taxonID	type	description	language	source
03B987DEFFC7FF8CEB4FF98FFDF5C541.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFC7FF8CEB4FF98FFDF5C541.taxon	materials_examined	Material examined MOROCCO – Chtouka Aït Baha Province • 1 ♀; Aït Baha, near Ichengline; 30.11556 ° N, 9.23131 ° W; 324 m a. s. l.; 20 Apr. 2025; N. Ouakri leg.; by hand, Argan grove, under stone in retreat; CSL, MOR _ 1973.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFC5FF82E8A3F910FAA6C181.taxon	description	Fig. 2, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFC5FF82E8A3F910FAA6C181.taxon	materials_examined	Material examined MOROCCO – Chtouka Aït Baha Province • 1 ♀; Aït Baha, near Aït Ben Ali; 30.10399 ° N, 9.21561 ° W; 530 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by hand, Argan grove, under stone; CSL, MOR _ 1707.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCBFF86EB40FCD0FE28C28D.taxon	description	urn: lsid: zoobank. org: act: F 7026 DE 1 - 81 B 6 - 4 D 50 - 9 E 98 - D 230 AAC 4 FE 3 C Figs 3 – 4, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCBFF86EB40FCD0FE28C28D.taxon	diagnosis	Diagnosis The new species resembles Cesonia aspida Chatzaki, 2002 in the conformation of the genitalia, but males can be distinguished by the hook-like RTA. The embolus is long, evenly tapered and curved outwards, consisting of a single branch, tip not bifid and the conductor lacks a sclerotized ventral tip. In C. aspida the RTA is claw-like, the embolus small and curved inwards and the slender side branch has a bifid tip. The conductor has a sclerotized ventral tip (cf. Fig. 3 C – H and Chatzaki et al. 2002: 626, figs 63 – 65). The females of the two species differ in the relative position of the spermathecae. In C. africana Lecigne & Ouakri sp. nov., they are separated by about a third of their diameter, but in C. aspida by about 0.56 × of their diameter. The globular lobes are smaller than the spermathecae in C. africana but in C. aspida they are larger than the spermathecae (cf. Fig. 4 C – F and Chatzaki et al. 2002: 626, figs 66 – 67). Moreover, the lateral heads are larger in the new species. Finally, the two species can also be distinguished by the shape of the dark median band on the abdomen, which in C. africana Lecigne & Ouakri sp. nov. tapers towards the spinnerets but is expanded posteriorly in C. aspida and the two light dorsal strips which are not connected at the posterior end whereas connected at the posterior end in C. aspida (cf. Fig. 3 A and Chatzaki et al. 2002: 627, fig. 68).	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCBFF86EB40FCD0FE28C28D.taxon	etymology	Etymology We name these specimens of both sexes as the first representative of the genus Cesonia in Africa.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCBFF86EB40FCD0FE28C28D.taxon	materials_examined	Type material Holotype MOROCCO – Agadir Ida-Outanane Prefecture • ♂ (left pedipalp detached); Agadir, Anza; 30.47808 ° N, 9.66300 ° W; 80 m a. s. l.; 22 Mar. 2025; S. Lecigne and N. Ouakri leg.; sparse Argan grove and Euphorbia L., on arid rocky ground, under stones; by hand; SMF, MOR _ 1677. Paratypes MOROCCO – Agadir Ida-Outanane Prefecture • 1 ♂ (abdomen and epigyne separated, abdomen lost); same data as for holotype; SMF, MOR _ 1677 • 1 ♀; same data as for holotype; 5 Jun. 2025; N. Ouakri leg.; SMF, MOR _ 1963.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCBFF86EB40FCD0FE28C28D.taxon	description	Description The specimens feature the characters listed by Chatzaki (2002: 626), i. e., dark longitudinal bands covering the entire length of the abdomen dorsally and laterally, separated by intervening light bands (see Fig. 3 A compared with Chatzaki 2002: 627, fig. 68); PME closer to PLE than to each other; embolus close to the membranous conductor, protruding beyond the tegulum. We therefore assign these specimens to the genus Cesonia. Male (holotype; Fig. 3) MEASUREMENTS. Total length 6.35; carapace length 2.83, width 2.35, CL / CW 1.20; scutum length 0.9. COLOUR OF LIVE SPECIMEN (Fig. 3 A). Carapace dorsally with three light and two dark longitudinal strips, the light median band joining the two light marginal bands posteriorly. Legs dark brown, covered with white hairs. Abdomen with two light dorsal strips not connected at posterior end, one median and two lateral longitudinal dark strips, one on each side. COLOUR IN ETHANOL (Fig. 3 B). Chelicerae brown, distal part paler. Sternum dark brown to brown. Legs dark brown except sides of Fe I, yellowish and all metatarsi and tarsi paler. Abdomen ventrally pale brown. CARAPACE. PME closer to PLE than to each other. Chelicerae: outer cheliceral margin armed with three small teeth, increasing in size distally. LEG SPINATION. Tibiae I – II, 3 pairs of ventral spines; metatarsi I – II, 1 pair of ventral, proximal spines, other segments with variable number of spines, more numerous on legs III – IV. ABDOMEN. Anterior scutum occupying 27 – 28 % of its length. PALP (Fig. 3 C – H). Tibial apophysis short, hook-like, supported by rounded protrusion (Fig. 3 G, RTA). Subtegulum striated on its prolateral side, ending behind conductor in broad flat process, curved and pointed (Fig. 3 H, St). Embolus fairly long, evenly tapered, slightly curved outwards, extending beyond edge of cymbium, at about 12 o’clock position. Conductor, membranous projection close to embolus. Female (paratype; Fig. 4); as in male except as noted MEASUREMENTS. Total length 6.33; carapace length 2.65, width 2.18, CL / CW 1.21. COLOUR IN ETHANOL (Fig. 4 B). Sternum brown, central area paler; chelicerae pale brown. EPIGYNE / VULVA (Fig. 4 C – F). Inner margins sclerotized, S-shaped (Fig. 4 F, dotted arrows). Median part forming a translucent V-shaped plate reaching lateral margins (Fig. 4 F, black arrow). Copulatory openings located laterally. Spermathecae rounded, separated by about ⅓ of their diameter, surmounted by two lateral heads and then by two globular lobes in contact with each other, inclined towards each other (Fig. 4 G, L). Variation MEASUREMENTS. Male (n = 2): total length 5.40 – 6.35, carapace length 2.53 – 2.83, width 2.05 – 2.35, CL / CW 1.20 – 1.23; scutum length 0.80 – 0.90.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCFFF86EB62FDE7FAB8C659.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCFFF86EB62FDE7FAB8C659.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 13 Mar. 2022; N. Ouakri leg.; by hand, Argan grove and Euphorbia, under stone in retreat; CSL, MOR _ 1546 • 1 ♀; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 21 Jan. 2024; N. Ouakri leg.; by hand, Argan grove, palm trees and olive trees, under stone; CSL, MOR _ 1500 • 1 ♀; same data as for preceding; 18 Feb. 2024; N. Ouakri leg.; by hand, Argan grove, on ground; CSL, MOR _ 1503. – Chtouka Aït Baha Province • 1 ♂; Aït Baha, near Aït Ben Ali; 30.10399 ° N, 9.21561 ° W; 530 m a. s. l.; 23 Oct. 2022; N. Ouakri leg.; by hand, Argan grove, under stone; CSL, MOR _ 1518 • 1 ♀; same data as for preceding; 15 Mar. 2024; N. Ouakri leg.; by hand, Argan grove, under stone with its egg sac in retreat; CSL, MOR _ 1527 • 1 ♀; same data as for preceding; 19 Mar. 2025; S. Lecigne leg.; by hand, Argan grove, under stone; CSL, MOR _ 1787 • 1 ♀; Aït Baha, near Dam Ahl Souss; 30.06279 ° N, 9.12406 ° W; 619 m a. s. l.; 11 Jan. 2022; N. Ouakri leg.; by hand, Argan grove, under stone; CSL, MOR _ 1519 • 1 ♀; same data as for preceding; 4 Feb. 2024; N. Ouakri leg.; by hand, Argan grove, under stone with its egg sac in retreat; CSL, MOR _ 1499.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCFFF87EB0AFA53FEDEC253.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCFFF87EB0AFA53FEDEC253.taxon	materials_examined	Material examined MOROCCO – Chtouka Aït Baha Province • 1 ♀; Aït Baha, near Ichengline; 30.11556 ° N, 9.23131 ° W; 324 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by hand, Argan grove, under stone; CSL, MOR _ 1704.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCEFF87E8ABFE49FAFAC737.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCEFF87E8ABFE49FAFAC737.taxon	materials_examined	Material examined MOROCCO – Chtouka Aït Baha Province • 1 ♂; Aït Baha, near Aït Ben Ali; 30.10399 ° N, 9.21561 ° W; 530 m a. s. l.; 11 Dec. 2022; N. Ouakri leg.; by hand, Argan grove, on ground; CSL, MOR _ 1510 • 1 ♀; same data as for preceding; CSL, MOR _ 1564 • 1 ♂; same data as for preceding; CSL, MOR _ 1753 • 1 ♂; same data as for preceding; CSL, MOR _ 1757 • 1 ♀; same data as for preceding; CSL, MOR _ 1971.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCEFF87EB19FB6CFD5FC411.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCEFF87EB19FB6CFD5FC411.taxon	materials_examined	Material examined MOROCCO – Chtouka Aït Baha Province • 1 ♀; Aït Baha, near Aït Ben Ali; 30.10399 ° N, 9.21561 ° W; 530 m a. s. l.; 11 Sep. 2022; N. Ouakri leg.; by hand, Argan grove, under stone; CSL, MOR _ 1520 • 1 ♀; same data as for preceding; MOR _ 1559 • 1 ♀; same data as for preceding; CSL, MOR _ 1562 • 1 ♂; same data as for preceding; CSL, MOR _ 1542 • 1 ♀; same data as for preceding.; 23 Oct. 2022; N. Ouakri leg.; by hand, Argan grove, under stone; CSL, MOR _ 1509.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCDFF84EB1AFEADFBB5C1E5.taxon	description	Fig. 5, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCDFF84EB1AFEADFBB5C1E5.taxon	materials_examined	Material examined MOROCCO – Chtouka Aït Baha Province • 1 ♀; Aït Baha, near Aït Ben Ali; 30.10399 ° N, 9.21561 ° W; 530 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by hand, Argan grove, under stone; CSL, MOR _ 1713.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCDFF84EB1AFEADFBB5C1E5.taxon	distribution	Distribution So far, Scotophaeus validus (Lucas, 1846) is known from southern Europe and northern Africa including Morocco, Algeria, Tunisia and Libya (Nentwig et al. 2025; WSC 2025).	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCCFF85EB0FFEADFC00C04C.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCCFF85EB0FFEADFC00C04C.taxon	materials_examined	Material examined MOROCCO – Chtouka Aït Baha Province • 1 ♀; Aït Baha, near Aït Ben Ali; 30.10399 ° N, 9.21561 ° W; 530 m a. s. l.; 25 Dec. 2022; N. Ouakri leg.; by hand, Argan grove, on ground; CSL, MOR _ 1962.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCCFF85EB67FC26FDAFC6C5.taxon	description	Fig. 6, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCCFF85EB67FC26FDAFC6C5.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Alma; 30.51117 ° N, 9.59297 ° W; 410 m a. s. l.; 2 Jul. 2022; N. Ouakri leg.; by hand, Argan grove and olive trees, under stone; CSL, MOR _ 1744. – Chtouka Aït Baha Province • 1 ♂; Aït Baha, near Aït Ben Ali; 30.10399 ° N, 9.21561 ° W; 530 m a. s. l.; 26 May. 2024; N. Ouakri leg.; by hand, Argan grove, under stone, in retreat; CSL, MOR _ 1525.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCCFF99EB50F9E9FB06C282.taxon	description	Fig. 7, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCCFF99EB50F9E9FB06C282.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 19 Mar. 2023; N. Ouakri leg.; by hand, Argan grove, palm trees and olive trees, under stone; CSL, MOR _ 1531 • 1 ♂; Tinfoul; 30.51986 ° N, 9.35192 W; 337 m a. s. l.; 21 Jan. 2024; N. Ouakri leg.; by hand, Argan grove, palm trees and olive trees, on ground; CSL, MOR _ 1496. – Chtouka Aït Baha Province • 1 ♀; Aït Baha, near Ichengline; 30.11556 ° N, 9.23131 ° W; 324 m a. s. l; 11 Dec. 2022; N. Ouakri leg.; by hand, Argan grove, under stone; CSL, MOR _ 1517. Other material examined Zelotes oryx (Simon 1880) ALGERIA? – Biskra? • 1 ♂, 1 ♀; (remark: without specified locality, probably the type series from Algeria, Biskra (Bosmans, 1995, pers. comm.), examination based on photos); repos. ident., MNHNP AR 1992.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFCCFF99EB50F9E9FB06C282.taxon	discussion	Remarks The original description of Zelotes oryx (Simon, 1880) is very brief and provides no useful details on the conformation of the bulbus nor any illustrations. Denis (1952) does not provide any further details, but does produce two figures based on specimens from the MNHNP. It is not clear whether these specimens correspond to those in Simon’s collection. However, his figures (Denis 1952: 121, figs 13 – 15, partly reproduced here in Fig. 7 E, J )) are consistent with the male (MNHNP AR 1992) whose photos are presented (Fig. 7 D, K). The original description of Zelotes tragicus (O. Pickard-Cambridge 1872) focuses mainly on somatic characters. Regarding the palpus, only the tibial apophysis is described, but the drawing shows only the ventral view and remains imprecise. However, the descriptions of the tibial apophyses of both species are convergent: regarding Z. oryx “ … apophyse (vue de profil) presque aussi longue que l’article, très-épaisse, non atténuée, légèrement dirigée en bas, tronquée très obliquement avec l’angle supérieur prolongé en pointe aiguë relevée en haut et un peu infléchie en dehors ” (Simon 1880) [“ ... the apophysis (viewed in profile) is almost as long as the article, very thick, not attenuated, slightly directed downwards, truncated very obliquely with the upper angle prolonged into a sharp point raised at the top and slightly bent outwards ”]; regarding Z. tragicus ‘ … this is broad and strong at its commencement, but at about the middle of its length from the underside it goes off abruptly into a tapering sharp-pointed form on the upper-side, with a rather upward direction’ (O. Pickard-Cambridge 1872). Specimens of both sexes sampled during the survey in the vicinity of Agadir (Tinfoul and Aït Baha) were compared with what we consider to be the type series of Zelotes oryx. Direct examination of the type series was not possible (loan and examination not permitted); however, we were able to study photographs of the specimens (Fig. 7 B – D, K). They clearly match the figures of Zelotes tragicus according to Alioua & Bosmans (2024) and Levy (1998) (Fig. 7 I), specifically the shape of the tibial apophysis, broad and robust (Fig. 7 D – F, RTA) and the spiral embolus (Fig. 7 H – K, see E). Thus, we consider Zelotes oryx as a junior synonym of Zelotes tragicus. The epigyne of the female illustrated by Marinaro (1968) belongs to another, unknown species.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD0FF99E8A9FDDDFE6AC79A.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD0FF99E8A9FDDDFE6AC79A.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♂; Alma; 30.51117 ° N, 9.59297 ° W; 410 m a. s. l.; 2 Jul. 2022; N. Ouakri leg.; by hand, Argan grove and olive trees, under stone; CSL, MOR _ 1556 • 1 ♂; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 14 Aug. 2022; N. Ouakri leg.; by hand, Argan grove and Euphorbia; under stone; CSL, MOR _ 1513. – Chtouka Aït Baha Province • 1 ♂, 2 juvs; Aït Baha, near Ichengline; 30.11556 ° N, 9.23131 ° W; 324 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by hand, Argan grove, under stone; CSL, MOR _ 1705.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD0FF9EEB7AFAD6FC1DC2AF.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD0FF9EEB7AFAD6FC1DC2AF.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 5 ♀♀, 1 ♂; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 22 Mar. 2025; S. Lecigne leg.; by hand, sparse Argan grove on arid rocky soil, under stone; CSL, MOR _ 1678 • 2 ♀♀; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; Argan grove, palm trees and olive trees, under stone; CSL, MOR _ 1721 • 1 ♀; same data as for preceding; CSL, MOR _ 1774 • 1 ♀; same data as for preceding; N. Ouakri leg.; Argan grove, palm trees and olive trees, under stone; CSL, MOR _ 1977.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD7FF9FEB65FB10FEF3C25C.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD7FF9FEB65FB10FEF3C25C.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♂; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 13 Nov. 2022; N. Ouakri leg.; by hand, Argan grove, palm trees and olive trees, on the ground; CSL, MOR _ 1552 • 1 ♀; same data as for preceding; 21 Jan. 2024; N. Ouakri leg.; by hand, Argan grove and olive trees, on the ground; CSL, MOR _ 1498 • 1 ♀; same data as for preceding; under stone; CSL, MOR _ 1504.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD7FF9EE8BAFDC4FE50C740.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD7FF9EE8BAFDC4FE50C740.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 5 ♀♀; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 22 Mar. 2025; S. Lecigne leg.; by hand, sparse Argan grove on arid rocky soil, under stone; CSL, MOR _ 1686 • 2 ♀♀; same data as for preceding; 1 Apr. 2025; N. Ouakri leg.; by hand, Argan grove and Euphorbia, under stone; CSL, MOR _ 1969.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD6FF9FEB0BFE0BFD8EC7E3.taxon	description	Fig. 8, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD6FF9FEB0BFE0BFD8EC7E3.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♂; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 4 Dec. 2022; N. Ouakri leg.; by hand, Argan grove and Euphorbia, in a burrow under stone; CSL, MOR _ 1537.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD6FF9FEB0BFE0BFD8EC7E3.taxon	discussion	Remarks Regarding the identification of the male specimen from Anza: in addition to the two keels i. e., superior prolateral and apical, we note the presence of the large tooth (an angular process = third keel) in the subapical part of the embolus in N. uncinata which does not occur in other members of the BE group, defined by Decae (2012) (after Zonstein 2017: 155 ‒ 156). The species is new to Morocco.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD6FF9CEB65FA7FFB30C41F.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD6FF9CEB65FA7FFB30C41F.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 28 May 2025; N. Ouakri leg.; by hand, Argan grove and Euphorbia, under stone; CSL, MOR _ 1968 • 1 ♀, 1 ♂; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by hand, Argan grove, palm trees and olive trees, under stone; CSL, MOR _ 1724.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD6FF9CEB65FA7FFB30C41F.taxon	discussion	Remarks Oecobius machadoi was recently reported for the first time from Morocco, i. e., Marrakech (Lecigne et al. 2025). Its range extends southward with the present record in Agadir Ida Outanane.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD4FF9DEB02FC3FFD90C559.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD4FF9DEB02FC3FFD90C559.taxon	materials_examined	Material examined MOROCCO – Chtouka Aït Baha Province • 1 ♂; Aït Baha, near Ichengline; 30.11556 ° N, 9.23131 ° W; 324 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by hand, Argan grove, under stone; CSL, MOR _ 1706.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD4FF9DEB19FE8AFACBC023.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD4FF9DEB19FE8AFACBC023.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 22 Mar. 2025; S. Lecigne leg.; by hand, sparse Argan grove on arid rocky soil, between stones, on web, with cocoon; CSL, MOR _ 1689.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD4FF92EB31F96EFE2CC562.taxon	description	Fig. 9, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFD4FF92EB31F96EFE2CC562.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♂; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 28 May 2025; N. Ouakri leg.; by hand, Argan grove and Euphorbia, on ground; CSL, MOR _ 1975 • 1 ♂; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 12 May 2024; N. Ouakri leg.; by hand, Argan grove, palm trees and olive trees, under stone; CSL, MOR _ 1767 • 2 ♂♂ same data as for preceding; 19 Mar. 2025; S. Lecigne leg.; by hand Argan grove, palm trees and olive trees, on ground and under stone; CSL, MOR _ 1727.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDBFF93EB1EF938FC3AC144.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDBFF93EB1EF938FC3AC144.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 3 ♂♂, 1 juv; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 22 Mar. 2025; S. Lecigne leg.; by hand, sparse Argan grove on arid rocky soil, under stone; CSL, MOR _ 1694.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDAFF96EB6FFD5DFC8BC611.taxon	description	urn: lsid: zoobank. org: act: 6 AA 3 C 964 - 3 A 75 - 4 D 08 - 9492 - 568 C 002 D 4 E 57 Figs 10 – 11, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDAFF96EB6FFD5DFC8BC611.taxon	diagnosis	Diagnosis The new species most resembles Menemerus davidi Prószyński & Wesołowska, 1999 but the male differs in the shape of the palpal retrolateral lobe, large and not bilobed in M. currens Lecigne sp. nov. The palpal ventral tibial apophysis is bent retrolaterally in the new species and pointed ventrally in M. davidi. Menemerus currens lacks both palpal dorsal tibial apophysis and basal femoral process; M. davidi has a flat dorsal tibial apophysis and a big process at base of femur (cf. Fig. 10 B – F and Wesołowska 1999: 280, figs 78 – 85). The female of the new species can be distinguished by the presence of two large posterolateral epigynal teeth, absent in those of M. davidi. The spermathecae of M. currens are higher than wide, subcircular in M. davidi (cf. Fig. 11 D – G; Wesołowska 1999: 281, figs 86 – 88).	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDAFF96EB6FFD5DFC8BC611.taxon	etymology	Etymology The specific epithet of this new species refers to the behaviour of the specimens observed in their environment, fleeing by running quickly on the ground rather than jumping.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDAFF96EB6FFD5DFC8BC611.taxon	materials_examined	Type material Holotype MOROCCO – Agadir Ida-Outanane Prefecture • ♂ (left pedipalp detached); Agadir, Anza; 30.47808 ° N, 9.66300 ° W; 80 m a. s. l.; 22 Mar. 2025; S. Lecigne leg.; sparse Argan grove and Euphorbia, on arid rocky ground, under stone; by hand; SMF, MOR _ 1684. Paratype MOROCCO – Agadir Ida-Outanane Prefecture • 1 ♀ (abdomen and epigyne separated; abdomen damaged); same data as for holotype; SMF, MOR _ 1684.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDAFF96EB6FFD5DFC8BC611.taxon	description	Description Male (holotype; Fig. 10) MEASUREMENTS. Total length 6.80; carapace length 3.00, width 2.29, CL / CW 1.31; PLE – PLE 1.50. COLOUR IN ETHANOL (Fig. 10 A). Carapace dark brown with broad median band of long white flattened hairs, widening anteriorly to anterior eye line, edged with orange hairs. Marginal bands of white hairs narrowing anteriorly and interrupted at level of PLE. Clypeus with white hairs, thicker under AME; anterior eyes surrounded by long orange setae. Sternum pale brown. Legs yellowish except coxae, trochanters and femora of first legs brown but femora distally paler; tibiae and metatarsi of first legs darker; retrolateral side of femora of second pair of legs with large greyish marks. Pedipalps brown, femora dorsally covered with long white hairs, denser distally. Abdomen yellowish with two lateral wide longitudinal dark strips, converging at rear towards spinnerets, but without joining. CARAPACE. Chelicerae: outer cheliceral margin armed with two teeth, distal one largest. LEG SPINATION. Tibiae II and metatarsi I – II, with 2 pairs of ventral spines; other segments with variable number of spines. PALP (Fig. 10 B – G). Ventral apophysis directed outwards, tip slightly curved ventrally (Fig. 10 E). Retrolateral side of palpal femur with sub-circular depression and ventro-apical area of femur pointed and sclerotized (Fig. 10 G). Ventro-retrolateral edge of tegulum straight. Embolus short and stout, strongly sclerotized, slightly and evenly curved outwards, oriented towards 1 o’clock position (Fig. 10 C, F). Female (paratype; Fig. 11); as in male except as noted MEASUREMENTS. Total length 9.20; carapace length 3.05, width 2.30, CL / CW 1.33. COLOUR ON LIVE SPECIMEN (Fig. 11 A). Silvery appearance of broad median white band on carapace and abdomen, contrasting with dark brown lateral stripes. COLOUR IN ETHANOL (Fig. 11 B – C). Legs yellowish with greyish areas giving vaguely ringed appearance, particularly on tibiae and metatarsi of third and fourth legs. Pedipalps yellow, tibiae and tarsi with long, sparse white hairs, mainly on sides. LEG SPINATION. Metatarsi I – II, 2 pairs of ventral spines; other segments with variable number of spines. EPIGYNE / VULVA (Fig. 11 D – G). Epigyne wide, with large atria, provided with two large blunt posterolateral teeth (Fig. 11 F, ET). Internal structure strongly sclerotized (Fig. 11 E); spermathecae (Sp) higher than wide (Fig. 11 G).	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDFFF97EB47FA8AFB8DC076.taxon	description	Fig. 12, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDFFF97EB47FA8AFB8DC076.taxon	materials_examined	Type material Holotype NIGER – Sahara Gangana • ♀ (examination based on photos); MNHNP, jar 2662 (” X. 37; rec. Berland, det. J. Millot ”). Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by hand, Argan grove, palm trees and olive trees, under stone; CSL, MOR _ 1730.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDFFF97EB47FA8AFB8DC076.taxon	discussion	Remarks The specimen found in Tinfoul corresponds to the original description (Berland & Millot 1941). However, the lateral edges of the epigyne of our specimen (Fig. 12 C vs Berland & Millot 1941: 307, fig. 8 c) do not show any apparent convergence, as is the case for the type specimen. The description by these authors does not detail the internal genital structures. The photos of the vulva of the Egyptian specimen (El-Helbawy et al. 2025: fig. 3 b – c) do not provide sufficient detail of the internal structures to provide certainty. Nevertheless, we were able to compare our specimen both with another individual from Morocco featuring the same convex shape of the lateral margins of the epigyne as the type drawn by Berland & Millot (1941), and with the holotype itself (see supra material examined). Our two specimens from Morocco display the same conformation of the vulva. Consequently, we assign the specimen from Tinfoul to Mogrus sahariensis.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDDFF95EB04FEADFDE5C1DE.taxon	description	Fig. 13, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDDFF95EB04FEADFDE5C1DE.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 19 Mar. 2023; N. Ouakri leg.; by hand, Argan grove, palm trees and olive trees, under stone; CSL, MOR _ 1758 • 1 ♀; same data as for preceding; 18 Feb. 2024; N. Ouakri leg.; by hand, Argan grove, palm trees and olive trees, on ground; CSL, MOR _ 1523 • 1 ♀; same data as for preceding; 23 Jun. 2024; N. Ouakri. Leg.; Argan grove and olive trees, under stone in retreat; CSL, MOR _ 1522 • 1 ♀; same data as for preceding; 19 Mar. 2025; S. Lecigne leg.; Argan grove, palm trees and olive trees, under stone; CSL, MOR _ 1726.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDCFF95EB06FCD4FC5BC7EA.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDCFF95EB06FCD4FC5BC7EA.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by beating thorny bushes in Argan grove; CSL, MOR _ 1733.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDCFF95EB09FA80FED7C4F3.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♂ sub.; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by beating thorny bushes in Argan grove; CSL, MOR _ 1732.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFDCFF95EB09FA80FED7C4F3.taxon	discussion	Remark Its discovery in Tinfoul Argan grove constitutes the second citation of the species in Morocco since Simon (1909).	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE3FFAAEB4FFE8AFB9BC711.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Aghroud; 30.62920 ° N, 9.82606 ° W; 24 m a. s. l.; 3 Mar. 2024; N. Ouakri leg.; by hand, Argan grove and Euphorbia, under stone; CSL, MOR _ 1506 • 1 ♀, 1 ♂; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 22 Mar. 2025; S. Lecigne leg.; by hand, sparse Argan grove on arid rocky soil, under stone; CSL, MOR _ 1693. – Chtouka Aït Baha Province • 2 ♀♀; Aït Baha, near Aït Ben Ali; 30.10399 ° N, 9.21561 ° W; 530 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by hand; Argan grove, under stone; CSL, MOR _ 1710 • 1 ♀; Same data as for preceding; CSL, MOR _ 1785.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE3FFAAEB4FFE8AFB9BC711.taxon	distribution	Distribution and habitat Scytodes velutina Heineken & Lowe, 1832 is a Mediterranean species and even reaching the Cape Verde Islands. For the distribution and habitat of the species, see Le Peru (2011).	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE3FFAAEB03FB40FABDC5E9.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE3FFAAEB03FB40FABDC5E9.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 juv.; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 1 Apr. 2025; N. Ouakri leg.; by hand, Argan grove, palm trees and olive trees, under stone in retreat; CSL, MOR _ 1964. – Chtouka Aït Baha Province • 1 juv.; Aït Baha, near Ichengline; 30.11556 ° N, 9.23131 ° W; 324 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by hand, Argan grove, under stone; CSL, MOR _ 1700 • 1 ♀; same data as for preceding; 10 May 2025; N. Ouakri leg.; by hand, Argan grove, under stone; CSL, MOR _ 1966.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE2FFABEB35FE8AFB8FC0B5.taxon	description	Fig. 14, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE2FFABEB35FE8AFB8FC0B5.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♂; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 13 Mar. 2022; N. Ouakri leg.; by hand, Argan grove and Euphorbia, under stone in a silk retreat; CSL, MOR _ 1563.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE2FFABEB35FC3AFD3EC4B2.taxon	description	Figs 14 E – 15, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE2FFABEB35FC3AFD3EC4B2.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 22 Mar. 2025; S. Lecigne leg.; grove with sparse Argan, on arid rocky soil, by hand, under stone; CSL, MOR _ 1679 • 1 ♀; same data as for preceding; 28 May 2025; N. Ouakri leg.; by hand, Argan grove and Euphorbia, under stone in its retreat; CSL, MOR _ 1965. – Chtouka Aït Baha Province • 1 ♀; Aït Baha, near Aït Ben Ali; 30.10399 ° N, 9.21561 ° W; 530 m a. s. l.; 23 Oct. 2022; N. Ouakri leg.; by hand, Argan grove, under stone in its retreat; CSL, MOR _ 1549 • 1 ♀; Aït Baha, near Dam Ahl Souss; 30.06279 ° N, 9.12406 ° W; 619 m a. s. l.; 11 Jan. 2022; N. Ouakri leg.; by hand, Argan grove, under stone in a burrow; CSL, MOR _ 1548.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE7FFAEEB68FEACFE68C0AB.taxon	description	Fig. 16 A, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE7FFAEEB68FEACFE68C0AB.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Aghroud; 30.62920 ° N, 9.82606 ° W; 24 m a. s. l.; 19 May 2024; N. Ouakri leg.; by hand, Argan grove and Euphorbia, under stone, in its retreat; CSL, MOR _ 1768.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE7FFAFEB29FBDAFDA7C117.taxon	description	Fig. 16 B – C, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE7FFAFEB29FBDAFDA7C117.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Azrarag; 30.46878 ° N, 9.46733 ° W; 231 m a. s. l.; 6 Feb. 2022; N. Ouakri leg.; by hand, Argan grove, under stone in a silk retreat; CSL, MOR _ 1521.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE6FFACEB79FD44FC78C13E.taxon	description	Fig. 17, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE6FFACEB79FD44FC78C13E.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♂; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 13 Mar. 2022; N. Ouakri leg.; by hand, Argan grove and Euphorbia, in a burrow under stone; CSL, MOR _ 1742.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE5FFACEB7CFD2AFD40C7C9.taxon	description	Fig. 18, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE5FFACEB7CFD2AFD40C7C9.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 2 ♀♀; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by hand, Argan grove, palm trees and olive trees, under stone; CSL, MOR _ 1780 • 1 ♀; same data as for preceding; 10 Apr. 2025; N. Ouakri leg.; by hand, Argan grove, palm trees and olive trees, under stone; CSL, MOR _ 1980.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE5FFACEB71FAA3FEF3C480.taxon	description	Fig. 19, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE5FFACEB71FAA3FEF3C480.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by beating, thorny bushes in Argan grove; CSL, MOR _ 1735.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE4FFA2EB0FF9F6FACEC5B2.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFE4FFA2EB0FF9F6FACEC5B2.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 5 ♀♀, 2 ♂♂, 2 juvs; Anza; 30.47800 ° N, 9.66281 ° W; 41 m a. s. l.; 22 Mar. 2025; S. Lecigne leg.; by hand, Sparse Argan grove and Euphorbia, on arid rocky soil, under stone; CSL, MOR _ 1688 • 1 ♀, Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by hand, Argan grove, palm trees and olive trees, under stone; CSL, MOR _ 1720.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFEBFFA3EB14F902FE74C181.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFEBFFA3EB14F902FE74C181.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♀; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 1 Apr. 2025; N. Ouakri leg.; by hand, Argan grove, palm trees and olive trees, in litter under stone; CSL, MOR _ 1974. – Chtouka Aït Baha Province • 1 ♀, 1 ♂; Aït Baha, near Aït Ben Ali; 30.10399 ° N, 9.21561 ° W; 530 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by hand, Argan grove, on arid rocky ground, under stone; CSL, MOR _ 1782.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFEAFFA3EB66FCD1FC40C69D.taxon	description	Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFEAFFA3EB66FCD1FC40C69D.taxon	materials_examined	Material examined MOROCCO – Chtouka Aït Baha Province • 1 ♂; Aït Baha, near Ichengline; 30.11556 ° N, 9.23131 ° W; 324 m a. s. l.; 23 Oct. 2022; N. Ouakri leg.; by hand, Argan grove, in litter under stone; CSL, MOR _ 1511.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFEAFFA0EB1FF9CEFCDDC191.taxon	description	Fig. 20, Table 2	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
03B987DEFFEAFFA0EB1FF9CEFCDDC191.taxon	materials_examined	Material examined MOROCCO – Agadir Ida Outanane Prefecture • 1 ♂; Tinfoul; 30.51986 ° N, 9.35192 ° W; 337 m a. s. l.; 19 Mar. 2025; S. Lecigne leg.; by beating, thorny bushes in Argan grove; CSL, MOR _ 1734.	en	Ouakri, Najat, Lecigne, Sylvain, Oualid, Jaouad Abou, Dugon, Michel (2026): Spiders from the Arganeraie Biosphere Reserve of Souss-Massa region (Morocco), and descriptions of two new species (Arachnida: Araneae: Gnaphosidae, Salticidae). European Journal of Taxonomy 1051: 1-50, DOI: 10.5852/ejt.2026.1051.3261, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3261/14417
