taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
7B87110B6FD857F5943EACD002AA22F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724883	https://doi.org/10.3897/dez.73.188790.figure2	Figure 2. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), female, Sabah (Malaysia). A. Habitus; B. Head, frontal view; C. Body, lateral view; D. Labels. (Photos by Benjamin W. Price).	Figure 2. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), female, Sabah (Malaysia). A. Habitus; B. Head, frontal view; C. Body, lateral view; D. Labels. (Photos by Benjamin W. Price).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
7B87110B6FD857F5943EACD002AA22F4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724896	https://doi.org/10.3897/dez.73.188790.figure15	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
AEBAAD821E1D58209AF34FEE9A63C142.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724882	https://doi.org/10.3897/dez.73.188790.figure1	Figure 1. Habitus of Nuglerus spp., females. A. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, Coimbatore (India); B. N. insignis Banks, 1939, stat. rev., Palawan (Philippines); C. Neotype of Bofia maculata Navás, 1915 (herein designate) (current name: Nuglerus maculatus (Navás, 1915 )), Hoa Binh (Vietnam); D. N. maculatus (Navás, 1915), Yunnan (China); E. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); F. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); G. N. shihongliangi Zheng, Badano & Liu, sp. nov., Bali Is. (Indonesia); H. Nuglerus sp., East Kalimantan (Indonesia).	Figure 1. Habitus of Nuglerus spp., females. A. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, Coimbatore (India); B. N. insignis Banks, 1939, stat. rev., Palawan (Philippines); C. Neotype of Bofia maculata Navás, 1915 (herein designate) (current name: Nuglerus maculatus (Navás, 1915 )), Hoa Binh (Vietnam); D. N. maculatus (Navás, 1915), Yunnan (China); E. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); F. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); G. N. shihongliangi Zheng, Badano & Liu, sp. nov., Bali Is. (Indonesia); H. Nuglerus sp., East Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
AEBAAD821E1D58209AF34FEE9A63C142.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724884	https://doi.org/10.3897/dez.73.188790.figure3	Figure 3. Nuglerus guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India). A. Habitus; B. Head, frontal view; C. Head and thorax, dorsal view; female terminalia (D, E): D. Lateral view; E. Ventral view. Abbreviations of terms of wing venation: A: anal veins; abl: anterior Banksian line C: costa; CuA: cubitus anterior; CuP: cubitus posterior; MA: media anterior; MP: media posterior; RA: radius anterior; RP: radius posterior; Sc: subcostal. Abbreviations of terms of terminalia: ect: ectoprocts, gx: gonocoxites, pp: pregenital plate.	Figure 3. Nuglerus guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India). A. Habitus; B. Head, frontal view; C. Head and thorax, dorsal view; female terminalia (D, E): D. Lateral view; E. Ventral view. Abbreviations of terms of wing venation: A: anal veins; abl: anterior Banksian line C: costa; CuA: cubitus anterior; CuP: cubitus posterior; MA: media anterior; MP: media posterior; RA: radius anterior; RP: radius posterior; Sc: subcostal. Abbreviations of terms of terminalia: ect: ectoprocts, gx: gonocoxites, pp: pregenital plate.	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
AEBAAD821E1D58209AF34FEE9A63C142.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724896	https://doi.org/10.3897/dez.73.188790.figure15	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
7B14FB59817451CBAFB71F01CC3DAC10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724882	https://doi.org/10.3897/dez.73.188790.figure1	Figure 1. Habitus of Nuglerus spp., females. A. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, Coimbatore (India); B. N. insignis Banks, 1939, stat. rev., Palawan (Philippines); C. Neotype of Bofia maculata Navás, 1915 (herein designate) (current name: Nuglerus maculatus (Navás, 1915 )), Hoa Binh (Vietnam); D. N. maculatus (Navás, 1915), Yunnan (China); E. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); F. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); G. N. shihongliangi Zheng, Badano & Liu, sp. nov., Bali Is. (Indonesia); H. Nuglerus sp., East Kalimantan (Indonesia).	Figure 1. Habitus of Nuglerus spp., females. A. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, Coimbatore (India); B. N. insignis Banks, 1939, stat. rev., Palawan (Philippines); C. Neotype of Bofia maculata Navás, 1915 (herein designate) (current name: Nuglerus maculatus (Navás, 1915 )), Hoa Binh (Vietnam); D. N. maculatus (Navás, 1915), Yunnan (China); E. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); F. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); G. N. shihongliangi Zheng, Badano & Liu, sp. nov., Bali Is. (Indonesia); H. Nuglerus sp., East Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
7B14FB59817451CBAFB71F01CC3DAC10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724885	https://doi.org/10.3897/dez.73.188790.figure4	Figure 4. Holotype of Nuglerus insignis Banks, 1939, stat. rev., female, Los Baños (the Philippines). A. Forewing; B. Hindwing; C. Labels; D. Head and thorax, lateral view; E. Head, frontal view; F. Head and thorax, dorsal view. (Photos by Crystal A. Maier).	Figure 4. Holotype of Nuglerus insignis Banks, 1939, stat. rev., female, Los Baños (the Philippines). A. Forewing; B. Hindwing; C. Labels; D. Head and thorax, lateral view; E. Head, frontal view; F. Head and thorax, dorsal view. (Photos by Crystal A. Maier).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
7B14FB59817451CBAFB71F01CC3DAC10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724886	https://doi.org/10.3897/dez.73.188790.figure5	Figure 5. Nuglerus insignis Banks, 1939, stat. rev., female, Palawan (the Philippines). A. Habitus; B. Head, frontal view; C. Head and thorax, dorsal view; female terminalia (D, E): D. Lateral view; E. Ventral view.	Figure 5. Nuglerus insignis Banks, 1939, stat. rev., female, Palawan (the Philippines). A. Habitus; B. Head, frontal view; C. Head and thorax, dorsal view; female terminalia (D, E): D. Lateral view; E. Ventral view.	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
7B14FB59817451CBAFB71F01CC3DAC10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724896	https://doi.org/10.3897/dez.73.188790.figure15	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
7B14FB59817451CBAFB71F01CC3DAC10.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724897	https://doi.org/10.3897/dez.73.188790.figure16	Figure 16. Molecular phylogeny of Nuglerus and selected species of Myrmeleontidae based on COI barcoding sequence data. Tree topologies are identical among the Maximum Likelihood inference (ML) / Neighbour-joining analyses (NJ).	Figure 16. Molecular phylogeny of Nuglerus and selected species of Myrmeleontidae based on COI barcoding sequence data. Tree topologies are identical among the Maximum Likelihood inference (ML) / Neighbour-joining analyses (NJ).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
8CC9C99BA89758468E7621344659A288.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724882	https://doi.org/10.3897/dez.73.188790.figure1	Figure 1. Habitus of Nuglerus spp., females. A. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, Coimbatore (India); B. N. insignis Banks, 1939, stat. rev., Palawan (Philippines); C. Neotype of Bofia maculata Navás, 1915 (herein designate) (current name: Nuglerus maculatus (Navás, 1915 )), Hoa Binh (Vietnam); D. N. maculatus (Navás, 1915), Yunnan (China); E. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); F. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); G. N. shihongliangi Zheng, Badano & Liu, sp. nov., Bali Is. (Indonesia); H. Nuglerus sp., East Kalimantan (Indonesia).	Figure 1. Habitus of Nuglerus spp., females. A. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, Coimbatore (India); B. N. insignis Banks, 1939, stat. rev., Palawan (Philippines); C. Neotype of Bofia maculata Navás, 1915 (herein designate) (current name: Nuglerus maculatus (Navás, 1915 )), Hoa Binh (Vietnam); D. N. maculatus (Navás, 1915), Yunnan (China); E. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); F. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); G. N. shihongliangi Zheng, Badano & Liu, sp. nov., Bali Is. (Indonesia); H. Nuglerus sp., East Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
8CC9C99BA89758468E7621344659A288.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724887	https://doi.org/10.3897/dez.73.188790.figure6	Figure 6. Nuglerus maculatus (Navás, 1915), female. A. Habitus, Yunnan (China); B. Head, frontal view, neotype, Hoa Binh (Vietnam); C. Head and thorax, dorsal view, Yunnan (China); female terminalia (D, E), Yunnan (China): D. Lateral view; E. Ventral view.	Figure 6. Nuglerus maculatus (Navás, 1915), female. A. Habitus, Yunnan (China); B. Head, frontal view, neotype, Hoa Binh (Vietnam); C. Head and thorax, dorsal view, Yunnan (China); female terminalia (D, E), Yunnan (China): D. Lateral view; E. Ventral view.	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
8CC9C99BA89758468E7621344659A288.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724896	https://doi.org/10.3897/dez.73.188790.figure15	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
D1C929B8CE89560CAC0A26C5B79022EE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724882	https://doi.org/10.3897/dez.73.188790.figure1	Figure 1. Habitus of Nuglerus spp., females. A. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, Coimbatore (India); B. N. insignis Banks, 1939, stat. rev., Palawan (Philippines); C. Neotype of Bofia maculata Navás, 1915 (herein designate) (current name: Nuglerus maculatus (Navás, 1915 )), Hoa Binh (Vietnam); D. N. maculatus (Navás, 1915), Yunnan (China); E. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); F. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); G. N. shihongliangi Zheng, Badano & Liu, sp. nov., Bali Is. (Indonesia); H. Nuglerus sp., East Kalimantan (Indonesia).	Figure 1. Habitus of Nuglerus spp., females. A. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, Coimbatore (India); B. N. insignis Banks, 1939, stat. rev., Palawan (Philippines); C. Neotype of Bofia maculata Navás, 1915 (herein designate) (current name: Nuglerus maculatus (Navás, 1915 )), Hoa Binh (Vietnam); D. N. maculatus (Navás, 1915), Yunnan (China); E. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); F. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); G. N. shihongliangi Zheng, Badano & Liu, sp. nov., Bali Is. (Indonesia); H. Nuglerus sp., East Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
D1C929B8CE89560CAC0A26C5B79022EE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724896	https://doi.org/10.3897/dez.73.188790.figure15	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
D1C929B8CE89560CAC0A26C5B79022EE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724888	https://doi.org/10.3897/dez.73.188790.figure7	Figure 7. Nuglerus maizuqii Zheng, Badano & Liu, sp. nov., holotype, female, Pahang (Malaysia). A. Habitus; B. Head, frontal view; C. Head and thorax, dorsal view; female terminalia (D, E): D. Lateral view; E. Ventral view.	Figure 7. Nuglerus maizuqii Zheng, Badano & Liu, sp. nov., holotype, female, Pahang (Malaysia). A. Habitus; B. Head, frontal view; C. Head and thorax, dorsal view; female terminalia (D, E): D. Lateral view; E. Ventral view.	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
D1C929B8CE89560CAC0A26C5B79022EE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724889	https://doi.org/10.3897/dez.73.188790.figure8	Figure 8. Habitus of Nuglerus maizuqii Zheng, Badano & Liu, sp. nov., 3 rd instar larva, Pahang (Malaysia). A. Dorsal view; B. Ventral view; C. Lateral view.	Figure 8. Habitus of Nuglerus maizuqii Zheng, Badano & Liu, sp. nov., 3 rd instar larva, Pahang (Malaysia). A. Dorsal view; B. Ventral view; C. Lateral view.	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
D1C929B8CE89560CAC0A26C5B79022EE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724890	https://doi.org/10.3897/dez.73.188790.figure9	Figure 9. Nuglerus maizuqii Zheng, Badano & Liu, sp. nov., 3 rd instar larva, Pahang (Malaysia). A. Head, dorsal view; B. Thorax, dorsal view; C. Terminal of abdomen, ventral view. Abbreviations of terms of larval characters: 1 t: first tooth; 2 t: second tooth; 3 t: third tooth; Ims: interdental mandibular setae; Do: dolichaster; Tsp: thoracic setiferous processes; As 9: abdominal segment 9; Wdr: weakly developed rastrum; Wdds: weakly developed digging setae.	Figure 9. Nuglerus maizuqii Zheng, Badano & Liu, sp. nov., 3 rd instar larva, Pahang (Malaysia). A. Head, dorsal view; B. Thorax, dorsal view; C. Terminal of abdomen, ventral view. Abbreviations of terms of larval characters: 1 t: first tooth; 2 t: second tooth; 3 t: third tooth; Ims: interdental mandibular setae; Do: dolichaster; Tsp: thoracic setiferous processes; As 9: abdominal segment 9; Wdr: weakly developed rastrum; Wdds: weakly developed digging setae.	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
D1C929B8CE89560CAC0A26C5B79022EE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724892	https://doi.org/10.3897/dez.73.188790.figure11	Figure 11. Nuglerus maizuqii Zheng, Badano & Liu, sp. nov., living adult, Pahang (Malaysia). A. Habitus; B. Distal wings of living adult; C. THickened markings on forewing poststigmal area, lateral view; D. Thickened markings on hindwing poststigmal area, lateral view. (Photos by Yuchen Zheng).	Figure 11. Nuglerus maizuqii Zheng, Badano & Liu, sp. nov., living adult, Pahang (Malaysia). A. Habitus; B. Distal wings of living adult; C. THickened markings on forewing poststigmal area, lateral view; D. Thickened markings on hindwing poststigmal area, lateral view. (Photos by Yuchen Zheng).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
AA00FF173354504AA70387F33348654C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724892	https://doi.org/10.3897/dez.73.188790.figure11	Figure 11. Nuglerus maizuqii Zheng, Badano & Liu, sp. nov., living adult, Pahang (Malaysia). A. Habitus; B. Distal wings of living adult; C. THickened markings on forewing poststigmal area, lateral view; D. Thickened markings on hindwing poststigmal area, lateral view. (Photos by Yuchen Zheng).	Figure 11. Nuglerus maizuqii Zheng, Badano & Liu, sp. nov., living adult, Pahang (Malaysia). A. Habitus; B. Distal wings of living adult; C. THickened markings on forewing poststigmal area, lateral view; D. Thickened markings on hindwing poststigmal area, lateral view. (Photos by Yuchen Zheng).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
57C3E3AB25065CA286806052221207B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724893	https://doi.org/10.3897/dez.73.188790.figure12	Figure 12. Nuglerus pustulatus (McLachlan, 1867), female. A, B, F. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka; C, D, E, G – J. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (now junior synonym of N. pustulatus); A, C. Habitus; B, D. Labels; E, F. Head, frontal view; G. Head and thorax, dorsal view; female terminalia (H, I): H. Lateral view; I. Ventral view; J. Head and thorax, lateral view.	Figure 12. Nuglerus pustulatus (McLachlan, 1867), female. A, B, F. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka; C, D, E, G – J. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (now junior synonym of N. pustulatus); A, C. Habitus; B, D. Labels; E, F. Head, frontal view; G. Head and thorax, dorsal view; female terminalia (H, I): H. Lateral view; I. Ventral view; J. Head and thorax, lateral view.	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
57C3E3AB25065CA286806052221207B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724896	https://doi.org/10.3897/dez.73.188790.figure15	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
57C3E3AB25065CA286806052221207B9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724892	https://doi.org/10.3897/dez.73.188790.figure11	Figure 11. Nuglerus maizuqii Zheng, Badano & Liu, sp. nov., living adult, Pahang (Malaysia). A. Habitus; B. Distal wings of living adult; C. THickened markings on forewing poststigmal area, lateral view; D. Thickened markings on hindwing poststigmal area, lateral view. (Photos by Yuchen Zheng).	Figure 11. Nuglerus maizuqii Zheng, Badano & Liu, sp. nov., living adult, Pahang (Malaysia). A. Habitus; B. Distal wings of living adult; C. THickened markings on forewing poststigmal area, lateral view; D. Thickened markings on hindwing poststigmal area, lateral view. (Photos by Yuchen Zheng).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
C3BD6428B7AA5E8AB053AC9E475587CD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724882	https://doi.org/10.3897/dez.73.188790.figure1	Figure 1. Habitus of Nuglerus spp., females. A. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, Coimbatore (India); B. N. insignis Banks, 1939, stat. rev., Palawan (Philippines); C. Neotype of Bofia maculata Navás, 1915 (herein designate) (current name: Nuglerus maculatus (Navás, 1915 )), Hoa Binh (Vietnam); D. N. maculatus (Navás, 1915), Yunnan (China); E. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); F. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); G. N. shihongliangi Zheng, Badano & Liu, sp. nov., Bali Is. (Indonesia); H. Nuglerus sp., East Kalimantan (Indonesia).	Figure 1. Habitus of Nuglerus spp., females. A. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, Coimbatore (India); B. N. insignis Banks, 1939, stat. rev., Palawan (Philippines); C. Neotype of Bofia maculata Navás, 1915 (herein designate) (current name: Nuglerus maculatus (Navás, 1915 )), Hoa Binh (Vietnam); D. N. maculatus (Navás, 1915), Yunnan (China); E. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); F. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); G. N. shihongliangi Zheng, Badano & Liu, sp. nov., Bali Is. (Indonesia); H. Nuglerus sp., East Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
C3BD6428B7AA5E8AB053AC9E475587CD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724893	https://doi.org/10.3897/dez.73.188790.figure12	Figure 12. Nuglerus pustulatus (McLachlan, 1867), female. A, B, F. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka; C, D, E, G – J. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (now junior synonym of N. pustulatus); A, C. Habitus; B, D. Labels; E, F. Head, frontal view; G. Head and thorax, dorsal view; female terminalia (H, I): H. Lateral view; I. Ventral view; J. Head and thorax, lateral view.	Figure 12. Nuglerus pustulatus (McLachlan, 1867), female. A, B, F. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka; C, D, E, G – J. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (now junior synonym of N. pustulatus); A, C. Habitus; B, D. Labels; E, F. Head, frontal view; G. Head and thorax, dorsal view; female terminalia (H, I): H. Lateral view; I. Ventral view; J. Head and thorax, lateral view.	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
C3BD6428B7AA5E8AB053AC9E475587CD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724895	https://doi.org/10.3897/dez.73.188790.figure14	Figure 14. Nuglerus shihongliangi Zheng, Badano & Liu, sp. nov., holotype, male, Bali Is. (Indonesia). A. Habitus; B. Head, frontal view; C. Head and thorax, dorsal view; male terminalia (D, E): D. Lateral view; E. Ventral view; male genitalia (F – J): F. Ventral view; G. Anteroventral view; H. Caudal view; I. Lateral view; J. Dorsal view. Abbreviations of terms of terminalia: ect: ectoprocts, gst: gonostylus, gx: gonocoxites, gp: gonapophyses, S 9: sternum 9.	Figure 14. Nuglerus shihongliangi Zheng, Badano & Liu, sp. nov., holotype, male, Bali Is. (Indonesia). A. Habitus; B. Head, frontal view; C. Head and thorax, dorsal view; male terminalia (D, E): D. Lateral view; E. Ventral view; male genitalia (F – J): F. Ventral view; G. Anteroventral view; H. Caudal view; I. Lateral view; J. Dorsal view. Abbreviations of terms of terminalia: ect: ectoprocts, gst: gonostylus, gx: gonocoxites, gp: gonapophyses, S 9: sternum 9.	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
C3BD6428B7AA5E8AB053AC9E475587CD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724894	https://doi.org/10.3897/dez.73.188790.figure13	Figure 13. Nuglerus ruzzieri Badano, Zheng & Liu, sp. nov., holotype, female, Sulawesi (Indonesia). A. Habitus; B. Head, frontal view; C. Head and thorax, dorsal view; female terminalia (D, E): D. Lateral view; E. Ventral view. Abbreviations of terms of terminalia: ect: ectoprocts, gst: gonostylus, gx: gonocoxites, pp: pregenital plate.	Figure 13. Nuglerus ruzzieri Badano, Zheng & Liu, sp. nov., holotype, female, Sulawesi (Indonesia). A. Habitus; B. Head, frontal view; C. Head and thorax, dorsal view; female terminalia (D, E): D. Lateral view; E. Ventral view. Abbreviations of terms of terminalia: ect: ectoprocts, gst: gonostylus, gx: gonocoxites, pp: pregenital plate.	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
C3BD6428B7AA5E8AB053AC9E475587CD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724896	https://doi.org/10.3897/dez.73.188790.figure15	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
C3BD6428B7AA5E8AB053AC9E475587CD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724897	https://doi.org/10.3897/dez.73.188790.figure16	Figure 16. Molecular phylogeny of Nuglerus and selected species of Myrmeleontidae based on COI barcoding sequence data. Tree topologies are identical among the Maximum Likelihood inference (ML) / Neighbour-joining analyses (NJ).	Figure 16. Molecular phylogeny of Nuglerus and selected species of Myrmeleontidae based on COI barcoding sequence data. Tree topologies are identical among the Maximum Likelihood inference (ML) / Neighbour-joining analyses (NJ).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
5428A63CA37555C3BC41FB93C7FA856D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724882	https://doi.org/10.3897/dez.73.188790.figure1	Figure 1. Habitus of Nuglerus spp., females. A. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, Coimbatore (India); B. N. insignis Banks, 1939, stat. rev., Palawan (Philippines); C. Neotype of Bofia maculata Navás, 1915 (herein designate) (current name: Nuglerus maculatus (Navás, 1915 )), Hoa Binh (Vietnam); D. N. maculatus (Navás, 1915), Yunnan (China); E. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); F. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); G. N. shihongliangi Zheng, Badano & Liu, sp. nov., Bali Is. (Indonesia); H. Nuglerus sp., East Kalimantan (Indonesia).	Figure 1. Habitus of Nuglerus spp., females. A. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, Coimbatore (India); B. N. insignis Banks, 1939, stat. rev., Palawan (Philippines); C. Neotype of Bofia maculata Navás, 1915 (herein designate) (current name: Nuglerus maculatus (Navás, 1915 )), Hoa Binh (Vietnam); D. N. maculatus (Navás, 1915), Yunnan (China); E. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); F. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); G. N. shihongliangi Zheng, Badano & Liu, sp. nov., Bali Is. (Indonesia); H. Nuglerus sp., East Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
5428A63CA37555C3BC41FB93C7FA856D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724895	https://doi.org/10.3897/dez.73.188790.figure14	Figure 14. Nuglerus shihongliangi Zheng, Badano & Liu, sp. nov., holotype, male, Bali Is. (Indonesia). A. Habitus; B. Head, frontal view; C. Head and thorax, dorsal view; male terminalia (D, E): D. Lateral view; E. Ventral view; male genitalia (F – J): F. Ventral view; G. Anteroventral view; H. Caudal view; I. Lateral view; J. Dorsal view. Abbreviations of terms of terminalia: ect: ectoprocts, gst: gonostylus, gx: gonocoxites, gp: gonapophyses, S 9: sternum 9.	Figure 14. Nuglerus shihongliangi Zheng, Badano & Liu, sp. nov., holotype, male, Bali Is. (Indonesia). A. Habitus; B. Head, frontal view; C. Head and thorax, dorsal view; male terminalia (D, E): D. Lateral view; E. Ventral view; male genitalia (F – J): F. Ventral view; G. Anteroventral view; H. Caudal view; I. Lateral view; J. Dorsal view. Abbreviations of terms of terminalia: ect: ectoprocts, gst: gonostylus, gx: gonocoxites, gp: gonapophyses, S 9: sternum 9.	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
5428A63CA37555C3BC41FB93C7FA856D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1724896	https://doi.org/10.3897/dez.73.188790.figure15	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	Figure 15. Distal wings of Nuglerus spp., females. A – E, K – O. Forewing; F – J, P – T. Hindwing; A, F. Holotype of Acratoleon dispar Banks, 1934 (current name: Nuglerus dispar comb. nov.), Sabah (Malaysia); B, G. N. guimaraesae Zheng, Badano & Liu, sp. nov., holotype, female, Coimbatore (India); C, H. N. insignis Banks, 1939, stat. rev., Palawan (the Philippines); D, I. N. maculatus (Navás, 1915), Yunnan (China); E, J. N. maizuqii Zheng, Badano & Liu, sp. nov., Pahang (Malaysia); K, P. Holotype of Glenurus pustulatus McLachlan, 1867, Sri Lanka (current name: N. pustulatus); L, Q. Holotype of Nuglerus scalaris Navás, 1912, with wrong type locality (“ Bresil ”) (current status: a junior synonym of N. pustulatus); M, R. N. ruzzieri Badano, Zheng & Liu, sp. nov., holotype, Sulawesi (Indonesia); N, S. N. shihongliangi Zheng, Badano & Liu, sp. nov., holotype, Bali Is. (Indonesia); O, T. Nuglerus sp., E. Kalimantan (Indonesia).	2026-07-27	Zheng, Yuchen;Badano, Davide;Liu, Xingyue		Zenodo	biologists	Zheng, Yuchen;Badano, Davide;Liu, Xingyue			
