identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E19CB8B3C12D5A6CA61B60EF5990BF18.text	E19CB8B3C12D5A6CA61B60EF5990BF18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptinaria parana Pilsbry 1926	<div><p>Leptinaria parana Pilsbry, 1926</p><p>Fig. 3 E</p><p>Leptinaria parana: Simone and Martins 2020: 19, fig. 5 [see this article for full synonymy]; Salvador et al. 2024: 151.</p><p>Material examined.</p><p>LES 0031622, LES 0031653 (Canaã dos Carajás, cave SB_0049; JE Gallão, JS Gallo, DF Torres &amp; VF Sperandei leg., 16. ii. 2022) • LES 0031655 (Parauapebas, cave N 1_0174; JE Gallão &amp; DB Ribeiro leg., 04. iv. 2023) .</p><p>GenBank acc. nr.</p><p>PZ 576040 (COI), PZ 574757 (16 S), PZ 577562 (ITS 2 + 28 S) (LES 0031622).</p><p>Identification.</p><p>This species can be identified by its small size (~ 5.5 mm height, ~ 2 mm width, ~ 6 whorls), elongated spire, and faint growth lines (Pilsbry 1926).</p><p>Distribution.</p><p>Known only from the type locality in Pará (Pilsbry 1906), interpreted as a suburb of Belém (see discussion in Simone and Martins 2020). Records of L. parana beyond the type locality are likely incorrect. Based on the image provided in their work, the occurrence of L. parana in Peru reported by Ituarte et al. (2008) likely corresponds to a different species. The record of L. parana from the state of Santa Catarina, southern Brazil, reported by Agudo-Padrón (2018) and follow-up works (Agudo-Padrón 2022, 2024), with authorship erroneously attributed to “ Pilbry, 1906 ”, is likewise dubious. The vouchers could not be traced; the illustration provided by Agudo-Padrón (2018: 58, fig. 2) is inconclusive (showing only a dorsal view of the shell) and likely corresponds to other congeners known from that area (see Simone 2006). The present record extends the species’ range by ca 550 km south of its previous record in Belém.</p><p>Remarks.</p><p>In the studied caves, L. parana was found in the twilight zone (in leaf litter) and the aphotic zone (under rocks, on very humid unconsolidated substrate). Furthermore, the species has also been recorded in anthropically disturbed areas in the nearby municipality of Parauapebas (Paulo Ricardo S. Coelho, pers. comm. 23 March 2026, species identity confirmed via photographs).</p></div>	https://treatment.plazi.org/id/E19CB8B3C12D5A6CA61B60EF5990BF18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo B.;Ferreira-Santos, Laura;Cavallari, Daniel C.;Gallão, Jonas E.;Bichuette, Maria E.	Salvador, Rodrigo B., Ferreira-Santos, Laura, Cavallari, Daniel C., Gallão, Jonas E., Bichuette, Maria E. (2026): Terrestrial gastropods from caves in Serra dos Carajás, eastern Brazilian Amazon, with the description of two new species (Eupulmonata, Stylommatophora). ZooKeys 1287: 213-237, DOI: 10.3897/zookeys.1287.194544
4AA0A7767D9655D3AA8D877084AC4266.text	4AA0A7767D9655D3AA8D877084AC4266.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leptinaria unilamellata (d'Orbigny 1838)	<div><p>Leptinaria unilamellata (d’Orbigny, 1838)</p><p>Fig. 3 F</p><p>Leptinaria lamellata: Morretes 1959: 132; Salgado and Coelho 2003: 155; Simone 2006: 186, fig. 678; Salvador et al. 2021: 5.</p><p>Leptinaria unilamelata [sic]: Oliveira et al. 1981: 327.</p><p>Leptinaria unilamellata: Simone 2006: 186, fig. 684; Breure et al. 2016: 35, tab. 3, fig. 14; Alexandre et al. 2017: 34, tabs 1–2, fig. 2 K; Breure et al. 2020: 27, figs 1–5 [see this article for full synonymy]; Lima et al. 2021: 274, fig. 3 T – V; Rangel et al. 2021: 4, tab. 1, fig. 2 b; Silva et al. 2021: 48, fig. 4 G, H; Salvador et al. 2024: 151.</p><p>Material examined.</p><p>LES 0031620, LES 0031621, LES 0031623 (Canaã dos Carajás, cave SB_0049; JE Gallão, JS Gallo, DF Torres &amp; VF Sperandei leg., 16. ii. 2022) • LES 0031629 (Parauapebas, cave N 3_0026; JE Gallão, JS Gallo, DF Torres &amp; VF Sperandei leg., 18. ii. 2022) • LES 0031643 (Parauapebas, cave N 3_0026; JE Gallão, TS Gallo, DF Torres &amp; JEM Santos leg., 16. viii. 2022) • LES 0031655 (Canaã dos Carajás, cave SB_0049; JE Gallão &amp; DB Ribeiro leg., 04. iv. 2023) • LES 0031657 (Canaã dos Carajás, cave SB_0094; JE Gallão &amp; DB Ribeiro leg., 06. iv. 2023) • LES 0031648, LES 0031650 (Canaã dos Carajás, cave SB_0049; JE Gallão &amp; DB Ribeiro leg., 12. iv. 2023) • LES 0031648 (Canaã dos Carajás, cave SB_0051; JE Gallão &amp; DB Ribeiro leg., 12. iv. 2023) .</p><p>GenBank acc. nr.</p><p>PZ 574924 (16 S), PZ 577563 (ITS 2 + 28 S) (LES 0031650).</p><p>Identification.</p><p>Leptinaria unilamellata is a very widespread species in the Neotropics, characterised by the single lamella that gives it its name, and known from both natural and anthropically disturbed areas. It is thought to represent a species complex, but genetic and morphometric studies would be required to clarify that. In any case, the species shows conchological variation, with most shells being smooth, while axially ribbed specimens are uncommon but well known. These two shell morphs are known to occur in the same place in southeastern Brazil (DCC, pers. obs.), and both were found in the present material.</p><p>Distribution.</p><p>Nicaragua to southern Brazil (Simone 2006; Delannoye et al. 2015; Silva et al. 2021).</p><p>Remarks.</p><p>In the studied cave, Leptinaria unilamellata was found in the aphotic zone. Individuals were found under rocks on humid unconsolidated substrate, among roots, or on bat guano (mixed guano of frugivorous and insectivorous bats). This species has also been recorded in anthropically disturbed areas in the nearby municipality of Parauapebas in Pará (Paulo Ricardo S. Coelho, pers. comm., species identity confirmed via photographs).</p></div>	https://treatment.plazi.org/id/4AA0A7767D9655D3AA8D877084AC4266	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo B.;Ferreira-Santos, Laura;Cavallari, Daniel C.;Gallão, Jonas E.;Bichuette, Maria E.	Salvador, Rodrigo B., Ferreira-Santos, Laura, Cavallari, Daniel C., Gallão, Jonas E., Bichuette, Maria E. (2026): Terrestrial gastropods from caves in Serra dos Carajás, eastern Brazilian Amazon, with the description of two new species (Eupulmonata, Stylommatophora). ZooKeys 1287: 213-237, DOI: 10.3897/zookeys.1287.194544
C201E9A6AF1D561E9426335EF2F585C6.text	C201E9A6AF1D561E9426335EF2F585C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rectobelus arsayae Salvador & Ferreira-Santos & Cavallari & Gallão & Bichuette 2026	<div><p>Rectobelus arsayae sp. nov.</p><p>Fig. 3 A – C</p><p>Type material.</p><p>Holotype LES 0031661 .</p><p>Type locality.</p><p>Brazil, Pará state, Canaã dos Carajás municipality, cave SB_0022 (GEM 1442) (JE Gallão &amp; DB Ribeiro leg., 06. iv. 2023).</p><p>Diagnosis.</p><p>Shell nearly cylindrical in shape, with ca 10 whorls and ca 10 mm in length. The apex is not as narrow as in congeners and the protoconch is only slightly inflated relative to the first teleoconch whorls. Presence of very faint columellar fold, extending internally into the shell throughout the length of the body whorl. Presence of two internal palatal lamellae in sequence; the external-most one is ~ ¹ / 3 whorl deep into the shell, and the second one is ~ ¹ / 2 whorl in distance from the first one.</p><p>Description.</p><p>Shell small (shell height = 10.2 mm; shell width = 1.9 mm), multispiral (10 ¹ / 2 whorls), narrow, elongated, nearly cylindrical. Shell colour whitish yellow to light ochre. Protoconch (~ 2 whorls) bulbous, smooth. Teleoconch smooth, except for prosocline growth lines. Whorls increase regularly in height and only slightly in width. Suture well-marked, but not deep. Aperture quadrangular. Peristome simple. Weak columellar fold present on body whorl. Two internal palatal lamellae present on the body and penultimate whorl, spaced ~ ¹ / 2 whorl apart.</p><p>Distribution.</p><p>Known only from the type locality.</p><p>Habitat.</p><p>Found in the twilight zone of the cave, under a rock. The specimen is likely autochthonous, given that the fragility of the shell would not allow extensive transport.</p><p>Etymology.</p><p>Considering that the type locality is a cave in Canaã dos Carajás, the specific epithet honours Arsay, the Canaanite (Ugaritic) goddess of the underworld (an equivalent to the Hurrian Allani and the Mesopotamian Ereshkigal).</p><p>Remarks.</p><p>Only a single shell of this species was found during the study, despite extensive searches. Nevertheless, it is biogeographically highly isolated (both in physical distance and in biome) and exhibits a distinct different morphology from other species in the genus, allowing us to confident assign it to a new species. Rectobelus arsayae sp. nov. can be readily diagnosed from its congeners by its more cylindrical shell, whereas the other species exhibit more conical shell profiles. As the spire of R. arsayae sp. nov. is broader compared e to the more tapering spire of its congeners, its protoconch seems less inflated. Its shell is larger than that of the Peruvian R. silvaevagus comb. nov. (~ 8.5 mm shell height and 9 ¹ / 2 whorls) and much smaller than the Argentine R. birabeni, including the newly synonymized R. mariaisabelae comb. nov. (~ 14.5–16.0 mm shell height and 11–12 whorls). It is of similar size to the Venezuelan R. rectus (~ 9.0– 10.5 mm shell height and 10–11 whorls) and the sinistrally-shelled Brazilian subfossil R. levogyrus (~ 10.5–11.0 mm, shell height and 8–10 whorls).</p><p>The shell of Rectobelus arsayae sp. nov. also exhibits a very faint columellar fold, which extends internally along the entire length of the body whorl, as visible in the tomography images (Fig. 3 C). That feature was previously known only in R. birabeni but, as explained above, this could be a variable character. Furthermore, tomography images revealed the presence of two internal palatal lamellae that appear in sequentially during shell growth (Fig. 3 C). The outermost lamella is ~ ¹ / 3 whorl deep into the shell from the aperture, and the second one is ~ ¹ / 2 whorl in distance from the first. Again, a similar feature has only been reported in R. birabeni, but other species have not been examined for juveniles or using tomography in the adult shell.</p><p>The present record greatly extends the known distribution of the genus in South America and also represents the first report of modern Rectobelus in Brazil ( R. levogyrus is known from archaeological settings only; Simone and D’ávila 2019).</p></div>	https://treatment.plazi.org/id/C201E9A6AF1D561E9426335EF2F585C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo B.;Ferreira-Santos, Laura;Cavallari, Daniel C.;Gallão, Jonas E.;Bichuette, Maria E.	Salvador, Rodrigo B., Ferreira-Santos, Laura, Cavallari, Daniel C., Gallão, Jonas E., Bichuette, Maria E. (2026): Terrestrial gastropods from caves in Serra dos Carajás, eastern Brazilian Amazon, with the description of two new species (Eupulmonata, Stylommatophora). ZooKeys 1287: 213-237, DOI: 10.3897/zookeys.1287.194544
8C1EC4F407DE567EB170FFCBCBD679B7.text	8C1EC4F407DE567EB170FFCBCBD679B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rectobelus Baker 1927	<div><p>Genus Rectobelus Baker, 1927</p><p>Obeliscus (Rectobelus) Baker, 1927: 6 .</p><p>Obeliscus (Microbeliscus) Weyrauch, 1964: 40 [type species, by original designation: Obeliscus (Microbeliscus) silvaevagus Weyrauch, 1964].</p><p>Obeliscus (Nannobeliscus) Weyrauch, 1967: 458 [new name for Obeliscus (Microbeliscus) Weyrauch, 1964 non Sandberger, 1875].</p><p>Type species.</p><p>Obeliscus (Rectobelus) rectus Baker, 1927 [Venezuela, La Fría], by original designation.</p><p>Included species.</p><p>Rectobelus birabeni (Hylton Scott, 1946) [= Nannobeliscus mariaisabelae Miquel &amp; Jaime, 2018, syn. nov.], Rectobelus levogyrus Simone &amp; D’ávila, 2019, Rectobelus rectus Baker, 1927, Rectobelus silvaevagus (Weyrauch, 1964), comb. nov.</p><p>Diagnosis.</p><p>Small shell (~ 10–15 mm), multispiral (8–12 whorls), elongated and very narrow (conico-elongated to nearly cylindrical). Protoconch bulbous. Shell smooth, except for growth lines. Aperture quadrangular, with simple peristome. Weak columellar fold may be present in adults. Internal palatal lamellae may be present.</p><p>Distribution.</p><p>Venezuela (Baker 1927), northern Brazil (Pará state; this study), central Peru (Weyrauch 1964, 1967), and northern Argentina (Jujuy province; Hylton Scott 1946; Miquel and Jaime 2018). It is also known from Rondônia state, western Brazil; however, this record is based exclusively on shells from archaeological contexts (shell mounds dated ca 4500 BP), and it is currently unclear whether the species is still extant (Simone and D’ávila 2019).</p><p>Remarks.</p><p>Rectobelus has been described as a “ section ” of Obeliscus Beck, 1837 and has been since recognised as either its subgenus or synonym (Baker 1927; Zilch 1960), or as a synonym of Ischnocion Pilsbry, 1907 (Hausdorf et al. 2012). Alternatively, it has been considered a genus on its own (Schileyko 1999), a classification recently supported by the revision of Simone and D’ávila (2019). Similarly, Nannobeliscus has been deemed a subgenus of Obeliscus (Weyrauch 1964, 1967), a synonym of Ischnocion together with Rectobelus (Hausdorf et al. 2012), or a separate genus (Schileyko 1999; Miquel and Jaime 2018). The species of both genera are extremely similar in shell shape and size, the known anatomical differences are minimal (see below), and the distributions of both genera are consistent with one another, including a co-occurrence at the same locality in northern Argentina. Thus, herein we reinforce that Nannobeliscus is a synonym of Rectobelus (cf. Hausdorf et al. 2012). That results in the new combination of the type species of Nannobeliscus as Rectobelus silvaevagus (Weyrauch, 1964), comb. nov.</p><p>Rectobelus birabeni and Nannobeliscus mariaisabelae were described from the same province in Argentina and have indistinguishable shells, exhibiting the same shape, size (~ 14.5–16.0 mm), and number of whorls (11–12), as well as having identical radulae (Hylton Scott 1946; Miquel and Jaime 2018; Simone and D’ávila 2019). The only conchological differences reported were the presence of a palatal lamella in some juveniles of the former and a weak columellar fold in adults of the latter (Miquel and Jaime 2018), neither of which is supported here. Firstly, as described by Hylton Scott (1946) based on a series of juvenile and adult specimens, and reinforced here through CT - scan imaging (see below), the palatal lamellae are present at certain stages during the animal’s growth. As the snail grows, the lamellae become internalized in the shell and are, therefore not visible in adults. Secondly, the columellar fold was also reported by Hylton Scott (1946) in some specimens of R. birabeni . As argued by Simone and D’ávila (2019), the columellar fold is variable within many species of South American achatinids and is not particularly reliable for diagnosis by itself. Finally, the subtle anatomical differences noted by Miquel and Jaime (2018) in the size and shape of the penis and spermatheca duct are very tenuous and known to vary intraspecifically, as well as seasonally, in stylommatophorans (Baur and Baur 2017). As such, Nannobeliscus mariaisabelae should be considered a junior synonym of R. birabeni .</p></div>	https://treatment.plazi.org/id/8C1EC4F407DE567EB170FFCBCBD679B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo B.;Ferreira-Santos, Laura;Cavallari, Daniel C.;Gallão, Jonas E.;Bichuette, Maria E.	Salvador, Rodrigo B., Ferreira-Santos, Laura, Cavallari, Daniel C., Gallão, Jonas E., Bichuette, Maria E. (2026): Terrestrial gastropods from caves in Serra dos Carajás, eastern Brazilian Amazon, with the description of two new species (Eupulmonata, Stylommatophora). ZooKeys 1287: 213-237, DOI: 10.3897/zookeys.1287.194544
A35F2D59EF11564FBBDEA9B084EEB5DB.text	A35F2D59EF11564FBBDEA9B084EEB5DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scolodonta carajaensis Salvador & Ferreira-Santos & Cavallari & Gallão & Bichuette 2026	<div><p>Scolodonta carajaensis sp. nov.</p><p>Figs 2 E – F, 4 A – C</p><p>Type material.</p><p>Holotype LES 0031619 . Paratypes LES 0031618 (collected together with holotype) • CMRP 1300 (ex LES 0031640) (Parauapebas, cave N 3_0003; JE Gallão, TS Gallo, DF Torres &amp; JEM Santos leg., 13. viii. 2022) .</p><p>Type locality.</p><p>Brazil, Pará state, Canaã dos Carajás municipality, FLONA de Carajás, Serra Sul, cave S 11 D_0001 (GEM 00650) (JE Gallão, JS Gallo, FD Torres &amp; VF Sperandei leg., 11. ii. 2022).</p><p>Diagnosis.</p><p>Spire elevated; body whorl short, with rounded profile (lacking a marked angulation); aperture D-shaped, rounded; peristome simple.</p><p>Description.</p><p>Shell small (~ 6 mm wide), discoid, with ~ 4 ¹ / 2 whorls; glossy and translucent, tan in colour. Shell smooth except for fine growth lines. Protoconch seemingly has 1 1 / 2 whorl, but transition to teleoconch is indistinct. Spire slightly elevated; spire whorls with a weakly convex profile. Suture well marked, slightly deep. Whorls increase regularly in both width and height. Body whorl profile rounded. Aperture D-shaped, rounded. Insertion of aperture on body whorl just 0.1–0.2 mm over the median region of whorl. Peristome simple, sharp. Umbilicus wide, deep. Shell measurements (holotype): 4 ¹ / 2 whorls, shell height = 3.0 mm, shell width = 6.0 mm.</p><p>Soft body, orange-red (Fig. 2 E, F); some parts (middle of the eyestalks, tip of lower tentacles, border of the foot, edge of the mantle) are slightly paler and less reddish. Head-foot, marked by longitudinal rows of minute and more darkly pigmented spots. Head-foot narrow, with very narrow “ tail ”; eyestalks proportionately long (around the same size as frontal end of head-foot). Mantle and soft parts are partially visible through the shell.</p><p>GenBank acc. nr.</p><p>PZ 576041 (COI), PZ 575549 (16 S) (LES 0031619); PZ 576042 (COI), PZ 575550 (16 S), PZ 575551 (ITS 2 + 28 S) (CMRP 1300).</p><p>Distribution.</p><p>Known from caves in the municipalities of Canaã dos Carajás (type locality) and Parauapebas, in Serra dos Carajás.</p><p>Habitat.</p><p>Found from the entrance to the aphotic zones of the caves. Live individuals were found in very humid microhabitats: under rock, in an area with plenty of organic matter and plant roots (in Canaã dos Carajás), and on unconsolidated substrate close to water drip (in Parauapebas).</p><p>Etymology.</p><p>The specific epithet alludes to the study area, Serra dos Carajás.</p><p>Remarks.</p><p>Scolodonta is primarily identified by its glossy and often translucent shell (Hausdorf 2006), which is slightly sturdier-looking than those of other glossy-shelled scolodontids (e. g., Happia Bourguignat, 1890, Systrophiella Baker, 1925). The lack of shell sculpture (except for fine growth lines) and the weakly elevated spire are also diagnostic of the genus (Hausdorf 2006).</p><p>We are confident in assigning our specimens to a new species, even though two species of “ Scolodonta ” are already known from Pará state. Dohrn (1882) described both Scolodonta nitidula (Dohrn, 1882) and Scolodonta amazonica (Dohrn, 1882) (originally allocated to Ammonoceras Pfeiffer, 1855) from the margins of the Amazon River in Pará. While his descriptions are informative, the original study lacks illustrations. Neither species has been figured since, and the type material is likely lost (Dance 1966). Nevertheless, based on the descriptions and discussion in Dohrn (1882), his S. nitidula is likely synonymous with Tamayoa decolorata (see below), which has a smaller and more rounded shell, as well as a yellow head-foot. On the other hand, S. amazonica is described as having a punctiform and barely visible umbilicus (Dohrn 1882), which is immediately inconsistent with Scolodonta carajaensis sp. nov. Furthermore, S. amazonica has a reflexed peristome around the columella and an even rounder aperture in comparison to S. nitidula / T. decolorata (Dohrn 1882), which is again inconsistent with the new species.</p><p>Scolodonta carajaensis sp. nov. can be readily distinguished from other Brazilian Scolodontidae . Scolodonta bounobaena (d’Orbigny, 1835) is similar in overall shell shape, but it is slightly smaller, with thicker walls; also, it has a stronger angulation in the median-apical region of the body whorl, and a lightly thickened peristome. Scolodonta interrupta (Suter, 1900) is similar in size but has a much more raised spire and a more circular aperture. Furthermore, the periodical interruptions seen in the shell of S. interrupta are more reminiscent of members of the family Streptaxidae, which could indicate an incorrect allocation (Salvador 2021). Scolodonta spirorbis (Deshayes, 1850) is much smaller, with a fragile discoid shell, and very likely does not belong in Scolodonta (Roosen et al., unpub. data).</p><p>The occurrence of this species in more than one cave across the two municipalities suggests that the species is more widespread on the surface across Serra dos Carajás. Such an established epigean population is supported by genetic evidence; there is a 100 % match of both COI and 16 S sequences of the holotype and paratype, one from each municipality, indicating interconnectivity and gene flow. Furthermore, no obvious troglomorphisms were observed; the translucent shell is very common in multiple members of this family (Roosen and Breure 2024 a, 2024 b; Roosen et al. 2025 a, 2025 b) and cannot be considered a troglomorphism per se.</p></div>	https://treatment.plazi.org/id/A35F2D59EF11564FBBDEA9B084EEB5DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo B.;Ferreira-Santos, Laura;Cavallari, Daniel C.;Gallão, Jonas E.;Bichuette, Maria E.	Salvador, Rodrigo B., Ferreira-Santos, Laura, Cavallari, Daniel C., Gallão, Jonas E., Bichuette, Maria E. (2026): Terrestrial gastropods from caves in Serra dos Carajás, eastern Brazilian Amazon, with the description of two new species (Eupulmonata, Stylommatophora). ZooKeys 1287: 213-237, DOI: 10.3897/zookeys.1287.194544
4E758E13E50C56F88A41661DB4CC4A04.text	4E758E13E50C56F88A41661DB4CC4A04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Systrophia siolii Haas 1955	<div><p>Systrophia siolii Haas, 1955</p><p>Fig. 4 D</p><p>Systrophia (Systrophia) siolii Haas, 1955: 103, figs 4–6; Solem 1967: 129.</p><p>Systrophia siolii: Richardson 1989: 137; Salgado and Coelho 2003: 169; Simone 2006: 223, fig. 845; Salvador et al. 2024: 153.</p><p>Material examined.</p><p>LES 0031645 (Canaã dos Carajás, Base ICMBio Gavião Real, Águas Claras; JE Gallão, JS Gallo, DF Torres &amp; JEM Santos leg., viii. 2022) .</p><p>Identification.</p><p>The discoid, smooth shell (~ 4 mm height, ~ 9.5 mm width, ~ 8 whorls), with tightly coiled whorls and the body whorl suddenly expands near the aperture, is consistent with Systrophia siolii, known from elsewhere in Pará state (Simone 2006).</p><p>Previously known distribution.</p><p>Known only from the type locality, Serra Formosa near Mulata, Monte Alegre, Pará (Haas 1955; Simone 2006). This species was found in the woods of the epigean zone near the caves, but was not found inside any caves per se. In any event, the present record extends its range ca 630 km southeast of its previous record in Monte Alegre. Furthermore, the species has also been found in anthropically disturbed areas in the nearby municipality of Parauapebas (Paulo Ricardo S. Coelho, pers. comm. 23 March 2026, species identity confirmed via photographs).</p></div>	https://treatment.plazi.org/id/4E758E13E50C56F88A41661DB4CC4A04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo B.;Ferreira-Santos, Laura;Cavallari, Daniel C.;Gallão, Jonas E.;Bichuette, Maria E.	Salvador, Rodrigo B., Ferreira-Santos, Laura, Cavallari, Daniel C., Gallão, Jonas E., Bichuette, Maria E. (2026): Terrestrial gastropods from caves in Serra dos Carajás, eastern Brazilian Amazon, with the description of two new species (Eupulmonata, Stylommatophora). ZooKeys 1287: 213-237, DOI: 10.3897/zookeys.1287.194544
C27DBC1DCFD95FABAA8B6D673D63A3A1.text	C27DBC1DCFD95FABAA8B6D673D63A3A1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tamayoa decolorata (Drouet 1859)	<div><p>Tamayoa decolorata (Drouët, 1859)</p><p>Figs 2 D, 4 E</p><p>Zonites decoloratus Drouët, 1859: 50, pl. 1, figs 3–5.</p><p>Helix decolorata: Pfeiffer 1868: 169.</p><p>Zonites (Hyalinia) decolorata: Tryon 1886: 166, pl. 52, figs 69–71.</p><p>Scolodonta (Happia) decolorata: Kobelt 1905–1906: 68, pl. 51, figs 14, 15.</p><p>Scolodonta decolorata: Kobelt 1910: 146.</p><p>Happia decolorata: Baker 1925: 23; Richardson 1989: 119.</p><p>Tamayoa (Tamayops) decolorata: Tillier 1980: 106, pl. 5, fig. 1, figs 86–91.</p><p>Tamayoa (Tamayoa) decolorata: Schileyko 2000: 759, fig. 988 A – C.</p><p>Happiella decolorata: Chase and Robinson 2001: 49; Rosenberg and Muratov 2006: 140, tab. 2.</p><p>Tamayoa decolorata: Massemin et al. 2009: 423, text fig., pl. 11, fig. B; Robinson et al. 2009: 640, figs 6 F, 9 B; Delannoye et al. 2015: 282, pl. 62; Charles 2016: 50, pl. 2, fig. 5; Sei et al. 2017: 705, tab. 2; Salvador 2021: 66; González-Guillén and Teruel 2022: 111; Wagner 2022: 24; Georgiev and Mishev 2025: 4.</p><p>Material examined.</p><p>LES 0031616 (Parauapebas, cave N 1_0174; ME Bichuette, JE Gallão &amp; DF Torres leg., 22. viii. 2021) • LES 0031624, LES 0031625 (Canaã dos Carajás, cave SB_0049; JE Gallão, JS Gallo, DF Torres &amp; VF Sperandei leg., 16. ii. 2022) • LES 0031656 (Parauapebas, cave N 1_0172; JE Gallão &amp; DB Ribeiro leg., 05. iv. 2023) • LES 0031630 (Parauapebas, cave N 1_0174; JE Gallão, JS Gallo, DF Torres &amp; VF Sperandei leg., 12. ii. 2022) • LES 0031654 (Parauapebas, cave N 1_0174; JE Gallão &amp; DB Ribeiro leg., 04. iv. 2023) • LES 0031626, LES 0031627 (Parauapebas, cave N 1_0208; JE Gallão, JS Gallo, DF Torres &amp; VF Sperandei leg., 19. ii. 2022) • LES 0031638, LES 0031642 (Parauapebas, cave N 3_0003; JE Gallão, JS Gallo, DF Torres &amp; JEM Santos leg., 13. viii. 2022) • LES 0031617 (Parauapebas, cave N 3_0023; ME Bichuette, JE Gallão &amp; DF Torres leg., 25. viii. 2021) • LES 0031628 (Parauapebas, cave N 3_0026; JE Gallão, JS Gallo, DF Torres &amp; VF Sperandei leg., 18. ii. 2022) • LES 0031631 (Parauapebas, cave S 11 D_0013; JE Gallão, JS Gallo, DF Torres &amp; VF Sperandei leg., 15. ii. 2022) • LES 0031659, LES 0031660 (Canaã dos Carajás, cave SB_0022; JE Gallão &amp; DB Ribeiro leg., 06. iv. 2023) • LES 0031651, LES 0031652 (Canaã dos Carajás, cave SB_0049; JE Gallão &amp; DB Ribeiro leg., 12. iv. 2023) • LES 0031632, LES 0031633, LES 0031634 (Canaã dos Carajás, cave SB_0051; JE Gallão, JS Gallo, DF Torres &amp; VF Sperandei leg., 16. ii. 2022) • LES 0031644 (Canaã dos Carajás, cave SB_0051; JE Gallão, JS Gallo, DF Torres &amp; JEM Santos leg., 17. viii. 2022) • LES 0031649 (Canaã dos Carajás, cave SB_0051; JE Gallão &amp; DB Ribeiro leg., 12. iv. 2023) • LES 0031658 (Canaã dos Carajás, cave SB_0094; JE Gallão &amp; DB Ribeiro leg., 06. iv. 2023) • LES 0031646, LES 0031647 (Canaã dos Carajás, cave ST_0016; JE Gallão &amp; DB Ribeiro leg., 13. iv. 2023) .</p><p>Identification.</p><p>The smooth shell, its size (~ 4 mm height, ~ 7 mm width, ~ 4 whorls), and profile, the slight constriction of the body whorl close to the aperture, and the whitish-yellow body (Fig. 2 D), with a distinct yellow tint on the visible mantle area, all support its identification as Tamayoa decolorata (Massemin et al. 2009; Delannoye et al. 2015). The present specimens could also fit in the description of Scolodonta nitidula (Dohrn, 1882) from the margins of the Amazon River in Pará. However, the original description lacks illustrations, the species has never illustrated since, and the type material is likely lost (Dance 1966). Considering the comparisons made by Dohrn (1882) between his species and Tamayoa decolorata and Hapiella surinamensis (Pfeiffer, 1872), it is very likely that his Scolodonta nitidula is synonymous with Tamayoa decolorata .</p><p>Distribution.</p><p>This species is known from French Guiana and is considered to have been introduced to the Antilles: Barbados, Dominica, Guadeloupe, Jamaica, and Martinique (Delannoye et al. 2015). The present record is the first report of this species from Brazil, although it is not the first time a land snail previously known from French Guiana has been recorded in northern Brazil (Salvador et al. 2018, 2020). A previous record of T. decolorata from southeastern Brazil (Esteves et al. 2025) has been considered erroneous (Salvador et al. 2026).</p><p>Remarks.</p><p>In the studied caves, T. decolorata was found from the entrance zone all to the aphotic zone. Individuals were found under rocks on humid or dry unconsolidated substrate, among roots, or rarely over bat guano (mixed guano of frugivorous and insectivorous bats). This species has also been recorded in anthropically disturbed areas in the nearby municipality of Parauapebas in Pará (Paulo Ricardo S. Coelho, pers. comm. 23 March 2026, species identity confirmed via photographs).</p></div>	https://treatment.plazi.org/id/C27DBC1DCFD95FABAA8B6D673D63A3A1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Salvador, Rodrigo B.;Ferreira-Santos, Laura;Cavallari, Daniel C.;Gallão, Jonas E.;Bichuette, Maria E.	Salvador, Rodrigo B., Ferreira-Santos, Laura, Cavallari, Daniel C., Gallão, Jonas E., Bichuette, Maria E. (2026): Terrestrial gastropods from caves in Serra dos Carajás, eastern Brazilian Amazon, with the description of two new species (Eupulmonata, Stylommatophora). ZooKeys 1287: 213-237, DOI: 10.3897/zookeys.1287.194544
