eyespots. It differs from T. tiani and T. nanxingensis by its shorter and thicker pedipalpal chela (length 1.50 mm (♂), 1.58 mm (♀), 7.14 (♂), 6.58 (♀) × as long as broad compared to length 1.79 mm (♀), 9.42 (♀) × as long as broad in T. tiani and length 1.92–1.97 mm (♂), 1.85 mm (♀), 7.04–7.11 (♂), 7.12 (♀) × as long as broad in T. nanxingensis) and the position of trichobothrium (sb slightly closer to st than to b compared to sb closer to b than st in T. tiani and T. nanxingensis) (Hou et al. 2023b; Sun et al. 2025)</p><p>Distribution</p><p>Known only from the type locality, Laqiu Cave (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E451FF8C6D40FE97FC31FA81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E456FF866D74FAD2FBF3FABF.text	096987E3E456FF866D74FAD2FBF3FABF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius longulimembrum Sun & Guo & Zhang 2026	<div><p>Tyrannochthonius longulimembrum sp. nov.</p><p>urn:lsid:zoobank.org:act: 4304E479-6BBA-47D9-B60C-83FC270A6091</p><p>Figs 1N, 16–20</p><p>Diagnosis</p><p>(♀) Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed, triangular; posterior margin of carapace with 2 setae; tergites I–III each with two setae, tergite IV with four setae. Pedipalps slender, femur 7.69 (♀) × as long as broad, length 1.23 (♀); chela 8.25 (♀) × as long as broad, length 1.65 (♀); both chelal fingers with intercalary teeth; sb midway between st and b.</p><p>Etymology</p><p>The specific name is a noun derived from the Latin words ‘ longulus ’ and ‘ membrum ’, meaning ‘rather long’ and ‘member’, which refers to the elongated pedipalps and legs.</p><p>Vernacular name</p><p>Khƌ伪$.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♀; Guangxi, Baise City, Tiandong County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.39363&amp;materialsCitation.latitude=23.518475" title="Search Plazi for locations around (long 107.39363/lat 23.518475)">Silin Town</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.39363&amp;materialsCitation.latitude=23.518475" title="Search Plazi for locations around (long 107.39363/lat 23.518475)">Guangyang Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.39363&amp;materialsCitation.latitude=23.518475" title="Search Plazi for locations around (long 107.39363/lat 23.518475)">Guangyang Cave</a>; 23.518475° N, 107.393634° E; 363 m a.s.l.; 6 Oct. 2019; Zegang Feng and Lingchen Zhao leg.; MHBU, Ps.- MHBU-GX2019100601.</p><p>Description</p><p>Adult female (Figs 16–20)</p><p>COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale.</p><p>CEPHALOTHORAX (Figs 17E, 18A). Carapace nearly subquadrate, 0.96 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome pointed, triangular; without eyes or eyespots; with 18 setae arranged s4s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with 10 terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 18D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5.</p><p>CHELICERA (Figs 17D, 18B). Almost as long as carapace, 2.30 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 17 teeth, distal one largest; movable finger with 15 contiguous small teeth; galea completely vestigial (Fig. 17D). Serrula exterior with 27 and serrula interior with 11 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 18C).</p><p>PEDIPALP (Figs 17A–B, F, 18E, 19A–B). Trochanter 1.65, femur 7.69, patella 2.56, chela 8.25, hand 3.05 × as long as broad; femur 2.67 × as long as patella; movable chelal finger 1.72 × as long as hand and 0.64 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, t slightly distal to it and est distal to b (Figs 17A, 19A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 17B, 19B). Both chelal fingers with row of teeth, spaced regularly along the margin, teeth smaller distally and proximally: fixed finger with 31 well-spaced, and pointed teeth and 28 intercalary microdenticles; movable finger with 20 well-spaced, pointed teeth, plus 15 intercalary microdenticles and 15 vestigial, rounded and contiguous basal teeth.</p><p>OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 2: 4: 4: 4: 5: 5: 5: 4: T2T: 0. Sternal chaetotaxy IV–XII: 12:8: 9: 9:10:9: 9: -:2. Genital region: sternite II with 10 setae scattered on median area, genital opening slit-like, sternite III with row of 12 setae (Fig. 17C).</p><p>LEGS (Fig. 19C–D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.91 × as long as patella; tarsus 2.00 × as long as tibia. Leg IV: femoropatella 4.30 × as long as deep; tibia 6.09 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 3.33 × as long as deep (TS = 0.33), telotarsus 15.00 × as long as deep and 2.50 × as long as basitarsus (TS = 0.33). Setae of leg I (trochanter to tibia) 3:12: 10: 10, setae of leg IV (trochanter to basitarsus) 1: 3: 7: 10: 10. Arolium not divided, slightly shorter than the simple claws.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses)</p><p>Female</p><p>Body length 2.04. Pedipalps: trochanter 0.33/0.20 (1.65), femur 1.23/0.16 (7.69), patella 0.46/0.18 (2.56), chela 1.65/0.20 (8.25), hand 0.61/0.20 (3.05), movable chelal finger length 1.05. Chelicera 0.76/0.33 (2.30), movable finger length 0.40. Carapace 0.66/0.69 (0.96). Leg I: trochanter 0.19/0.15 (1.27), femur 0.67/0.09 (7.44), patella 0.35/0.08 (4.38), tibia 0.32/0.06 (5.33), tarsus 0.64/0.06 (10.67). Leg IV: trochanter 0.21/0.15 (1.40), femoropatella 0.99/0.23 (4.30), tibia 0.67/0.11 (6.09), basitarsus 0.30/0.09 (3.33), telotarsus 0.75/0.05 (15.00).</p><p>Remarks</p><p>Tyrannochthonius longulimembrum sp. nov. most closely resembles T. ganshuanensis in the presence of two setae on tergites I–II, carapace without eyes or eyespots and both chelal fingers with intercalary teeth, but differs in several characters as follows: longer and thinner pedipalpal chela (length 1.65 mm (♀), 8.25 (♀) × as long as broad compared to length 1.21–1.44 mm (♀), 6.90–7.30 (♀) × as long as broad in T. ganshuanensis) and longer and thinner pedipalpal femur (length 1.23 mm (♀), 7.69 (♀) × as long as broad compared to length 0.95–0.97 mm (♀), 0.95–0.97 (♀) × as long as broad in T. ganshuanensis) (Mahnert 2009).</p><p>Distribution</p><p>Known only from the type locality, Guangyang Cave (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E456FF866D74FAD2FBF3FABF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E45CFFBB6D51FAF4FC37F887.text	096987E3E45CFFBB6D51FAF4FC37F887.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius miridentatus Sun & Guo & Zhang 2026	<div><p>Tyrannochthonius miridentatus sp. nov.</p><p>urn:lsid:zoobank.org:act: 77F8DD22-F3A8-451C-BCA5-079F34901B51</p><p>Figs 1G, 21–24</p><p>Diagnosis</p><p>(♀) Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small and pointed, triangular; posterior margin of carapace with 2 setae; tergites I–IV each with four setae. Pedipalps slender, femur 5.50–5.85 (♀) × as long as broad, length 0.76–0.77 mm (♀); chela 5.60–6.00 (♀) × as long as broad, length 1.12–1.14 mm (♀); both chelal fingers without intercalary teeth; sb slightly closer to st than to b.</p><p>Etymology</p><p>The specific name is an adjective derived from a combination of the Latin words ‘ mirus ’ and ‘ dentatus ’, meaning ‘remarkable’ and ‘toothed’, respectively, which refers to the distinctly heteromorphic chelal teeth.</p><p>Vernacular name</p><p>异Ṅƌ伪$.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♀; Guangxi, Hechi City, Fengshan County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.03722&amp;materialsCitation.latitude=24.565643" title="Search Plazi for locations around (long 107.03722/lat 24.565643)">Fengcheng Town</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.03722&amp;materialsCitation.latitude=24.565643" title="Search Plazi for locations around (long 107.03722/lat 24.565643)">Songren Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=107.03722&amp;materialsCitation.latitude=24.565643" title="Search Plazi for locations around (long 107.03722/lat 24.565643)">Xi’an Cave</a>; 24.565644° N, 107.037218° E; 526 m a.s.l.; 5 Oct. 2019; Zegang Feng and Lingchen Zhao leg.; MHBU, Ps.- MHBU-GX2019100501.</p><p>Paratypes</p><p>CHINA • 2 ♀♀; same data as for holotype; MHBU, Ps.- MHBU-GX2019100502–03 .</p><p>Description</p><p>Adult females (Figs 21–24)</p><p>COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale.</p><p>CEPHALOTHORAX (Figs 22E, 23A). Carapace nearly subquadrate, 0.94–0.96 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small and pointed, triangular; without eyes or eyespots; with 18 setae arranged s4s: 4: 4: 2:2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with seven terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 23D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5.</p><p>CHELICERA (Figs 22D, 23B). Almost as long as carapace, 1.92–2.38 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 10–11 teeth, distal one largest; movable finger with 12–14 contiguous small teeth; galea completely vestigial (Fig. 22D). Serrula exterior with 20–27 and serrula interior with 12 blades. Rallum with nine blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 23C).</p><p>PEDIPALP (Figs 22A–B, F, 23E, 24A–B). Trochanter 1.64–1.69, femur 5.50–5.85, patella 1.88–2.00, chela 5.60–6.00, hand 1.90–2.05 × as long as broad; femur 2.41–2.53 × as long as patella; movable chelal finger 1.87–2.03 × as long as hand and 0.73–0.77 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb slightly closer to st than to b; b and t situated subdistally, it slightly distal to t; est situated at same level as t, and distal to b (Figs 22A, 24A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 22B, 24B). Both chelal fingers with a row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 23–25 well-spaced, and pointed macrodenticles; movable finger with 30–32 contiguous remarkly retrorse and pointed teeth, plus 5–6 vestigial, rounded and contiguous basal teeth.</p><p>OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4:4: 4:4: 4:4: 4–5: 5–6:5–7: 4: T2T:0. Sternal chaetotaxy IV–XII: 12: 9–12:10–11:9–10:10: 10–11: 9–10:-: 2. Genital region: sternite II with 10 setae scattered on median area, genital opening slit-like, sternite III with row of 10–12 setae (Fig. 22C).</p><p>LEGS (Fig. 24C–D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.61–1.87 × as long as patella; tarsus 2.19–2.47 × as long as tibia. Leg IV: femoropatella 3.45– 3.72 × as long as deep; tibia 5.50–5.88 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 3.00–4.00 × as long as deep (TS = 0.29–0.33), telotarsus 10.20–12.25 × as long as deep and 2.40–2.72 × as long as basitarsus (TS = 0.26–0.31). Setae of leg I (trochanter to tibia) 3–4: 10– 11: 10: 9–15, setae of leg IV (trochanter to basitarsus) 2: 3–4: 6–7: 11–12: 9–10. Arolium not divided, slightly shorter than simple claws.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses)</p><p>Females</p><p>Body length 1.52–1.92. Pedipalps: trochanter 0.22–0.23/0.13–0.14 (1.64–1.69), femur 0.76–0.77/0.13– 0.14 (5.50–5.85), patella 0.30–0.32/0.15–0.17 (1.88–2.00), chela 1.12–1.14/0.19–0.20 (5.60–6.00), hand 0.38–0.39/0.19–0.20 (1.90–2.05), movable chelal finger length 0.73–0.77. Chelicera 0.46–0.50/0.21–0.24 (1.92–2.38), movable finger length 0.26–0.28. Carapace 0.46–0.47/0.49 (0.94–0.96). Leg I: trochanter 0.15–0.16/0.09–0.11 (1.36–1.67), femur 0.42–0.44/0.07–0.08 (5.50–6.14), patella 0.23–0.26/0.06 (3.83– 4.33), tibia 0.19–0.22/0.05–0.06 (3.67–4.20), tarsus 0.46–0.49/0.04–0.05 (9.2–11.75). Leg IV: trochanter 0.19–0.21/0.12–0.14 (1.50–1.75), femoropatella 0.65–0.69/0.18–0.20 (3.45–3.72), tibia 0.44–0.47/0.08 (5.50–5.88), basitarsus 0.18–0.20/0.05–0.06 (3.00–4.00), telotarsus 0.48–0.51/0.04–0.05 (10.20–12.25).</p><p>Remarks</p><p>Tyrannochthonius miridentatus sp. nov. differs from all other hypogean species of the genus except T. quattuor by the movable finger of pedipalpal chela exhibits a continuous and strongly retrorse teeth, except for the few most distal ones, sb slightly closer to st than to b, the presence of four setae on tergites I–IV, epistome triangular and carapace without eyes or eyespots. It differs from T. quattuor by its shorter and thicker pedipalpal chela (length 1.12–1.14 mm (♀), 5.60–6.00 (♀) × as long as broad compared to length 1.74–1.86 mm (♀), 6.89–7.25 (♀) × as long as broad in T. quattuor) and shorter and thicker pedipalpal femur (length 0.76–0.77 mm (♀), 5.50–5.85 (♀) × as long as broad compared to compared to length 1.24–1.32 mm (♀), 7.33–7.75 (♀) × as long as broad in T. quattuor) (Hou et al. 2023b).</p><p>Distribution</p><p>Known only from the type locality, Xi’an Cave (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E45CFFBB6D51FAF4FC37F887	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB86ED6FE97FD90FBCD.text	096987E3E462FFB86ED6FE97FD90FBCD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius rudongyanensis Sun, Guo & Zhang 2025	<div><p>Tyrannochthonius rudongyanensis Sun, Guo &amp; Zhang, 2025</p><p>Fig. 1B</p><p>Tyrannochthonius rudongyanensis Sun, Guo &amp; Zhang, 2025: 117–123, figs 2–5.</p><p>Diagnosis and description</p><p>For details see Sun et al. (2025).</p><p>Vernacular name</p><p>乳洞岩ƌ伪$.</p><p>Material examined</p><p>Holotype</p><p>CHINA • ♂; Guangxi, Guilin City, Xing’an County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.620255&amp;materialsCitation.latitude=25.574192" title="Search Plazi for locations around (long 110.620255/lat 25.574192)">Rudongyan Cave</a>; 25.574192° N, 110.620256° E; 240 m a.s.l.; 10 Oct. 2019; Zegang Feng and Lingchen Zhao leg.; MHBU, Ps.- MHBU-GX2019101001.</p><p>Paratypes</p><p>CHINA • 2 ♀♀; same data as for holotype; MHBU, Ps.- MHBU-GX2019101002–03; CHINA • 3 ♀♀; Guangxi, Guilin City, Xing’an County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.620255&amp;materialsCitation.latitude=25.574192" title="Search Plazi for locations around (long 110.620255/lat 25.574192)">Rudongyan Cave</a>; 25.570469° N, 110.627489° E; 213 m a.s.l.; 4 Feb. 2023; Xiangbo Guo, Jianzhou Sun, Tao Zheng and Songtao Shi leg.; MHBU, Ps.- MHBU-GX2023020401–03 .</p><p>Distribution</p><p>Rudongyan Cave (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E462FFB86ED6FE97FD90FBCD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E462FFB26D58FB05FC1BFD26.text	096987E3E462FFB26D58FB05FC1BFD26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius shenlongensis Sun & Guo & Zhang 2026	<div><p>Tyrannochthonius shenlongensis sp. nov.</p><p>urn:lsid:zoobank.org:act: 3F46460A-87B2-4B81-B1B3-F241F830565D</p><p>Figs 1F, 25–29</p><p>Diagnosis</p><p>(♂ ♀) Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, triangular; posterior margin of carapace with 2 setae; tergites I–II each with two setae, tergites III–IV each with four setae. Pedipalps slender, femur 6.36 (♂), 6.00–6.17 (♀) × as long as broad, length 0.70 (♂), 0.66–0.74 (♀); chela 6.71 (♂), 6.71–6.80 (♀) × as long as broad, length 0.0.94 (♂), 0.94–1.02 (♀); both chelal fingers with intercalary teeth; sb closer to st than b.</p><p>Etymology</p><p>Named after the type locality, Shenlong Cave.</p><p>Vernacular name</p><p>DZAEƌ伪$.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♂; Guangxi, Hechi City, Huanjiang Maonan Autonomous County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.38596&amp;materialsCitation.latitude=24.773163" title="Search Plazi for locations around (long 108.38596/lat 24.773163)">Dacai Township</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.38596&amp;materialsCitation.latitude=24.773163" title="Search Plazi for locations around (long 108.38596/lat 24.773163)">Shenlong Cave</a>; 24.773162° N, 108.385965° E; 201.2 m a.s.l.; 31 Jul. 2021; Zegang Feng, Hongru Xu, Liu Fu and Nana Zhan leg.; MHBU, Ps.- MHBU-GX2021073101.</p><p>Paratypes</p><p>CHINA • 2 ♀♀; same data as for holotype; MHBU, Ps.- MHBU-GX2021073102–03 .</p><p>Description</p><p>Adult male (Figs 25–29)</p><p>COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale.</p><p>CEPHALOTHORAX (Figs 26D, 27A). Carapace nearly subquadrate, 1.05 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, triangular; carapace without eyes or eyespots; with 18 setae arranged s4s: 4: 4:2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to the sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with 8–10 terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 27D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5.</p><p>CHELICERA (Figs 26C, 27B). Almost as long as carapace, 2.44 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13 teeth, distal one largest; movable finger with 13 contiguous small teeth; galea completely vestigial (Fig. 26C). Serrula exterior with 23 and serrula interior with 13 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 27C).</p><p>PEDIPALP (Figs 26A–B, E, 27E, 28A–B). Trochanter 1.64, femur 6.36, patella 2.17, chela 6.71, hand 2.50 × as long as broad; femur 2.69 × as long as patella; movable chelal finger 1.74 × as long as hand and 0.65 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb closer to st than b; b and t situated subdistally, t slightly distal to it and est slightly distal to b (Figs 26A, 28A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 26B, 28B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 32 well-spaced, slightly retrorse and pointed teeth and 19 intercalary microdenticles; movable finger with 24 almost contiguous, slightly retrorse and pointed teeth, plus nine intercalary microdenticles and six vestigial, rounded and contiguous basal teeth.</p><p>OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 2: 4: 4: 4: 4: 5: 5: 5: 4: T2T: 0. Sternal chaetotaxy IV–XII: 12: 7:7: 9:8:9: 9:-: 2. Genital region: sternite II with nine setae scattered on median area, genital opening slit-like, sternite III with row of 16 setae (Fig. 26G).</p><p>LEGS (Fig. 28C–D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.94 × as long as patella; tarsus 2.11 × as long as tibia. Leg IV: femoropatella 2.94 × as long as deep; tibia 4.63 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 2.83 × as long as deep (TS = 0.21), telotarsus 14.00 × as long as deep and 2.47 × as long as basitarsus (TS = 0.33). Setae of leg I (trochanter to tibia) 4: 11: 11: 11, setae of leg IV (trochanter to basitarsus) 3:3: 6: 10:8. Arolium not divided, slightly shorter than simple claws.</p><p>Adult females (paratypes; Figs 25B, 26G)</p><p>Mostly same as males; tergal chaetotaxy I–XII: 2: 2: 4: 4: 4: 4: 4: 5: 5: 4: T2T: 0; sternal chaetotaxy IV–XII: 11–12:7–8: 7–8:8:8–9: 9:9: -:2. Genital region: sternite II with nine setae scattered on median area, sternite III with row of 10–12 setae.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses)</p><p>Male</p><p>Body length 1.40. Pedipalps: trochanter 0.20/0.11 (1.82), femur 0.54/0.10 (5.40), patella 0.24/0.12 (2.00), chela 0.79/0.13 (6.08), hand 0.25/0.13 (1.94), movable chelal finger length 0.55. Chelicera 0.40/0.19 (2.10), movable finger length 0.23. Carapace 0.38/0.45 (0.84). Leg I: trochanter 0.12/0.09 (1.33), femur 0.29/0.05 (5.80), patella 0.14/0.05 (2.80), tibia 0.17/0.04 (4.25), tarsus 0.32/0.04 (8.00). Leg IV: trochanter 0.17/0.11 (1.54), femoropatella 0.48/0.19 (2.52), tibia 0.32/0.08 (4.00), basitarsus 0.14/0.06 (2.33), telotarsus 0.34/0.03 (11.33).</p><p>Females</p><p>Body length 1.42–1.45. Pedipalps: trochanter 0.20–0.21/0.11 (1.82–1.91), femur 0.55–0.58/0.10–0.12 (4.83–5.50), patella 0.27/0.12 (2.25), chela 0.82–0.83/0.16 (5.13–5.19), hand 0.26–0.27/0.16 (1.63– 1.69), movable chelal finger length 0.57. Chelicera 0.43–0.45/0.22 (1.95–2.04), movable finger length 0.24–0.25. Carapace 0.39–0.48/0.48–0.49 (0.81–0.98). Leg I: trochanter 0.11–0.13/0.08–0.10 (1.30– 1.38), femur 0.29–0.31/0.06 (4.83–5.17), patella 0.15–0.17/0.05–0.06 (2.83–3.00), tibia 0.16–0.17/0.04 (4.00–4.25), tarsus 0.34–0.35/0.04 (8.50–8.75). Leg IV: trochanter 0.13–0.18/0.10–0.11 (1.30–1.64), femoropatella 0.48–0.49/0.17–0.18 (2.72–2.82), tibia 0.32–0.35/0.08 (4.00–4.38), basitarsus 0.15– 0.16/0.06 (2.50–2.67), telotarsus 0.34–0.35/0.03–0.04 (8.50–11.67).</p><p>Remarks</p><p>Tyrannochthonius shenlongensis sp. nov. most closely resembles T. akaelus in the both chelal fingers with intercalary teeth, tergites I–II each with two setae, tergites III–IV each with four setae, tergite X with four setae; epistome small, triangular and carapace without eyes or eyespots, but differs in several characters as follows: shorter and thicker pedipalpal chela (length 0.94–1.02 mm (♀), 6.71–6.80 (♀) × as long as broad compared to length 1.28 mm (♀), 7.53 (♀) × as long as broad in T. akaelus) and shorter and thicker pedipalpal femur (length 0.66–0.74 mm (♀), 6.00–6.17 (♀) × as long as broad compared to length 0.90 mm (♀), 6.42 (♀) × as long as broad in T. akaelus) and the position of trichobothrium (sb closer to st than b compared to sb situated midway between st and b in T. akaelus) (Mahnert 2009).</p><p>Distribution</p><p>Known only from the type locality, Shenlong Cave (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E462FFB26D58FB05FC1BFD26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB36E8AFE97FDB4FC20.text	096987E3E469FFB36E8AFE97FDB4FC20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius tiani Sun, Guo & Zhang 2025	<div><p>Tyrannochthonius tiani Sun, Guo &amp; Zhang, 2025</p><p>Fig. 1D</p><p>Tyrannochthonius tiani Sun, Guo &amp; Zhang, 2025: 123–127, figs 6–8.</p><p>Diagnosis and description</p><p>For details see Sun et al. (2025).</p><p>Vernacular name</p><p>田氏ƌ伪$.</p><p>Material examined</p><p>Holotype</p><p>CHINA • ♀; Guangxi, Hechi City, Huanjiang County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.03124&amp;materialsCitation.latitude=25.18312" title="Search Plazi for locations around (long 108.03124/lat 25.18312)">The Mulun National Nature Reserve</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.03124&amp;materialsCitation.latitude=25.18312" title="Search Plazi for locations around (long 108.03124/lat 25.18312)">Ganxiao Cave</a>; 25.18312° N, 108.03124° E; 695 m a.s.l.; 8 Dec. 2023; Mingyi Tian leg.; MHBU, Ps.- MHBU-GX2023120801.</p><p>Distribution</p><p>Ganxiao Cave (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E469FFB36E8AFE97FDB4FC20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E469FFB56D55FC70FC23FDB1.text	096987E3E469FFB56D55FC70FC23FDB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius xianguensis Sun & Guo & Zhang 2026	<div><p>Tyrannochthonius xianguensis sp. nov.</p><p>urn:lsid:zoobank.org:act: 714F40F5-E81B-4AC8-89BF-8C6191CC42DA</p><p>Figs 1K, 30–34</p><p>Diagnosis</p><p>(♀) Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome small, triangular; posterior margin of carapace with 2 setae; tergites I–II each with four setae, tergites III–IV each with more four setae. Pedipalps slender, femur 4.89 (♀) × as long as broad, length 0.44 (♀); chela 5.25 (♀) × as long as broad, length 0.63 (♀); both chelal fingers without intercalary teeth; sb closer to st than b.</p><p>Etymology</p><p>Named after the type locality, Xiangu Cave.</p><p>Vernacular name</p><p>仙Rƌ伪$.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♀; Guangxi, Hechi City, Yizhou District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.71361&amp;materialsCitation.latitude=24.501837" title="Search Plazi for locations around (long 108.71361/lat 24.501837)">Xiayan Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.71361&amp;materialsCitation.latitude=24.501837" title="Search Plazi for locations around (long 108.71361/lat 24.501837)">Xiangu Cave</a>; 24.501836° N, 108.713608° E; 124 m a.s.l.; 5 Oct. 2019; Zegang Feng and Lingchen Zhao leg.; MHBU, Ps.- MHBU-GX2019100501.</p><p>Description</p><p>Adult female (Figs 30–34)</p><p>COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale.</p><p>CEPHALOTHORAX (Figs 31E, 32A). Carapace nearly subquadrate, 0.94 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, triangular; without eyes or eyespots; with 18 setae arranged s4s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with a rounded anteromedial process; coxae II with six or eight terminally indented coxal spines, set as oblique and arched row, central spines slightly longer than others (Fig. 32D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5.</p><p>CHELICERA (Figs 31D, 32B). Almost as long as carapace, 1.83 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13 teeth, distal one largest; movable finger with nine contiguous small teeth; galea completely vestigial (Fig. 31D). Serrula exterior with 18 and serrula interior with 12 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 32C).</p><p>PEDIPALP (Figs 31A–B, F, 32E, 33A–B). Trochanter 1.75, femur 4.89, patella 2.11, chela 5.25, hand 1.67 × as long as broad; femur 2.32 × as long as patella; movable chelal finger 2.25 × as long as hand and 0.71 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb closer to st than b; b and t situated subdistally, t slightly distal to it and est distal to b (Figs 31A, 33A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 31B, 33B). Both chelal fingers with row of teeth, spaced regularly along the margin, teeth smaller distally and proximally: fixed finger with 25 well-spaced, slightly retrorse and pointed teeth; movable finger with 11 well-spaced, pointed teeth, plus 12 vestigial, rounded and contiguous basal teeth.</p><p>OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 5: 6: 6: 5: 6: 5: 5: 4: T2T: 0. Sternal chaetotaxy IV–XII: 10:11:9: 9:8: 9: 7:-: 2. Genital region: sternite II with 10 setae scattered on median area, genital opening slit-like, sternite III with row of 12 setae (Fig. 31C).</p><p>LEGS (Fig. 33C–D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.92 × as long as patella; tarsus 2.08 × as long as tibia. Leg IV: femoropatella 2.42 × as long as deep; tibia 4.20 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 3.00 × as long as deep (TS = 0.32), telotarsus 8.00 × as long as deep and 2.00 × as long as basitarsus (TS = 0.29). Setae of leg I (trochanter to tibia) 2: 10: 9: 11, setae of leg IV (trochanter to basitarsus) 2: 3: 6: 11: 10. Arolium not divided, slightly shorter than simple claws.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses)</p><p>Female</p><p>Body length 1.04. Pedipalps: trochanter 0.14/0.08 (1.75), femur 0.44/0.09 (4.89), patella 0.19/0.09 (2.11), chela 0.63/0.12 (5.25), hand 0.20/0.12 (1.67), movable chelal finger length 0.45. Chelicera 0.33/0.18 (1.83), movable finger length 0.18. Carapace 0.32/0.34 (0.94). Leg I: trochanter 0.09/0.07 (1.29), femur 0.23/0.04 (5.75), patella 0.12/0.04 (3.00), tibia 0.12/0.03 (4.00), tarsus 0.25/0.03 (8.33). Leg IV: trochanter 0.11/0.07 (1.57), femoropatella 0.29/0.12 (2.42), tibia 0.29/0.12 (2.42), basitarsus 0.12/0.04 (3.00), telotarsus 0.24/0.03 (8.00).</p><p>Remarks</p><p>Tyrannochthonius xianguensis sp. nov. most closely resembles T. xinzhaiensis by the tergites I–II each with four setae, both chelal fingers without intercalary teeth, trichobothrium sb closer to st than b and carapace without eyes or eyespots, but differs in several characters as follows: shorter and thicker pedipalpal chela (length 0.63 mm (♀), 5.25 (♀) × as long as broad compared to length 1.61–1.64 mm (♀), 6.42–6.44 (♀) × as long as broad in T. xinzhaiensis) and shorter and thicker pedipalpal femur (length 0.44 mm (♀), 4.89 (♀) × as long as broad compared to length 1.14–1.18 mm (♀), 6.71–6.77 (♀) × as long as broad in T. xinzhaiensis) and tergites III–IV each with more four setae (vs tergites III–IV each with four setae in T. xinzhaiensis) (Li 2022).</p><p>Distribution</p><p>Known only from the type locality, Xiangu Cave (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E469FFB56D55FC70FC23FDB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E46FFFAF6D60FDC2FBF5FA51.text	096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius xiannvyanensis Sun & Guo & Zhang 2026	<div><p>Tyrannochthonius xiannvyanensis sp. nov.</p><p>urn:lsid:zoobank.org:act: 732B6785-C1B6-4F71-B9D1-526839238FCB</p><p>Figs 1J, 35–39</p><p>Diagnosis</p><p>(♂ ♀) Moderately sized troglomorphic species with elongated appendages; four eyes, anterior pair of eyes reduced, posterior pair of eyespots; anterior margin of carapace thin, finely denticulated, epistome small, rounded and obtuse; posterior margin of carapace with 2 setae; tergites I–IV each with four setae. Pedipalps slender, femur 6.53–6.80 (♂), 6.75–7.27 (♀) × as long as broad, length 0.98–1.02 (♂), 1.08–1.09 (♀); chela 7.21–7.33 (♂), 6.77–6.91 (♀) × as long as broad, length 1.32–1.37 (♂), 1.45–1.49 (♀); both chelal fingers with intercalary teeth; sb midway between st and b.</p><p>Etymology</p><p>Named after the type locality, Xiannvyan Cave.</p><p>Vernacular name</p><p>仙kƌ伪$.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♂; Guangxi, Hechi City, Yizhou District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.56743&amp;materialsCitation.latitude=24.48801" title="Search Plazi for locations around (long 108.56743/lat 24.48801)">Qingyuan Town</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.56743&amp;materialsCitation.latitude=24.48801" title="Search Plazi for locations around (long 108.56743/lat 24.48801)">Yankou Village</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.56743&amp;materialsCitation.latitude=24.48801" title="Search Plazi for locations around (long 108.56743/lat 24.48801)">Xiannvyan Cave</a>; 24.488011° N, 108.567428° E; 150 m a.s.l.; 4 Oct. 2019; Zegang Feng and Lingchen Zhao leg.; MHBU, Ps.- MHBU-GX2019100401.</p><p>Paratypes</p><p>CHINA • 1 ♂, 2 ♀♀; same data as for holotype; MHBU, Ps.- MHBU-GX2019100402–04 .</p><p>Description</p><p>Adult male (Figs 35–39)</p><p>COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale.</p><p>CEPHALOTHORAX (Figs 36D, 37A). Carapace nearly subquadrate, 1.04 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, rounded and obtuse; four eyes, anterior pair of eyes reduced, posterior pair of eyespots; with 18 setae arranged s4s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to the setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with a rounded anteromedial process; coxae II with 12 terminally indented coxal spines on each side, set as an oblique and arched row, central spines slightly longer than others (Fig. 37D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5.</p><p>CHELICERA (Figs 36C, 37B). Almost as long as carapace, 1.79–1.86 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 9–12 teeth, distal one largest; movable finger with 10–15 contiguous small teeth; galea completely vestigial (Fig. 36C). Serrula exterior with 20–22 and serrula interior with 12–13 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 37C).</p><p>PEDIPALP (Figs 36A–B, F, 37E, 38A–B). Trochanter 1.63–1.81, femur 6.53–6.80, patella 2.29–2.69, chela 7.21–7.33, hand 2.50–2.68 × as long as broad; femur 2.37–2.51 × as long as patella; movable chelal finger 1.71–1.93 × as long as hand and 0.64–0.65 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb midway between st and b; b and t situated subdistally, it slightly distal to t and t distal to est; est slightly distal to b (Figs 36A, 38A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 36B, 38B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 30–34 well-spaced, slightly retrorse and pointed teeth and 25– 26 intercalary microdenticles; movable finger with 20–24 almost contiguous, slightly retrorse and pointed teeth, plus 9– 14 intercalary microdenticles and 6–10 vestigial, rounded and contiguous basal teeth.</p><p>OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4–5: 5–6: 5–6: 4–6: T2T: 0. Sternal chaetotaxy IV–XII: 12: 8–10:6–8: 6–8: 7–9:9: 9: -:2. Genital region: sternite II with 9–10 setae scattered on median area, genital opening slit-like, sternite III with row of 14–16 setae (Fig. 36G).</p><p>LEGS (Fig. 38C–D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.93–2.00 × as long as patella; tarsus 2.25–2.32 × as long as tibia. Leg IV: femoropatella 2.79– 2.90 × as long as deep; tibia 5.27–5.36 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 2.67–2.78 × as long as deep (TS = 0.24–0.33), telotarsus 10.83–13.00 × as long as deep and 2.60–2.71 × as long as basitarsus (TS = 0.18–0.23). Setae of leg I (trochanter to tibia) 3: 10–11: 7–8: 8–10, setae of leg IV (trochanter to basitarsus) 1–2: 3: 6: 8–9: 8–9. Arolium not divided, slightly shorter than simple claws.</p><p>Adult females (paratypes; Figs 35B, 36H)</p><p>Mostly same as males; tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4: 5: 6: 4: T2T: 0; sternal chaetotaxy IV–XII: 10–12: 10–11: 6–8: 8–9: 9–10: 9–10: 9: -:2. Genital region: sternite II with 10 setae scattered on median area, sternite III with row of 12 setae.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses)</p><p>Males</p><p>Body length 1.85–2.13. Pedipalps: trochanter 0.26–0.29/0.16 (1.63–1.81), femur 0.98–1.02/0.15 (6.53– 6.80), patella 0.39–0.43/0.16–0.17 (2.29–2.69), chela 1.32–1.37/0.18–0.19 (7.21–7.33), hand 0.45– 0.51/0.18–0.19 (2.50–2.68), movable chelal finger length 0.87. Chelicera 0.52–0.54/0.29 (1.79–1.86), movable finger length 0.30. Carapace 0.58–0.59/0.56–0.57 (1.04). Leg I: trochanter 0.18–0.21/0.14–0.15 (1.29–1.40), femur 0.52–0.56/0.09 (5.78–6.22), patella 0.26–0.29/0.07–0.08 (3.63–3.71), tibia 0.25– 0.28/0.06 (4.17–4.67), tarsus 0.58–0.63/0.05–0.06 (10.50–11.60). Leg IV: trochanter 0.26–0.27/0.16– 0.18 (1.44–1.69), femoropatella 0.81–0.90/0.29–0.31 (2.79–2.90), tibia 0.58–0.59/0.11 (5.27–5.36), basitarsus 0.24–0.25/0.09 (2.67–2.78), telotarsus 0.65/0.05–0.06 (10.83–13.00).</p><p>Females</p><p>Body length 1.90–2.26. Pedipalps: trochanter 0.29–0.32/0.17 (1.71–1.88), femur 1.08–1.09/0.15–0.16 (6.75–7.27), patella 0.43–0.45/0.18–0.19 (2.37–2.39), chela 1.45–1.49/0.21–0.22 (6.77–6.91), hand 0.53–0.59/0.21–0.22 (2.52–2.68), movable chelal finger length 0.92–0.94. Chelicera 0.59–0.61/0.30– 0.32 (1.84–2.03), movable finger length 0.30–0.31. Carapace 0.63–0.64/0.60–0.61 (1.03–1.07). Leg I: trochanter 0.21–0.22/0.16 (1.32–1.38), femur 0.55–0.56/0.10 (5.50–5.60), patella 0.28–0.29/0.08 (3.50– 3.63), tibia 0.28–0.29/0.07 (4.00–4.14), tarsus 0.63–0.65/0.06 (10.50–10.83). Leg IV: trochanter 0.27– 0.28/0.17–0.18 (1.5–1.65), femoropatella 0.90–0.92/0.30–0.31 (2.96–3.00), tibia 0.53–0.60/0.10–0.12 (5.00–5.30), basitarsus 0.27/0.09 (3.00), telotarsus 0.67–0.68/0.05–0.06 (11.33–13.40).</p><p>Remarks</p><p>Tyrannochthonius xiannvyanensis sp. nov. most closely resembles T. fenghuangensis sp. nov. in the position of trichobothrium sb (sb midway between st and b), the shape of epistome (small, rounded and obtuse) and carapace with four eyes, anterior pair of eyes reduced, posterior pair of eyespots, but differs in several characters as follows: longer and thinner pedipalpal chela (length 1.32–1.37 mm (♂), 1.45–1.49 mm (♀), 7.21–7.33 (♂), 6.77–6.91 (♀) × as long as broad compared to length 0.79 mm (♂), 0.82–0.83 mm (♀), 6.08 (♂), 5.13–5.19 (♀) × as long as broad in T. fenghuangensis) and longer and thinner pedipalpal femur (length 0.98–1.02 mm (♂), 1.08–1.09 (♀), 6.53–6.80 (♂), 6.75–7.27 (♀) × as long as broad compared to length 0.54 mm (♂), 0.55–0.58 mm (♀), 5.40 (♂), 4.83–5.50 (♀) × as long as broad in T. fenghuangensis).</p><p>Distribution</p><p>Known only from the type locality, Xiannvyan Cave (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E46FFFAF6D60FDC2FBF5FA51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E475FFA66D2FFA62FBFCFEC0.text	096987E3E475FFA66D2FFA62FBFCFEC0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius xiaoi Sun & Guo & Zhang 2026	<div><p>Tyrannochthonius xiaoi sp. nov.</p><p>urn:lsid:zoobank.org:act: DECB1715-F328-41F5-971E-22E85EF9BF78</p><p>Figs 1A, 40–44</p><p>Diagnosis</p><p>(♂ ♀) Moderately sized troglomorphic species with elongated appendages; four eyes, anterior pair of eyes reduced, posterior pair of eyespots; anterior margin of carapace thin, finely denticulated, epistome small, triangular; posterior margin of carapace with 2 setae; tergites I–IV each with four setae. Pedipalps slender, femur 4.54–4.75 (♂), 4.57–4.92 (♀) × as long as broad, length 0.56–0.59 (♂), 0.64 (♀); chela 5.23–5.67 (♂), 5.00–5.28 (♀) × as long as broad, length 0.84–0.89 (♂), 0.95 (♀); both chelal fingers with intercalary teeth; sb closer to st than b.</p><p>Etymology</p><p>This species is named after Jiewen Xiao, who participated in field work and collected some of the specimens.</p><p>Vernacular name</p><p>NJ氏ƌ伪$.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♂; Guangxi, Guilin City, Xing’an County, cave without name (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=110.62379&amp;materialsCitation.latitude=25.581533" title="Search Plazi for locations around (long 110.62379/lat 25.581533)">Unnamed Cave</a> 1); 25.581534° N, 110.62378374° E; 229 m a.s.l.; 30 Nov. 2023; Jiewen Xiao leg.; MHBU, Ps.- MHBU-GX2023113002.</p><p>Paratypes</p><p>CHINA • 1 ♂, 2 ♀♀; same data as for holotype; MHBU, Ps.- MHBU-GX2023113003–05 .</p><p>Description</p><p>Adult males (Figs 40–44)</p><p>COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale.</p><p>CEPHALOTHORAX (Figs 41D, 42A). Carapace nearly subquadrate, 0.89–0.90 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, triangular; four eyes, anterior pair of eyes reduced, posterior pair of eyespots; with 18 setae arranged s4s: 4: 4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta less than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with 9–10 terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 42D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5.</p><p>CHELICERA (Figs 41C, 42B). Almost as long as carapace, 1.75–1.83 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 12–13 teeth, distal one largest; movable finger with 12 contiguous small teeth; galea completely vestigial (Fig. 41C). Serrula exterior with 18–19 and serrula interior with 12–14 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 42C).</p><p>PEDIPALP (Figs 41A–B, G, 42E, 43A–B). Trochanter 1.67–1.93, femur 4.54–4.75, patella 2.00–2.23, chela 5.23–5.67, hand 1.65–1.73 × as long as broad; femur 1.93–2.11 × as long as patella; movable chelal finger 2.21–2.31 × as long as hand and 0.70–0.71 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb closer to st than b; b and t situated subdistally, it slightly distal to t; est situated at same level as t, and distal to b (Figs 41A, 43A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 41B, 43B). Both chelal fingers with a row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 24 well-spaced, slightly retrorse and pointed teeth and 17– 22 intercalary microdenticles; movable finger with 10–13 almost contiguous, slightly retrorse and pointed teeth, plus 9– 11 intercalary microdenticles and 9–13 vestigial, rounded and contiguous basal teeth.</p><p>OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4–5: 4–5: 4–6: 6: 6: 6: 5: T2T: 0. Sternal chaetotaxy IV–XII: 12–15: 9–10: 9–10: 9–14: 9–12: 9–13: 9: -: 2. Genital region: sternite II with 7–10 setae scattered on median area, genital opening slit-like, sternite III with row of 16–18 setae (Fig. 41H).</p><p>LEGS (Fig. 43C–D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 1.56–2.00 × as long as patella; tarsus 1.83–2.00 × as long as tibia. Leg IV: femoropatella 2.35– 3.13 × as long as deep; tibia 2.85–4.00 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 2.14–2.67 × as long as deep (TS = 0.33–0.38), telotarsus 8.03–8.75 × as long as deep and 2.06– 2.27 × as long as basitarsus (TS = 0.23–0.27). Setae of leg I (trochanter to tibia) 3–4: 9–10: 7–10: 9–12, setae of leg IV (trochanter to basitarsus) 1–3: 3: 5–6: 8:8–9. Arolium not divided, slightly shorter than simple claws.</p><p>Adult females (paratypes; Figs 40B, 41F, I)</p><p>Mostly same as males; tergal chaetotaxy I–XII: 4: 4:4:5–6: 6: 6: 6:6: 6: 5:T2T: 0; sternal chaetotaxy IV–XII: 14: 11: 11: 13: 14:13: 9: -: 2. Genital region: sternite II with 10 setae scattered on median area, sternite III with row of 12 setae.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses)</p><p>Males</p><p>Body length 1.30–1.66. Pedipalps: trochanter 0.20–0.22/0.11–0.12 (1.67–1.93), femur 0.56–0.59/0.12– 0.13 (4.54–4.75), patella 0.28–0.29/0.13–0.14 (2.00–2.23), chela 0.84–0.89/0.15–0.17 (5.23–5.67), hand 0.26–0.28/0.15–0.17 (1.65–1.73), movable chelal finger length 0.59–0.62. Chelicera 0.41–0.44/0.23– 0.24 (1.75–1.83), movable finger length 0.24–0.26. Carapace 0.41–0.50/0.46–0.48 (0.89–0.90). Leg I: trochanter 0.13–0.15/0.08–0.09 (1.42–1.87), femur 0.28–0.31/0.05–0.07 (4.29–5.43), patella 0.12– 0.18/0.05–0.06 (2.50–3.60), tibia 0.13–0.18/0.04–0.06 (2.82–4.25), tarsus 0.33–0.35/0.04 (8.25–9.03). Leg IV: trochanter 0.15–0.19/0.11–0.12 (1.30–1.64), femoropatella 0.45–0.55/0.15–0.20 (2.35–3.13), tibia 0.25–0.36/0.09 (2.85–4.00), basitarsus 0.12–0.16/0.05–0.07 (2.14–2.67), telotarsus 0.31–0.35/0.04 (8.03–8.75).</p><p>Females</p><p>Body length 1.53–1.80. Pedipalps: trochanter 0.23–0.24/0.13 (1.77–1.91), femur 0.64/0.13–0.14 (4.57– 4.92), patella 0.25–0.32/0.12–0.15 (2.07–2.13), chela 0.95/0.18–0.19 (5.00–5.28), hand 0.31/0.18–0.19 (1.63–1.72), movable chelal finger length 0.65–0.66. Chelicera 0.48–0.49/0.27 (1.78–1.81), movable finger length 0.29. Carapace 0.47/0.54–0.56 (0.84–0.87). Leg I: trochanter 0.14–0.15/0.10–0.12 (1.25– 1.41), femur 0.33–0.35/0.06–0.07 (5.00–5.39), patella 0.15–0.18/0.06 (2.36–3.00), tibia 0.20/0.05 (3.76–4.00), tarsus 0.36–0.37/0.04–0.05 (7.40–9.53). Leg IV: trochanter 0.18–0.20/0.13–0.15 (1.33– 1.38), femoropatella 0.53–0.55/0.12–0.19 (2.89–4.41), tibia 0.32–0.36/0.08–0.09 (4.00–4.15), basitarsus 0.15–0.17/0.06–0.06 (2.50–2.86), telotarsus 0.33–0.38/0.04 (8.25–9.67).</p><p>Remarks</p><p>Tyrannochthonius xiaoi sp. nov. differs from all other hypogean species of the genus from China except T. duo, T. fenghuangensis sp. nov. and T. xiannvyanensis sp. nov. by the four eyes, anterior pair of eyes reduced, posterior pair of eyespots. It differs from T. duo, T. fenghuangensis and T. xiannvyanensis by its shorter and thicker pedipalpal chela (length 0.84–0.89 mm (♂), 0.56–0.59 mm (♀), 5.23–5.67 (♂), 4.54– 4.75 (♀) × as long as broad compared to length 1.11 mm (♂), 1.17–1.18 mm (♀), 6.94 (♂), 5.85–6.56 (♀) × as long as broad in T. duo, and 1.32–1.37 mm (♂), 1.45–1.49 mm (♀), 7.21–7.33 (♂), 6.77–6.91 (♀) × as long as broad in T. xiannvyanensis), the shape of epistome (epistome small, triangular compared to epistome small, rounded and obtuse in T. fenghuangensis and T. xiannvyanensis) and the position of trichobothrium (sb closer to st than b compared to sb midway between st and b in T. fenghuangensis and T. xiannvyanensis and sb closer to b than to st in T. duo) (Hou et al. 2023b).</p><p>Ecology</p><p>These specimens were collected under mud inside the cave (Fig. 44).</p><p>Distribution</p><p>Known only from the type locality, Unnamed Cave 1 (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E475FFA66D2FFA62FBFCFEC0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFA66EB0FE10FDDBFBA6.text	096987E3E47CFFA66EB0FE10FDDBFBA6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius yanwuensis Sun, Guo & Zhang 2025	<div><p>Tyrannochthonius yanwuensis Sun, Guo &amp; Zhang, 2025</p><p>Fig. 1E</p><p>Tyrannochthonius yanwuensis Sun, Guo &amp; Zhang, 2025: 127–131, figs 9–11.</p><p>Diagnosis and description</p><p>For details see Sun et al. (2025).</p><p>Vernacular name</p><p>岩屋ƌ伪$.</p><p>Material examined</p><p>Holotype</p><p>CHINA • ♂; Guangxi, Baise City, Leye County, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.4315&amp;materialsCitation.latitude=24.8239" title="Search Plazi for locations around (long 106.4315/lat 24.8239)">Yanwu Cave</a>; 24.823900° N, 106.431500° E; 695 m a.s.l.; 28 Apr. 2024; Sunbin Huang, Mingzhi Zhao, Yan Li and Rong Chen leg.; MHBU, Ps.- MHBU-GX2024042801.</p><p>Distribution</p><p>Yanwu Cave (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E47CFFA66EB0FE10FDDBFBA6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E47CFFD96D67FBEAFC07FE48.text	096987E3E47CFFD96D67FBEAFC07FE48.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius yilingyanensis Sun & Guo & Zhang 2026	<div><p>Tyrannochthonius yilingyanensis sp. nov.</p><p>urn:lsid:zoobank.org:act: D3D473B6-ED3C-4AA5-8FFE-5F3D0773CDFB</p><p>Figs 45–49</p><p>Diagnosis</p><p>(♂ ♀) Moderately sized troglomorphic species with elongated appendages; four eyes, anterior pair of eyes reduced, posterior pair of eyespots; anterior margin of carapace thin, finely denticulated, epistome small, triangular; posterior margin of carapace with 2 setae; tergites I–IV each with four setae. Pedipalps slender, femur 4.54–4.75 (♂), 4.57–4.92 (♀) × as long as broad, length 0.56–0.59 (♂), 0.64 (♀); chela 5.23–5.67 (♂), 5.00–5.28 (♀) × as long as broad, length 0.84–0.89 (♂), 0.95 (♀); both chelal fingers with intercalary teeth; sb closer to st than b.</p><p>Etymology</p><p>Named after the type locality, Yilingyan Cave.</p><p>Vernacular name</p><p>伊岭岩ƌ伪$.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♂; Guangxi, Nanning City, Wuming District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.29476&amp;materialsCitation.latitude=23.041315" title="Search Plazi for locations around (long 108.29476/lat 23.041315)">Yilingyan Cave</a>; 23.041315° N, 108.294759° E; 96 m a.s.l.; 8 Oct. 2019; Zegang Feng and Lingchen Zhao leg.; MHBU, Ps.- MHBU-GX2019100801.</p><p>Paratypes</p><p>CHINA • 1 ♂, 2 ♀♀; same data as for holotype; MHBU, Ps.- MHBU-GX2019100802–04 .</p><p>Description</p><p>Adult males (Figs 45–49)</p><p>COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale.</p><p>CEPHALOTHORAX (Figs 46D, 47A). Carapace nearly subquadrate, 0.89–0.90 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, triangular; four eyes, anterior pair of eyes reduced, posterior pair of eyespots; with 18 setae arranged s4s: 4:4: 2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I with rounded anteromedial process; coxae II with 7–8 terminally indented coxal spines on each side, set as oblique and arched row, central spines slightly longer than others (Fig. 47D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5.</p><p>CHELICERA (Figs 46C, 47B). Almost as long as carapace, 2.04–2.14 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13–14 teeth, distal one largest; movable finger with 13–15 contiguous small teeth; galea completely vestigial (Fig. 46C). Serrula exterior with 16–21 and serrula interior with 10–12 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 47C).</p><p>PEDIPALP (Figs 46A–B, F, 47E, 48A–B). Trochanter 1.83–1.92, femur 5.15–5.67, patella 1.93–2.21, chela 6.47–6.80, hand 2.20–2.33 × as long as broad; femur 2.19–2.31 × as long as patella; movable chelal finger 1.94–1.97 × as long as hand and 0.67 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb closer to st than b; b and t situated subdistally, it situated at same level as t; est slightly distal to b (Figs 46A, 48A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 46B, 48B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 28–33 well-spaced, slightly retrorse and pointed teeth and 26– 27 intercalary microdenticles; movable finger with 15–20 almost contiguous, slightly retrorse and pointed teeth, plus 15– 16 intercalary microdenticles and 14–20 vestigial, rounded and contiguous basal teeth.</p><p>OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 4: 4: 4: 4: 4: 4: 4: 5: 5: 4: T2T: 0. Sternal chaetotaxy IV–XII: 12: 10: 9: 9: 9: 9: 9: -: 2. Genital region: sternite II with nine setae scattered on median area, genital opening slit-like, sternite III with row of 14 setae (Fig. 46G).</p><p>LEGS (Fig. 48C–D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 2.05–2.17 × as long as patella; tarsus 2.00–2.21 × as long as tibia. Leg IV: femoropatella 2.60– 2.86 × as long as deep; tibia 4.67–5.00 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 3.00–3.33 × as long as deep (TS = 0.24), telotarsus 10.83–13.00 × as long as deep and 2.28– 2.35 × as long as basitarsus (TS = 0.32–0.33). Setae of leg I (trochanter to tibia) 3–4: 11–12: 8:9–12, setae of leg IV (trochanter to basitarsus) 2–3: 3: 5–8: 9–10: 8–9. Arolium not divided, slightly shorter than simple claws.</p><p>Adult females (paratypes; Figs 45B, 46H)</p><p>Mostly same as males; tergal chaetotaxy I–XII: 4: 4: 4: 5–6: 6: 6: 6: 6: 6:5:T2T:0; sternal chaetotaxy IV– XII: 14: 11: 11:13: 14: 13: 9: -: 2. Genital region: sternite II with 10 setae scattered on median area, sternite III with row of 12 setae.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses)</p><p>Males</p><p>Body length 1.39–1.45. Pedipalps: trochanter 0.22–0.23/0.12 (1.83–1.92), femur 0.67–0.68/0.12–0.13 (5.15–5.67), patella 0.29–0.31/0.14–0.15 (1.93–2.21), chela 0.97–1.02/0.15 (6.47–6.80), hand 0.33– 0.35/0.15 (2.20–2.33), movable chelal finger length 0.65–0.68. Chelicera 0.47–0.49/0.22–0.24 (2.04– 2.14), movable finger length 0.24–0.25. Carapace 0.44–0.45/0.50–0.51 (0.88). Leg I: trochanter 0.14– 0.15/0.09 (1.56–1.67), femur 0.39–0.41/0.07 (5.57–5.86), patella 0.18–0.20/0.06–0.07 (2.57–3.33), tibia 0.19–0.21/0.04–0.05 (3.80–5.25), tarsus 0.42/0.04 (10.50). Leg IV: trochanter 0.18–0.20/0.11 (1.64– 1.82), femoropatella 0.63–0.65/0.22–0.25 (2.60–2.86), tibia 0.42–0.45/0.09 (4.67–5.00), basitarsus 0.20–0.21/0.06–0.07 (3.00–3.33), telotarsus 0.47–0.48/0.04 (11.75–12.00).</p><p>Females</p><p>Body length 1.39–1.53. Pedipalps: trochanter 0.25/0.13–0.14 (1.78–1.92), femur 0.72–0.76/0.14–0.15 (5.07–5.14), patella 0.32–0.33/0.16–0.17 (1.94–2.00), chela 1.06–1.09/0.18–0.19 (5.58–6.05), hand 0.36–0.38/0.18–0.19 (1.89–2.11), movable chelal finger length 0.71–0.73. Chelicera 0.52–0.55/0.25– 0.27 (2.04–2.08), movable finger length 0.27–0.29. Carapace 0.47–0.54/0.54–0.56 (0.84–1.00). Leg I: trochanter 0.14–0.17/0.10–0.12 (1.40–1.42), femur 0.40–0.43/0.07–0.08 (5.38–5.71), patella 0.21– 0.22/0.07 (3.00–3.14), tibia 0.22/0.05–0.06 (3.67–4.40), tarsus 0.44–0.45/0.04–0.05 (9.00–11.00). Leg IV: trochanter 0.20–0.21/0.13–0.14 (1.50–1.54), femoropatella 0.66–0.68/0.23–0.24 (2.83–2.87), tibia 0.46/0.08–0.10 (4.60–5.75), basitarsus 0.21/0.07–0.08 (2.63–3.00), telotarsus 0.49/0.04–0.05 (9.80– 12.25).</p><p>Remarks</p><p>Tyrannochthonius yilingyanensis sp. nov. most closely resembles T. xiaoi sp. nov. in the both chelal fingers with intercalary teeth, trichobothrium sb closer to st than b, epistome small, triangular, tergites I–IV each with four setae and four eyes, anterior pair of eyes reduced, posterior pair of eyespots, but differs in several characters as follows: longer and thinner pedipalpal chela (length 0.97–1.02 mm (♂), 1.06–1.09 mm (♀), 6.47–6.80 (♂), 5.58–6.05 (♀) × as long as broad compared to length 0.84–0.89 mm (♂), 0.95 mm (♀), 5.23–5.67 (♂), 5.00–5.28 (♀) × as long as broad in T. xiaoi) and longer and thinner pedipalpal femur (length 0.67–0.68 mm (♂), 0.72–0.76 (♀), 5.15–5.67 (♂), 5.07–5.14 (♀) × as long as broad compared to length 0.56–0.59 mm (♂), 0.64 mm (♀), 4.54–4.75 (♂), 4.57–4.92 (♀) × as long as broad in T. xiaoi).</p><p>Distribution</p><p>Known only from the type locality, Yilingyan Cave (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E47CFFD96D67FBEAFC07FE48	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E403FFD36D1BFD88FBFCFC1D.text	096987E3E403FFD36D1BFD88FBFCFC1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius yui Sun & Guo & Zhang 2026	<div><p>Tyrannochthonius yui sp. nov.</p><p>urn:lsid:zoobank.org:act: 219044D3-5956-477A-B9C0-CF290452B267</p><p>Figs 1M, 50–54</p><p>Diagnosis</p><p>(♂) Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed, triangular; posterior margin of carapace with 2 setae; tergites I–II each with two setae, tergite III–IV each with four setae. Pedipalps slender, femur 5.00 (♂) × as long as broad, length 0.40 (♂); chela 6.44 (♂) × as long as broad, length 0.58 (♂); both chelal fingers without intercalary teeth; sb closer to st than b.</p><p>Etymology</p><p>This species is named after Kun Yu, who collected this specimen.</p><p>Vernacular name</p><p>Ḕ氏ƌ伪$.</p><p>Type material</p><p>Holotype</p><p>CHINA • ♂; Guangxi, Liuzhou City, Yufeng District, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.49899&amp;materialsCitation.latitude=24.20261" title="Search Plazi for locations around (long 109.49899/lat 24.20261)">Liyong Town</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.49899&amp;materialsCitation.latitude=24.20261" title="Search Plazi for locations around (long 109.49899/lat 24.20261)">Unnamed Cave</a> 2; 24.20260909° N, 109.49899591° E; 94 m a.s.l.; 9 Aug. 2023; Kun Yu leg.; MHBU, Ps.- MHBU-GX2023080901.</p><p>Description</p><p>Adult male (Figs 50–54)</p><p>COLOR. Generally pale yellow, chelicerae, carapace, pedipalps and tergites slightly darker black, soft parts pale.</p><p>CEPHALOTHORAX (Figs 51E, 52A). Carapace nearly subquadrate, 0.94 × as long as broad, weakly constricted basally; posterior region with squamous sculpturing laterally, other area smooth, without furrows; anterior margin slightly serrate; epistome small, triangular; without eyes or eyespots; with 18 setae arranged s4s: 4: 4:2: 2, most setae acuminate, sturdy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, first and second pair situated middle and flank to setae of ocular row, third pair situated lateral to sole pair of setae of posterior row. Manducatory process with two acuminate distal setae, anterior seta more than ½ as long as medial seta; apex of coxa I rounded anteromedial process; coxae II with seven or eight terminally indented coxal spines, set as oblique and arched row, central spines slightly longer than others (Fig. 52D); intercoxal tubercle absent; chaetotaxy of coxae: P 3, I 3, II 4, III 5, IV 5.</p><p>CHELICERA (Figs 51D, 52B). Almost as long as carapace, 1.93 × as long as broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal setae shorter than others; movable finger with one medial seta. Cheliceral hand with moderate wrinkle on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with nine teeth, distal one largest; movable finger with 10 contiguous small teeth; galea completely vestigial (Fig. 51D). Serrula exterior with 17 and serrula interior with 11 blades. Rallum with eight blades, distal one longest, with fine barbules and slightly set apart from other blades, latter tightly grouped and with long pinnae, some of which subdivided (Fig. 52C).</p><p>PEDIPALP (Figs 51A–B, F, 52E, 53A–B). Trochanter 1.75, femur 5.00, patella 2.00, chela 6.44, hand 2.11 × as long as broad; femur 2.22 × as long as patella; movable chelal finger 2.05 × as long as hand and 0.67 × as long as chela. Setae generally long and acuminate. Chelal hand not constricted towards fingers, apodeme complex of movable chelal finger slightly sclerotized. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib and isb situated close together, submedially on dorsum of chelal hand; eb, esb and ist at base of fixed chelal finger; esb slightly distal eb and ist slightly distal to esb; it slightly distal to est, situated subdistally; et slightly near to tip of fixed chelal finger, slightly close to chelal teeth; dx situated distal to et; sb closer to st than b; b and t situated subdistally, it slightly distal to t; est situated at same level as t and distal to b (Figs 51A, 53A). Microsetae (chemosensory setae) present on dorsum of chelal hand (Figs 51B, 53B). Both chelal fingers with row of teeth, spaced regularly along margin, teeth smaller distally and proximally: fixed finger with 25 well-spaced, slightly retrorse and pointed teeth and 12 intercalary microdenticles; movable finger with 10 almost contiguous, slightly retrorse and pointed teeth, plus nine intercalary microdenticles and nine vestigial, rounded and contiguous basal teeth.</p><p>OPISTHOSOMA. Generally typical, pleural membrane finely granulated. All tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I–XII: 2: 4: 4: 4: 4: 5: 5: 5: 5: 4: T2T: 0. Sternal chaetotaxy IV–XII: 12: 8: 7: 7: 6: 7: 7: -: 2. Genital region: sternite II with 10 setae scattered on median area, genital opening slit-like, sternite III with row of 14 setae (Fig. 51C).</p><p>LEGS (Fig. 53C–D). Fine granulation present on anterodorsal faces of femur IV and patella IV. Leg I: femur 2.09 × as long as patella; tarsus 2.00 × as long as tibia. Leg IV: femoropatella 2.57 × as long as deep; tibia 5.00 × as long as deep; with basal tactile setae on both tarsal segments: basitarsus 3.00 × as long as deep (TS = 0.29), telotarsus 8.33 × as long as deep and 2.08 × as long as basitarsus (TS = 0.32). Setae of leg I (trochanter to tibia) 4: 10: 8: 9, setae of leg IV (trochanter to basitarsus) 2: 3: 6: 9: 10. Arolium not divided, slightly shorter than simple claws.</p><p>Dimensions (length/breadth or, in the case of the legs, length/depth in mm; ratios in parentheses)</p><p>Male</p><p>Body length 0.98. Pedipalps: trochanter 0.14/0.08 (1.75), femur 0.40/0.08 (5.00), patella 0.40/0.08 (5.00), chela 0.58/0.09 (6.44), hand 0.19/0.09 (2.11), movable chelal finger length 0.39. Chelicera 0.29/0.15 (1.93), movable finger length 0.17. Carapace 0.31/0.33 (0.94). Leg I: trochanter 0.08/0.04 (2.00), femur 0.23/0.04 (5.75), patella 0.11/0.04 (2.75), tibia 0.12/0.03 (4.00), tarsus 0.24/0.03 (8.00). Leg IV: trochanter 0.11/0.08 (1.38), femoropatella 0.36/0.14 (2.57), tibia 0.25/0.05 (5.00), basitarsus 0.12/0.04 (3.00), telotarsus 0.25/0.03 (8.33).</p><p>Remarks</p><p>Tyrannochthonius yui sp. nov. most closely resembles T. yanwuensis by the tergite I with two setae, tergite III–IV and X each with four setae, both chelal fingers with intercalary teeth and carapace without eyes or eyespots, but differs in several characters as follows: tergite II with four setae (vs tergite II with two setae in T. yanwuensis), shorter pedipalpal chela (length 0.58 mm (♂) compared to length 1.31 mm (♂) in T. yanwuensis) and shorter pedipalpal femur (length 0.40 mm (♂) compared to length 0.94 mm (♂) in T. yanwuensis) (Sun et al. 2025).</p><p>Ecology</p><p>The specimen was collected from bat guano inside the cave (Fig. 54).</p><p>Distribution</p><p>Known only from the type locality, Unnamed Cave 2 (Guangxi, China).</p></div>	https://treatment.plazi.org/id/096987E3E403FFD36D1BFD88FBFCFC1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
096987E3E409FFD76FC5FC29FA83FA41.text	096987E3E409FFD76FC5FC29FA83FA41.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tyrannochthonius Chamberlin 1929	<div><p>Key to the species of Tyrannochthonius Chamberlin, 1929 from China (subspecies not included)</p><p>1. Carapace with eyes or eyespots ........................................................................................................ 2</p><p>– Carapace without eyes or eyespots ................................................................................................. 12</p><p>2. Trichobothrium sb closer to b than to st or midway between st and b ............................................. 3</p><p>– Trichobothrium sb closer to st than to b ........................................................................................... 6</p><p>3. Trichobothrium sb midway between st and b; epistome small, rounded and obtuse ....................... 4</p><p>– Trichobothrium sb closer to b than to st; epistome pointed, triangular ............................................ 5</p><p>4. Pedipalpal chela length 0.72 mm (♂), 0.82–0.83 mm (♀), 6.08 (♂), 5.13–5.19 (♀) × as long as broad; palpal femur length 0.54 mm (♂), 0.55–0.58 mm (♀), 5.40 (♂), 4.83–5.50 (♀) × as long as broad ........................................................................................................ T. fenghuangensis sp. nov.</p><p>– Pedipalpal chela length 1.32–1.37 mm (♂), 1.45–1.49 mm (♀), 7.21–7.33 (♂), 6.77–6.91 (♀) × as long as broad; palpal femur length 0.98–1.02 mm (♂), 1.08–1.09 mm (♀), 6.53–6.80 (♂), 6.75–7.27 (♀) × as long as broad ............................................................................... T. xiannvyanensis sp. nov.</p><p>5. Pedipalpal chela length 1.11 mm (♂), 1.17–1.18 mm (♀), 6.94 (♂), 5.85–6.56 (♀) × as long as broad; palpal femur length 0.76 mm (♂), 0.79–0.81 mm (♀), 5.85 (♂), 5.79–6.08 (♀) × as long as broad .............................................................................................. T. duo Hou, Feng &amp; Zhang, 2023</p><p>– Pedipalpal chela length 1.60–1.78 mm (♂), 1.57–1.64 mm (♀), 7.27–7.42 (♂), 6.28–6.83 (♀) × as long as broad; palpal femur length 1.10–1.21 mm (♂), 1.10–1.18 mm (♀), 6.72–6.88 (♂), 6.47–6.94 (♀) × as long as broad ............................................................ T. infirmus Hou, Feng &amp; Zhang, 2023</p><p>6. Carapace with 16 setae ......................................................................... T. japonicus Ellingsen, 1907</p><p>– Carapace with 18 setae ..................................................................................................................... 7</p><p>7. Tergites I and II each with six setae; both chelal fingers without intercalary teeth ............................ ........................................................................................................ T. pachythorax Redikorzev, 1938</p><p>– Tergites I and II each with four setae; both chelal fingers with intercalary teeth ............................. 8</p><p>8. Carapace with two anterior corneate eyes only ......................... T. capito Hou, Feng &amp; Zhang, 2023</p><p>– Carapace with four eyes .................................................................................................................... 9</p><p>9. Carapace with four corneate eyes .................................................................. T. robustus Beier, 1951</p><p>– Carapace with four eyes, anterior pair of eyes well-developed, posterior pair of eyespots ........... 10</p><p>10. Epistome small, rounded and obtuse ................................................................ T. baiseensis sp. nov.</p><p>– Epistome triangular ..........................................................................................................................11</p><p>11. Pedipalpal chela length 0.84–0.89 mm (♂), 0.95 mm (♀), 5.23–5.67 (♂), 5.00–5.28 (♀) × as long as broad; palpal femur length 0.56–0.59 mm (♂), 0.64 mm (♀), 4.54–4.75 (♂), 4.57–4.92 (♀) × as long as broad .............................................................................................................. T. xiaoi sp. nov.</p><p>– Pedipalpal chela length 0.97–1.02 mm (♂), 1.06–1.09 mm (♀), 6.47–6.80 (♂), 5.58–6.05 (♀) × as long as broad; palpal femur length 0.67–0.68 mm (♂), 0.72–0.76 mm (♀), 5.15–5.67 (♂), 5.07–5.14 (♀) × as long as broad ................................................................................. T. yilingyanensis sp. nov.</p><p>12. At least one finger of chela with intercalary teeth .......................................................................... 13</p><p>– Neither chelal finger with intercalary teeth .................................................................................... 37</p><p>13. Only one finger of chela with intercalary teeth .............................................................................. 14</p><p>– Both chelal fingers with intercalary teeth ....................................................................................... 18</p><p>14. Only movable finger of chela with intercalary teeth .............. T. calvatus Hou, Feng &amp; Zhang, 2023</p><p>– Only fixed finger of chela with intercalary teeth ............................................................................ 15</p><p>15. Trichobothrium sb midway between st and b ................ T. parcidentatus Hou, Feng &amp; Zhang, 2023</p><p>– Trichobothrium sb closer to b than to st ......................................................................................... 16</p><p>16. Epistome absent ............................................................................ T. zhai Gao, Zhang &amp; Chen, 2020</p><p>– Epistome triangular ......................................................................................................................... 17</p><p>17. Tergites I–IV each with four setae ........................................ T. chixing Gao, Wynne &amp; Zhang, 2018</p><p>– Tergites I–IV each with two setae ............................................ T. gracilis Hou, Feng &amp; Zhang, 2023</p><p>18. Trichobothrium sb closer to st than to b ......................................................................................... 19</p><p>– Trichobothrium sb closer to b than to st or midway between st and b ........................................... 27</p><p>19. Epistome small, rounded and obtuse .............................................................................................. 20</p><p>– Epistome pointed, triangular ........................................................................................................... 21</p><p>20. Body size 1.62–1.69 (♀) mm; pedipalpal chela 6.07–6.20 (♀) × as long as broad; tergite V with four setae, tergite X with three setae; fixed finger of pedipalpal chela with 25– 26 intercalary teeth; rallum with eight blades ...................................................................... T. arificus Hou, Feng &amp; Zhang, 2023</p><p>– Body size 1.29–1.33 (♀) mm; pedipalpal chela 5.69–5.75 (♀) × as long as broad; tergite V with five setae, tergite X with five setae; fixed finger of pedipalpal chela with 14 intercalary teeth; rallum with seven blades ................................................................... T. yanshanensis Hou, Feng &amp; Zhang, 2023</p><p>21. Tergite II with two setae ................................................................................................................. 22</p><p>– Tergite II with four setae ................................................................................................................. 24</p><p>22. Tergite IV with two setae ..................................................... T. qilinensis Hou, Feng &amp; Zhang, 2023</p><p>– Tergite IV with four setae ............................................................................................................... 23</p><p>23. Pedipalpal chela length 1.28 mm (♀), 7.53 (♀) × as long as broad; palpal femur length 0.90 mm (♀), 6.42 (♀) × as long as broad ........................................................................ T. akaelus Mahnert, 2009</p><p>– Pedipalpal chela length 0.94–1.02 mm (♀), 6.71–6.80 (♀) × as long as broad; palpal femur length 0.66–0.74 mm (♀), 6.00–6.17 (♀) × as long as broad ................................ T. shenlongensis sp. nov.</p><p>24. Pedipalpal chela length min. 1.40 mm (♂); palpal femur length min. 0.90 mm (♂) ..................... 25</p><p>– Pedipalpal chela length less than 1.40 mm (♂); palpal femur length less than 0.90 mm (♂) ........ 26</p><p>25. Pedipalpal chela length 1.75–1.76 mm (♂), 1.77–1.87 mm (♀), 7.29–7.65 (♂), 7.38–7.79 (♀) × as long as broad; palpal femur length 1.20–1.25 mm (♂), 1.25–1.26 mm (♀), 6.58–6.67 (♂), 6.63–6.94 (♀) × as long as broad ............................................................... T. acutus Hou, Feng &amp; Zhang, 2023</p><p>– Pedipalpal chela length 1.45–1.46 mm (♂), 7.63–7.66 (♂) × as long as broad; palpal femur length 0.97–0.99 mm (♂), 6.06–6.07 (♂) × as long as broad ................................ T. yamuheensis Li, 2022</p><p>26. Pedipalpal chela length 1.21–1.27 mm (♂), 1.24–1.29 mm (♀), 6.68–6.72 (♂), 6.20–6.79 (♀) × as long as broad; palpal femur length 0.83–0.87 mm (♂), 0.86–0.89 mm (♀), 5.80–5.93 (♂), 5.93–6.14 (♀) × as long as broad ............................................................ T. hispidus Hou, Feng &amp; Zhang, 2023</p><p>– Pedipalpal chela length 1.01–1.09 mm (♂), 1.21–1.25 mm (♀), 5.61–5.66 (♂), 6.37–6.40 (♀) × as long as broad; palpal femur length 0.73–0.76 mm (♂), 0.80–0.83 mm (♀), 4.87–4.90 (♂), 5.33–5.37 (♀) × as long as broad .......................................................................... T. huilongshanensis Li, 2022</p><p>27. Trichobothrium sb midway between st and b ................................................................................. 28</p><p>– Trichobothrium sb closer to b than to st ......................................................................................... 32</p><p>28. Tergite I with two setae ................................................................................................................... 29</p><p>– Tergite I with four setae .................................................................................................................. 30</p><p>29. Tergites II and X with four setae; pedipalpal chela length 0.58 mm (♂); palpal femur length 0.40 mm (♂) ................................................................................................................................. T. yui sp. nov.</p><p>– Tergites II and X with two setae; pedipalpal chela length 1.31 mm (♂); palpal femur length 0.94 mm (♂) ....................................................................................... T. yanwuensis Sun, Guo &amp; Zhang, 2025</p><p>30. Epistome small, rounded and obtuse; pedipalpal chela length 0.74–0.76 mm (♂), 0.78–0.79 mm (♀); palpal femur length 0.48–0.51 mm (♂), 0.53–0.55 mm (♀) .............................................................. ............................................................................................. T. breviculus Hou, Feng &amp; Zhang, 2023</p><p>– Epistome small, pointed; pedipalpal chela length min. 1.40 mm (♂), 1.40 mm (♀); palpal femur length min. 0.95 mm (♂), 1.00 mm (♀) ......................................................................................... 31</p><p>31. Pedipalpal chela length 1.80–1.98 mm (♂), 1.93–2.00 mm (♀), 8.25–8.57 (♂), 7.41–7.42 (♀) × as long as broad; palpal femur length 1.23–1.36 mm (♂), 1.35–1.37 mm (♀), 8.50–8.79 (♂), 8.06–8.44 (♀) × as long as broad .................................................................. T. altus Hou, Feng &amp; Zhang, 2023</p><p>– Pedipalpal chela length 1.43–1.49 mm (♂), 1.44 mm (♀), 6.81–7.45 (♂), 6.55 (♀) × as long as broad; palpal femur length 0.97–1.02 mm (♂), 1.00– 1.02 mm (♀), 6.47–6.80 (♂), 6.67–6.80 (♀) × as long as broad ............................................................................... T. maculosus Hou, Feng &amp; Zhang, 2023</p><p>32. Tergites I and II each with two setae .............................................................................................. 33</p><p>– Tergites I and II each with four setae .............................................................................................. 34</p><p>33. Pedipalpal chela length 1.21–1.44 mm (♀), 6.90–7.30 (♀) × as long as broad; palpal femur length 0.95–0.97 mm (♀), 5.90–6.70 (♀) × as long as broad ................. T. ganshuangensis Mahnert, 2009</p><p>– Pedipalpal chela length 1.65 mm (♀), 8.25 (♀) × as long as broad; palpal femur length 1.23 mm (♀), 7.69 (♀) × as long as broad ................................................................... T. longulimembrum sp. nov.</p><p>34. The movable finger of the pedipalpal chela with almost upright teeth ............................................... ...................................................................................... T. rudongyanensis Sun, Guo &amp; Zhang, 2025</p><p>– The movable finger of the pedipalpal chela with almost retrorse teeth .......................................... 35</p><p>35. Tergite X with two setae ............................................. T. babaowanensis Hou, Feng &amp; Zhang, 2023</p><p>– Tergite X with four or five setae ..................................................................................................... 36</p><p>36. The movable finger of the pedipalpal chela with slightly retrorse teeth.............................................. .......................................................................................................... T. antridraconis Mahnert, 2009</p><p>– The movable finger of the pedipalpal chela with remarkly retrorse teeth ........................................... ....................................................................................... T. multidentatus Hou, Feng &amp; Zhang, 2023</p><p>37. Tergite I with two setae ................................................................................................................... 38</p><p>– Tergite I with four setae .................................................................................................................. 47</p><p>38. Trichobothrium sb closer to b than to st or midway between st and b ............................................ 39</p><p>– Trichobothrium sb closer to st than to b ......................................................................................... 45</p><p>39. Trichobothrium sb midway between st and b ................................................................................. 40</p><p>– Trichobothrium sb closer to b than to st ......................................................................................... 42</p><p>40. Tergite III with four setae ................................................. T. brevispinus Hou, Feng &amp; Zhang, 2023</p><p>– Tergite III with two setae ................................................................................................................ 41</p><p>41. Tergite IV with four setae; epistome small, rounded and obtuse T. harveyi Gao, Zhang &amp; Chen, 2020</p><p>– Tergite IV with two setae; epistome absent ................................................... T. dongjiensis Li, 2022</p><p>42. Tergite III with three or four setae; tergite X with four setae .............................................................. ............................................................................................... T. oblongus Hou, Feng &amp; Zhang, 2023</p><p>– Tergite III with two setae; tergite X with two setae ........................................................................ 43</p><p>43. Epistome small, rounded .................................................... T. multicaves Hou, Feng &amp; Zhang, 2023</p><p>– Epistome absent .............................................................................................................................. 44</p><p>44. Pedipalpal chela 8.44–9.06 (♂), 7.75–8.33 (♀) × as long as broad; coxae II with 12–13 terminally indented coxal spines ............................................................... T. umidus Hou, Feng &amp; Zhang, 2023</p><p>– Pedipalpal chela 7.45–7.95 (♂), 7.52 (♀) × as long as broad; coxae II with 8–9 terminally indented coxal spines .............................................................................. T. pinguis Hou, Feng &amp; Zhang, 2023</p><p>45. Epistome pointed, triangular; tergite X with two setae ........... T. pandus Hou, Feng &amp; Zhang, 2023</p><p>– Epistome absent; tergite X with four setae ..................................................................................... 46</p><p>46. Chelal fixed finger with 20–21 teeth, movable finger with 25–26 teeth; rallum with seven blades ... ................................................................................................... T. planus Hou, Feng &amp; Zhang, 2023</p><p>– Chelal fixed finger with 30–32 teeth, movable finger with 25–26 teeth; rallum with eight blades .... .................................................................................................... T. pictus Hou, Feng &amp; Zhang, 2023</p><p>47. Trichobothrium sb closer to b than to st or midway between st and b ........................................... 48</p><p>– Trichobothrium sb closer to st than to b ......................................................................................... 50</p><p>48. Trichobothrium sb midway between st and b ................................................... T. laqiuensis sp. nov.</p><p>– Trichobothrium sb closer to b than to st ......................................................................................... 49</p><p>49. Chelal movable finger with slightly heterodont teeth; pedipalpal chela length 1.85 mm (♀), 7.12 (♀) × as long as broad; palpal femur length 1.24 mm (♀), 7.75 (♀) × as long as broad ................... ........................................................................................ T. nanxingensis Hou, Feng &amp; Zhang, 2023</p><p>– Chelal movable finger with remarkly heterodont teeth; pedipalpal chela length 1.79 mm (♀), 9.42 (♀) × as long as broad; palpal femur length 1.32 mm (♀), 9.43 (♀) × as long as broad ................... ........................................................................................................ T. tiani Sun, Guo &amp; Zhang, 2025</p><p>50. Epistome small, rounded and obtuse .............................................................................................. 51</p><p>– Epistome small, pointed .................................................................................................................. 52</p><p>51. Pedipalpal chela length 1.61–1.64 mm (♂), 1.56–1.58 mm (♀), 7.00–7.07 (♂), 8.67–8.69 (♀) × as long as broad; palpal femur length 1.16–1.19 mm (♂), 1.11–1.17 mm (♀), 8.92–8.95 (♂), 8.54–8.59 (♀) × as long as broad ..................................................................................... T. huaerensis Li, 2022</p><p>– Pedipalpal chela length 2.14–2.34 mm (♂), 2.37–2.46 mm (♀), 6.69–7.13 (♂), 6.15–6.58 (♀) × as long as broad; palpal femur length 1.58–1.68 mm (♂), 1.67–1.75 mm (♀), 9.29–9.33 (♂), 8.75–8.79 (♀) × as long as broad .................................................................. T. latus Hou, Feng &amp; Zhang, 2023</p><p>52. The movable finger of the pedipalpal chela with continuous and strongly retrorse teeth, except for the few most distal ones .................................................................................................................. 53</p><p>– The movable finger of the pedipalpal chela with well-spaced, slightly or upright teeth ................ 54</p><p>53. Pedipalpal chela length 1.12–1.14 mm (♀), 5.60–6.00 (♀) × as long as broad; palpal femur length 0.76–0.77 mm (♀), 5.50–5.85 (♀) × as long as broad .................................. T. miridentatus sp. nov.</p><p>– Pedipalpal chela length 1.74–1.86 mm (♀), 6.89–7.25 (♀) × as long as broad; palpal femur length 1.24–1.32 mm (♀), 7.33–7.75 (♀) × as long as broad ........... T. quattuor Hou, Feng &amp; Zhang, 2023</p><p>54. Pedipalpal chela length 0.63 mm (♀), 5.25 (♀) × as long as broad; palpal femur length 0.44 mm (♀), 4.89 (♀) × as long as broad ............................................................................ T. xianguensis sp. nov.</p><p>– Pedipalpal chela length 1.61–1.64 mm (♀), 6.42–6.44 (♀) × as long as broad; palpal femur length 1.14–1.18 mm (♀), 6.71–6.77 (♀) × as long as broad ................................. T. xinzhaiensis Li, 2022</p></div>	https://treatment.plazi.org/id/096987E3E409FFD76FC5FC29FA83FA41	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng	Sun, Jianzhou, Guo, Xiangbo, Zhang, Feng (2026): A review of the genus Tyrannochthonius (Pseudoscorpiones: Chthoniidae) from Guangxi, China, with the description of eleven new species. European Journal of Taxonomy 1058: 1-81, DOI: 10.5852/ejt.2026.1058.3278, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3278/14472
