taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
096987E3E443FF936D40FCD1FC39F9D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E443FF936D40FCD1FC39F9D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376216/files/figure.png	https://doi.org/10.5281/zenodo.20376216	Fig. 2. Tyrannochthonius baiseensis sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX2021072601), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX2021072602), dorsal view.	Fig. 2. Tyrannochthonius baiseensis sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX2021072601), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX2021072602), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E443FF936D40FCD1FC39F9D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376222/files/figure.png	https://doi.org/10.5281/zenodo.20376222	Fig. 3. Tyrannochthonius baiseensis sp. nov. A–G. Holotype, ♂ (Ps.-MHBU-GX2021072601). H. Paratype, ♀ (Ps.-MHBU-GX2021072602). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Carapace, dorsolateral view, indicate strongly weakly eyes (red arrow). F. Left pedipalp (minus chela), dorsal view. G. Male genital area, ventral view. H. Female genital area, ventral view.	Fig. 3. Tyrannochthonius baiseensis sp. nov. A–G. Holotype, ♂ (Ps.-MHBU-GX2021072601). H. Paratype, ♀ (Ps.-MHBU-GX2021072602). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Carapace, dorsolateral view, indicate strongly weakly eyes (red arrow). F. Left pedipalp (minus chela), dorsal view. G. Male genital area, ventral view. H. Female genital area, ventral view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E443FF936D40FCD1FC39F9D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376226/files/figure.png	https://doi.org/10.5281/zenodo.20376226	Fig. 4. Tyrannochthonius baiseensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2021072601). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	Fig. 4. Tyrannochthonius baiseensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2021072601). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E443FF936D40FCD1FC39F9D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376228/files/figure.png	https://doi.org/10.5281/zenodo.20376228	Fig. 5. Tyrannochthonius baiseensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2021072601) A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	Fig. 5. Tyrannochthonius baiseensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2021072601) A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E443FF936D40FCD1FC39F9D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376230/files/figure.png	https://doi.org/10.5281/zenodo.20376230	Fig. 6. Dayan Cave, type locality of Tyrannochthonius baiseensis sp. nov. A–B. T. baiseensis in its natural environment. C. Entrance. D. Inside the cave entrance.	Fig. 6. Dayan Cave, type locality of Tyrannochthonius baiseensis sp. nov. A–B. T. baiseensis in its natural environment. C. Entrance. D. Inside the cave entrance.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E449FF906EB3F919FD9FFDFD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376236/files/figure.png	https://doi.org/10.5281/zenodo.20376236	Fig. 7. Tyrannochthonius fenghuangensis sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX2021072801), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX2021072802), dorsal view.	Fig. 7. Tyrannochthonius fenghuangensis sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX2021072801), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX2021072802), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376238/files/figure.png	https://doi.org/10.5281/zenodo.20376238	Fig. 8. Tyrannochthonius fenghuangensis sp. nov. A–G. Holotype, ♂. (Ps.-MHBU-GX2021072801). H. Paratype, ♀ (Ps.-MHBU-GX2021072802). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Carapace, dorsolateral view, indicate strongly weakly eyes (red arrow). F. Left pedipalp (minus chela), dorsal view. G. Male genital area, ventral view. H. Female genital area, ventral view.	Fig. 8. Tyrannochthonius fenghuangensis sp. nov. A–G. Holotype, ♂. (Ps.-MHBU-GX2021072801). H. Paratype, ♀ (Ps.-MHBU-GX2021072802). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Carapace, dorsolateral view, indicate strongly weakly eyes (red arrow). F. Left pedipalp (minus chela), dorsal view. G. Male genital area, ventral view. H. Female genital area, ventral view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376240/files/figure.png	https://doi.org/10.5281/zenodo.20376240	Fig. 9. Tyrannochthonius fenghuangensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2021072801). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	Fig. 9. Tyrannochthonius fenghuangensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2021072801). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376244/files/figure.png	https://doi.org/10.5281/zenodo.20376244	Fig. 10. Tyrannochthonius fenghuangensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2021072801). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	Fig. 10. Tyrannochthonius fenghuangensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2021072801). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E44AFF8A6D6DFD35FBFCFEC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376246/files/figure.png	https://doi.org/10.5281/zenodo.20376246	Fig. 11. Unnamed Cave 3, type locality of Tyrannochthonius fenghuangensis sp. nov. A. T. fenghuangensis in its natural environment. B. Entrance.	Fig. 11. Unnamed Cave 3, type locality of Tyrannochthonius fenghuangensis sp. nov. A. T. fenghuangensis in its natural environment. B. Entrance.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E451FF8C6D40FE97FC31FA81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E451FF8C6D40FE97FC31FA81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376250/files/figure.png	https://doi.org/10.5281/zenodo.20376250	Fig. 12. Tyrannochthonius laqiuensis sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX18051101), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX18051102), dorsal view.	Fig. 12. Tyrannochthonius laqiuensis sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX18051101), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX18051102), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E451FF8C6D40FE97FC31FA81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376252/files/figure.png	https://doi.org/10.5281/zenodo.20376252	Fig. 13. Tyrannochthonius laqiuensis sp. nov. A–F. Holotype, ♂ (Ps.-MHBU-GX18051101). G. Paratype, ♀ (Ps.-MHBU-GX18051102). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Left pedipalp (minus chela), dorsal view. F. Male genital area, ventral view. G. Female genital area, ventral view.	Fig. 13. Tyrannochthonius laqiuensis sp. nov. A–F. Holotype, ♂ (Ps.-MHBU-GX18051101). G. Paratype, ♀ (Ps.-MHBU-GX18051102). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Left pedipalp (minus chela), dorsal view. F. Male genital area, ventral view. G. Female genital area, ventral view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E451FF8C6D40FE97FC31FA81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376256/files/figure.png	https://doi.org/10.5281/zenodo.20376256	Fig. 14. Tyrannochthonius laqiuensis sp. nov., holotype, ♂ (Ps.-MHBU-GX18051101). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	Fig. 14. Tyrannochthonius laqiuensis sp. nov., holotype, ♂ (Ps.-MHBU-GX18051101). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E451FF8C6D40FE97FC31FA81.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376260/files/figure.png	https://doi.org/10.5281/zenodo.20376260	Fig. 15. Tyrannochthonius laqiuensis sp. nov., holotype, ♂ (Ps.-MHBU-GX18051101). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	Fig. 15. Tyrannochthonius laqiuensis sp. nov., holotype, ♂ (Ps.-MHBU-GX18051101). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E456FF866D74FAD2FBF3FABF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E456FF866D74FAD2FBF3FABF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376264/files/figure.png	https://doi.org/10.5281/zenodo.20376264	Fig. 16. Tyrannochthonius longulimembrum sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100601), dorsal view.	Fig. 16. Tyrannochthonius longulimembrum sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100601), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E456FF866D74FAD2FBF3FABF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376266/files/figure.png	https://doi.org/10.5281/zenodo.20376266	Fig. 17. Tyrannochthonius longulimembrum sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100601). A. Left chela, lateral view. B. Left chela, dorsal view. C. Female genital area, ventral view. D. Left chelicera, dorsal view. E. Carapace, dorsal view. F. Left pedipalp (minus chela), dorsal view.	Fig. 17. Tyrannochthonius longulimembrum sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100601). A. Left chela, lateral view. B. Left chela, dorsal view. C. Female genital area, ventral view. D. Left chelicera, dorsal view. E. Carapace, dorsal view. F. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E456FF866D74FAD2FBF3FABF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376270/files/figure.png	https://doi.org/10.5281/zenodo.20376270	Fig. 18. Tyrannochthonius longulimembrum sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100601). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	Fig. 18. Tyrannochthonius longulimembrum sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100601). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E456FF866D74FAD2FBF3FABF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376274/files/figure.png	https://doi.org/10.5281/zenodo.20376274	Fig. 19. Tyrannochthonius longulimembrum sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100601). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	Fig. 19. Tyrannochthonius longulimembrum sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100601). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E456FF866D74FAD2FBF3FABF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376277/files/figure.png	https://doi.org/10.5281/zenodo.20376277	Fig. 20. Guangyang Cave, type locality of Tyrannochthonius longulimembrum sp. nov. A. Entrance. B. Inside the cave entrance.	Fig. 20. Guangyang Cave, type locality of Tyrannochthonius longulimembrum sp. nov. A. Entrance. B. Inside the cave entrance.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E45CFFBB6D51FAF4FC37F887.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E45CFFBB6D51FAF4FC37F887.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376279/files/figure.png	https://doi.org/10.5281/zenodo.20376279	Fig. 21. Tyrannochthonius miridentatus sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501), dorsal view.	Fig. 21. Tyrannochthonius miridentatus sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E45CFFBB6D51FAF4FC37F887.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376281/files/figure.png	https://doi.org/10.5281/zenodo.20376281	Fig. 22. Tyrannochthonius miridentatus sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501). A. Left chela, lateral view. B. Left chela, dorsal view. C. Female genital area, ventral view. D. Left chelicera, dorsal view. E. Carapace, dorsal view. F. Left pedipalp (minus chela), dorsal view.	Fig. 22. Tyrannochthonius miridentatus sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501). A. Left chela, lateral view. B. Left chela, dorsal view. C. Female genital area, ventral view. D. Left chelicera, dorsal view. E. Carapace, dorsal view. F. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E45CFFBB6D51FAF4FC37F887.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376287/files/figure.png	https://doi.org/10.5281/zenodo.20376287	Fig. 23. Tyrannochthonius miridentatus sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	Fig. 23. Tyrannochthonius miridentatus sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E45CFFBB6D51FAF4FC37F887.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376289/files/figure.png	https://doi.org/10.5281/zenodo.20376289	Fig. 24. Tyrannochthonius miridentatus sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	Fig. 24. Tyrannochthonius miridentatus sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E462FFB86ED6FE97FD90FBCD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E462FFB26D58FB05FC1BFD26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E462FFB26D58FB05FC1BFD26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376291/files/figure.png	https://doi.org/10.5281/zenodo.20376291	Fig. 25. Tyrannochthonius shenlongensis sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX2021073101), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX2021073102), dorsal view.	Fig. 25. Tyrannochthonius shenlongensis sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX2021073101), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX2021073102), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E462FFB26D58FB05FC1BFD26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376293/files/figure.png	https://doi.org/10.5281/zenodo.20376293	Fig. 26. Tyrannochthonius shenlongensis sp. nov. A–F. Holotype, ♂ (Ps.-MHBU-GX2021073101). G. Paratype, ♀ (Ps.-MHBU-GX2021073102). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Left pedipalp (minus chela), dorsal view. F. Male genital area, ventral view. G. Female genital area, ventral view.	Fig. 26. Tyrannochthonius shenlongensis sp. nov. A–F. Holotype, ♂ (Ps.-MHBU-GX2021073101). G. Paratype, ♀ (Ps.-MHBU-GX2021073102). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Left pedipalp (minus chela), dorsal view. F. Male genital area, ventral view. G. Female genital area, ventral view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E462FFB26D58FB05FC1BFD26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376295/files/figure.png	https://doi.org/10.5281/zenodo.20376295	Fig. 27. Tyrannochthonius shenlongensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2021073101). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	Fig. 27. Tyrannochthonius shenlongensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2021073101). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E462FFB26D58FB05FC1BFD26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376299/files/figure.png	https://doi.org/10.5281/zenodo.20376299	Fig. 28. Tyrannochthonius shenlongensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2021073101). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	Fig. 28. Tyrannochthonius shenlongensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2021073101). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E462FFB26D58FB05FC1BFD26.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376303/files/figure.png	https://doi.org/10.5281/zenodo.20376303	Fig. 29. Shenlong Cave, type locality of Tyrannochthonius shenlongensis sp. nov. A–B, D–E. T. shenlongensis in its natural environment. C. Entrance.	Fig. 29. Shenlong Cave, type locality of Tyrannochthonius shenlongensis sp. nov. A–B, D–E. T. shenlongensis in its natural environment. C. Entrance.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E469FFB36E8AFE97FDB4FC20.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E469FFB56D55FC70FC23FDB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E469FFB56D55FC70FC23FDB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376305/files/figure.png	https://doi.org/10.5281/zenodo.20376305	Fig. 30. Tyrannochthonius xianguensis sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501), dorsal view.	Fig. 30. Tyrannochthonius xianguensis sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E469FFB56D55FC70FC23FDB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376307/files/figure.png	https://doi.org/10.5281/zenodo.20376307	Fig. 31. Tyrannochthonius xianguensis sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501). A. Left chela, lateral view. B. Left chela, dorsal view. C. Female genital area, ventral view. D. Left chelicera, dorsal view. E. Carapace, dorsal view. F. Left pedipalp (minus chela), dorsal view.	Fig. 31. Tyrannochthonius xianguensis sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501). A. Left chela, lateral view. B. Left chela, dorsal view. C. Female genital area, ventral view. D. Left chelicera, dorsal view. E. Carapace, dorsal view. F. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E469FFB56D55FC70FC23FDB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376309/files/figure.png	https://doi.org/10.5281/zenodo.20376309	Fig. 32. Tyrannochthonius xianguensis sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	Fig. 32. Tyrannochthonius xianguensis sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E469FFB56D55FC70FC23FDB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376311/files/figure.png	https://doi.org/10.5281/zenodo.20376311	Fig. 33. Tyrannochthonius xianguensis sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	Fig. 33. Tyrannochthonius xianguensis sp. nov., holotype, ♀ (Ps.-MHBU-GX2019100501). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E469FFB56D55FC70FC23FDB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376315/files/figure.png	https://doi.org/10.5281/zenodo.20376315	Fig. 34. Xiangu Cave, type locality of Tyrannochthonius xianguensis sp. nov. A–B. Entrance. C. Area where the specimen of T. xianguensis was collected. D. T. xianguensis, ♀ (Ps.-MHBU-GX2019100501) in its natural environment.	Fig. 34. Xiangu Cave, type locality of Tyrannochthonius xianguensis sp. nov. A–B. Entrance. C. Area where the specimen of T. xianguensis was collected. D. T. xianguensis, ♀ (Ps.-MHBU-GX2019100501) in its natural environment.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376319/files/figure.png	https://doi.org/10.5281/zenodo.20376319	Fig. 35. Tyrannochthonius xiannvyanensis sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX2019100401), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX2019100403), dorsal view.	Fig. 35. Tyrannochthonius xiannvyanensis sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX2019100401), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX2019100403), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376323/files/figure.png	https://doi.org/10.5281/zenodo.20376323	Fig. 36. Tyrannochthonius xiannvyanensis sp. nov. A–F. Holotype, ♂ (Ps.-MHBU-GX2019100401). G. Paratype, ♀ (Ps.-MHBU-GX2019100403). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Carapace, dorsolateral view, indicate eyespot (red arrow). F. Left pedipalp (minus chela), dorsal view. G. Male genital area, ventral view. H. Female genital area, ventral view.	Fig. 36. Tyrannochthonius xiannvyanensis sp. nov. A–F. Holotype, ♂ (Ps.-MHBU-GX2019100401). G. Paratype, ♀ (Ps.-MHBU-GX2019100403). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Carapace, dorsolateral view, indicate eyespot (red arrow). F. Left pedipalp (minus chela), dorsal view. G. Male genital area, ventral view. H. Female genital area, ventral view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376325/files/figure.png	https://doi.org/10.5281/zenodo.20376325	Fig. 37. Tyrannochthonius xiannvyanensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2019100401). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	Fig. 37. Tyrannochthonius xiannvyanensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2019100401). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376327/files/figure.png	https://doi.org/10.5281/zenodo.20376327	Fig. 38. Tyrannochthonius xiannvyanensis sp. nov., holotype ♂ (Ps.-MHBU-GX2019100401). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	Fig. 38. Tyrannochthonius xiannvyanensis sp. nov., holotype ♂ (Ps.-MHBU-GX2019100401). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E46FFFAF6D60FDC2FBF5FA51.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376331/files/figure.png	https://doi.org/10.5281/zenodo.20376331	Fig. 39. Xiannvyan Cave, type locality of Tyrannochthonius xiannvyanensis sp. nov. A–B. T. xiannvyanensis in its natural environment.	Fig. 39. Xiannvyan Cave, type locality of Tyrannochthonius xiannvyanensis sp. nov. A–B. T. xiannvyanensis in its natural environment.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376333/files/figure.png	https://doi.org/10.5281/zenodo.20376333	Fig. 40. Tyrannochthonius xiaoi sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX2023113002), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX2023113004), dorsal view.	Fig. 40. Tyrannochthonius xiaoi sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX2023113002), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX2023113004), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376335/files/figure.png	https://doi.org/10.5281/zenodo.20376335	Fig. 41. Tyrannochthonius xiaoi sp. nov. A–E, G–H. Holotype, ♂ (Ps.-MHBU-GX2023113002). F, I–J. Paratype, ♀ (Ps.-MHBU-GX2023113004). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Carapace, dorsolateral view, indicate eyespot (red arrow). F. Carapace (dorsolateral view), indicate eyespot (red arrow). G. Left pedipalp (minus chela), dorsal view. H. Male genital area, ventral view. I. Female genital area, ventral view.	Fig. 41. Tyrannochthonius xiaoi sp. nov. A–E, G–H. Holotype, ♂ (Ps.-MHBU-GX2023113002). F, I–J. Paratype, ♀ (Ps.-MHBU-GX2023113004). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Carapace, dorsolateral view, indicate eyespot (red arrow). F. Carapace (dorsolateral view), indicate eyespot (red arrow). G. Left pedipalp (minus chela), dorsal view. H. Male genital area, ventral view. I. Female genital area, ventral view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376339/files/figure.png	https://doi.org/10.5281/zenodo.20376339	Fig. 42. Tyrannochthonius xiaoi sp. nov., holotype, ♂ (Ps.-MHBU-GX2023113002). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	Fig. 42. Tyrannochthonius xiaoi sp. nov., holotype, ♂ (Ps.-MHBU-GX2023113002). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376341/files/figure.png	https://doi.org/10.5281/zenodo.20376341	Fig. 43. Tyrannochthonius xiaoi sp. nov., holotype, ♂ (Ps.-MHBU-GX2023113002). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	Fig. 43. Tyrannochthonius xiaoi sp. nov., holotype, ♂ (Ps.-MHBU-GX2023113002). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E475FFA66D2FFA62FBFCFEC0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376343/files/figure.png	https://doi.org/10.5281/zenodo.20376343	Fig. 44. Unnamed Cave 1, type locality of Tyrannochthonius xiaoi sp. nov. A. Inside the cave entrance. B. Areas where specimens of T. xiaoi were collected.	Fig. 44. Unnamed Cave 1, type locality of Tyrannochthonius xiaoi sp. nov. A. Inside the cave entrance. B. Areas where specimens of T. xiaoi were collected.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E47CFFA66EB0FE10FDDBFBA6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E47CFFD96D67FBEAFC07FE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376347/files/figure.png	https://doi.org/10.5281/zenodo.20376347	Fig. 45. Tyrannochthonius yilingyanensis sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX2019100801), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX2019100803), dorsal view.	Fig. 45. Tyrannochthonius yilingyanensis sp. nov. A. Holotype, ♂ (Ps.-MHBU-GX2019100801), dorsal view. B. Paratype, ♀ (Ps.-MHBU-GX2019100803), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E47CFFD96D67FBEAFC07FE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376349/files/figure.png	https://doi.org/10.5281/zenodo.20376349	Fig. 46. Tyrannochthonius yilingyanensis sp. nov. A–G. Holotype, ♂ (Ps.-MHBU-GX2019100801). H. Paratype, ♀ (Ps.-MHBU-GX2019100803). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Carapace, dorsolateral view, indicate strongly weakly eyes (red arrow). F. Left pedipalp (minus chela), dorsal view. G. Male genital area, ventral view. H. Female genital area, ventral view.	Fig. 46. Tyrannochthonius yilingyanensis sp. nov. A–G. Holotype, ♂ (Ps.-MHBU-GX2019100801). H. Paratype, ♀ (Ps.-MHBU-GX2019100803). A. Left chela, lateral view. B. Left chela, dorsal view. C. Left chelicera, dorsal view. D. Carapace, dorsal view. E. Carapace, dorsolateral view, indicate strongly weakly eyes (red arrow). F. Left pedipalp (minus chela), dorsal view. G. Male genital area, ventral view. H. Female genital area, ventral view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E47CFFD96D67FBEAFC07FE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376351/files/figure.png	https://doi.org/10.5281/zenodo.20376351	Fig. 47. Tyrannochthonius yilingyanensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2019100801). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	Fig. 47. Tyrannochthonius yilingyanensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2019100801). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E47CFFD96D67FBEAFC07FE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376353/files/figure.png	https://doi.org/10.5281/zenodo.20376353	Fig. 48. Tyrannochthonius yilingyanensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2019100801). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	Fig. 48. Tyrannochthonius yilingyanensis sp. nov., holotype, ♂ (Ps.-MHBU-GX2019100801). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E47CFFD96D67FBEAFC07FE48.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376355/files/figure.png	https://doi.org/10.5281/zenodo.20376355	Fig. 49. Yilingyan Cave, type locality of Tyrannochthonius yilingyanensis sp. nov. A. Inside the cave entrance. B. Entrance. C. Area where specimens of T. yilingyanensis were collected. D. T. yilingyanensis in its natural environment. E–F. Cave environment.	Fig. 49. Yilingyan Cave, type locality of Tyrannochthonius yilingyanensis sp. nov. A. Inside the cave entrance. B. Entrance. C. Area where specimens of T. yilingyanensis were collected. D. T. yilingyanensis in its natural environment. E–F. Cave environment.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376212/files/figure.png	https://doi.org/10.5281/zenodo.20376212	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	Fig. 1. Study area and species composition of the genus Tyrannochthonius Chamberlin, 1929 in karst caves of Guangxi Zhuang Autonomous Region. Green represents known species, red represents new species. A. Unnamed Cave 1 (T. xiaoi sp. nov.). B. Rudongyan Cave (T. rudongyanensis Sun, Guo & Zhang, 2025). C. Maomaotou Cave (T. chixingi Gao, Wynne & Zhang, 2018). D. Ganxiao Cave (T. tiani Sun, Guo & Zhang, 2025). E. Yanwu Cave (T. yanwuensis Sun, Guo & Zhang, 2025). F. Shenlong Cave (T. shenlongensis sp. nov.). G. Xian Cave (T. miridentatus sp. nov.). H. Dayan Cave (T. baiseensis sp. nov.). I. Laqiu Cave (T. laqiuensis sp. nov.). J. Xiannvyan Cave (T. xiannvyanensis sp. nov.). K. Xiangu Cave (T. xianguensis sp. nov.). L. Unnamed Cave 3 (T. fenghuangensis sp. nov.). M. Unnamed Cave 2 (T. yui sp. nov.). N. Guangyang Cave (T. longulimembrum sp. nov.). O. Yilingyan Cave (T. yilingyanensis sp. nov.).	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376357/files/figure.png	https://doi.org/10.5281/zenodo.20376357	Fig. 50. Tyrannochthonius yui sp. nov., holotype, ♂ (Ps.-MHBU-GX2023080901), dorsal view.	Fig. 50. Tyrannochthonius yui sp. nov., holotype, ♂ (Ps.-MHBU-GX2023080901), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376361/files/figure.png	https://doi.org/10.5281/zenodo.20376361	Fig. 51. Tyrannochthonius yui sp. nov., holotype, ♂ (Ps.-MHBU-GX2023080901). A. Left chela, lateral view. B. Left chela, dorsal view. C. Male genital area, ventral view. D. Left chelicera, dorsal view. E. Carapace, dorsal view. F. Left pedipalp (minus chela), dorsal view.	Fig. 51. Tyrannochthonius yui sp. nov., holotype, ♂ (Ps.-MHBU-GX2023080901). A. Left chela, lateral view. B. Left chela, dorsal view. C. Male genital area, ventral view. D. Left chelicera, dorsal view. E. Carapace, dorsal view. F. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376363/files/figure.png	https://doi.org/10.5281/zenodo.20376363	Fig. 52. Tyrannochthonius yui sp. nov., holotype, ♂ (Ps.-MHBU-GX2023080901). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	Fig. 52. Tyrannochthonius yui sp. nov., holotype, ♂ (Ps.-MHBU-GX2023080901). A. Carapace, dorsal view. B. Left chelicera, dorsal view, with details of teeth. C. Rallum. D. Coxal spines on coxae II, ventral view. E. Left pedipalp (minus chela), dorsal view.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376365/files/figure.png	https://doi.org/10.5281/zenodo.20376365	Fig. 53. Tyrannochthonius yui sp. nov., holotype, ♂ (Ps.-MHBU-GX2023080901). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	Fig. 53. Tyrannochthonius yui sp. nov., holotype, ♂ (Ps.-MHBU-GX2023080901). A. Left chela, lateral view, with details of teeth and trichobothrial pattern. B. Left chela, dorsal view. C. Leg I, lateral view. D. Leg IV, lateral view. Abbreviations: see Material and methods.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
096987E3E403FFD36D1BFD88FBFCFC1D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/20376367/files/figure.png	https://doi.org/10.5281/zenodo.20376367	Fig. 54. Unnamed Cave 2, type locality of Tyrannochthonius yui sp. nov. A. The environment outside the cave (red arrow pointing to the cave entrance). B. Entrance. C. Inside the cave entrance. D. Cave tunnel junction with red arrow indicating the path to the specimen collection site. E–F. Area where the holotype (Ps.-MHBU-GX2023080901) of T. yui was collected.	Fig. 54. Unnamed Cave 2, type locality of Tyrannochthonius yui sp. nov. A. The environment outside the cave (red arrow pointing to the cave entrance). B. Entrance. C. Inside the cave entrance. D. Cave tunnel junction with red arrow indicating the path to the specimen collection site. E–F. Area where the holotype (Ps.-MHBU-GX2023080901) of T. yui was collected.	2026-05-21	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng		Zenodo	biologists	Sun, Jianzhou;Guo, Xiangbo;Zhang, Feng			
