identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
2567A40CB2525D0AA26E0E0CFFA1B323.text	2567A40CB2525D0AA26E0E0CFFA1B323.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mysteriohelota Hsiao 2026	<div><p>Genus Mysteriohelota gen. nov.</p><p>Figs 1, 2, 3, 4</p><p>Type species.</p><p>Mysteriohelota metallicus sp. nov. from the Cretaceous Kachin amber (Cenomanian, ca. 99 Ma) of northern Myanmar, by monotypy.</p><p>Diagnosis.</p><p>Body small (≤ 4.0 mm) (usually ≥ 6.5 mm in extant genera). Head elongate, longer than wide (transverse in Burmahelota Liu, Ślipiński, Ren &amp; Pang, Mesohelotopsis Kirejtshuk, Metahelotella, Neohelota and Trihelota). Frontoclypeal suture present (absent in other helotid genera). Neck broad (narrow in Lobatihelota). Antennal club ca. 2.0 times as long as wide (usually as long as wide in Neohelota while ca. 1.5 times as long as wide in Afrohelotina Kirejtshuk, Palaeohelota Liu, Ślipiński, Ren &amp; Pang, Strophohelota Kirejtshuk, Helota and Trihelota). Antennal grooves well-developed, extending beyond eyes (short and indistinct in Lobatihelota and Trihelota). Pronotum with length distinctly shorter than wide (as long as wide in Burmahelota while longer than wide in Lobatihelota and Trihelota). Pronotal lateral sides distinctly widening posteriad (arcuate in Burmahelota and Mesohelotopsis while subparallel-sided in Lobatihelota and Trihelota). Pronotal posterior margin deeply bisinuate, forming well-developed median lobe (subtruncate, with four weakly developed sinuations in Strophohelota). Pronotal disc glabrous, covered with sparse and fine punctation (surface rugose, irregularly and coarsely punctate, with raised patches in Helota). Prosternum unmodified (with paired short carinae in Trihelota). Prosternal process with apex truncate (usually strongly expanded, with apex broadly emarginate in extant genera and Lobatihelota). Mesoventrite unmodified (with paired carinae continuous with the sides of prosternal process). Exposed portions of metepisterna separating metacoxae from elytra (metacoxae extending laterally to meet elytra in Trihelota). Elytra glabrous, lacking elevated costae (with elevated costae in Afrohelotina, Helota, Lobatihelota and Palaeohelota). Elytral striae punctate, punctation indistinct, sparse and shallow (distinctly developed, deep and coarse in Afrohelotina, Helota, Metahelotella, Mesohelotopsis, Neohelota and Strophohelota). Elytra without coloured spots (present in Burmahelota, Helota, Neohelota and Trihelota). Tibiae subparallel-sided (widening apicad in Lobatihelota). Tarsomeres not lobed, IV only slightly shorter than III (I – III bilobed, with densely setose pads, IV extremely shorter and smaller than III in Lobatihelota).</p><p>Etymology.</p><p>The generic epithet is a combination of the Greek word μῠστήρῐον [mystery, enigma], referring to its unique morphology compared with other helotid beetles (e. g. presence of frontoclypeal suture), and Helota, the type genus of the family. Gender feminine.</p></div>	https://treatment.plazi.org/id/2567A40CB2525D0AA26E0E0CFFA1B323	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hsiao, Yun	Hsiao, Yun (2026): Quantitative morphological analysis reveals a greater morphological diversity in Cretaceous Helotidae (Coleoptera, Nitiduloidea), with description of a new genus and species from mid-Cretaceous Kachin amber. Fossil Record 29 (1): 171-183, DOI: 10.3897/fr.29.185700
044B8D7AEA6F52008E2E9E0D7318BBD9.text	044B8D7AEA6F52008E2E9E0D7318BBD9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mysteriohelota metallicus Hsiao 2026	<div><p>Mysteriohelota metallicus sp. nov.</p><p>Figs 1, 2, 3, 4</p><p>Etymology.</p><p>The specific epithet is derived from its green metallic shining head and pronotum.</p><p>Holotype.</p><p>No. YH-COL 00005, deposited in the National Museum of Natural Science (NMNS), Taichung, Taiwan; mostly complete and well-preserved adult, with abdomen and left hind leg missing .</p><p>Type locality.</p><p>Hukawng Valley, Kachin State, northern Myanmar.</p><p>Type stratum.</p><p>Kachin amber (earliest Cenomanian, ca. 99 Ma; Shi et al. 2012).</p><p>Diagnosis.</p><p>Same as genus.</p><p>Description.</p><p>Body length: 3.89 mm (from clypeus to elytral apex); width: 1.33 mm (at the widest point of elytra). Body (Figs 1, 2) elongate, long oval, 2.9 times as long as wide, lateral sides moderately narrowing posteriad, surface sparsely and shallowly punctate. Body and legs uniformly dark brown to black, head and pronotum with green to blue metallic lustre.</p><p>Head (Figs 1, 3 A – D) elongate, 1.1 times as long as wide, widest across eyes, abruptly constricted immediately behind eyes forming broad neck and extremely short temples. Frontoclypeal suture present. Clypeus broadly rounded apically. Mandibles broad, strongly curved. Terminal maxillary palpomere fusiform and elongate, widest medially. Antennal insertions located anteriorly of eyes, dorsally largely concealed. Antenna 11 - segmented, with well-developed and compact 3 - segmented club 1.9 times as long as wide, extending to apical fourth of pronotum, sparsely setose; approximate ratio of antennomere lengths as follows: 15.7: 10.0: 10.0: 8.6: 7.1: 5.7: 5.7: 5.7: 7.9: 7.9: 12.9; scape enlarged and globular, 1.8 times as long as wide; pedicel stout, 1.8 times as long as wide, 0.7 times as wide as scape; antennomere III – IV clavate, subequal in width, III 2.8 times as long as wide, IV 2.4 times as long as wide, both 0.6 times as wide as pedicel; V slightly widening apicad, 1.3 times as long as wide, 0.8 times as wide as pedicel; VI – VII moderately widening apicad, both 1.1 times as long as wide, both 0.9 times as wide as pedicel; VIII distinctly widening apicad, 0.7 times as long as wide, 1.4 times as wide as pedicel; IX – XI of subequal width, forming a compact club with dense short setae, widest at X, 2.6 times as wide as pedicel, IX – X extremely widening apicad, IX 0.6 times as long as wide, X 0.5 times as long as wide, XI with apex broadly rounded, 0.9 times as long as wide. Eyes large and hemispherical, distinctly protruding dorsally and laterally. Surface distinctly and sparsely punctate. Ventral side of head with short antennal grooves extending beyond eyes.</p><p>Pronotum (Figs 1, 3 A, B) roundly trapezoidal, 0.8 times as long as wide, longest medially, moderately widening posteriad, 1.5 times as wide as head. Anterior angles broadly rounded. Lateral pronotal carina with distinct bead. Pronotal disc moderately convex and glabrous, surface sparsely and shallowly punctate. Posterior margin deeply bisinuate, median lobe well-developed and rounded. Posterior angles strongly acute and distinctly projecting, pointing laterally at approximately 45 ° from body axis. Mesoscutellar shield minute, subtriangular, widest in anterior fourth. Prosternum (Figs 2, 3 C – F) flattened, area in front of procoxae 2.3 times as long as maximum procoxal diameter. Notosternal sutures complete. Hypomeron somewhat impressed posteriorly for reception of profemur. Procoxal cavities oval, widely separated, externally closed, interprocoxal distance 1.7 times as long as maximum procoxal diameter; protrochantin concealed. Prosternal process wide, subparallel-sided, apically truncate, broadly overlapping mesoventrite. Mesoventrite (Figs 2, 3 E, F) separated by complete sutures from mesepisterna. Mesocoxae longitudinally oval, widely separated, intermesocoxal distance 1.4 times as long as maximum mesocoxal diameter. Mesometaventral junction externally as nearly straight line. Metaventrite (Figs 2, 3 E, F) medially weakly depressed, with complete discrimen and transverse katepisternal suture. Metanepisternum extremely long and narrow. Metacoxae oval, horizontally oriented, not extending to meet elytra, oval, widely separated, intermetacoxal distance 1.8 times as long as maximum metacoxal diameter; metacoxal plates very weakly developed.</p><p>Elytra (Figs 1, 4) elongate, long oval, 1.7 times as long as wide, widest at elytral humeri, moderately narrowing posteriad. Elytral humeri obtusely rounded, without pointed humeral projection. Elytral surface glabrous, lacking costae, with extremely sparse and shallow punctate striae. Elytra unicolour, without coloured spots. Elytral apices with an apical indentation with a stout seta. Epipleura (Fig. 2) extending to apical fifth of elytra, widest basally.</p><p>Legs (Figs 1, 2, 3, 4) elongate and slender. Trochanterofemoral joint distinctly oblique. Femora widened, ventrally grooved for reception of tibiae. Tibiae slender, subparallel-sided, protibiae slightly bent in holotype, terminating with a pair of small tibial spurs and a small brush of setae. Tarsal formula 5-5 - 5, not lobed, with elongate setae ventrally; tarsomere IV only slightly shorter than III; V longer than the preceding four segments combined; tarsal adhesive soles absent. Claws simple, not toothed. Empodium well-developed, bisetose.</p></div>	https://treatment.plazi.org/id/044B8D7AEA6F52008E2E9E0D7318BBD9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Hsiao, Yun	Hsiao, Yun (2026): Quantitative morphological analysis reveals a greater morphological diversity in Cretaceous Helotidae (Coleoptera, Nitiduloidea), with description of a new genus and species from mid-Cretaceous Kachin amber. Fossil Record 29 (1): 171-183, DOI: 10.3897/fr.29.185700
